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Home , Ahasverus (beetle), Bean weevil, Callosobruchus phaseoli, Cartodere, Cartodere constricta, Deathwatch beetle

Abound: A survey of beetles relevant to control professionals

PJ Liesch UW Diagnostic Lab [email protected], Twitter: @WiBugGuy

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Topics Covered in Today’s Presentation

1) General introduction to beetles

2) Survey of beetles relevant to the industry • Examined records from the UW Insect Diagnostic Lab over the last 5+ years

• Identified top ~24 beetles associated with structural pest control in Wisconsin and Upper Midwest • in/around structures, common ID requests at IDL, and species commonly misidentified

• Slides will emphasize diagnostics and biology of species with relevant notes about management

• Detailed information on size, appearance, key ID features, biology, and look-alikes

• Slides available on UW Insect Diagnostic Lab website: insectlab.russell.wisc.edu (Resources > Presentation Notes)

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2 Meet the Beetles: Order Coleoptera

• First appeared ~300 MYA

• Largest order of (and ) on the planet: • Worldwide: 176 Families & ~400,000 species worldwide • US: 130 Families & ~25,000 species • Wisconsin: 100+ Families & ~5,000 - 6,000 beetles species

• Nearly 1 out of every 4 extant species is a type of beetle! Vertebrates 3% • Undergo complete metamorphosis: > > > adult Other Animals 19% • Hard body (typically); possess protective wing covers elytra

• Can vary greatly in size and appearance! Crustaceans Other Insects 4% 48% Why are beetles tough to ID? Arachnids • So many species! 6% Beetles • Often tiny (<1/4”) 21% • Many look similar

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Ecological Importance: What do beetles do? In short…lots of things

Goldenrod pollinating • Predators goldenrod flower

• Pollinators

• Plant/crop pests

Silphid larvae on Jewelry made from • Scavengers/decomposers dead animal Buprestid elytra

• Food for other animals as food Dung beetles • Culturally important

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4 Beetles in Structural Pest Control: Their dark side…

• Nuisance invaders/incidentals

• Stored product pests / scavengers Asian lady beetles as a Stored product nuisance invader Fabric/Museum pests Clemson University - USDA Cooperative • Extension Slide Series, Bugwood.org Damage

• Wood borers & firewood pests Fabric damage

Clemson University - USDA Cooperative John C. French Sr., Retired, Universities: Auburn, GA, Extension Slide Series, Bugwood.org Clemson and U of MO, Bugwood.org 5

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Beetle References For Pest Control Professionals Very detailed coverage of stored product pests Good value: $$ Good coverage of pest species For pest control professional Great value: $ User friendly Authoritative work! For pest control professional Expensive: $$$$ Very technical Very detailed coverage of Doesn’t cover biology/ beetles and other pests management Costly: $$$ Loaded with photos For pest control professional Good value: $$ User friendly Mostly covers non-pests

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6 General Info on Pest Name of Beetle Family Name, Scientific Name

0 ½ 1 Size: Inches ± #” Millimeters #-# mm 0 10 20 30

Key ID feature(s): • Important features to help you identify this beetle

Notes: • Described important aspects of biology • Implications for scouting & management also mentioned when appropriate

Potential Look-alikes: • Any common look-alikes?

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Nuisance invader (Autumn)

Multicolored Asian Lady Beetle (Larva) Multicolored Asian Lady Beetle Family : Harmonia axyridis Multicolored Asian Lady Beetle (Adult) 0 ½ 1 Size: Inches ±1/4” Millimeters David Cappaert, Bugwood.org 5-8 mm 0 10 20 30 Overwintering cluster of MALBs Key ID feature(s): Distinctive! • Convex shape • Usually orange with black spots or simply orange • Black W shape on white pronotum • Larvae: spiky, black & orange

Notes: • Non-native species from east Asia • Overwinter in cracks on cliffs in native range or in other protected locations • Larvae are beneficial predators

Potential Look-alikes: • Other lady beetles, but this is our only species that will invade in significant numbers

Pest and Image Library, Bugwood.org PJ Liesch, UW Insect Diagnostic Lab 8

8 Nuisance invader (Autumn) Elm Family Chrysomelidae: Xanthogaleruca luteola

0 ½ 1 Elm Leaf Beetle (adult) Size: Inches ±1/4” Millimeters 5-8 mm 0 10 20 30

Key ID feature(s): • Oblong shape • Yellowish body with black stripes • Black spots on pronotum Elm Leaf Beetle (larvae & plant damage) Notes: Pest of elm trees • Pest of elm trees; larvae skeletonize foliage • Adults seek overwintering sites in fall

Whitney Cranshaw, Colorado State Potential Look-alikes: University, Bugwood.org • Other leaf beetles, but this is the only species that will invade structures in notable numbers • Ex. striped cucumber beetles (crop/garden pest) • Ex. Western corn rootworm (crop/garden pest)

William M. Ciesla, Forest Health Management International, Bugwood.org 9

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General ID & Broad-Nosed Weevils: General ID Family & Subfamily

General Appearance (Adult) 0 ½ 1 Size: Inches Varies, usually: 1/8-3/8” Millimeters 3-9 mm 0 10 20 30

Key ID feature(s): • Oblong or pear-shaped bodies • Color variable: black, brown, reddish, grey, pale • Long snout (weevils); shorter snout (broad-nosed) • Elbowed antennae (similar to ) Pest and Diseases Image Library, Bugwood.org Notes: Broad-Nosed Weevil (Curculionidae: Entiminae) • Life histories vary significantly by species • Some notable stored product pests & nuisance invaders

Potential Look-alikes: • Some stored product pests due to similar size/color

Kent Loeffler, Cornell University, Bugwood.org 10

10 Nuisance invader (Warm-season) Black Vine Weevil & Root Weevil sulcatus & O. ovatus Strawberry Root Weevil () Black Vine Weevil (Otiorhynchus sulcatus)

Key ID feature(s): • Pear-shaped bodies; elbowed antennae • BVW: Black w/pale spots on elytra • SRW: Reddish brown; “wrinkled” pronotum • BVW notably larger than SRW

Notes: • Both species native to Europe • Associated with plants (larvae feed on roots) • BVW strongly associated with yews

Potential Look-alikes: • Unlikely to misidentify • A few other broad-nosed weevils can invade structures; will have similar body shape and general appearance but are often pale/grey Kent Loeffler, Cornell University, Bugwood.org Pest and Diseases Image Library, Bugwood.org

0 ½ 1 0 ½ 1 Size: Inches Size: Inches ±3/8” ± 3/16” Millimeters Millimeters 7.5-9 mm 4-5.5 mm 0 10 20 30 0 10 20 30 11

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Nuisance invader (Moisture/) Stored Product Pest (Fungal Feeder) (Adults) Foreign Grain Beetle Family : advena

0 ½ 1 Size: Inches ±1/8” Millimeters 2-3 mm 0 10 20 30

Key ID feature(s): • Small size • Reddish brown or dark beige color • Clubbed antennae • “Shoulder pads” on pronotum A group of Foreign Grain Beetles

Notes: • Unusual association w/new construction • Can indicate plumbing/moisture issues • Can be associated with musty grains

Potential Look-alikes: • Other stored product pests due to similar size/color Emilie Bess, USDA APHIS PPQ, Bugwood.org

Phil Pellitteri, UW Insect Diagnostic Lab

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12 Nuisance invader (Moisture/Fungus) Stored Product Pest (Fungal Feeder)

Plaster Beetles & Relatives Minute Brown Scavenger Beetle Plaster Beetle ( filum) Family Lathridiidae; several species ( constricta)

0 ½ 1 Size: Inches 1/16”+ Millimeters 1.5-2.5 mm 0 10 20 30

Key ID feature(s): • Some of the smallest beetles you’ll likely encounter • Brown or reddish brown color • Clubbed antennae • Unique shape of pronota • Golf-ball-like divots (punctures) on elytra

Notes: • Unusual association w/new construction • Can indicate plumbing/moisture issues • Can be associated with musty grains

Potential Look-alikes: • Other stored product pests due to similar size/color

Udo Schmidt, Wikipedia Natasha Wright, Braman & Pest Elimination, Bugwood.org

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General ID Nuisance invader (Incidental)

Size: ~1/4” Corn Beetle Ground Beetles ( lecontei) Family Carabidae; many species General Anatomy of a 0 ½ 1 Size: Inches Varies, usually: “Pincher Beetle” 3/16-1” + Millimeters (Scarites sp.) www.opsu.edu 4-25+ mm 0 10 20 30

Key ID feature(s): • Long, slender legs and antennae (no club) • Oval/slender bodies Ground Beetle (Harpalus sp.) • Enlarged trochanters at base of hind legs • Size, color, appearance varies by species

Notes: • Common, widespread family • Most species are beneficial predators • Some can be strongly attracted to lights • Can’t survive indoors long

Potential Look-alikes: • Sometimes mistaken for roaches • Scarites spp. can be mistaken for stag beetles Judy Gallagher, Wikipedia

Siga, Wikipedia Size: ~1/2 - 3/4” Size: ~3/4 - 1” +

Mohammed El Damir, Bugwood.org 14

14 Stored Product Pest (Primary/Internal Feeder)

Granary Weevil & Relatives ( oryzae) Family Curculionidae: Sitophilus granarius Granary Weevil (Sitophilus granaries) 0 ½ 1 Size: Inches Varies, usually: 1/8”+ Millimeters 3-4.5 mm 0 10 20 30

Key ID feature(s): Jennifer C. Girón Duque, University of Kansas, Bugwood.org • Oblong bodies • Reddish brown or brown color • Prominent “snout” Weevil (Sitophilus zeamais) • Elbowed antennae • Golf-ball-like punctures on body

Notes: • Associated with whole stored grains • Occasionally associated with processed foods (pasta)

Potential Look-alikes: • Other stored product pests due to similar size/color Gary Alpert, Harvard University, Bugwood.org Pest and Diseases Image Library, Bugwood.org

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Stored Product Pest (Primary/Internal Feeder) Lesser Grain Borer Family : Rhyzopertha dominica

0 ½ 1 Lesser Grain Borer (Rhyzopertha dominica) Size: Inches Varies, usually: ~ 1/8” Millimeters Lesser Grain Borer Infestation 2-3 mm 0 10 20 30

Key ID feature(s): • Cylindrical bodies • Reddish brown color • “Hood” over head • Clubbed antennae w/ 3 distinct segments in club • Golf-ball-like punctures on body Pest and Diseases Image Library, Bugwood.org

Notes: Lesser Grain Borer (Rhyzopertha dominica) • Associated with whole stored grains, esp. rice, wheat, barley, etc. Clemson University - USDA Cooperative Extension Slide Series , Bugwood.org Potential Look-alikes: • Other stored product pests due to similar size/color • Bark beetles due to size/shape/color; but these wouldn’t be in grains

Gary Alpert, Harvard University, Bugwood.org 16

16 General ID

Pea & Bean Weevils (Seed Beetles) Seed Beetle Infestation Family Chrysomelidae: subfamily Bruchinae General Body Shape 0 ½ 1 Size: Inches Varies, usually: ~ 1/8” + Millimeters 2.5+ mm 0 10 20 30

Key ID feature(s): • Rounded, convex bodies • “Swollen” femurs Carl Davies, CSIRO via Wikipedia • Color pattern from tiny setae (“hairs”) • Short elytra; exposed pygidium View of Pygidium • Emarginate (notched) eyes Pest and Diseases Image Library, Bugwood.org Emarginate Eyes Notes: • Associated with legumes (peas, beans, etc.)

Potential Look-alikes: • Distinctive in appearance and habits

Pest and Diseases Image Library, Bugwood.org Pest and Diseases Image Library, Bugwood.org 17

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Stored Product Pest (Primary/Internal Feeder) Pea & Bean Weevils (Seed Beetles) Family Chrysomelidae: subfamily Bruchinae

0 ½ 1 Weevil Size: Inches ( phaseoli) Varies, usually: ~ 1/8” + Millimeters Pea Weevil ( pisorum) 3+ mm 0 10 20 30

Mexican ( subfasciatus)

www.padil.gov.au

Natasha Wright, Braman Termite & Pest Elimination, Bugwood.org Pest and Diseases Image Library, Bugwood.org : Associated with peas, lentils, Notes: Associated with peas and lentils Notes Notes: Associated with beans , mung beans

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18 Stored Product Pest (Secondary/External Feeder) Red (Tribolium castaneum) Confused Flour Beetle (Tribolium confusum) Flour Beetles Family Tenebrionidae, Tribolium spp.

0 ½ 1 Size: Inches 1/8” + Millimeters 2.5-4.5 mm 0 10 20 30

Key ID feature(s): • Small size, fairly flat • Reddish brown color • Ridge in front of eye • RFB: Distinct, 3-segmented antennal club • CFB: Gradual antennal club

Notes: Natasha Wright, Braman Termite & Pest • Scavengers of grains; common in processed foods Elimination, Bugwood.org Pest and Diseases Image Library, Bugwood.org • Red flour beetle can fly, CFB can’t fly Confused Flour Beetle Antenna

Potential Look-alikes: • Other stored product pests due to similar size/color • Related flour beetle species—examine under microscope!

Pest and Diseases Image Library, Bugwood.org Pest and Diseases Image Library, Bugwood.org 19

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Stored Product Pest (Secondary/External Feeder) Sawtoothed & Merchant Grain Beetles Family Silvanidae: surinamensis & O. mercator

Sawtoothed Grain Beetle 0 ½ 1 Merchant Grain Beetle (Oryzaephilus surinamensis) Size: Inches () ~1/8” Millimeters 2.5-3.5 mm 0 10 20 30

Key ID feature(s): • Slender bodies, fairly flat • Reddish brown or brown color • Serrated pronotum (“saw teeth”) • Trapezoid-shaped heads Pest and Diseases Image Library, Bugwood.org • Sawtoothed grain beetle: long “temple” • Merchant grain beetle: short “temple” Merchant Grain Beetle Head Notes: • Scavengers of grains; common in processed foods • Merchant grain beetle can fly, SGB can’t fly

Potential Look-alikes:

• Other stored product pests due to similar Pest and Diseases Image Library, Bugwood.org size/color Gary Alpert, Harvard University, Bugwood.org Pest and Diseases Image Library, Bugwood.org 20

20 Stored Product Pest (Secondary/External Feeder) Scavenger Infestation in Bait Drugstore & Cigarette Beetles Station Family : Stegobium paniceum & serricorne Mohammed El Damir, Bugwood.org Cigarette Beetle (Lasioderma serricorne) 0 ½ 1 (Stegobium paniceum) Size: Inches 1/16-1/8” Millimeters 1.5-3.5 mm 0 10 20 30

Key ID feature(s): • Rounded, convex bodies • “Hooded” head • Body covered with fine setae (“hairs”) • Drugstore beetle: lines on elytra; oddly clubbed antennae • Cigarette beetle: no lines on elytra; serrate antennae

Notes: • Very wide feeding habits—just about any type of

dried plant material: foods, herbs/spices, dried Pest and Diseases Image Library, Bugwood.org flowers, , etc. Natasha Wright, Braman Termite & Pest Elimination, Bugwood.org Cigarette Beetle Antenna Drugstore Beetle Antenna Potential Look-alikes: • Warehouse beetles due to size/shape Death-Watch and Beetles of Wisconsin R.A. Arango & Death-Watch and Spider Beetles of Wisconsin R.A. Arango & D.K. Young. USDA Technical Report FPL–GTR–209 D.K. Young. USDA Technical Report FPL–GTR–209 21

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Stored Product Pest (Secondary/External Feeder) Scavenger Fabric/Museum Pest

Dermestes Larva Larder Beetle Family : lardarius

0 ½ 1 Size: Inches Larder Beetle (Adult) ±1/4” 5-8 mm Millimeters 0 10 20 30 Mohammed El Damir, Bugwood.org Larval Urogomphi Key ID feature(s): • Oval body • Black with beige band across elytra • Covered with fine “hairs” (setae) • Clubbed antennae • “Hairy” larvae have rear-curving horns (urogomphi) Hide Beetle Larder Beetle Black Larder Beetle at end of body () () ()

Notes: Minor tunneling caused by larvae seeking pupation spots • One of commonest Dermestids in homes • Likes protein: fur, feathers, sinews, dead insects, wool, , etc. • Common foods: dried goods, meats/cheeses

Potential Look-alikes: Udo Schmidt via Wikipedia • Adults not easily confused with other beetles Larvae could be confused with other Dermestids • Mohammed El Damir, Bugwood.org 22

22 Stored Product Pest (Secondary/External Feeder) Scavenger Fabric/Museum Pest

Larder Beetle Relative: Hide Beetle Hide Beetle (Adult) Family Dermestidae: Dermestes maculatus

0 ½ 1 Size: Inches 1/4-3/8” 5.5-10 mm Millimeters 0 10 20 30

Key ID feature(s): • Oval body • Black (dorsal), white underside • Covered with fine “hairs” (setae) Ventral • Clubbed antennae Dorsal • “Hairy” larvae have forward-curving horns (urogomphi) at end of body Joseph LaForest, University of Georgia, Bugwood.org

Dermestes Larva Notes: Larval Urogomphi • Likes protein: fur, feathers, sinews, dead insects, wool, silk, etc. • Common foods: dried goods, meats/cheeses • Used in museums to clean skeletal remains

Potential Look-alikes: Larder beetle (adults colored differently; larval Hide Beetle Larder Beetle Black Larder Beetle • Mohammed El Damir, Bugwood.org urogomphi curve in different directions) (Dermestes maculatus) (Dermestes lardarius) (Dermestes ater) 23

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Stored Product Pest (Secondary/External Feeder) Scavenger Fabric/Museum Pest Warehouse Beetle (Larvae) Warehouse Beetles Family Dermestidae: variabile

0 ½ 1 Size: Inches Warehouse Beetle (Adult) ±1/8” 1.8-3.5 mm Millimeters 0 10 20 30

Key ID feature(s): • Small size, convex body • Brownish color with “wavy pattern” on elytra Pest and Diseases Image Library, Bugwood.org • Covered with fine “hairs” (setae) Up-close of larval Setae • Clubbed antennae • “Hairy” larvae, with long setae at back end of body

Notes: • One of commonest Dermestids in homes • Likes protein: fur, feathers, sinews, dead insects, wool, silk, etc. • Common foods: grain-based items, dried goods, etc.

Potential Look-alikes: • Adults: other Trogoderma sp. • Adults: Drugstore/cigarette beetles due to size/shape Pest and Diseases Image Library, Bugwood.org

• Larvae: other Dermestid larvae Pest and Diseases Image Library, Bugwood.org 24

24 Stored Product Pest (Secondary/External Feeder) Scavenger Fabric/Museum Pest (Trogoderma granarium) Warehouse Beetle Relatives Family Dermestidae: Trogoderma spp.

Other Trogoderma spp. 0 ½ 1 Size: Inches ±1/8” 1.8-3.5 mm Millimeters 0 10 20 30

Key ID feature(s): • Same general appearance as Warehouse beetles • Differ in color pattern and subtle features of the antennae, antennal cavity, etc. Natasha Wright, Braman Termite & Pest Elimination, Bugwood.org

Notes: • Most have similar habits to warehouse beetles Khapra Beetle Infestation • Khapra beetle (T. granarium) is one of most significant stored product pests worldwide

Potential Look-alikes: • Adults: Warehouse beetles • Adults: drugstore/cigarette beetles due to size/shape • Larvae: other Dermestids Ministry of Agriculture and Regional Development, Bugwood.org Comparison of females of some Trogoderma non‐granarium species: (A) T. angustum; (B) of Trogoderma spp. in stored T. glabrum; (C) T. grassmani; (D) T. inclusum; (E) T. ornatum; (F) T. simplex; (G) T. sternale; (H) T. variabile; (I) T. versicolor. Scale bar = 2 mm. Tomasz Klejdysz, Instytut Ochrony Roślin products should be closely examined – Państwowy Instytut Badawczy, Poznań, Poland 25

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Stored Product Pest (Secondary/External Feeder) Scavenger Fabric/Museum Pest

Anthrenus larva Family Dermestidae: verbasci

0 ½ 1 Varied Carpet Beetle (Adult) Size: Inches ±1/10” 2-3 mm Millimeters 0 10 20 30

Key ID feature(s): Entomart via Wikipedia • Oval, egg-like body • Limbs can be tucked in when disturbed • Pale, mottled pattern on body created by small, flat “scales” Larval damage in museum setting • “Hairy” larvae, with two “tufts” at end of body

Notes: • One of commonest Dermestids in homes • Likes protein: fur, feathers, sinews, dead insects, wool, silk, flower , etc. • Common foods: grain-based items, dried goods, etc.

Potential Look-alikes: • Other Dermestids Ryan Jones, Colonial Williamsburg Foundation, Bugwood.org

Natasha Wright, Braman Termite & Pest Elimination, Bugwood.org 26

26 Stored Product Pest (Secondary/External Feeder) Scavenger Fabric/Museum Pest Damage to Wool Sweater Family Dermestidae: unicolor

0 ½ 1 Varied Carpet Beetle (Adult) Size: Inches 1/8-3/16” 3-5 mm Millimeters 0 10 20 30

Key ID feature(s): • Oval, seed-like body • Entirely brownish or blackish • Antennal club long and slender • “Carrot-like” larvae slender and “hairy”; long filaments at end of body Clemson University - USDA Cooperative Extension Slide Series, Bugwood.org

Notes: • Less common that other Dermestids Attagenus larva • Likes protein: fur, feathers, sinews, dead insects, wool, silk, etc. • Can infest food items: grain-based items, dried goods Joseph Berger, Bugwood.org

Potential Look-alikes: • Adults: Warehouse beetles, cigarette/drugstore • Larvae: other Dermestids Palosirrka via Wikipedia 27

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Wood-Borer Firewood Pest Old World Lyctus Beetle (Lyctus brunneus) Powderpost Beetles Families Lyctidae & Anobiidae (now Ptinidae)

0 ½ 1 Lyctid PPB Damage Size: Inches 1/8-5/16” Pest and Diseases Image Library, Bugwood.org 3-8 mm Millimeters 0 10 20 30 Furniture Beetle ( punctatum)

Key ID feature(s): • Slender bodies • “Hooded” heads (Anobiid PPB) • Small holes in wood (~1/16 - 1/8”) • Frass: fine powder (Lyctids); small, loosely- Photo submission at UW Insect Diagnostic Lab packed clumps (Anobiids) • Larvae: small, pale, grub-like Pest and Diseases Image Library, Bugwood.org

Notes: (Hemicoelus sp.) • Common wood-boring pests • Can reinfest under the right conditions Beetle Type Wood Attacked • Differ in types of wood they infest Lyctid PPB • New Hardwoods New softwoods Potential Look-alikes: • Anobiid PPB Old hardwoods Stored product pests (flour beetles, etc.) • • Old softwoods • Drugstore/cigarette beetles • Bark beetles • Pest and Diseases Image Library, Bugwood.org 28

28 Wood-Borer (Adult) Wharf Borer Family : melanura

0 ½ 1 Size: Inches Wharf Borer (Larva) ± 1/2” ~12 mm Millimeters 0 10 20 30

Key ID feature(s): • Slender bodies • Beige color with black at tips of elytra • Larva slender, pale with 3 pairs true legs, 2 pairs of prolegs; up to 1.5” long

Notes: • Uncommon wood-boring pests • Associated with very damp wood: wharfs, docks, water leaks indoors, etc.

Potential Look-alikes: M. O'Donnell and A. Cline, Wood Boring Beetle • Longhorned beetles Families, USDA APHIS PPQ, Bugwood.org

Udo Schmidt via Wikipedia

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General ID

Longhorned beetle Larva Longhorned Beetles (Roundheaded Borers) (aka “roundheaded borer”)

Family Cerambycidae Longhorned Beetle Anatomy

Size: 0 ½ 1 1 ½ 2 Inches 2+ Inches Varies by Millimeters Species 0 10 20 30 40 50

Key ID feature(s): • Slender bodies, often large (3/4”+) • Lobed tarsi • Long antennae (usually >1/2 body length) • Emarginate (“notched”) eyes • Larvae pale, usually legless, round • Create round or oval holes ~1/4-1/2”

Notes: • Common wood-boring pests • Usually associated with dead/dying trees • Common sources: firewood, “rustic” furniture, or new log cabins • A few can be associated with structural wood

Potential Look-alikes: Wharf borer (which has distinctive color) Gerald J. Lenhard, Louisiana • From: A Field Guide to Northeastern Longhorned Beetles, Douglas Yanega State University, Bugwood.org 30

30 Wood-Borer Firewood Pest Old House Borer ( bajulus)

Natasha Wright, Longhorned Beetles Braman Termite & Pest Elimination, Family Cerambycidae; Many(!) species Bugwood.org Size: Up to 3/4”

Cedar Tree Borer Brown Prionid Pine Sawyer ( ligneus) (Orthosoma brunneum) (Monochamus scutellatus) Redheaded Ash Borer Borer (Neoclytus acuminatus) (Megacyllene robinia) Size: ~3/4” Size: Up to 2”

Size: ~1”

USDA Screening Aid for the Cerambycidae of Size: 3/4-1” Western North America Size: ~1/2” Natasha Wright, Braman Termite & USDA Screening Aid for the Cerambycidae of Pest Elimination, Bugwood.org Western North America Steven Valley, Oregon Department of Agriculture, Nathan Lord, Longicorn ID, USDA APHIS PPQ, Bugwood.org Bugwood.org 31

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Questions & Discussion What do you want to know about beetles?

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