Arthropoda Selecta 27(1): 95–105 © ARTHROPODA SELECTA, 2018

Arachnid types in the Zoological Museum of the Moscow State University. II. Aranei 1 (Arachnida)

Òèïû ïàóêîîáðàçíûõ â Çîîëîãè÷åñêîì ìóçåå ÌÃÓ. II. Aranei 1 (Arachnida)

Kirill G. Mikhailov Ê.Ã. Ìèõàéëîâ

Zoological Museum MGU, Bolshaya Nikitskaya Str. 2, Moscow 125009 Russia. E-mail: [email protected] Зоологический музей МГУ, ул. Большая Никитская, 2, Москва 125009 Россия.

KEY WORDS: arachnids, Araneae, spiders, museum collections, types, holotypes, paratypes. КЛЮЧЕВЫЕ СЛОВА: паукообразные, Araneae, пауки, музейные коллекции, типы, голотипы, паратипы.

ABSTRACT: A catalogue of 32 holotypes, 165 numerous inventory books on diverse invertebrate paratypes and two syntypes belonging to 50 spider groups (see Fig. 2 for Aranei), and (3) type index cards species in seven families (Agelenidae, Araneidae, (Fig. 3). Unfortunately, only a small part of this docu- Linyphiidae, Liocranidae, Lycosidae, Nesticidae, Sal- mentation has been digitized to date (see Mikhailov, ticidae) from the collection of the Zoological Museum 2016b, for further details). of the Moscow State University is provided. Other In Moscow, the spider collection is the largest one depositories housing the remaining types of the listed among all Russian and ex-USSR natural history muse- species are given as well. ums, being even larger than those both of the Zoologi- How to cite this article: Mikhailov K.G. 2018. cal Institute, Russian Academy of Sciences in St. Pe- Arachnid types in the Zoological Museum of the Mos- tersburg, and of the Institute of Animal Systematics cow State University. II. Aranei 1 (Arachnida) // Ar- and Ecology, Siberian Branch, Russian Academy of thropoda Selecta. Vol.27. No.1. P.95–105. doi: Sciences in Novosibirsk. In the Museum, the spider 10.15298/arthsel. 27.1.12 type collection (Fig. 4) is kept separately from the main collection of this group (Fig. 5). РЕЗЮМЕ: Представлен список 32 голотипов, The following indices are used for arachnids in the 165 паратипов и 2 синтипов, относящихся к 50 ви- registers of the Zoological Museum of the Moscow дам пауков (Aranei). Они принадлежат к 7 семей- State University: Ta — spiders (Aranei, or Araneae), ствам (Agelenidae, Araneidae, Linyphiidae, Liocra- Tb — scorpions (Scorpiones), Tc — solpugids (Solif- nidae, Lycosidae, Nesticidae, Salticidae) и хранятся в ugae), Td — mites and ticks (Acari), Tk — harvestmen Зоологическом музее МГУ. Также перечислены (Opiliones), Tl — pseudoscorpions (Pseudoscorpiones), другие хранилища, в которых размещены другие Tm — whip scorpions (Uropygi; currently no types). типы указанных видов. Altogether, the types of 50 species representing seven families (Agelenidae, Araneidae, Linyphiidae, Introduction Liocranidae, Lycosidae, Nesticidae, Salticidae) are list- ed below, including 32 holotypes, 165 paratypes and This paper is the second in the series cataloging the two syntypes; 96 accession numbers have been allocat- arachnid types deposited in the Zoological Museum of ed. The Museum card index of the spider type collec- the Moscow State University, starting with the spider tion is arranged alphabetically by genus name as origi- type collection, the largest one in the Museum. As nally described: Acantholycosa F. Dahl, 1908, Aeluril- indicated earlier [Mikhailov, 2016b], Museum’s inver- lus Simon, 1884, Agyneta Hull, 1911, etc. The follow- tebrate type collections include holo-, para- and syn- ing list of types presents only species belonging to the types of over 3,000 species, of which some 1,000 spe- genera whose names start from ‘A’. cies belong to Aranei. The Museum’s arachnid collec- tion was founded in the 1860s and currently contains Methods more than 200,000 arachnid specimens, exclusive of Acari [Mikhailov, 2016a]. The following sequence of data is accepted in the Traditionally, at least since the 1930s, the follow- type catalogue: species name in the original descrip- ing handwritten documentation has been accepted in tion; a reference to the first description; transcription the Museum: (1) departmental register (Fig. 1), (2) of data labels; list of other collections where the re- 96 K.G. Mikhailov

Fig. 1. An acquisition book of the ZMMU Invertebrate Department. Рис. 1. Книга поступлений отдела беспозвоночных Зоологического музея МГУ. maining types of the respective species are kept; re- abouts of other types, except those that are deposited in marks; current taxonomic status of the species. the ZMMU, is given on the basis of original descrip- When necessary, label abbreviations are put in an- tions. In the ZMMU, museum accession numbers for gled brackets, <>. Author’s comments to the labels are Aranei are indexed with ‘Ta’ which is put at the begin- given in square brackets, []. An English translation of ning of museum labels. Labels are transcribed literally, the original Russian labels is provided in figure brack- with mistakes being indicated in comments. ets, {}. The following translations of the administrative units Museums or personal collections are abbreviated as are used hereinafter: Area for “область, oblast”, Dis- follows: ISEA — Institute of Animal Systematics and trict for “район, rayon”, Province for “край, kray”. Ecology, Siberian Branch, Russian Academy of Sci- The nomenclature follows WSC [2018]. ences (Novosibirsk, Russia, curator G.N. Azarkina); JW — personal collection of Jörg Wunderlich (Germa- Type catalogue ny); MMUE — Manchester Museum, University of Manchester (UK, curator D.V. Logunov); NHRS — aborigenica Zyuzin et Marusik, 1988, Acantholy- Naturhistoriska Riksmuseet, Stockholm (Sweden, cu- cosa rator G. Lindberg); PSU — Department of Invertebrate Zyuzin, Marusik, 1988: 1083–1084, figs. Zoology, Perm State University (Russia, curator S.L. Ta-7342. Paratype $, Приморье, пос. Хасан {Primorie, Esyunin); SMF — Senckenberg Museum (Frankfurt Khasan}, 20.VIII.1981, А.К. Панютин {A.K. Panyutin}. am Main, Germany, curator P. Jäger); YM — personal The holotype and other paratypes are kept in ZIN. collection of Yuri M. Marusik (Magadan, Russia and In the original label, the collector name is misspelled as “Пантюлин {Pantyulin}”. Turku, Finland); ZIN — Zoological Institute, Russian Current status. Acantholycosa aborigenica Zyuzin et Academy of Sciences (St. Petersburg, Russia, curator Marusik, 1988 (Lycosidae). V.A. Krivokhatky); ZMMU — Zoological Museum, Moscow State University (Moscow, Russia, curator affinisoides Tanasevitch, 1984, Agyneta (Aprola- K.G. Mikhailov); ZMUT — Zoological Museum, Uni- gus) versity of Turku (Finland, curator S. Koponen). Where- Tanasevitch, 1984: 49–50, рис.3 (А–Ж). Arachnid types in the Zoological Museum of the Moscow State University. II. Aranei 1 97

Ta-4197. Holotype #, Эвенкия, р. Кочечум (правый приток <р.> Нижней Тунгуски), 40 км выше устья, лист- венничная тайга, подстилка {Evenkiya, Kochechum River (right tributary of Nizhnaya Tunguska River, 40 km up- stream of the mouth, larch forest, litter}, 3.VIII.1979, А.А. Вахрушев {A.A. Vakhrushev}. Other types are kept in ZIN. Current status. Agyneta affinisoides Tanasevitch, 1984 (Linyphiidae).

agrestis Ponomarev, 2007, Agroeca Ponomarev, 2007: 94–95, fig. 37. Ta-6880. Holotype $, Казахстан, Атырауская обл., пос. Кульсары, в квартире {Kazakhstan, Atyrau Area, Kulsary, indoor}, 12.05.1986, В. Романов {V. Romanov}. One paratype ($) is not yet transferred to ZMMU. Current status. Agroeca agrestis Ponomarev, 2007 (Li- ocranidae).

altaiensis Marusik, Azarkina et Koponen, 2004, Acantholycosa Marusik et al., 2004: 105, figs 1–9, 56, map 1. Ta-7538. Paratype $, Altai Province, Charyshskoe Dis- tr., Tigirek Mt. Range, Uba Pass, kurums [=scree], 50°53′N, 83°48′E, 27.07.1999, G. Azarkina et D. Ryzhkov. The holotype and other paratypes are kept in ISEA. Current status. Acantholycosa altaiensis Marusik, Az- arkina et Koponen, 2004 (Lycosidae).

Fig. 2. ZMMU registers for Aranei: above — the old register (1931–1984), below — the new one (2009–). ater Kroneberg, 1875, Aelurops Kroneberg, 1875: 50–51, tab. V, fig. 38. Рис. 2. Инвентарные книги пауков (Aranei) Зоологического Ta-1051. Holotype # [without palps], Туркестанская музея МГУ: сверху — старая книга (1931–1984), снизу — новая книга (с 2009 г.). Ученая Экспедиция Императорского Общества Люби- 98 K.G. Mikhailov

Fig. 3. An example of a ZMMU type index card, recto and verso. Рис. 3. Образец типовой карточки Зоологического музея МГУ, спереди и сзади. Arachnid types in the Zoological Museum of the Moscow State University. II. Aranei 1 99

Fig. 4. The type collection of Aranei in ZMMU. Photo by Fig. 5. The main collection of Aranei in ZMMU. Photo by A.V. A.V. Sysoev. Sysoev. Рис. 4. Типовая коллекция пауков (Aranei) Зоологичес- Рис. 5. Основная коллекция пауков (Aranei) Зоологического кого музея МГУ. Фото А.В. Сысоева. музея МГУ. Фото А.В. Сысоева.

телей Естествознания. <А.П.> Федченко {Turkestan Sci- Types are kept in ZMMU only. entific Expedition by the Imperator Society for Friends of In the original description, the misspelled name ‘azerba- Natural Sciences. A.P. Fedchenko}. 14. Ташкент {Tash- jdzanica’ is used in figure captions and a general species list. kent}. 4.III. Current status. Synonymized with Aelurillus concolor The species was described from the holotype only. Kulczyński, 1901 (Salticidae) [Azarkina, 2003, initially with Current status. Aelurillus ater (Kroneberg, 1875) (Salti- A. muganicus Dunin, 1984; see Azarkina, Mirshamsi, 2014 cidae). for further details].

azarkinae Marusik et Omelko, 2011, Acantholycosa brutus Wesołowska, 1996, Aelurillus Marusik, Omelko, 2011: 2–5, figs 1–5, 8–12, 26–28. Wesołowska, 1996: 23–24, figs 4A, B. Ta-7714. Holotype #, Russia, Maritime Prov., Lazovskii Ta-7501. Holotype $, USSR, Turkmenia, South Ustyurt, Distr., Sestra Mt., 43°31′52.23″N, 134°02′49.44″E, scree, Kaplankyrsky Reserve, 2.04.<19>85, leg. L.A. Mitroshina. 20.06.2005, M.M. Omelko. The species was described from the holotype only. Ta-7715. Paratype $, Russia, Maritime Prov., Lazovskii Current status. Aelurillus brutus Wesołowska, 1996 (Sal- Distr., Sestra Mt., 43°31′52.23″N, 134°02′49.44″E, scree, ticidae). 20.06.2005, M.M. Omelko. Other types are kept in the collection of Gornotaezhnaya bulavintsevi Tanasevitch, 2016, Agyneta Station (Ussuriysk Distr., Maritime Prov.). Tanasevitch, 2016: 75, figs 1–8. Current status. Acantholycosa azarkinae Marusik et Ta-7750. Holotype #, Якутия, окр. пос. Тикси {Yaku- Omelko, 2011 (Lycosidae). tia, nr. Tiksi}, VIII.1985, leg. BulavintsevV. The species was described from the holotype only. azerbajdzanicus Dunin, 1984, Aelurillus Current status. Agyneta bulavintsevi Tanasevitch, 2016 Dunin, 1984: 51, fig. 11. (Linyphiidae). Ta-6790. Holotype $, Azerbaijan, Baku, Zykh, 8.06.1979, coll. P.M. Dunin. caucasica Guseinov, Marusik et Koponen, 2005, Ta-6791. Paratype $, Azerbaijan, Baku, Zykh, 8.06.1979, Agelescape coll. P.M. Dunin. Guseinov et al., 2005: 157, figs 9–12, 69–71, 105. Ta-6792. Paratype $, Azerbaijan, Apsheron Peninsula, Ta-7506. Holotype $, CN Azerbaijan, Ismailly Distr., Dyubendy, 21.05.1977, coll. P.M. Dunin. env. of Khanaya Vill., 700 m, 8.07.2001, E.F. Guseinov. 100 K.G. Mikhailov

Ta-7507. Paratypes 2 $$, CN Azerbaijan, Ismailly Dis- helvenacius Logunov, 1993, Aelurillus tr., env. of Khanaya Vill., 700 m, 1.10.2003, E.F. Guseinov. Logunov, 1993: 47–49, fig. 1a,b. Types are kept in ZMMU only. Ta-4779. Holotype $, Монголия, Эхийн-Гол {Mongo- Current status. Agelescape caucasica Guseinov, Marusik lia, Ekhiin-Gol}, оазис {oasis}, сухой чий {dry Achna- et Koponen, 2005 (Agelenidae). therum}, 30.VII.1983, собр. Н.<Б.> Ананьева {coll. N.B. Ananieva}. cuneata Tanasevitch, 2014, Agyneta The species was described from the holotype alone. Tanasevitch, 2014: 77, figs 1–9. Current status. Aelurillus helvenacius Logunov, 1993 Ta-7696. Holotype #, Амурская обл., Селемджинский (Salticidae). р-н, о-в на р. Бысса, 2 км выше устья руч. Кукуя {Amur Area, Selemdzhinsk Distr., an island on Byssa River, 2 km khakassica Marusik, Azarkina et Koponen, 2004, upstream of Kukuy Creek’s mouth}, 6.VI.2007, Е.<М.> Acantholycosa Веселова, А.<Б.> Рывкин {E.M. Veselova, A.B. Ryvkin}. Marusik et al., 2004: 119, figs 100–103, map 1. The species was described from the holotype only. Ta-7541. Paratypes 10 ##, Russia, SW Khakassia, Current status. Agyneta cuneata Tanasevitch, 2014 Abakan Mt. Range, Choosek Mt. Range, 20 km SSE of (Linyphiidae). Mrassu, talus, 1600–1800 m, 7–19.07.1999, D. Lomakin. The holotype and other paratypes are kept in ISEA. dubatolovi Azarkina, 2003, Aelurillus Current status. Acantholycosa khakassica Marusik, Az- Azarkina, 2003: 95–97, figs 29–50, Table 1. arkina et Koponen, 2004 (Lycosidae). Ta-6586. Paratypes 5 ##, 1 $, Северный Таджикистан, хр. Моголтау {northern Tadzhikistan, Mogoltau Mt. Range}, kopetdaghensis Tanasevitch, 1989, Agyneta (Meio- подгорная равнина {foothill plain}, 27.02.1983, coll. А.Б. neta) Ненилин {A.B. Nenilin}. Tanasevitch, 1989: 89–91, figs 1–5. Ta-6587. Paratypes 2 ##, 1 $, Туркмения, Ц<ентр>. Ta-5430. Holotype #, Туркмения, <Юго->Зап. Ко- Копетдаг, ~20 км Ю Геок-Тепе, гора Душак {Turkmenia, петдаг, ущ. Айдере {Turkmenia, SW Kopetdagh, Aidere Central Kopetdagh, ca 20 km S of Geok-Tepe, Dushak Mt.}, Canyon}, в подстилке грецкого ореха {Juglans regia, lit- 2300 м {m}, юж. склон {S slope}, арчёвое редколесье ter}, 16.II–11.III.1979, собр. В.Я. Фет {coll. V.Ya. Fet}. {sparse Juniperus stand}, 20.04.1989, coll. К.Г. Михайлов Ta-5431. Paratypes 2 ##, Туркмения, <Юго->Зап. {K.G. Mikhailov}. Копетдаг, ущ. Айдере {Turkmenia, SW Kopetdagh, Aidere Ta-6588. Paratype $, Uzbekistan, Kuraminskii Mt. Canyon}, подстилка {litter}, 18.III–8.IV.1979, собр. В.Я. Range, Bash-Kul, 12.04.1985, coll. S.V. Ovchinnikov. Фет {coll. V.Ya. Fet}. Ta-6589. Paratype $, Казахстан, Джезказганская обл., Other types are kept in SMF. Балхашск. р-н, пос. Хасхабье {Kazakhstan, Dzhezkazghan Current status. Agyneta kopetdaghensis Tanasevitch, Area, Balkhash Distr., Khaskhabye}, VI.1988, О. Ильченко 1989 (Linyphiidae). {O. Ilchenko}. Other types are kept in ISEA (holotype, paratypes), kurchumensis Marusik, Azarkina et Koponen, 2004, MMUE and NHRS. Acantholycosa Current status. Aelurillus dubatolovi Azarkina, 2003 (Sal- Marusik et al., 2004: 119, figs 82–83, map 1. ticidae). Ta-6777. Paratype $, Kazakhstan, East-Kazakhstan Area, Kurchum Mt. Range, Kurchum River upper flow, 23.08.1990, dudkorum Marusik, Azarkina et Koponen, 2004, V.K. Zinchenko. Acantholycosa Other types are kept in ISEA (holotype) and JW (now in Marusik et al., 2004: 113–115, figs 59, 73–79, 84–85, map 1. SMF). Ta-7539. Holotype #, Altai, 50 km W of Kosh-Agach, Current status. Synonymized with Acantholycosa katun- ca. 20–25 km W of Bel’tir, Taltura (Chagan-Uzun) River ensis Marusik, Azarkina et Koponen, 2004 (Lycosidae) [Fo- canyon, 2900–3100 m, mountain stone steppe, 26–28.07. michev et al., 2016]. 1999, V.V. Glupov. Ta-7540. Paratypes 3 $$, Altai, 50 km W of Kosh- laimonasi Tanasevitch, 2006, Agyneta Agach, ca. 20–25 km W of Bel’tir, Taltura (Chagan-Uzun) Tanasevitch, 2006: 29–30, figs 1–7. River canyon, 2900–3100 m, mountain stone steppe, 26– Ta-7192. Holotype #, Russia, Khabarovsk Province, 28.07.1999, V.V. Glupov. Verkhnebureinsky District, Bureinsky N, Other types are kept in ISEA and ZMUT. upper reaches of Pravaya Bureya River, source of Lednik- Current status. Acantholycosa dudkorum Marusik, Az- ovyi River (ca. 52°08′N, 134°26′E), montane tundra, 30.VI. arkina et Koponen, 2004 (Lycosidae). 2000, L. Trilikauskas. The species was described from the holotype alone. dunini Guseinov, Marusik et Koponen, 2005, Age- Current status. Agyneta laimonasi Tanasevitch, 2006 lescape (Linyphiidae). Guseinov et al., 2005: 158, figs 1, 2, 6–8. Ta-7508. Holotype #, CW Azerbaijan, Khojavend (=Ga- laniger Logunov et Marusik, 2000, Aelurillus drut) Distr., env. of Azykh Vill., 12.07.1977, P.M. Dunin. Logunov, Marusik, 2000: 265, figs 1–6. Ta-7509. Paratype $, CW Azerbaijan, Khojavend (=Ga- Ta-6304. Paratypes 3 ##, 2 $$, Kazakhstan, Pavlodar drut) Distr., env. of Azykh Vill., 12.07.1977, P.M. Dunin. Area, ~1 km W of Ermak (=Aksu), 14–28.06.1998, leg. Types are kept in ZMMU only. A.V. Gromov. Current status. Agelescape dunini Guseinov, Marusik et Other types are kept in ISEA (holotype, paratypes) and Koponen, 2005 (Agelenidae). NHRS. Arachnid types in the Zoological Museum of the Moscow State University. II. Aranei 1 101

Current status. Aelurillus laniger Logunov et Marusik, Ta-6708. Paratype #, Iran, Fars Prov., Haft-barm, 52° 2000 (Salticidae). 15′E, 29°45′N, 24.05.2000, Yu. Marusik. Other types are kept in ISEA, MMUE, YM (holotype, levii Tanasevitch, 1984, Agyneta (Aprolagus) now in SMF) and Dept. of Agricultural Zoology, Plant Pests Tanasevitch, 1984: 50–51, рис. 4 (А–З). and Diseases Research Institute (Tehran, Iran). Ta-4198. Holotype #, Енисей, Мирное {Enisei, Mir- Current status. Aelurillis marusiki Azarkina, 2003 (Sal- noe} (62°20′ сев.шир. {N}), пойменные кустарниковые ticidae). заросли {floodplain shrubs}, 9.VIII.1979, leg. К.Ю. Еськов {K.Yu. Eskov}. metatarsalis Tanasevitch, 2014, Agyneta Ta-4199. Paratypes 3 ##, 10 $$, Енисей, Мирное Tanasevitch, 2014: 78–79, figs 10–17. {Enisei, Mirnoe}, тальники {willows}, 9.VIII.1979, leg. Ta-7697. Holotype #, Респ. Алтай, Чойский р-н, окр. К.Ю. Еськов {K.Yu. Eskov}. горы Балтырган {Altai Republic, Choya Distr., environs of Ta-4200. Paratype $, Енисей, Мирное {Enisei, Mir- Baltyrgan Mt.}, 2100 м {m}, 4.8.2001, Н.В. Левина {N.V. noe}, в речных наносах {in river drifts}, 5.VIII.1978, leg. Levina}. К.Ю. Еськов {K.Yu. Eskov}. The species was described from the holotype alone. Ta-4201. Paratypes 2 ##, Енисей, Мирное, левый берег Current status. Agyneta metatarsalis Tanasevitch, 2014 Енисея {Enisei, Mirnoe, Enisei Rever left bank}, тальники (Linyphiidae). {willows}, 23.V.1978, leg. К.Ю. Еськов {K.Yu. Eskov}. Ta-7245. Paratypes 2 ##, 4 $$, тальники {willows}, minimontanus Azarkina, 2002, Aelurillis 9.VIII.1979, К.Ю. Еськов {K.Yu. Eskov}. Azarkina, 2002: 257–258, figs 54–63. Types are kept in ZMMU only. Ta-6709. Paratypes 1 #, 1 $, IN-14 India, Himachal Current status. Agyneta levii Tanasevitch, 1984 (Linyphi- Pradesh, Sissu Vill., 32°28.1′N, 77°07.9′E, 3150–3500 m, idae). 8–10.06.1999, Yu. Marusik. Other types are kept in ISEA, MMUE and YM (holo- levinae Marusik, Azarkina et Koponen, 2004, Acan- type, now in SMF). tholycosa Current status. Aelurillis minimontanus Azarkina, 2003 Marusik et al., 2004: 112, figs 61–65, map 1. (Salticidae). Ta-6775. Holotype #, Altai, Katun’ Reserve, kurums [=scree], traps, 11.07.2000, N. Levina. muganicus Dunin, 1984, Aelurillus Ta-7542. Paratype #, Atlai, Sredneye Multinskoye Lake, Dunin, 1984: 50–51, figs 12, 13. ca. 30 km SE of Ust-Koksa, 1600–2200 m, 30.07–5.08.1994, Ta-6788. Paratype $, Азербайджан, о-в Глиняный, S.I. Golovatch, A.B. Ryvkin. напротив п. Алят {Azerbaijan, Glinyanyi Is., opposite Alyat Types are kept in ZMMU only. Vil.}, 15.07.1979, сб. П.М. Дунин {coll. P.M. Dunin}. Current status. Acantholycosa levinae Marusik, Azarki- Ta-6789. Paratypes 4 ##, Azerbaijan, Apsheron Pen- na et Koponen, 2004 (Lycosidae). insula, Bina, 3.10.1973, coll. P.M. Dunin. Whereabouts of other types (holotype # and paratype $) is not known. levyi Guseinov, Marusik et Koponen, 2005, Age- In the original description, the misspelled name A. mugan- lescape ica is used in figure captions and a general species list. Guseinov et al., 2005: 158, figs 3–5. Current status. Synonymized with Aelurillus concolor Ta-7510. Holotype #, CN Azerbaijan, Ismailly Distr., Kulczyński, 1901 (Salticidae) [Azarkina, Mirshamsi, 2014]. env. of Khanaya Vill., 700 m, 06.2002, E.F. Guseinov. The species was described from the holotype alone. nataliae Esyunin et Efimik, 1998, Aituaria Current status. Agelescape levyi Guseinov, Marusik et Esyunin, Efimik, 1998: 147, figs 1–2, 4–7. Koponen, 2005 (Agelenidae). Ta-6232. Holotype #, Оренбургская область, Куван- дыкский р-н, пос. Айтуарка {Orenburg Area, Kuvandyk logunovi Marusik, Azarkina et Koponen, 2004, Distr., Aituarka}, в помещении {indoor}, IX.1996, собр. Acantholycosa Н.С. Мазура {coll. N.S. Mazura}. Marusik et al., 2004: 131, figs 88–91, 129, 173–180, map 1. The species was described from the holotype alone. Ta-6770. Paratype #, Altai, 50 km W of Kosh-Agach, Current status. Synonymized with Aituaria pontica ca. 20–25 km W of Bel’tir, Taltura (Chagan-Uzun) River (Spassky, 1932) (Nesticidae) [Esyunin, 2017]. canyon, 2200–2300 m, stony debris on the upper border of Larix forest, 25–30.06.1999, D. Logunov. oligerae Marusik, Azarkina et Koponen, 2004, Ta-6771. Paratype $, Altai, Chiri Lake, 1800–2000 m, Acantholycosa 30.07.1997, A. Tanasevitch. Marusik et al., 2004: 126–128, figs 19–20, 128, 152–161, Other types are kept in ISEA (holotype, paratypes) and map 2. ZMUT. Ta-6772. Holotype #, Russia, Maritime Province, Lazo Current status. Acantholycosa logunovi Marusik, Az- Reserve, Sukhoi Klyuch site, edge of stony debris, on slope, arkina et Koponen, 2004 (Lycosidae). 22.06.1981, T. Oliger. Ta-6773. Paratypes 2 $$, Russia, Maritime Province, marusiki Azarkina, 2002, Aelurillis Lazo Reserve, Sukhoi Klyuch site, edge of stony debris, on Azarkina, 2002: 255–257, figs 44–53. slope, 22.06.1981, T. Oliger. Ta-6707. Paratype $, Iran, Fars Prov., bridge on Ghara- Ta-6774. Paratype #, Russia, Maritime Province, Lazo Aghatch River, 52°13′E, 29°49′N, 24.05.2000, Yu. Reserve, Sukhoi Klyuch site, mixed forest, N exposed slope, Marusik. 23.06.1981, T. Oliger. 102 K.G. Mikhailov

Other types are kept in JW (now in SMF) and ZMUT. River, 40 km upstream of the mouth, larch forest, litter}, Current status. Acantholycosa oligerae Marusik, Az- гарь, подстилка {burnt forest, litter}, 5.VIII.1978, А.А. arkina et Koponen, 2004 (Lycosidae). Вахрушев {A.A. Vakhrushev}. Ta-4194. Paratype $, Эвенкия, р. Кочечум (правый paraplumalis Marusik, Azarkina et Koponen, 2004, приток Нижней Тунгуски), 40 км выше устья {Evenkiya, Acantholycosa Kochechum River (right tributary of Nizhnaya Tunguska Marusik et al., 2004: 112, figs 38–43, 49–52, 55, map 1. River, 40 km upstream of the mouth}, гарь, подстилка Ta-6758. Holotype #, Altai, Choya Distr., Boityrgan {burnt forest, litter}, 5.VIII.1978, А.А. Вахрушев {A.A. Mt., stony mountain tundra, 12.08.2001, N. Levina. Vakhrushev}. Ta-6759. Paratype $, Altai, Choya Distr., Boityrgan Mt., Types are kept in ZMMU only. stony mountain tundra, 12.08.2001, N. Levina. Current status. Agyneta pseudosaxatilis Tanasevitch, Ta-6760. Paratypes 2 $$, Altai, Choya Distr., Boityrgan 1984 (Linyphiidae). Mt., mountain meadow, 13.08.2001, N. Levina. Ta-6761. Paratype $, Altai, Choya Distr., Boityrgan Mt., ripariensis Tanasevitch, 1984, Agyneta (Aprolagus) mountain meadow, 13.08.2001, N. Levina. Tanasevitch, 1984: 48–49, ðèñ. 2 (À–Å). Ta-6762. Paratype $, Altai, Choya Distr., Boityrgan Mt., Ta-4195. Holotype #, Полярный Урал (вост. макро- stony debris, 4.08.2001, N. Levina. склон), долина р. Собь, 20 км к западу от пос. Харп {Polar Ta-6763. Paratypes 4 $$, Altai, Choya Distr., Boityrgan Urals (eastern macroslope), Sob’ River valley, 20 km W of Mt., stony tundra, 12.08.2001, N. Levina. Kharp}, участок берега, занятый ерниковой (Betula nana) Ta-6764. Paratypes 5 $$, Altai, Choya Distr., Boityrgan мохово-лишайниковой ассоциацией, подстилка {river bank Mt., mountain meadow, 13.08.2001, N. Levina. with dwarf birch (Betula nana) moss-lichen association}, Ta-6765. Paratypes 2 $$, Altai, Choya Distr., Boityrgan 19.VIII.1983, leg. А.В. Танасевич {A.V. Tanasevitch}. Mt., stony tundra, 12.08.2001, N. Levina. Ta-4196. Paratypes 2 ##, 5 $$, Коми АССР, Вор- Ta-6766. Paratype #, Altai, Choya Distr., Boityrgan кутинский р-н, окр. пос. Мульда {Komi Republic, Vor- Mt., forest tundra, 14.08.2001, N. Levina. kuta Distr., nr. Mul’da}, тундра, склон холма, кустар- Ta-6767. Paratypes 2 $$, Altai, Choya Distr., Boityrgan ничковая мохово-лишайниковая ассоциация {tundra, hill Mt., stony tundra, 12.08.2001, N. Levina. slope, shrubby moss-lichen association}, 11.IX.1983, leg. Ta-6768. Paratype $, Altai, Choya Distr., Boityrgan Mt., А.В. Танасевич {A.V. Tanasevitch}. stony tundra, 12.08.2001, N. Levina. Ta-7955. Paratypes 1 #, 2 $$, Полярный Урал, долина Ta-6769. Paratype $, Altai, Choya Distr., Boityrgan Mt., р. Собь, 20 км З от п. Харп, берег р. Собь {Polar Urals, mountain meadow, 13.08.2001, N. Levina. Sob’ River valley, 20 km W of Kharp, Sob’ River bank}, Ta-7543. Paratype $, Altai, Choya Distr., Boityrgan Mt., подстилка {litter}, 19.8.1983, leg. А.<В.> Танасевич {A.V. forest tundra and mountain tundra, 14.08.2001, N. Le- Tanasevitch}. vina. Other types are kept in ZIN. Other types are kept in JW (now in SMF), MMUE, Current status. Agyneta ripariensis Tanasevitch, 1984 NHRS, ZMUT. (Linyphiidae). Current status. Acantholycosa paraplumalis Marusik, Azarkina et Koponen, 2004 (Lycosidae). saaristoi Tanasevitch, 2000, Agyneta Tanasevitch, 2000: 206–209, figs 48–68. petrophila Marusik, Azarkina et Koponen, 2004, Ta-7843. Paratype #, КЗО [=Кзыл- Acantholycosa Ординская обл.], Яныкурганский р-н, 35 км NNO г. Яны- Marusik et al., 2004: 119, figs 104–107, map 1. курган, хр. Каратау {Kzyl-Orda Area, Yanykurgan Distr., Ta-7544. Paratypes 10 ##, Russia, Khakassia, West- 35 km NNE of Yanykurgan, Karatau Mt. Ridge}, 15.6. ern Sayan, 50 km SSW of Tashtyp, central part of Khansyn <19>89 <, leg. A.A. Zyuzin>. Mt. Range, scree, 1800–2100 m, 11–24.07.2000, D. Lo- Other types are kept in ISEA, PSU and ZMUT. makin. The holotype and several paratypes are not yet trans- The holotype and other paratypes are kept in ISEA. ferred by the author to ZMMU. Current status. Acantholycosa petrophila Marusik, Az- Current status. Agyneta saaristoi Tanasevitch, 2000 arkina et Koponen, 2004 (Lycosidae). (Linyphiidae).

plumalis Marusik, Azarkina et Koponen, 2004, sakhalinensis Saaristo et Marusik, 2004, Agy- Acantholycosa phantes Marusik et al., 2004: 109–112, figs 30–37, 44–48, map 1. Saaristo, Marusik, 2004: 79, figs 7, 12–13, 19–21. Ta-6757. Paratypes 3 $$, Altai, Teletskoye Lake, Chiri Ta-7240. Holotype #, Sakhalin Island, central part, Le- Vil., scree, 30.07.1997, A. Tanasevitch. onidovka River (right tributary of Poronai River), 30 km The holotype and other paratypes are kept in ISEA. downstream from Ulyanovka River mouth, 49°15.092′N, Current status. Acantholycosa plumalis Marusik, Az- 142°43.889′E, 4–5 August 2001, Y.M. Marusik. arkina et Koponen, 2004 (Lycosidae). Ta-7241. Paratypes 2 $$, 4 juv., Sakhalin Island, central part, Leonidovka River (right tributary of Poronai River), 30 pseudosaxatilis Tanasevitch, 1984, Agyneta (Apro- km downstream from Ulyanovka River mouth, 49°15.092′N, lagus) 142°43.889′E, 4–5 August 2001, Y.M. Marusik. Tanasevitch, 1984: 47–48, рис. 1 (А–З). Ta-7242. Paratype $, Sakhalin Island, central part, Le- Ta-4193. Holotype #, Эвенкия, р. Кочечум (правый onidovka River (right tributary of Poronai River), 30 km приток Нижней Тунгуски), 40 км выше устья {Evenkiya, downstream from Ulyanovka River mouth, 49°15.092′N, Kochechum River (right tributary of Nizhnaya Tunguska 142°43.889′E, 4–5 August 2001, Y.M. Marusik. Arachnid types in the Zoological Museum of the Moscow State University. II. Aranei 1 103

Other types are kept in JW and ZMUT. sundukovi Marusik, Azarkina et Koponen, 2004, Current status. Agyphantes sakhalinensis Saaristo et Acantholycosa Marusik, 2004 (Linyphiidae). Marusik et al., 2004: 128, figs 162–167, map 2. Ta-7546. Holotype #, Russia, Maritime Province, Lazo sayanensis Marusik, Azarkina et Koponen, 2004, Reserve, Amerika kordon, 43°16′N, 134°03′E, 14–17.05. 1999, Yu. Sundukov. Acantholycosa The species was described from the holotype only. Marusik et al., 2004: 107–109, figs 10–14, map 1. Current status. Acantholycosa sundukovi Marusik, Az- Ta-7545. Paratype #, Krasnoyarsk Province, Yerma- arkina et Koponen, 2004 (Lycosidae). kovski Distr., Abakan–Kyzyl Hwy 612th km, Oiskoye Lake, 5.07.1989, D.L. Grodnitskiy. The holotype is kept in ISEA. talyshica Guseinov, Marusik et Koponen, 2005, Current status. Acantholycosa sayanensis Marusik, Az- Agelescape arkina et Koponen, 2004 (Lycosidae). Guseinov et al., 2005: 158–159, figs 13–16. Ta-7519. Holotype $, SE Azerbaijan, Lerik Distr., env. of Lyalyakeran Vill., 1600 m, 23.08.1995, E.F. Guseinov. serpentina Guo et Zhang, 2010, Aculepeira Paratypes (7 $$) have not yet been transferred to ZMMU. Guo, Zhang, 2010: 261–263, figs 1–5. Current status. Agelescape talyshica Guseinov, Marusik Ta-7700. Paratype $, Qiyi Farm (29°38′N, 91°02′E), et Koponen, 2005 (Agelenidae). Lhasa City, Tibet Autonomous Region, 30 July 2002, Feng Zhang leg. tianschanica Tanasevitch, 1989, Agyneta (Meio- The holotype and other paratypes are kept in the Muse- neta) um of Hebei University (Baoding, China). Tanasevitch, 1989: 92, figs 6–11. Current status. Aculepeira serpentina Guo et Zhang, Ta-5436. Holotype #, Киргизия, Сары-Челекский 2010 (Araneidae). запов., Каратоко {Kirghizia, Sary-Chelekskiy Reserve, Kara- toko}, сухой альп<ийский> луг, под камнями {dry alpine spirembolus Marusik, Azarkina et Koponen, 2004, meadow, under stones}, 3250 м {m}, 21.IX.1983, собр. Acantholycosa А.Б. Рывкин {coll. A.B. Ryvkin}. Marusik et al., 2004: 125–126, figs 143–146, map 1. Ta-5437. Paratypes 1 #, 2 $$, Киргизия, Сары- Ta-6776. Paratype #, Altai, Kholzun Mt. Range, upper Челекский запов., Каратоко {Kirghizia, Sary-Chelekskiy reaches of left tributary of Bannaya River, 2000–2250 m, Reserve, Karatoko}, сухой альп<ийский> луг, под камнями mountain tundra, 13–14.06.1999, A. Dudko, R. {dry alpine meadow, under stones}, 3250 м {m}, 21.IX.1983, Dudko. собр. А.Б. Рывкин {coll. A.B. Ryvkin}. The holotype is kept in ISEA. Ta-5438. Paratypes 1 #, 16 $$, Зап. Тянь-Шань, Фер- Current status. Acantholycosa spirembolus Marusik, ганский хр., окр. пос. Арсланбоб {W Tien-Shang, Fer- Azarkina et Koponen, 2004 (Lycosidae). ghanskiy Mt. Range, environs of Arslanbob}, сухой суб- альп<ийский> луг, под камн<ями> {dry alpine meadow, under stones}, 2200 м {m}, 1.X.1983, собр. К.Ю. Еськов subnivalis Tanasevitch, 1989, Agyneta (Meioneta) {coll. K.Yu. Eskov}. Tanasevitch, 1989: 91–92, figs 16–18. Ta-5439. Paratype $ [epigyne only], Зап. Тянь-Шань, Ta-5432. Holotype #, Киргизия, Киргизский хр., пер. Ферганский хр., окр. пос. Арсланбоб {W Tien-Shang, Тюя-Ашу {Kirghizia, Kirghizsky Mt. Ridge, Tyuya-Ashu Ferghanskiy Mt. Range, environs of Arslanbob}, сухой суб- Pass}, 3000–3400 м {m}, 6–8.VIII.1984, собр. С.Л. Зон- альп<ийский> луг, под камн<ями> {dry alpine meadow, штейн {coll. S.L. Zonstein}. under stones}, 2200 м {m}, 1.X.1983, собр. К.Ю. Еськов Ta-5433. Paratype # [palp only], Киргизия, Центр. {coll. K.Yu. Eskov}. Тянь-шань, сырты, окр. метеост. “Тянь-Шань” {Kirghi- Other types are kept in SMF. zia, Central Tien-Shang, syrts, environs of Tien-Shang me- Current status. Agyneta tianschanica Tanasevitch, 1989 teostation}, 3700–3900 м {m}, 15–25.vii.1983, собр. Б. (Linyphiidae). Павлюк {coll. B. Pavlyuk}. Ta-5434. Paratype #, Киргизия, Центр. Тянь-шань, tibialis Tanasevitch, 2005, Agyneta сырты, окр. метеост. “Тянь-Шань” {Kirghizia, Central Tanasevitch, 2005: 165–166, figs 1–8. Tien-Shang, syrts, environs of Tien-Shang meteostation}, Ta-7827. Holotype #, Алтай, Загриха {Altais, Za- 3700–3900 м {m}, 15–25.vii.1983, собр. Б. Павлюк {coll. grikha}, 1700 м {m}, 27.6.[20]00, Г.<Н.> Азаркина {G.N. B. Pavlyuk}. Azarkina}. Ta-5435. Paratype #, <Таджикистан,> Туркестанский The types should be kept in ZMMU only; one paratype хребет, Кусавли-сай {Tadjikistan, Turkestanskiy Mt. Ridge, (#) should still be transferred to ZMMU. Kusavli-Sai}, арчевник, под камн<ями> {Juniperus stand, Current status. Agyneta tibialis Tanasevitch, 2005 (Liny- under stones}, 2400–2700 м {m}, 29.IX.1970, собр. Е.М. phiidae). Андреева {coll. E.M. Andreeva}. Ta-7956. Paratypes 3 $$, <Киргизия,> Заалайский хр., uzbekistanica Tanasevitch, 1984, Agyneta (Meio- сев. склон, Бордоба {Kirghizia, Zaalayskiy Mt. Range, N neta) slope, Bordoba}, 3500 м {m}, 8.X-<19>70, Е.<М.> Анд- Tanasevitch, 1984: 51–52, ðèñ. 5 (À–Ç). реева {E.M. Andreeva}. Ta-4202. Holotype #, Узбекская ССР, Ташкентская Types are kept in ZMMU only. область, Бостанлыкский р-н, близ поселка Сиджак {Us- Current status. Agyneta subnivalis Tanasevitch, 1989 bek SSR, Tashkent Area, Bostanlyk Distr., near Sidzhak}, (Linyphiidae). склон горы (2000 м над уровнем моря), арчевое ред- 104 K.G. Mikhailov

колесье, под камнями {mountain slope (2000 m a.s.l.), G. Krivosheina (Moscow, Russia) are highly appreciated. Juniperus sparse stand, under stones}, 10.IV.1982, leg. А.В. This paper is supported by the Russian Science Foundation Танасевич {A.V. Tanasevitch}. Project No. 14-50-00029. The curatorial work with museum Other types are kept in ZIN. registers and other paper documentation (card indexes, etc.) Current status. Agyneta uzbekistanica Tanasevitch, 1984 was supported by the Moscow State University Project No. (Linyphiidae). AAAA-A16-116021660077-3.

variegatus Kroneberg, 1875, Aelurops References Kroneberg, 1875: 51, tab. V, fig. 39. Ta-1049. Syntype $, Туркестанская Ученая Azarkina G.N. 2002. New and poorly known species of the genus Экспедиция Императорского Общества Любителей Aelurillus Simon, 1884 from Central Asia, Asia Minor and the Естествознания. <А.П.> Федченко{Turkestan Scientific eastern Mediterranean // Bull. Brit. arachnol. Soc. Vol.12. Expedition by the Imperator Society for Friends of Natural Pt.6. P.249–263. Sciences. A.P. Fedchenko}. 16. Самарканд {Samarkand}. Azarkina G.N. 2003. Aelurillus ater (Kroneberg, 1875) and related Ta-1050. Syntypes 1 # [with the palp detached], 1 $, species of jumping spiders in the fauna of Middle Asia and the Туркестанская Ученая Экспедиция Императорского Об- Caucasus (Aranel: Salticidae) // Arthropoda Selecta. Vol.11(for щества Любителей Естествознания. <А.П.> Федченко 2002). No.1. P.89–107. {Turkestan Scientific Expedition by the Imperator Society Azarkina G.N., Mirshamsi O. 2014. 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Miscellaneous notes on Palae- The type locality is situated in East Kazakhstan Area arctic Salticidae (Arachnida: Aranei) // Arthropoda Selecta. (Kazakhstan) rather than in Russia, as stated in the original Vol.8 (for 1999). No.4. P.263–292. description. Marusik Yu.M., Azarkina G.N., Koponen S. 2004. A survey of Current status. Acantholycosa zinchenkoi Marusik, Az- East Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa and related new genera // Arthropoda Selecta. Vol.12 (for arkina et Koponen, 2004 (Lycosidae). 2003). No.2. P.101–148. Marusik Yu.M., Koponen S., Vinokurov N.N., Nogovitsyna S.N. zonsteini Marusik et Omelko, 2017, Acantholy- 2002. Spiders (Aranei) from the northernmost forest-tundra of cosa northeastern Yakutia (70°35´ N, 134°34´ E) with description Marusik, Omelko, 2017: 597–599, figs 1–9. of three new species // Arthropoda Selecta. Vol.10 (for 2001). Ta-8006. Holotype #, Khabarovsk Prov., Slavyanka, No.4. P.351–370. 1990, S. Golovatch. Marusik Yu.M., Omelko M.M. 2011. A survey of East Palaearctic The species was described from the holotype only. Lycosidae (Araneae). 7. A new species of Acantholycosa Dahl, Current status. Acantholycosa zonsteini Marusik et Om- 1908 from the Russian Far East // ZooKeys. Vol.79. P.1–10. elko, 2017 (Lycosidae). Marusik Yu.M., Omelko M.M. 2017. A new species of Acantholy- cosa baltoroi-group (Araneae: Lycosidae, Pardosinae) from . the Russian Far East // Zootaxa. Vol.4232. No.4. P.597–600. Acknowledgements. I am cordially obliged to my col- Mikhailov K.G. 2016a. Arachnid types in the Zoological Museum, league Dmitry V. Logunov (Manchester, UK) for reviewing Moscow State University. I. Opiliones (Arachnida) // Arthro- the manuscript and linguistic help. Comments from Galina poda Selecta. Vol.25. No.3. P.327–334. Arachnid types in the Zoological Museum of the Moscow State University. II. Aranei 1 105

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