Pterygota Winged Insects
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The Mitochondrial Genomes of Palaeopteran Insects and Insights
www.nature.com/scientificreports OPEN The mitochondrial genomes of palaeopteran insects and insights into the early insect relationships Nan Song1*, Xinxin Li1, Xinming Yin1, Xinghao Li1, Jian Yin2 & Pengliang Pan2 Phylogenetic relationships of basal insects remain a matter of discussion. In particular, the relationships among Ephemeroptera, Odonata and Neoptera are the focus of debate. In this study, we used a next-generation sequencing approach to reconstruct new mitochondrial genomes (mitogenomes) from 18 species of basal insects, including six representatives of Ephemeroptera and 11 of Odonata, plus one species belonging to Zygentoma. We then compared the structures of the newly sequenced mitogenomes. A tRNA gene cluster of IMQM was found in three ephemeropteran species, which may serve as a potential synapomorphy for the family Heptageniidae. Combined with published insect mitogenome sequences, we constructed a data matrix with all 37 mitochondrial genes of 85 taxa, which had a sampling concentrating on the palaeopteran lineages. Phylogenetic analyses were performed based on various data coding schemes, using maximum likelihood and Bayesian inferences under diferent models of sequence evolution. Our results generally recovered Zygentoma as a monophyletic group, which formed a sister group to Pterygota. This confrmed the relatively primitive position of Zygentoma to Ephemeroptera, Odonata and Neoptera. Analyses using site-heterogeneous CAT-GTR model strongly supported the Palaeoptera clade, with the monophyletic Ephemeroptera being sister to the monophyletic Odonata. In addition, a sister group relationship between Palaeoptera and Neoptera was supported by the current mitogenomic data. Te acquisition of wings and of ability of fight contribute to the success of insects in the planet. -
An IPM Pocket Guide for Weed Identification in Nurseries and Landscapes Produced by Michigan State University Extension
Back to table of contents Back to index 2020 PDF An IPM Pocket Guide for Weed Identification in Nurseries and Landscapes Produced by Michigan State University Extension Contents Index of weeds (clickable) ........................ 2-8 Weed identification ............................... 9-183 Using this scouting guide..........................184 Weed classification ...................................185 Grass and broadleaf characteristics . 186-189 Glossary ........................................... 190-192 Credits and acknowledgments ......... 193-194 Suggested reading ...................................195 1 Back to table of contents Back to index Index By common name Annual bluegrass ...........................................32-33 Annual sowthistle................................................73 Asiatic (common) dayflower ..........................18-19 Barnyardgrass ....................................................27 Bindweed, field ..........................................101-103 Bindweed, hedge .......................................102-103 Birdseye pearlwort ..............................................98 Birdsfoot trefoil...........................................109-110 Bittercress, hairy ............................................83-85 Bittercress, smallflowered...................................85 Black medic ............................................... 112-113 Blackseed plantain ...........................................150 Bluegrass, annual ..........................................32-33 Brambles ...................................................164-165 -
A New Insect Trackway from the Upper Jurassic—Lower Cretaceous Eolian Sandstones of São Paulo State, Brazil: Implications for Reconstructing Desert Paleoecology
A new insect trackway from the Upper Jurassic—Lower Cretaceous eolian sandstones of São Paulo State, Brazil: implications for reconstructing desert paleoecology Bernardo de C.P. e M. Peixoto1,2, M. Gabriela Mángano3, Nicholas J. Minter4, Luciana Bueno dos Reis Fernandes1 and Marcelo Adorna Fernandes1,2 1 Laboratório de Paleoicnologia e Paleoecologia, Departamento de Ecologia e Biologia Evolutiva, Universidade Federal de São Carlos (UFSCar), São Carlos, São Paulo, Brazil 2 Programa de Pós Graduacão¸ em Ecologia e Recursos Naturais, Centro de Ciências Biológicas e da Saúde, Universidade Federal de São Carlos (UFSCar), São Carlos, São Paulo, Brazil 3 Department of Geological Sciences, University of Saskatchewan, Saskatoon, Saskatchewan, Canada 4 School of the Environment, Geography, and Geosciences, University of Portsmouth, Portsmouth, Hampshire, United Kingdom ABSTRACT The new ichnospecies Paleohelcura araraquarensis isp. nov. is described from the Upper Jurassic-Lower Cretaceous Botucatu Formation of Brazil. This formation records a gigantic eolian sand sea (erg), formed under an arid climate in the south-central part of Gondwana. This trackway is composed of two track rows, whose internal width is less than one-quarter of the external width, with alternating to staggered series, consisting of three elliptical tracks that can vary from slightly elongated to tapered or circular. The trackways were found in yellowish/reddish sandstone in a quarry in the Araraquara municipality, São Paulo State. Comparisons with neoichnological studies and morphological inferences indicate that the producer of Paleohelcura araraquarensis isp. nov. was most likely a pterygote insect, and so could have fulfilled one of the Submitted 6 November 2019 ecological roles that different species of this group are capable of performing in dune Accepted 10 March 2020 deserts. -
Bramshill Site of Special Scientific Interest
The Dragonflies of Bramshill Site of Special Scientific Interest Freshwater Habitats Trust Author Ken Crick Forward Bramshill Site of Special Scientific Interest (SSSI) is a Flagship Pond Site. Part of a network of the very best of Britain’s ponds; sites of exceptional importance for freshwater wildlife and some of our finest freshwater habitats. The Flagship sites can be a single special pond, or more commonly group of ponds, selected because they support rich, often irreplaceable, communities and species at risk of extinction. They represent some of the least impacted, most diverse pond habitats remaining in the country. Many of our nation’s most beautiful and biodiverse waterbodies have degraded irrevocably, and it’s critically important that the remaining sites are well protected and well managed. In 2015, with funding from the Heritage Lottery Fund, Freshwater Habitats Trust launched the Flagship Ponds project, Mats of Water Crowfoot flower on Bramshill working with land managers and community groups to ensure that the most Plantation’s Longwater. critical pond sites in Britain were protected for the long term. This book has been published with the aim of enabling people visiting this, Introduction immensely important Flagship Pond Site in Northern Hampshire, to identify the dragonflies and damselflies they encounter - by reference to a simple text This nationally important Site of managed by Forestry Commission and in places subsequent backfilling Special Scientific Interest (SSSI) England (FCE), please see the site with landfill, Bramshill SSSI has and photographs. It should also inform those visiting the site of the location is notified as such in part for its map on page 6 which depicts the through a combination of careful of the majority of freshwater habitats. -
State of New York City's Plants 2018
STATE OF NEW YORK CITY’S PLANTS 2018 Daniel Atha & Brian Boom © 2018 The New York Botanical Garden All rights reserved ISBN 978-0-89327-955-4 Center for Conservation Strategy The New York Botanical Garden 2900 Southern Boulevard Bronx, NY 10458 All photos NYBG staff Citation: Atha, D. and B. Boom. 2018. State of New York City’s Plants 2018. Center for Conservation Strategy. The New York Botanical Garden, Bronx, NY. 132 pp. STATE OF NEW YORK CITY’S PLANTS 2018 4 EXECUTIVE SUMMARY 6 INTRODUCTION 10 DOCUMENTING THE CITY’S PLANTS 10 The Flora of New York City 11 Rare Species 14 Focus on Specific Area 16 Botanical Spectacle: Summer Snow 18 CITIZEN SCIENCE 20 THREATS TO THE CITY’S PLANTS 24 NEW YORK STATE PROHIBITED AND REGULATED INVASIVE SPECIES FOUND IN NEW YORK CITY 26 LOOKING AHEAD 27 CONTRIBUTORS AND ACKNOWLEGMENTS 30 LITERATURE CITED 31 APPENDIX Checklist of the Spontaneous Vascular Plants of New York City 32 Ferns and Fern Allies 35 Gymnosperms 36 Nymphaeales and Magnoliids 37 Monocots 67 Dicots 3 EXECUTIVE SUMMARY This report, State of New York City’s Plants 2018, is the first rankings of rare, threatened, endangered, and extinct species of what is envisioned by the Center for Conservation Strategy known from New York City, and based on this compilation of The New York Botanical Garden as annual updates thirteen percent of the City’s flora is imperiled or extinct in New summarizing the status of the spontaneous plant species of the York City. five boroughs of New York City. This year’s report deals with the City’s vascular plants (ferns and fern allies, gymnosperms, We have begun the process of assessing conservation status and flowering plants), but in the future it is planned to phase in at the local level for all species. -
Dragonfly News 66
Dragonfly News 66 The Magazine of the British Dragonfly Society Autumn 2014 www.british-dragonflies.org.uk Meet the new BDS Chairman, How many Willow Emeralds are David Chelmick ovipositing? Dragonfly hunting....in Sweden? Andy Holt’s unique larval portraits How tatty can a dragonfly be and still fly? Dragonfly News 66 The Magazine of the British Dragonfly Society Published twice a year, in April and October, Dragonfly News covers all aspects of the British Dragonfly Society’s field, recording, monitoring, research, conservation and social activities, as well as information from the wider dragonfly, natural history and conservation world. The emphasis is on dragonflies recorded in the UK. The British Dragonfly Society aims to promote and encourage the study, conservation and understanding of dragonflies and their natural habitats, especially in the UK, and to raise public awareness of dragonflies. Dragonfly News is edited & designed by: Trustees & Officers of the BDS Mark Tyrrell, 8 Warwick Close, Raunds, Chairman: David Chelmick Northants., NN9 6JH Tel. Vice-Chairman: Vacant e-mail: Secretary: Henry Curry, 23 Bowker Way, Whittlesey, Peterborough, PE7 1PY. Tel. Deadlines for inclusion of copy: Spring 31 January Treasurer: Brian Walker, 49 Roman Way, Wantage, Autumn 31 July Oxfordshire, OX12 9YF. Tel. Advertising Rates: Trustees: David Goddard, Stuart Irons, Mick Parfitt. £15 for small-ad (text only); £40 for quarter- Journal Editor: Peter Mill, 8 Cookridge Grove, LEEDS, page; £60 for half-page; £100 for full-page. LS16 7LH. Shop Manager: Lynn Curry, 23 Bowker Way, Whittlesey, Peterborough, PE7 1PY Tel. © British Dragonfly Society 2014 All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted, in any Dragonfly Conservation Group (DCG) form or by any means, electronic, mechanical, photocopying, Convenor: Dave Smallshire, 8, Twindle Beer, Chudleigh, Newton recording or otherwise, without the permission of the British Abbot, Devon, TQ13 0JP. -
Driven Color Lightness and Body Size Variation Scale to Local Assemblages of European Odonata but Are Modified by Propensity for Dispersal
Received: 2 April 2020 | Revised: 26 June 2020 | Accepted: 29 June 2020 DOI: 10.1002/ece3.6596 ORIGINAL RESEARCH Temperature-driven color lightness and body size variation scale to local assemblages of European Odonata but are modified by propensity for dispersal Daniel Acquah-Lamptey1 | Martin Brändle1 | Roland Brandl1 | Stefan Pinkert1,2 1Faculty of Biology, Department of Ecology – Animal Ecology, Philipps-Universität Abstract Marburg, Marburg, Germany 1. Previous macrophysiological studies suggested that temperature-driven color 2 Ecology & Evolutionary Biology, Yale lightness and body size variations strongly influence biogeographical patterns in University, New Haven, CT, USA ectotherms. However, these trait–environment relationships scale to local assem- Correspondence blages and the extent to which they can be modified by dispersal remains largely Daniel Acquah-Lamptey, Faculty of Biology, Department of Ecology – Animal Ecology, unexplored. We test whether the predictions of the thermal melanism hypoth- Philipps-Universität Marburg, Karl-von- esis and the Bergmann's rule hold for local assemblages. We also assess whether Frisch-Straße 8, 35043 Marburg, Germany. Email: [email protected] these trait–environment relationships are more important for species adapted to less stable (lentic) habitats, due to their greater dispersal propensity compared to those adapted to stable (lotic) habitats. 2. We quantified the color lightness and body volume of 99 European dragon- and damselflies (Odonata) and combined these trait information with survey data for 518 local assemblages across Europe. Based on this continent-wide yet spatially explicit dataset, we tested for effects temperature and precipitation on the color lightness and body volume of local assemblages and assessed differences in their relative importance and strength between lentic and lotic assemblages, while ac- counting for spatial and phylogenetic autocorrelation. -
Dragonflies and Damselflies in Your Garden
Natural England works for people, places and nature to conserve and enhance biodiversity, landscapes and wildlife in rural, urban, coastal and marine areas. Dragonflies and www.naturalengland.org.uk © Natural England 2007 damselflies in your garden ISBN 978-1-84754-015-7 Catalogue code NE21 Written by Caroline Daguet Designed by RR Donnelley Front cover photograph: A male southern hawker dragonfly. This species is the one most commonly seen in gardens. Steve Cham. www.naturalengland.org.uk Dragonflies and damselflies in your garden Dragonflies and damselflies are Modern dragonflies are tiny by amazing insects. They have a long comparison, but are still large and history and modern species are almost spectacular enough to capture the identical to ancestors that flew over attention of anyone walking along a prehistoric forests some 300 million river bank or enjoying a sunny years ago. Some of these ancient afternoon by the garden pond. dragonflies were giants, with This booklet will tell you about the wingspans of up to 70 cm. biology and life-cycles of dragonflies and damselflies, help you to identify some common species, and tell you how you can encourage these insects to visit your garden. Male common blue damselfly. Most damselflies hold their wings against their bodies when at rest. BDS Dragonflies and damselflies belong to Dragonflies the insect order known as Odonata, Dragonflies are usually larger than meaning ‘toothed jaws’. They are often damselflies. They are stronger fliers and referred to collectively as ‘dragonflies’, can often be found well away from but dragonflies and damselflies are two water. When at rest, they hold their distinct groups. -
Is Ellipura Monophyletic? a Combined Analysis of Basal Hexapod
ARTICLE IN PRESS Organisms, Diversity & Evolution 4 (2004) 319–340 www.elsevier.de/ode Is Ellipura monophyletic? A combined analysis of basal hexapod relationships with emphasis on the origin of insects Gonzalo Giribeta,Ã, Gregory D.Edgecombe b, James M.Carpenter c, Cyrille A.D’Haese d, Ward C.Wheeler c aDepartment of Organismic and Evolutionary Biology, Museum of Comparative Zoology, Harvard University, 16 Divinity Avenue, Cambridge, MA 02138, USA bAustralian Museum, 6 College Street, Sydney, New South Wales 2010, Australia cDivision of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024, USA dFRE 2695 CNRS, De´partement Syste´matique et Evolution, Muse´um National d’Histoire Naturelle, 45 rue Buffon, F-75005 Paris, France Received 27 February 2004; accepted 18 May 2004 Abstract Hexapoda includes 33 commonly recognized orders, most of them insects.Ongoing controversy concerns the grouping of Protura and Collembola as a taxon Ellipura, the monophyly of Diplura, a single or multiple origins of entognathy, and the monophyly or paraphyly of the silverfish (Lepidotrichidae and Zygentoma s.s.) with respect to other dicondylous insects.Here we analyze relationships among basal hexapod orders via a cladistic analysis of sequence data for five molecular markers and 189 morphological characters in a simultaneous analysis framework using myriapod and crustacean outgroups.Using a sensitivity analysis approach and testing for stability, the most congruent parameters resolve Tricholepidion as sister group to the remaining Dicondylia, whereas most suboptimal parameter sets group Tricholepidion with Zygentoma.Stable hypotheses include the monophyly of Diplura, and a sister group relationship between Diplura and Protura, contradicting the Ellipura hypothesis.Hexapod monophyly is contradicted by an alliance between Collembola, Crustacea and Ectognatha (i.e., exclusive of Diplura and Protura) in molecular and combined analyses. -
Dragonflies of La Brenne & Vienne
Dragonflies of La Brenne & Vienne Naturetrek Tour Report 13 - 20 June 2018 Dainty White-faced Darter (Leucorrhinia caudalis) male Yellow-spotted Emerald (Somatochlora flavomaculata) male Report and images by Nick Ransdale Naturetrek Mingledown Barn Wolf's Lane Chawton Alton Hampshire GU34 3HJ UK T: +44 (0)1962 733051 E: [email protected] W: www.naturetrek.co.uk Tour Report Dragonflies of La Brenne & Vienne Tour participants: Nick Ransdale (leader) with six Naturetrek clients Summary This two-centre holiday in central-western France gave an excellent insight into not only the dragonflies but also the abundant butterflies, birds and other wildlife of the region. The first two days were spent in the southern Vienne before we moved to the bizarre landscape of the Pinail reserve, and finally to Mezieres where we spent three days in the Brenne - ‘land of a thousand lakes’. This year's tour started on the cool side at 17-18°C, but settled into a pattern that proved to be ideal for finding and photographing odonata. Due to the sharp eyes, flexibility and optimism of group members, the tour was a resounding success, scoring a total of 44 species (tour average 41), equalling the tour record. The emphasis here is always on getting good, diagnostic views for all participants, something we achieved for all but one species. It was a good year for 'sets' of species this year, with both pincertails, four emerald dragonflies and both whiteface species. Added to this were five fritillary butterfly species, both Emperors (Purple and Lesser Purple), and an outstanding two clearwing moths – both Hornet and Firey. -
Predictive Modelling of Spatial Biodiversity Data to Support Ecological Network Mapping: a Case Study in the Fens
Predictive modelling of spatial biodiversity data to support ecological network mapping: a case study in the Fens Christopher J Panter, Paul M Dolman, Hannah L Mossman Final Report: July 2013 Supported and steered by the Fens for the Future partnership and the Environment Agency www.fensforthefuture.org.uk Published by: School of Environmental Sciences, University of East Anglia, Norwich, NR4 7TJ, UK Suggested citation: Panter C.J., Dolman P.M., Mossman, H.L (2013) Predictive modelling of spatial biodiversity data to support ecological network mapping: a case study in the Fens. University of East Anglia, Norwich. ISBN: 978-0-9567812-3-9 © Copyright rests with the authors. Acknowledgements This project was supported and steered by the Fens for the Future partnership. Funding was provided by the Environment Agency (Dominic Coath). We thank all of the species recorders and natural historians, without whom this work would not be possible. Cover picture: Extract of a map showing the predicted distribution of biodiversity. Contents Executive summary .................................................................................................................... 4 Introduction ............................................................................................................................... 5 Methodology .......................................................................................................................... 6 Biological data ................................................................................................................... -
Arthropods of Elm Fork Preserve
Arthropods of Elm Fork Preserve Arthropods are characterized by having jointed limbs and exoskeletons. They include a diverse assortment of creatures: Insects, spiders, crustaceans (crayfish, crabs, pill bugs), centipedes and millipedes among others. Column Headings Scientific Name: The phenomenal diversity of arthropods, creates numerous difficulties in the determination of species. Positive identification is often achieved only by specialists using obscure monographs to ‘key out’ a species by examining microscopic differences in anatomy. For our purposes in this survey of the fauna, classification at a lower level of resolution still yields valuable information. For instance, knowing that ant lions belong to the Family, Myrmeleontidae, allows us to quickly look them up on the Internet and be confident we are not being fooled by a common name that may also apply to some other, unrelated something. With the Family name firmly in hand, we may explore the natural history of ant lions without needing to know exactly which species we are viewing. In some instances identification is only readily available at an even higher ranking such as Class. Millipedes are in the Class Diplopoda. There are many Orders (O) of millipedes and they are not easily differentiated so this entry is best left at the rank of Class. A great deal of taxonomic reorganization has been occurring lately with advances in DNA analysis pointing out underlying connections and differences that were previously unrealized. For this reason, all other rankings aside from Family, Genus and Species have been omitted from the interior of the tables since many of these ranks are in a state of flux.