A'merican Diptrkera

A'MERICAN DIPTrkERA

J3Y C. IL CURRAN

BULLETIN OF T'HEE AMERIC,AN,"MUSEUM OF NATURAL' HISTORY -VOL. LXXX, ART. II1, pp. 51-41 New York Issued Septemnber 28, 1942 Article IIL.-AMERICAN DIPTERA

BY C. H. CURRAN

In addition to the descriptions of a num- 10.-Posterior femora with conspicuous ber of new species there are notes on certain pile.femoratus Karsch. forms and keys to most of the genera in Posterior femora without conspicu- which species are described. Most of the ous pile.. .fumipennis Westwood. new forms have come to light during the 11.-Posterior tibiae black apically. . .. 12. study of material received for identifica- Posterior tibiae with the apical third tion. Unless otherwise noted the types yellow.... lugubris Osten Sacken. are in The American Museum of Natural 12.-Abdomen brownish red, the base History. brown above...... columbianus Karsch. Bombyliidae Abdomen black on apical third or more.13. SYSTROPUS WIEDEMANN 13.-Posterior tibiae wholly black..... 14. KEY TO NEOTROPICAL SPECIES Posterior tibiae whitish at the base...... foenoidesWestwood. 1.-Anterior fen;ora all yellow .....2. 14.-A yellow spot on the pleura in front of Anterior femora partly black or the root of the wings.. brownish...... 9. ....dolorosus Wiedemann. 2.-Posterior tarsi black basally.....3. Pleura yellow only above the front Posterior tarsi yellow basally...... 5. coxae. 15. 3.-Posterior legs wholly black...... 15.-Fifth abdominal segment wholly red- ...... nitidus Wiedemann. dish. mars, n. sp. Posterior legs not wholly black....4. Fifth abdominal segment black on its

4.-Four small roundish black spots be- whole length...... 16. tween the hind coxae and base of 16.-Tip of apical cell about one-third its abdomen. greatest width.... geijskesi, n. sp. ... 9 quadrimaculatus Williston. Tip of apical cell more than one-

Two large and two small black spots half its greatest width ...... 17. on this area...... 17.-Wings yellowish brown from base to ...... e cerdo Osten Sacken. apical crossvein, the discal and 5.-Mesonotum black...... 6. anal cells hyaline......

Mesonotum dark red...... dimidiatus,n. sp...... pulcherWilliston. Wings tinged with brown, the costal 6.-Posterior femora broadly yellow api- border darker. .willistoni, n. sp. cally...... 7. Posterior femora black apically.... Systropus mars, new species ...... rufiventris Osten Sacken. Black, the thorax with yellow markings, the 7.-Antennae yellow ...... sallei Costa. abdomen with the fifth segment wholly reddish. Antennae black...... 8. Length, 16 mm. 8.-Pleura blackish...... FEMALE.-Sides of oral margin reddish yellow, wholly the antennal prominence orange above; face and similis Williston. frontal triangle sericeous, the occiput cinereous. Pleura with yellow and red mark- Antennae black, the first segment about as long ings. rogersi Osten Sacken. as the head-height, a little shorter than the apical two, the third about twice as long as the second. 9.-Wings clouded with brown before the Thorax black, the metasternum brownish, an apex...... 10. obscure reddish spot above the posterior coxae, Wings not so clouded ...... Il a large pale yellow spot above the front coxae; ...... 51 52 Bulletin American Museum of Natural History [VTol. LXXX a bioad yellow stripe, broadened at each end, Thorax black, a large pale yellowish spot extends from the humeri almost to the base of above the front coxae, a small spot on the the wings and the posterior calli are yellowish. humeri and another in front of the wings reddish The pleura are mostly thinly sericeous, the meso- yellow; mesonotum rather brownish pollinose notum with obscure tiny blackish hairs; scu- on the disc except for a pair of weakly metallic tellum all black, with extremely short silvery vittae, the pleura mostly whitish pollinose. hair. Hair on the dorsum black, on the pleura and Legs blackish, the anterior femora with a large scutellum whitish, the scutellum with several brownish red area below on the basal half, the very short erect coarse hairs at the apex. aniterior four tibiae brownish posteriorly and Legs black; posterior surface of the front four weakly sericeous, their tarsi rather reddish yellow tibiae and most of the first segment of their tarsi and sericeous on the basal half of the first seg- and the apex of their femora posteriorly, pale ment posteriorly. yellowish and with silvery reflections in inost Wings with brown tinge, the costal border lights. narrowly darkened on the basal three-fourths. Wings tinged with dark browin, the costal Apical cell slightly narrowed apically. Squamae border rather broadly brown. Squarnae yellow. yellow. Halteres brown with the apical third Halteres brown, the anterior surface yellow on of the knob yellow. the apical half. Abdomen black, the lateral third of the second Abdomen black, the sides broadly pale yellow- to fourth segments and the whole of the fifth ish from the base of the second to the basal reddish, the red color extending weakly onto the fifth of the sixth segment. Hair short, black, sixth segment, the black stripe on the fourth white basally, and yellow on the yellow areas. segment rather weak. Hair very short, black, The broad ventral margins of the tergites are dis- rather golden on the reddish areas, white basally. tinctly darkened. The apex of the abdomen is HoLOTYPE.-Female, Pie de Palo, San moderately broadened. Juan, Argentina. HOLOTYPE.-Male, Kabelstation (Para- This species most closely approaches the maribo), Surinam, September 21-27, 1938 Nearctic macer Loew than any of the others (D. C. Geijskes). known from South America. It differs from conopoides Kunckel (from Argentina) in Systropus willistoni, new species many respects but may be readily dis- Black, the thorax and abdomen with yellow tinguished by the wvholly red fifth abdomi- markings; wings cinereous hyaline, narrowly nal segment and the much less expanded ab- darkened in front. Length, 18 mm. FEMALE.-Sides of oral margin yellow; face domen. and frontal triangle black, silvery pollinose, an orange triangle surrounding the antennae. First Systropus conopoides Kunckel antennal segment three-fourths as long as the KtTNCICE;L, 1905, BuIl. Sc. France et Belg., head-height, the others missing. XXXIX, P. 147.-1904, Compt. Rend. Acad. Thorax black, large pale yellowv triangles on Sci. Paris, CXXXVIII, p. 1623 (notes). the humeri and in front of the wings and a large This species is illustrated by the author spot above the front coxae; mesonotum with a and compared pair of narrowly separated metallic vittae bear- with foenoides WNestwood ing white pollen, the pleura with silvery white from Mexico. The description is not pollen. Hair black, white on the pleura. Scu- sufficiently clear to permit inclusion of the tellum with whitish pollen and hair on the apical species in the key but none of the species two-fifths. Legs black, the apex of the anterior four before me agrees with it. No name was femora posteriorly, posterior surface of their applied to the species in 1904, but its tibiae and most of the posterior surface of the biology was discussed. first segment of their tarsi yellow and covered with silvery pollen. Systropus geijskesi, new species Wings tinged with brown, darkened along the costa; apical cell more than half as wide at the Black, the thorax and abdomen with yellowish tip as at the widest part. Squamae yellow. markings; apical cell strongly narrowed at the Halteres reddish, the knob yellow in front, black apex. Length, 19 mm. behind. MALE.-Sides of oral margin pale yellow, the Abdomen black, the sides of the second to face and front black, silvery white pollinose, fourth segments and the base of the fifth broadly the vertical triangle reddish; occiput white yellow. Hair black, white on the yellow aieas pollinose. Antennae black, the first segment and on the lateral third of the first segment. three-fourths as long as the head-height, not Apex of abdomen scarcely quite so long as the apical two segments com- widened. bined, the third segnment alnmost three times as HOLOTYPE.-Female, Chapada, Brazil, long as the second. December (Williston Collection). 1942] Curran, American Diptera 53 There is a second specimen that is prob- Femora black or castaneous behind. .3. ably a male, but the apical abdominal 3.-Wings brown in front, dark grayish or segments are missing. The third antennal smoky behind, with a yellowish segment is about twice as long as the stripe extending from the anal angle second; the white hair on the scutellum is to the discal cell (Guiana) ...... limited to the lateral third; otherwise it ...... cayennensis Fabricius agrees with the type. Wings more tawny, scarcely darkened along the costa, the apex and pos- Systropus dimidiatus, new species terior border dark (Peru)...... with yellow markings; wings yellowish Black, ...... inca,n. sp. brown on the basal half except posteriorly. reddish.. 5. Length, 18 mm. 4.-Femora wholly ...... FEMALE.-Eyes narrowly separated; face and Femora extensively black or castane- front black, silvery pollinose; sides of oral margin ous...... 6. yellow, but brownish near the eyes. Antennal 5.-Tarsi with golden yellow hair above, swelling reddish yellow, the antennae missing. Thorax black, the pleura with silvery pollen the bristles black (Brazil) ...... and white hair. A large pale yellow spot above ...... coarctatum Perty. the front coxae, the posterior calli yellow and the Tarsi, except the first segment of the large triangle on the humeri and in front of the hind pair, with black hair and wings rather broadly connected along the sides of the mesonotum. The mesonotal vittae are bristles above (Surinam)...... not strong, not metallic and conspicuously sepa- ...... auricinctumSchiner rated by a weak grayish pollinose line; mesonotal 6.-Posterior tarsi yellow haired above . 7. hair black, that on the scutellum white on the Posterior tarsi with black hair above apical two-fifths, the scutellum without erect black setulae apically. except on part of the first segment. . Legs black; apex of the anterior four femora ...... bassleri, n. sp. posteriorly, posterior surface of their tibiae and 7.-Central American species...... most of the posterior surface of the first segment ...... secabilisWalker. of their tarsi pale yellow and silvery pollinose. Wings yellowish brown in front as far as the Cuban species ...... annulatum Bigot. anterior crossvein and in the second basal cell, otherwise cinereous hyaline. Squamae yellow. Blepharepium vorax, new species Halteres brownish red, the knob almost all reddish yellow but brown basally. Legs wholly reddish; mesonotum dark Abdomen black, the sides of the second to with blackish sides; wings dull orange with dark and with short apex and broad posterior border. Length, 18 to fourth segments broadly reddish 19 mm. whitish hair, the hair black, white on the sides MALE.-Face, cheeks and anterior border of of the first segment. Apex of abdomen moder- the front reddish in ground color, the face and ately broadened. front whitish pollinose; front shining black, HOLOTYPE.-Female, Chapada, Brazil, with brown pollen on the sides and vertex. January (Williston Collection). Occiput black, the pollen somewhat brassy but broadly grayish white along the orbits, the hair This is the only American species I have black. Mystax whitish, composed of more than seen in which the eyes are distinctly sepa- twenty bristles. Palpi reddish, with black hair rated. The character may not apply to on the apical half. Antennae reddish, black the male. haired, the third segment long and tapering apically. Asilidae Mesonotum dull dark reddish, a sublateral interrupted stripe and the posterior border BLEPHAREPIuM RONDANI black; a pair of narrow median vittae, which unite behind, and two lateral spots very pale TABLE OF SPECIES brassy yellowish; hair sparse, black. Pleura dull blackish, the propleura, a broad band on the 1.-Anterior coxae with wholly pale hair mesopleura and upper part of the sternopleura and bristles ...... 4. and another below the squamae reddish, the Anterior coxae with bristly black pollen pale brassy yellow; lateral slopes of the hair and some pale bristles api- metanotum yellowish in ground color, separated 2. from the pale pollinose median area by a broad cally...... brown stripe. Scutellum reddish yellow with the 2.-Legs wholly reddish (Peru)...... narrow base and sides brown. Hair on sterno- ...... vorax,n. sp. pleura black above, yellow below. 54 Bulletin American Museum of Natural History [VTol. LXXX

Legs reddish, the trochanters brown or black; Palpi reddish yellow, the apical half brown below, hair black, golden yellow on the broad antero- the hair mostly black. Proboscis reddish to pale ventral surface of the front tibiae and on the castaneous. posterior and ventral surfaces of the posterior Thorax black in ground color, the humeri tibiae except at the base. paler. Mesonotum with a brownish vitta that Wings dull orange or brownish yellow with the is divided anteriorly and bordered with brownish broad apex and posterior border dark, smoky or yellow; the lateral spots, scutellum and noto- brownish. Knob of halteres brown. pleural spots brassy yellow, the pleura with the Abdomen blackish, the apex of the first seg- usual pale pollinose bands of brassy yellow, that ment, the second except basally and the sides on the mesopleura broadly white in front, the and apex of the third reddish, the apices of the propleura usually without conspicuous pale following segments similarly colored. Hair pollen. Hair black. Scutellum yellow in ground black, yellow on the tips of the two apical seg- color. ments and on the genitalia. Pollen cinereous Legs dark reddish, the coxae brown in ground yellow basally, more golden yellow beyond the color, the femora and tibiae blackish or dark third segment, the fourth to sixth segments with castaneous posteriorly and ventrally, the pos- large lateral triangles of brown pollen basally terior tibiae rarely almost all blackish. Hair on each side. Second and third sternites reddish black, tawny on the ventral surface of the pos- with yellow hair, the apical sternites and the hair terior tibiae and anteroventrally on the front mostly reddish yellow. Genitalia reddish, the tibiae. base shining black below. Wings rather variable in color, often bright FEMALE.-The basal segment of the ovipositor orange yellow with the broad apex and posterior is broadly reddish apically and bears tawny hair; border smoky and the costal border darkened the apical sternites have only black pile. basally, or the pale area luteous, rarely poorly male, and allotype, defined. Halteres with brown knobs. TYPES.-Holotype, Abdomen black, the apices of the second and female, La Merced, Peru. following segments reddish and with bright red- This species seems to be close to subcon- dish yellow pollen; basal segments with cinere- tractus WValker, from "Amazon District," ous yellow pollen, the following with brown pollen but the color of the abdomen does not except apically, the apical segments with more or agree. less broad preapical bands of rather olivaceous pollen. Hair black, rarely a few yellow hairs Blepharepium cayennensis Fabricius on the sides of the apical segment. Second ster- Asilus cayennensis FABRICIUS, 1787, Mantissa nite mostly reddish, the basal sternites with Ins., II, p. 360. yellow hair. Genitalia partly reddish, sometimes with wholly pale hair, but the base usually bears A shining blackish species with golden or black hairs. brassy yellow pollen, the three basal ab- FEMALE.-The genital segment bears only dominal segments with yellow pollinose pale hair. fasciae. The legs are blackish, the femora and tibiae dark reddish in front. The knob TYPEs.-Holotype, male, and allotype, of the halteres varies from yellow to brown. female, Iquitos, Peru, March 13, 1924 (H. The color of the wNing is distinctive. Bassler). Paratypes, six males and five I have before me twenty-four specimens females from the following localities in of both sexes from various localities in Peru: Iquitos, March 13 and 22; Rio British Guiana. Morona, January 12; middle Rio Ucayali, September 20; Rio Huallaga, September Blepharepium inca, new species 15; Rio Santiago, November 11 and 12; Black, wings luteous to bright orange with the Achinamiza, December 21; La Merced and broad apex and posterior border dark. Length, Lacorezza, Puntamayo District, August. 18 to 24 mm. This species is most nearly related to MALE.-Face usually brownish in ground cayennensis Fabricius, but it does not have color with the margins broadly reddish or yellowish, rarely almost all reddish yellow, the cheeks the pale streak in the wing, and the anten- brownish and with brown pollen; face and lower nae are almost always darker. Despite the half of the front very pale brassy yellow polli- variation in wing color it appears certain nose, almost whitish. Front shining black, the that all the material belongs to a single sides broadly opaque. Occiput black, the orbits broadly grayish yellow, the central portion with species. rich brownish yellow to yellowish brown pollen. Hair black. Antennae usually dark reddish, the Blepharepium bassleri, new species third segment often brown on the apical half or Femora bicolored; front coxae wholly pale more, all the segments rarely brown or blackish. haired; tarsi black haired; wings luteous or pale 1942] Curran, American Diptera 55 brownish orange with weak dark posterior border Blepharepium coarctatum Perty and apex. Length, 18 to 20 mm. FEMALE.-Face and cheeks reddish, the face Laphria coarctata PERTY, 1830-34, Delect. with a roughened ferruginous stripe on the upper Animal. Arctic. Brasil, p. 181 (f.). half; pollen brassy yellowish, brighter on the The wings are brownish with an almost occiput; upper part of front shining black with hyaline median stripe as in cayennensis, a narrow band of brown pollen at the vertex and along the orbits. Bristles black but a number of the second and third abdominal segments fine yellow ones on the lower part of the occiput. reddish and the legs wholly reddish. The Antennae brownish red, the third segment some- tarsal hair is all yellowish. times mostly brown. Palpi dark reddish, the I have seen coarctatum only from Brazil. hair yellow, partly black on the apical half. It is represented in the Museum Collection Thorax blackish in ground color, the humeri and lateral margins of the mesonotum more or by two males and three females from the less reddish, the propleura partly reddish, and Williston Collection. there is a large spot on the upper part of the sternopleura that sometimes extends onto the Blepharepium secabilis Walker mesopleura; lateral slopes of the mesonotum usually reddish and usually a reddish spot be- Dasypogon secabilis WALKER, 1860, Tr. Ent. low the posterior spiracle. The mesonotum has Soc. London, p. 276. the usual dark and brassy yellow stripes and the The mesonotum is almost uniformly pleura the usual brown bands. Scutellum brassy colored, the outer dark vittae rarely with yellow above, brownish luteous on the upper or part of the apex and with yellow pollen below. dark brown cast, usually reddish ferrugi- Legs reddish, the femora daik brown or cas- nous. The anterior femora are usually all taneous posteriorly and ventrally, the tibiae reddish, the posterior four black on the dark behind except apically. Hair black, yellow basal half or more, the black rarely reduced, on the anterior coxae. seldom almost absent on the middle legs. Wings brownish yellow to dusky orange, the Hair on the tarsi all yellowish. apex and posterior border broadly darker but less contrasting than in inca. Knob of halteres Eight specimens from Mexico (Williston black or reddish. Collection) were reported upon in the Abdomen black, the apices of the segments Biologia. There is a female from Honduras, narrowly reddish, the pale color expanded on the October, 1928, and two females, Fort sides of the second segment. Hair black, Randolph, Panama Canal Zone, January yellow on the first segment, broad apex of the second, basal three sternites and genitalia. 23, 1929 (Curran). The Panamanian specimens have the dark stripes more conspic- TYPEs.-Holotype, female, Rio Mara- uous than the others. non, Peru, August 28, 1923; paratypes, female, Rio Santiago, Peru, November 11, Blepharepium annulatum Bigot 1924, and female, Iquitos, Peru, March Senobasis annulata BIGOT, 1856, in Sagra, 19, 1924, all collected by H. Bassler. Hist. Isla de Cuba, Part 2, VII, p. 331 (f.). The single damaged female before me is Blepharepium auricinctum Schiner very similar to secabilis, but the dark stripes on the mesonotum are blackish. I can see Senobasis auricincta SCHINER, 1867, Verh. 371. no other differences worthy of note. Per- Zool.-bot. Ges. Wien, XVII, p. haps both names apply to the same species, Bromley, without seeing specimens, but it is impossible to decide without more placed this species as a synonym of cayen- material. nensis Fabricius. It is a very different appearing species with wholly reddish legs, DORYCLUS JAENNICKE as pointed out by Schiner in his comparison All the described species in this genus with secabilis Walker. There is evidently have been placed as synonyms of distendens an error in Schiner's description since he Wiedemann, but it seems almost certain states that the posterior claws are minute. that they are distinct, even though the de- He undoubtedly refers to the pulvilli. I scriptions leave much to be desired. The have before me one male and two females four species before me are easily separated collected by Dr. D. C. Geijskes in Surinam and show differences in both color and in June and July. structure. 56 Bulletin American Museum of Natural History [Vol. LXXX

TABLE OF SPECIES partly reddish, the wings clear hyaline. Length, about 12 mm. 1.-WVings hyaline, at most slightly dark- MALE.-Front shining reddish brown, the ened along the veins ...... 2. face more reddish; hair black, yellowish on the Wings rather extensively brownish.... upper half of the occiput and vertex; vertex white pollinose, the face with a yellowish tinged ...... 3. pollinose triangle on each side. Antennae rusty 2.-Legs wholly black; metanotum smooth reddish, the hair black. Palpi reddish basally, (Brazil).... distendens Wiedemann. becoming brown at the apex, the hair black, Femora ferruginous; metanotum very yellow on the basal half of the lower surface. Thorax shining black, with brownish red areas strongly roughened in the middle as follows: a broad band covering the ptero- (Panama) ...... panamensis, n. sp. pleura and upper part of the sternopleura, a tri- 3.-Thorax and abdomen brownish or cas- angle below the halteres, the anterior and pos- taneous; wings yellowish with very terior borders of the mesonotum and the lateral margins behind the roots of the wings and the extensive brown coloring (see Cur- scutellum. The mesonotum has a pair of approx- ran, 1934, North American Diptera, imate grayish vittae on the anterior three- Asilidae, Fig. 122) (Brazil)...... fourths and a pair of broader lateral vittae on ...... crassitarsis the posterior three-fourths; the sides of the meta- Macquart. notum and metasternum are also grayish polli- Abdomen blue or violaceous; wings nose, the mesonotum strongly roughened in the hyaline basally ...... 4. middle. Hair black on the mesonotum, else- 4.-Basal antennal segments black (Brazil) where cinereous and very fine. Legs black, the anterior four femora, the ...... varipennis Walker. under half of the hind pair and the ventral sur- Antennae reddish ...... 5. face of all the tibiae, except apically, brownish 5.-Legs brown, the front ones black; hind red. tibiae testaceous; palpi black Wings hyaline, the veins brown. Squamae (Mexico)... cyaneiventris Macquart. white, the halteres brownish. Abdomen metallic dark blue; hair pale brassy Legs reddish, the anterior tibiae and all yellowish, the middle of the apical four segments the tarsi black; palpi ferruginous and the genitalia with black hair. (Surinam) ...... l atipes WVulp. HOLOTYPE.-Male, Corozal, Canal Zone, January 16, 1929 (Curran). Doryclus distendens Wiedemann Asilus distendens WIEDEMANN, 1828, Ausser. Doryclus crassitarsis Macquart Zweifi., I, p. 571. Megapoda crassitarsis MACQUART, 1846, Dipt. This species is black, the abdomen viola- Exot., Suppl. I, p. 70 (f.). ceous; the wings hyaline with black veins which are bordered with faint clouding. This is the species illustrated in my The median pollinose vittae on the mesono- North American Diptera under the name tum are narrow, the outer ones wide and distendens. It is very different from that interrupted at the suture. The hair in front species and may be readily recognized by of the halteres is black on the lower half the ferruginous coloration. The pollinose and whitish above. The pollinose spots on stripes on the mesonotum are very wide the face are unusually broad and not tri- and only narrowly separated from each angular. It is the only species described other. There is a pair from Chapada, with the legs wholly black. None of the Brazil (Williston Collection). names placed as synonyms of distendens in catalogues belongs here. I have before me Doryclus latipes Wulp a single female from Nova Teutonia, Brazil, WULP, 1870, Tijd. v. Ent., XIII, p. 216 (f.). collected by Fritz Plaumann. The type, The face is ferruginous, the front shining from Brazil, was a male. brown. The pollinose vittae on the meso- Hermann states that guntheri Arribal- notum are united for much of their length zaga is the same. and the pleura is largely ferruginous, as are the sides of the mesonotum and scutellum. Doryclus panamensis, new species The legs are reddish with the tarsi and tip Black, the abdomen metallic blue, the legs of the hind tibiae black. The wing pattern 1942] Curran, American Diptera 57 is very similar to crassitarsis, but the pale Legs black, the trochanters, bases and apices areas are much more nearly hyaline. of the femora and the basal fourth or less of the tibiae pale orange; tarsi brownish red. Hair One female, Botanical Garden, Para- and bristles whitish. Pulvilli only about one- maribo, Surinam, May 10, 1941 (D. C. third as long as the claws. Geijskes). Wings cinereous hyaline, the veins brown. This does not agree with the description Squamae and halteres yellowish. Abdomen cinereous pollinose, the segments ap- of varipennis Walker or cyaneiventris Mac- pearing darker on the basal half in some lights. quart, but it is possible that it is the same Genitalia short, shining blackish. Hair whitish. as one of them. It is not impossible that FEMALE.-The legs are miostly reddish yellow: all three names represent a single species. front femora with narrow black band, the middle pair about half black, the posterior pair two- LBSTOMYIA WILLISTON thirds black. Tibiae with the bases more broadly yellowish and with reddish apices, the Since the publication of a key to the spe- tarsi reddish. Abdomen with the sixth and cies in American Museum Novitates, No. following segments shining black and with the 487 (1931), one species has been described bases of the second to fifth segments broadly bare except laterally. One female has the legs and another new species is now before me. much more extensively reddish (the anterior four femora wholly reddish), and has six scutellar TABLE OF SPECIES bristles. I.-Scutellar bristles black...... 2. TYPEs.-Holotype, male, allotype, fe- Scutellar bristles whitish...... 3. male, and paratypes, five males and two 2.-Coxae and base of femora black in females, Tempe, Arizona, April (D. K. ground color (Arizona) ...... Duncan)...... atripes Wilcox. This species is smaller, more slender and Coxae and base of femora reddish in paler in color than any of the others and ground color (California)...... has shorter pulvilli...... fraudiger...... Williston. 3.-Femora and tibiae black above, reddish SMERINGOLAPHRIA HERMANN below (Wyoming).. strigipes Curran. HE:RMANN, 1912, Nova Acta Acad. Caes. Femora black annulate or all reddish Leop.-Carol. Germ. Nat. Cur., XCVI, p. 226...... 4.Panamasitus CURRAN, 1930, Amer. Mus. 4.-Scutellum with six (rarely eight) bris- Novitates, No. 425, p. 20. tles; male genitalia reddish, twice as Hermann described the genus, basing it long as wide; anterior femora of on a species from Bolivia. Panamasilus female than half was described from a series of specimens more black (Cali- collected on Barro Colorado Island, fornia, Wyoming)...... Panama Canal Zone. The two species are ...... sabulorum Osten Sacken. Scutellum with four (rarely six) bris- easily distinguished as follows: tles; male genitalia black, not half so a.-Wings with large preapical brown long as wide; anterior femora of spot and a small one near the female at most with black median middle; mesonotum ferruginous.. band (Arizona). unicolor, n. sp...... maculipennis Macquart. b.-Wings uniformly yellowish brown; Lestomyia unicolor, new species mesonotum almost all black in Black, thickly cinereous pollinose, legs partly ground color...... white. reddish, the bristles mostly Length, 6.5 ...... panamensisCurran. to 9 mm. MALE.-Head cinereous, the hair and bristles white; mouth and proboscis shining black. Smeringolaphria maculipennis Macquart Antennae black, very thinly pollinose. Face Laphria maculipennis MACQUART, 1846, Dipt. gently gibbous on the lower half. Exot., Suppl. I, p. 73. Thorax cinereous pollinose, the mesonotum pictipennis HERMANN, 1912, loc. cit. with very approximate median and incomplete I have before me a specimen from Suri- sublateral dark vittae; bristles white, the dorso- nam centrals almost all black. The pollen of the (Kabelstation, September 21-27, 1938) middle of the mesonotum and scutellum has a collected by Dr. D. C. Geijskes which yellowish tinge. agrees with the descriptions. The species 58 Bulletin American Museum of Natural History [Vol. LXXX is apparently widespread in South America. Mallophora fairchildi, new species Dr. Bromley has called my attention to the Face wholly pollinose; abdomen without pale some white above synonymy. It is quite certain that pollen; scutellum black haired with hair at the base laterally. Length, 18 mm. the specimen before me is maculipennis, and MALE.-Head black, reddish only above the the description given by Hermann agrees anterior oral margin; face grayish yellow polli- in all details. The species is placed in Nusa nose, with black hair and bristles, some white in the Kertesz Catalogue. hairs on the upper part of the face and some white bristles in the mystax immediately above the oral margin, front with brown pollen and Mallophora geijskesi, new species black hair; occiput white pollinose and pilose, Black, the legs chiefly reddish yellow; pile with black bristles behind the eyes. Antennae -mostly yellow and tawny; face wholly pollinose. black, black haired, the arista considerably Length, 18 mm. longer than the elongate third antennal seg- MALE.-Head black in ground color, the face ment. Palpi black and with black hair. reddish yellow, with ochraceous pollen, the orbits Thorax black, with brownish pollen that be- paler; pile reddish yellow, the front with black comes more grayish on the lower half of the hair; bristles on the cheeks, on the border of pleura; mesonotum with four blackish or dark the mystax and on the upper posterior orbits brownish vittae. Hair black, white on the pleura black; frontal pollen reddish yellow, that on the but almost all black on the mesopleura; scutel- occiput pale yellowish. Antennae black, the lum with a few white hairs laterally. base of the second segment reddish, the hair Legs orange, the anterior four femora mostly black; arista slightly longer than the elongate shining black in front; tarsi black, the first third segment. Palpi black, the hair coarse, segment of the anterior four mostly reddish tawny. above, the apex of the posterior tibiae narrowly Thorax blackish, tawny pollinose, the mesono- black. Femora with black hair but with yellow tum with four brownish vittae, the median pair hair above except at the apex; anterior four narrow and short. Pile tawny, becoming yellow tibiae with black hair above and below, and on the lower part of the pleura, the disc of the reddish yellow hair in front and behind; posterior mesonotum with coarse black hair. Scutellum tibiae with yellowish hair ventrally and on most with one or two pairs of black bristles. of the posterior surface. Anterior tarsi with Legs reddish, the anterior four femora mostly whitish hair and reddish yellow bristles pos- shining black in front, the posterior tarsi with teriorly, the anterior bristles black; middle tarsi the basal four segments black, not conspicuously with black hair, the posterior pair black haired, expanded, the hair black, longer and white on but the upper surface of the second to fifth seg- more than the anterior half of the apical three ments bears increasingly long white hair dor- segments, the apical segment without black hair. sally, and there is a narrow stripe of white hair Pile of the legs tawny, black on the under sur- on the dorsal surface of the first segment toward face of the anterior four femora; not erect on the anterior edge; all the hair on the first seg- the posterior tibiae. ment is appressed. Wings strongly tinged with brown. Squamae Wings tinged with brown, the costal border reddish yellow; halteres reddish with brown tinge pale brownish. Squamae with white fringe. on apical half. Halteres reddish with brown knob. Abdomen black, the sides rather broadly Abdomen black, opaque, the venter sub- tawny to brownish gray pollinose, the pollen opaque. Hair whitish on the first segment, on expanding to form narrow bands on the apices the venter and on the lateral margins of the of the second to fourth segments; pile tawny, second to fifth tergites, expanded to form whitish black on the sixth and following segments, and fasciae on the second to fourth segments but on the broad apex of the fifth except laterally. these segments with fine black bristles laterally. Genitalia shining black, black haired, the basal Genitalia shining black, the basal half above third above with thick white appressed hair. with dense white appressed hair, the hair and Venter yellowish brown pollinose and tawny bristles otherwise black. pilose. HOLOTYPE.-Male, El Valle, Cocle Prov., HOLOTYPE.-Male, Brownsberg, Suri- Panama, April 13, 1941 (G. B. Fairchild). nam, September 17, 1938 (D. C. Geijskes). This species traces to ada Curran (coup- This species belongs to the second group let 29) but may be at once distinguished by and traces to couplet 18 in my recently the absence of pale pollen on the abdomen published key (1941, Jour. N. Y. Ent. Soc., and the white instead of yellow pile. Like XLIX, p. 270). It most nearly approaches the preceding species it belongs in group II. zita Curran, but the posterior tarsi are not strongly broadened, and the white hair is Dolichopidae on the anterior half of the segments and is LAXINA CURRAN longer but is not particularly dense. CUREAN, 1934, Fam. Gen. N. A. Dipt., p. 230. 19421 (.urran, American Diptera g9

This genuis was established t-o ineludle First genital segment Nith fine, little those species fo-rmerly placed in (ondylo- conLspicuous bristles ...... 11. stylus, Psilopiis, Sciapts, etc., having the 9.-The genital lamellae bear three long, lower part of the face haired. The genotype ObtuUse spines onI inner surface is patibldatus Say, w-hich ldiffers fronm all the apically (Florida to South America other species in havting the upper half of the and W-est Indies) (tenuitarsus Van face haireL. With the exception of palibu- Duzee) ... graenichtri V'atn Duzee. talus the genus is an artificial one, since 10.-Genital lamellae long, strong, bare on there are species renmaining in (Condylostyhts basal hallf, the spines strongly which agree in all other respects with some swollen apically (Guiana) ...... of the species included in Laxina. How- ...... cancer Van Duzee. ever, the separation of this group of species Genital laiiellae finer, with fine hair is very conivenient alnd aidls greatly in their on whole length, the apical spines recognition. Up to the present time there weakly swollen (eastern United has been no listing of the species of Laxina, States) ...... caudatuts Wiedemann. and I take this opportunity to present a key 11.-Genitalia with a pair of bristles longer to the nmales and discuss the various forms. than the genitalia and two or three The females of a number of the species pairs of shorter ones, all directed cannot be separated at the present time. backwards (eastern states)...... ctannectans,n. sp. TABLE OF SPECIES Genital bristles much shorter than the genitalia ...... 12. MALES 12.-North American species.13. I.-Femora black or greeniish ...... 3. South American species (Chile, Ar- Femora yellowish ...... 2. gentina). ancminatus Van Duzee. 2.-Apical segment of front and middle 13.-First segment of the middle tarsi with tarsi whitish (eastern U.nited States) antero-dorsal row of cilia composed ...... banksiV-an Duzee. of sulbrect bristles (United States) Apical segment of all the tarsi blackish ...... cocxkerelliVan Duzee. (United States). .flazipes Aldrich. First segment wiith shorter, less erect 3.-Wings hyaline ...... 4. bristly hairs (U1nited States) (scrobi- Wings with two brown, anteriorly nator Loew)...... connected crossbands on the apical ...... nigrofemoratus WValker. half (UI:nited States) patibidatus Say. 4.-Face with white hairs only ...... 6. Laxina patibulatus Say Face with the upper hair black, or all Dolichopus patibidatus SAY, 1823, Jour. Acad. black. . Nat. Sci. Phila., III, p. 87 (Florida). 5.-Facial pile all black (Nexico). Psilopus amatus W,ALKER, 1849. List Dipt., ...... occidentaiis Bigot. III, p. 648 (New York). PsAilopus carolinensis BIGOT, 188S, BUI. SoC. Facial pile white on lower half (Brazil) Ent. France, p. xxix (Carolina)...... ornfticauda Van Duzee. Psi2opus infiCitus WALKER, 1849, List Dipt., 6.-First segment of niiddle tarsi with row III, p. 649 (Mexico). of closely placed, very strongly This is one of the commonest and most curved short bristles below ...... 8. widely distributed species in the genus. It Middle tarsi wiithout such bristles. . 7. is readily recognized by the banded wings. 7.-Apex of middle tibia with very long It is the genotype of Laxina and differs spine below (United States)...... from all the other species in having hairs . calcaratuts Loew. over practically all the face, the hairs being Middle tibia wNithout spine (Pennsyl- limited to the lower half in the others. vania, eastern states)...... inermis Loew. Laxina flavipes Aldrich 8.-First genital segnient bearing two Psiopus flasipes ALDRICH, 1904, Tr. Amer. rows of extremely long bristles. . . 9. Ent. Soc., XXX, p. 284. 60 Bulletin American Museum of Natural History [Vol. LXX

Psilopus fermoratus SAY, 1829 (not 1823), Laxina ornaticauda Van Duzee Jour. Acad. Nat. Sci. Phila., VI, p. 361. Condylostylus ornaticauda VAN DUZEE, 1931, Originally described from South Dakota Amer. Mus. Novitates, No. 483, p. 9 (f.). and Massachusetts. There is a male from The types were from the Williston Collec- Ramsey, New Jersey, June 18, 1921. This tion, and the same is true of the Brazilian species is rare in collections. It is like specimen included by Aldrich in barbatus. inermis Loew, except that the femora are The only means of separating the two yellowvish. species is by the color of the hair on the face, which is more than half white in orna- Laxina banksi Van Duzee ticauda. In the females of both species the Sciapus banksi VAN DuzEn, 1915, Ent. News, front coxae are almost all green in front, XXVI, p. 23. reddish posteriorly. Described from Virginia and New York and not since recorded. It differs from Laxina caudatus Wiedemann Jiavipes in having the apical segments of the Psilopus caudatus WIEDEMANN, 1830, Ausser. Zweifl. II, p. 224 (United States). anterior four tarsi whitish. Psilopus virgo WIEDEMANN, 1830, loc. cit. Psilopus cacsdatulus LoEw, 1861, Neue Beitr., Laxina calcaratus Loew VIII, p. 79. Psilopus calcaratus LoEw, 1861, Neue Beitr., L. caudatus is apparently confined to VIII, p. 93 (Carolina). North America and probably does not ex- Differs from all the species with hyaline tend south beyond Mexico. I have speci- wings and black femora (except inermis) mens from the area between Florida and by lacking the curved bristles on the middle Canada, but other references refer to tarsi. It differs from inermis by the pres- graenicheri and ornaticauda Van Duzee. ence of a very long, strong, suberect bristle The spedies figured by Becker from Brazil on the lower apex of the middle tibia. Our apparently was ornaticauda, although it is material is from New York, Illinois and possible that it is smaragdulus Wiedemann, Indiana. known only from a female from Uruguay. In this group of species there is a group of long bristles apparently attached to each Laxina inermis Loew side of the first genital segment, the bristles Psilopus inermis LoEw, 1861, Neue Beitr., very long above and becoming short below VIII, p. 93 (Pennsylvania). and forming a more or less fan-like row. Lacks the strong bristle on the middle The species are very similar in all respects tibiae but otherwise similar to calcaratus. but show differences in the male genitalia Our specimens are from New York, Massa- and the color of the front coxae in the fe- chusetts, North Carolina and Wisconsin. males. In the female of caudatus the anterior coxae are yellow except a small basal Laxina occidentalis Bigot spot on the outer side, a characteristic of Psilopus occidentalis BIGOT, 1888, Bull. Soc. most of the Nearctic species. In the Neo- Ent. France, p. xxix. tropical species the anterior coxae are usu- Psilopus barbatus ALDRICH, 1901, Biol. Centr. Amer. Dipt., I, p. 359. ally green on the basal half or more. PARENT, 1932, Ann. Soc. Sci. Brux., LII, p. 224. Laxina cancer Van Duzee Condylostylus cancer VAN DuZEE, 1934, Bull. This is the only species in which the hair Amer. Mus. Nat. Hist., LXVI, p. 366 (f.). on the face is all black. In ornaticauda Van ?Psilopus caudatulus ALDRICH, 1896, Tr. Ent. Duzee the hair is black above and white Soc. London, p. 343. below. Aldrich described barbatus from Described from a single male from Mexico and Brazil, but it is possible that Georgetown, British Guiana. It differs the specimen from the latter country is re- from caudatus in having the outer genital ferable to the following species, provided lamellae broader and very much longer, the two are distinct. curved inwardly apically and with longer, 1942] Curran, Amwerican Diptera 61 stouter, more capitate, backwardly directed with white hair. Middle tibiae with very long, two erect, apical bristle below, the first segment of spines on the inner apex. There are their tarsi with a series of anterior and posterior, females from St. Vincent and one from short, curved spines on ventral surface, the pos- Colombia that I believe belong here. The terior row double and with an antero-dorsal row anterior coxae are green except at the tips. of downwardly curved cilia. Posterior tibiae blackish apically, the apical segment of their tarsi They might, however, be ornaticauda since distinctly broadened. the female of that species bears only wvhite Wings hyaline. Squamal cilia black. Halteres hair on the face. yellow. Abdomen green, the basal half more or less Laxina graenicheri Van Duzee strongly bluish or violaceous, the apical segments with broad, dull black basal fascia. Genitalia Condylostylus graenicheri VAN DrZEE, 1927, brown, the basal segment bearing three or four Ent. News, XXXV,III, p. 73 (Florida). bristles on each side, the upper one very long Psiloputs caztdatus ALDRICH, 1901, Biol. Centr. and strong, the others not so long as the geni- Amer. Dipt., I, p. 360. talia, outer lamellae small, brown, lying close Psilopus caudatulits ALDRICH, 1902, Kans. to the genital segment. Univ. Sci. Bull., I, p. 94. FEDMALE.-Similar to related females but ap- PARENT, 1933, Bull. Ann. Soc. Enit. Belg., parently with a slightly larger green spot on the LXXIII, p. 175 (f.) (Cocas Island). outer surface of the front coxae at the base. Condylostylus tenuimanus VAN DrzEE, 1931, Amer. Mus. Novitates, No. 484, p. 2 (f.) (Hon- TYPEs.-Holotype, male, and allotype, duras). female, Avon Old Farms, Connecticut, This species is not rare in Florida, Cen- June 16 and 18, 1929 (Curran). Para- tral America and some of the West Indies, types, male, Black Mountains, North but it is not the species recorded from St. Carolina, June 24-30, 1906 (W\. Beuten- Vincent by Aldrich under the name cauda- muller) and male, Waverley, Massachu- tulus. The female has the front coxae setts. green on about the basal half, while the The type male was determined as cauda- male bears three stout, weakly clavate tus, the female as scrobinator and one para- spines on an inner lobe of the outer genital type as caudatulus. The species is inter- forceps. It is easily recognized by the mediate between nigrofemoratus and cau- genital characters. datus, but the genital lamellae are almost The type of tenuimanus is a poorly pre- as in nigrofemnoratus. served specimen, and Van Duzee was misled because the long bristles on the geni- Laxina nigrofemoratus Walker were all broken off. As a result he Psilopus nigrofemoratuts WALKER, 1849, List talia Dipt. Brit. Mus., III, p. 650 (North America, considered it to be related to scrobinator Nova Scotia). Loew. The genitalia are identical with a Psilopus scrobinator LoEw, 1861, Neue Beitr., paratype of graenicheri in the Museum VIII, P. 91. ?Condylostylus acuminatus VAN DUZEE, 1920, Collection. Dipt. Patagonia, V, p. 89 (Argentina). Laxina connectans, new species There can be no doubt about the syn- Metallic green, wings hyaline; basal genital onomy of scrobinator and nigrofemoratus, segment with a single pair of long bristles and but acu?ninatus may not belong here. several short ones; femora black. Length, 4 to Parent, who examined the types of Walker 5 mm. and Van Duzee, stated that they were the MALE.-Face more or less bluish or violaceous, rather thickly, white pollinose; front sometimes same, but Van Duzee indicated a slight mostly bluish, its hair white. Antennae black, difference. The statement by Parent the longest bristle on the second segment longer leaves no doubt about scrobinator, a com- than the antennae. Proboscis reddish yellow, mon species in Canada and the United palpi black, with white hair and two or three black bristles. States. The species (if distinct) belonging Thorax green to violaceous, the pleura white to this group have very small, triangular, pollinose. Five pairs of dorsocentrals; two pairs outer genital lamellae, a very long erect of large scutellars. bristle on the apex of the middle tibiae and Legs green, the tibiae, tips of the femora and on most of the first segment of the front tarsi two series of short, stout, curved spines yellowish; coxae and lower surface of the femora the basal segment of the middle tarsi. The 62 Bulletin American Museum of Natural History [Vol. LXXX number of these spines varies slightly. All mens from North and South America is of the species lack the long bristles on the necessary before the relationship of the basal segment of the genitalia, the bristles forms can be established. present being short, a single pair rarely equaling the length of the genitalia. Laxina smaragdulus Wiedemann Psilopus smaragdulus WIEDEMt.NN, 1830, Laxina cockerelli Van Duzee Ausser. Zweifl., II, p. 225. Psilopus cockerelli VAN DUZER, 1927, Ent. This species was described from a female News, XXXVI, p. 73 (Colorado). from Uruguay, and it cannot be placed at This form, according to the description, the present time. If the front coxae are differs from nigrofemoratus only in having wholly yellow (and the same is true of an antero-dorsal ciliation composed of fine, acuminatus Van Duzee) it would be the short bristles on the first segment of the same as acuminatus, but if the front coxae middle tarsi. One of our specimens was are greenish it would probably be ornati- determined by Van Duzee who stated that caudus Van Duzee. There can be little the ciliation was not present in scrobinator. doubt that it is the same as one or the other Other specimens determined by Van Duzee of these species. See discussion under as scrobinator have the ciliation as fully acuminatus. developed as in the specimen determined as cockerelli, and all the specimens under Laxina femoratus Say scrobinator in the collection show a ciliation, Dolichopus femoratus SAY, 1823, Jour. Acad. but in some the hairs are much less erect, Nat. Sci. Phila., III, p. 86 (Pennsylvania). but apparently not less numerous nor no- Say described the male from Pennsyl- ticeably shorter. I think that Mr. Van vania and later mentioned a specimen from Duzee failed to notice this character and Indiana which is undoubtedly flavipes Al- was therefore misled into describing cocker- drich. Wiedemann had a specimen re- elli. About two-thirds of our material ceived from Say which he described but from various localities would be cockerelli, which was not found in the Wiedemann while the others, most of which can be only Collection. This may have been the type, doubtfully assigned, approach his concep- and the identification of the species must tion of scrobinator. I believe that both rest upon the location of this specimen. names apply to the same species and that It is safe to presume that femoratus is the cockerelli should be placed as a synonym of same as either inermis Loew or nigro- nigrofemoratus Walker. femoratus Walker. Had Wiedemann mentioned the ornamentation of the middle Laxina acuminatus Van Duzee tarsi in caudatus it would be reasonably Condylostylus acuminatus VAN DuzzEa, 1930, safe to assume that the tarsi were not orna- Dipt. Patagonia, V, p. 89 (Argentina). mented in femoratus, and the name could Parent, who examined the type, placed then be applied to inermis with safety. The this as a synonym of nigrofemoratus Walker. fact that no mention is made of this char- Mr. Van Duzee stated that it differed from acter makes femoratus unrecognizable. scrobinator Loew in having longer and more Both inermis and nigrofemoratus are com- abundant hair on the lower part of the mon in Pennsylvania. apical abdominal segments. Since he considered that scrobinator lacked ciliation on the first segment of the middle tarsi it is Conopidae possible that acuminatus is different. If STYLOGASTER MACQUART there is not a row of short antero-dorsal MACQUART, 1835, Hist. Nat. Dipt., II, p. 38. bristles the species is distinct. Mr. Van This genus has been reviewed by Aldrich' Duzee had only a single male. Should the (1930) and Lopes2 (1937), and both dis- female have the anterior coxae extensively cussed the habits of the species. An un- would be dis- greenish the species certainly 1 Proc. U. S. N. M., LXXVIII, Art. 9, pp. 1-27. tinct. Comparison of a series of speci- 2 Arch. Inst. Bi;O. Veg. (Brazil), III, pp. 257-293. 1942] Curran, American Diptera 63 fortunate error has crept into the literature Middle tibiae with short sparse hair through the use of the words "army ants" on whole length...... 5. in the note forwarded to Dr. Aldrich. It is 4.-First segment of middle tarsi with an true that I observed and captured speci- anterior swelling before the middle mens over moving columns of army ants bearing six erect, black spinose of the genus Eciton, but most of the speci- hairs in a row (Brazil) ...... mens captured were associated with "ant ...... ornatipes Kroeber. armies"-that is, enormous numbers of ants Middle tarsi simple, the first segment much smaller than most Eciton. It would not swollen (Costa Rica)...... scarcely be true that Stylogaster could not ...... decorataAldrich. be an internal parasite of ants as large as 5.-Basal segment of the middle tarsi Eciton, but they could not be of the ants normal.. 6. comprising the armies I observed most fre- Basal segment of the middle tarsi quently. In the third paragraph of my with long bristles apically, the note published by Dr. Aldrich (p. 5) the second with dense hair on basal words "army ants" should be changed to half.. tarsata Lopes. "ant armies" in order to present a true 6.-Posterior tibiae with white or yellow picture. I may further add that I do not preapical band...... 7. believe these were actually ant armies on Posterior tibiae dark yellow, the the war path. It is much more likely that apical fifth brown (Panama)...... they were entire ant colonies moving to a ...... indistincta Aldrich. new home. 7.-Posterior tibiae black, with broad In the Aldrich key to females, S. speciosa base and preapical band pale .. .8. (couplet 7) belongs in the first division of Posterior tibiae with the apex black, the genus and will not trace out, while in the dark portion before the pre- couplet 7 of the Lopes key, ethiopa Town- apical band reddish brown, not send does not belong but actually traces to black (Brazil). .longicornis Lopes. couplet 8. Sr. Lopes correctly placed 8.-Third antennal segment only a little speciosa, and the position of ethiopa is cor- longer than the second on the outer rected in the following key. side (Brazil) ...... nigrifrons Lopes. Although most of the species are repre- Thirdsegmenttwiceaslongassecond on sented in the Museum Collection it has outer side (Guiana) geijskesi, n. sp. been necessary to rely upon descriptions 9.-First abdominal segment broadly in order to include some of the species, and yellow at the sides ...... 12. free use has been made of the keys in the First abdominal segment dark in color above mentioned works. The excellent laterally, at least strongly stained illustrations by Lopes will greatly facili- with brown...... 10. tate the identification of the species. 10.-Apical sternite with cluster of black setulae on each side of a broad deep TABLE OF SPECIES median emargination (Central MALES America) .... minuta Townsend. l.-Ocellar triangle long, almost or quite Apical sternite without setulae... 11. reaching the frontal lunule ...... 2. 11.-Middle third of posterior femora and Ocellar triangle short, rarely extend- preapical fifth of their tibiae whitish ing beyond the middle of the front (Panama) ... . banksi Aldrich...... 9. Posterior femora and tibiae almost 2.-Ocellar triangle opaque or scarcely unicolorous brownish (Central shining (eastern United States) .... America) .... apicalis Aldrich...... neglecta Williston. 12.-Third antennal segment three times Ocellar triangle polished ...... 3. as long as the second on the outer 3.-Middle tibiae with dense black hair on side; bristles on sides of basal two the apical half of the ventral sur- abdominal segments wholly pale face ...... 4. (Guiana) .... fasciata Aldrich. 64 Bulletin American Museum of Natural History [Vol. IXXX Third segment not twice as long as the a little beyond the middle of the second;. bristles on basal abdominal front...... 11. segments mostly black...... 13. 2.-Ocellar triangle polished ...... 4. 13.-Sides of the fourth abdominal tergite Ocellar triangle subopaque, scarcely greatly produced as ventral lobe shining...... 3. and bearing bristles posteriorly 3.-Ovipositor almost three times as long (Guiana). breviventris Aldrich. as the abdomen, with preapical Sides of abdominal segments normal . yellow band (Brazil)......

...... 14...... macruraLopes. 14.-Front, except the lateral margins, Ovipositor little more than twice as uniformly dull brownish red, the long as the abdomen, the apex very orbits brown. (Panama)...... broadly yellow (United States)...... currani Aldrich...... neglecta...... W.illiston. Front differently colored...... 15. 4.-Bristles of the first abdominal seg- 15.-Last section of fourth vein nearly ment black...... 5. straight, the apical cell narrowing Bristles of the first abdominal segmnent from the posterior crossvein to the all yellowish...... 6. apex...... 16. 5.-Apical segment of ovipositor narrowly Last section of fourth vein very yellow at base; white band of hind strongly curved, the apical cell wide tibiae almost all black haired to well beyond the posterior cross- (Peru) - .- peruviana Aldrich. vein...... 18. Apical segment of ovipositor yellow- 16.-Fifth sternite bearing two groups of ish on basal half; white band of black setulae before the apex (Cen- hind tibiae pale pilose except below tral America) ....pectinata Aldrich. (Brazil)... australis Lopes. Fifth sternite without such setulae... 6.-Posterior crossvein almost parallel ...... 17. with the wing margin; posterior 17.-Fifth sternite with a row of back- tibiae with white annulus...... 7. wardly directed fine bristles on the Posterior crossvein nearly transverse; apex (Guatemala)...... hind tibiae not white annulate ...... bequaerti, n. sp. (Panama).... panamensis Aldrich. Fifth sternite without bristles apically 7.-Apical segment of the ovipositor black (Guiana, Costa Rica)...... pilose on whole length .... 8...... rectinervis Aldrich. Apical segment of the ovipositor white 18.-Anterior coxae with long, soft yellow haired on basal half (Panama).... hair in front and behind at apex ...... speciosa Aldrich...... 19. 8.-Anterior and middle trochanters yel-

Anterior coxae with black hair in low...... 9. front at apex (Central and South Anterior and middle trochanters cas- America).. stylosa Townsend. taneous (Brazil)... . tarsata Lopes. 19.-Mesopleura mostly brown, antennae 9.-Third vein ending well before the tip broadly black apically on upper of the wing...... 10. half; abdomen with blackish Third vein ending very slightly be- fasciae (Brazil). .stylatus Fabricius. fore the tip of the wing (Guate- Mesopleura reddish yellow, antennae mala) .. ... azteca, n. sp. all reddish yellow; abdomen with 10.-White preapical band on hind tibiae weak brownish fasciae (North with black hair only above (Central America) .... biannulatus Say. America)..... ethiopa Townsend. White band with only a few white FEMALES hairs (Brazil).. ornatipes Kroeber. 1.-Ocellar triangle long, almost or quite 11.-econd and third abdominal segments reaching the frontal lunule...... 2. entirely yellow (Bolivia) ...... Ocellar triangle extending at most to .r.*-.abdominalisKroeber. 1942] Curran, American Diptera 65

Second and third segments with at The type male is in the American least pale brown fasciae...... 12. Museum collection, having been donated 12.-Second and third segments with only by the New York Zoological Society narrow brownish posterior fasciae through Dr. William Beebe, together with (United States) .... biannulata Say. the types of breviventris. Through a mis- Abdominal segments withbroadblack- understanding Dr. Aldrich recorded the ish fasciae or mostly black..... 13. types as being in the Canadian National 13.-First abdominal segment with black Collection. In both these species the ab- bristles laterally...... 15. domen is short and broad. First segment with only yellow S. rectinervis is peculiar in that the pos- bristles ...... 14. terior tibiae are produced as a strong, tri- 14.-Second segment of ovipositor white angular spur at the apex in the male. I do and with white hair on basal half not have a female before me so do not (Panama) ...... speciosa Aldrich. know whether the character occurs also in Second segment of ovipositor black on this sex. basal half (Brazil)...... I strongly suspect that only the type ...... stylata Fabricius. male belongs to this species; the male from 15.-Front brownish red, the orbits brown; Costa Rica in Dr. A. L. Melander's collec- ocellar triangle scarcely passing the tion is probably a distinct species, and the anterior ocellus (Panama)...... absence of the yellow preapical band on ...... currani Aldrich. the posterior tibiae of the Kartabo female Front black at the vertex and some- indicates that the specimen represents a times on whole length ...... 16. distinct form. I have not seen any other 16.-Second segment of ovipositor white species in which the two sexes displayed except at base (Panama)...... such differences in leg coloration. If the ...... apicalis Aldrich. female of rectinervis has a pale preapical Second segment of ovipositor not fascia it will trace to banksi Aldrich and not white...... 17. to rectinervis as in the above key. 17.-Posterior tibiae with broad whitish or Stylogaster geijskesi, new species pale yellowish preapical band Black and yellow; hind femora without long (Panama) .... . banksi Aldrich. hair; front wholly black; antennae elongate. Posterior tibiae not conspicuously Length, 8 mm. banded with pale color, the apex MALE.-Front and occiput black, the front opaque, the ocellar triangle extending almost to sometimes broadly black.. 1.l8. the yellow lunule, broad and shining, the pollen 18.-Ocellar triangle extending only on the occiput golden yellow, becoming white slightly beyond the ocelli and below. Face, cheeks and lower occiput yellow, rounded in front (Central America) silvery white pollinose. Antennae yellow, the third segment orange, broadly black above, al- ...stylosa Townsend. most three times as long as the second on the Ocellar triangle extending to near the outer side, the second black above; arista wholly middle of the front ...... 19. black. Proboscis wholly black, the basal sec- 19.-Pleura wholly yellowish ...... 20. tion a little longer than the abdomen. Thorax yellow; mesonotum black, the humeri Mesopleura shining brown (Central yellow; posterior calli and a transverse prescutel- America) .... minuta Townsend. lar spot luteous; notopleura black, the pleura 20.-Fourth abdominal segment wholly with a broad brown band extending over the concealed by the third laterally mesopleura and onto the sternopleura. Hair black. Scutellum brownish, paler on the sides, (Guiana) .... breviventris Aldrich. with thin white pollen. Fourth abdominal segment visible on Legs yellow, simple; posterior femora with its whole length (Guiana, Costa very broad basal and preapical black bands, Rica).... rectinervis Aldrich. their tibiae black with broad basal and preapical yellow bands, the tarsi wholly black. Hair black, short, yellow on the anterior four tibiae Stylogaster rectinervis Aldrich except on the anterior surface of the middle pair ALDRICH, 1930, Proc. U. S. N. M., LXXVIII, and the basal half of the posterior surface of the Art. 9, p. 23. front pair. Bulletin American Museum of Natural History [Vol. LXXX

Wings tinged with brown except posteriorly; the basal segments and base of the terminal seg- apical cell long but only a little nirrower than ment rather broadly reddish, the hair all black. the submarginal cell, the last section of the fourth Sa. Emilia Po- vein carried well out in line with the preceding HOLOTYPE.-Female, section before curving forward, the third vein chuta, Guatemala, February-March, 1931 ending conspicuously before the tip of the wing. (J. Bequaert). Knob of halteres yellow, the stem brown. Abdomen dark rusty reddish, becoming brown Stylogaster bequaerti, new species apically, the first segment black with the sides Dark brownish and yellow, the pale ab- yellow, the second with a small yellow spot in dominal spots narrowly connected in front; front on each side; second segment with rather posterior femora with long pile. Length, 5.5 narrow white pollinose basal fascia, the third mm. to fifth each with a broad cinereous pollinose MALE.-Face, cheeks and posterior oral mar- basal fascia, the ground color partly black gin yellow, silvery white pollinose. Front opa- beneath them. Hair black; yellow on the sides que brown, the ocellar triangle shining, extending of the first segment and on the pollinose fasciae to the middle of the front, its border reddish laterally. Genitalia mostly reddish yellow, the yellow. Occiput black, white pollinose and with hair black. All the sternites apparently absent, white hair. Antennae short, reddish, the first but there may be a very small fifth one. segment yellow, the third mostly blackish, with HOLOTYPE.-Male, Serabreck, Surinam, a large oval spot basally above and most of the lower border broadly reddish. Proboscis with May, 1941 (Hermans), received from Dr. almost the basal half and labellae yellow. D. C. Geijskes. Pleura and humeri yellow, the dorsum brown, a broad brown band extending over the meso- Stylogaster azteca, new species pleura; posterior calli luteous. Hair black. Legs yellow; posterior coxae shining brown Black and yellow, the abdomen with broadly except on front, their femora brownish red, interrupted pale fasciae. Length, about 6 mm. yellow below, with a very broad preapical brown FEMALE.-Face, cheeks and posterior oral band and a broad basal band that leaves only border pale yellow, silvery white pollinose; the narrow base pale; posterior tibiae and tarsi front and occiput black, the occiput with white brownish red, the tarsi becoming brown apically; pollen and hair. The front becomes brown hair black, yellow on the posterior surface of the anteriorly, is opaque laterally, with the orbits middle tibiae, and on the front tibiae except white on more than the anterior half, the ocellar basally and on the basal half of the anterior sur- triangle shining, broad and reaching the lunula face and on the posterior surface of the first seg- broadly. Antennae short, pale orange, the basal ment of the anterior four tarsi. The front coxae segment yellow, the second brown above, three- have long black hair on the apex in front and fourths as long as the third, the third broadly behind. brown above except basally. Proboscis black, Wings cinereous hyaline; apical cell narrow, only the labellae yellow. the third vein ending a little before the tip of Pleura and humeri pale yellow, a broad brown the wing, the apical section of the fourth vein band extending across the mesopleura and onto gently curved. Knob of halteres blackish, the the upper part of the sternopleura. Dorsum of stem reddish yellow. the thorax shining black, the scutellum and the Abdomen brown, broadly yellow on the sides, posterior border of the mesonotum, broadly ex- the second to fourth segments with yellow fasciae panded in the middle, luteous, the scutellum that are narrow in the middle or weakly inter- darkened on the disc. Hair shining brown in rupted, those on the third and fourth segments some lights, the bristles black. very broad, the first and fourth narrow. Hair Legs pale yellow, the posterior coxae partly all black. Genitalia brown above, yellow below, shining reddish brown; posterior femora reddish where there is some yellow hair. Apex of fifth yellow with two broad black bands, their tibiae sternite with row of backwardly directed con- reddish yellow with the broad apex black and a fine black bristles, and without other narrow brown band beyond the middle, the tiguous posterior tarsi wholly black. Hair black, short, hair or setulae. yellow on the middle tibiae except in front and HOLOTYPE.-Male, Guatalon, Sa. Ade- on the anterior tibiae beyond the basal third. laida, Guatemala, March-April, 1931 (J. Wings cinereous hyaline; third vein ending Bequaert). only slightly before the wing-tip, the apical cell narrow, the last section of the fourth vein Lauxanlldae gently curved. Halteres yellow. Abdomen dark brown with four pairs of sub- PHYSEGENUA MACQUART triangular reddish yellow spots forming broadly A study of the material in the collection interrupted fasciae on the second to fifth seg- indicates that a number of species have ments, the sides of the first segment yellow. Hair black, yellow on the sides of the first seg- been confused under the name vittata ment and on the yellow spots. Ovipositor black, Macquart. I have retained the name 1942] Curran, American Diptera 67 vittata for one of the species, but it is pos- vittata, but it is closer to it than the other sible that examination of the type may species. prove that the species is not in the collection. Physegenua ferruginea Schiner The following key will separate the SCHINER, 1868, Novara, p. 277. forms before me. Ferruginous, the front wNith three black spots, mesonotum with two or four obscure, TABLE OF SPECIES narrow, darker vittae; mesopleura with a 1.-Face with black markings ...... 2. subrectangular, opaque black spot below Face without black markings ...... 6. the bristle; apical abdominal segnment with 2.-Face with a median black vitta below. . median black spot; anterior tibiae and

...... 3. tarsi, an incomplete preapical band on the Face without a median black vitta front femora and the posterior tibiae black- (Brazil, Panama) . vittata Macquart. ish, the middle tibiae mostly brownish. 3.-Sternopleura with broad blackish stripe A series of both sexes from Brazil. or mostly blackish ...... 4. It is possible that more than one species Sternopleura wholly pale; mesopleura will agree with the original description. with moderately large opaque black The opaque black spot on the pleura usually spot behind (West Indies) ...... partly surrounds the bristle, and usually ...obscuripennis...... Bigot. only the dorsocentral vittae are distinct. 4.-Front with a slender brown median They are rarely brownish. vitta continuous with the facial vitta Physegenua banksi, new species (Brazil) ... lineata, n. sp. Front without median vitta ...... 5. Agrees with ferruginea but has only a faint indication of a small opaque spot below the meso- 5.-Mesopleura with a large, opaque black pleural bristle and well separated from it. spot below the bristle (Central Length, about 4.5 mm. America) ...... centralis, n. sp. TYPEs.-Holotype, male, and allotype, Mesopleura with a very weak opaque female, Frijoles, Canal Zone, July 10, 1924 blackish spot (South America) ...... (N. Banks), in Museum of Comparative ...... eronis, n. sp. Zoology. 6.-Mesopleura with opaque black spot Unfortunately the male abdomen is adjacent to the bristle (Brazil) ...... damaged and it is not advisable to spread ...... ferruginea Schiner. the genitalia which are no doubt distinct Mesopleura without distinct spot from ferruginea. This may be concluded (Panama) ...... banksi, n. sp. from the fact that in the vittata group the absence of the black pleural spot is ac- Physegenua vittata Macquart companied by genitalic differences. MACQUART, 1847, Dipt. Exot., Suppl. III, p. 60 (f.). Physegenua lineata, new species This species has apparently not been Ferruginous, front yellowish with median correctly identified since it was described. brown vitta. Length, 4 to 5 mm. Macquart makes no mention of a median MALE.-Front yellowish, the ocellar triangle and a spot at the base of the anterior bristles black stripe on the face, a character pos- black; a linear brown line extends from the sessed by all the forms previously identi- ocelli, passes between the antennae and joins fied as vittata. There is a specimen collected the median black facial vitta. Occiput reddish by Nathan Banks at Canal yellow; a broad black stripe extends over the Frijoles, Zone, sides of the face from the eye to the groove just July 10, 1924, that agrees with the descrip- above the oral margin. Face, polished, trans- tion, and I am considering that it is vittata. lucent, the vitta moderately broad. Antennae The black stripes on the sides of the face reddish, the third segment usually darkened are broader than in allied and the apically; arista short plumose. Palpi yellowish. forms, Thorax with a pair of blackish or dark ferrugin- mesonotal vittae are stronger and united ous vittae along the dorsocentral lines; pleura posteriorly where they cover the disc of with broad brown stripe along the upper margin the scutellum. It is possible that this is not in front of the wings, the sternopleura almost all 68 Bulletin American Mu-seum of Natural History [V-ol. LXXX blackish or brownish, and there is a large brown Lauxania variegata LoEw, 1861, Cent., I, No. spot in front of the halteres. Free border of the 83. scutellum yellowish. There is just a trace of an This species, which resembles eronis, is opaque spot on the mesopleura. Legs reddish yellow; anterior tibiae and tarsi readily recognized by the wholly pale brown to black, their femora with a preapical sternopleura. It is the only species seen black band. from the West Indies and is well repre- Wings brownish in front, broadly so apically. sented in the collection. all Squamae with brownish border and fringe. Apparently Halteres reddish yellow. West Indian records of vittata refer to Abdomen with a black spot on the apex of the obscuripennis. sixth segment, and the ventral surface in large part black or brown. PSEUDOGRIPHONEURA HENDEL FEMALE.-Differs only sexually. This genus is well represented in the TYPEs.-Holotype, male, and allotype, American tropics. It is close to Xeno- female, Corumba, Brazil (Williston Collec- chaetina Malloch but lacks setulae on the tion). Paratypes, three females, Corumba, second vein. It is readily separated from and four males and seven females, Bauru, Neogriphoneura by the presence of only two Sao Paulo, December 4, 1919 (Cornell Uni- strong dorsocentrals, rarely a third weak versity Expedition). one as in Pseudocaliope. There is only one This species is easily recognized by the sternopleural bristle. vittate front. The following key includes the species known to me, but there may be others de- Physegenua eronis, new species scribed. Ferruginous, with black and brown markings. Length, 4 to 4.5 mm. TABLE OF SPECIES This species is similar in color to the preceding but lacks the brown line on the front. The facial 1.-Legs wholly pale, yellowish or whitish vitta is about five times as long as wide. In the ...... 30. female the anterior and ventral edges of the Legs in part black or brown, at least sternopleura are pale. the front tarsi brownish ...... 2. TYPEs.-Holotype, male, allotype, fe- 2.-Mesonotum with two or more vittae, male, and paratypes, two males and three if with median and lateral vittae, females, Brazil (Williston Collection), male the median one rarely weak, the and three females, British Guiana, and one vittae sometimes formed of rows of female from Paramaribo, Surinam, in the spots ...... 3. Cornell University Collection. Mesonotum not vittate...... 14. 3.-Mesonotum grayish, with two or four Physegenua centralis, new species longitudinal brown vittae ...... 4. Ferruginous, with black and brown markings. Mesonotum differently colored.... 8. Length, 4.5 mm. MALE.-Similar to lineata but without the 4.-Apices of abdominal segments broadly median frontal vitta; mesopleura with a moder- whitish pollinose ...... 5. ately large opaque black spot well below the Abdomen without white pollinose bristle; and the black on the sternopleura is fasciae...... 6. much less extensive, leaving the broad anterior border reddish. The black stripe on the side of 5.-Abdominal segments broadly reddish the face extends little more than half way to the yellow apically (Central and South oral margin. America) ...... variata Hendel. TYPEs.-Holotype, male, British Hon- Abdominal segments wholly dark in duras, November, 1928; allotype, female, ground color, the apices broadly Penas, Bauderas Bay, Mexico, November silvery gray pollinose (Venezuela). 15, 1937. Paratype, male, La Ceiba, ...... cinerellaHendel. Honduras, October 10, 1916. 6.-Wings cinereous hyaline (Costa Rica, Panama) .... hyalipennis Malloch. Physegenua obscuripennis Bigot Wings partly brownish ...... 7. Siomyza obscuripennis BIGOT, 1857, in Sagra, 7.-Wings strongly tinged with brown, Hist. Nat. Cuba, VII, p. 826. darkened in front, a pale median .1942] Curran, American Diptera 69

vitta on the basal-half (Peru) ...... Disc of basal segments and their ....melanoptera...... Hendel. bases yellowish red, strongly con- Wings brown along the costa and trasting with the blackish areas apically, cinereous posteriorly (Panama) .... zeteki, n. sp. (Brazil) ...... cormoptera Hendel. 19.-Middle tibiae black on apical two- 8.-Mesonotum shining rusty reddish, thirds...... 20. witb six shining brown vittae Middle tibiae with the apical fifth (Guiana) ...... diversa Curran. black (Panama).... siesta, n. sp. Thorax differently colored ...... 9. 20.-Front shining black, the anterior 9.-Mesonotum opaque black, with four border yellowish (Panama) ...... white vittae, the median ones ...... l...... Iutzi,n. sp. bordering the scutellum (West Front opaque, grayish (Panama)....

...... n. sp. Indies) ...... albovittata Loew. .rana, Mesonotum differently colored .... 10. 21.-Middle of face shining black ...... 22. 10.-Mesonotum with three brown or Middle wholly pollinose ...... 28. cinereous vittae ...... 11. 22.-Front with opaque black markings. . . Mesonotum with numerous brown ...... 23. spots forming lines (Puerto Rico) Front without black markings (Flor- vittifacies Curran. ida) ...... incongrua Malloch. 11.-Tibiae pale yellow with the base and 23.-Front with a rather quadrate opaque

apex black ...... 12. black marking on anterior half or Tibiae brown with the base obscurely more ...... 26.

reddish or with yellow band .. 13. The opaque black marking is wholly 12.-The brown median vitta extends over or almost wholly divided by the the disc of the scutellum (Costa frontal triangle ...... 24. of Rica)...... marmorata Malloch. 24.-Opaque black markings in the form The scutellum is wholly cinereous two triangles, the orbits polished

(Guiana) . .... claripennis Curran. (eastern United States) ......

13.-Mesonotum cinereous, with median ...... intermediaMalloch. and lateral brown vittae (Florida). . The opaque triangles are separated

...... cineracea Coquillett. by a fine line anteriorly, the orbits Mesonotum brown with narrow cine- not wholly shining ...... 25.

reous vittae ...... altera, n. sp. 25.-The opaque blackish spot along the 14.-Tibiae and tarsi almost all whitish... orbits separated from the median

..... 21. black spots (Texas)...... Anterior tibiae brownish or their tarsi ...... signatifronsCoquillett.

all brown .... 15. The opaque black spots contiguous, 15.-Mesonotum shining ferruginous or the orbits shining only at the bases reddish yellow; femora blackish of the bristles (eastern United (Guiana) .....luteipennis Curran. States) ...... scutellata Malloch. Mesonotum blackish in ground color 26.-The opaque black spot is separated ...... 16. from the anterior ocellus by more 16.-Base of wing not or scarcely dark- than the length of the ocellar tri-

ened ...... 18. angle ...... 27. Base of wing broadly blackish.1....17. The opaque spot is much less widely 17.-Thorax opaque blackish (Panama).. separated from the anterior ocellus

...... aries, n. sp. (eastern United States and Canada)

Thorax subshining blackish brown or ...... gracilipesLoew. ferruginous (Guiana)...... 27.-Middle of parafrontal strongly acicu-

...... nigra Curran. late or weakly striate (Florida) ....

18.-Abdomen almost uniformly colored, ...... Joridensis,n. sp. paler areas not strongly contrast- Parafrontal smoothly polished 19. ing...... (Panama)...... proana, n. sp. 70 Bulletin American Museum of Natural History (Vol. LXXX

28.-Polished parafrontal stripe not inter- markings; thorax with narrow cinereous vittae. rupted between the Length, 4.25 mm. bristles MALE.-Head a reddish n. black, triangle on each (Panama)...... anora, sp. side of the anterior border of the front. Front a Polished parafrontal stripe distinctly little wider than long, brownish gray, the vertex interrupted between the bristles. . . and parafrontal stripes more ochraceous, the ...... 29. latter with a narrow brownish stripe above, the anterior angles whitish; above the white spot 29.-Mesonotum with rather thin cinereous an opaque black triangle that extends more than pollen (Panama) ....ludens, n. sp. half way to the vertex along each orbit. Anterior Mesonotum with thick ochraceous frontal much shorter than the posterior; ocellars pollen (Panama)... aurescens, n. sp. long. Occiput grayish brown, the orbits whitish below. Face cinereous pollinose, with a pair of 30.-Mesonotum blackish in ground color transverse, dorsally convex spots just below the ...... 31. middle and a large orbital triangle above opaque Mesonotum pale, rarely with brown brown. Basal antennal segments black, the vittae...... 34. third missing. Palpi black. Thorax brownish, with brown 31.-Mesonotum black...... pollen, the shining 32. mesonotum grayish behind and with three nar- Mesonotum thickly pollinose .. .. 33. row gray vittae on the posterior three-fourths 32.-Median frontal vitta shining black, and indications of an interrupted lateral stripe, the orbits opaque (Costa Rica) .... inside of which is a more blackish stripe; pleura with a cinereous area in front and a ...... lateralis large smaller Malloch. one below the squamae. Scutellum very gently Median frontal vitta mostly opaque convex, brown pollinose. black (Costa Rica)...... Legs blackish; tibiae with a narrow, subbasal ...... aliena...... Malloch. pale yellow band; basal segments of the pos- 33.-Base of scutellum terior four tarsi dull yellowish. very broadly viola- Wings with luteous tinge, the veins luteous. ceous (Panama) ...... viola, n. sp. Squamae brown. Halteres yellow. Scutellum wholly grayish (Panama). . Abdomen shining brown, the genitalia of ...... pallipes Malloch. medium size. 34.-Mesonotum with three or five weak HOLOTYPE.-Male (Williston Collec- brownish vittae (Brazil, Colombia) tion). The specimen is probably from ...... elegantula Frey. Brazil. Mesonotum without distinct vittae...... 35.Pseudogriphoneura aries, new species 35.-Face with a black median spot below Black, the abdomen with reddish markings; (W¶Test Indies)....anomala Curran. thorax opaque; legs partly yellow; base of wings Face without black spot below.... 36. black. Length, 3 mm. MALE.-Head black; front wider than long 36.-Scutellum wholly pale...... 38. dull black, a median linear triangle in front of Scutellum with one or two black spots the ocelli and the front border rather reddish, the apically...... 37. anterior border with cinereous pollen laterally; 37.-Scutellum with two black spots bristles arising from bare spots; anterior orbitals much smaller than the posterior; ocellars mod- apically (Panama).lina, n. sp. erately long. Lower occipital and the facial Scutellum with a single black spot orbits whitish pollinose; face dull brownish (Brazil) ...... willistoni, n. sp. black, with slight grayish sheen above in some 38.-Antennae mostly deep black...... 39. lights. Antennae blackish, the third segment reddish basally; arista long plumose. Palpi Antennae mostly reddish, the third black. segment partly reddish brown Thorax opaque black, the mesonotum with (Brazil)...... plana, n. sp. brown tinge in some lights and slightly slaty 39.-Antennae all black (Panama)...... green laterally, with a pair of very obscure darker vittae, the pleura more brownish white pollinose ...... celeste, n. sp. in front. Scutellum slightly convex, with brown Third antennal segment reddish on pollen apically. basal half (Panama)...... Legs blackish; basal three tarsal segments ...... picticornis, n. sp. yellowish; posterior four tibiae yellowish on apical half or more. Pseudogriphoneura Wings cinereous hyaline, the base broadly altera, new species black. Squamae brown. Halteres yellow. Dark brownish; face and front with opaque Abdomen black, the disc reddish, the apices 1942] Curran, American Diptera 71 of the third and following segments reddish pollinose, as wide as long, narrowing anteriorly; yellow. Genitalia small reddish yellow. Abdo- anterior orbitals much shorter than the posterior; men with brownish pollen, the sides of the third ocellars weak; verticals long and strong. Oc- and fourth segments silvery white. ciput brown pollinose. Face gently receding, scarcely convex, the pollen brownish gray, ci- HOLOTYPE.-Male, Barro Colorado nereous along the orbits. Antennae black; arista Island, Canal Zone, January 19, 1929 very short plumose. Palpi black. The anterior (Curran). border of the front is reddish yellow. This species is much smaller than nigra Thorax shining black, brown pollinose; humeral area dark ferruginous. Scutellum gently Curran and is very much darker. P. nigra convex. is somewhat shining and more brownish. Legs black; bases of posterior four tibiae reddish yellow, their tarsi somewhat reddish zeteki, new species basally; front tibiae conspicuously swollen and Pseudogriphoneura appressed. Blackish, the abdomen partly reddish, the legs Wings cinereous with luteous tinge, the veins partly yellow. Length, 3 to 4 mm. luteous. Squamae brownish. Halteres yellow. MALE.-Front and upper half of the occiput Abdomen shining black with some castaneous black, the face and lower half of the occiput areas on the disc. Genitalia small, black. mostly reddish brown to brownish red. Front higher than wide, opaque, thinly brownish ci- HOLOTYPE.-Male, El Volcan, Chiriqui, nereous pollinose, a subshining stripe connecting Panama, February 22, 1936 (F. E. Lutz). the orbitals, the upper orbits broadly brown; has paired castaneous anterior orbital much shorter than the posterior; The abdomen ocellars short. Occiput with white pollen along spots on the intermediate segments, the the lower orbits. Face cinereous pollinose, very spots basal and rounded behind. The pale slightly convex. Antennae brownish red, the areas probably vary in extent. arista long plumose. Palpi black. Thorax brownish, the mesonotum black except laterally, brown pollinose, the pleura with Pseudogriphoneura siesta, new species hair some reddish areas, the propleura yellow; Ferruginous and black, the legs partly yellow- black. Scutellum black, brown pollinose. ish. Length, 3.5 mm. Legs black, the apices of the femora reddish bases MxLLE.-Head dark brown; front shining, with or yellowish; middle tibiae and tarsi, the brown pollen, slightly wider than long; anterior and apices of the hind tibiae and the basal four tibiae orbital much shorter than the posterior, ocellars tarsal segments yellowish, the posterior minute; anterior border more or less reddish. mostly brownish yellow or red, the apical seg- Face and occiput brown pollinose, the orbits ments of the front tarsi rather reddish; tro- Front cinereous; face scarcely convex. Antennae chanters and posterior four coxae reddish. black, the base of the third segment reddish; tibiae not swollen. arista moderately plumose. Wings cinereous, the veins luteous. Squamae dark ferruginous, the pleura becoming with browni border and fringe, the halteres yellow. Thorax reddish yel- dark below; pollen brown; scutellum gently Abdomen black with reddish and convex. low marlings; disc of second to fourth segments posterior four tarsi yellowish, with a broad Legs black; broadly reddish, the fifth and sixth with dark apex; middle tibiae yellowish'with the median yellow vitta, the bases of the third to broad- apical fifth black; posterior tibiae yellowish sixth segments pale fasciate, the bands with the basal fourth black and an incomplete ened and white pollinose laterally and usually ante- sixth subbasal brown band on the ventral half; connected on the fourth to segments. rior tibiae rather strongly swollen. Venter reddish yellow. Genitalia large, mostly Wings with a slight brown tinge, the veins reddish. luteous. Squamae with brown border and fringe. FEMAiLE.-The front slightly wider, almost as Halteres yellow. wide as long; all the tarsi yellowish. Abdomen castaneous with the apices of the seg- TYPES.-Holotype, male, allotype, fe- ments black, or practically all black. male, and paratypes, three males and five FEMALE.-The front tibiae and tarsi are females, Barro Colorado, Canal Zone, slightly less swollen. December 21, 1928, to February 14, 1929 TYPEs.-Holotype, male, allotype, fe- (Curran), the holotype and allotype, male, Barro Colorado, February 14, 1929; December 22 and January 7, respectively. paratype, male, Corozal, Canal Zone, Janu- ary 21, 1929 (Curran). Pseudogriphoneura lutzi, new species Shining blackish; face pollinose; legs partly Pseudogriphoneura rana, new species yellowish. Length, 3.25 mm. MALE.-Head black, the front obscurely Black, tibiae pale basally; wings mostly LXXX 72 Bulletin American Museum of Natural History [Vol. resembles brownish; third antennal segment with reddish While this species closely basal spot above. Length, 4 to 4.25 mm. gracilipes Loew in color it may be readily FEMALE.-Head black, front longer than wide, by the smaller opaque black through distinguished gray pollinose, a greenish tinge showing spot on the frontal vitta, the roughened the pollen; vertex and parafrontal lines brown- geni- ish, the bristles arising from bare spots; anterior parafrontals and the very different frontals much smaller than the posterior; talia. ocellars short. Lower occiput and face thickly cinereous white pollinose. Antennae black, the new species third segment with a conspicuous reddish spot Pseudogriphoneura proana, above at the base; arista long plumose. Palpi Shining black; tibiae and tarsi whitish; tho- black. rax with bicolored pollen. Length, 3.5 mm. Thorax black; mesonotum with a broad MALE-Head black; front a little wider band of pale brownish pollen in the middle, ex- than long, shining, the anterior border white tending over the scutellum, bordered with pollinose laterally, the anterior three-fifths of grayish, outside of which there is a broad, almost the frontal vitta opaque black; anterior orbital bare, shining stripe, the lateral margins whitish. much smaller than the posterior; ocellars very Pleura with cinereous pollen, but there is a broad small. Occiput dull black, the orbits cinereous brown stripe extending the whole length near below. Face very gently convex, the orbits the upper margin. broadly white. Antennae reddish, the third Legs black; tips of the femora, basal fifth or segment elongate, mostly black; arista long less of the tibiae and the basal two tarsal seg- plumose. Palpi black. ments yellowish, the tarsi brown apically. Thorax black, with brown pollen; a broad Abdomen shining black or dark castaneous, band of whitish pollen extends across the middle obscurely brownish pollinose. of the pleura and onto the mesonotum in front TYPEs.-Holotype, female, and para- of the suture, and the posterior border of the Canal mesonotuin is broadly einereous on the sides. type, female, Barro Colorado Island, Scutellum brown pollinose, the base broadly Zone, February 14 and January 11, 1929 whitish. (Curran). Coxae and femora black; trochanters and apices of the femora reddish yellow; tibiae and floridensis, new tarsi whitish, the tarsi yellowish apically; front Pseudogriphonleura tibiae swollen, of almost uniform width except species on the strongly narrowed basal fifth. Black, tibiae yellow; front with opaque black Wings with luteous tinge and with luteous spot. Length, 3.5 mm. veinls. Squainae brownish. Halteres yellow. MALE.-FroInt shining, the anterior border Abdomeni shiniing black, thinly brown pol- opaque black, the median vitta opaque black on linose, the genitalia rather small. more than the anterior half, the dull area trans- FEMALE. Front tibiae slender. verse above and widely separated from the an- TYPEs.-Holotype, male, allotype, fe- terior ocellus; parafrontals with distinct, fine Barro Colorado ridges on the median half, anterior orbital much male, and paratype, male, shorter than the posterior. Ocellars short. Island, Canal Zone, January 7, 1929 Occiput brown pollinose below. Face shining (Curran). black, the orbits broadly white. Antennae This species is very similar in appear- reddish, the third segment mostly brown; are arista long plumose. Palpi black. ance to floridlensis, lut the parafrontals Thorax black, with brown pollen; a broad smooth, the front tibiae of the male are band of whitish pollen extends acioss the pleura more uniformly swollen, and the genitalia and well onto the mesonotum in front of the are quite suture, while the broad posterior border of the dlifferent. mesonotum bears cinereous or yellowish tinged pollen, the pollen on the apical portion of the Pseudogriphoneura anora, new species flat, dull black scutellum brown. Black, th1C abdomnern inostly castaneous or Coxae and femora black, the tips of the femora, yellow. Length, 3.5 the tarsi some- dark ferruginous; tibia.e the tibiae and tarsi pale yellow, to 4 imIil. what darkened apically. Tibiae all slender. a little wider the veins yellowish. MALE.-Head black, the front Wings with luteous tinge, than long, dull, thiiinly gray pollinose, a broad Squamae and halteres reddish yellow. of the with luteous parafrontal stripe including the bases Wings with luteous tinge and orbitals, the arnterior oIrbital strong; ocellars veins. Squamae brownish. Halteres yellow. polli- brownish short. Posterior orbits and face cinereous Abdomen shining black, with thin nose; face very slightly cornvex. Antennae red- pollen. Genitalia of moderate size. dish browrn, the base of the third seginent paler; HOLOTYPE.-Male, Jacksonville, Florida, arista long plumose. Palpi black. March, 1911. Thorax black, rather thickly cinereous polli- 19421 Curran, American Diptera 73 nose but still somewhat shining; scutellum densely ochraceous pollinose, somewhat brassy flattened. or weakly golden. Legs black, the tibiae and basal three tarsal Legs black, the tibiae reddish yellow; tarsi segments yellow; front tibiae a little swollen. yellow. Hair black, yellow anteriorly, on the front tibiae Wings with luteous tinge, the veins luteous. and tarsi. Squamae brown. Halteres yellow. Wings with brown tinge, the veins luteous. Abdomen black, with brassy ochraceous pollen Squamae brown. Halteres reddish or yellowish. except basally. Abdomen ferruginous to castaneous, the tips of the segments blackish. Genitalia brownv, of HOLOTYPnE.-Female, Barro Colorado medium size. Island, Canal Zone, December 24, 1928 FEMALE.-Differs only sexually. (Curran). TYPES.-Holotype, male, allotype, female, paratypes, male and female, Corozal, Pseudogriphoneura viola, new species Canal Zone, January 16, 1929, and one Mostly reddish yellow, the mesonotum black; male, January 22, 1929 (Curran). legs wholly pale. Length, 3.5 mm. MALE.-Front conspicuously longer than wide, black and reddish in ground color, with Pseudogriphoneura ludens, new species cinereous pollen and appearing slate colored; anterior orbital shorter than posterior; ocellars Black, tibiae and tarsi yellow; wings brown- short. Occiput dark, thickly grayish white ish. Length, 4 to 4.5 mm. pollinose. Cheeks brown except along the MALEC.-Head black, cinereous pollinose, the orbits. Face mostly dark in ground color, facial and posterior orbits more whitish. Front thickly whitish pollinose; gently convex. An- longer than wide, appearing gray from frontal tennae yellowish; arista brown, long plumose. view; anterior orbital shorter than posterior; Palpi brown. ocellars short. Occiput with brownish pollen. Thorax yellowish, the mesonotum brown; a Face very gently convex. Antennae brownish, middle the basal segments and base of the third reddish broad median brownish band across the a the below; arista short plumose. Palpi black. of the pleura and large brownish spot below squamae; mesonotum brown pollinose, the Thorax black, cinereous white pollinose, the median fourth densely cinereous pollinose except mesonotum less thickly so and with brownish just before the scutellum; scutellum cinereous tinge on the disc. Scutellum flat, slate-gray, with the base broadly brownish in the middle. with the base broadly violaceous brown except Legs black, the tibiae and basal three tarsal on the sides. Legs yellow, anterior tibiae and tarsi with segments yellow; anterior tibiae and first tarsal white hair. segment yellow haired in front. Wings with luteous tinge and with luteous Wings brownish, pale basally. Squamae veins. Squamae with brown border and fringe. brown. Halteres yellow. Halteres yellow. Abdomen shining brown or castaneous, with Abdomen yellowish red; genitalia rather large. narrow dark median vitta and sides. Genitalia small. There is thin brown pollen and stains that may indicate that there are sometimes brownish TYPEs.-Holotype, male, and paratype, bands. male, Barro Colorado Island, Canal Zone, HOLOTYPE.-Male, Barro Colorado January 7 and February 21, 1929 (Curran). Island, Canal Zone, December 22, 1928 (Curran). Pseudogriphoneura aurescens, new species Pseudogriphoneura lina, new species Black, the mesonotum with bright ochraceous Rusty reddish; legs wholly pale; scutellum pollen; tibiae reddish yellow. Length, 4.5 mm. with black spot on each side. Length, 3 to 3.5 FEMALE.-Head blackish. Front about as mm. wide as long, with ochraceous pollen, but there MALE.-Front a little wider than long, dull is a broad brownish band just before the reddish rusty reddish, with paler median stripe and an- anterior margin; anterior orbital much shorter terior margin; anterior orbital much shorter than the posterior; no shining parafrontal than the posterior; ocellars short; the orbitals stripes; ocellars minute. Occiput brown polli- arise from shining spots. Occipital and facial nose, the orbits more whitish below. Face gray orbits with white pollen; face slightly convex, pollinose, the orbits whitish on reddish ground, with brownish yellow pollen; parafacials marked not convex. Antennae blackish, the third seg- below by a brown line along the inner margin. ment brownish red; arista short plumose. Antennae reddish, the third segment darker Palpi black. on the upper half; arista black, long plumose. Thorax black, dark brown pollinose, the Palpi reddish brown. mesonoturn and scutellum, except on the sides, Thorax rusty reddish, with brownish gray 74 Bulletin American Museum of Natural History [Vol. LXxx pollen above and cinereous pollen on the pleura; Squamae with brownish border and fringe. scutellum flattened, dull reddish, a dull blackish Halteres reddish yellow. spot below and surrounding each apical bristle. Abdomen rusty reddish yellow. In the speci- Legs reddish yellow; front tibiae not swollen. men before me it is variously stained with Wings with luteous tinge, the veins luteous. blackish or brown, but it seems likely that the Squamae brownish. Halteres reddish yellow. abdomen is normally pale, without dark mark- Abdomen pale rusty reddish, the apex paler. ings. Genitalia small. HOLOTYPE.-Female, France Field FEMALE.-Differs only sexually. Canal Zone, January 18, 1929 (Curran). TYPEs.-Holotype, male, and allotype, female, Barro Colorado Island, Canal Zone, February 13, 1929; paratypes, seven males Pseudogriphoneura picticornis, new and eight females, Barro Colorado Island, species December 22, 1928, to 1929 Rusty reddish yellow, antennae black and February 21, orange; legs yellowish. Length, 4 mm. (Curran). MALE.-Head rusty reddish, with brownish yellow pollen. Front about as long as wide, Pseudogriphoneura willistonii new with shining vitta extending from the anterior species orbital to the vertical bristles; anterior orbital shorter than the posterior; ocellars short. Rusty reddish yellow, the scutellum with a Antennae dull black, the basal half of the third small apical blackish spot. Length, 4 to 4.25 segment dull orange; arista short plumose. mm. Palpi black, reddish below on more than the MALE.-Front a little longer than wide, basal half. yellowish red, usually somewhat golden about Thorax rusty reddish, slightly shining, the the ocelli; anterior orbital not much shorter dorsum with grayish brown pollen, the pleura than the posterior; ocellars moderately long. more cinereous, seutellum flat, brownish yellow Occiput and face cinereous white pollinose, the pollinose. middle of the face usually brownish in ground Legs reddish yellow, front tibiae not con- color. Antennae pale reddish; arista moder- spicuously swollen. ately long plumose. Palpi reddish brown. Wings einereous hyaline; veins dark luteous. Thorax rusty reddish yellow; not shining; Squamae with brown border and fringe. Hal- mesonotum with dull yellowish red pollen, the teres reddish yellow. pleura white pollinose. Scutellum flattened, the Abdomen shining rusty reddish, obscurely upper surface mostly reddish brown, or at least pollinose, the apices of the segments narrowly darker than the border; the apex with a small brownish; a narrow brown mnedian vitta extends blackish spot below that is carried narrowly up- from the base of the second segment to the ward to join the dark disc. apex. The tip of the abdomnen is partly con- Legs reddish yellow. cealed, but the specimen appears to be a male. Wings with luteous tinge, the veins yellowish. The genitalia are small. Abdomen with cinereous pollen having violaceous tinge, and with a narrow, sometimes weak, HOLOTYPE.-Male, Barro Colorado median brown vitta on the third and following Island, Canal Zone, December 22, 1928 segments. Genitalia small. (Curran). TYPEs.-Holotype, male, and paratypes, three males, Chapada, Brazil (Willi- ston Collection). Pseudogriphoneura plana, new species Rusty reddislh yellow; a third, weak dorsocentral. Length, 3.25 mrim. Pseudogriphoneura celeste, new species MALE.-Front about as long as wide, dull Rusty reddish yellow; face pollinose; anten- rusty reddish yellow, the vertex and frontal nae wholly black. Length, 3 mm. stripes paler; an obscure, mediani pale vitta in FEMALE. Front shining, a little wider than front of the ocelli; orbitals of almost equal long, the anterior border yellowish; anterior length; oeellars moderately long. Occiput and orbital about half as long as the posterior; ocel- face with yellowish pollen. Antennae reddish, lars moderately long. Occiput and facial orbits the third segment more or less browrnish above with whitish pollen; face yellowish, scarcely except basally; arista long pubescent. Palpi convex. Antennae wholly black. Palpi black reddish. with reddish base. Thorax with brownish yellow pollen above Thorax rusty reddish, the pleura paler and and thin whitish pollen on the pleura. Scutel- with thin yellowish pollen. Scutellum flattened. lum flattened. Legs reddish yellow, the hair all black; tarsi Legs reddish yellow. darkened apically. Wings with luteous tinge and darker veins. Wings cinereous hyaline; veins luteous. Squamae and halteres yellowish. 1942] Curran, American Diptera 75

Abdomen with brownish yellow pollen, the TABLE OF SPECIES apices of the segments narrowly brownish, and there is a trace of a slender median brown 1.-A large black spot on the posterior of vitta. Genitalia of moderate size. mesonotum and base of scutellum... TYPEs.-Holotype, male, Rio de Janeiro, ...... timida, n. sp. and paratypes, two males, Chapada, Brazil No black on the thorax...... 2. (Williston Collection). 2.-Front with a slender median brown The paratypes are slightly teneral. This stripe...... striga, n. sp. species possibly does not belong in this Front without dark stripe ...... 3. genus, but it seems to fit here better than 3.-Crossveins brown and distinctly in any other described. clouded ...... tertia, n. sp. Crossveins yellowish...... T HAAEONIA, NEW GENUS ...... sordida W iedemann Related to Xenochaetina Malloch. Face shining, convex, abruptly receding below; front Neogriphoneura timida, new species longer than wide, gently convex; anterior or- Length, 4 to 4.5 mm. bitals almost as strong as the posterior, not or MALE.-Front pale rusty reddish yellow to scarcely convergent, both moderately short; yellow, shining, the ocellar triangle brown; ocellars small, reclinate. Two pairs of dorso- ocellars absent; occiput and face yellowish, the centrals, no intra-alar, one sternopleural. Legs, face sometimes mostly whitish, the orbits abdomen and wings as in Xenochaetina. with white pollen. Antennae pale orange, the GENOTYPE.-Physogenua nigra Williston. arista black. Palpi yellowish. Thorax yellow, rusty reddish yellow above; The single specimen of this species be- mesonotum with a black or brown median fore me is one of the cotypes from St. stripe beginning well behind the front margin Vincent. It is very similar to Xenochaetina and usually tapering posteriorly, and a large black posterior spot that extends well onto flavipennis Fabricius, but the anterior the scutellum and is rather orbicular in shape. orbital bristles are a little longer, and the Scutellum flat, yellow. ocellar bristles are reclinate instead of Legs reddish yellow. proclinate. There are two small bristles Wings with luteous tinge, the veins yellow. Squamae with brown fringe. Halteres yellow- before the furcation of the second and third ish. veins. In Physegenua the anterior orbital Abdomen yellowish, the upper surface mostly bristles are more hair-like and strongly rusty reddish yellow. Genitalia small. convergent, the oral margin is not sepa- FEMALE.-Agrees except sexually. rated from the facial convexity by a groove, TYPEs.-Holotype, male, and allotype, and the wing veins are bare. female, Patilla Point, Panama, February It is possible that some of the cotypes 1, 1929, and paratypes, four males and five of P. nigra belong with flavipennis; if all females, same locality, January 15 and but the one before me do, the present speci- February 1, 1929, and one female, Corozal, men may be considered the type. Other- Canal Zone, January 22, 1929 (Curran). wise the type should be selected from material in the British Museum. Neogriphoneura striga, new species Rusty reddish yellow above, paler below, NEOGRIPHONEURA MALLOCH front with a fine median brown line. Length; 4.5 to 5 mm. This genus is characterized by the pres- MALE.-Front a little wider than long, shining ence of a single sternopleural, three strong rusty reddish yellow, more yellow in front and dorsocentrals, broad front, and the anterior behind; yellowish pollinose from anterior view; orbitals are very strong and somewhat con- ocellar triangle and an incomplete, slender median vitta brown; orbitals of about equal vergent, though inclined toward the vertex. size; ocellars absent. Occiput and face yellow- The anterior portion of the front is de- ish, with whitish pollen, the middle of the face pressed, but there is some variation in this shining, moderately convex below. Antennae All the have the arista orange; arista black, long plumose. Palpi yel- respect. species lowish. long rayed above and short rayed below. Thorax rusty yellowish, rusty reddish yellow The species before me are included in the above, the pleura with thin whitish pollen, the following key. All are rusty reddish yellow. dorsum shining; scutellum flattened. [Vol. LXXX 76 Bulletin American Museum of Natural History and they may Legs pale reddish yellow. not always well developed, Wings with slight luteous tinge; veins lute- be little stronger than in some species of and ous. Squamae with rusty brownish border Minettia. However, the wings are usually golden fringe. Halteres yellow. elongate, more obtusely rounded Abdomen yellowish, pale rusty reddish yel- more from brown low above, with yellowish gray pollen apically and have a characteristic frontal view. Genitalia small. pattern. I have seen only two species, in- FEMALE.-Differs only sexually. cluding the one described below. male, allotype, fe- TYPEs.-Holotype, OF SPECIES male, and paratypes, four females, Barro TABLE Colorado Island, Canal Zone, December 1 .-No hyaline spot in the submarginal cell...... 2. 24, 1928, to February 18, 1929 (Curran), Brazil One or more hyaline spots in the sub- and male and female, Chapada, 4. (Williston Collection). The holotype is marginal cell. very obtusely rounded apically dated February 18, 1929, the allotype, 2.-Wings 3. December 24, 1928; the other specimens ...... were taken on January 11 and February Wings somewhat pointed apically (Mexico) ...... vergens Giglio-Tos. 15 and 16. spot I can see no difference in the Brazilian 3.-Posterior crossvein with hyaline con- America, Panama) specimens, but they are not in perfect anteriorly (Southdistinctissima Schiner. dition. The front in this species is less ...... concave and less rounded in front than in No hyaline spot on the crossvein sordida, but this condition varies some- (Mexi.co). Palans Giglio-Tos. what. 4.-Only one hyaline spot in the submarginal cell...... 5. Neogriphoneura tertia, new species Three or more hyaline spots in the submarginal cell (Mexico)...... Rusty reddish yellow, paler below; crossveins Hendel. blackish. Length, 4 to 4.5 mm...... caloptera MALE.-Front conspicuously wider than 5.-A hyaline spot in the apical cell just long, moderately depressed anteriorly, shining, beyond the posterior crossvein (West the pollen brownish yellow from anterior view; Williston. anterior margin yellow; ocellars minute. Occi- Indies) .. angttstipennis put and parafacials white pollinose; face yellow- No hyaline spot neair the posterior ish. Antennae pale orange; arista black, long crossvein in the apical cell...... 6. plumose. Palpi yellowish. with rows of brown yellow above, yellowish 6.-Abdomen three Thorax rusty reddish spots (Brazil) . excepta Walker. below, shining, the pleura with some thin white pollen. Abdomen wholly yellow (Panaima)...... Legs rusty yellowish. ..banksi,n. sp...... Wings with luteous tinge; veins yellowish, the crossveins blackish and narrowly clouded. Giglio-Tos Squamae yellow, the inner part of the fringe Chaetocoelia palans brown. Halteres yellow. GIGLTO-ToS, 1895, R. Acad. Sci. Torino, Abdomen rusty reddish yellow, more yellow- XLV, p. 51 (f.). ish below and apically; cinereous pollinose from This is the type of the genus. The ab- anterior view. FEMALE.-Differs only sexually; the terminal clomen is yellow basally ard brown apically. segments are pale yellowish. Known only from Mexico. TYPEs.-Holotype, male, Patilla Point, January 15, 1929 (Curran); allo- Chaetocoelia vergens Giglio-Tos Panama, l. type, female, and paratypes, male and GIGLIo-Tos, 1895, Aca1d. Sci. Torino, female, Chapada, Brazil (Williston Collec- XLV, p. 51 (f.). tion). Hendel has placed this Cas a synonym of distinctissima Schiner, and this may be cor- CHAETOCOELIA GIGLIO-Tos rect. However, the wings alre much more in This genus is not easily separated from pointed (according to the figure) than Minettia Desvoidy because the tubercles any of the other species. If the outline of from which the frontal bristles arise are the wing is aecurately (Irawn the species 1942] Curran, American Diptera 77 may not belong in this genus. The name and basal corners. Three pairs of dorsocentrals, the intra-alar strong. should be retained until Mexican speci- Legs reddish yellow. mens are available for study. Wings brown, hyaline posteriorly and with three clear spots apically in the brown field. Squamae brown. Halteres brownish yellow. Chaetocoelia distinctissima Schiner Abdomen slightly shining yellow. Genitalia Sapromya distinctissima SCEINER, 1868, No- small. vara, p. 280. FEMALE.-Differs sexually. HENDEL, 1907, Wiener Ent. Zeitung, XXVI, p. 229 (f.). TYPEs.-Holotype, male, and allotype, "South Amer- female, Barro Colorado Island, Canal Originally described from 21 and 15, 1929 (Curran). have me a specimen col- Zone, February ica." I before Paratypes, male, Barro Colorado, Febru- lected by Nathan Banks at Fort Sherman, (Curran), and four males and Canal Zone, July 7, 1924, which agrees with ary 21, 1929 description. eight females, Barro Colorado, June 20 to the July 21, 1924 (N. Banks). This species is quite distinct from the Chaetocoelia caloptera Hendel others and is easily recognized by the yellow HENDEL, 1907, Wiener Ent. Zeitung, XXVI, abdomen. The face of the female is some- p. 229 (f.). times paler than described and more macu- Distinguished by having more clear late. The wing and profile of the head are spots in the wing than other species. illustrated in my "North American Dip- Known only from Mexico. tera," pp. 316 and 320. In the typical species the ocellar bristles are rather long. Chaetocoelia angustipennis Williston Sapromyza angustipennis WILLISTON, 1896, XENOPTERELLA MALLOCH Tr. Ent. Soc. London, p. 381 (f.). The The only known species belonging to this Known only from St. Vincent. from Mexico City. I are characteristic. genus was described wing markings have before me a species from Colorado. The two are separable as follows: Chaetocoelia excepta Walker TIrypeta excepta WALKER, 1852, Dipt. Saund- 1.-Entire costal border broadly brown, ers., p. 387 (f.). the crossveins bordered with brown Most closely related to the following ...... costalis, n. sp. species, but the abdomen bears three series Costal border not wholly brown, the of brown spots. Known only from Brazil. wing with transverse fasciae ...... obliqua Malloch. Chaetocoelia banksi, new species Thorax brown; abdomen wholly yellow; Xenopterella obliqua Malloch wings with three hyaline spots in the brown MALLOCE, 1926, Proc. U. S. N. M., LXVIII, field. Length, 3.5 to 4 mm. Art. 21, p. 3 (f.). MALE-.Head brownish or brownish red, the lower half of the occiput and the face with cinere- Rusty reddish yellow, with black and ous white pollen, the facial orbits, oral margin brown markings, the ocellar triangle (expanded in the middle) reddish or reddish black. The wing is illustrated by Malloch yellow in ground color, the face usually with a in "North American Diptera." pair of brown spots below that may be united and above. Front longer than wide, slightly widening above; anterior orbitals a little more than half Xenopterella costalis, new species as long as the posterior, usually slightly curveed crossveins inward; ocellars short; orbitals arising from Rusty reddish yellow and black; shininig spots. Antennae brownish red or dull transverse. Length, 4.5 mm. haired. MALE.-Front wider than long, blackish, the orange; arista mostly black, very short large, Palpi reddish. broad orbits and vertex reddish, a very brown, rather dull, the pleura transverse opaque black spot in front of the Thorax reddish with whitish orbitals with some cinereous pollen in front and below; ocelli, the orbits pollen; Scutel- strong, both situated on the posterior half of the mesonotum obsoletely vittate in front. above, lum rich reddish brown, with paler median line front; ocellars long. Occiput black 78 Bulletin American Museum of Natural History [Vol. LXXX yellowish and with white pollen on the lower anterior half. Front not three times as wide as half, reddish behind the vertex. Face reddish the ocellar triangle, strongly widening anteri- yellow, with an obscure brownish spot in the orly. Antennae blackish to brownish red, the middle below and a large, opaque black spot on arsista yellowish and plumose on the apical each side above, the parafacials very broad and half, blackish and pectinate on-the basal half. white pollinose. Antennae reddish; arista The carina separating the sides of the antennal black, broadened and flattened, tapering, with depressions is high and sometimes little concave rather thick short hair. Palpi reddish, with in profile on the lower half, but this varies in the black hair. specimens before me. Thorax rusty yellowish, the dorsum rusty Thorax black, rather thinly cinereous polli- reddish with almost the lateral third brownish; nose, the pleura with dense white hair, the a broad median vitta of thin whitish pollen; dorsum with appressed black hair except for a pleura with broad brownish border below the white lateral patch above the base of the wings wings and squamae. Scutellum reddish yellow, and sometimes a few white hairs at the outer flattened. Two sternopleurals; three dorso- end of the suture. There are two bare blackish centrals. spots near the middle on the anterior half of the Legs reddish yellow, the tarsi somewhat dark- pleura and a tuft of black hairs above. The ened, the front pair mostly brownish. scutellum is black haired, at most three or four Wings cinereous hyaline, the broad anterior whitish hairs being present on each side near the border and apex brown; crossveins strongly apex. bordered with brown, the border on the poste- Legs black, and with black hair, the anterior rior crossvein triangular, broad posteriorly. femora densely short white haired posteriorly, Squamae brown. Halteres yellowish. and there are some white hairs basally on the Abdomen shining black, somewhat reddish middle femora; all the femora are silvery white basally. Genitalia of medium size, reddish. pollinose below on about the basal half. HOLOTYPE.-Male, Pagosa Wings light brownish, the hair on the squa- Springs, mae all white. Colorado, June 22-24, 1919 (F. E. Lutz). Abdomen metallic steel blue or with greenish This may possibly represent a distinct reflection, the base black, the intermediate seg- genus, but I prefer to place it in Xenop- ments densely cinereous pollinose, with some terella. The genotype apparently does not large irregularly placed shining spots, with a narrow median bare vitta and the narrow apices have a strap-like arista, but this may be a of the segments incompletely shining; the apical sexual character. segment is about half shining above, the base being widely pollinose laterally and narrowly Cuterebridae so in the middle. The amount of pollen on the apical segment varies, and when the large bare The following species, reared from lar- spots are confluent with the posterior shining vae found in jack-rabbits collected in Ore- portions the segment may be more than half gon, is strikingly different from related shining. The sternites are black haired, while the inner edges of the tergites bear white hair forms, and it does not seem possible that except at the base. it can be only a color variety of princeps FEMALE.-Agrees with the male, but the front Austen. The species was included in a is about five times as wide as the ocellar triangle. small collection received from Dr. Wm. L. TYPEs.-Holotype, male, Paisley, Ore- Jellison, to whom the type specimens will be gon, larva in jack-rabbit, August 25, 1935, returned, a paratype being deposited in adult killed and pinned December 7, 1935, the collection of the Museum. collected by G. M. Kohls, reared by W. L. Jellison; allotype, female, larva, Paisley, Cuterebra jellisoni, new species Oregon, August 25, 1935, adult March 18, Black, with cinereous pollen, the abdomen with metallic blue or bluish green reflections 1936; paratype, male, larva, Paisley, Au- in some lights; intermediate abdominal seg- gust 25, 1935, adult January 4, 1936, both ments almost all cinereous pollinose; scutellum collected and reared as was the type. with black hair only. Length, 19 to 21 mm. According to Townsend's classification MALE.-Head black, densely cinereous pollinose, the front almost bare but with two orbital this species would belong to Bogeria Aus- pollinose spots below, the lower one sometimes ten, but I do not see how the genus can be connected with the spot above the outer base maintained and believe all of our species of the antennae. On the face there are three should be placed in Cuterebra. The shape bare spots, a large broad transverse one above, a small one below the eye and one half-way be- of the front, used by Townsend to separate tween this and the oral margin. Pile white, Cuterebra and Bogeria, depends greatly black on the front but mostly yellowish on the upon drying and is too variable to be of 1942] Curran, American Diptera 79 even specific value. The only character From a study of the material before me I upon which Bogeria could be maintained feel that all the above names apply to is the non-broadened male tarsi, and I do species belonging to a single genus. The not attach great importance to this. presence or absence of hairs on the para- In Volume II of the "Manual of Myi- facials is not a stable character in this ology" Townsend placed his C. lepusculi group, and the same applies to pile or weak as a synonym of Bogeria princeps Austen, bristles on the prosternum. The charac- but later, in Volume VI, he was inclined to ters by which Hystricia should be recog- believe that the three species, ruficrus Aus- nized are: the almost wholly pilose squa- ten and the two above named, might all mae and the produced oral margin, with be forms of the same species. Unless there the vibrissae situated well above it. The is great variation this seems unlikely. C. type of Hystricia (amaena Macquart) will princeps has the abdomen almost all polli- not trace out in Townsend's "Manual," nose above, while in lepusculi it is pollinose the species tracing to Hystriciopsis Town- only on the sides and apical segment. The send. The number of sternopleurals varies distinctness of ruficrus is scarcely question- from one to three and the postsutural able. In these three forms the scutellum of dorsocentrals from one to four. Since the the male is bordered below by a conspicuous dorsocentrals and acrosticals are often fine fringe of rather long white pile, a character they are subject to variation in the same that is strikingly absent in jellisoni, new species and sometimes in the same speci- species. men. For convenience in separating the species The following key separates the species comprising this group, the following key is before me: presented: TABLE OF SPECIEs TABLE OF SPECIES 1.-Parafacials with hair on upper half or 1.-Basal half or more of the femora red- more ...... 2. dish.. .. ruficrus Austen. Parafacials with at most two or three Femora black, in part densely white hairs above ...... 12. pollinose ...... 2. 2.-Femora and tibiae black, tarsi white. . 2.-Face with only two shining black spots ...... 3. below the large upper spot ...... 3. Femora practically all reddish or Face with three shining black spots yellowish, or the tibiae pale ...... 4. below.princeps Austen. 3.-Mesonotum grayish pollinose; tibiae 3.-Dorsum of the abdomen mostly free of not silvery...... condor, n. sp. pollen and shining; scutellum of Mesonotum with obscure brownish male with the free border white pollen; tibiae silvery ...... haired .. l epusculi Townsend...... albimana, n. sp. Dorsum of abdomen chiefly cinereous 4.-Abdomen orange to yellowish, some- pollinose; scutellum wholly black times shining, with black markings haired in both sexes...... 9...... jellisoni,...... n. sp. Abdomen dark reddish to black.... 5. 5.-Squamae blackish ...... 6. Tachinidae Squamae whitish .... browni, n. sp. HYSTRICIA MACQUART 6.-Abdomen with interrupted reddish MACQUART, 1843, Dipt. Exot., II, Part 3, p. 43. and dull black fasciae...... Bombyliopsis TOWNSEND, 1915, Proc. Biol...... humeralis,n. sp. Soc. Wash., XXXVIII, p. 23. Abdomen not strikingly bicolored. . 7. Engelomyia TOWNSEND, 1931, Rev. Ent., 7.-Two sternopleurals .....vultur, n. sp. I, p. 349. One Hystriciella ENGEL, 1920, Zool. Jahrb., XLIII, sternopleural...... 8. p. 311 (preoccupied). 8.-Tibiae yellow haired. flavitibia, n. sp. Hystriciopsis TOWNSEND, 1914, Ins. Ins. Tibiae black haired...... Mens.. II, p. 85...... nigrotibiata, n. sp. 80 Bulletin American Museum of Natural History [Vol. LXXX

9.-Abdomen clothed with long orange given in the key will separate it. The two hair ...... rufohirta, n. sp. basal antennal segments are pale reddish. Abdomen with some yellow pile which There is one male before me from Eslava, is short and sparse, the hairs mostly Federal District, Mexico. black on apical segments ...... 10. 10.-Black parafacial hairs limited to the Hystricia amaena Macquart upper half.... abrupta Wiedemann. MACQUART, 1843, Dipt. Exot., II, Part 3, p. Black parafacial hairs extending al- 44. most to the vibrissae ...... 11. Wulp separated this species from micans 11.-Fourth abdominal segment with broad by the color of the antennae and tarsi, both band of yellow pile basally; outer of which were said to be black. There is a forceps of male genitalia with two female before me from El Volcan, Chiriqui, spines at the apex... micans Wulp. February 22, 1936 (W. J. Gertsch), which Fourth segment without pale pile; has the second and third antennal seg- outer forceps with single curved ments black, but the tarsi are only slightly apical spine... abrupta Wiedemann. darkened. The parafacials are almost 12.-Abdomen orange with small black wholly bare, and the species is readily sepa- spots in the middle and on the sides rated from micans by this character...... amaena Macquart. This specimen agrees with Macquart's Abdomen differently colored ...... 13. original description except that he describes 13.-Abdomen black, rather shining...... the apical tarsal segments as blackish. This ...... obscuraTownsend. is the type of Hystricia. Abdomen differently colored ..... 14. 14.-Tarsi black...... 15. Tarsi yellow...... Hystricia condor, new species ...... Calohystricia gertschi, n. sp. Black, the thorax with pollen; tarsi whitish. Length, 10 to 12 mm. 15.-Abdomen shining castaneous...... MALE.-Head black, with olive gray pollen; ...... vargas, n. sp. occiput and lower cheeks whitish pilose, the Abdomen ochraceous pollinose...... hairs and bristles black. Front half as wide as ...... n. one eye, the vitta reddish brown; nine or ten laxa, sp. pairs of frontals, the ocellars long but weak, verticals cruciate, the outer verticals absent; Hystricia abrupta Wiedemnann cheeks two-thirds as wide as the eye-height. Tachina abrupta WIEDEMANN, 1830, Ausser. Face angularly concave, the oral margin as Zweifl., II, p. 293. prominent as the antennal base, parafacials slightly narrowing below, not so wide as the A very common species in the United third antennal segment, with hair on more than States and Canada and undoubtedly occur- the upper half. Antennae black, the second ring in northern Mexico. The hair on the segment partly obscure reddish, the third parafacials is usually coarse and black widest apically, the tip broadly rounded; arista black. Apical section of proboscis about as above and very fine and yellow below, but long as the eye-height; palpi long, reddish, the in some specimens the hair is all black. apex very broadly black above, the hair black. This is the type of Bombyliopsis Town- Pile of the eyes pale, tawny on the middle por- send. The only difference of note between tion. Thorax black, conspicuously olive gray polli- this species and amaena is the presence of nose, the mesonotum with four narrow, inter- more hairs on the parafacials, and I do not rupted dark vittae. Hair black, yellow on the believe the character is of generic value. sternum; dorsocentrals 3-3, acrosticals 3-2; sternopleurals 2-1, the bristles fine. Scutel- Hystricia micans Wulp lum brown pollinose, bearing numerous spines and black hair. WULP, 1888, Biol. Centr. Amer. Dipt., I, p. Legs black, narrow knees and trochanters 16 (f.). reddish, the tarsi pale yellow; coxae gray This species is rather similar to the com- pollinose, hair black, pubescence on front and mon Nearctic abrupta Wiedemann and hind tibiae, on the under surface of the first segment of the front tarsi and on the ventral particularly to those forms in which the and posterior surfaces of the hind tarsi yellow; parafacial hair is all black. The characters coxae with fine yellow hair. 1942] Curran, American Diptera 81

Wings brown, the base blackish. Squamae Hystricia browni, new species brown, with black hair; halteres brown. Blackish, the abdomen brownish red, with Abdomen blackish, with brown pollen and whitish pollen; squamae grayish white, with appearing dark castaneous, the genitalia black pile. Length, 12 mm. shining black, the hair all black. Abdomen FEMALE.-Head blackish, the cheeks in front heavily spined, the bases of the segments broadly and the face immediately below the antennae bare but bearing black hair; apex of fourth seg- reddish; pollen olive gray, cinereous on the ment bearing a row of ordinary bristles; sides of middle of the face, olive brown on the front. venter bare on basal half. Front half as wide as one eye, with two rows of FEMALE.-Front almost three-fourths as frontals and one proclinate orbital on each side, wide as eye; apical four segments of the front the inner row interrupted in the middle; ocellars tarsi broadened and pale pubescent beneath; long and fine. Occiput and lower cheeks with disc of scutellum more reddish. whitish pile. Oral margin not so prominent as TYPEs.-Holotype, male, and allotype, the antennal base; parafacials haired on their Prov- whole length. Antenna reddish, the third seg- female, Hacienda Talahua, Bolivar ment black, about twice as long as wide. Palpi ince, Ecuador, May 4 and April 26, 1939 reddish. Eyes reddish yellow pilose. (F. M. Brown). Thorax blackish, the sides of the mesonotum, This species seems to approach Engelo- scutellum and some pleural areas brownish red; mesonotum olive brown pollinose, the vittae myia Townsend but does not fully agree narrow and weak. Acrosticals 1-2, dorsocen- with the characters given for the genus. trals 3-2, one sternopleural. Hair yellow, black on the dorsum and on most of the mesopleura. Hystricia albimana, new species Scutellum with stout spines. Legs reddish, black haired, the fine hair on the Blackish; thorax brown pollinose; tibiae coxae, trochanters and under surface of the silvery; tarsi pale yellowish. Length, 12 mm. femora basally ye}low. Apical four segments MALE.-Head black, brown pollinose; face of the front tarsi conspicuously broadened. paler above, cheeks olive gray, the occiput more Wings dark brown, the base blackish. Hal- brownish yellow and with yellow pile which teres brown with reddish apex. becomes whitish below. Front one-third as Abdomen dull brick red, with cinereous white wide as eye, with about ten frontals and a second and brownish pollen. The pale pollen covers row of three opposite the antennae, the upper most of the venter and forms two broad longi- ones fine; ocellars long and fine; hair long; tudinal bands on the dorsum that broaden on outer verticals absent. Face deeply concave, the apices of the segments and converge posteri- the oral margin not so prominent as the an- orly, beginning at the base of the second segment tennal base; cheeks with sparse, coarse hair, and continuing to the middle of the fourth. the oral margin bearing some fine yellow hairs; Abdomen with spines, the tip with ordinary parafacials haired on almost the whole length. bristles; on the second segment the spines form Antennae black, the second segment partly median and lateral patches. brownish red; third segment broad, obtusely rounded apically, a little more than twice as HOLOTYPE.-Female, Minza Ridge, Vol- long as wide. Eyes with brownish pile. Palpi can Tungurahua, Ecuador, April 8, 1939 reddish, becoming black apically. (F. M. Brown). Thorax black, brown pollinose, with four weak dark vittae. Hair black. Acrosticals 1-1; dorsocentrals 2-2; one sternopleural. Scutel- Hystricia humeralis, new species lum darker than the mesonotum, with black Black, the abdomen largely reddish; tarsi pale hair, heavy spines and two or three lateral yellow. Length, 12 mm. bristles. MALE.-Head black, the pollen olive brown Legs black, the knees, trochanters and broad on the front, olive gray on the occiput and apices of the femora below reddish; tibiae dull cheeks, brown on the lower face and parafacials silvery pollinose, their pubescence pale yellow. and brownish yellow on the middle of the face. Tarsi pale yellow, the hair and bristles whitish. Front two-fifths as wide as eye; about ten pairs * Wings dark brown, the base blackish. Squa- of frontals and three in a secondary row opposite mae and halteres brown, the former with black the antennae; ocellars fine. Pile of occiput and pile. lower cheeks bright yellowish, rather golden. Abdomen blackish, the sides somewhat brown- Oral margin not so prominent as the antennal ish red, the pollen brownish. Second segment base; parafacials haired on whole length. An- spinose in the middle and laterally, the third and tennae blackish, the basal segments in part dull fourth spinose except basally, the fourth with reddish; third segment not twice so long as apical bristles. Genitalia shining black basally. wide, the apex oblique with rounded corners. Palpi reddish. Eyes with brown, yellow tipped HOLOTYPE.-Male, Hacienda Talahua, pile. Bolivar Province, Ecuador, April 29, 1939 Thorax black; humeri reddish; pollen dark (F. M. Brown). brown, on the mesopleura somewhat olivaceous; 82 Bulletin American Museum of Natural History [Vol. LXXX

vittae inconspicuous. Hair black. Scutellum Hystricia flavitibia, new species dark brown, with stout spines. Acrosticals 3-0; dorsocentrals 3-2; one sternopleural. Black, the abdomen mostly castaneous; legs Legs reddish, hair black; tarsi pale yellow and reddish. Length, 14 mm. with yellow hair and bristles. MALE.-Head black, brownish pollinose, the Wings dark brown, the base blackish. Squa- front, occiput and cheeks more olive gray; hair mae and halteres brown, the former with black black. Front one-third as wide as eye; nine or pile. ten pairs of frontals and a secondary row of five below; ocellars very fine. Occipital pile red- Abdomen black and reddish (in life the reddish yellow. Face rather deeply concave, the rish color is rather strongly coppery or violace- oral margin practically as prominent as the ous). First segment black; second dull black antennal base; parafacials rather wide, haired on the median third and with very large lateral on whole length. Antennae black, the basal black spot that extends along the posterior segments brownish red, the third widest api- border of the venter; third segment with cally, the apex obtuse. Palpi reddish basally, smaller median spot, but it is connected posteri- becoming black apically, the hair black. Eyes orly to the lateral black spot; fourth with a with brown pile. longitudinal median spot and the posterior Thorax black, thinly grayish brown pollinose, border broadly black. Hair black. Spines the vittae very obscure; hair black. Bristles numerous; median spine bunch on second seg- on disc of thorax fine; acrosticals 1-1; dorso- ment not connected with lateral spines; apical centrals 1-2; one sternopleural. Scutellum segment with ordinary bristles apically. Geni- brownish, brown pollinose; with many black talia reddish, the basal segment shining black. spines. There is a trace of pollen on the reddish areas, Legs reddish, the coxae mostly blackish or but it is not much paler than the ground color. darkened; hair and bristles black, but all the HOLOTYPE.-Male, Hacienda Talahua, hair and bristles of the tarsi and the short Bolivar Province, Ecuador, April 26, 1939 bristles and the hair on the tibiae yellowish. (F. M. Brown). Wings blackish brown. Squamae brown, with black hair. Halteres reddish. Hystricia vultur, new species Abdomen brownish red or pale castaneous, the Black, with olive gray pollen; tarsi pale yel- hair black; pollen grayish brown above, pale low; two sternopleurals. Length, 10 mm. below, visible only in some views; a broad FEMALE.-Head black, with olivaceous pol- median vitta of black that expands on the len, the middle of the face brownish. Front fourth segment. The spines on the middle of two-fifths as wide as eye, with eight to ten pairs the second segment are broadly separated of frontals in single row, and one proclinate orbi- from the lateral ones; some bristles apically tal; ocellars fine. Occiput very pale yellowish on the fourth segment. Genitalia mostly black, pilose. Cheeks less than half the eye-height. the basal segment with a number of spinose Oral margin almost as prominent as the anten- bristles. nal base; parafacials haired to below the middle. HOLOTYPE.-Male, Hacienda Talahua, Antennae black, the basal segments partly Bolivar Province, Ecuador, May 4, 1939 brownish red; third segment broad, obtusely rounded apically. Palpi reddish, with black (F. M. Brown). hair and bristles. Thorax black, rather thinly einereous polli- Hystricia nigrotibiata, new species nose, the black vittae moderately wide but Blackish, the abdomen dull castaneous, legs poorly developed. Acrosticals 3-1; dorso- reddish, the tarsi yellow. Length, 15 mm. centrals 3-2;. two sternopleurals; hair wholly MALE.-Head black, with brownish olivaceous black. Scutellum brownish, gray pollinose, pollen, the face mostly reddish brown and with with numerous black spines. brownish pollen. Front slightly more than one- Legs reddish, the coxae more or less tinged third the eye-width; twelve to fourteen pairs with brown; hair black; tarsi yellow and with of frontals and a secondary row below; ocellars wholly yellow hair and bristles. little stronger than the adjacent hairs. Pile of Wings dark brownish. Squamae reddish the occiput bright reddish yellow. Parafacials brown, with black pile. Halteres reddish wide, haired on the whole length. Face concave, brown. the oral margin not so prominent as the anten- Abdomen blackish, apparently with brown nal base. Antennae black, the basal segments pollen and two wide cinereous vittae; it has reddish, the third moderately wide, obtuse api- been moistened, and the color of the pollen cally. Palpi reddish, becoming black apically, cannot be discerned. Second to fourth segments the hair wholly black. Eyes brown pilose. with numerous spines except on the base, the Thorax dull black, with thin brown pollen fourth with many bristles apically. and black hair, the bristles on the disc of the HOLOTYPE.-Female, El Volcan, Chiri- mesonotum fine. Acrosticals, 1-1; dorsocentrals, 1-2; one sternopleural. Scutellum brown- qui, Panama, February 26, 1936 (NV. J. ish and with dark brown pollen; clothed with Gertsch). many spines. 1942] Curran, Ame,rican Diptera 83

Legs reddish, the femora more or less broadly male, same locality, April 26, 1939 (F. M. black basally, the tarsi yellow. Hair and Brown). bristles black, yellow on the tarsi. Wings blackish brown. Squamae black, with black hair. Halteres brownish red. Hystricia vargas, new species Abdomen dull castaneous, with large, median, Black; thorax grayish pollinose; abdomen united dark spots on the second and third seg- shining castaneous; tarsi black. Length, 12 ments and the apex of the fourth black; pollen mm. brownish, obscure. The median cluster of FEMALE.-Head black, cinereous pollinose. spines on the second segment is connected Front three-fifths to one-half as wide as eye; along the posterior margin with those on the with eight to ten pairs of frontals, the upper sides; fourth segment with bristles apically. three reclinate, and a secondary row of four or Genitalia black and reddish. five below, the ocellars moderately strong; HOLOTrYPE.-Male, Minza Ridge, Volcan two pairs of orbitals. Occiput and lower cheek with cinereous pile. Oral margin much less Tungurahua, Ecuador, April 8, 1939 (F. prominent than antennal base; parafacials with M. Brown). only a few short hairs above. Antennae black; third segment a little longer than the second, Hystricia rufohirta, new species obtusely rounded apically. Palpi reddish apically, blackish on the basal half, the hair wholly Blackish, the abdomen reddish with black black. Eyes with short yellowish pile. vitta, and wholly tawny pilose; legs reddish. Thorax black, rather thickly cinereous polli- Length, 10 to 11 mm. nose, the vittae distinct; hair black, pale brassy MALE.-Head blackish, olive gray pollinose, yellow on the pleura. Acrosticals 3-3; dorso- the middle of the face reddish and with grayish centrals 3-4; sternopleurals 2-1. Scutellum brown pollen. Front about one-third as wide as reddish brown, with cinereous pollen and many eye, with eight to ten pairs of frontals, the lower spines. ones weak, but there is a secondary row of five Legs brownish red, the femora with large strong ones below; ocellars poorly differentiated. blackish areas apically on under surface, tarsi Occipital pile reddish yellow, brassy below and black. Hair black, yellow on the coxae and on the lower half of the cheeks. Oral margin base of femora below. not so prominent as the antennal base; para- Wings strongly tinged with blackish brown, facials moderately wide, haired on whole length. the base darker. Squamae brownish, with Antennae reddish, the third segment black, black pile. Halteres reddish yellow. broad, obtuse apically. Palpi reddish, the hair Abdomen shining wine red or castaneous, black. Eyes with yellowish pile. wholly without pollen. Second segment almost Thorax gray pollinose laterally, brownish wholly clothed with spines that become short above, the vittae obscure. Acrosticals 1-0 or anteriorly; third segment with discal and apical 0-0; dorsocentrals 2-2; two sternopleurals; rows of spines that are connected laterally, the hair black, yellow on the pleura, scutellum and discal row sometimes partly double; fourth posterior calli. Scutellum reddish, with few segment with two discal rows of spines and with spines. apical bristles; hair black, the base of the under Legs reddish, with black hair and bristles, surface with pale yellow pile medianly. the coxae and bases of the femora on the under surface yellow pilose. TYPEs.-Holotype, female, and para- Wings very strongly tinged with blackish types, two females, Nova Teutonia, Brazil, brown. Squamae translucent grayish brown, May 27 and June 1, 3,1939 (F. Plaumann). with pale border, the pile yellow. Halteres reddish. Abdomen bright orange red, with median Hystricia laxa, new species rather narrow black vitta composed of triangles Blackish, the abdomen brown with ochrace- on the second and third segments and a stripe ous pollen above; legs black; prosternum usu- on the fourth; hair and fine bristles reddish. ally haired. Length, 11.5 to 14 mm. Second segment with median patch of spines MALE.-Head black, cinereous yellow polli- connected along the posterior border with four nose, the front more grayish. Front one-sixth or five forming a lateral patch: third and fourth to one-fifth as wide as eye; eleven to fourteen segments with discal and apical rows of spines, pairs of frontals and a secondary row below; the third segment with two or three additional ocellars moderately long and strong. Pile of ones on the middle. Genitalia wholly reddish. occiput and lower cheeks pale yellowish. Oral FEMALE.-Front half as wide as eye; anterior margin considerably less prominent than the tarsi strongly broadened; genitalia mostly antennal base; parafacials bare. Antennae shining black. black, the third segment obtuse apically, about as long as the second. Palpi brown with some- TYPEs.-Holotype, male, and allotype, what reddish apex. female, Hacienda Talahua, Bolivar Prov- Thorax black, with yellowish brown pollen ince, Ecuador, April 29, 1939; paratype, which becomes grayish on the lower part of the 84 Bulletin American Museum of Natural History [Vol. LXXX pleura; vittae conspicuous, moderately wide; may exist in the same species. Whether hair black, yellow on the sternopleura; acrosticals 3-3; dorsocentrals 3-4; sternopleurals 2-1. the genus is distinct from Lasiona Wulp Scutellum brownish red, ochraceous pollinose, cannot be determined without examination with numerous spines. of specimens of that genus. Lasiona does Legs black, the tips of the tarsi reddish; hair appear to be considerably narrower, more black, partly yellow on the coxae and bases of the femora. like Macromya. In his description of Wings brownish. Squamae brown, with pale Lasiona WVulp states that the cheeks are border and black pile. Halteres brownish, the narrow, but the illustration shows them to stem mostly reddish. be wide. Also, Lasiona is said to have only Abdomen shining brown, the dorsum thickly ochraceous pollinose, but the sides and apex are fine hair on the cheeks, while there are some shining and thinly brown pollinose. Second bristles in both Macromya and Calohy- segment with numerous median spines broadly stricia. connected with the lateral spines, third and fourth segments with spines except basally, Calohystricia gertschi, new species the fourth with bristles apically. Genitalia brownish. Reddish, the mesonotum black, parafacials bare; tarsi yellow. Length, 14 mm. TYPEs.-Holotype, male, and paratypes, FEMALE.-Head reddish, the front and occiput twelve males, El Volcan, Chiriqui, Feb- blackish, pollen brassy yellow, more grayish on ruary 18, 20, and March 1, 1936 (W. J. the occiput, becoming brown on the upper part of the front. Front two-fifths as wide as eye; Gertsch), the holotype collected March 1. six or seven pairs of frontals and a secondary row In this species most of the specimens of three or four below; two pairs of orbitals; have the prosternum pilose, but the ocellars moderately strong. Occipital pile amount of pile varies, and some have it yellowish; cheeks with coarse, black hair and a few bristles near the middle. Face retreating bare. The vibrissae are situated well above to the moderately produced oral margin, with a the oral margin, thus distinguishing the conspicuous median ridge; parafacials bare. species from Calohystricia. Antennae reddish, the third segment mostly black and almost twice as long as the second. Proboscis short, with large, fleshy reddish yellow CALOHYSTRICIA TOWNSEND labellae; palpi reddish. TOWNSEND, 1931, Rev. Ent., I, p. 350. Thorax reddish, the mesonotum black except laterally, thickly cinereous yellow pollinose, the This genus is most closely related to vittae moderately wide and conspicuous; acros- Macromya Desvoidy, differing in having ticals 3-3; dorsocentrals 3-4; sternopleurals the squamae haired on the outer half in- 2-1; pleura yellowish pollinose; hair all black. stead of the outer Scutellum reddish, yellowish pollinose, with only along border, and many spines. the presence of three sternopleurals. It Legs reddish, with black hair, the tibiae and forms a natural connecting link between tarsi with reddish yellow hair, the tarsi with pale Hystricia and Macromya. The only de- bristles. scribed species is velutina from Wings brownish. Squamae brown with yel- Wulp, low border, with short black pile on more than Costa Rica. It has the abdomen velvety the outer posterior half. Halteres reddish yel- black with rather narrow apical white low. pollinose crossbands on the segments. Abdomen broad, dull brownish red above, While the species will trace out to Calo- dark orange red below; second segment with a median black vitta, the third with a black spot hystricia in Townsend's "Manual" the key posteriorly; bases of the segments narrowly is not sufficient for its distinction. Both yellow pollinose, and there is an incomplete Macromya and Calohystricia have the white pollinose line on the middle of the second vibrissae practically level with the oral to fourth segments. Second segment with a patch of spines in the middle not connected with margin, while in the other American genera the lateral spines; third with about ten discal with pilose squamae the vibrissae are spines in the middle, the fourth with two discal situated well above the more produced oral rows of spines and with bristles apically. margin. Some species of Hystricia (Engel- HOLOTYPE.-Female, El Volcan, Chiri- omyia) have the prosternum haired to qui, Panama, February 22, 1936 (W. J. greater or less degree, but this character Gertsch).