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Carpenter 1999 EL PLESIOSAURIO DE VILLA DE LEYVA (BOYACÁ, COLOMBIA)
Boletín de Geología Vol. 23, No. 38, Enero-Junio de 2001 Callawayasaurus colombiensis (Welles) Carpenter 1999 EL PLESIOSAURIO DE VILLA DE LEYVA (BOYACÁ, COLOMBIA). ¿UN NUEVO ESPÉCIMEN? Jerez Jaimes, J. H1.; Narváez Parra, E. X1. RESUMEN En los depósitos del Cretácico (Aptiano) de Villa de Leyva se han reportado dos especies de plesiosaurios, un pliosaurio Kronosaurus boyacensis Hampe 1992, y un plesiosaurio Callawayasaurus colombiensis (Welles) Carpenter, 1999 (= Alzadasaurus colombiensis Welles, 1962). Se realiza la determinación de un espécimen de elasmosaurio encontrado por los pobladores de la zona rural de Villa de Leyva en 1999 con base en material fotográfico del mismo, siendo muy probable que corresponda a la especie Callawayasaurus colombiensis (Welles) Carpenter, 1999. Palabras Claves: Cretácico, Plesiosaurios, Villa de Leyva. ABSTRACT In the deposits of the Cretaceous (Aptian) of Villa de Leyva two plesiosaurs species have been reported, a pliosaur Kronosaurus boyacensis Hampe 1992, and a plesiosaur Callawayasaurus colombiensis (Welles) Carpenter, 1999 (= Alzadasaurus colombiensis Welles, 1962). We carried out the determination of elasmosaur specimen found by the inhabitants of the rural area of Villa de Leyva in 1999, on the basis of photographic material of it. Probably it corresponds to the Callawayasaurus colombiensis specie (Welles) Carpenter, 1999. Key Words: Cretaceous, Plesiosaurs, Villa de Leyva. 1Biólogos, Calle 10A # 24-68 Bucaramanga, Santander (Colombia). Correo electrónico: [email protected] Callawayasaurus -
Estimating the Evolutionary Rates in Mosasauroids and Plesiosaurs: Discussion of Niche Occupation in Late Cretaceous Seas
Estimating the evolutionary rates in mosasauroids and plesiosaurs: discussion of niche occupation in Late Cretaceous seas Daniel Madzia1 and Andrea Cau2 1 Department of Evolutionary Paleobiology, Institute of Paleobiology, Polish Academy of Sciences, Warsaw, Poland 2 Independent, Parma, Italy ABSTRACT Observations of temporal overlap of niche occupation among Late Cretaceous marine amniotes suggest that the rise and diversification of mosasauroid squamates might have been influenced by competition with or disappearance of some plesiosaur taxa. We discuss that hypothesis through comparisons of the rates of morphological evolution of mosasauroids throughout their evolutionary history with those inferred for contemporary plesiosaur clades. We used expanded versions of two species- level phylogenetic datasets of both these groups, updated them with stratigraphic information, and analyzed using the Bayesian inference to estimate the rates of divergence for each clade. The oscillations in evolutionary rates of the mosasauroid and plesiosaur lineages that overlapped in time and space were then used as a baseline for discussion and comparisons of traits that can affect the shape of the niche structures of aquatic amniotes, such as tooth morphologies, body size, swimming abilities, metabolism, and reproduction. Only two groups of plesiosaurs are considered to be possible niche competitors of mosasauroids: the brachauchenine pliosaurids and the polycotylid leptocleidians. However, direct evidence for interactions between mosasauroids and plesiosaurs is scarce and limited only to large mosasauroids as the Submitted 31 July 2019 predators/scavengers and polycotylids as their prey. The first mosasauroids differed Accepted 18 March 2020 from contemporary plesiosaurs in certain aspects of all discussed traits and no evidence Published 13 April 2020 suggests that early representatives of Mosasauroidea diversified after competitions with Corresponding author plesiosaurs. -
A Revision of the Classification of the Plesiosauria with a Synopsis of the Stratigraphical and Geographical Distribution Of
LUNDS UNIVERSITETS ARSSKRIFT. N. F. Avd. 2. Bd 59. Nr l. KUNGL. FYSIOGRAFISKA SÅLLSKAPETS HANDLINGAR, N. F. Bd 74. Nr 1. A REVISION OF THE CLASSIFICATION OF THE PLESIOSAURIA WITH A SYNOPSIS OF THE STRATIGRAPHICAL AND GEOGRAPHICAL DISTRIBUTION OF THE GROUP BY PER OVE PERSSON LUND C. W. K. GLEER UP Read before the Royal Physiographic Society, February 13, 1963. LUND HÅKAN OHLSSONS BOKTRYCKERI l 9 6 3 l. Introduction The sub-order Plesiosauria is one of the best known of the Mesozoic Reptile groups, but, as emphasized by KuHN (1961, p. 75) and other authors, its classification is still not satisfactory, and needs a thorough revision. The present paper is an attempt at such a revision, and includes also a tabular synopsis of the stratigraphical and geo graphical distribution of the group. Some of the species are discussed in the text (pp. 17-22). The synopsis is completed with seven maps (figs. 2-8, pp. 10-16), a selective synonym list (pp. 41-42), and a list of rejected species (pp. 42-43). Some forms which have been erroneously referred to the Plesiosauria are also briefly mentioned ("Non-Plesiosaurians", p. 43). - The numerals in braekets after the generic and specific names in the text refer to the tabular synopsis, in which the different forms are numbered in successional order. The author has exaroined all material available from Sweden, Australia and Spitzbergen (PERSSON 1954, 1959, 1960, 1962, 1962a); the major part of the material from the British Isles, France, Belgium and Luxembourg; some of the German spec imens; certain specimens from New Zealand, now in the British Museum (see LYDEK KER 1889, pp. -
Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships
438 Article 438 by Saswati Bandyopadhyay1* and Sanghamitra Ray2 Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships 1Geological Studies Unit, Indian Statistical Institute, 203 B. T. Road, Kolkata 700108, India; email: [email protected] 2Department of Geology and Geophysics, Indian Institute of Technology, Kharagpur 721302, India; email: [email protected] *Corresponding author (Received : 23/12/2018; Revised accepted : 11/09/2019) https://doi.org/10.18814/epiiugs/2020/020028 The twelve Gondwanan stratigraphic horizons of many extant lineages, producing highly diverse terrestrial vertebrates India have yielded varied vertebrate fossils. The oldest in the vacant niches created throughout the world due to the end- Permian extinction event. Diapsids diversified rapidly by the Middle fossil record is the Endothiodon-dominated multitaxic Triassic in to many communities of continental tetrapods, whereas Kundaram fauna, which correlates the Kundaram the non-mammalian synapsids became a minor components for the Formation with several other coeval Late Permian remainder of the Mesozoic Era. The Gondwana basins of peninsular horizons of South Africa, Zambia, Tanzania, India (Fig. 1A) aptly exemplify the diverse vertebrate faunas found Mozambique, Malawi, Madagascar and Brazil. The from the Late Palaeozoic and Mesozoic. During the last few decades much emphasis was given on explorations and excavations of Permian-Triassic transition in India is marked by vertebrate fossils in these basins which have yielded many new fossil distinct taxonomic shift and faunal characteristics and vertebrates, significant both in numbers and diversity of genera, and represented by small-sized holdover fauna of the providing information on their taphonomy, taxonomy, phylogeny, Early Triassic Panchet and Kamthi fauna. -
How Plesiosaurs Swam: New Insights Into Their Underwater Flight Using “Ava”, a Virtual Pliosaur
Preprints (www.preprints.org) | NOT PEER-REVIEWED | Posted: 9 October 2019 doi:10.20944/preprints201910.0094.v1 How Plesiosaurs Swam: New Insights into Their Underwater Flight Using “Ava”, a Virtual Pliosaur Max Hawthorne1,*, Mark A. S. McMenamin 2, Paul de la Salle3 1Far From The Tree Press, LLC, 4657 York Rd., #952, Buckingham, PA, 18912, United States 2Department of Geology and Geography, Mount Holyoke College, South Hadley, Massachusetts, United States 3Swindon, England *Correspondence: [email protected]; Tel.: 267-337-7545 Abstract Analysis of plesiosaur swim dynamics by means Further study attempted to justify the use of all four flippers of a digital 3D armature (wireframe “skeleton”) of a simultaneously via the use of paddle-generated vortices, pliosauromorph (“Ava”) demonstrates that: 1, plesiosaurs which require specific timing to achieve optimal additional used all four flippers for primary propulsion; 2, plesiosaurs thrust. These attempts have largely relied on anatomical utilized all four flippers simultaneously; 3, respective pairs studies of strata-compressed plesiosaur skeletons, and/or of flippers of Plesiosauridae, front and rear, traveled through preconceived notions as pertains to the paddles’ inherent distinctive, separate planes of motion, and; 4, the ability to ranges of motion [8, 10-12]. What has not been considered utilize all four paddles simultaneously allowed these largely are the opposing angles of the pectoral and pelvic girdles, predatory marine reptiles to achieve a significant increase in which strongly indicate varied-yet-complementing relations acceleration and speed, which, in turn, contributed to their between the front and rear sets of paddles, both in repose and sustained dominance during the Mesozoic. -
A New Species of Cyclotosaurus (Stereospondyli, Capitosauria) from the Late Triassic of Bielefeld, NW Germany, and the Intrarelationships of the Genus
Foss. Rec., 19, 83–100, 2016 www.foss-rec.net/19/83/2016/ doi:10.5194/fr-19-83-2016 © Author(s) 2016. CC Attribution 3.0 License. A new species of Cyclotosaurus (Stereospondyli, Capitosauria) from the Late Triassic of Bielefeld, NW Germany, and the intrarelationships of the genus Florian Witzmann1,2, Sven Sachs3,a, and Christian J. Nyhuis4 1Department of Ecology and Evolutionary Biology, Brown University, Providence, G-B204, RI 02912, USA 2Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, 10115 Berlin, Germany 3Naturkundemuseum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany 4Galileo-Wissenswelt, Mummendorferweg 11b, 23769 Burg auf Fehmarn, Germany aprivate address: Im Hof 9, 51766 Engelskirchen, Germany Correspondence to: Florian Witzmann (fl[email protected]; fl[email protected]) Received: 19 January 2016 – Revised: 11 March 2016 – Accepted: 14 March 2016 – Published: 23 March 2016 Abstract. A nearly complete dermal skull roof of a capi- clotosaurus is the sister group of the Heylerosaurinae (Eo- tosaur stereospondyl with closed otic fenestrae from the mid- cyclotosaurus C Quasicyclotosaurus). Cyclotosaurus buech- dle Carnian Stuttgart Formation (Late Triassic) of Bielefeld- neri represents the only unequivocal evidence of Cycloto- Sieker (NW Germany) is described. The specimen is as- saurus (and of a cyclotosaur in general) in northern Germany. signed to the genus Cyclotosaurus based on the limited con- tribution of the frontal to the orbital margin via narrow lat- eral processes. A new species, Cyclotosaurus buechneri sp. nov., is erected based upon the following unique combina- 1 Introduction tion of characters: (1) the interorbital distance is short so that the orbitae are medially placed (shared with C. -
Mosasaurs Continuing from Last Time…
Pliosaurs and Mosasaurs Continuing From Last Time… • Pliosauridae: the big marine predators of the Jurassic Pliosauridae • Some of the largest marine predators of all time, these middle Jurassic sauropterygians include such giants as Kronosaurus, Liopleurodon, Macroplata, Peloneustes, Pliosaurus, and Brachauchenius Pliosaur Mophology • While the number of cervical vertebrae is less than in plesiosaurs, there is still variation: Macroplata (29) vs. Kronosaurus (13) Pliosaur Morphology • Larger pliosaurs adopted a more streamlined body shape, like modern whales, with a large skull and compact neck, and generally the hind limbs were larger than the front, while plesiosaurs had larger forelimbs Pliosaur Morphology • Powerful limb girdles and large (banana sized) conical teeth helped pliosaurs eat larger, quicker prey than the piscivorous plesiosaurs Liopleurodon • NOT 25 m long in general (average of 40 feet), though perhaps certain individuals could reach that size, making Liopleurodon ferox the largest carnivore to ever live • Recent skull studies indicate that Liopleurodon could sample water in stereo through nostrils, locating scents much as we locate sound Cretaceous Seas • Breakup of Gondwana causes large undersea mountain chains to form, raising sea levels everywhere • Shallow seas encourage growth of corals, which increases calcium abundance and chalk formation • Warm seas and a gentle thermal gradient yield a hospitable environment to rays, sharks, teleosts, and the first radiation of siliceous diatoms Kronosaurus • Early Cretaceous -
Anatomy and Relationships of the Triassic Temnospondyl Sclerothorax
Anatomy and relationships of the Triassic temnospondyl Sclerothorax RAINER R. SCHOCH, MICHAEL FASTNACHT, JÜRGEN FICHTER, and THOMAS KELLER Schoch, R.R., Fastnacht, M., Fichter, J., and Keller, T. 2007. Anatomy and relationships of the Triassic temnospondyl Sclerothorax. Acta Palaeontologica Polonica 52 (1): 117–136. Recently, new material of the peculiar tetrapod Sclerothorax hypselonotus from the Middle Buntsandstein (Olenekian) of north−central Germany has emerged that reveals the anatomy of the skull and anterior postcranial skeleton in detail. Despite differences in preservation, all previous plus the new finds of Sclerothorax are identified as belonging to the same taxon. Sclerothorax is characterized by various autapomorphies (subquadrangular skull being widest in snout region, ex− treme height of thoracal neural spines in mid−trunk region, rhomboidal interclavicle longer than skull). Despite its pecu− liar skull roof, the palate and mandible are consistent with those of capitosauroid stereospondyls in the presence of large muscular pockets on the basal plate, a flattened edentulous parasphenoid, a long basicranial suture, a large hamate process in the mandible, and a falciform crest in the occipital part of the cheek. In order to elucidate the phylogenetic position of Sclerothorax, we performed a cladistic analysis of 18 taxa and 70 characters from all parts of the skeleton. According to our results, Sclerothorax is nested well within the higher stereospondyls, forming the sister taxon of capitosauroids. Palaeobiologically, Sclerothorax is interesting for its several characters believed to correlate with a terrestrial life, although this is contrasted by the possession of well−established lateral line sulci. Key words: Sclerothorax, Temnospondyli, Stereospondyli, Buntsandstein, Triassic, Germany. -
Late Cretaceous) of Morocco : Palaeobiological and Behavioral Implications Remi Allemand
Endocranial microtomographic study of marine reptiles (Plesiosauria and Mosasauroidea) from the Turonian (Late Cretaceous) of Morocco : palaeobiological and behavioral implications Remi Allemand To cite this version: Remi Allemand. Endocranial microtomographic study of marine reptiles (Plesiosauria and Mosasauroidea) from the Turonian (Late Cretaceous) of Morocco : palaeobiological and behavioral implications. Paleontology. Museum national d’histoire naturelle - MNHN PARIS, 2017. English. NNT : 2017MNHN0015. tel-02375321 HAL Id: tel-02375321 https://tel.archives-ouvertes.fr/tel-02375321 Submitted on 22 Nov 2019 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. MUSEUM NATIONAL D’HISTOIRE NATURELLE Ecole Doctorale Sciences de la Nature et de l’Homme – ED 227 Année 2017 N° attribué par la bibliothèque |_|_|_|_|_|_|_|_|_|_|_|_| THESE Pour obtenir le grade de DOCTEUR DU MUSEUM NATIONAL D’HISTOIRE NATURELLE Spécialité : Paléontologie Présentée et soutenue publiquement par Rémi ALLEMAND Le 21 novembre 2017 Etude microtomographique de l’endocrâne de reptiles marins (Plesiosauria et Mosasauroidea) du Turonien (Crétacé supérieur) du Maroc : implications paléobiologiques et comportementales Sous la direction de : Mme BARDET Nathalie, Directrice de Recherche CNRS et les co-directions de : Mme VINCENT Peggy, Chargée de Recherche CNRS et Mme HOUSSAYE Alexandra, Chargée de Recherche CNRS Composition du jury : M. -
A BRACHYOPID TEMNOSPONDYL from the LOWER CYNOGNATHUS ASSEMBLAGE ZONE in the NORTHERN KAROO BASIN, SOUTH AFRICA by Ross J. Damian
Palaeont. afr., 38, 57-69 (2002) A BRACHYOPID TEMNOSPONDYL FROM THE LOWER CYNOGNATHUS ASSEMBLAGE ZONE IN THE NORTHERN KAROO BASIN, SOUTH AFRICA by Ross J. Damiani and Ashleigh M. Jeannot Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa ABSTRACT A new brachyopid temnospondyl is described from the Early to Middle Triassic Cynognathus Assemblage Zone of the upper Beaufort Group, Karoo Basin of South Africa. It is the fourth named brachyopid from the Karoo and the first from the northern part of the basin. Despite the incomplete nature of the holotype skull, the new brachyopid apparently shows closest affinities to Batrachosuchus watsoni. However, differences in the width of the sensory sulci, the absence of a transverse occipital sulcus, and the presence of a unique narial morphology, warrants separation at the species level. The holotype skull also provides insight into the morphology of the ventral surface of the skull roof and the configuration of the bones between the orbit and the nostril. A referTed right mandibular ramus, the most complete yet recovered of a brachyopid, also shows several unique features. A reconsideration of the taxonomy of the brachyopid genus Batrachosuchus reveals that Batrachosuchus watsoni possesses several characters distinct from the type species, Batrachosuchus browni, and is thus transfened to a new genus. In addition, 'Batrachosuchus' henwoodi and Batrachosuchus concordi probably do not pertain to the genus Batrachosuchus. Brachyopid diversity in the Karoo is exceeded only by the Mastodonsauridae and Rhinesuchidae, and they may eventually prove to be important aids in the biostratigraphy of the Cynognathus Assemblage Zone. -
Functional Morphology of Stereospondyl Amphibian Skulls
Functional Morphology of Stereospondyl Amphibian Skulls Samantha Clare Penrice Doctor of Philosophy School of Life Sciences College of Science July 2018 Functional morphology of stereospondyl amphibian skulls Stereospondyls were the most diverse clade of early tetrapods, spanning 190 million years, with over 250 species belonging to eight taxonomic groups. They had a range of morphotypes and have been found on every continent. Stereospondyl phylogeny is widely contested and repeatedly examined but despite these studies, we are still left with the question, why were they so successful and why did they die out? A group-wide analysis of functional morphology, informing us about their palaeobiology, was lacking for this group and was carried out in order to address the questions of their success and demise. Based on an original photograph collection, size independent skull morphometrics were used, in conjunction with analyses of the fossil record and comparative anatomy, to provide a synthesis of the functional morphology of stereospondyl amphibians. Stereospondyls originated in the Carboniferous and most taxonomic groups were extinct at the end of the Triassic. The early Triassic had exceptionally high numbers of short- lived genera, in habitats that were mostly arid but apparently experienced occasional monsoon rains. Genera turnover slowed and diversity was stable in the Middle Triassic, then declined with a series of extinctions of the Late Triassic. Stereospondyls showed the pattern of ‘disaster’ taxa: rapidly diversifying following a mass extinction, spreading to a global distribution, although this high diversity was relatively short-lived. Geometric morphometrics on characteristics of the skull and palate was carried out to assess general skull morphology and identified the orbital position and skull outline to be the largest sources of skull variation. -
Reptiles Fósiles De Colombia Un Aporte Al Conocimiento Y a La Enseñanza Del Patrimonio Paleontológico Del País
Reptiles Fósiles de Colombia Un aporte al conocimiento y a la enseñanza del patrimonio paleontológico del país Luis Gonzalo Ortiz-Pabón Universidad Pedagógica Nacional Facultad de Ciencia y tecnología Departamento de biología Bogotá D.C. 2020 Reptiles Fósiles de Colombia Un aporte al conocimiento y a la enseñanza del patrimonio paleontológico del país Luis Gonzalo Ortiz-Pabón Trabajo presentado como requisito para optar por el título de: Licenciado en Biología Directora: Heidy Paola Jiménez Medina MSc. Línea de Investigación: Educación en Ciencias y formación Ambiental Grupo de Investigación: Educación en Ciencias, Ambiente y Diversidad Universidad Pedagógica Nacional Facultad de Ciencia y tecnología Departamento de biología Bogotá D.C. 2020 Dedicatoria A mi mami, quien ha estado acompañándome y apoyándome en muchos de los momentos definitivos de mi vida, además de la deuda que tengo con ella desde 2008. “La ciencia no es una persecución despiadada de información objetiva. Es una actividad humana creativa, sus genios actúan más como artistas que como procesadores de información” Stephen J. Gould Agradecimientos En primera instancia agradezco a todos los maestros y maestras que fueron parte esencial de mi formación académica. Agradecimiento especial al Ilustrador y colega Marco Salazar por su aporte gráfico a la construcción del libro Reptiles Fósiles de Colombia, a Oscar Hernández y a Galdra Films por su valioso aporte en la diagramación y edición del libro, a Heidy Jiménez quien fue mi directora y guía en el desarrollo de este trabajo y a Vanessa Robles, quien estuvo acompañando la revisión del libro y este escrito, además del apoyo emocional brindado en todo momento.