IAPT/IOPB Chromosome Data 8

TAXON 58 (4) • November 2009: 1281–1289 Marhold (ed.) • IAPT/IOPB chromosome data 8

IOPB COLUMN

Edited by Karol Marhold & Ilse Breitwieser

IAPT/IOPB chromosome data 8

Edited by Karol Marhold

Victor V. Chepinoga, Aleksandr A. Gnutikov, Amoria montana (L.) Soják, 2n = 16; Russia, Irkutskaya Oblast’, Ilya V. Enushchenko & Sergey A. Rosbakh AG & A. Hoff C510. Amoria repens (L.) C. Presl, 2n = 32; Russia, Irkutskaya Oblast’, Department of Botany & Genetics, Irkutsk State University, AG & IE С451. 1 Karl Marks Str., 664003, Irkutsk, Russia. Victor.Chepinoga@ Astragalus olchonensis Gontsch., 2n = 16; Russia, Irkutskaya gmail.com Oblast’, VCH, A. Verkhozina & E. Kuznetsova С634. Lathyrus humilis (Ser.) Spreng., 2n = 14; Russia, Irkutskaya All materials CHN; collectors: VCH = V.V. Chepinoga, AG = Oblast’, AG & VCH C185. A.A. Gnutikov, IE = I.V. Enuschchenko; vouchers in IRKU. Lathyrus pratensis L., 2n = 14; Russia, Irkutskaya Oblast’, AG & IE C474. This study was supported by Grants no. 05-05-64061, 07-04- Melilotoides platycarpos (L.) Soják, 2n = 16; Russia, Irkutskaya 00610 from Russian Fund for Basic Research, RNP.2.2.1.1.7334 Oblast’, AG & IE C535. and RNP.2.2.3.1.4667 from Ministry of Education and Science of Trifolium pratense L., 2n = 14; Russia, Irkutskaya Oblast’, AG Russian Federation. & IE С457. Vicia amoena Fisch., 2n = 14; Russia, Irkutskaya Oblast’, AG CYPERACEAE & IE C472. Blysmus rufus (Huds.) Link, 2n = 40; Russia, Buryatiya, AG & Vicia cracca L., 2n = 14; Russia, Irkutskaya Oblast’, AG & V. IE С357; Russia, Chitinskaya Oblast’, AG C605. Mozer C156. Bolboschoenus planiculmis (F. Schmidt) T.V. Egorova, 2n = 52; Vicia unijuga A. Br., 2n = 12; Russia, Irkutskaya Oblast’, AG Russia, Chitinskaya Oblast’, AG C607. & IE C471. Carex appendiculata (Trautv. & C.A. Mey.) Kük., 2n = 78; Rus- sia, Chitinskaya Oblast’, AG & IE C388. NITRARIACEAE Carex atherodes Spreng., 2n = ca. 78; Russia, Chitinskaya Nitraria sibirica Pall., 2n = 60; Russia, Buryatia, VCH C642. Oblast’, AG C419. Carex bohemica Schreb., 2n = ca. 62; Russia, Chitinskaya Oblast’, POACEAE AG & IE C413. Calamagrostis obtusata Trin., 2n = 28; Russia, Tyva, E. Veder- Carex duriuscula C.A. Mey., 2n = 62; Russia, Buryatiya, AG nikova C643. & IE C363. Eragrostis pilosa (L.) P. Beauv., 2n = 40; Russia, Chitinskaya Carex enervis C.A. Mey., 2n = 44–46; Russia, Buryatiya, AG Oblast’, AG & IE C610. & IE C359. Festuca ovina L., 2n = 28; Russia, Irkutskaya Oblast’, IE C623. Carex juncella (Fr.) Th. Fr., 2n = 52; Russia, Irkutskaya Oblast’, Milium effusum L., 2n = 28; Russia, Tyva, E. Vedernikova C635. AG & IE C464. Poa irсutica Roshev., 2n = 28; Russia, Irkutskaya Oblast’, IE C624. Carex reptabunda (Trautv.) V.I. Krecz., 2n = 44; Russia, Chitins- Poa alpina L., 2n = 28; Russia, Irkutskaya Oblast’, IE C622. kaya Oblast’, AG & IE C379. Setaria pumila (Poir.) Roem. & Schult., 2n = 36; Russia, Carex vesicata Meinsh., 2n = 54; Russia, Chitinskaya Oblast’, Irkutskaya Oblast’, IE C639. AG & IE C389, C390. Eleocharis acicularis (L.) Roem. & Schult., 2n = 18; Russia, RANUNCULACEAE Chitinskaya Oblast’, AG & IE C393. Adonis sibirica (Patrin ex DC.) Ledeb., 2n = 16; Russia, Irkut- Eleocharis palustris (L.) Roem. & Schult., 2n = 16; Russia, Chi- skaya Oblast’, IE C437. tinskaya Oblast’, AG & IE C392. Anemonastrum crinitum (Juz.) Holub, 2n = 14; Russia, Irkutskaya Scirpus radicans Schkuhr, 2n = 56; Russia, Chitinskaya Oblast’, Oblast’, AG & VCH C141. AG & IE C398. Batrachium circinatum (Sibth.) Spach, 2n = 16; Russia, Chi- tinskaya Oblast’, AG C614. FABACEAE Halerpestes salsuginosa (Pall. ex Georgi) Greene, 2n = 48; Rus- Amoria hybrida (L.) C. Presl, 2n = 16; Russia, Irkutskaya Oblast’, sia, Buryatiya, AG & IE C360, AG & IE C559. AG & A. Hoff C509. Halerpestes sarmentosa (Adams) Kom., 2n = 16; Russia, Chi-

All materials for the chromosome column should be submitted electronically to: Karol Marhold, [email protected] (Insti- tute of Botany, Slovak Academy of Sciences, SK-845 23 Bratislava, Slovakia, and Department of Botany, Charles University, CZ 128-01 Prague, Czech Republic). The full version of this contribution is available in the online edition of TAXON. The following citation format is recommended: Baltisberger, M. & Voelger, M. 2006. Sternbergia sicula. In: Marhold, K. (ed.), IAPT/IOPB chromosome data 1. Taxon 55: 444, E2.

1281 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: 1281–1289

tinskaya Oblast’, AG & IE C376; Russia, Buryatiya, AG & IE CHENOPODIACEAE С355, AG & VCH C560. Salsola kali L., 2n = 36; Morocco, RV 10901 & CHO 5349. Ranunculus chinensis Bunge, 2n = 16; Russia, Chitinskaya Oblast’, AG C420. CISTACEAE Ranunculus gmelinii DC., 2n = 16; Russia, Chitinskaya Oblast’, Fumana laevipes (L.) Spach, 2n = 32; Morocco, RV 11239 & AG & IE C414; Russia, Irkutskaya Oblast’, AG & IE С459. CHO 5687. Ranunculus longicaulis C.A. Mey., 2n = 32; Russia, Chitinskaya Helianthemum apeninnum (L.) Mill., 2n = 20; Morocco, RV 10721 Oblast’, AG & IE С427. & CHO 5169, RV 11031 & CHO 5479. Ranunculus polyanthemos L., 2n = 16; Russia, Irkutskaya Oblast’, Helianthemum papillare Boiss., 2n = 20; Morocco, RV 11669 & AG & IE С499. CHO 6117. Ranunculus radicans C.A. Mey., 2n = 16; Russia, Chitinskaya Oblast’, AG C592. COMPOSITAE Ranunculus repens L., 2n = 32; Russia, Irkutskaya Oblast’, AG Erigeron bonariensis L. [≡ Conyza bonariensis (L.) Cronquist], & A. Hoff C518; Russia, Chitinskaya Oblast’, AG & IE C382. 2n = 36; Morocco, RV 11480 & CHO 5928. Ranunculus reptans L., 2n = 32; Russia, Irkutskaya Oblast’, AG & IE С449; Russia, Chitinskaya Oblast’, AG & IE C429, AG CRUCIFERAE & IE C588. **Alyssum simplex Rudolphi, 2n = 32; Morocco, RV 10709 & Ranunculus sceleratus L., 2n = 64; Russia, Chitinskaya Oblast’, CHO 5157. AG & IE С400, AG & IE C401. Anastatica hierochuntica L., 2n = 22; Morocco, 7 Jun 1993 CHO & RV s.n. Biscutella baetica Boiss. & Reut., 2n = 16; Morocco, RV 9858 & Robert Vogt1 & Christoph Oberprieler2 CHO 4306, RV 10103 & CHO 4551, RV 10190 & CHO 4638, RV 10786 & CHO 5234, RV 11191 & CHO 5639. 1 Botanischer Garten und Botanisches Museum Berlin- **Brassica fruticulosa subsp. cossoniana (Boiss.& Reut.) Maire, Dahlem, Freie Universität Berlin, Königin-Luise-Str. 6–8, 2n = 16; Morocco, RV 10946 & CHO 5394, RV 11534 & CHO 14195 Berlin, Germany. [email protected] (author for cor- 5982. respondence) Carrichtera annua (L.) DC., 2n = 16; Morocco, RV 10950 & CHO 2 Institut für Botanik, Universität Regensburg, Universitätsstr. 5398, RV 11130 & CHO 5578. 31, 93053 Regensburg, Germany Diplotaxis harra (Forssk.) Boiss. subsp. harra, 2n = 26; Morocco, RV 10540 & CHO 4988. This contribution is ninth in the series entitled “Chromosome Eruca vesicaria subsp. sativa (Mill.) Thell., 2n = 22; Morocco, numbers of North African phanerogams.” RV 10934 & CHO 5382, RV 11044 & CHO 5492. *Kremeriella cordylocarpus (Coss. & Durieu) Maire, 2n = 22; All materials CHN; collectors: RV = R. Vogt, CHO = Ch. Morocco, RV 11173 & CHO 5621. Oberprieler; vouchers in B. Matthiola parviflora (Schousb.) R. Br., 2n = 14; Morocco, RV 11127 & CHO 5575. * The chromosome number of the marked species has been *Muricaria prostrata (Desf.) Desv., 2n = 26; Morocco, RV 10685 counted for the first time. & CHO 5133. ** The chromosome number of the marked species is deviat- Rorippa nasturtium-aquaticum (L.) Hayek [ = Nasturtium of- ing from former reports. ficinale R. Br.], 2n = 32; Morocco, RV 10102 & CHO 4550. Rapistrum rugosum (L.) All. subsp. rugosum, 2n = 16; Morocco, BORAGINACEAE RV 11146 & CHO 5594. Cynoglossum creticum Mill., 2n = 24; Morocco, RV 11547 & Sisymbrium irio L., 2n = 14; Morocco, RV 11297 & CHO 5745. CHO 5995. Cynoglossum dioscoridis Vill., 2n = 24; Morocco, 7 Jun 1993 EUPHORBIACEAE CHO & RV s.n. Euphorbia exigua L. subsp. Exigua, 2n = 24; Morocco, RV 10764 & CHO 5212. CARYOPHYLLACEAE Euphorbia helioscopia L. subsp. helioscopia, 2n = 42; Morocco, Rhodalsine geniculata (Poir.) F.N. Williams [≡ Minuartia genicu- RV 11604 & CHO 6052. lata (Poir.) Thell.], 2n = 18; Morocco, RV 11296 & CHO 5744, Euphorbia hirsuta L., 2n = 14; Morocco, RV 10195 & CHO 4643. RV 10945 & CHO 5393. Euphorbia medicaginea Boiss., 2n = 16; Morocco, RV 10917 & Silene aellenii Sennen [ = Silene divaricata Lag., nom. illeg.], 2n CHO 5365. = 24; Morocco, RV 11211 & CHO 5659. *Euphorbia resinifera Berg, 2n = ca. 112; Morocco, RV 11940. Silene colorata Poir., 2n = 24; Morocco, RV 10877 & CHO 5325. Euphorbia segetalis L., 2n = 16; Morocco, RV 9878 & CHO 4326. Silene obtusifolia Willd., 2n = 24; Morocco, RV 9921 & CHO Euphorbia terracina L., 2n = 18; Morocco, RV 10931 & CHO 4369, RV 9926 & CHO 4374. 5379. Silene ramosissima Desf., 2n = 24; Morocco, RV 10925 & CHO Mercurialis ambigua L. f. [≡ Mercurialis annua subsp. ambigua 5373. (L. f.) Arcang.], 2n = 48; Morocco, RV 10753 & CHO 5201, Silene secundiflora Otth, 2n = 24; Morocco, RV 11188 & CHO RV 11178 & CHO 5626, RV 11300 & CHO 5748, RV 11550 & 5636, RV 11018 & CHO 5466. CHO 5998. Silene tridentata Desf., 2n = 24; Morocco, RV 10855b & CHO 5303b. FUMARIACEAE Spergularia diandra (Guss.) Boiss., 2n = 18; Morocco, RV 11116 Fumaria bastardii Boreau, 2n = 48; Morocco, RV 10322 & CHO & CHO 5564. 4770.

1282 TAXON 58 (4) • November 2009: 1281–1289 Marhold (ed.) • IAPT/IOPB chromosome data 8

GERANIACEAE Asterolinum linum-stellatum (L.) Duby, 2n = 20; Morocco, RV Erodium chium (L.) Willd., 2n = 20; Morocco, RV 10843 & CHO 10851 & CHO 5299. 5291, RV 10875a & CHO 5323a, RV 11168 & CHO 5616, RV 11344 & CHO 5792, RV 11576 & CHO 6024. RANUNCULACEAE *Erodium crenatum Pomel, 2n = 20; Morocco, RV 10303 & CHO Ranunculus paludosus Poir., 2n = 16; Morocco, RV 9966 & CHO 4751. 4414. Erodium glaucophyllum (L.) L´Hér., 2n = 20; Tunisia, RV 12754 Ranunculus spicatus subsp. blepharicarpos (Boiss.) Grau, 2n = & CHO 7059, CHO & RV. 16; Morocco, RV 11398 & CHO 5846, Cubr 34703. *Erodium guttatum (Desf.) Willd., 2n = 20; Morocco, RV 10587a & CHO 5035a, 26 Sep 1994 RV & CHO, RV 10722 & CHO RUBIACEAE 5170, RV 11732 & CHO 6180. Galium verrucosum Huds., 2n = 22; Morocco, RV 10282 & CHO Erodium laciniatum (Cav.) Willd. subsp. laciniatum, 2n = 20; 4730, RV 11064 & CHO 5512. Morocco, RV 10875b & CHO 5323b. Valantia muralis L., 2n = 18; Morocco, RV 9882 & CHO 4330. Erodium laciniatum subsp. pulverulentum (Boiss.) Batt., 2n = 20; Morocco, RV 11364 & CHO 5812. SAPOTACEAE Erodium moschatum (L.) L’Hér., 2n = 20; Morocco, RV 9992 & *Argania spinosa (L.) Skeels, 2n = 20; Morocco, 14 Apr 1994 RV CHO 4440. & CHO s.n., Cubr 40606. Erodium salzmannii Delile, 2n = 20; Morocco, RV 10186 & CHO 4634, RV 10206 & CHO 4654, RV 11865 & CHO 6313. SCROPHULARIACEAE Geranium purpureum Vill., 2n = 32; Morocco, RV 11363 & CHO *Misopates microcarpum (Pomel) D.A. Sutton [≡ Antirrhinum 5811. microcarpum Pomel], 2n = 16; Morocco, RV 10958 & CHO Geranium rotundifolium L., 2n = 26; Morocco, RV 11345 & CHO 5406, RV 11242 & CHO 5690. 5793. Scrophularia laevigata Vahl, 2n = 58; Morocco, RV 11522 & CHO 5970. LABIATAE *Rosmarinus eriocalyx var. pallescens (Maire) Upson & Jury, 2n UMBELLIFERAE = 24; Morocco, RV 11434 & CHO 5882. Bifora testiculata (L.) Roth, 2n = 22; Morocco, RV 10267 & CHO Salvia aegyptiaca L., 2n = 26; Morocco, RV 11078 & CHO 5526. 4715. Salvia verbenaca L., 2n = 24; Morocco, RV 10635 & CHO 5083. *Bupleurum lateriflorum Coss., 2n = 28; Morocco, RV & CHO. Daucus carota L. subsp. carota, 2n = 18; Morocco, RV 10203 LEGUMINOSAE & CHO 4651. Astragalus epiglottis L., 2n = 16; Morocco, RV 11199 & CHO 5647. Pseudorlaya pumila (L.) Grande, 2n = 16; Morocco, RV 10884 Tripodion tetraphyllum (L.) Fourr. [≡ Anthyllis tetraphylla L.], & CHO 5332. 2n = 16; Morocco, 7 Jun 1993 RV & CHO s.n. VALERIANACEAE LILIACEAE Centranthus calcitrapae (L.) Dufr., 2n = 32; Morocco, RV 11320 Asphodelus refractus Boiss., 2n = 52; Morocco, RV 10458 & & CHO 5768, RV 11995. CHO 4906. Fedia cornucopiae (L.) Gaertn., 2n = 32; Morocco, RV 11541 & Asphodelus tenuifolius Cav., 2n = 28; Morocco, RV 11743 & CHO CHO 5989. 6191. Valerianella discoidea (L.) Loisel., 2n = 14; Morocco, RV 11051 & CHO 5499. LINACEAE Linum strictum L., 2n = 18; Morocco, RV 10960 & CHO 5408. Julia V. Shner, Michael G. Pimenov & Eugene V. Kljuykov MALVACEAE *Malva aegyptia L., 2n = 24; Morocco, RV 10673 & CHO 5121. Botanical Garden of Moscow State University, Moscow, 119899, Russia. [email protected] (author for correspondence) PAPAVERACEAE Papaver rhoeas L., 2n = 14; Morocco, RV 11102 & CHO 5550. All materials CHN; collectors P&K = M.G. Pimenov & E.V. Kljuykov, Pimenov & al. = M.G. Pimenov, E.V. Kljuykov, Hu Zhi- PLANTAGINACEAE Hao & Liu QiXin; vouchers in MW. Plantago afra L., 2n = 12; Morocco, RV 10944 & CHO 5392, RV 11050 & CHO 5498. This study was supported by grants 04-04-48146 and 07-04- Plantago albicans L., 2n = 10; Morocco, RV 11065 & CHO 5513. 00270 from the Russian Foundation for Basic Research (RFBR).

POLYGONACEAE APIACEAE Emex spinosa (L.) Campd., 2n = 20; Morocco, RV 11917 & CHO Acronema tenerum (DC.) Edgew., 2n = 18; India, Uttaranchal, 6365. P&K 235; India, Sikkim, P&K 61. Rumex vesicarius L., 2n = 18; Morocco, RV 10383 & CHO 4831, Angelica pubescens Maxim., 2n = 22; Japan, 11 Aug 2004 M.G. RV 10563 & CHO 5011, RV 11029 & CHO 5477. Pimenov s.n. Bupleurum candollei Wall. ex DC., 2n = 16; India, Sikkim, P&K PRIMULACEAE 57. Anagallis arvensis L., 2n = 20; Morocco, RV 10941 & CHO 5389. 2n Bupleurum gracillimum Klotzsch, 2n = 14; India, Himachal = 40; Morocco, RV 10175 & CHO 4623, RV 11664 & CHO 6112. Pradesh, P&K 66.

1283 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: 1281–1289

Bupleurum marginatum DC., 2n = 16; India, Uttaranchal, P&K Suaeda corniculata (C.A. Mey.) Bunge, 2n = 54; Russia, Golub 7, 10. 761 (NS), L 434, 490 (NS), 18 Sep 2002, Osipov s.n. (NS). Bupleurum mucronatum Wight & Arn., n = 7; India, Tamil Nadu, Suaeda kulundensis Lomon. & Freitag, 2n = 72; Russia, Golub P&K 58. 762 (NS); L 79, 80, 85, 740, 742, 755 (NS); K 101b, 103, 123, Heracleum hemsleyanum Diels, 2n = 22; China, Sichuan, Pi- 135 (NS); K & A 4 (NS); K & S 133 (NS); A & al. 303 (NS); menov & al. 327. 17 Sep 1999, A & al. s.n. (NS); Korolyuk 241 (NS); Zolotov Heracleum nubigenum C.B. Clarke, 2n = 44; India, Sikkim, P&K 58. 16 (NS). Kazakhstan: L 51a, 194b, 205. 2n = 90; Russia, L 25, Hymenidium brunonis (DC.) Lindl., n = 11; India, Himachal 51b, 69, 79b, 85b, 386, s.n. (NS); 22 Sep 1999, A & al. s.n. Pradesh, P&K 205. (NS); K 86, 107, 151 (NS); 2 Sep. 1999, K s.n. (NS); K & S 80 Hymenidium decurrens (Franch.) Pimenov & Kljuykov, 2n = 18; (NS); 28 Sep 2002, Grebenyuk s.n. (NS). China, Yunnan, Pimenov & al. 560. Suaeda maritima (L.) Dumort. s.l. 2n = 36; Italy, F 34.1007, Kailashia robusta Pimenov & Kljuykov, 2n = 22; India, Utta- 34.1011,2, 34.1021, 34.1031,3, 34.1096–8 (KAS); Germany, Oct ranchal, P&K 16. 2006, Prinz s.n. (KAS). Kedarnatha meifolia Pimenov & Kljuykov, 2n = 22; India, Suaeda patagonica Speg., 2n = 54; Argentina, 9 Feb 2003, Schrö- Uttaranchal, P&K 236. der s.n. (KAS). Ligusticopsis coniifolia (DC.) Pimenov & Kljuykov, 2n = 22; Suaeda salsa (L.) Pall., 2n = 36; Ukraine, F 33.219, 33.252, 33.277 India, Uttaranchal, P&K 25. (KAS); Romania, Oct 2003, Fagaras s.n. (KAS); 27 Sep 2003, Ligusticopsis wallichiana (DC.) Pimenov & Kljuykov, 2n = 22; Miele s.n. (KAS). India, Sikkim, P&K 56. Suaeda sibirica Lomon. & Freitag, 2n = 72; Russia, L 46, 59, Melanosciadium bipinnatum (Shan RenHwa & Pu FaTing) Pi- 284, 285, 286, 287, 299, 335, 336, 341, 366, 373, 379, 426, menov & Kljuykov, 2n = 22; China, Sichuan, Pimenov & al. 190. 431, 434, 438, 440, 445, 449, 477, 482, 489, 502, 521, 601, 606, Osmorhiza aristata (Thunb.) Rydb., 2n = 22; India, Uttaranchal, 609, 610, 612, 616, 618, 621, 626, 638, 643 (NS); 28 Sep 2002, P&K 228. Osipov s.n. (NS); Pimpinella acuminata (Edgew.) C.B. Clarke, 2n = 20; India, Himachal Pradesh, P&K 167. Pimpinella katrajensis R.S. Rao & Hemadri, n = 9; India, Nina S. Probatova1, Vitaly P. Seledets2, Elvira G. Rudyka1, Maharashtra, P&K 33. Aleksandr A. Gnutikov3, Zoya V. Kozhevnikova1 Pimpinella wallichiana (Miq. ex Hohen.) Gandhi, n = 10. India, & Vyacheslav Yu. Barkalov1 Karnataka, P&K 38. Semenovia thomsonii (C.B. Clarke) Manden., 2n = 22; India, 1 Institute of Biology & Soil Science, Far East Branch of Rus- Himachal Pradesh, P&K 150. sian Academy of Sciences, 159 Stoletya Prospect, 690022, Seseli nudum (Lindl.) Pimenov & Kljuykov, 2n = 22; India, Vladivostok, Russia. [email protected] (author for Uttaranchal, P&K 8. correspondence) Sinocarum wolffianum (Fedde ex H. Wolff) P.K. Mukh. & Con- 2 Pacific Institute of Geography, Far East Branch of Russian stance, 2n = 22; India, Sikkim, P&K 62. Academy of Sciences, 7 Radio Str., 690041, Vladivostok, Tetrataenium aquilegifolium (C.B.Clarke) Manden., 2n = 44; In- Russia dia, Maharashtra, P&K 2, 31. 3 Baikal Research Center, Irkutsk State University, 5–10 Karl Tetrataenium ligusticifolium (Wight) Manden. ( = T. candollea- Marx Str., 664003, Irkutsk, Russia num (Wight & Arn.) Manden.), n = 22; India, Tamil Nadu, Kerala, P&K 89, 93. All materials CHN; collectors: NP = N. Probatova, VS = Tetrataenium rigens (DC.) Manden.; 2n = 22; India, Karnataka, V. Seledets; vouchers in VLA. P&K 37. Tongoloa gracilis H. Wolff, 2n = 22; China, Sichuan, Pimenov This study was supported by Grants nos. 04-04-49750 and & al. 299. 07-04-00610 (to N.S. Probatova), from Russian Fund for Basic Trachyspermum stictocarpum (C.B. Clarke) H. Wolff, n = 11; Research (RFBR). India, Maharashtra, P&K 14. Vanasushava pedata (Wight) P.K. Mukh. & Constance, n = 23; ALLIACEAE India, Tamil Nadu, P&K 72. Allium anisopodium Ledeb., 2n = 16; Russia, Primorskii Krai, Barkalov 9221. Allium condensatum Turcz., 2n = 16; Russia, Primorskii Krai, Maria Lomonosova1 & Helmut Freitag2 Barkalov 9174. Allium macrostemon Bunge, 2n = 32; Russia, Primorskii Krai, 1 Central Siberian Botanical Garden, Siberian Branch of the NP & VS 9608. Russian Academy of Science, Novosibirsk 630090, Russia. Allium maximowiczii Regel, 2n = 16; Russia, Amurskaya oblast’, [email protected] (author for correspondence) Kudrin 11216. 2 Plant Taxonomy and Morphology, Institute of Biology, Uni- Allium ochotense Prokh., 2n = 32; Russia, Kamchatskii Krai, versity of Kassel, 34132 Kassel, Germany Barkalov 7369, Khabarovskii Krai, Barkalov 9907. Allium prokhanovii Vved., 2n = 16; Russia, Primorskii Krai, All materials CHN; collectors: A = Artemov, F = Freitag, K = Lapenko 11095. Krasnoborov, L = Lomonosova, S = Shaulo. Allium senescens L., 2n = 16; Russia, Primorskii Krai, Barkalov 8846, Nechaev 10609. 2n = 32; Russia, Primorskii Krai, CHENOPODIACEAE Barkalov 9215, Verkholat 10388. Suaeda aegyptiaca (Hasselq.) Zohary, 2n = 18; Jordan, F Allium strictum Schrad., 2n = 32; Russia, Kamchatskii Krai, 30.120/2001 (KAS). Barkalov 7372.

1284 TAXON 58 (4) • November 2009: 1281–1289 Marhold (ed.) • IAPT/IOPB chromosome data 8

AMARANTHACEAE Lychnis fulgens Fisch. ex Curt., 2n = 24; Russia, Primorskii Krai, Amaranthus blitoides S. Watson, 2n = 32; Russia, Krasnodarskii Shatalova 7677. Krai, NP & VS 11286. Spergula arvensis L., 2n = 18; Russia, Karelia, NP & VS 11195.

ASCLEPIADACEAE CHENOPODIACEAE Pycnostelma paniculata (Thunb.) Makino, 2n = 22; Russia, Pri- Atriplex micrantha C.A. Mey., 2n = 18; Russia, Krasnodarskii morskii Krai, Verkholat 10314. Krai, NP & VS 11270. Chenopodium foliosum Asch., 2n = 18; Russia, Irkutskaya ASTERACEAE Oblast’, Repina 11058. Artemisia glomerata Ledeb., 2n = 36; Russia, Kamchatskii Krai, Chenopodium polyspermum L., 2n = 18; Russia, Krasnodarskii Yakubov 8441. [Error: “2n = 18” — Probatova & al., Taxon 57: Krai, NP & VS 11271. 1269, E5. 2008.] Chenopodium urbicum L., 2n = 18; Russia, Krasnodarskii Krai, Artemisia integrifolia L., 2n = 36; Russia, Primorskii Krai, NP & VS 11275. Barkalov 9816. Corispermum nitidum Schult., 2n = 18; Russia, Krasnodarskii Bidens cernua L., 2n = 24; Russia, Primorskii Krai, Barkalov 8859. Krai, NP & VS 11186. Bidens frondosa L., 2n = 48; Russia, Krasnodarskii Krai, NP & Corispermum ulopterum Fenzl, 2n = 18; Russia, Buryatia, Ka- VS 11295, Saratovskaya Oblast’, NP & VS 11194. zanovskii 11057. Bidens tripartita L., 2n = 48; Russia, Krasnodarskii Krai, NP Salsola pontica (Pall.) Degen, 2n = 18; Russia, Krasnodarskii & VS 11298. Krai, NP & VS 11269. Centaurea diffusa Lam., 2n = 18; Russia, Krasnodarskii Krai, Teloxys aristata (L.) Moq., 2n = 18; Russia, Irkutskaya Oblast’, Manilo 11187. NP & VS 11120. Centaurea frygia L., 2n = 22; Russia, Karelia, NP & VS 11199 Chondrilla juncea L., 2n = 20; Russia, Krasnodarskii Krai, NP CHLORANTHACEAE & VS 11227. Chloranthus japonicus Siebold, 2n = 30; Russia, Primorskii Krai, Cichorium intybus L., 2n = 18; Russia, Krasnodarskii Krai, NP Barkalov 7798. & VS 11274. Crepis setosa Haller f., 2n = 8, 16; Russia, Krasnodarskii Krai, CONVALLARIACEAE NP & VS 11272. Maianthemum bifolium (L.) F.W. Schmidt, 2n = 54; Russia, Cyclachaena xanthiifolia (Nutt.) Fresen., 2n = 36; Russia, Sara- Khabarovskii Krai, Kozhevnikov 9913. tovskaya Oblast’, NP & VS 11188. Hieracium umbellatum L., 2n = 18; Russia, Karelia, NP & VS 11207. CONVOLVULACEAE Inula oculus-christi L., 2n = 32; Russia, Krasnodarskii Krai, Calystegia dahurica (Herb.) Choisy, 2n = 22; Russia, Primorskii NP & VS 11282. Krai, Lapenko 11050. Lapsana communis L., 2n = 14; Russia, Karelia, NP & VS 11201, Leningradskaya Oblast’, NP & VS 11170. CRASSULACEAE Phalacroloma septentrionale (Fern. & Wieg.) Tzvelev, 2n = 27; Sedum acre L., 2n = 16; Russia, Karelia, NP & VS 11175. Russia, Krasnodarskii Krai, NP & VS 11289. Sonchus asper (L.) Hill, 2n = 18; Russia, Krasnodarskii Krai, EUPHORBIACEAE Manilo 11172. Acalypha australis L., 2n = 20; Russia, Krasnodarskii Krai, NP Tripleurospermum inodorum (L.) Sch. Bip., 2n = 36; Russia, & VS 11189. Karelia, NP & VS 11177, Leningradskaya Oblast’, NP & VS 11180. FABACEAE Lespedeza bicolor Turcz., 2n = 22; Russia, Amurskaya Oblast’, BORAGINACEAE NP & VS 11239. Amsinckia micrantha Suksd., 2n = 16; Russia, Magadanskaya Sophora flavescens Sol., 2n = 18; Russia, Amurskaya Oblast’, Oblast’, Lyssenko 11240. Timchenko 11077. Trifolium repens L., 2n = 32; Russia, Amurskaya Oblast’, Tim- BRASSICACEAE chenko 11072. Barbarea arcuata (Opiz ex J. & C. Presl) Rchb., 2n = 16; Russia, Vicia nervata Sipl., 2n = 24; Russia, Irkutskaya Oblast’, Kaza- Krasnodarskii Krai, NP & VS 11297. novskii 11225. Erysimum amurense Kitag., 2n = 12; Russia, Khabarovskii Krai, Barkalov 11325. FUMARIACEAE Lepidium densiflorum Schrad., 2n = 32; Russia, Amurskaya Corydalis repens Mandl & Muehld., 2n = 16; Russia, Primorskii Oblast’, Timchenko 11228. Krai, Volynets 11317. Corydalis speciosa Maxim., 2n = 16; Russia, Primorskii Krai, CANNABACEAE Lapenko 11075. Humulus lupulus L., 2n = 20; Russia, Krasnodarskii Krai, NP & VS 11293. HEMEROCALLIDACEAE Hemerocallis minor Mill., 2n = 22; Russia, Primorskii Krai, CARYOPHYLLACEAE Barkalov 9231. Dianthus versicolor Fisch. ex Link, 2n = 30; Russia, Primorskii Krai, Barkalov 9815. HYPERICACEAE Gypsophila perfoliata L., 2n = 34; Russia, Krasnodarskii Krai, Hypericum perforatum L., 2n = 32; Russia, Karelia, Kravchenko NP & VS 11284. 11204.

1285 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: 1281–1289

IRIDACEAE Agrostis capillaris L., 2n = 28; Russia, Karelia, NP & VS 11251, Iris setosa Pall. ex Link, 2n = 38; Russia, Kamchatskii Krai, 11306. Barkalov 7368, 7653. Agrostis clavata Trin., 2n = 42; Russia, Amurskaya Oblast’, Tim- chenko 11067, Magadanskaya Oblast’, Lyssenko 10993. LAMIACEAE Agrostis scabra Willd., 2n = 42; Russia, Primorskii Krai, Calamintha nepeta (L.) Savi, 2n = 24; Russia, Krasnodarskii Barkalov 8826. Krai, NP & VS 7447. Agrostis trinii Turcz., 2n = 28; Russia, Primorskii Krai, VS Glechoma longituba (Nakai) Kuprian., 2n = 18; Russia, Primor- 10001. skii Krai, Lapenko 11032. 2n = 36; Russia, Primorskii Krai, Alopecurus aequalis Sobol., 2n = 14; Russia, Irkutskaya Oblast’, Lapenko 10962 (VLA). Chepinoga & al. 10516. Lamium album L., 2n = 18; Russia, Moskovskaya Oblast’, NP Alopecurus arundinaceus Poir., 2n = 28; Russia, Primorskii Krai, & VS 11166. Lapenko 11258. Lycopus europaeus L., 2n = 22; Russia, Krasnodarskii Krai, NP Anisantha sterilis (L.) Nevski, 2n = 14; Russia, Krasnodarskii & VS 11179. Krai, NP & VS 11208. Lycopus lucidus Turcz. ex Benth., 2n = 22; Russia, Primorskii Anthoxanthum odoratum L., 2n = 20; Russia, Krasnoyarskii Krai, Krai, Barkalov 8853. Yakubov 11320. Prunella asiatica Nakai, 2n = 28; Russia, Primorskii Krai, Shata- Apera maritima Klokov, 2n = 14; Russia, Krasnodarskii Krai, lova 7671. NP & VS 11303. Scutellaria krasevii Kom. & I. Schischk., 2n = 16; Russia, Pri- Arthraxon centrasiaticus (Griseb.) Gamajun., 2n = 36; Russia, morskii Krai, Barkalov 8867. Primorskii Krai, Lapenko 11256. Beckmannia syzigachne (Steud.) Fernald, 2n = 14; Russia, Altai, LILIACEAE Olonova 10539, Magadanskaya Oblast’, NP & VS 7131, 7194, Fritillaria camschatcensis (L.) Ker-Gawl., 2n = 24; Russia, Ka- Primorskii Krai, Shatalova 7673. mchatskii Krai, Barkalov 7366. Bromopsis inermis (Leyss.) Holub, 2n = 56; Russia, Amurskaya Lilium debile Kittlitz, 2n = 24; Russia, Kamchatskii Krai, Oblast’, Timchenko 11329. Barkalov 7364. Bromus squarrosus L., 2n = 14; Russia, Volgogradskaya Oblast’, Lilium distichum Nakai, 2n = 24; Russia, Khabarovskii Krai, Yurtzeva 10485. 2n = 14+0–4B; Russia, Krasnodarskii Krai, Barkalov 9916. NP & VS 11292. Calamagrostis arundinacea (L.) Roth, 2n = 28; Russia, Karelia, LIMONIACEAE NP & VS 11324, Khakassia, Yakubov 11244. Limonium caspium (Willd.) Gams, 2n = 18; Russia, Krasnodarskii Calamagrostis epigeios (L.) Roth, 2n = 28; Russia, Karelia, NP Krai, NP & VS 11288. & VS 11254. Calamagrostis extremiorientalis (Tzvelev) Prob., 2n = 28; Russia, MALVACEAE Amurskaya Oblast’, Timchenko 11159. Abutilon theophrasti Medik., 2n = 28; Russia, Krasnodarskii Calamagrostis monticola V. Petrov ex Kom., 2n = 28; Russia, Krai, NP & VS 11285. Khabarovskii Krai, Yermoshkin 9759. Althaea hirsuta L., 2n = 42; Russia, Krasnodarskii Krai, NP & Calamagrostis obtusata Trin., 2n = 28; Russia, Krasnoyarskii VS 11331. Krai, Yakubov 11249. Hibiscus trionum L., 2n = 56; Russia, Krasnodarskii Krai, NP Сatabrosa aquatica (L.) P. Beauv., 2n = 20; Russia, Altai, Olo- & VS 11184. nova 10545. Malva mohileviensis Downar, 2n = 56; Russia, Irkutskaya Oblast’, Сinna latifolia (Trevir.) Griseb., 2n = 28; Russia, Magadanskaya NP & VS 11015. Oblast’, Mochalova 7243. Dactylis glomerata L., 2n = 28; Russia, Krasnoyarskii Krai, ONAGRACEAE Yakubov 11218. Epilobium maximowiczii Hausskn., 2n = 36; Russia, Primorskii Digitaria ciliaris (Retz.) Koeler, 2n = 36; Russia, Krasnodarskii Krai, Barkalov 7510. Krai, NP & VS 11183. Echinochloa oryzicola (Vasinger) Vasinger, 2n = 36; Russia, Pri- ORCHIDACEAE morskii Krai, Lukacheva 9625, 9630, 9631. Oreorchis patens (Lindl.) Lindl., 2n = 48; Russia, Primorskii Echinochloa spiralis Vasinger, 2n = 36; Russia, Krasnodarskii Krai, Barkalov 7767. Krai, Manilo 11168, Primorskii Krai, Lukacheva 9629. Elymus caninus (L.) L., 2n = 28; Russia, Karelia, NP & VS 11308, OXALIDACEAE Krasnoyarskii Krai, Yakubov 11127. Xanthoxalis corniculata (L.) Small, 2n = 24; Russia, Amurskaya Elymus charkeviczii Prob, 2n = 28; Russia, Yakutia, NP & VS Oblast’, NP & VS 10692. 3753. Elymus fibrosus (Schrenk) Tzvelev, 2n = 28; Russia, Irkutskaya PINACEAE Oblast’, NP & VS 10726, Buryatia, NP 6918. Picea koraiensis Nakai, 2n = 24; China, Heilongjiang Province, Elymus franchetii Kitag., 2n = 42; Russia, Khabarovskii Krai, Il’ina 11304. NP & VS 7083. Elymus hyperarcticus (Polunin) Tzvelev, 2n = 28; Russia, POACEAE Magadanskaya Oblast’, Petrovskii 11-64. Achnatherum sibiricum (L.) Keng ex Tzvelev, 2n = 24; Russia, Elymus novae-angliae (Scribn.) Tzvelev, 2n = 28; Russia, Amur- Tuva, Yakubov 11213, 11217. skaya Oblast’, NP & VS 10727, Khabarovskii Krai, Van 11327. Agropyron desertorum (Fisch. ex Link) Schult., 2n = 14; Russia, Elymus sibiricus L., 2n = 28; Russia, Altai, Olonova 10531; Rus- Saratovskaya Oblast’, NP & VS 11305. sia, Tuva, Yakubov 11211.

1286 TAXON 58 (4) • November 2009: 1281–1289 Marhold (ed.) • IAPT/IOPB chromosome data 8

Elytrigia pontica (Podp.) Holub, 2n = 42; Russia, Krasnodarskii Puccinellia hauptiana V.I. Krecz., 2n = 28; Russia, Magadans- Krai, NP & VS 11301. kaya Oblast’, NP & VS 7264. Elytrigia repens (L.) Nevski, 2n = 42; Russia, Krasnodarskii Puccinellia macranthera V.I. Krecz., 2n = 56; Russia, Buryatia, Krai, NP & VS 11281, Saratovskaya Oblast’, NP & VS 11311, Chepinoga & Dulepova 10852. Amurskaya Oblast’, Timchenko 11070, 11071. Puccinellia tenuiflora (Griseb.) Scribn. & Merr., 2n = 28; Russia, Eragrostis amurensis Prob., 2n = 40; Russia, Chitinskaya Oblast’, Buryatia, Chepinoga & Dulepova 10855. NP & VS 11265. Schedonorus pratensis (Huds.) P. Beauv., 2n = 14; Russia, Kare- Festuca altaica Trin., 2n = 28; Russia, Altai, Olonova 10536. lia, NP & VS 11316. Festuca sphagnicola B. Keller, 2n = 28; Russia, Krasnoyarskii Setaria pumila (Poir.) Roem. & Schult., 2n = 36; Russia, Kras- Krai, Yakubov 11220, 11245, 11247, 11248. nodarskii Krai, NP & VS 11171. Glyceria triflora (Korsh.) Kom., 2n = 20; Russia, Krasnoyarskii Setaria verticillata (L.) P. Beauv., 2n = 36; Russia, Saratovskaya Krai, Yakubov 11131, Khabarovskii Krai, Van 11318. Oblast‘, NP & VS 11310. Hierochloë glabra Trin., 2n = 28; Russia, Amurskaya Oblast’, Setaria viridis (L.) P. Beauv., 2n = 18; Russia, Krasnodarskii NP & VS 11104. Krai, Manilo 11273. Hordeum jubatum L., 2n = 28; Russia, Irkutskaya Oblast’, NP Stipa capillata L., 2n = 44; Russia, Tuva, Yakubov 11233. & VS 11106. Trisetum altaicum Roshev., 2n = 14; Russia, Buryatia, Ved- Hordeum murinum L., 2n = 14; Russia, Krasnodarskii Krai, ernikova 11230. Manilo 11277. Trisetum mongolicum (Hult.) Peschkova, 2n = 28; Russia, Altai, Koeleria altaica (Domin) Krylov, 2n = 14; Russia, Altai, Olo- Olonova 10533. nova 10530. Trisetum sibiricum Rupr., 2n = 14; Russia, Tuva, Yakubov 11224. Koeleria cristata (L.) Pers., 2n = 14; Russia, Krasnodarskii Krai, NP & VS 11302. POLYGONACEAE Leersia oryzoides (L.) Sw., 2n = 48; Russia, Krasnodarskii Krai, Polygonum calcatum Lindm., 2n = 20; Russia, Magadanskaya NP & VS 11181. Oblast’, NP & VS 7188. Leymus angustus (Trin.) Pilg., 2n = 84; Russia, Tuva, Yakubov Polygonum maritimum L., 2n = 20; Russia, Krasnodarskii Krai, 11221. NP & VS 11290. Lolium perenne L., 2n = 14; Russia, Krasnodarskii Krai, NP & Rumex amurensis F. Schmidt ex Maxim., 2n = 20; China, NP VS 11312. & VS 8152. Panicum ruderale (Kitag.) Chang, 2n = 36; Russia, Irkutskaya Rumex crispus L., 2n = 40; Russia, Khabarovskii Krai, Barkalov Oblast’, Chepinoga & Enuschenko 10880. 10080. Phleum nodosum L., 2n = 14; Russia, Krasnodarskii Krai, NP Rumex patientia L., 2n = 40; Russia, Primorskii Krai, Barkalov 8831. & VS 11299. Rumex pseudonatronatus (Borb.) Borb. ex Murb., 2n = 40; Rus- Phleum phleoides (L.) H. Karst., 2n = 28; Russia, Tuva, Yakubov sia, Khabarovskii Krai, Barkalov 10081. 11215. Rumex sylvestris (Lam.) Wallr., 2n = 20; Russia, Krasnodarskii Poa alpigena (Blytt) Lindm., 2n = ca. 56; Russia, Magadanskaya Krai, NP & VS 11276. Oblast’, Lyssenko 10655; 2n = 56; Russia, Arkhangel’skaya Oblast’, Timofeeva & Kutenkov 11173. PRIMULACEAE Poa altaica Trin., 2n = 28; Russia, Altai, Olonova 10544. Lysimachia barystachys Bunge, 2n = 24; Russia, Primorskii Krai, Poa angustifolia L., 2n = 56; Russia, Saratovskaya Oblast’, NP & Lapenko 11048. VS 11323, Chitinskaya Oblast’, Tsyrenova 10818. [Probatova Androsace amurensis Prob., 2n = 20; Russia, Irkutskaya Oblast’, & al., Taxon 57: 1273, E15. 2008, as “Poa pratensis”.] Kazanovskii 11059. Poa attenuata Trin., 2n = 28; Russia, Altai, Olonova 10542. Androsace septentrionalis L., 2n = 20; Russia, Primorskii Krai, Poa botryoides (Trin. ex Griseb.) Kom., 2n = 28; Russia, Tuva, Barkalov 8943. Yakubov 11226. Poa bulbosa L., 2n = 28; Russia, Krasnodarskii Krai, NP & VS RANUNCULACEAE 11300. Anemonidium dichotomum (L.) Holub, 2n = 28; Russia, Primor- Poa compressa L., 2n = 42; Russia, Karelia, NP & VS 11203. skii Krai, Barkalov 9236. Poa krylovii Reverd., 2n = 42; Russia, Tuva, Yakubov 11212. Consolida paniculata (Host) Schur, 2n = 16; Russia, Krasnodar- Poa lapponica Prokudin, 2n = 28; Russia, Karelia, NP & VS 11262. skii Krai, NP & VS 11192. Poa palustris L., 2n = 28; Russia, Amurskaya Oblast’, Tim- Ranunculus chinensis Bunge, 2n = 16; Russia, Primorskii Krai, chenko 11065, Khabarovskii Krai, Van 11267, Primorskii Barkalov 8828. Krai, Barkalov 7518. Ranunculus gmelinii DC., 2n = 16; Russia, Primorskii Krai, Poa sibirica Roshev., 2n = 14; Russia, Krasnoyarskii Krai, Barkalov 7520. Yakubov 11135, 11250. Ranunculus japonicus Thunb., 2n = 14; Russia, Primorskii Krai, Poa skvortzovii Prob., 2n = 28; Russia, Amurskaya Oblast’, Tim- Lapenko 11033. chenko 11068, Khabarovskii Krai, Van 11268. 2n = 42; Russia, Ranunculus repens L., 2n = 32; Russia, Amurskaya Oblast’, Tim- Khabarovskii Krai, NP & VS 3261. chenko 11069, Khabarovskii Krai, Barkalov 9897, Primorskii Poa sugawarae Ohwi, 2n = 28; Russia, Sakhalinskaya Oblast’, Krai, Barkalov 9230. Barkalov 8596. Poa trivialis L., 2n = 14; Russia, Karelia, NP & VS 11196. ROSACEAE Psathyrostachys juncea (Fisch.) Nevski, 2n = 14; Russia, Kras- Duchesnea indica (Andr.) Focke, 2n = 42; U.S.A., Texas, Rudyka noyarskii Krai, Yakubov 11229. 9724. Puccinellia distans (Jacq.) Parl., 2n = 42; Russia, Primorskii Krai, Fragaria mandshurica Staudt, 2n = 14; Russia, Khabarovskii NP & VS 9607. Krai, Barkalov 11360.

1287 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: 1281–1289

Rosa rugosa Thunb., 2n = 14; Russia, Primorskii Krai, Sha- Mohammad Mehdi Dehshiri & Rezvan Bohloli tokhina 58. Spiraea media F. Schmidt, 2n = 18; Russia, Irkutskaya Oblast’, Department of Biology, Faculty of Science, Islamic Azad NP & VS 11243. University (Boroujerd Branch), Boroujerd, Lorestan, Iran. [email protected] (author for correspondence) SAXIFRAGACEAE Bergenia crassifolia (L.) Fritsch, 2n = 34; Russia, Buryatia, All materials CHN; vouchers in TARI. Chepinoga 9422. This study was supported by the grant no. 11230506852009, SCROPHULARIACEAE from Islamic Azad University (Boroujerd Branch) for Basic Re- Kickxia elatine (L.) Dumort., 2n = 18; Russia, Krasnodarskii search (IAUB). Krai, NP & VS 11296. Limosella aquatica L., 2n = 40; Russia, Primorskii Krai, Barkalov FABACEAE 9271. Astragalus (sect. Incani) curvirostris Boiss. subsp. curvirostris, Linaria genistifolia (L.) Mill., 2n = 12; Russia, Krasnodarskii 2n = 16; Iran, Lorestan, M.M. Dehshiri & R. Bohloli 91609, Krai, NP & VS 11283. M.M. Dehshiri & R. Bohloli 91610. Linaria vulgaris L., 2n = 12; Russia, Krasnodarskii Krai, Manilo Astragalus (sect. Incani) cyclophyllos Beck, 2n = 16; Iran, Lores- 11193, Irkutskaya Oblast’, NP & VS 10734. tan, M.M. Dehshiri & R. Bohloli 91611. Verbascum thapsus L., 2n = 36; Russia, Karelia, NP & VS Astragalus (sect. Onobrychoidei) effusus Bunge, 2n = 32; Iran, 11198. Lorestan, M.M. Dehshiri & R. Bohloli 91612, M.M. Dehshiri Veronica daurica Steven, 2n = 34; Russia, Primorskii Krai, La- & R. Bohloli 91613. penko 11096. Astragalus (sect. Incani) kabutarlanensis Dehshiri & Maas- soumi, 2n = 16; Iran, Lorestan, M.M. Dehshiri & R. Bohloli TRILLIACEAE 91614. Trillium camschatcense Ker-Gawl., 2n = 10; Russia, Kamchatskii Astragalus (sect. Incani) supervisus (Kuntze) Sheld., 2n = 16; Krai, Barkalov 7365. Iran, Lorestan, M.M. Dehshiri & R. Bohloli 91615.

VIOLACEAE Viola aff. acuminata Ledeb., 2n = 20; Russia, Primorskii Krai, Johan J. Spies1, Paula Spies1, Adre Minnaar1, Susan M.C. Barkalov 9223. Reinecke1, Hanno (J.L.) du Preez1 & Riana Kleynhans1, 2

1 Department of Genetics (64), University of the Free State, María Santos-Vicente, Enrique Rico & M. Montserrat P.O. Box 339, Bloemfontein 9300, South Africa. spiesjj.sci@ Martínez-Ortega ufs.ac.za (author for correspondence) 2 ARC Roodeplaat-Vegetable and Ornamental Plant Institute, Departmento de Botánica, Universidad de Salamanca, 37007 Private Bag X293, 0001 Pretoria, South Africa. Salamanca, Spain. [email protected] (author for correspondence) All materials CHN; vouchers BLFU.

All materials CHN; collectors: AA = A. Abad, CM = HYACINTHACEAE C. Molina, LD = L. Delgado, LG = L.P. Gavilán, LM = L.M. Lachenalia algoensis Schönland, 2n = 14; South Africa, Spies Muñoz-Centeno, MO = M.M. Martínez-Ortega, MS = M. Santos- 7988. Vicente, JA = J.A. Sánchez-Agudo, SA = S. Andrés; vouchers in Lachenalia aloides var. aurea (Lindl.) Engl., 2n = 14; South Af- SALA. rica, Spies 7060. Lachenalia aloides var. luteola G.D. Duncan, 2n = 14; South This study was supported by the Spanish Dirección General de Africa, Spies 7061. Investigación (Ministerio de Ciencia y Tecnología and Ministerio Lachenalia aloides var. quadricolor (Jacq.) Engl., 2n = 14; South de Educación y Ciencia) through the projects Flora Iberica VI Africa, Spies 7062. (REN2002-04634-C05-02; grant ref. BES-2003-2213 to M. Santos Lachenalia aloides var. vanzyliae W.F. Barker, 2n = 28; South Vicente) and Flora Iberica VII (CGL2005-05471-C04-03) and by Africa, Spies 7063. the Junta de Castilla y León through project SA142A08. We greatly Lachenalia arbuthnotiae W.F. Barker, 2n = 14, n = 7; South Af- acknowledge this support. rica, Spies 8054. Lachenalia attenuata W.F. Barker ex. G.D. Duncan, 2n = 14; ASTERACEAE South Africa, Spies 8164. Inula bifrons L., 2n = 16; Spain, MS 948 & LD; France, MS 777 Lachenalia bolusii W.F. Barker, 2n = 18; South Africa, Spies 7987. & LD. Lachenalia bulbifera (Cyr.) Engl., 2n = 28; South Africa, Spies Inula conyzae (Griess.) Meikle, 2n = 32; Spain, MS 597 & LG, 7064. MS 599 & LD. Lachenalia carnosa Bak., 2n = 16; South Africa, Spies 6996, Inula ensifolia L., 2n = 16; Italy, MO, LM, LD & MS 600. 6998. n = 8+0–2B; South Africa, Spies 6934. Inula helenium L., 2n = 20; Spain, MS 957, LD & CM. Lachenalia comptonii W.F. Barker, 2n = 20; South Africa, Spies Inula helvetica Grauer, 2n = 16; Spain, JA, MS 407 & SA, MS 8135. 752, MO, SA & AA. Lachenalia contaminata Ait., 2n = 14; South Africa, Spies 7989. Lachenalia doleritica G.D. Duncan, 2n = 18; South Africa, Spies 7985.

1288 TAXON 58 (4) • November 2009: 1281–1289 Marhold (ed.) • IAPT/IOPB chromosome data 8

Lachenalia elegans W.F. Barker, 2n = 28, n = 14; South Africa, of Shanghai Subject Chief Scientist (no. 08XD14016), and 211 Spies 8014, 8015, 8016, 8018, 8020, 8021, 8022, 8023, 8025, Project. 8026, 8027. 2n = 42, n = 21; South Africa, Spies 8019, 8024. Lachenalia elegans W.F. Barker var. elegans, 2n = 14; South FRULLANIACEAE Africa, Spies 8152. Frullania ericoides (Nees) Nees & Mont., n = 8 (♂); China, R.-L. Lachenalia elegans var. suaveolens W.F. Barker, 2n = 28; South Zhu & al. 2006042502. Africa, Spies 8165. Frullania fengyangshanensis R.L. Zhu & M.L. So, n = 9 (♀); Lachenalia fistulosa Bak., 2n = 14; South Africa, Spies 6936, China, R.-L. Zhu & al. 2006072907; n = 8 (♂); China, R.-L. 8056, 8057, 8058, 8060, 8063, 8136. Zhu & al. 2006072901. Lachenalia hirta (Thunb.) Thunb., 2n = 22; South Africa, Spies Frullania moniliata (Reinw. & al.) Mont., n = 9; China, R.-L. 6858, 6859, 7381. Zhu & al. 2006072801. Lachenalia juncifolia Bak., 2n = 22; South Africa, Spies 8166. Frullania motoyana Steph., n = 9; China, R.-L. Zhu & al. Lachenalia liliflora Jacq., 2n = 16; South Africa, Spies 8163. 2003120206. Lachenalia mathewsii W.F. Barker, 2n = 14; South Africa, Spies Frullania muscicola Steph., n = 9; China, R.-L. Zhu & al. 8150. 2004051309, R.-L. Zhu 20040703. Lachenalia maximiliani Schltr. ex W.F. Barker, 2n = 16; South Frullania tamarisci (L.) Dumort., n = 9; China, R.-L. Zhu & al. Africa, Spies 8153. 2004-9307a. Lachenalia mediana Jacq., 2n = 18, n = 9; South Africa, Spies 7086, 7087. GEOCALYCACEAE Lachenalia mediana var. rogersii (Bak.) W.F. Barker, n = 13; Heteroscyphus coalitus (Hook.) Schiffn., n = 9; China, R.-L. Zhu South Africa, Spies 7079. & al. 2006073102. Lachenalia mutabilis Sweet, 2n = 12, n = 6; South Africa, Spies Heteroscyphus planus (Mitt.) Schiffn., n = 9; China, R.-L. Zhu 8001, 8006, 8007, 8008. 2n = 14, n = 7; South Africa, Spies & al. 20040503004, R.-L. Zhu & al. 2006073105. 8000, 8002, 8003, 8004, 8005, 8009, 8010, 8011, 8012, 8013. Lophocolea bidentata (L.) Dumort., n = 18; China, R.-L. Zhu & Lachenalia namaquensis Schltr. ex W.F. Barker, 2n = 16, n = 8; al. 2006073002. n = 9; China, R.-L. Zhu & al. 2003112603. South Africa, Spies 6969, 6970. Lophocolea ciliolata (Nees) Gottsche, n = 9; China, R.-L. Zhu & Lachenalia orchioides (L.) Ait., 2n = 14, n = 7; South Africa, al. 2005090405, R.-L. Zhu & al. 2003112802. Spies 8068, 8069, 8071, 8072, 8073, 8074, 8075, 8079, 8080, 8082, 8083, 8084, 8085, 8086, 8089, 8090, 8091, 8092, 8093, HAPLOMITRIACEAE 8094, 8095, 8097, 8098, 8099, 8100, 8101. Haplomitrium mnioides (Lindb.) R.M. Schust., n = 9; China, R.-L. Lachenalia orthopetala Jacq., 2n = 16; South Africa, Spies 7986. Zhu & al. 20049706. Lachenalia pallida Ait., 2n = 16; South Africa, Spies 7089, 8157. Lachenalia peersii Marloth ex. W.F. Barker, 2n = 14; South Af- JACKIELLACEAE rica, Spies 8159. Jackiella javanica Schiffn., n = 8; China, R.-L. Zhu & al. Lachenalia purpureo-caerulea Jacq., 2n = 16; South Africa, 2005090103; Singapore, R.-L. Zhu 20070124-12. Spies 6939. Lachenalia pusilla Jacq., 2n = 18; South Africa, Spies 8155. JUNGERMANNIACEAE Lachenalia reflexa Thunb., 2n = 14; South Africa, Spies 7090, Hattoria yakushimensis (Horik.) R.M. Schust., n = 9; China, R.-L. 7091. Zhu & al. 20049509. Lachenalia rosea Andr., 2n = 28; South Africa, Spies 8167, 8168. Nardia assamica (Mitt.) Amakawa, n = 8+m; China, R.-L. Zhu Lachenalia rubida Jacq., 2n = 14; South Africa, Spies 7093, & al. 2006072910. 8156. Lachenalia splendida Diels, 2n = 16, n = 8; South Africa, Spies LEPIDOLAENACEAE 6796, 6977. Trichocoleopsis sacculata (Mitt.) S. Okamura, n = 9; China, R.-L. Lachenalia unifolia Jacq., 2n = 22, n = 11; South Africa, Spies Zhu & al. 2006073007. 6861, 6863, 6864, 6866, 6867, 6871, 6872, 6874, 6878, 6879, 6881, 6889, 6891, 6892, 6898. LEPIDOZIACEAE Lachenalia cf. unifolia, 2n = 22; South Africa, Spies 6899, 6902. Bazzania tridens (Reinw. & al.) Trevis., n = 10; China, R.-L. Zhu Lachenalia violacea Jacq., n = 7; South Africa, Spies 7101, 7104. & al. 2004051312 , R.-L. Zhu & al. 2004051313, R.-L. Zhu & al. 2003112701.

Huan Hang1, Min Zheng1,2 & Rui-Liang Zhu1 LOPHOZIACEAE Plicanthus birmensis (Steph.) R.M. Schust., n = 9; China, R.-L. 1 Department of Biology, School of Life Science, East China Zhu & al. 20049307. Normal University, 3663 Zhong Shan North Road, Shanghai 200062, China. [email protected] (author for corre- PORELLACEAE spondence) Porella chinensis (Steph.) S. Hatt., n = 8; China, R.-L. Zhu & 2 School of Environmental Studies, China University of Geo- al. 200502114. sciences, 388 Lumo Road, Wuhan 430074, China Porella densifolia (Steph.) S. Hatt., n = 8; China, R.-L. Zhu & al. 2006042606. All materials CHN, vouchers in HSNU. Porella obtusata var. macroloba (Steph.) S. Hatt. & M.X. Zhang, n = 8, China, R.-L. Zhu & al. 2006072814. This study was supported by the National Natural Science Porella ulophylla (Steph.) S. Hatt, n = 8; China, R.-L. Zhu & al. Foundation of China (nos. 30825004, 30670155), the Program 2006042503, R.-L. Zhu & al. 2005042801.

1289 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

IOPB COLUMN

Edited by Karol Marhold & Ilse Breitwieser

IAPT/IOPB chromosome data 8

Edited by Karol Marhold

Victor V. Chepinoga, Aleksandr A. Gnutikov, Olovyanninskii Raion, 14 km NNE of Olovyannaya, right riverside Ilya V. Enushchenko & Sergey A. Rosbakh of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, 19 Jun 2007, Gnutikov & Enushchenko C413 (IRKU). Department of Botany & Genetics, Irkutsk State University, 1 Karl Marks Str., 664003, Irkutsk, Russia. Victor.Chepinoga@ Carex duriuscula C.A. Mey. gmail.com ▼**2n = 62, CHN. Russia, Buryatiya Republic, Mukhorshi- birskii Raion, 15 km NNW of Mukhorshibir’, close to waterbody * First chromosome count for the species. on one of inflows of Tugnui river, in steppe, 51°11′N, 107°16′E, 14 ** New chromosome number (cytotype) for the species. Jun 2007, Gnutikov & Enushchenko C363 (IRKU). ▼ First chromosome count from an East Siberian accession. Carex enervis C.A. Mey. This study was supported by Grants no. 05-05-64061, 07-04- *2n = 44–46, CHN. Russia, Buryatiya Republic, Mukhorshi- 00610 from Russian Fund for Basic Research, RNP.2.2.1.1.7334 birskii Raion, 15 km NNW of Mukhorshibir’, waterbody on one and RNP.2.2.3.1.4667 from Ministry of Education and Science of of inflows of Tugnui river, on the shore, 51°11′N, 107°16′E, 13 Jun Russian Federation. 2007, Gnutikov & Enushchenko C359 (IRKU).

CYPERACEAE Carex juncella (Fr.) Th. Fr. Blysmus rufus (Huds.) Link ▼**2n = 52, CHN. Russia, East Siberia, Irkutskaya Oblast’, ▼2n = 40, CHN. Russia, East Siberia, Buryatiya Republic, Tulunskii Raion, 2 km S of Krasnoozerskii village, right riverside Tarbagataiskii Raion, Tarbagatai village, right riverside of Kui- of Iya river, opposite of Evdokimovskii village, in water, 54°13′N, tunka river, 51°28′N, 107°22′E, 13 Jun 2007, Gnutikov & Enush- 100°42′E, 01 Jul 2007, Gnutikov & Enushchenko C464 (IRKU). chenko С357 (IRKU); Russia, East Siberia, Chitinskaya Oblast’, Uletovskii Raion, 4 km N of Ablatuiskii Bor village, bitterly-salt Carex reptabunda (Trautv.) V.I. Krecz. lake Selitryanoe, solonetz meadow at the shore, 51°13′34″N, *2n = 44, CHN. Russia, East Siberia, Chitinskaya Oblast’, 112°14′16″E, 10 Aug 2007, Gnutikov C605 (IRKU). Uletovskii Raion, 4 km N of Ablatuiskii Bor village, bitterly- salt lake Selitryanoe, solonetz meadow along the shoreline, Bolbosc hoenus planiculmis (F. Schmidt) T.V. Egorova 51°13′34″N, 112°14′16″E, 15 Jun 2007, Gnutikov & Enushchenko ▼2n = 52, CHN. Russia, East Siberia, Chitinskaya Oblast’, C379 (IRKU). Uletovskii Raion, 4 km N of Ablatuiskii Bor village, bitterly- salt lake Selitryanoe, solonetz meadow at the shore, 51°13′34″N, Carex vesicata Meinsh. 112°14′16″E, 10 Aug 2007, Gnutikov C607 (IRKU). *2n = 54, CHN. Russia, East Siberia, Chitinskaya Oblast’, Olovyanninskii Raion, 14 km NNE of Olovyannaya, right riverside Carex appendiculata (Trautv. & C.A. Mey.) Kük. of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, 17 Jun ▼2n = 78, CHN. Russia, East Siberia, Chitinskaya Oblast’, 2007, Gnutikov & Enushchenko C389, C390 (IRKU). Olovyanninskii Raion, 14 km NNE of Olovyannaya, right riverside of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, 17 Jun Eleocharis acicularis (L.) Roem. & Schult. 2007, Gnutikov & Enushchenko C388 (IRKU). ▼2n = 18, CHN. Russia, East Siberia, Chitinskaya Oblast’, Olovyanninskii Raion, 14 km NNE of Olovyannaya, right riverside Carex atherodes Spreng. of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, 17 Jun ▼**2n = ca. 78, CHN. Russia, East Siberia, Chitinskaya 2007, Gnutikov & Enushchenko C393 (IRKU). Oblast’, Aginskii Buryatskii autonomic Region, Mogoytuyskii Raion, 1 km NW of Ononsk village, right riverside of Onon river, Eleocharis palustris (L.) Roem. & Schult. in water, 51°00′N, 115°34′E, 19 Jun 2007, Gnutikov C419 (IRKU). 2n = 16, CHN. Russia, East Siberia, Chitinskaya Oblast’, Olovyanninskii Raion, 14 km NNE of Olovyannaya, right river- Carex bohemica Schreb. side of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, ▼2n = ca. 62, CHN. Russia, East Siberia, Chitinskaya Oblast’, 17 Jun 2007, Gnutikov & Enushchenko C392 (IRKU).

E1 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Scirpus radicans Schkuhr Oka River, close to willow’s thickets, 53°36′N, 101°50′E, 07 Jul ▼2n = 56, CHN. Russia, East Siberia, Chitinskaya Oblast’, 2005, Gnutikov & V. Mozer C156 (IRKU). Olovyanninskii Raion, 14 km NNE of Olovyannaya, right riverside of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, 17 Jun Vicia unijuga A. Br. 2007, Gnutikov & Enushchenko C398 (IRKU). 2n = 12, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tu- lunskii Raion, 2 km N of Krasnoozerskii village, right riverside of FABACEAE Iya River, on the shore, 54°17′N, 100°42′E, 02 Jul 2007, Gnutikov Amoria hybrida (L.) C. Presl & Enushchenko C471 (IRKU). 2n = 16, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tai- shetskii Raion, 5 km NE of Staryi Akul’shet village, left riverside NITRARIACEAE of Biryusa river, 56°02′N, 98°02′E, 07 Jul 2007, Gnutikov & A. Hoff Nitraria sibirica Pall. C509 (IRKU). ▼2n = 60, CHN. Russia, East Siberia, Buryatia Republic, Barguzinskii Raion, to the North of Dushelan village, surroundings Amoria montana (L.) Soják of Bol’shoe Alginskoe lake, solonchak, 53°38′23″N, 109°56′19″E, ▼2n = 16, CHN. Russia, East Siberia, Irkutskaya Oblast’, 30 Jul 2004, Chepinoga C642 (IRKU). Taishetskii Raion, 5 km NE of Staryi Akul’shet village, left riverside of Biryusa river, 56°02′N, 98°02′E, 07 Jul 2007, Gnutikov & POACEAE A. Hoff C510 (IRKU). Calamagrostis obtusata Trin. 2n = 28, CHN. Russia, East Siberia, Tyva Republic, Todzhin- Amoria repens (L.) C. Presl skii Raion, East Sayan mountains, left riverside of Arzhaan-Khem 2n = 32, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tu- river, the shore of mineral spring Zhoigan, 52°33′N, 98°45′E, 10 lunskii Raion, 2 km W of Uygat village, floodplain of left riverside Aug 2007, E. Vedernikova C643 (IRKU). Kirei river, on the field road, 54°05′N, 100°34′E, 30 Jun 2007, Gnutikov & Enushchenko С451 (IRKU). Eragrostis pilosa (L.) P. Beauv. ▼2n = 40, CHN. Russia, East Siberia, Chitinskaya Oblast’, Astragalus olchonensis Gontsch. 5 km SW of Cheremkhovo village, left riverside of Ingoda river, *2n = 16, CHN. Russia, East Siberia, Irkutskaya Oblast’, 51°26′03″N, 112°40′43″E, 10 Aug 2007, Gnutikov & Enushchenko Olkhonskii Raion, Baikal lake, west coast of Olkhon island, C610 (IRKU). Peschanaya bay, on the sand dunes, 53°17′00″N, 107°35′00″E, Festuca ovina L. 30 Aug 2004, Chepinoga, A. Verkhozina & E. Kuznetsova С634 2n = 28, CHN. Russia, East Siberia, Irkutskaya Oblast’, Slu- (IRKU). dyanskii Raion, Khamar-Daban mountains, valley of Sludyanka river, hill’s slope by pathway to the weather station, 51°36′N, Lathyrus humilis (Ser.) Spreng. 103°38′E, 15 Jun 2008, Enushchenko C623 (IRKU). 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Ziminskii Raion, Batama village, larch-pine forest, 53°54′N, Milium effusum L. 101°36′E, 10 Jul 2005, Gnutikov & Chepinoga C185 (IRKU). ▼2n = 28, CHN. Russia, East Siberia, Tyva Republic, Todzhinskii Raion, East Sayan mountains, the small island on Lathyrus pratensis L. Arzhaan-Khem river, in willow’s thickets, 52°33′N, 98°45′E, 10 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tu- Aug 2007, E. Vedernikova C635 (IRKU). lunskii Raion, 2 km N of Krasnoozerskii village, right riverside of Iya River, on the shore, 54°17′N, 100°42′E, 02 Jul 2007, Gnutikov Poa irсutica Roshev. & Enushchenko C474 (IRKU). 2n = 28, CHN. Russia, East Siberia, Irkutskaya Oblast’, Slu- dyanskii Raion, Khamar-Daban mountains, valley of Sludyanka Melilotoides platycarpos (L.) Soják river, by pathway to the weather station, 51°36′N, 103°38′E, 14 Jun ▼2n = 16, CHN. Russia, East Siberia, Irkutskaya Oblast’, 2008, Enushchenko C624 (IRKU). Taishetskii Raion, Shelaevo village, the shore of Solonetskoe lake, on the road 56°56′N, 97°40′E, 10 Jul 2007, Gnutikov & Enush- Poa alpina L. chenko C535 (IRKU). 2n = 28, CHN. Russia, East Siberia, Irkutskaya Oblast’, Sludy- anskii Raion, Khamar-Daban mountains, valley of Sludyanka river, Trifolium pratense L. by pathway to the weather station, 51°36′N, 103°38′E, 15 Jun 2008, 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tu- Enushchenko C622 (IRKU). lunskii Raion, 2 km W of Uygat village, left riverside of Kirei river, on the meadow, 54°05′N, 100°34′E, 30 Jun 2007, Gnutikov Setaria pumila (Poir.) Roem. & Schult. & Enushchenko С457 (IRKU). ▼2n = 36, CHN. Russia, East Siberia, Irkutskaya Oblast’, Angarsk city, surroundings of Elovskoe reservoir, in the forest Vicia amoena Fisch. by the road to dacha’s village «Utyes», 52°29′N, 103°48′E, 01 Sep 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tu- 2005, Enushchenko C639 (IRKU). lunskii Raion, 2 km N of Krasnoozerskii village, right riverside of Iya River, on the shore, 54°17′N, 100°42′E, 02 Jul 2007, Gnutikov RANUNCULACEAE & Enushchenko C472 (IRKU). Adonis sibirica (Patrin ex DC.) Ledeb. 2n = 16, CHN. Russia, East Siberia, Irkutskaya Oblast’, south- Vicia cracca L. western part of Angarsk city, right riverside of Kitoi river, sur- 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Zim- roundings of camping “Leninets”, 52°29′N, 103°48′E, 03 Jun 2007, inskii Raion, 3 km SSW of Novoletniki village, left riverside of Enushchenko C437 (IRKU).

E2 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

Anemonastrum crinitum (Juz.) Holub Ranunculus repens L. 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Ust’- 2n = 32, CHN. Russia, East Siberia, Irkutskaya Oblast’, Taish- Ordynskii Buryatskii Autonomous Region, Ekhirit-Bulagatskii etskii Raion, Shitkino village, lake in the village, on the moist Raion, 4 km E of Kharat village, floodplain of Kharat river, border shore, 56°22′N, 98°20′E, 08 Jul 2007, Gnutikov & A. Hoff C518 of birch-pine forest, 52°44′N, 105°04′E, 17 Jun 2005, Gnutikov & (IRKU); Russia, East Siberia, Chitinskaya Oblast’, Chitinskii Chepinoga C141 (IRKU). Raion, nearly from estuary of Kruchina river (left inflow of Ingoda river), in coastal shrubs, 51°50′N, 113°44′E, 15 Jun 2007, Gnutikov Batrachium circinatum (Sibth.) Spach & Enushchenko C382 (IRKU). ▼2n = 16, CHN. Russia, East Siberia, Chitinskaya Oblast’, 5 km SW of Cheremkhovo village, left riverside of Ingoda river, Ranunculus reptans L. 51°26′N, 112°40′E, 11 Aug 2007, Gnutikov C614 (IRKU). 2n = 32, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tulunskii Raion, 2 km W of Uygat village, floodplain of left riv- Halerpestes salsuginosa (Pall. ex Georgi) Greene erside Kirei river, on the field road, 54°05′N, 100°34′E, 30 Jun 2n = 48, CHN. Russia, East Siberia, Buryatiya Republic, 2007, Gnutikov & Enushchenko С449 (IRKU); Russia, East Sibe- Mukhorshibirskii Raion, 15 km NNW of Mukhorshibir’, waterbody ria, Chitinskaya Oblast’, Khilokskii Raion, 5 km SW Khokhotui on one of inflows of Tugnui river, on the shore, 51°11′N, 107°16′E, village, on moist shore, 51°15′N, 109°31′E, 21 Jun 2007, Gnutikov 13 Jun 2007, Gnutikov & Enushchenko C360 (IRKU); Russia, East & Enushchenko C429 (IRKU); Russia, East Siberia, Chitinskaya Siberia, Buryatiya Republic, Mukhorshibirskii Raion, 3 km S of Oblast’, Uletovskii Raion, Goreka village, left riverside of Goreka Sagan-Nur village, salt lake Olon-Sheber, in shallow water, 51°18′N, river, moist shore, 51°04′N, 111°59′E, 07 Aug 2007, Gnutikov & 108°25′E, 05 Aug 2007, Gnutikov & Enushchenko C559 (IRKU). Enushchenko C588 (IRKU).

Halerpestes sarmentosa (Adams) Kom. Ranunculus sceleratus L. 2n = 16, CHN. Russia, East Siberia, Chitinskaya Oblast’, Ule- 2n = 64, CHN. Russia, East Siberia, Chitinskaya Oblast’, tovskii Raion, 4 km N of Ablatuiskii Bor village, bitterly-salt lake Olovyanninskii Raion, 8 km SSE of Olovyannaya, near Topolyovka Selitryanoe, solonetz meadow on the shore, 51°13′N, 112°14′E, 15 Jun village, flood-land meadow of Ingoda river, 50°53′N, 115°37′E, 18 2007, Gnutikov & Enushchenko C376 (IRKU); Russia, East Siberia, Jun 2007, Gnutikov & Enushchenko С400 (IRKU); Russia, East Buryatiya Republic, Tarbagataiskii Raion, Tarbagatai village, right Siberia, Chitinskaya Oblast’, Olovyanninskii Raion, 4 km W of riverside of Kuitunka river, 51°28′N, 107°22′E, 13 Jun 2007, Gnuti- Yasnogorsk, left riverside of Onon river, moist shore, 51°52′N, kov & Enushchenko С355 (IRKU); Russia, East Siberia, Buryatiya 115°38′E, 18 Jun 2007, Gnutikov & Enushchenko C401 (IRKU). Republic, Mukhorshibirskii Raion, 3 km S of Sagan-Nur village, salt lake Olon-Sheber, in shallow water, 51°18′N, 108°25′E, 05 Aug 2007, Gnutikov & Chepinoga C560 (IRKU). Robert Vogt1 & Christoph Oberprieler2

Ranunculus chinensis Bunge 1 Botanischer Garten und Botanisches Museum Berlin- ▼2n = 16, CHN. Russia, East Siberia, Chitinskaya Oblast’, Dahlem, Freie Universität Berlin, Königin-Luise-Str. 6–8, Olovyanninskii Raion, 14 km NNE of Olovyannaya, right riverside 14195 Berlin, Germany. [email protected] (author for cor- of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, 19 Jun respondence) 2007, Gnutikov C420 (IRKU). 2 Institut für Botanik, Universität Regensburg, Universitätsstr. 31, 93053 Regensburg, Germany Ranunculus gmelinii DC. 2n = 16, CHN. Russia, East Siberia, Chitinskaya Oblast’, This contribution is ninth in the series entitled “Chromosome Olov yanninskii Raion, 14 km NNE of Olovyannaya, right river- numbers of North African phanerogams.” (For previous contributions side of Onon river, opposite of Zugol village, 51°02′N, 115°38′E, see Oberprieler & Vogt, 1993, 1996; Vogt & Oberprieler, 1993, 1994, 19 Jun 2007, Gnutikov & Enushchenko C414 (IRKU); Russia, East 2008; Hellwig & al., 1994; Kilian & al., 1995; Scholz & al., 1998;) Siberia, Irkutskaya Oblast’, Tulunskii Raion, 2 km W of Uygat village, left riverside floodplain Kirei river, on the shore, 54°05′N, BORAGINACEAE 100°34′E, 30 Jun 2007, Gnutikov & Enushchenko С459 (IRKU). Cynoglossum creticum Mill. 2n = 24, CHN. Morocco, Monts des Beni-Snassen, Gorges Ranunculus longicaulis C.A. Mey. du Zegzel between the villages of Trashroute and Moulay Ahmed, 2n = 32, CHN. Russia, East Siberia, Chitinskaya Oblast’, slopes of the gorge below and above road, riverbed, 450–550 m, Chitinskii Raion, 5 km N of Lesnoi Godordok, near Staraya Kuka 34°49′N, 2°22′W, 10 May 1993, Vogt 11547 & Oberprieler 5995 (B). village, floodplain of Ingoda river, 51°45′N, 113°04′E, 20 Jun 2007, Gnutikov & Enushchenko С427 (IRKU). Cynoglossum dioscoridis Vill. 2n = 24, CHN. Morocco, Anti-Atlas, Aït-Baha-Valley, Tioulit Ranunculus polyanthemos L. NW, Madao NW, fallow land, marl lime, 1,350 m, 4.1992, Gruber 2n = 16, CHN. Russia, East Siberia, Irkutskaya Oblast’, Tulun- (seed samples acquired from the Botanic Garden Salzburg [sub skii Raion, 2 km W of Uygat village, left riverside Kirei river, on Cynoglossum creticum Mill.]), cultivated at the Botanic Garden the meadow, 54°05′N, 100°34′E, 30 Jun 2007, Gnutikov & Enush- Berlin-Dahlem, 7 Jun 1993, Oberprieler & Vogt s.n. (B). chenko С499 (IRKU). CARYOPHYLLACEAE Ranunculus radicans C.A. Mey. Rhodalsine geniculata (Poir.) F.N. Williams [≡ Minuartia 2n = 16, CHN. Russia, East Siberia, Chitinskaya Oblast’, Ule- geniculata (Poir.) Thell.] tovskii Raion, Nikolaevskoe village, the moist shore of Nikolaevs- 2n = 18, CHN. Morocco, Monts des Beni-Snassen, track koe lake, 51°03′N, 111°44′E, 08 Aug 2007, Gnutikov C592 (IRKU). from Beni Ammar to Taforalt via Tanezzert, settlements around

E3 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Tanezzert between Sidi-Yahia-Ben-Ahmed and Tagliat, roadsides, CISTACEAE field margins, 450 m, 34°48′N, 2°36′W, 8 May 1993, Vogt 11296 Fumana laevipes (L.) Spach & Oberprieler 5744 (B); Morocco, Monts des Beni-Snassen, road 2n = 32, CHN. Morocco, Monts des Beni-Snassen, track from 5311 between Mechrâ-Homadi and Berkane, slopes along the road Beni Ammar to Taforalt via Talezzert, mountain ridge between ca. 1 km E of the junction with road S 412 (Mechrâ-Homadi–El- Djebel Astar-ach-Chaib and Oued Tlata-at-Talremt ca. 3.5 km SE Aioun), Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, of the marabout “Sidi Boubker” in Beni Ammar, pastures and 6 May 1993, Vogt 10945 & Oberprieler 5393 (B). Tetraclinis woodland, 340–370 m, 34°48′N, 2°38′W, 8 May 1993, Vogt 11239 & Oberprieler 5687 (B). Silene aellenii Sennen [= Silene divaricata Lag., nom. illeg.] 2n = 24, CHN. Morocco, Monts des Beni-Snassen, track from Helianthemum apeninnum (L.) Mill. Beni Ammar to Taforalt via Talezzert, mountain ridge between 2n = 20, CHN. Morocco, Tell Atlas, Col de Jerada on road P 19 Djebel Astar-ach-Chaib and Oued Tlata-at-Talremt ca. 3.5 km SE between Aïn-Beni-Mathar and Oujda, roadsides ca. 4 km N of pass, of the marabout “Sidi Boubker” in Beni Ammar, pastures and 1,120 m, 34°21′N, 2°03′W, 4 May 1993, Vogt 10721 & Oberprieler Tetraclinis woodland, 340–370 m, 34°48′N, 2°38′W, 8 May 1993, 5169 (B); Morocco, Monts des Beni-Snassen, road 5311 between Vogt 11211 & Oberprieler 5659 (B). Mechrâ-Homadi and Berkane, valley SE point 351 m ca. 2 km E Mechrâ-Homadi, Tetraclinis articulata woodland, limestone cliffs, Silene colorata Poir. dry oued, 200–250 m, 34°45′N, 2°47′W, 7 May 1993, Vogt 11031 2n = 24, CHN. Morocco, Mediterranean coast W of Saidia, & Oberprieler 5479 (B). surroundings of the mouth of Oued Moulouya ca. 10 km W Saidia, dunes ca. 1 km inland, 10 m, 35°07′N, 2°18′W, 5 May 1993, Vogt Helianthemum papillare Boiss. 10877 & Oberprieler 5325 (B). 2n = 20, CHN. Morocco, Monts des Beni-Snassen, Gorges du Zegzel between the villages of Moulay Ahmed and Takerboust, Silene obtusifolia Willd. riverbed and roadsides, 270–300 m, 34°50′N, 2°22′W, 10 May 2n = 24, CHN. Morocco, Tangier Peninsula, Cap Ras Ciress, 1993, Vogt 11669 & Oberprieler 6117 (B). surroundings of the lighthouse 9,7 km NE of Ksar-es-Seghir, costal rocks, moist places in Pinus woodland, 5–60 m, 35°55′N, 5°28′W, COMPOSITAE 20 Apr 1993, Vogt 9921 & Oberprieler 4369 (B), Vogt 9926 & Erigeron bonariensis L. [≡ Conyza bonariensis (L.) Cron- Oberprieler 4374 (B). quist] 2n = 36, CHN. Morocco, Monts des Beni Snassen, Gorges du Silene ramosissima Desf. Oued Zegzel between Djebel Israne and Djebel Achaoun E Taforalt, 2n = 24, CHN. Morocco, Rharb, road 2301 between Nador S-facing mountain slopes of Djebel Israne near the “Grotte des and Kenitra, ca. 10 km N Benmansoun, dunes with Eucalyptus Pigeons”, 690 m, 34°50′N, 2°24′W, 9 May 1993, Vogt 11480 & plantation, 20 m, 34°40′N, 6°23′W, 25 Apr 1993, Vogt 10925 & Oberprieler 5928 (B). Oberprieler 5373 (B). CRUCIFERAE Silene secundiflora Otth **Alyssum simplex Rudolphi 2n = 24, CHN. Morocco, Monts des Beni-Snassen, track from 2n = 32, CHN. Morocco, Tell Atlas, Col de Jerada on road Beni Ammar to Taforalt via Talezzert, Oued Tlata-at-Talremt ca. P 19 between Aïn-Beni-Mathar and Oujda, roadsides ca. 4 km 2.5 km S of the marabout “Sidi Boubker”, roadsides, limestone N of pass, 1,120 m, 34°21′N, 2°03′W, 4 May 1993, Vogt 10709 & cliffs, 220 m, 34°48′N, 2°39′W, 7 May 1993, Vogt 11188 & Ober- Oberprieler 5157 (B). prieler 5636 (B); Morocco, Monts des Beni-Snassen, road 5311 between Mechrâ-Homadi and Berkane, valley SE point 351 m ca. 2 Anastatica hierochuntica L. km E Mechrâ-Homadi, Tetraclinis articulata woodland, limestone 2n = 22, CHN. Morocco, Oued Ziz, Rissani W 3 km, Ham- cliffs, dry oued, 200–250 m, 34°45′N, 2°47′W, 7 May 1993, Vogt mada, limestone, 850 m, Mar 1992, Gruber (seed sample ac- 11018 & Oberprieler 5466 (B). quired from the Botanic Garden Salzburg), cultivated at the Bo- tanic Garden Berlin-Dahlem, 7 Jun 1993, Oberprieler & Vogt Silene tridentata Desf. s.n. (B). 2n = 24, CHN. Morocco, Monts des Beni-Snassen, Taforalt, N-facing slopes of Djebel Achouan ca. 1 km E Taforalt, stony slopes Biscutella baetica Boiss. & Reut. and limestone rocks, 850–900 m, 34°49′N, 2°24′W, 4 May 1993, 2n = 16, CHN. Morocco, Tangier Peninsula, road 8303 from Vogt 10855b & Oberprieler 5303b (B). Souk-Tleta-Taghramet to Ceuta (Sebta), limestone rocks above Souk-Tleta-Taghramet and around quarry not far from the sum- Spergularia diandra (Guss.) Boiss. mit of the pass, 450–500 m, 35°48′N, 5°27′W, 17 Apr 1993, Vogt 2n = 18, CHN. Morocco, Monts des Beni-Snassen, road 5311 9858 & Oberprieler 4306 (B); Morocco, Jebala, Djebel Sougna between Mechrâ-Homadi and Berkane, ca. 2 km as the crow flies near road S 603 between Chefchaouene and Ksar-el-Kebir, W of NE Mechrâ-Homadi, field margins, 380 m, 34°47′N, 2°45′W, 7 village Doumse ca. 14 km E Tanakoub, river banks, field margins, May 1993, Vogt 11116 & Oberprieler 5564 (B). and sources, 850 m, 35°05′N, 5°24′W, 23 Apr 1993, Vogt 10103 & Oberprieler 4551 (B); Morocco, Rharb, road S 216 between CHENOPODIACEAE Arbaoua and Moulay Bousselham, ca. 3,4 km W of junction with Salsola kali L. road to Lalla-Rhano and Ksar-el-Kebir, ungrazed field margin, 2n = 36, CHN. Morocco, Mediterranean coast W of Saidia, 10 m, 34°51′N, 6°10′W, 24 Apr 1993, Vogt 10190 & Oberprieler surroundings of the mouth of Oued Moulouya ca. 10 km W Saidia, 4638 (B); Morocco, Monts des Beni-Snassen, Taforalt, N-facing dunes ca. 1 km inland, 10 m, 35°07′N, 2°18′W, 5 May 1993, Vogt slopes of Djebel Achouan ca. 1 km E Taforalt, stony slopes and 10901 & Oberprieler 5349 (B). limestone rocks, 850–900 m, 34°49′N, 2°24′W, 4 May 1993, Vogt

E4 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

10786 & Oberprieler 5234 (B); Morocco, Monts des Beni-Snassen, Rorippa nasturtium-aquaticum (L.) Hayek [ = Nasturtium track from Beni Ammar to Taforalt via Talezzert, Oued Tlata-at- officinale R. Br.] Talremt ca. 2.5 km S of the marabout “Sidi Boubker”, roadsides, 2n = 32, CHN. Morocco, Jebala, Djebel Sougna near road limestone cliffs, 220 m, 34°48′N, 2°39′W, 7 May 1993, Vogt 11191 S 603 between Chefchaouene and Ksar-el-Kebir, W of village & Oberprieler 5639 (B). Doumse ca. 14 km E Tanakoub, river banks, field margins, and sources, 850 m, 35°05′N, 5°24′W, 23 Apr 1993, Vogt 10102 & **Brassica fruticulosa subsp. cossoniana (Boiss.& Reut.) Oberprieler 4550 (B). Maire 2n = 16, CHN. Morocco, Monts des Beni-Snassen, road 5311 Rapistrum rugosum (L.) All. subsp. rugosum between Mechrâ-Homadi and Berkane, slopes along the road ca. 1 2n = 16, CHN. Morocco, Monts des Beni-Snassen, road 5311 km E of the junction with road S 412 (Mechrâ-Homadi–El-Aioun), between Mechrâ-Homadi and Berkane, Beni Ammar, surroundings Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, 6 May of Oued Lefrane near Beni Ammar, road embankments, hedges, 1993, Vogt 10946 & Oberprieler 5394 (B); Morocco, Monts des 140 m, 34°50′N, 2°41′W, 7 May 1993, Vogt 11146 & Oberprieler Beni-Snassen, road through the gorge of Oued Zegzel between 5594 (B). Taforalt and Berkane, surroundings of the settlement of Trashroute, field margins, hedges, bushes, 590 m, 34°48′N, 2°22′W, 10 May Sisymbrium irio L. 1993, Vogt 11534 & Oberprieler 5982 (B). 2n = 14, CHN. Morocco, Monts des Beni-Snassen, track from Beni Ammar to Taforalt via Tanezzert, settlements around Carrichtera annua (L.) DC. Tanezzert between Sidi-Yahia-Ben-Ahmed and Tagliat, roadsides, 2n = 16, CHN. Morocco, Monts des Beni-Snassen, road 5311 field margins, 450 m, 34°48′N, 2°36′W, 8 May 1993, Vogt 11297 between Mechrâ-Homadi and Berkane, slopes along the road ca. & Oberprieler 5745 (B). 1 km E of the junction with road S 412 (Mechrâ-Homadi–El- Aioun), Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, EUPHORBIACEAE 6 May 1993, Vogt 10950 & Oberprieler 5398 (B); Morocco, Monts Euphorbia exigua L. subsp. exigua des Beni-Snassen, road 5311 between Mechrâ-Homadi and Ber- 2n = 24, CHN. Morocco, Monts des Beni-Snassen, Taforalt, kane, ca. 2 km as the crow flies NE Mechrâ-Homadi, field margins, N-facing slopes of Djebel Achouan ca. 1 km E Taforalt, stony slopes 380 m, 34°47′N, 2°45′W, 7 May 1993, Vogt 11130 & Oberprieler and limestone rocks, 850–900 m, 34°49′N, 2°24′W, 4 May 1993, 5578 (B). Vogt 10764 & Oberprieler 5212 (B).

Diplotaxis harra (Forssk.) Boiss. subsp. harra Euphorbia helioscopia L. subsp. helioscopia 2n = 26, CHN. Morocco, Prov. Figuig, road P 32 between 2n = 42, CHN. Morocco, Monts des Beni Snassen, Gorges Bouarfa and Figuig, north-facing slopes of Djebel Grouz ca. 23 du Zegzel between the village of Trashroute and Takerboust, sur- km W Figuig, mountains ca. 6 km S of road P 32, rocks and stony roundings of Moulay Ahmed, wet embankments, roadsides, field slopes, 1,100–1,300 m, 32°08′N, 1°24′W, 1 May 1993, Vogt 10540 margins, 380–400 m, 34°50′N, 2°21′W, 10 May 1993, Vogt 11604 & Oberprieler 4988 (B). & Oberprieler 6052 (B).

Eruca vesicaria subsp. sativa (Mill.) Thell. Euphorbia hirsuta L. 2n = 22, CHN. Morocco, Mediterranean coast NW Oujda, 2n = 14, CHN. Morocco, Rharb, Moulay Bousselham, la- Ras-el-Ma, surroundings of lighthouse at Ras Kebdana, maritime guna Merdja Zerga, moist meadows and marshes, 0–5 m, 34°53′N, sands, 40 m, 35°09′N, 2°25′W, 5 May 1993, Vogt 10934 & Ober- 6°16′W, 25 Apr 1993, Vogt 10195 & Oberprieler 4643 (B). prieler 5382 (B); Morocco, Monts des Beni-Snassen, road 5311 between Mechrâ-Homadi and Berkane, valley SE point 351 m ca. 2 Euphorbia medicaginea Boiss. km E Mechrâ-Homadi, Tetraclinis articulata woodland, limestone 2n = 16, CHN. Morocco, Rharb, road 2301 between Nador cliffs, dry oued, 200–250 m, 34°45′N, 2°47′W, 7 May 1993, Vogt and Kenitra, ca. 10 km N Benmansoun, dunes with Eucalyptus 11044 & Oberprieler 5492 (B). plantation, 20 m, 34°40′N, 6°23′W, 25 Apr 1993, Vogt 10917 & Oberprieler 5365 (B). *Kremeriella cordylocarpus (Coss. & Durieu) Maire 2n = 22, CHN. Morocco, Monts des Beni-Snassen, track from *Euphorbia resinifera Berg Beni Ammar to Taforalt via Talezzert, Oued Tlata-at-Talremt ca. 2n = ca. 112, CHN. Morocco, Middle Atlas, road S 508 be- 2.5 km S of the marabout “Sidi Boubker”, roadsides, limestone tween Demnate and Azilal, ca. 10 km W Azilal, stony slopes, 1,200 cliffs, 220 m, 34°48′N, 2°39′W, 7 May 1993, Vogt 11173 & Ober- m, 31°57′N, 6°41′W, 23 May 1993, Vogt 11940 (B). prieler 5621 (B). Euphorbia segetalis L. Matthiola parviflora (Schousb.) R. Br. 2n = 16, CHN. Morocco, Tangier Peninsula, road 8303 from 2n = 14, CHN. Morocco, Monts des Beni-Snassen, road 5311 Souk-Tleta-Taghramet to Ceuta (Sebta), limestone rocks above between Mechrâ-Homadi and Berkane, ca. 2 km as the crow flies Souk-Tleta-Taghramet and around quarry not far from the summit NE Mechrâ-Homadi, field margins, 380 m, 34°47′N, 2°45′W, 7 May of the pass, 450–500 m, 35°48′N, 5°27′W, 17 Apr 1993, Vogt 9878 1993, Vogt 11127 & Oberprieler 5575 (B). & Oberprieler 4326 (B).

*Muricaria prostrata (Desf.) Desv. Euphorbia terracina L. 2n = 26, CHN. Morocco, Eastern plateaus, road P 19 between 2n = 18, CHN. Morocco, Mediterranean coast NW Oujda, Bouarfa and Oujda, ca. 20 km S Aïn-Beni-Mathar, field margins, Ras-el-Ma, surroundings of lighthouse at Ras Kebdana, maritime roadsides, railroad track, 970 m, 33°44′N, 2°00′W, 3 May 1993, sands, 40 m, 35°09′N, 2°25′W, 5 May 1993, Vogt 10931 & Ober- Vogt 10685 & Oberprieler 5133 (B). prieler 5379 (B).

E5 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Mercurialis ambigua L. f. [≡ Mercurialis annua subsp. am- 2 May 1993, Vogt 10587a & Oberprieler 5035a (B), cultivated at the bigua (L. f.) Arcang.] Botanic Garden Berlin-Dahlem, 26 Sep 1994, Vogt & Oberprieler 2n = 48, CHN. Morocco, Monts des Beni-Snassen, Taforalt, s.n. (B); Morocco, Tell Atlas, Col de Jerada on road P 19 between N-facing slopes of Djebel Achouan ca. 1 km E Taforalt, stony slopes Aïn-Beni-Mathar and Oujda, roadsides ca. 4 km N of pass, 1,120 and limestone rocks, 850–900 m, 34°49′N, 2°24′W, 4 May 1993, Vogt m, 34°21′N, 2°03′W, 4 May 1993, Vogt 10722 & Oberprieler 5170 10753 & Oberprieler 5201 (B); Morocco, Monts des Beni-Snassen, (B); Morocco, Djebel Siroua, road P 32 between Tazenakht and track from Beni Ammar to Taforalt via Talezzert, Oued Tlata-at- Taliouine, ca. 5.3 km E Taliouine, roadsides, 1,320 m, 30°30′N, Talremt ca. 2.5 km S of the marabout “Sidi Boubker”, roadsides, 7°55′W, 13 May 1993, Vogt 11732 & Oberprieler 6180 (B). limestone cliffs, 220 m, 34°48′N, 2°39′W, 7 May 1993, Vogt 11178 & Oberprieler 5626 (B); Morocco, Monts des Beni-Snassen, track Erodium laciniatum (Cav.) Willd. subsp. laciniatum from Beni Ammar to Taforalt via Tanezzert, settlements around 2n = 20, CHN. Morocco, Mediterranean coast W of Saidia, Tanezzert between Sidi-Yahia-Ben-Ahmed and Tagliat, roadsides, surroundings of the mouth of Oued Moulouya ca. 10 km W Saidia, field margins, 450 m, 34°48′N, 2°36′W, 8 May 1993, Vogt 11300 & dunes ca. 1 km inland, 10 m, 35°07′N, 2°18′W, 5 May 1993, Vogt Oberprieler 5748 (B); Morocco, Monts des Beni-Snassen, Gorges 10875b & Oberprieler 5323b (B). du Zegzel between the villages of Trashroute and Moulay Ahmed, slopes of the gorge below and above road, riverbed, 450–550 m, Erodium laciniatum subsp. pulverulentum (Boiss.) Batt. 34°49′N, 2°22′W, 10 May 1993, Vogt 11550 & Oberprieler 5998 (B). 2n = 20, CHN. Morocco, Monts des Beni-Snassen, track from Beni Ammar to Taforalt via Tanezzert, mountain slopes and lime- FUMARIACEAE stone cliffs between the settlements Tagliat and Wawizaght E of Fumaria bastardii Boreau Tanezzert, 450–550 m, 34°48′N, 2°34′W, 8 May 1993, Vogt 11364 2n = 48, CHN. Morocco, Middle Atlas, Massif du Kandar S & Oberprieler 5812 (B). Sefrou, road P 20 between Sefrou and Boulmane, limestone cliffs near the street ca. 7,5 km S of Sefrou, 1,150 m, 33°45′N, 4°51′W, Erodium moschatum (L.) L’Hér. 28 Apr 1993, Vogt 10322 & Oberprieler 4770 (B). 2n = 20, CHN. Morocco, Tangier Peninsula, road P 32 between Tangier and Tetouan, around the junction with road 8302 GERANIACEAE to Melloussa, field margins, 70 m, 35°40′N, 5°41′W, 21 Apr 1993, Erodium chium (L.) Willd. Vogt 9992 & Oberprieler 4440 (B). 2n = 20, CHN. Morocco, Monts des Beni-Snassen, Taforalt, N-facing slopes of Djebel Achouan ca. 1 km E Taforalt, stony slopes Erodium salzmannii Delile and limestone rocks, 850–900 m, 34°49′N, 2°24′W, 4 May 1993, 2n = 20, CHN. Morocco, Rharb, road S 216 between Arba- Vogt 10843 & Oberprieler 5291 (B); Morocco, Mediterranean coast oua and Moulay Bousselham, ca. 3,4 km W of junction with road W of Saidia, surroundigs of the mouth of Oued Moulouya ca. 10 km to Lalla-Rhano and Ksar-el-Kebir, ungrazed field margin, 10 m, W Saidia, dunes ca. 1 km inland, 10 m, 35°7′N, 2°18′W, 5 May 34°51′N, 6°10′W, 24 Apr 1993, Vogt 10186 & Oberprieler 4634 1993, Vogt 10875a & Oberprieler 5323a (B); Morocco, Monts (B); Morocco, Rharb, Moulay Bousselham, lagune Merdja Zerga, des Beni-Snassen, track from Beni Ammar to Taforalt via Talez- moist meadows and marshes, 0–5 m, 34°53′N, 6°16′W, 25 Apr 1993, zert, Oued Tlata-at-Talremt ca. 2.5 km S of the marabout “Sidi Vogt 10206 & Oberprieler 4654 (B); Morocco, Anti-Atlas, Col du Boubker”, roadsides, limestone cliffs, 220 m, 34°48′N, 2°39′W, Kerdous, mountain slopes E of hotel at Col du Kerdous, rocks, 7 May 1993, Vogt 11168 & Oberprieler 5616 (B); Morocco, Monts Cupressus plantations, 1,200–1,250 m, 29°35′N, 9°27′W, 17 May des Beni-Snassen, track from Beni Ammar to Taforalt via Tane- 1993, Vogt 11865 & Oberprieler 6313 (B). zzert, mountain slopes and limestone cliffs between the settlements Tagliat and Wawizaght E of Tanezzert, 450–550 m, 34°48′N, Geranium purpureum Vill. 2°34′W, 8 May 1993, Vogt 11344 & Oberprieler 5792 (B); Morocco, 2n = 32, CHN. Morocco, Monts des Beni-Snassen, track from Monts des Beni-Snassen, Gorges du Zegzel between the villages Beni Ammar to Taforalt via Tanezzert, mountain slopes and lime- of Trashroute and Moulay Ahmed, slopes of the gorge below and stone cliffs between the settlements Tagliat and Wawizaght E of above road, riverbed, 450–550 m, 34°49′N, 2°22′W, 10 May 1993, Tanezzert, 450–550 m, 34°48′N, 2°34′W, 8 May 1993, Vogt 11363 Vogt 11576 & Oberprieler 6024 (B). & Oberprieler 5811 (B).

*Erodium crenatum Pomel Geranium rotundifolium L. 2n = 20, CHN. Morocco, Middle Atlas, Massif du Kandar S 2n = 26, CHN. Morocco, Monts des Beni-Snassen, track from Sefrou, road P 20 between Sefrou and Boulmane, limestone cliffs Beni Ammar to Taforalt via Tanezzert, mountain slopes and lime- near the street ca. 7.5 km S of Sefrou, 1,150 m, 33°45′N, 4°51′W, stone cliffs between the settlements Tagliat and Wawizaght E of 28 Apr 1993, Vogt 10303 & Oberprieler 4751 (B). Tanezzert, 450–550 m, 34°48′N, 2°34′W, 8 May 1993, Vogt 11345 & Oberprieler 5793 (B). Erodium glaucophyllum (L.) L’Hér. 2n = 20, CHN. Tunisia, Gouvernorat de Gafsa, Tunisie du Sud, LABIATAE road 201 between Redeyef and Tamerza, ca. 3 km NW Redeyef, *Rosmarinus eriocalyx var. pallescens (Maire) Upson & Jury road embankments, sand, 680 m, 9 May 1994, Vogt 12754 & Ober- 2n = 24, CHN. Morocco, Monts des Beni-Snassen, track be- prieler 7059 (B), cultivated at the Botanic Garden Berlin-Dahlem, tween Tanezzert and Taforalt, valley of Oued Tazzemoute, sur- 17 Aug 1995, Oberprieler & Vogt s.n. (B). roundings of the pass ca. 4.4 km W Taforalt, road embankments, maquis, 900–950 m, 34°38′N, 2°27′W, 9 May 1993, Vogt 11434 & *Erodium guttatum (Desf.) Willd. Oberprieler 5882 (B). 2n = 20, CHN. Morocco, Prov. Figuig, mountains W of Djebel Maiz near road P 32 between Bouarfa and Figuig, ca. 55 km WNW Salvia aegyptiaca L. Figuig, dry oued, stony slopes, and rocks, 1,200 m, 32°14′N, 1°44′W, 2n = 26, CHN. Morocco, Monts des Beni-Snassen, road 5311

E6 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

between Mechrâ-Homadi and Berkane, valley SE point 351 m ca. between Mechrâ-Homadi and Berkane, slopes along the road ca. 2 km E Mechrâ-Homadi, Tetraclinis articulata woodland, lime- 1 km E of the junction with road S 412 (Mechrâ-Homadi–El- stone cliffs, dry oued, 200–250 m, 34°45′N, 2°47′W, 7 May 1993, Aioun), Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, Vogt 11078 & Oberprieler 5526 (B). 6 May 1993, Vogt 10944 & Oberprieler 5392 (B); Morocco, Monts des Beni-Snassen, road 5311 between Mechrâ-Homadi and Ber- Salvia verbenaca L. kane, valley SE point 351 m ca. 2 km E Mechrâ-Homadi, Tetracli- 2n = 24, CHN. Morocco, Prov. Figuig, road P 19 between nis articulata woodland, limestone cliffs, dry oued, 200–250 m, Bouarfa and Oujda, ca. 5 km N Tendrara, roadsides in a Pinus 34°45′N, 2°47′W, 7 May 1993, Vogt 11050 & Oberprieler 5498 (B). plantation, 1,300 m, 33°08′N, 1°59′W, 2 May 1993, Vogt 10635 & Oberprieler 5083 (B). Plantago albicans L. 2n = 10, CHN. Morocco, Monts des Beni-Snassen, road 5311 LEGUMINOSAE between Mechrâ-Homadi and Berkane, valley SE point 351 m ca. Astragalus epiglottis L. 2 km E Mechrâ-Homadi, Tetraclinis articulata woodland, lime- 2n = 16, CHN. Morocco, Monts des Beni-Snassen, track from stone cliffs, dry oued, 200–250 m, 34°45′N, 2°47′W, 7 May 1993, Beni Ammar to Taforalt via Talezzert, mountain ridge between Vogt 11065 & Oberprieler 5513 (B). Djebel Astar-ach-Chaib and Oued Tlata-at-Talremt ca. 3.5 km SE of the marabout “Sidi Boubker” in Beni Ammar, pastures and POLYGONACEAE Tetraclinis woodland, 340–370 m, 34°48′N, 2°38′W, 8 May 1993, Emex spinosa (L.) Campd. Vogt 11199 & Oberprieler 5647 (B). 2n = 20, CHN. Morocco, Prov. Agadir, Oued Massa, road P 30 between Agadir and Tiznit, ca. 25 km N Tiznit, river banks Tripodion tetraphyllum (L.) Fourr. [≡ Anthyllis tetraphylla L.] and roadsides, 29°54′N, 9°38′W, 19 May 1993, Vogt 11917 & Ober- 2n = 16, CHN. Morocco, Taliouine NE 15 km, heavily grazed prieler 6365 (B). Argania plantation, Trockenfeldbau, limestone and fanglomerates, 980 m, 4.1992, Gruber (seed sample acquired from Botanic Garden Rumex vesicarius L. Salzburg), cultivated at the Botanic Garden Berlin-Dahlem, 7 Jun 2n = 18, CHN. Morocco, Great Atlas, Gorges du Ziz between 1993, Vogt & Oberprieler s.n. (B). Rich and Er-Rachidia, surroundings of Tunnel du Legionaires 21.1 km S Rich, 1,280 m, 32°10′N, 4°23′W, 29 Apr 1993, Vogt LILIACEAE 10383 & Oberprieler 4831 (B); Morocco, Prov. Figuig, road P 32 Asphodelus refractus Boiss. between Bouarfa and Figuig, north-facing slopes of Djebel Grouz 2n = 52, CHN. Morocco, Prov. d’Er-Rachidia, road P 32 be- ca. 23 km W Figuig, mountains ca. 6 km S of road P 32, rocks and tween Er-Rachidia and Boudnib, ca. 19.2 km E of junction to Er- stony slopes, 1,100–1,300 m, 32°08′N, 1°24′W, 1 May 1993, Vogt foud, dry oued with Ziziphus, 1,030 m, 31°55′N, 4°08′W, 30 Apr 10563 & Oberprieler 5011 (B); Morocco, Monts des Beni-Snassen, 1993, Vogt 10458 & Oberprieler 4906 (B). road 5311 between Mechrâ-Homadi and Berkane, valley SE point 351 m ca. 2 km E Mechrâ-Homadi, Tetraclinis articulata wood- Asphodelus tenuifolius Cav. land, limestone cliffs, dry Oued, 200–250 m, 34°45′N, 2°47′W, 2n = 28, CHN. Morocco, Anti-Atlas, road S 509 between Aït-Ba- 7 May 1993, Vogt 11029 & Oberprieler 5477 (B). hia and Tafraoute, ca. 15.7 km SE Aït-Bahia, roadside, rocks, 900 m, 29°57′N, 9°01′W, 14 May 1993, Vogt 11743 & Oberprieler 6191 (B). PRIMULACEAE Anagallis arvensis L. LINACEAE 2n = 20, CHN. Morocco, Monts des Beni-Snassen, road 5311 Linum strictum L. between Mechrâ-Homadi and Berkane, slopes along the road ca. 2n = 18, CHN. Morocco, Monts des Beni-Snassen, road 5311 1 km E of the junction with road S 412 (Mechrâ-Homadi–El- between Mechrâ-Homadi and Berkane, slopes along the road ca. Aioun), Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, 1 km E of the junction with road S 412 (Mechrâ-Homadi–El- 6 May 1993, Vogt 10941 & Oberprieler 5389 (B). Aioun), Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, 6 May 1993, Vogt 10960 & Oberprieler 5408 (B). 2n = 40, CHN. Morocco, Rharb, road S 216 between Arba- oua and Moulay Bousselham, ca. 3,4 km W of junction with road MALVACEAE to Lalla-Rhano and Ksar-el-Kebir, ungrazed field margin, 10 m, *Malva aegyptia L. 34°51′N, 6°10′W, 24 Apr 1993, Vogt 10175 & Oberprieler 4623 (B); 2n = 24, CHN. Morocco, Eastern plateaus, road P 19 between Morocco, Monts des Beni-Snassen, Gorges du Zegzel between the Bouarfa and Oujda, ca. 20 km S Aïn-Beni-Mathar, field margins, villages of Moulay Ahmed and Takerboust, riverbed and road- roadsides, railroad track, 970 m, 33°44′N, 2°00′W, 3 May 1993, sides, 270–300 m, 34°50′N, 2°22′W, 10 May 1993, Vogt 11664 & Vogt 10673 & Oberprieler 5121 (B). Oberprieler 6112 (B).

PAPAVERACEAE Asterolinum linum-stellatum (L.) Duby Papaver rhoeas L. 2n = 20, CHN. Morocco, Monts des Beni-Snassen, Taforalt, 2n = 14, CHN. Morocco, Monts des Beni-Snassen, road 5311 N-facing slopes of Djebel Achouan ca. 1 km E Taforalt, stony slopes between Mechrâ-Homadi and Berkane, ca. 2 km as the crow flies and limestone rocks, 850–900 m, 34°49′N, 2°24′W, 4 May 1993, NE Mechrâ-Homadi, field margins, 380 m, 34°47′N, 2°45′W, 7 Vogt 10851 & Oberprieler 5299 (B). May 1993, Vogt 11102 & Oberprieler 5550 (B). RANUNCULACEAE PLANTAGINACEAE Ranunculus paludosus Poir. Plantago afra L. 2n = 16, CHN. Morocco, Tangier Peninsula, road 8302 be- 2n = 12, CHN. Morocco, Monts des Beni-Snassen, road 5311 tween Ksar-es-Seghir and route P 38 (Tangier–Tetouan), ca. 1 km N

E7 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Melloussa, field margins and stony embankments, 180 m, 35°43′N, *Bupleurum lateriflorum Coss. 5°38′W, 20 Apr 1993, Vogt 9966 & Oberprieler 4414 (B). 2n = 28, CHN. Morocco, Great Atlas, Tizi-n-Test – Pass, 1991, APE 1050, Gardner, Jury & Ait-Lafkih (seed sample acquired from Ranunculus spicatus subsp. blepharicarpos (Boiss.) Grau Botanic Garden Reading), cultivated at the Botanic Garden Berlin- 2n = 16, CHN. Morocco, Monts des Beni-Snassen, track be- Dahlem, 14 Apr 1994, Vogt & Oberprieler s.n. (B), 26 Sep 1994, tween Tanezzert and Taforalt, immediate E Oulad Yahya, road- Vogt & Oberprieler s.n. (B). sides, field margins, rocks, 760 m, 34°38′N, 2°29′W, 8 May 1993, Vogt 11398 & Oberprieler 5846 (B), cultivated at the Botanic Gar- Daucus carota L. subsp. carota den Berlin-Dahlem, 21 Mar 1996, Cubr 34703 (B). 2n = 18, CHN. Morocco, Rharb, Moulay Bousselham, lagune Merdja Zerga, moist meadows and marshes, 0–5 m, 34°53′N, RUBIACEAE 6°16′W, 25 Apr 1993, Vogt 10203 & Oberprieler 4651 (B). Galium verrucosum Huds. 2n = 22, CHN. Morocco, Plaine du Sais, road 3360 between Pseudorlaya pumila (L.) Grande Aïn-Cheggag and road P 24 (Fes–Ifrane), roadsides and fieldmar- 2n = 16, CHN. Morocco, Mediterranean coast W of Saidia, gins 2.6 km SW of junction with P 24, 620 m, 33°59′N, 5°00′W, 27 surroundings of the mouth of Oued Moulouya ca. 10 km W Saidia, Apr 1993, Vogt 10282 & Oberprieler 4730 (B); Morocco, Monts des dunes ca. 1 km inland, 10 m, 35°07′N, 2°18′W, 5 May 1993, Vogt Beni-Snassen, road 5311 between Mechrâ-Homadi and Berkane, 10884 & Oberprieler 5332 (B). valley SE point 351 m ca. 2 km E Mechrâ-Homadi, Tetraclinis articulata woodland, limestone cliffs, dry oued, 200–250 m, 34°45′N, VALERIANACEAE 2°47′W, 7 May 1993, Vogt 11064 & Oberprieler 5512 (B). Centranthus calcitrapae (L.) Dufr. 2n = 32, CHN. Morocco, Monts des Beni-Snassen, track from Valantia muralis L. Beni Ammar to Taforalt via Tanezzert, mountain slopes and lime- 2n = 18, CHN. Morocco, Tangier Peninsula, road 8303 from stone cliffs between the settlements Tagliat and Wawizaght E of Souk-Tleta-Taghramet to Ceuta (Sebta), limestone rocks above Tanezzert, 450–550 m, 34°48′N, 2°34′W, 8 May 1993, Vogt 11320 Souk-Tleta-Taghramet and around quarry not far from the summit & Oberprieler 5768 (B); Morocco, Middle Atlas, Aïn Leuh, road of the pass, 450–500 m, 35°48′N, 5°27′W, 17 Apr 1993, Vogt 9882 S 303 between Aïn Leuh and Source de l’Oum-er-Rbia, immedi- & Oberprieler 4330 (B). ate S of Aïn Leuh, 1,510 m, 33°17′N, 5°21′W, 28 May 1993, Vogt 11995 (B). SAPOTACEAE *Argania spinosa (L.) Skeels Fedia cornucopiae (L.) Gaertn. 2n = 20, CHN. Morocco, Simou S 4 km, Callitris-Argania- 2n = 32, CHN. Morocco, Monts des Beni-Snassen, Gorges Juniperus phoenicea woodland, heavily grazed, marl lime, 430 m, du Zegzel between the villages of Trashroute and Moulay Ahmed, 3.1992, leg. Gruber, Fürnkranz & Strobl s.n. (seed sample acquired slopes of the gorge below and above road, riverbed, 450–550 from Botanic Garden Salzburg), cultivated at the Botanic Garden m, 34°49′N, 2°22′W, 10 May 1993, Vogt 11541 & Oberprieler Berlin-Dahlem, 14 Apr 1994, Vogt & Oberprieler s.n. (B), 28 May 5989 (B). 2003, Cubr 40606 (B). Valerianella discoidea (L.) Loisel. SCROPHULARIACEAE 2n = 14, CHN. Morocco, Monts des Beni-Snassen, road 5311 *Misopates microcarpum (Pomel) D.A. Sutton [≡ Antirrhinum between Mechrâ-Homadi and Berkane, valley SE point 351 m ca. microcarpum Pomel] 2 km E Mechrâ-Homadi, Tetraclinis articulata woodland, lime- 2n = 16, CHN. Morocco, Monts des Beni-Snassen, road 5311 stone cliffs, dry oued, 200–250 m, 34°45′N, 2°47′W, 7 May 1993, between Mechrâ-Homadi and Berkane, slopes along the road ca. Vogt 11051 & Oberprieler 5499 (B). 1 km E of the junction with road S 412 (Mechrâ-Homadi–El- Aioun), Tetraclinis articulata woodland, 200 m, 34°43′N, 2°48′W, Acknowledgements 6 May 1993, Vogt 10958 & Oberprieler 5406 (B); Morocco, Monts We would like to express our sincere thanks to Mrs M. Lüchow des Beni-Snassen, track from Beni Ammar to Taforalt via Talez- and Mrs P. Schirarend for their excellent technical assistance. zert, mountain ridge between Djebel Astar-ach-Chaib and Oued Thanks for the determination of our collections of Erigeron are Tlata-at-Talremt ca. 3.5 km SE of the marabout “Sidi Boubker” due to Dr. K.-P. Buttler, Frankfurt am Main. in Beni Ammar, pastures and Tetraclins woodland, 340–370 m, 34°48′N, 2°38′W, 8 May 1993, Vogt 11242 & Oberprieler 5690 (B). Literature cited Hellwig, F., Oberprieler, C., Vogt, R. & Wagenitz, G. 1994. Scrophularia laevigata Vahl Chromosome numbers of North African phanerogams. 2n = 58, CHN. Morocco, Monts des Beni Snassen, Gorges III. Some counts in Centaurea (Compositae, Cardueae). du Oued Zegzel between Djebel Israne and Djebel Achaoun E Willdenowia 24: 249–254. Taforalt, S-facing mountain slopes of Djebel Israne near the “Grotte Kilian, N., Oberprieler, C. & Vogt, R. 1995. Chromosome des Pigeons”, 690 m, 34°50′N, 2°24′W, 9 May 1993, Vogt 11522 & numbers of North African phanerogams. V. Some counts Oberprieler 5970 (B). in Launaea (Compositae, Lactuceae). Willdenowia 25: 273–281. UMBELLIFERAE Oberprieler, C. & Vogt, R. 1993. Chromosome numbers of Bifora testiculata (L.) Roth North African phanerogams. II. Willdenowia 23: 211–238. 2n = 22, CHN. Morocco, Plaine du Sais, road 3360 between Oberprieler, C. & Vogt, R. 1996. Chromosome numbers of Aïn-Cheggag and road P 24 (Fes–Ifrane), roadsides and fieldmar- North African phanerogams. VI. Some counts in Legumi- gins 2.6 km SW of junction with P 24, 620 m, 33°59′N, 5°00′W, nosae. Willdenowia 25: 669–680. 27 Apr 1993, Vogt 10267 & Oberprieler 4715 (B). Scholz, H., Oberprieler, C. & Vogt, R. 1998. Chromosome

E8 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

numbers of North African phanerogams. VII. Some notes *Heracleum nubigenum C.B. Clarke on North African Gramineae. Lagascalia 20: 265–275. 2n = 44, CHN. India, Sikkim, E Himalaya, valley of upper Vogt, R. & Oberprieler, C. 1993. Chromosome numbers of Teesta River, near Lachung, 27°31′N, 88°44′E, 2,500 m, 15 Oct North African phanerogams. I. Fl. Medit. 3: 187–210. 2003, M.G. Pimenov & E.V. Kljuykov 58 (MW). Vogt, R. & Oberprieler, C. 1994. Chromosome numbers of North African phanerogams. IV. Candollea 49: 549–570. *Hymenidium brunonis (DC.) Lindl. Vogt, R. & Oberprieler, C. 2008. Chromosome numbers of n = 11, CHN. India, Himachal Pradesh, Shimla distr., northern North African phanerogams. VIII. More counts in Com- slope of Choordar Mt. (S of Darain village), 30°52′N, 77°30′E, positae. Willdenowia 38: 497–519. 3,640 m, 13 Sep 2000, M.G. Pimenov & E.V. Kljuykov 205 (MW).

*Hymenidium decurrens (Franch.) Pimenov & Kljuykov Julia V. Shner, Michael G. Pimenov & Eugene V. Kljuykov 2n = 18, CHN. China, Yunnan, Tali Co, Diancang Shan Mts., eastern slope, above Yangbi, rich mixed forests, 25°42′N, 100°06′E, Botanical Garden of Moscow State University, Moscow, 119899, 3,400 m, 04 Oct 1998, M.G. Pimenov, E.V. Kljuykov, Hu ZhiHao Russia. [email protected] (author for correspondence) & Liu QiXin 560 (MW).

* First chromosome count for the species. **Kailashia robusta Pimenov & Kljuykov ** First chromosome count for the genus. 2n = 22, CHN. India, Uttaranchal, Uttarkashi distr., W Hi- *** New number for the species. malaya, valley of upper Bhagirathi River, near Bhojbasa, 30°54′N, 79°03′E, 3,850 m (classical locality), 06 Oct 2003, M.G. Pimenov This study was supported by grants 04-04-48146 and 07-04- & E.V. Kljuykov 16 (MW). 00270 from the Russian Foundation for Basic Research (RFBR). **Kedarnatha meifolia Pimenov & Kljuykov APIACEAE 2n = 22, CHN. India, Uttaranchal, Chamoli distr., Main Hi- Acronema tenerum (DC.) Edgew. malayan Range, basin of Alaknanda River, valley of Hem Ganga 2n = 18, CHN. India, Uttaranchal, Chamoli distr., Main Hi- near Ghangaria, 30°37′N, 79°34′E, 3,300 m (classical locality), 18 malayan Range, basin of Alaknanda River, valley of Hem Ganga Sep 2000. M.G. Pimenov & E.V. Kljuykov 236 (MW). near Ghangaria, 30°37′N, 79°34′E, 3,300 m, 18 Sep 2000, M.G. Pimenov & E.V. Kljuykov 235 (MW); India, Sikkim, E Himalaya, *Ligusticopsis coniifolia (DC.) Pimenov & Kljuykov valley of upper Teesta River near Yumthang, 27°33′N, 88°43′E, 2n = 22, CHN. India, Uttaranchal state, Chamoli distr., 3,400 m, 16 Oct 2003, M.G. Pimenov & E.V. Kljuykov 61 (MW). W Himalaya, valley of upper Mandakini River, between Gaurikund and Kedarnath, near Rampara, 30°41′N, 79°03′E, 2,800 m, 07 Oct Angelica pubescens Maxim. 2003, M.G. Pimenov & E.V. Kljuykov 25 (MW). 2n = 22, CHN. Japan, near Kyoto, Kurama-dera, 35°11′N, 135°38′E, 11 Aug 2004, Pimenov s.n. (MW). Ligusticopsis wallichiana (DC.) Pimenov & Kljuykov 2n = 22, CHN. India, Sikkim, E Himalaya, valley of upper Bupleurum candollei Wall. ex DC. Teesta River, near Lachung, 27°31′N, 88°44′E, 2,500 m, 15 Oct 2n = 16, CHN. India, Sikkim, E Himalaya, valley of upper 2003, M.G. Pimenov & E.V. Kljuykov 56 (MW). Teesta River, near Lachung, 27°31′N, 88°44′E, 2,500 m, 03 Oct 2003, M.G. Pimenov & E.V. Kljuykov 57 (MW). **Melanosciadium bipinnatum (Shan RenHwa & Pu FaTing) Pimenov & Kljuykov ***Bupleurum gracillimum Klotzsch 2n = 22, CHN. China, Sichuan, route Hongyuan-Barkam, val- 2n = 14, CHN. India, Himachal Pradesh, riv. Tsarap between ley of Suome River, near Maerkang, 31°55′N, 102°39′E, 2,800 m, Kumjung and Serai, 32°49′N, 77°26′E, 4,620 m, 29 Aug 2000, M.G. 16 Sep 1998, M.G. Pimenov, E.V. Kljuykov, Hu ZhiHao & Liu QiXin Pimenov & E.V. Kljuykov 66 (MW), 190 (MW).

Bupleurum marginatum DC. Osmorhiza aristata (Thunb.) Rydb. 2n = 16, CHN. India, Uttaranchal, Uttarkashi distr., W Hi- 2n = 22, CHN. India, Uttaranchal, between Bhyundar and malaya, foothills, valley of Bhagirathi River, near Sukhi, 30°53′N, Ghangaria, 18 Sep 2000, M.G. Pimenov & E.V. Kljuykov 228 (MW). 78°40′E, 2,040 m, 01 Oct 2003, M.G. Pimenov & E.V. Kljuykov 7 (MW); India, Uttaranchal, Uttarkashi distr., W Himalaya, val- Pimpinella acuminata (Edgew.) C.B. Clarke ley of upper Bhagirathi River, between Gangotri and Choibasa, 2n = 20, CHN. India, Himachal Pradesh, between Naggar and 31°00′N, 78°58′E, 3,060 m, 02 Oct 2003, M.G. Pimenov & E.V. Banjar, forest margin, 31°33′N, 77°20′E, 1,910 m, 11 Sep 2000, Kljuykov 10 (MW). M.G. Pimenov & E.V. Kljuykov 167 (MW).

Bupleurum mucronatum Wight & Arn. *Pimpinella katrajensis R.S. Rao & Hemadri n = 7, CHN. S India, Tamil Nadu, distr. Nilgiris, Coonor, n = 9, CHN. S India, Maharashtra, Satara distr., between Pan- Dolphin’s Nose, 11°21′N, 76°51′E, 1,800–1,900 m, 09 Nov 2006, chgani and Wai, 17°56′N, 73°50′E, 1,150–1,275 m, 02 Nov 2006, M.G. Pimenov & E.V. Kljuykov 58 (MW). M.G. Pimenov & E.V. Kljuykov 33 (MW).

Heracleum hemsleyanum Diels Pimpinella wallichiana (Miq. ex Hohen.) Gandhi 2n = 22, CHN. China, Sichuan, between Yajiang and Litang, n = 10, CHN. S India, Karnataka, distr. Kodagu, near Madik- 90 km road station, 29°59′N, 100°52′E, 4,100 m, 23 Sep 1998, eri, 12°24′N, 75°44′E, 1,190–1,200 m, 05 Nov 2006, M.G. Pimenov M.G. Pimenov, E.V. Kljuykov, Hu ZhiHao & Liu QiXin 327(MW). & E.V. Kljuykov 38 (MW).

E9 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

*Semenovia thomsonii (C.B. Clarke) Manden. 2n = 18, CHN. Jordan, Azraq oasis 90 km E of Amman, Frei- 2n = 22, CHN. India, Himachal Pradesh, Spiti, inter Hansa tag 30.120 (KAS). and Kiato ca. 9 km E of Lasar, 32°26′N, 77°44′E, 4,100 m, 09 Sep 2000, M.G. Pimenov & E.V. Kljuykov 150 (MW). Suaeda corniculata (C.A. Mey.) Bunge s.str. 2n = 54, CHN. Russia, Samara obl., 20 km N of Tolyatti, *Seseli nudum (Lindl.) Pimenov & Kljuykov Nizhn. Sancheleevo, Golub 761 (NS); Russia, Buryat Republic, 2n = 22, CHN. India, Uttaranchal, Uttarkashi distr., W Hi- Gusinoozersk distr., Selenduma, Lomonosova 434 (NS); Russia, malaya, valley of upper Bhagirathi River, between Gangotri and Buryat Republic, Gusinoozersk distr., Sulfat’noe lake, 18 Sep 2002, Choibasa, 31°00′N, 78°58′E, 3,060 m, 02 Oct 2003, M.G. Pimenov Osipov s.n. (NS); Russia, Chita obl., Onon distr., Ulan-Nur lake, & E.V. Kljuykov 8 (MW). Lomonosova 490 (NS).

*Sinocarum wolffianum (Fedde ex H. Wolff) P.K. Mukh. & Suaeda kulundensis Lomon. & Freitag Constance 2n = 72, CHN. Russia, Samara obl., 20 km N of Tolyatti, 2n = 22, CHN. India, Sikkim, E Himalaya, valley of upper Nizhn. Sancheleevo, Golub 762 (NS); Russia, Kurgan obl., Teesta River, near Yumthang, 27°33′N, 88°43′E, 3,400 m, 16 Oct Vargashi, Lomonosova 740 (NS); Russia, Kurgan obl., Makushino, 2003, M.G. Pimenov & E.V. Kljuykov 62 (MW). Lomonosova 742 (NS); Russia, Kurgan obl., Medvezhie lake, Lomonosova 755 (NS); Russia, Novosibirsk obl., Karasuk distr., ***Tetrataenium aquilegifolium (C.B. Clarke) Manden. Troitskoe, Lomonosova 79, 80 (NS); Russia, Novosibirsk obl., 2n = 44, CHN. S India, Maharashtra, Raigad distr., Matheran, Ust’-Tarsk distr., Pokrovka, Krasnoborov 123 (NS); Russia, No- Nigam’s point. 29 Oct 2006, M.G. Pimenov & E.V. Kljuykov 2 vosibirsk obl., Kuibyshev distr., Aul-Bergul, Krasnoborov & Arte- (MW); S India, Maharashtra, Satara distr., Mahabaleshwar hill mov 4 (NS); Russia, Altaisk. Krai, Shipunovo distr., 196 km from station, Cannought Peak. 17°56′N, 73°40′E, 1,440 m, 01 Nov 2006, Barnaul to Rubtsovsk, Lomonosova 85 (NS); Russia, Altaisk. Krai, M.G. Pimenov & E.V. Kljuykov 31 (MW). Shipunovo distr., Korobeinikovo, Zolotov 16 (NS); Russia, Altaisk. Krai, Volchikha distr., Ust’-Kormikha, 17 Sep 1999, Artemov & ***Tetrataenium ligusticifolium (Wight) Manden. ( = T. canal. s.n. (NS); Russia, Altaisk. Krai, Rubtsovsk distr., 17 km W of dolleanum (Wight & Arn.) Manden.) Veseloyarsk, Krasnoborov 103 (NS); Russia, Altaisk. Krai, Lok- n = 22, CHN. S India, Tamil Nadu, distr. Dingigul, Kodai- tevsk distr., Noven’koe, Krasnoborov 101b (NS); Russia, Altaisk. kanal, Observatory Rd. 10°14′N, 77°28′E, 1,800 m, 11 Nov 2006, Krai, Kulunda distr., Mirabilit, Krasnoborov 135 (NS); Russia, M.G. Pimenov & E.V. Kljuykov 89 (MW); S India, Kerala, Idukki Altaisk. Krai, Kulunda distr., Kirei, Krasnoborov & Shaulo 133 distr., 6 km from Munnar, 10°03′N, 77°03′E, 1,350–1,380 m, 19 (NS); Russia, Altaisk. Krai, Burla distr., Novopeschanoe, Artemov Nov 2006, M.G. Pimenov & E.V. Kljuykov 93 (MW). & al. 303 (NS); Russia, Altaisk. Krai, Klyuchi distr., Petukhi, Ko- rolyuk 241 (NS); Kazakhstan, Taldy-Kurgan obl., Motai, Lomono- ***Tetrataenium rigens (DC.) Manden. sova 51a (NS); Kazakhstan, East-Kazakhstan obl., Zharma distr., 2n = 22, CHN. S India, Karnataka, Kodagu distr., near Kakk- Salkintobe, Lomonosova 205 (NS); Kazakhstan, East-Kazakhstan abe vill., Honey Valley Estate, 12°13′N, 75°39′E, 910–930 m, 06 obl., Tarbagatai distr., Belaya Shkola, Lomonosova 194b (NS). Nov 2006, M.G. Pimenov & E.V. Kljuykov 37 (MW). 2n = 90, CHN. Russia, Novosibirsk obl., Kochki, Lomonosova *Tongoloa gracilis H. Wolff 386 (NS); Russia, Novosibirsk. obl., Karasuk distr., Troitskoe, Lo- 2n = 22, CHN. China, Sichuan, Kanding Co, near Tongolo monosova 69, 79b (NS); Russia, Novosibirsk obl., Karasuk distr., (Dongolou), 30°03′N, 101°29′E, 3,700 m, 21 Sep 1998, M.G. Pi- Khoroshee lake, 22 Sep 1999, Artemov & al. s.n. (NS); Russia, No- menov, E.V. Kljuykov, Hu ZhiHao & Liu QiXin 299 (MW). vosibirsk obl., Kyshtovka distr., Vorob’evo, Krasnoborov 107 (NS); Russia, Altaisk. krai, Shipunovo distr., 196 km from Barnaul to *Trachyspermum stictocarpum (C.B. Clarke) Rubtsovsk, Lomonosova 85b (NS); Russia, Altaisk. krai, Mikhailov H. Wolff distr., Malinovoe lake, Lomonosova 25 (NS); Russia, Altaisk. Krai, n = 11, CHN. S India, Maharashtra, Raigad distr., Matheran Uglovskoe distr., Simonovo, 2 Sep. 1999, Krasnoborov s.n. (NS); hill station, road to Garbut Point, 18°59′N, 73°17′E, 790–800 m, Russia, Altaisk. Krai, Kulunda distr., Shimolino, 31 Aug 1988, 30 Oct 2006, M.G. Pimenov & E.V. Kljuykov 14 (MW). Lomonosova s.n. (NS); Russia, Altaisk. Krai, Blagoveshchenka distr., Kuchukskoe lake, 28 Sep 2002, Grebenyuk s.n. (NS); Russia, Vanasushava pedata (Wight) P.K. Mukh. & Constance Altaisk. Krai, Pospelikha distr., Bol’shevka, Krasnoborov 86 (NS); n = 23, CHN. S India, Tamil Nadu, distr. Dingigul, between Russia, Altaisk. Krai, Khabary distr., Serp & Molot, Krasnoborov Kodaikanal and Pancarpuram, 10°14′N, 77°28′E, 1,800–1,820 m, 151 (NS); Russia, Altaisk. Krai, Aley, Krasnoborov & Shaulo 80 11 Nov 2006, M.G. Pimenov & E.V. Kljuykov 72 (MW). (NS); Kazakhstan, Taldy-Kurgan obl., Motai, Lomonosova 51b (NS).

Maria Lomonosova1, Helmut Freitag2 S. maritima (L.) Dumort. s.l. 2n = 36, CHN. Italy, Venice prov., 4 km N of Venice, Fre- 1 Central Siberian Botanical Garden, Siberian Branch of the itag 34.1096–8 (KAS); Italy, Grosseto prov., Laguna di Orbetello, Russian Academy of Science, Novosibirsk 630090, Russia. embankment Orbetello to Peninsula Mte. Argentario, Freitag [email protected] (author for correspondence) 34.1031,3 (KAS); Italy, Puglia prov., Gargano peninsula, Vieste 2 Plant Taxonomy and Morphology, Institute of Biology, port, Freitag 34.1007 (KAS); Italy, Puglia prov., 10 km S of Man- University of Kassel, 34132 Kassel, Germany. fredonia, near Mass. Piccardi, Freitag 34.1011,2, 34.1021 ; Germany, Saxony-Anhalt, 10 km W of Halle Teutschenthal, Oct 2006, Prinz CHENOPODIACEAE s.n. (KAS); Germany, Lower Saxony, Aurich distr., “Leybucht” in Suaeda aegyptiaca (Hasselq.) Zohary Western part of Ostfriesland, Oct 2006, Prinz s.n. (KAS).

E10 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

S. patagonica Speg. This study was supported by Grants nos. 04-04-49750 and 2n = 54, CHN. Argentina, Patagonia, Prov. Chubut, Peninsula 07-04-00610 (to N.S. Probatova), from Russian Fund for Basic Valdés, 9 Feb 2003, Schröder s.n. (KAS). Research (RFBR).

S. salsa (L.) Pall. ALLIACEAE 2n = 36, CHN. Ukraine, Crimea, S coast, 4 km N of Koktebel’, Allium anisopodium Ledeb. Barakol’ lake, Freitag 33.219 (KAS); Ukraine, E. Crimea, 25 km 2n = 16, CHN. Russia, Far East, Primorskii Krai, nature re- SSW of Kerch, Tobechic’ke lake, Freitag 33.252 (KAS); Ukraine, serve “Khankaiskii”, section Sosnovyi, Kazachii Bay, meadow- W. Crimea, 7 km W of Saki, Freitag 33.277 (KAS); Romania, steppe community, 19 May 2004, Barkalov 9221 (VLA). Black Sea coast, S of Danube delta, Chituk sandbank between the sea and Sinoe lake, Oct 2003, Fagaras s.n. (KAS); Romania, Allium condensatum Turcz. Transylvania, Turda, 27 Sep 2003, Miele s.n. (KAS). 2n = 16, CHN. Russia, Far East, Primorskii Krai, Ussuriiskii Raion, nature reserve “Ussuriiskii”, Zmejinaya Mt., on rocks, 14 Suaeda sibirica Lomon. & Freitag, May 2004, Barkalov 9174 (VLA). 2n = 72, CHN. Russia, Khakass Republic, Shira distr., Tus lake, Lomonosova 601 (NS); Russia, Khakass Republic, Beya distr., Allium macrostemon Bunge Gor’koe lake, Lomonosova 606, 626 (NS); Russia, Khakass Repub- 2n = 32, CHN. Russia, Far East, Primorskii Krai, Muravëv- lic, Ust’-Abakan distr., Krasnoe lake, Lomonosova, 609, 616, 618, Amurskii Peninsula, Ussuriiskii Gulf, Muraviinaya Bay, the 621 (NS); Russia, Khakass Republic, Ust’-Abakan distr., 20 km valley of Muraviinyi spring, near the nature monument “Cher- from Abakan to Askiz, Lomonosova 610, 612 (NS); Russia, Khakass nopikhtarnik”, 21 Oct 2004, Probatova & Seledets 9608 (VLA). Republic, Ust’-Abakan distr., Kamyzyak steppe, Ulug-Khol’ lake, Lomonosova 638, 643 (NS); Russia, Tuva Republic, Tes-Khem Allium maximowiczii Regel distr., O-Shynaa, Lomonosova 59 (NS); Russia, Tuva Republic, Tes- 2n = 16, CHN. Russia, Far East, Amurskaya oblast’, Arkha- Khem distr., Duz-Khol’ lake, 17 km NW of Erzin, Lomonosova 366 rinskii Raion, 8 km SE of Arkhara railway station, the bank of (NS); Russia, Tuva Republic, Tandin distr., Cheder lake, Lomono- Arkhara River, Salix community, 1 Jul 2008, Kudrin 11216 (VLA). sova 46 (NS); Russia, Tuva Republic, Tandin distr., Svatikovo lake, Lomonosova 335, 336, 341 (NS); Russia, Tuva Republic, Piy-Khem Allium ochotense Prokh. distr., salt lake E of Arzhaan, Lomonosova 373 (NS); Russia, Tuva 2n = 32, CHN. Russia, Far East, Kamchatskii Krai, Kam- Republic, Piy-Khem distr., near road Turan–Arzhaan, Lomonosova chatka Peninsula, ca. 20 km NE of Malki village, middle course 379 (NS); Russia, Irkutsk obl., Ust’-Ordynskoye distr., Lomonosova of Poperechnaya R., Betula ermanii forest, 28 Jul 1997, Barkalov 285, 286, 287, 299 (NS); Russia, Irkutsk obl.,Usole-Sibirskoe, Lo- 7369 (VLA); Russia, Far East, Khabarovskii Krai, Komsomol’skii monosova 284 (NS); Russia, Buryat Republic, Gusinoozersk distr., Raion, near the abandoned settlement Machtovyi, Machtovaya Sulfat’noe lake, Lomonosova 431, 28 Sep 2002, Osipov s.n. (NS); River, valley forest, 5 Aug 2005, Barkalov 9907 (VLA). Russia, Buryat Republic, Gusinoozersk distr., Borgoi steppe, Selen- duma, Lomonosova 434 (NS); Russia, Buryat Republic, Ulan-Ude Allium prokhanovii Vved. distr., Orongoi, Lomonosova 426 (NS); Russia, Buryat Republic, 2n = 16, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii Dzhida distr., Beloozersk, Lomonosova 438 (NS); Russia, Buryat Raion, 2.5 km SE of Kremovo village, slope of a hill, 8 Jul 2008, Republic, Dzhida distr., Nizhnee Beloe lake, Lomonosova 440 (NS); Lapenko 11095 (VLA). Russia, Buryat Republic, Barguzin distr., Suvo, Lomonosova 445 (NS); Russia, Buryat Republic, Barguzin distr., Uro, Lomonosova Allium senescens L. 449 (NS); Russia, Chita obl., Onon distr., Gashkoi lake, Lomono- 2n = 16, CHN. Russia, Far East, Primorskii Krai, west coast sova 477 (NS); Russia, Chita obl., Onon distr., Bulun-Sagan lake, of Khanka Lake, nature reserve “Khankaiskii”, section Sos- Lomonosova 482 (NS); Russia, Chita obl., Onon distr., Barun-Torei novyi, Przheval’skii Peninsula, grassland with shrubs, 21 Jun lake, Lomonosova 489 (NS); Russia, Chita obl., Onon distr., 35 km 2002, Barkalov 8846 (VLA); Russia, Far East, Primorskii Krai, from Solov’evsk to Borzya, Lomonosova 502 (NS); Russia, Chita Khankaiskii Raion, near Turii Rog settlement, 9 Jun 2007, Nechaev obl., Onon distr., Khuduktui lake, Lomonosova 521 (NS). 10609 (VLA).

2n = 32, CHN. Russia, Far East, Primorskii Krai, wildlife Nina S. Probatova1, Vitaly P. Seledets2, Elvira G. Rudyka1, reserve “Khankaiskii”, section Sosnovyi, Kazachii Bay, steppe- Aleksandr A. Gnutikov3, Zoya V. Kozhevnikova1 meadow community, 19 May 2004, Barkalov 9215 (VLA); Russia, & Vyacheslav Yu. Barkalov1 Far East, Amurskaya Oblast’, 6 km NW of Belogorsk, Tomi River, high terrace near the former river-bed, in vicinity of Tomskoe vil- 1 Institute of Biology & Soil Science, Far East Branch of Rus- lage, grass meadow-steppe community, 24 Aug 2006, Verkholat sian Academy of Sciences, 159 Stoletya Prospect, 690022, 10388 (VLA) Vladivostok, Russia. [email protected] (author for correspondence) Allium strictum Schrad. 2 Pacific Institute of Geography, Far East Branch of Russian 2n = 32, CHN. Russia, Far East, Kamchatskii Krai, Kam- Academy of Sciences, 7 Radio Str., 690041, Vladivostok, chatka Peninsula, ca. 20 km NE of Malki village, middle course of Russia Poperechnaya River, Betula ermanii forest, 28 Jul 1997, Barkalov 3 Baikal Research Center, Irkutsk State University, 5–10 Karl 7372 (VLA). Marx Str., 664003, Irkutsk, Russia AMARANTHACEAE * First chromosome count for the species. Amaranthus blitoides S. Watson ** New chromosome number (cytotype) for the species. 2n = 32, CHN. Russia, North Caucasus, Krasnodarskii Krai,

E11 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Cyclachaena xanthiifolia (Nutt.) Fresen. 11 Oct 2008, Probatova & Seledets 11286 (VLA). 2n = 36, CHN. Russia, Volga Region, Saratovskaya Oblast’, outskirts of Saratov city, Oktyabr’skoye canyon, waste place near ASCLEPIADACEAE the road, 5 Oct 2008, Probatova & Seledets 11188 (VLA). Pycnostelma paniculata (Thunb.) Makino 2n = 22, CHN. Russia, Far East, Primorskii Krai, Khanka Eupatorium cannabinum L. Lake, meadow on the lake terrace, 25 Jun 2006, Verkholat 10314 2n = 20, CHN. Russia, North Caucasus, Krasnodarskii Krai, (VLA). Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, 15 Oct 2008, Manilo 11190 (VLA). ASTERACEAE Artemisia glomerata Ledeb. Hieracium umbellatum L. 2n = 36, CHN. Russia, Far East, Kamchatskii Krai, Kam- 2n = 18, CHN. Russia, European part, Republic of Karelia, chatka Peninsula, Avachinskaya Sopka Volcano, southern slope, Pryazhinskii Raion, 4 km W of Kolatsel’ga village, Picea forest Lavovaya Pad’, subalpine zone, on volcanic scoria, 5 Sep 2000, with Vaccinium vitis-idaea and Calluna vulgaris, forest edge, 27 Yakubov 8441 (VLA). [Error: “2n = 18” — Probatova & al., Taxon Sep 2008, Probatova & Seledets 11207 (VLA). 57: 1269, E5. 2008.] Inula oculus-christi L. Artemisia integrifolia L. 2n = 32, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 36, CHN. Russia, Far East, Primorskii Krai, west coast of Starotitarovskii Raion, Tamanskii Peninsula, in vicinity of Solyonii Khanka Lake, nature reserve “Khankaiskii”, section Sosnovyi, Ka- village, salt depression on the shore of Akhtanizovskii estuary, zachii Bay, Salix spp. community, 2 Jul 2005, Barkalov 9816 (VLA). Tamarix-Phragmites-Puccinellia community, 12 Oct 2008, Pro- batova & Seledets 11282 (VLA). Bidens cernua L. 2n = 24, CHN. Russia, Far East, Primorskii Krai, Khanka Lapsana communis L. Lake, nature reserve “Khankaiskii”, Sosnovyi Isl., swampy plot, 2n = 14, CHN. Russia, European part, Republic of Karelia, 27 Jul 2002, Barkalov 8859 (VLA). Ladoga Lake, Valaam Isl., northeast slope of Mt. Favor, forest edge, roadside, 28 Sep 2008, Probatova & Seledets 11201 (VLA); Russia, Bidens frondosa L. Leningradskaya Oblast’, city of St.-Petersbourg, in the park of the 2n = 48, CHN. Russia, North Caucasus, Krasnodarskii Krai, V.L. Komarov Botanical Institute RAS, rare weed in plantations, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, 27 Oct 2008, Probatova & Seledets 11170 (VLA). 11 Oct 2008, Probatova & Seledets 11295 (VLA); Russia, Volga Region, Saratovskaya Oblast’, Saratov city, a rare weed along the Phalacroloma septentrionale (Fern. & Wieg.) Tzvelev road, 9 Oct 2008, Probatova & Seledets 11194 (VLA). 2n = 27, CHN. Russia, North Caucasus, Krasnodarskii Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Bidens tripartita L. 11 Oct 2008, Probatova & Seledets 11289 (VLA). 2n = 48, CHN. Russia, North Caucasus, Krasnodarskii Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Sonchus asper (L.) Hill 11 Oct 2008, Probatova & Seledets 11298 (VLA). 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Centaurea diffusa Lam. river bank, 15 Oct 2008, Manilo 11172 (VLA). 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Tripleurospermum inodorum (L.) Sch. Bip. 15 Oct 2008, Manilo 11187 (VLA). 2n = 36, CHN. Russia, Republic of Karelia, Ladoga Lake, Valaam Isl., slope of Mt. Favor, roadside, 28 Sep 2008, Probatova Centaurea frygia L. & Seledets 11177 (VLA); Russia, Leningradskaya Oblast’, city of 2n = 22, CHN. Russia, European part, Republic of Karelia, St.-Petersbourg, Petrogradskaya Storona district, quay of Karpovka Pryazhinskii Raion, Man’ga village, on the slope, 27 Sep 2008, River, a weed on the lawn, 27 Oct 2008, Probatova & Seledets Probatova & Seledets 11199 (VLA). 11180 (VLA).

Chondrilla juncea L. BORAGINACEAE **2n = 20, CHN. Russia, North Caucasus, Krasnodarskii Amsinckia micrantha Suksd. Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin 2n = 16, CHN. Russia, Far East, Magadanskaya Oblast’, River, 11 Oct 2008, Probatova & Seledets 11227 (VLA). Ol’skii Raion, Gadlya village, near a vegetable garden, 24 Aug 2006, Lyssenko 11240 (VLA). Cichorium intybus L. 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, BRASSICACEAE Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Barbarea arcuata (Opiz ex J. & C. Presl) Rchb. 11 Oct 2008, Probatova & Seledets 11274 (VLA). 2n = 16, CHN. Russia, North Caucasus, Krasnodarskii Krai, Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near Crepis setosa Haller f. Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11297 2n = 8, **16, CHN. Russia, North Caucasus, Krasnodarskii (VLA). Krai, Tamanskii Peninsula, Starotitarovskii Raion, between Solyo- nii and Starotitarovskaya villages, saline steppe community, 12 Oct Erysimum amurense Kitag. 2008, Probatova & Seledets 11272 (VLA). 2n = 12, CHN. Russia, Far East, Khabarovskii Krai, Solnech-

E12 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

nyi Raion, in vicinity of the railway station Sonakh, a rubbly- Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near Port melkozem slope, near the road, 21 Jul 2006, Barkalov 11325 (VLA). Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11186 (VLA).

Lepidium densiflorum Schrad. *Corispermum ulopterum Fenzl 2n = 32, CHN. Russia, Far East, Amurskaya Oblast’, outskirts 2n = 18, CHN. Russia, East Siberia, Republic of Buryatia, of Blagoveshchensk, on the way to Verkhne-Blagoveshchenskoe Barguzinskii Raion, Svyatoi Nos Peninsula, locality El’zikha, village, 25 Jul 2008, Timchenko 11228 (VLA). sandy shore of Kultuk Bay, 1 Sep 2003, Kazanovskii 11057 (VLA).

CANNABACEAE *Salsola pontica (Pall.) Degen Humulus lupulus L. 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 20, CHN. Russia, North Caucasus, Krasnodarskii Krai, Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11269 riverside shrubs, near a cottage, 11 Oct 2008, Manilo 11293 (VLA). (VLA).

CARYOPHYLLACEAE Teloxys aristata (L.) Moq. Dianthus versicolor Fisch. ex Link 2n = 18, CHN. Russia, East Siberia, Irkutskaya Oblast’, Ir- 2n = 30, CHN. Russia, Far East, Primorskii Krai, west lakeside kutsk city, Kirov square, a ruderal plant close to the building, 18 of Khanka Lake, nature reserve “Khankaiskii”, section Sosnovyi, Sep 2007, Probatova & Seledets 11120 (VLA). Kazachii Bay, meadow-steppe community, 2 Jul 2005, Barkalov 9815 (VLA). CHLORANTHACEAE Chloranthus japonicus Siebold Gypsophila perfoliata L. 2n = 30, CHN. Russia, Far East, Primorskii Krai, Shkotovskii **2n = 34, CHN. Russia, North Caucasus, Krasnodarskii Raion, near Anissimovka village, the valley of Smol’nyi spring, Krai, Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, Korean pine-broadleaved forest on the terrace, 14 Apr 1999, near Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets Barkalov 7798 (VLA). 11284 (VLA). CONVALLARIACEAE Lychnis fulgens Fisch. ex Curt. Maianthemum bifolium (L.) F.W. Schmidt 2n = 24, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii 2n = 54, CHN. Russia, Far East, Khabarovskii Krai, Ul’chskii Raion, outskirts of Grigorievka village, forest edge, 4 Jul 1998, Raion, 2–2.5 km W of Tabo Bay, east spur of Lobaz Mt., Picea and Shatalova 7677 (VLA). Abies forest, 6 Aug 2005, Kozhevnikov 9913 (VLA).

Spergula arvensis L. CONVOLVULACEAE 2n = 18, CHN. Russia, European part, Republic of Karelia, Calystegia dahurica (Herb.) Choisy Pitkyarantskii Raion, 10 km N of Pitkyaranta town, the edge of a 2n = 22, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii bog, 29 Sep 2008, Probatova & Seledets 11195 (VLA). Raion, SW of Novoshakhtinskii settlement, railbed slope, 6 Jul 2008, Lapenko 11050 (VLA). CHENOPODIACEAE Atriplex micrantha C.A. Mey. CRASSULACEAE 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, Sedum acre L. Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near 2n = 16, CHN. Russia, Republic of Karelia, in vicinity of Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11270 Ruskeala village, marble quarry, rocky slope, near path, 27 Sep (VLA). 2008, Probatova & Seledets 11175 (VLA).

Chenopodium foliosum Asch. EUPHORBIACEAE 2n = 18, CHN. Russia, East Siberia, Irkutskaya Oblast’, Ir- Acalypha australis L. kutsk city, Akademgorodok, as a weed on experimental plot of 2n = 20, CHN. Russia, North Caucasus, Krasnodarskii Krai, SIFIBR SO RAN, 29 Jun 2001, Repina 11058 (VLA). Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, 15 Oct 2008, Probatova & Seledets 11189 (VLA). Chenopodium polyspermum L. 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, FABACEAE Starotitarovskii Raion, Tamanskii Peninsula, in vicinity of Solyonii Lespedeza bicolor Turcz. village, salt depression on the shore of Akhtanizovskii estuary, 2n = 22, CHN. Russia, Far East, Amurskaya Oblast’, 40 km Tamarix-Phragmites-Puccinellia community, 12 Oct 2008, Pro- N of Blagoveshchensk city, wildlife protected area Mukhinka, batova & Seledets 11271 (VLA). mountainside, pine forest with Quercus mongolica, 30 Sep 2007, Probatova & Seledets 11239 (VLA). Chenopodium urbicum L. 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, Sophora flavescens Sol. Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near 2n = 18, CHN. Russia, Far East, Amurskaya Oblast’, 3–4 km Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11275 of Blagoveshchensk city, on the way to Verkhne-Blagoveshchens- (VLA). koe village, 25 Jul 2008, Timchenko 11077 (VLA).

Corispermum nitidum Schult. Trifolium repens L. 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 32, CHN. Russia, Far East, Amurskaya Oblast’, outskirts

E13 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

of Blagoveshchensk city, on the way to Verkhne-Blagoveshchens- Lycopus lucidus Turcz. ex Benth. koe village, 25 Jul 2008, Timchenko 11072 (VLA). 2n = 22, CHN. Russia, Far East, Primorskii Krai, nature reserve “Khankaiskii”, Przheval’skii Peninsula, on quagmire, 29 Jul Vicia nervata Sipl. 2002, Barkalov 8853 (VLA). 2n = 24, CHN. Russia, East Siberia, Irkutskaya Oblast’, Ol’khonskii Raion, near Sarma village, left riverside of the first Prunella asiatica Nakai right tributaryof Sarma River, 963 m, steppe community on a slope, 2n = 28, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii 29 Jul 2004, Kazanovskii 11225 (VLA). Raion, Grigor’evka village, artificial pond, meadow, 30 Jun 1989, Shatalova 7671 (VLA). FUMARIACEAE Corydalis repens Mandl & Muehld. Scutellaria krasevii Kom. & I. Schischk. 2n = 16, CHN. Russia, Far East, Primorskii Krai, Muravëv- 2n = 16, CHN. Russia, Far East, Primorskii Krai, nature re- Amurskii Peninsula, Vladivostok, near Akademgorodok, forest on serve “Khankaiskii”, Khanka Lake, Sosnovyi Isl., lakeside, 27 Jul the slope, 24 Apr 2009, Volynets 11317 (VLA). 2002, Barkalov 8867 (VLA).

Corydalis speciosa Maxim. LILIACEAE 2n = 16, CHN. Russia, Far East, Primorskii Krai, Chernigov- Fritillaria camschatcensis (L.) Ker-Gawl. skii Raion, railway station Buyanki, on the railroad embankment, 2n = 24, CHN. Russia, Far East, Kamchatskii Krai, Kam- 21 Jun 2008, Lapenko 11075 (VLA). chatka Peninsula, ca. 20 km NE of Malki village, middle course of Poperechnaya River, Betula ermanii forest, 28 Jul 1997, Barkalov HEMEROCALLIDACEAE 7366 (VLA). Hemerocallis minor Mill. 2n = 22, CHN. Russia, Far East, Primorskii Krai, nature re- Lilium debile Kittlitz serve “Khankaiskii”, section Sosnovyi, Przheval’skii Peninsula, 2n = 24, CHN. Russia, Far East, Kamchatskii Krai, Kam- among shrubs, 19 May 2004, Barkalov 9231 (VLA). chatka Peninsula, ca. 20 km north-eastwards of Malki village, middle course of Poperechnaya River, Betula ermanii forest, 28 HYPERICACEAE Jul 1997, Barkalov 7364 (VLA). Hypericum perforatum L. 2n = 32, CHN. Russia, Republic of Karelia, 7 km of Sortavala Lilium distichum Nakai town, arboretum “Dacha Vinera”, 27 Oct 2008, Kravchenko 11204 2n = 24, CHN. Russia, Far East, Khabarovskii Krai, (VLA). Komsomol’skii Raion, near the abandoned settlement Machtovyi, Machtovaya River, 5 Aug 2005, Barkalov 9916 (VLA). IRIDACEAE Iris setosa Pall. ex Link LIMONIACEAE 2n = 38, CHN. Russia, Far East, Kamchatskii Krai, Kam- Limonium caspium (Willd.) Gams chatka Peninsula, ca. 20 km NE of Malki village, middle course of 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, Poperechnaya River, downstream of the mouth of Zubastaya River, Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near Port meadow, 28 Jul 1997, Barkalov 7368, 7653 (VLA). Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11288 (VLA).

LAMIACEAE MALVACEAE Calamintha nepeta (L.) Savi Abutilon theophrasti Medik. 2n = 24, CHN. Russia, North Caucasus, Krasnodarskii Krai, **2n = 28, CHN. Russia, North Caucasus, Krasnodarskii Kamennoe More mountain Ridge, Lagonaki Plateau, ca. 1,800 m, Krai, Abinskii Raion, outskirts of Abinsk town, a weed in the val- in bush, 12 Sep 1991, Probatova & Seledets 7447 (VLA). ley of Abin River, 11 Oct 2008, Probatova & Seledets 11285 (VLA).

Glechoma longituba (Nakai) Kuprian. Althaea hirsuta L. 2n = 18, CHN. Russia, Far East, Primorskii Krai, Ussuriyskii **2n = 42, CHN. Russia, North Caucasus, Krasnodarskii Raion, 2 km S of Baranovskii railway station, railbed slope, 8 Jun Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin 2008, Lapenko 11032 (VLA). River, 11 Oct 2008, Probatova & Seledets 11331 (VLA).

2n = 36, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii Hibiscus trionum L. Raion, southwards of Novoshakhtinskii settlement, railbed slope, 2n = 56, CHN. Russia, Northern Caucasus, Krasnodarskii 15 May 2008, Lapenko 10962 (VLA). Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, river bank, 15 Oct 2008, Manilo 11184 (VLA). Lamium album L. 2n = 18, CHN. Russia, Moskovskaya Oblast’, city of Moscow, Malva mohileviensis Downar territory of the Main Botanical Garden RAS, along a foot-path, 29 **2n = 56, CHN. Russia, East Siberia, Irkutskaya Oblast’, Oct 2008, Probatova & Seledets 11166 (VLA). Irkutsk city, Pushkin Str., the waste place, 18 Sep 2007, Probatova & Seledets 11015 (VLA). Lycopus europaeus L. 2n = 22, CHN. Russia, North Caucasus, Krasnodarskii Krai, ONAGRACEAE Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Epilobium maximowiczii Hausskn. among riverside shrubs, 11 Oct 2008, Probatova & Seledets 11179 2n = 36, CHN. Russia, Far East, Primorskii Krai, Pogranich- (VLA). nyi Raion, in vicinity of the abandoned settlement Reshetnikovo,

E14 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

Mramornaya River valley, sandy-pebble shore, at the mouth of a River, temporary pond in the floodplain, 8 Jul 2005, Chepinoga spring, 22 Oct 1997, Barkalov 7510 (VLA). & al. 10516 (VLA).

ORCHIDACEAE Alopecurus arundinaceus Poir. Oreorchis patens (Lindl.) Lindl. 2n = 28, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii 2n = 48, CHN. Russia, Far East, Primorskii Krai, Shkotovskii Raion, W of Novoshakhtinskii settlement, at the bottom of the Raion, near Anissimovka village, the valley of Smol’nyi spring, railbed slope, 9 Jul 2008, Lapenko 11258 (VLA). Korean pine-broadleaved forest on the terrace, 14 Apr 1999, Barkalov 7767 (VLA). Anisantha sterilis (L.) Nevski 2n = 14, CHN. Russia, Northern Caucasus, Krasnodarskii OXALIDACEAE Krai, Tamanskii Peninsula, Starotitarovskii Raion, in vicinity of Xanthoxalis corniculata (L.) Small Solyonii village, salt depression on the shore of Akhtanizovskii es- 2n = 24, CHN. Russia, Far East, Amurskaya Oblast’, Bla- tuary, Tamarix-Phragmites-Puccinellia community, 12 Oct 2008, goveshchensk city, Relochnyi lane, as a weed of the lawn, 5 Oct Probatova & Seledets 11208 (VLA). 2007, Probatova & Seledets 10692 (VLA). Anthoxanthum odoratum L. PINACEAE 2n = 20, CHN. Russia, East Siberia, Krasnoyarskii Krai, in Picea koraiensis Nakai, 2n = 24, CHN. China, Heilongjiang vicinity of Tanzybei village, on the edge of a field, 3 Aug 2007, Province, Udal’anchi health resort, near health center, plantation Yakubov 11320 (VLA). of 30–40 years old, Sep 2008, Il’ina 11304 (VLA). Apera maritima Klokov POACEAE 2n = 14, CHN. Russia, North Caucasus, Krasnodarskii Krai, Achnatherum sibiricum (L.) Keng ex Tzvelev Tamanskii Peninsula, Starotitarovskii Raion, between Solyonii and 2n = 24, CHN. Russia, East Siberia, Republic of Tuva, Enissei Starotitarovskaya villages, salt depression, steppe, 12 Oct 2008, River, near the mouth of Sesserlik River, desertificated steppe on Probatova & Seledets 11303 (VLA). the watershed, 8 Aug 2007, Yakubov 11213 (VLA); Russia, East Siberia, Republic of Tuva, Enissei River, riverside near the mouth Arthraxon centrasiaticus (Griseb.) Gamajun. of Sesserlik River, valley Populus forest, 8 Aug 2007, Yakubov 2n = 36, CHN. Russia, Far East, Primorskii Krai, railway station 11217 (VLA). Amurskii Zaliv, waste ground, 24 Sep 2007, Lapenko 11256 (VLA).

Agropyron desertorum (Fisch. ex Link) Schult. Beckmannia syzigachne (Steud.) Fernald 2n = 14, CHN. Russia, Volga Region, Saratovskaya Oblast’, 2n = 14, CHN. Russia, West Siberia, Republic of Altai, south- outskirts of Saratov city, Oktyabr’skoye canyon, west slope, steppe east part, Kyzilchin locality, 10–11 Aug 2006, Olonova 10539 vegetation, 5 Oct 2008, Probatova & Seledets 11305 (VLA). (VLA); Russia, Far East, Magadanskaya Oblast’, Ten’kinskii Raion, Nel’koba village, roadside, 24 Aug 1993, Probatova & Agrostis capillaris L. Seledets 7131 (VLA); Russia, Far East, Magadanskaya Oblast’, 2n = 28, CHN. Russia, Republic of Karelia, Petrozavodsk city, Ten’kinskii Raion, 340 km of Ten’kinskaya route, IBPN biologi- lawn near the former Bank school building, 23 Sep 2008, Proba- cal station “Kontakt”, sample plot no. 35, 21 Aug 1993, Proba- tova & Seledets 11251 (VLA); Russia, European part, Republic tova & Seledets 7194 (VLA); Russia, Far East, Primorskii Krai, of Karelia, Valaam Isl., on the slope of Mt. Favor, 28 Sep 2008, Mikhailovskii Raion, Khanka Lake basin, 7 km of Grigorievka, Probatova & Seledets 11306 (VLA). Abramovka River near Abramovka village, river bank, 30 Jun 1998, Shatalova 7673 (VLA). Agrostis clavata Trin. 2n = 42, CHN. Russia, Far East, Amurskaya Oblast’, 3–4 km Bromopsis inermis (Leyss.) Holub of Blagoveshchensk, on the way to Verkhne-Blagoveshchenskoe 2n = 56, CHN. Russia, Far East, Amurskaya Oblast’, outskirts village, 25 Jul 2008, Timchenko 11067 (VLA); Russia, Far East, of Blagoveshchensk city, on the way to Verkhne-Blagoveshchens- Magadanskaya Oblast’, Srednekanskii Raion, wildlife reserve koe village, 25 Jul 2008, Timchenko 11329 (VLA). “Magadanskii”, left bank of Kolyma River, near Tyomnyi rivulet, 19 Aug 2007, Lyssenko 10993 (VLA). Bromus squarrosus L. 2n = 14, CHN. Russia, Volga Region, Volgogradskaya Oblast’, Agrostis scabra Willd. northern outskirts of Volgograd city, country place near GRES 2n = 42, CHN. Russia, Far East, Primorskii Krai, Khankaiskii settlement, 1 Aug 2006, Yurtzeva 10485 (VLA). Raion, in vicinity of Turii Rog settlement, 19 Jun 2002, Barkalov 8826 (VLA). 2n = 14+0–4B, CHN. Russia, North Caucasus, Krasnodarskii Krai, Tamanskii Peninsula, Starotitarovskii Raion, between Solyo- Agrostis trinii Turcz. nii and Starotitarovskaya villages, salt depression, steppe, 12 Oct 2n = 28, CHN. Russia, Far East, Primorskii Krai, Terneiskii 2008, Probatova & Seledets 11292 (VLA). Raion, Sikhote-Alinskii biosphere nature reserve, 25 km N of Ternei, locality Maissa, Yasnaya River, valley forest, 23 Sep 2005, Calamagrostis arundinacea (L.) Roth Seledets 10001 (VLA). 2n = 28, CHN. Russia, Republic of Karelia, Pitkyarantskii Raion, 20 km N of Sortavala town, Ladoga Lake, Kiryavalakhty Bay, Alopecurus aequalis Sobol. Picea forest on peninsula, 29 Sep 2008, Probatova & Seledets 11324; 2n = 14, CHN. Russia, East Siberia, Irkutskaya Oblast’, Zim- Russia, East Siberia, Republic of Khakassia, in vicinity of Dzhe- inskii Raion, 3 km SW of Novoletniki village, left side of Oka blakty, Pinus sylvestris forest, 14 Aug 2007, Yakubov 11244 (VLA).

E15 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Calamagrostis epigeios (L.) Roth of Yakutsk city, in vicinity of Mt. Chuchur-Muran, 27 Jul 1973, 2n = 28, CHN. Russia, Republic of Karelia, Ladoga L., Va- Probatova & Seledets 3753 (VLA). laam Isl., the slope of Mt. Favor, forest edge, near the road, 28 Sep 2008, Probatova & Seledets 11254 (VLA). Elymus fibrosus (Schrenk) Tzvelev 2n = 28, CHN. Russia, East Siberia, Irkutskaya Oblast’, Baikal Calamagrostis extremiorientalis (Tzvelev) Prob. L., 17 km NE of Listvyanka, near Bol’shie Koty village, Bol’shaya 2n = 28, CHN. Russia, Far East, Amurskaya Oblast’, 3–4 km Kotovka River, on pebbles, 12 Sep 2007, Probatova & Seledets of Blagoveshchensk, on the way to Verkhne-Blagoveshchenskoe 10726 (VLA); Russia, East Siberia, Republic of Buryatia, out- village, 25 Jul 2008, Timchenko 11159 (VLA). skirts of Ulan-Ude city, meadow-steppe vegetation, 12 Sep 1989, Probatova 6918 (VLA). Calamagrostis monticola V. Petrov ex Kom. 2n = 28, CHN. Russia, Far East, Khabarovskii Krai, Ul’chskii *Elymus franchetii Kitag. Raion, in vicinity of De-Castri settlement, summit of Mt. Arbat, 2n = 42, CHN. Russia, Far East, Khabarovskii Krai, in vicinity stony waste area, 20 Aug 2004, Yermoshkin 9759 (VLA). of the border of Bol’shekhekhtsirskii wildlife reserve, near the route Bychikha–Kazakevichevo, forest edge, 30 Aug 1993, Probatova Calamagrostis obtusata Trin. & Seledets 7083 (VLA). 2n = 28, CHN. Russia, East Siberia, Krasnoyarskii Krai, Za- padnyi Sayan mountain ridge, the valley of Omul R., Larix, Picea *Elymus hyperarcticus (Polunin) Tzvelev and Pinus forest, 7 Aug 2007, Yakubov 11249 (VLA). 2n = 28, CHN. Russia, Far East, Magadanskaya Oblast’, Vran- gel Isl., Mt. Tundrovaya, 18 Jun 1964, Petrovskii 11-64 (VLA). Сatabrosa aquatica (L.) P. Beauv. 2n = 20, CHN. Russia, West Siberia, Republic of Altai, south- Elymus novae-angliae (Scribn.) Tzvelev east part, Kyzilchin locality, 10–11 Aug 2006, Olonova 10545 (VLA). 2n = 28, CHN. Russia, Far East, Amurskaya Oblast’, Bla- goveshchensk city, roadside, 5 Oct 2007, Probatova & Seledets Сinna latifolia (Trevir.) Griseb. 10727 (VLA); Russia, Far East, Khabarovskii Krai, Solnechnyi 2n = 28, CHN. Russia, Far East, Magadanskaya Oblast’, Raion, Myao-Chan mountain ridge, in vicinity of Gornyi settle- Ol’skii Raion, in vicinity of the abandoned village Oyra, westwards ment, the bank of Silinka R., 5 Aug 2008, Van 11327 (VLA). of Armanj settlement, floodplain with Populus and Salix, 8 Sep 1995, Mochalova 7243 (VLA). Elymus sibiricus L. 2n = 28, CHN. Russia, West Siberia, Republic of Altai, Dactylis glomerata L. south-east part, Yuzhno-Chuyskii mountain ridge, Verkhnii Oyuk 2n = 28, CHN. Russia, East Siberia, Krasnoyarskii Krai, upper (Turoyik) Spring, upper mountain belt, 13 Aug 2006, Olonova course of Biryussa River, coniferous forest with tall herbs, 1 Aug 10531 (VLA); Russia, East Siberia, Republic of Tuva, Enissei 2007, Yakubov 11218 (VLA). River, near the mouth of Sesserlik River, slide-rocks, 10 Aug 2007, Yakubov 11211 (VLA). Digitaria ciliaris (Retz.) Koeler 2n = 36, CHN. Russia, North Caucasus, Krasnodarskii Krai, Elytrigia pontica (Podp.) Holub Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, 2n = 42, CHN. Russia, North Caucasus, Krasnodarskii Krai, river bank, 11 Oct 2008, Probatova & Seledets 11183 (VLA). Tamanskii Peninsula, Starotitarovskii Raion, between Solyonii and Starotitarovskaya villages, salt depression, steppe, 12 Oct 2008, Echinochloa oryzicola (Vasinger) Vasinger Probatova & Seledets 11301 (VLA). 2n = 36, CHN. Russia, Far East, Primorskii Krai, Khorol’skii Raion, Lugovoye, a weed of rice plantations, 17 Sep 2004, Elytrigia repens (L.) Nevski Lukacheva 9625 (VLA); Russia, Far East, Primorskii Krai, Spasskii 2n = 42, CHN. Russia, North Caucasus, Krasnodarskii Krai, Raion, near Novossel’skoye village, 17 Sep 2004, Lukacheva 9630 Abinskii Raion, outskirts of Abinsk town, the bank of Abin River (VLA); Russia, Far East, Primorskii Krai, Khankaiskii Raion, near (Kuban’ River basin), 15 Oct 2008, Manilo 11281 (VLA); Rus- Mel’gunovka village, 17 Sep 2004, Lukacheva 9631 (VLA). sia, Volga Region, Saratovskaya Oblast’, outskirts of Saratov city, Oktyabr’skoye canyon, west slope, steppe vegetation, 5 Oct 2008, Echinochloa spiralis Vasinger Probatova & Seledets 11311 (VLA); Russia, Far East, Amurskaya 2n = 36, CHN. Russia, North Caucasus, Krasnodarskii Krai, Oblast’, 3–4 km of Blagoveshchensk, on the way to Verkhne-Bla- Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, goveshchenskoe village, 25 Jul 2008, Timchenko 11070, 11071 (VLA). river bank, 15 Oct 2008, Manilo 11168 (VLA); Russia, Far East, Primorskii Krai, Khankaiskii Raion, near Vladimiro-Petrovka vil- Eragrostis amurensis Prob. lage, 17 Seр 2004, Lukacheva 9629 (VLA). 2n = 40; Russia, East Siberia, Chitinskaya Oblast’, railway station Khilok, as a weed on the edge of the platform, 17 Sep 2008, Elymus caninus (L.) L. Probatova & Seledets 11265 (VLA). 2n = 28, CHN. Russia, Republic of Karelia, Pitkyarantskii Raion, near Lyaskelya village, 27 Sep 2008, Probatova & Seledets Festuca altaica Trin. 11308 (VLA); Russia, East Siberia, Krasnoyarskii Krai, in vicinity 2n = 28, CHN. Russia, West Siberia, Republic of Altai, south- of Tanzybei settlement, coniferous forest on the slope, 3 Aug 2007, east part, Yuzhno-Chuyskii mountain ridge, the valley Akkol, up- Yakubov 11127 (VLA). per mountain belt, 12 Aug 2006, Olonova 10536 (VLA).

*Elymus charkeviczii Prob. Festuca sphagnicola B. Keller 2n = 28, CHN. Russia, Republic of Sakha-Yakutia, 10 km SW 2n = 28, CHN. Russia, East Siberia, Krasnoyarskii Krai,

E16 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

Zapadnyi Sayan mountain ridge, wildlife territory “Ergaki”, the Tamanskii Peninsula, Starotitarovskii Raion, between Solyonii and slope covered with mosses along the Abies forest edge, 4 Aug Starotitarovskaya villages, saline steppe community, 12 Oct 2008, 2007, Yakubov 11245 (VLA); Russia, East Siberia, Krasnoyarskii Probatova & Seledets 11299 (VLA). Krai, Zapadnyi Sayan mountain ridge, wildlife territory “Ergaki”, the stony slope covered with mosses in subalpine mountain belt, Phleum phleoides (L.) H. Karst. 4 Aug 2007, Yakubov 11220 (VLA); Russia, East Siberia, Kras- 2n = 28, CHN. Russia, East Siberia, Republic of Tuva, in noyarskii Krai, Zapadnyi Sayan mountain ridge, wildlife territory vicinity of Sesserlik village, steppe vegetation on the slope, 9 Aug “Ergaki”, the slope of Mt. Aerodrom, dwarf shrub–moss and lichen 2007, Yakubov 11215 (VLA). tundra, 5 Aug 2007, Yakubov 11247 (VLA); Russia, East Siberia, Krasnoyarskii Krai, wildlife territory “Ergaki”, the top of the Mt. Poa alpigena (Blytt) Lindm. Tushkanchik, dwarf shrub–lichen tundra, 11 Aug 2007, Yakubov 2n = ca. 56, CHN. Russia, Far East, Magadanskaya Oblast’, 11248 (VLA). Srednekanskii Raion, Kolymskoye village, meadow near the port, 26 Jul 2005, Lyssenko 10655 (VLA). Glyceria triflora (Korsh.) Kom. 2n = 20, CHN. Russia, East Siberia, Krasnoyarskii Krai, in 2n = 56, CHN. Russia, European part, Arkhangel’skaya vicinity of Tanzybei settlement, bank of the forest spring, 3 Aug Oblast’, Onezhskii Raion, Kozhozerskii wildlife park, 4.5 km SW 2007, Yakubov 11131 (VLA); Russia, Far East, Khabarovskii Krai, of the mouth of Boyan’ Spring, a bog, 6 Sep 2005, Timofeeva & Komsomol’skii Raion, Myao-Chan mountain ridge, the valley of Kutenkov 11173 (PTZ). Khudaki River, roadside on the slope, 16 Jul 2008, Van 11318 (VLA). Poa altaica Trin. Hordeum jubatum L. 2n = 28, CHN. Russia, West Siberia, Republic of Altai, 2n = 28, CHN. Russia, East Siberia, Irkutskaya Oblast’, Irkutsk southeast part, Yuzhno-Chuyskii mountain ridge, Verkhnii Oyuk city, roadside, 18 Sep 2007, Probatova & Seledets 11106 (VLA). (Turoyik) Spring, upper mountain belt, 13 Aug 2006, Olonova 10544 (VLA). Hordeum murinum L. 2n = 14, CHN. Russia, Northern Caucasus, Krasnodarskii Poa angustifolia L. Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin 2n = 56, CHN. Russia, Volga Region, Saratovskaya Oblast’, River, 15 Oct 2008, Manilo 11277 (VLA). outskirts of Saratov city, Oktyabr’skoye canyon, west slope, steppe community, 5 Oct 2008, Probatova & Seledets 11323 (VLA); Rus- Koeleria altaica (Domin) Krylov sia, East Siberia, Chitinskaya Oblast’, railway station Amazar, **2n = 14, CHN. Russia, West Siberia, Republic of Altai, south- along the railroad embankment, 17 Jun 2007, Tsyrenova 10818 east part, Kyzilchin locality, 10–11 Aug 2006, Olonova 10530 (VLA). (VLA). [Probatova & al., Taxon 57: 1273, E15. 2008, as “Poa pratensis”.] Koeleria cristata (L.) Pers. 2n = 14, CHN. Russia, North Caucasus, Krasnodarskii Krai, Poa attenuata Trin. Tamanskii Peninsula, Starotitarovskii Raion, between Solyonii and 2n = 28, CHN. Russia, West Siberia, Republic of Altai, south- Starotitarovskaya villages, salt depression, steppe, 12 Oct 2008, east part, Yuzhno-Chuyskii mountain ridge, the valley Akkol, up- Probatova & Seledets 11302 (VLA). per mountain belt, 12 Aug 2006, Olonova 10542 (VLA).

Leersia oryzoides (L.) Sw. Poa botryoides (Trin. ex Griseb.) Kom. 2n = 48, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 28, CHN. Russia, East Siberia, Republic of Tuva, water- Abinskii Raion, outskirts of Abinsk town, the bank of Abin River shed of Enissei River, near the mouth of Sesserlik River, desertifi- (Kuban’ River basin), 11 Oct 2008, Probatova & Seledets 11181 cated steppe, 8 Aug 2007, Yakubov 11226 (VLA). (VLA). Poa bulbosa L. Leymus angustus (Trin.) Pilg. 2n = 28, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 84, CHN. Russia, East Siberia, Republic of Tuva, riv- Tamanskii Peninsula, Starotitarovskii Raion, between Solyonii and erside of Enissei River, near the mouth of Sesserlik River, steppe Starotitarovskaya villages, salt depression, steppe, 12 Oct 2008, with Caragana, 8 Aug 2007, Yakubov 11221 (VLA). Probatova & Seledets 11300 (VLA).

Lolium perenne L. Poa compressa L. 2n = 14, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 42, CHN. Russia, Republic of Karelia, Sortavala town, Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near the bay of Ladoga Lake, stony slope near the bridge, 28 Sep 2008, Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11312 Probatova & Seledets 11203 (VLA). (VLA). *Poa krylovii Reverd. Panicum ruderale (Kitag.) Chang 2n = 42, CHN. Russia, East Siberia, Republic of Tuva, in vi- 2n = 36, CHN. Russia, East Siberia, Irkutskaya Oblast’, cinity of Sesserlik village, rocks on the slope, 9 Aug 2007, Yakubov Cheremkhovskii Raion, near Verkhnii Bulai village, 1,759th km 11212 (VLA). of Moskovskii route, on the railway embankment, 29 Aug 2006, Chepinoga & Enuschenko 10880 (VLA). *Poa lapponica Prokudin 2n = 28, CHN. Russia, Republic of Karelia, Petrozavodsk Phleum nodosum L. city, lawn near the former Bank School building, 23 Sep 2008, 2n = 14, CHN. Russia, North Caucasus, Krasnodarskii Krai, Probatova & Seledets 11262 (VLA).

E17 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Poa palustris L. Puccinellia tenuiflora (Griseb.) Scribn. & Merr. 2n = 28, CHN. Russia, Far East, Amurskaya Oblast’, 3–4 km 2n = 28, CHN. Russia, East Siberia, Republic of Buryatia, of Blagoveshchensk, on the way to Verkhne-Blagoveshchenskoe Bichurskii Raion, 9 km SW of Okino-Kluchi village, salt lake, village, 25 Jul 2008, Timchenko 11065 (VLA); Russia, Far East, 10 Jul 2006, Chepinoga & Dulepova 10855 (VLA). Khabarovskii Krai, Komsomol’skii Raion, Khudaki R. valley (Amur R. basin), roadside, 1 Jul 2008, Van 11267 (VLA); Russia, Schedonorus pratensis (Huds.) P. Beauv. Far East, Primorskii Krai, Pogranichnyi Raion, in vicinity of the 2n = 14, CHN. Russia, Republic of Karelia, Pitkyarantskii abandoned settlement Reshetnikovo, Mramornaya River valley, Raion, near Lyaskelya village, at the bottom of the slope, roadside, sandy-pebble shore, at the mouth of a spring, 22 Oct 1997, Barkalov 27 Sep 2008, Probatova & Seledets 11316 (VLA). 7518 (VLA). Setaria pumila (Poir.) Roem. & Schult. Poa sibirica Roshev. 2n = 36, CHN. Russia, North Caucasus, Krasnodarskii Krai, 2n = 14, CHN. Russia, East Siberia, Krasnoyarskii Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Zapadnyi Sayan mountain ridge, the valley of Omul R., Larix, river bank, on pebbles, 15 Oct 2008, Manilo 11171 (VLA). Picea and Pinus forest, 7 Aug 2007, Yakubov 11135 (VLA); Rus- sia, East Siberia, Krasnoyarskii Krai, Zapadnyi Sayan mountain Setaria verticillata (L.) P. Beauv. ridge, wildlife territory “Ergaki”, subalpine belt, stony slope, on 2n = 36, CHN. Russia, Volga Region, Saratovskaya Oblast’, the slide-rocks, 4 Aug 2007, Yakubov 11250 (VLA). outskirts of Saratov city, Oktyabr’skoye canyon, a lawn, near the road, 5 Oct 2008, Probatova & Seledets 11310 VLA). Poa skvortzovii Prob. 2n = 28, CHN. Russia, Far East, Amurskaya Oblast’, 3–4 km Setaria viridis (L.) P. Beauv. of Blagoveshchensk, on the way to Verkhne-Blagoveshchenskoe 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, village, 25 Jul 2008, Timchenko 11068 (VLA); Russia, Far East, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Khabarovskii Krai, Myao-Chan mountain ridge, left side of riverbank, 15 Oct 2008, Manilo 11273 (VLA). Tsirkul’ River valley, 3 km N of Eliberdan Mt., slide-rocks, 1 Aug 2008, Van 11268 (VLA). Stipa capillata L. 2n = 44, CHN. Russia, East Siberia, Republic of Tuva, Enissei 2n = 42, CHN. Russia, Far East, Khabarovskii Krai, 3–4 km River, near the mouth of Sesserlik River, steppe community with from Petropavlovka village, lakeside of Petropavlovskoe L., 4 Jul Caragana, 8 Aug 2007, Yakubov 11233 (VLA). 1971, Probatova & Seledets 3261 (VLA). Trisetum altaicum Roshev. Poa sugawarae Ohwi 2n = 14, CHN. Russia, East Siberia, Vostochnyi Sayan moun- 2n = 28, CHN. Russia, Far East, Sakhalinskaya Oblast’, North- tain ridge, Republic of Buryatia, Okinskii Raion, near Choigan- ern Sakhalin, Schmidt Peninsula, Bol’shaya Longri River, rubbly- Daban Pass, riverside of the confluent of Dunda-Gol River, on melkozem slope of a hill, 16 Aug 2001, Barkalov 8596 (VLA). rocks, 15 Aug 2007, Vedernikova 11230 (VLA).

Poa trivialis L. Trisetum mongolicum (Hult.) Peschkova 2n = 14, CHN. Russia, Republic of Karelia, Ladoga Lake, 2n = 28, CHN. Russia, West Siberia, Republic of Altai, Valaam Isl., slope of Mt. Favor, forest edge, along rivulet, 28 Sep southeast part, Yuzhno-Chuyskii mountain ridge, Verkhnii Oyuk 2008, Probatova & Seledets 11196 (VLA). (Turoyik) Spring, upper mountain belt, 13 Aug 2006, Olonova 10533 (VLA). Psathyrostachys juncea (Fisch.) Nevski 2n = 14, CHN. Russia, East Siberia, Krasnoyarskii Krai, Re- Trisetum sibiricum Rupr. public of Tuva, Enissei River, the slope near the mouth of Sesserlik 2n = 14, CHN. Russia, East Siberia, Republic of Tuva, near River, steppe community with Caragana, 8 Aug 2007, Yakubov Sesserlik village, rocky slope of a hill, 9 Aug 2007, Yakubov 11224 11229 (VLA). (VLA).

Puccinellia distans (Jacq.) Parl. POLYGONACEAE 2n = 42, CHN. Russia, Far East, Primorskii Krai, Muravëv- Polygonum calcatum Lindm. Amurskii Peninsula, Ussuriyskii Gulf, Muraviinaya Bay, sea 2n = 20, CHN. Russia, Far East, Magadanskaya Oblast’, coast, recreation area, 21 Oct 2004, Probatova & Seledets 9607 Ten’kinskii Raion, 332 km of Ten’kinskaya route, IBPN biologi- (VLA). cal station “Kontakt”, the bank of rivulet Kontaktovyi, on pebbles, 20 Aug 1993, Probatova & Seledets 7188 (VLA). Puccinellia hauptiana V.I. Krecz. 2n = 28, CHN. Russia, Far East, Magadanskaya Oblast’, Polygonum maritimum L. Ten’kinskii Raion, 332 km of Ten’kinskaya route, IBPN biologi- 2n = 20, CHN. Russia, North Caucasus, Krasnodarskii Krai, cal station “Kontakt”, bank of rivulet Kontaktovyi, 20 Aug 1993, Tamanskii Peninsula, Chushka spit, coast of the Sea of Azov, near Probatova & Seledets 7264 (VLA). Port Kavkaz, sand dunes, 12 Oct 2008, Probatova & Seledets 11290 (VLA). Puccinellia macranthera V.I. Krecz. **2n = 56, CHN. Russia, East Siberia, Republic of Buryatia, Rumex amurensis F. Schmidt ex Maxim. Bichurskii Raion, 2 km N of Okino-Kluchi village, flood-lands of 2n = 20, CHN. China, Heilongjiang Province, Kharbin city, Topka River (left confluent of Khilok River), saline meadow, 7 Jul sandbank of Sungari River, 2 Jul 2000, Probatova & Seledets 8152 2006, Chepinoga & Dulepova 10852 (VLA). (VLA).

E18 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

Rumex crispus L. Raion, 3 km W of Kremovo village, meadow, 15 Jun 2008, Lapenko **2n = 40, CHN. Russia, Far East, Khabarovskii Krai, 11033 (VLA). Komsomol’skii Raion, the abandoned settlement Machtovyi, Mach- tovaya River (the Amur River basin), wet meadow, 6 Aug 2005, Ranunculus repens L. Barkalov 10080 (VLA). 2n = 32, CHN. Russia, Far East, Amurskaya Oblast’, 3–4 km of Blagoveshchensk, on the way to Verkhne-Blagoveshchenskoe Rumex patientia L. village, 25 Jul 2008, Timchenko 11069 (VLA); Russia, Far East, 2n = 40, CHN. Russia, Far East, Primorskii Krai, Khankaiskii Khabarovskii Krai, Komsomol’skii Raion, in vicinity of abandoned Raion, west coast of Khanka Lake, nature reserve “Khankaiskii”, settlement Machtovyi, Machtovaya River (the Amur River basin), Sosnovyi area, Przheval’skii Peninsula, wet meadow, 21 Jun 2002, country road, 6 Aug 2005, Barkalov 9897 (VLA); Russia, Far East, Barkalov 8831 (VLA). Primorskii Krai, nature reserve “Khankaiskii”, section Sosnovyi, Przheval’skii Peninsula, sandbank of Khanka Lake, 19 May 2004, Rumex pseudonatronatus (Borb.) Borb. ex Murb. Barkalov 9230 (VLA). 2n = 40, CHN. Russia, Far East, Khabarovskii Krai,, Komsomol’skii Raion, the abandoned settlement Machtovyi, Mach- ROSACEAE tovaya River (the Amur River basin), wet meadow, 6 Aug 2005, Duchesnea indica (Andr.) Focke Barkalov 10081 (VLA). 2n = 42, CHN. U.S.A., Texas, Addison (near Dallas), as a weed on the lawn, 6 Apr 2005, Rudyka 9724 (VLA). Rumex sylvestris (Lam.) Wallr. **2n = 20, CHN. Russia, North Caucasus, Krasnodarskii Fragaria mandshurica Staudt Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin 2n = 14, CHN. Russia, Far East, Khabarovskii Krai, 15 km River, 15 Oct 2008, Manilo 11276 (VLA). NW of Komsomol’sk-na-Amure city, lower course of Tsirkul’ River, clearing, 12 Jul 2006, Barkalov 11360 (VLA). PRIMULACEAE Androsace amurensis Prob. Rosa rugosa Thunb. 2n = 20, CHN. Russia, East Siberia, Irkutskaya Oblast’, 2n = 14, CHN. Russia, Far East, Primorskii Krai, Khassan- Ol’khonskii Raion, lakeside of Baikal Lake, Cape Zunduk, beach skii Raion, Troitsa Bay, sandy beach, 19 Sep 2004, Shatokhina ridge, on pebbles, 3 Aug 2003, Kazanovskii 11059 (VLA). 58 (VLA).

Androsace septentrionalis L. Spiraea media F. Schmidt 2n = 20, CHN. Russia, Far East, Primorskii Krai, west coast 2n = 18, CHN. Russia, East Siberia, Irkutskaya Oblast’, Bai- of Khanka Lake, nature reserve “Khankaiskii”, section Sosnovyi, kal Lake, 17 km NE of Listvyanka, near Bol’shie Koty village, Przheval’skii Peninsula, sandy spit, Salix community, 21 Jun 2002, Pinus-Larix forest on the slope, 13 Sep 2007, Probatova & Seledets Barkalov 8943 (VLA). 11243 (VLA).

Lysimachia barystachys Bunge 2n = 24, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii Raion, south-eastwards of Novoshakhtinskii settlement, the railbed SAXIFRAGACEAE slope, 6 Jul 2008, Lapenko 11048 (VLA). Bergenia crassifolia (L.) Fritsch 2n = 34, CHN. Russia, East Siberia, Republic of Buryatia, RANUNCULACEAE Pribaikal’skii Raion, Turka village, the coast of Baikal Lake, Pinus- Anemonidium dichotomum (L.) Holub Bergenia forest, 11 Sep 2004, Chepinoga 9422 (VLA). 2n = 28, CHN. Russia, Far East, Primorskii Krai, nature reserve “Khankaiskii”, section Sosnovyi, Kazachii Bay, moist SCROPHULARIACEAE meadow on the edge of Salix community, 19 May 2004, Barkalov Kickxia elatine (L.) Dumort. 9236 (VLA). 2n = 18, CHN. Russia, North Caucasus, Krasnodarskii Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin River, Consolida paniculata (Host) Schur 11 Oct 2008, Probatova & Seledets 11296 (VLA). 2n = 16, CHN. Russia, North Caucasus, Krasnodarskii Krai, Tamanskii Peninsula, Starotitarovskii Raion, between Solyonii Limosella aquatica L. and Starotitarovskaya villages, saline steppe community, 12 Oct 2n = 40, CHN. Russia, Far East, Primorskii Krai, nature re- 2008, Probatova & Seledets 11192 (VLA), 21 Jun 2002, Barkalov serve “Khankaiskii”, section Sosnovyi, moist bank of Khanka 8828 (VLA). Lake, 20 Jun 2004, Barkalov 9271 (VLA).

Ranunculus chinensis Bunge Linaria genistifolia (L.) Mill. 2n = 16, CHN. Russia, Far East, Primorskii Krai, nature reserve 2n = 12, CHN. Russia, North Caucasus, Krasnodarskii Krai, “Khankaiskii”, Przheval’skii Peninsula, Barkalov 8828 (VLA). Tamanskii Peninsula, Chushka spit, between Port Kavkaz and Port Il’ich, near Dinskoi estuary (the Black Sea), 12 Oct 2008, Proba- Ranunculus gmelinii DC. tova & Seledets 11283 (VLA). 2n = 16, CHN. Russia, Far East, Primorskii Krai, west coast of Khanka Lake, Oct 1997, Barkalov 7520 (VLA). Linaria vulgaris L. 2n = 12, CHN. Russia, Northern Caucasus, Krasnodarskii Ranunculus japonicus Thunb. Krai, Abinskii Raion, outskirts of Abinsk town, the valley of Abin 2n = 14, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii River, 11 Oct 2008, Manilo 11193 (VLA); Russia, East Siberia,

E19 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Irkutskaya Oblast’, Baikal Lake, 17 km NE of Listvyanka, near L.P. Gavilán MS 597 (SALA 135 418) [Fig. 1B]; Spain, Ávila, Are- Bol’shie Koty village, Bol’shaya Kotovka River, on pebbles, 12 nas de San Pedro, river Pelayo, 40°12′N 05°07′W, 530 m, Alnus glu- Sep 2007, Probatova & Seledets 10734 (VLA). tinosa riparian forest, 10 Jul 2005, M. Santos-Vicente & L. Delgado MS 599 (SALA 135419) [Fig. 1C]. Verbascum thapsus L. Although this taxon has been widely studied in Europe from a 2n = 36, CHN. Russia, Republic of Karelia, in vicinity of karyological point of view (e.g., reports compiled by Bolkhovskikh Ruskeala village, marble quarry, rocky slope, near path, 27 Sep & al., 1969; van Loon, 1987; or Löve & Löve, 1961), these are the 2008, Probatova & Seledets 11198 (VLA). first populations studied in Spain.

Veronica daurica Steven Inula ensifolia L. 2n = 34, CHN. Russia, Far East, Primorskii Krai, Mikhailovskii **2n = 16, CHN. Italy, Udine, Gemona del Friuli, ascent to Raion, 3 km SE of Kremovo village, slope of a hill, 3 Aug 2008, Quarnan mountain, 46°17′N 13°08′E, 390 m, moist grazed land, Lapenko 11096 (VLA). calcareous soil, 17 Jul 2005, M.M. Martínez-Ortega, L.M. Muñoz- Centeno, L. Delgado & M. Santos-Vicente MS 600 (SALA 135420) TRILLIACEAE [Fig. 1D]. Trillium camschatcense Ker-Gawl. As for the preceding taxon, there are several previous chromo- 2n = 10, CHN. Russia, Far East, Kamchatskii Krai, Kam- some counts published for this species in Europe (see references in chatka Peninsula, ca. 20 km NE of Malki village, middle course of Bolkhovskikh & al., 1969, and Löve & Löve, 1961, among others). Poperechnaya River, Betula ermanii forest, 28 Jul 1997, Barkalov This represents the first chromosome number count for Italy and 7365 (VLA). coincides with previous counts.

VIOLACEAE Inula helenium L. Viola aff. acuminata Ledeb. **2n = 20, CHN. Spain, Soria, Abejar, towards Molinos de 2n = 20, CHN. Russia, Far East, Primorskii Krai, nature reserve Duero, 41°48′N 02°47′W, 1,100 m, moist places along roadside, “Khankaiskii”, section Sosnovyi, Przheval’skii Peninsula, Corylus 23 Aug 2007, M. Santos-Vicente, L. Delgado & C. Molina MS 957 heterophylla community, 19 May 2004, Barkalov 9223 (VLA). (SALA 135423) [Fig. 1E]. This is the first chromosome count of the species for the Ibe- rian Peninsula, which is in agreement with other counts reported María Santos-Vicente, Enrique Rico & M. Montserrat from Europe (e.g., Bolkhovskikh & al., 1969; Löve & Löve, 1961) Martínez-Ortega or China (Ma & al., 1985). No ploidy level variation has been found for this species, native from SE Europe and most probably Departamento de Botánica, Universidad de Salamanca, 37007 naturalized in the rest of its distribution area. Salamanca, Spain. [email protected] (author for correspondence) Inula helvetica Grauer * First chromosome count for the species. *2n = 16, CHN. Spain, Burgos, Jurisdicción de San Zadornil, ** New chromosome number (cytotype) for the species. San Millán de San Zadornil, river Omecillo, 42°51′N 03°09′W, 610 m, Populus riparian forest, 12 Jul 2004, J.A. Sánchez-Agudo, M. Santos- This study was supported by the Spanish Dirección General de Vicente & S. Andrés MS 407 (SALA 135417); Spain, Álava, Lantarón, Investigación (Ministerio de Ciencia y Tecnología and Ministerio Puentelarrá, river Ebro bank, 42°44′N 03°02′W, 463 m, Populus-Salix de Educación y Ciencia) through the projects Flora Iberica VI riparian forest, 30 Jun 2006, M. Santos-Vicente, M.M. Martínez- (REN2002-04634-C05-02; grant ref. BES-2003-2213 to M. Santos Ortega, S. Andrés & A. Abad MS 752 (SALA 135459) [Fig. 1F] Vicente) and Flora Iberica VII (CGL2005-05471-C04-03) and by We here report the first chromosome count for this taxon with the Junta de Castilla y León through project SA142A08. We greatly a scattered distribution over SW Europe. The chromosome number acknowledge this support. of the putative hybrid between I. helvetica and I. germanica L. (I. ×semiamplexicaulis Reut.) has been recently studied by Verlaque ASTERACEAE & Reduron (2008; 2n = 16) and matches up with the chromosome Inula bifrons L. number of the putative parental species; further studies should be **2n = 16, CHN. Spain, Segovia, Sacramenia, Coto de San carried out to clarify the taxonomic status of this taxon. Bernardo, 41°29′N 03°54′W, 900 m, clearings in Quercus faginea forest on calcareous soil, 10 Aug 2007, M. Santos-Vicente & L. Del- Literature cited gado MS 948 (SALA 135422); France, Alpes–de-Haute-Provence, Bolkhovskikh, Z., Grif, V., Matvejeva, T. & Zakharyeva, O. Reillanne, Pierrefeu farm, 43°53′N 05°38′E, 630 m, pasture in 1969. Khromosomnye Chisla Tsvetkovykh Rastenii [Chro- clearings of Quercus pubescens forest, 26 Jul 2006, M. Santos- mosome Numbers of Flowering Plants]. Nauka, Leningrad. Vicente & L. Delgado MS 777 (SALA 135421) [Fig. 1A]. Chouksanova, N.A., Sveshnikova, L.I. & Alexandrova, T.V. The chromosome numbers presented here agree with those 1968. Data on karyology of the family Compositae Giseke. reported for Bulgaria (Kuzmanov & Georgieva, 1983), Hungary Citologija 10: 198–206. (Sz.-Borsos, 1970) and Russia (Chouksanova & al., 1968). The Kuzmanov, B.A. & Georgieva, S. 1983. Asteraceae. In: Löve, first accession was collected in the only population of the species A. (ed.), IOPB Chromosome number reports, LXXXI. known in Spain. Taxon 32: 663–669 Loon, J.C. van. 1987. A Cytotaxonomical Atlas of the Balkan Inula conyzae (Griess.) Meikle Flora. Cramer, Berlín. **2n = 32, CHN. Spain, Burgos, Cillaperlata, El Castro, Löve, A. & Löve, D. 1961. Chromosome numbers of central 42°45′N 03°21′W, 840 m, Quercus faginea forest mixed with Buxus and northwest European plant species. Opera Bot. 5: 1–581. sempervirens, calcareous soil, 26 Jun 2005, M. Santos-Vicente & Ma, X.H., Qin, R.L. & Xing, W.B. 1985. Chromosome obser-

E20 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

FABACEAE Astragalus (sect. Incani) curvirostris Boiss. ssp. curvirostris *2n = 16, CHN. Iran, Lorestan, Boroujerd, East of Borou- jerd, 19 km from Boroujerd to Arak, before Zallian neck, Kabu- tarlan Village, 33°53′N, 48°44′E, 2,070 m, 21 Jul 2008, Dehshiri & Bohloli 91609 (TARI); Iran, Lorestan, Boroujerd, West of Borou- jerd, Kulidar Village, 33°53′N, 48°39′E, 2,100 m, 20 Jul 2008, Dehshiri & Bohloli 91610 (TARI).

Astragalus (sect. Incani) cyclophyllos Beck *2n = 16, CHN. Iran, Lorestan, Boroujerd, East of Borou- jerd, 19 km from Boroujerd to Arak, before Zallian neck, Kabu- tarlan Village, 33°53′N, 48°44′E, 2,070 m, 21 Jul 2008, Dehshiri & Bohluli 91611 (TARI).

Astragalus (sect. Onobrychoidei ) effusus Bunge *2n = 32, CHN. Iran, Lorestan, Boroujerd, East of Borou- jerd, 19 km from Boroujerd to Arak, before Zallian neck, Kabu- tarlan Village, 33°53′N, 48°44′E, 2,070 m, 21 Jul 2008, Dehshiri & Bohloli 91612 (TARI); Iran, Lorestan, Boroujerd, West of Borou- jerd, Kulidar Village, 33°53′N, 48°39′E, 2,100 m, 20 Jul 2008, Dehshiri &Bohloli 91613 (TARI).

Astragalus (sect. Incani) kabutarlanensis Dehshiri & Maassoumi *2n = 16, CHN. Iran, Lorestan, Boroujerd, East of Borou- jerd, 19 km from Boroujerd to Arak, before Zallian neck, Kabu- tarlan Village, 33°53′N, 48°44′E, 2,070 m, 21 Jul 2008, Dehshiri & Bohloli 91614 (TARI).

Astragalus (sect. Incani) supervisus (Kuntze) Sheld. *2n = 16, CHN. Iran, Lorestan, Boroujerd, West of Boroujerd, Kulidar Village, 33°53′N, 48°39′E, 2,100 m, 20 Jul 2008, Dehshiri & Bohloli 91615 (TARI).

Fig. 1. A, Inula bifrons, 2n = 16 (SALA 135421); B, I. conyzae, Johan J. Spies1, Paula Spies1, Adre Minnaar1, Susan M.C. 2n = 32 (SALA 135418); C, I. conyzae, 2n = 32 (SALA 135419); Reinecke1, Hanno (J.L.) du Preez1 & Riana Kleynhans1, 2 D, I. ensifolia, 2n = 16 (SALA 135420); E, I. helenium, 2n = 20 (SALA 135423); F, I. helvetica, 2n = 16 (SALA 135459). All of 1 Department of Genetics (64), University of the Free State, them mitotic metaphases. Scale bars = 10 μm. P.O. Box 339, Bloemfontein 9300, South Africa. spiesjj.sci@ ufs.ac.za (author for correspondence) 2 ARC Roodeplaat-Vegetable and Ornamental Plant Institute, vation of twenty species of drug plants in Xingjiang. Acta Private Bag X293, 0001 Pretoria, South Africa Bot. Boreal.-Occid. Sin. 5: 149–154. Sz.-Borsos, O. 1970. Contributions to the knowledge on the Degree reference system according to Edwards, D. & Leistner, chromosome numbers of phanerogams growing in Hun- O.A. 1971. A degree reference system for citing biological records gary and southeastern Europe. Acta Bot. Acad. Sci. Hung. in southern Africa. Mitt. Bot. Staatssamml. München 10: 501–509. 16: 255–265. Verlaque, R. & Reduron, J.-P. 2008. Inula ×semiamplexicau- HYACINTHACEAE lis Reut. In: Marhold, K. (ed.), IAPT/IOPB chromosome Lachenalia algoensis Schönland data 5. Taxon 57: 562, E24. 2n = 14, CHN. South Africa, Western Cape, 3422 (Mossel Bay): Sedgefield (-BB), Jan 2004, J.J. Spies 7988 (BLFU).

Mohammad Mehdi Dehshiri & Rezvan Bohloli Lachenalia aloides var. aurea (Lindl.) Engl. 2n = 14, CHN. South Africa, Western Cape, 3319 (Worcester): Department of Biology, Faculty of Science, Islamic Azad Bainskloof (-AC), 1983, J.J. Spies 7060 (BLFU). University (Boroujerd Branch), Boroujerd, Lorestan, Iran. [email protected] (author for correspondence) Lachenalia aloides var. luteola G.D. Duncan 2n = 14, CHN. South Africa, Western Cape, 3318 (Cape Town): * First chromosome count for the species. Voorberg jail, Porterville (-BB), 1988, J.J. Spies 7061 (BLFU).

This study was supported by the grant no. 11230506852009, Lachenalia aloides var. quadricolor (Jacq.) Engl. from Islamic Azad University (Boroujerd Branch) for Basic Re- 2n = 14, CHN. South Africa, Western Cape, 3318 (Cape search (IAUB). Town): Darling (-AD), 1983, J.J. Spies 7062 (BLFU).

E21 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

Lachenalia aloides var. vanzyliae W.F. Barker Lachenalia elegans var. suaveolens W.F. Barker 2n = 28, CHN. South Africa, Western Cape, 3318 (Cape 2n = 28, CHN. South Africa, Western Cape, 3219 (Wuppertal): Town): De Rust (-BD), 1983, J.J. Spies 7063 (BLFU). Cedarberg (-AC), Jan 2004, J.J. Spies 8165 (BLFU).

Lachenalia arbuthnotiae W.F. Barker Lachenalia fistulosa Baker 2n = 14, n = 7, CHN. South Africa, Western Cape, 3418 2n = 14, CHN. South Africa, Western Cape, 3319 (Worcester): (Simons Town): Steenbras River Mouth (-BB), 1983, J.J. Spies Klein Drakenstein (-CC), Jan 2004, J.J. Spies 8136 (BLFU); South 8054 (BLFU). Africa, Western Cape, 3418 (Simons Town): Houtbaai (-AB), J.J. Spies 6936 (1986), 8056 (1983), 8063 (1986) (BLFU), Dassenberg Lachenalia attenuata W.F. Barker ex G.D. Duncan (-AB), 1983, J.J. Spies 8057 (BLFU); South Africa, Western Cape, 2n = 14, CHN. South Africa, Northern Cape, 3220 (Suther- 3419 (Caledon): Caledon (-AB), 1983, J.J. Spies 8058 (BLFU); land): Agterkop (-AA), Jan 2004, J.J. Spies 8164 (BLFU). South Africa, Western Cape, 3419 (Caledon): Riviersonderend (-BB), 1983, J.J. Spies 8060 (BLFU). Lachenalia bolusii W.F. Barker 2n = 18, CHN. South Africa, Western Cape, 3219 (Wuppertal): Lachenalia hirta (Thunb.) Thunb. Bidouw (-AB), Jan 2004, J.J. Spies 7987 (BLFU). 2n = 22, CHN. South Africa, Western Cape, 3218 (Clanwil- liam): 3 km from Elandsbaai (-AD), 1983, J.J. Spies 6858 (BLFU); Lachenalia bulbifera (Cirillo) Engl. South Africa, Western Cape, 3219 (Wupppertal): Between Pakhuis 2n = 28, CHN. South Africa, Western Cape, 3419 (Caledon): Pass and Bidouw, Clanwilliam district (-AA), 1985, J.J. Spies 6859 Pearly Beach (-CB), 1987, J.J. Spies 7064 (BLFU). (BLFU); South Africa, Western Cape, 3219 (Wupppertal): 10 km north of Citrusdal (-CA), 1985, J.J. Spies 7381 (BLFU). Lachenalia carnosa Baker 2n = 16, CHN. South Africa, Northern Cape, 2917 (Spring- Lachenalia juncifolia Baker bok): Springbok (-DB), 1992, J.J. Spies 6998 (BLFU); South Af- 2n = 22, CHN. South Africa, Western Cape, 3319 (Worcester): rica, Northern Cape, 3018 (Kamiesberg): Road between Garies and Rawsonville (-CB), Jan 2004, J.J. Spies 8166 (BLFU). Leliefontein (-AB), 1986, J.J. Spies 6996. Lachenalia liliflora Jacq. n = 8+0–2B, CHN. South Africa, Northern Cape, 3018 (Ka- 2n = 16, CHN. South Africa, Western Cape, 3318 (Cape miesberg): Road between Garies and Leliefontein (-AB), 1986, J.J. Town): Stellenbosch (-DD), Jan 2004, J.J. Spies 8163 (BLFU). Spies 6934 (BLFU). Lachenalia mathewsii W.F. Barker Lachenalia comptonii W.F. Barker 2n = 14, CHN. South Africa, Western Cape, 3217 (Vreden- 2n = 20, CHN. South Africa, Northern Cape, 3220 (Suther- burg): Vredenburg (-DD), Jan 2004, J.J. Spies 8150 (BLFU). land): Komsberg (-DB), J.J. Spies 8135 (BLFU). Lachenalia maximiliani Schltr. ex W.F. Barker Lachenalia contaminata Aiton 2n = 16, CHN. South Africa, Western Cape, 3219 (Wuppertal): 2n = 14, CHN. South Africa, Western Cape, 3319 (Worcester): Bidouw (-AB), Jan 2004, J.J. Spies 8153 (BLFU). Rawsonville (-CB), Jan 2004, J.J. Spies 7989 (BLFU). Lachenalia mediana Jacq. Lachenalia doleritica G.D. Duncan 2n = 18, n = 9, CHN. South Africa, Western Cape, 3319 2n = 18, CHN. South Africa, Northern Cape, 3120 (Williston): (Worcester): Brandwag (-AC), 1996, J.J. Spies 7086 (BLFU); South Middelpos (-CC), Jan 2004, J.J. Spies 7985 (BLFU). Africa, Western Cape, 3319 (Worcester): Ceres (-AC), 1996, J.J. Spies 7087 (BLFU). Lachenalia elegans W.F. Barker 2n = 28, n = 14, CHN. South Africa, Northern Cape, 3119 Lachenalia mediana var. rogersii (Bak.) W.F. Barker (Calvinia): On top of Vanrhyns Pass (-AC), 1983, J.J. Spies 8014, n = 13, CHN. South Africa, Western Cape, 3318 (Cape Town): Jun 1992, 8020, 8021 (BLFU); South Africa, Northern Cape, 3119 Darling (-AD), 1986, J.J. Spies 7079 (BLFU). (Calvinia): Nieuwoudtville District (-AC), 1988, J.J. Spies 8018 (BLFU); South Africa, Western Cape, 3118 (Vanrhynsdorp): On Lachenalia mutabilis Sweet top of Gifberg (-DC), 1985, J.J. Spies 8015, 8016 (BLFU); South 2n = 12, n = 6, CHN. South Africa, Northern Cape, 3017 Africa, Western Cape, 3218 (Clanwilliam): Pakhuis Pass (-BB), (Hondeklipbaai): Soebatsfonteinroad (-BA), 1992, J.J. Spies 8001 1996, J.J. Spies 8022, 8023 (BLFU); South Africa, Western Cape, (BLFU); South Africa, Northern Cape, 3017 (Hondeklipbaai): 3319 (Worcester): Karoopoort (-BC), 1996, J.J. Spies 8025, 8026, 20 km from Kamieskroon, Leliefonteinroad (-BB), 1994, J.J. 8027 (BLFU). Spies 8007, 8008 (BLFU); Western Cape, 3318 (Cape Town): Langebaan Nature Reserve, Potsberg (-AA), 1992, J.J. Spies 8006 2n = 42, n = 21, CHN. South Africa, Northern Cape, 3119 (BLFU). (Calvinia): Calvinia (-AC), 1992, J.J. Spies 8019 (BLFU); South Africa, Western Cape, 3319 (Worcester): Karoopoort (-BC), 1996, 2n = 14, n = 7, CHN. South Africa, Northern Cape, 3017 J.J. Spies 8024 (BLFU). (Hondeklipbaai): 20 km from Kamieskroon, Leliefonteinroad (-BB), 1994, J.J. Spies 8000 (BLFU); South Africa, Western Cape, Lachenalia elegans W.F. Barker var. elegans 3218 (Clanwilliam): 5 km from Clanwilliam on N7 to Citrusdal 2n = 14, CHN. South Africa, Western Cape, 3219 (Wuppertal): (-BB), 1996, J.J. Spies 8009, 8010, 8011, 8012, 8013 (BLFU); South Wuppertal (-AC), Jan 2004, J.J. Spies 8152 (BLFU). Africa, Western Cape, 3218 (Clanwilliam): between Veldtdrift and Aurora (-CC), 1992, J.J. Spies 8003 (BLFU); South Africa, West-

E22 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

ern Cape, 3218 (Clanwilliam): Nuwerusdorp (-DC), 1992, J.J. Spies Lachenalia rubida Jacq. 8002 (BLFU); South Africa, Western Cape, 3318 (Cape Town): 2n = 14, CHN. South Africa, Western Cape, 3418 (Simons Langebaan Nature Reserve, Potsberg (-AA), 1992, J.J. Spies 8004, Town): 60 km from Faure turn off, Cape Town area (-BB), 1985, 8005 (BLFU). J.J. Spies 7093 (BLFU); South Africa, Western Cape, 3418 (Simons Town): Strand (-BB), Jan 2004, J.J. Spies 8156 (BLFU). Lachenalia namaquensis Schltr. ex W.F. Barker 2n = 16, n = 8, CHN. South Africa, Northern Cape, 2917 Lachenalia splendida Diels (Steinkopf): Springbok, (-BD), 1983, J.J. Spies 6969, 6970 2n = 16, n = 8, CHN. South Africa, Western Cape, 3118 (Vre- (BLFU). dendal): Vanrhynsdorp, (-CB), 1986, J.J. Spies 6977 (BLFU); South Africa, Western Cape, 3218 (Clanwilliam): Soutpanskoppies (-AB), Lachenalia orchioides (L.) Aiton 1983, J.J. Spies 6976 (BLFU). 2n = 14, n = 7, CHN. South Africa, Western Cape, 3218 (Clan- william): Pakhuis Pass (-BB), 1983, J.J. Spies 8086 (BLFU); South Lachenalia unifolia Jacq. Africa, Western Cape, 3318 (Cape Town): Stellenbosch (-DD), 1983, 2n = 22, n = 11, CHN. South Africa, Northern Cape, 3017 J.J. Spies 8084 (BLFU); South Africa, Western Cape, 3318 (Cape (Hondeklipbaai): On the way to Soebatsfontein in the Kam- Town): Helshoogte (-DD), 1983, J.J. Spies 8085 (BLFU); South Af- mieskroon district (-BA), 1985, J.J. Spies 6866 (BLFU); South Af- rica, Western Cape, 3318 (Cape Town): Klapmuts (-DD), J.J. Spies rica, Northern Cape, 3017 (Hondeklipbaai): Soebatsfonteinroad, 8089 (1990), 8090 (1992), 8092 (1992), 8093 (1992), 8095 (1992) 5 km from Kamieskroon (-BB), 1992, J.J. Spies 6878, 6879, 6881 (BLFU); South Africa, Western Cape, 3319 (Worcester): near Tul- (BLFU); South Africa, Western Cape, 3218 (Clanwilliam): Clanwil- bach (-AC), 1996, J.J. Spies 8071 (BLFU); South Africa, Western liam Botanical Gardens (-BB), 1983, J.J. Spies 6864 (BLFU); South Cape, 3319 (Worcester): Ceres Peak (-AD), 1996, J.J. Spies 8098, Africa, Western Cape, 3218 (Clanwilliam): Paleisheuwel (-BC), 8099 (BLFU); South Africa, Western Cape, 3319 (Worcester): 1983, J.J. Spies 6861 (BLFU); South Africa, Western Cape, 3219 Brandvlei Dam (-CB), 1996, J.J. Spies 8100, 8101 (BLFU); South (Wuppertal): 6 km from Wuppertal to Bidouw (-AC), 1983, J.J. Africa, Western Cape, 3319 (Worcester): along road to Ceres (-CB), Spies 6889 (BLFU); South Africa, Western Cape, 3318 (Cape 1992, J.J. Spies 8068, 8069 (BLFU); South Africa, Western Cape, Town): Darling (-AD), 1986, J.J. Spies 6867 (BLFU); South Africa, 3319 (Worcester): Du Toitskloof (-CB), J.J. Spies 8075 (1996), 8091 Western Cape, 3318 (Cape Town): Voorberg jail, Porterville (-BB), (1992) (BLFU); South Africa, Western Cape, 3319 (Worcester): 1988, J.J. Spies 6872 (BLFU); South Africa, Western Cape, 3318 Goudini (-CB), 1996, J.J. Spies 8073, 8079, 8080 (BLFU); South (Cape Town): Jonkershoek forestry station (-DD), 1990, J.J. Spies Africa, Western Cape, 3319 (Worcester): Rawsonville (-CB), 1996, 6874, 6891 (BLFU); South Africa, Western Cape, 3318 (Wuppertal): J.J. Spies 8072, 8074, 8082, 8083 (BLFU); South Africa, Western road between Klawer and Clanwilliam (-BB), 1992, J.J. Spies 6898 Cape, 3421 (Riversdale): Kanonkop (-BD), 1992, J.J. Spies 8094, (BLFU); South Africa, Western Cape, 3319 (Worcester): Witzen- 8097 (BLFU). berg (-AC), 1983, J.J. Spies 6863 (BLFU); South Africa, Western Cape, 3319 (Worcester): North of Gouda (-AC), 1988, J.J. Spies Lachenalia orthopetala Jacq. 6871 (BLFU); South Africa, Western Cape, 3319 (Worcester): on the 2n = 16, CHN. South Africa, Western Cape, 3319 (Worcester): Ceres side of Bainskloof Pass (-CA), 1992, J.J. Spies 6892 (BLFU). Rawsonville (-CB), Jan 2004, J.J. Spies 7986 (BLFU). Lachenalia cf. unifolia Jacq. Lachenalia pallida Aiton 2n = 22, CHN. South Africa, Western Cape, 3318 (Cape 2n = 16, CHN. South Africa, Western Cape, 3318 (Cape Town): Tienie Versfeld Nature Reserve, Darling (-AD), 1992, J.J. Town): Hopefield (-AB), Jan 2004, J.J. Spies 8157 (BLFU); South Spies 6899, 6902 (BLFU). Africa, Western Cape, 3318 (Cape Town): Potsdam (-DC), 1991, J.J. Spies 7089 (BLFU). Lachenalia violacea Jacq. n = 7, CHN. South Africa, Northern Cape, 3017 (Hondeklip- Lachenalia peersii Marloth ex W.F. Barker baai): Road between Garies and Kamieskroon (-BB), 1992, J.J. 2n = 14, CHN. South Africa, Western Cape, 3418 (Simons Spies 7101 (BLFU); South Africa, Northern Cape, 3017 (Hon- Town): Betty’s Bay (-BD), Jan 2004, J.J. Spies 8159 (BLFU). deklipbaai): Soebatsfontein road, 8 km from Kamieskroon (-BB), 1992, J.J. Spies 7104 (BLFU). Lachenalia purpureo-caerulea Jacq. 2n = 16, CHN. South Africa, Western Cape, 3218 (Clanwil- liam): Piekenierskloof Pass (-DB), 1983, J.J. Spies 6939 (BLFU). Huan Hang1, Min Zheng1,2 & Rui-Liang Zhu1

Lachenalia pusilla Jacq. 1 Department of Biology, School of Life Science, East China 2n = 18, CHN. South Africa, Western Cape, 3218 (Clanwil- Normal University, 3663 Zhong Shan North Road, Shanghai liam): Piketberg (-DA), Jan 2004, J.J. Spies 8155 (BLFU). 200062, China. [email protected] (author for correspondence) Lachenalia reflexa Thunb. 2 School of Environmental Studies, China University of Geo- 2n = 14, CHN. South Africa, Western Cape, 3322 (Oudt- sciences, 388 Lumo Road, Wuhan 430074, China shoorn): Between Oudtshoorn and Mossel Bay (-CC), 1987, J.J. Spies 7091 (BLFU); South Africa, Western Cape, 3418 (Simons The methods for the present study are the same as those de- Town): Dassenberg (-AB), 1983, J.J. Spies 7090 (BLFU). scribed in Zheng & Zhu (2008, 2009).

Lachenalia rosea Andrews * First chromosome count for the species. 2n = 28, CHN. South Africa, Western Cape, 3419 (Caledon): ** New chromosome number for the species. Caledon (-AB), Jan 2004, J.J. Spies 8167, 8168 (BLFU). ▼ First chromosome number from a Chinese accession.

E23 Marhold (ed.) • IAPT/IOPB chromosome data 8 TAXON 58 (4) • November 2009: E1–E25

This study was supported by the National Natural Science n = 8 (♂), CHN. China, Zhejiang, Longquan, Fengyangshan Foundation of China (nos. 30825004, 30670155), the Program of Nature Reserve, 1,520 m, 29 Jul 2006, R.-L. Zhu & al. 2006072901 Shanghai Subject Chief Scientist (no. 08XD14016), and 211 Project. (HSNU).

FRULLANIACEAE Frullania moniliata (Reinw. & al.) Mont. (Fig. 1C) Frullania ericoides (Nees) Nees & Mont. n = 9. CHN. China, Zhejiang, Longquan, Fengyangshan Nature n = 8 (♂). CHN. China, Zhejiang, Linan, Tianmushan Na- Reserve, 1,480 m, 28 Jul 2006, R.-L. Zhu & al. 2006072801 (HSNU). ture Reserve, 900 m, 25 Apr 2006, R.-L. Zhu & al. 2006042502 (HSNU). Frullania motoyana Steph. ▼n = 9, CHN. China, Hainan, Changjiang, Bawangling Nature *Frullania fengyangshanensis R.L. Zhu & M.L. So (Fig. 1A, B) Reserve, 1,200 m, 2 Dec 2003, R.-L. Zhu & al. 2003120206 (HSNU). n = 9 (♀), CHN. China, Zhejiang, Longquan, Fengyangshan Nature Reserve, 1,480 m, 29 Jul 2006, R.-L. Zhu & al. 2006072907 Frullania muscicola Steph. (HSNU). n = 9, CHN. China, Zhejiang, Linan, Tianmushan Nature Re- serve, 900 m, 13 May 2004, R.-L. Zhu & al. 2004051309 (HSNU); China, Zhejiang, Tonglu, Holiday Village, 800 m, 3 Jul 2004, R.-L. Zhu 20040703 (HSNU).

Frullania tamarisci (L.) Dumort. n = 9, CHN. China, Guangxi, Xingan, Maoershan Nature Reserve, 2,100 m, 3 Sep 2004, R.-L. Zhu & al. 2004-9307 (HSNU).

GEOCALYCACEAE Heteroscyphus coalitus (Hook.) Schiffn. n = 9, CHN. China, Zhejiang, Longquan, Fengyangshan Nature Reserve, 1,240 m, 31 Jul 2006, R.-L. Zhu & al. 2006073102 (HSNU).

Heteroscyphus planus (Mitt.) Schiffn. (Fig. 1D) ▼n = 9, CHN. China, Zhejiang, Shengzhou, Guimen, 350 m, 3 May 2004, R.-L. Zhu 20040503004 (HSNU); China, Zhejiang, Longquan, Fengyangshan Nature Reserve, 1,240 m, 31 Jul 2006, R.-L. Zhu & al. 2006073105 (HSNU).

Lophocolea bidentata (L.) Dumort. (Fig. 1E, F) ▼n = 18, CHN. China, Zhejiang, Longquan, Fengyangshan Nature Reserve, 1,460 m, 30 Jul 2006, R.-L. Zhu & al. 2006073002 (HSNU).

▼n = 9, CHN. China, Hainan, Changjiang, Bawangling Na- ture Reserve, 1,000 m, 26 Nov 2003, R.-L. Zhu & al. 2003112603 (HSNU).

Lophocolea ciliolata (Nees) Gottsche ▼n = 9, CHN. China, Hainan, Lingshui, Diaoluoshan Na- ture Reserve, 1,060 m, 4 Sep 2005, R.-L. Zhu & al. 2005090405 (HSNU); China, Hainan, Changjiang, Bawangling Nature Reserve, 1,050 m, 28 Nov 2003, R.-L. Zhu & al. 2003112802 (HSNU). Fig. 1. A & B, Frullania fengyangshanensis; A, mitotic metaphase, n = 9(♀) (R.-L. Zhu & al. 2006072907); B, HAPLOMITRIACEAE mitotic metaphase, n = 8 (♂) (R.-L. Zhu & al. 2006072901). Haplomitrium mnioides (Lindb.) R.M. Schust. C, Frullania moniliata, mitotic metaphase, n = 9 (R.-L. n = 9, CHN. China, Guangxi, Xingan, Maoershan Nature Zhu & al. 2006072801). D, Heteroscyphus planus, mitotic Reserve, 1,170 m, 7 Sep 2004, R.-L. Zhu & al. 20049706 (HSNU). metaphase, n = 9 (R.-L. Zhu 20040503004). E–F, Lophocolea bidentata; E, mitotic metaphase, n = 18 (R.-L. Zhu & al. JACKIELLACEAE 2006073002); F, mitotic metaphase, n = 9 (R.-L. Zhu & al. Jackiella javanica Schiffn. (Fig. 1G) 2003112603). G, Jackiella javanica, mitotic metaphase, n = **▼n = 8, CHN. China, Hainan, Lingshui, Diaoluoshan Nature 8 (R.-L. Zhu & al. 2005090103). H, Hattoria yakushimensis, Reserve, 920 m, 1 Sep 2005, R.-L. Zhu & al. 2005090103 (HSNU); mitotic metaphase, n = 9 (R.-L. Zhu & al. 20049509). I, Singapore, Upper Picrce Canal, 01°22′51.51″ N, 103°48′54.31″ E, Nardia assamica; mitotic metaphase, n = 9 (R.-L. Zhu & al. 24 Jan 2007, R.-L. Zhu 20070124-12 (HSNU). 2006072910). J, Bazzania tridens; mitotic metaphase, n = 10 (R.-L. Zhu & al. 2004051312). K, Plicanthus birmensis, JUNGERMANNIACEAE mitotic metaphase, n = 9 (R.-L. Zhu & al. 20049307). L, Hattoria yakushimensis (Horik.) R.M. Schust. (Fig. 1H) Porella obtusata var. macroloba, mitotic metaphase, n = 8 ▼n = 9, CHN. China, Guangxi, Xingan, Maoershan Nature (R.-L. Zhu & al. 2006072814). Scale bar = 5 μm. Reserve, 2020m, 5 Sep 2004, R.-L. Zhu & al. 20049509 (HSNU).

E24 TAXON 58 (4) • November 2009: E1–E25 Marhold (ed.) • IAPT/IOPB chromosome data 8

Nardia assamica (Mitt.) Amakawa (Fig. 1I) n = 8, CHN. China, Zhejiang, Shengzhou, Guimen, 350 m, 11 ▼n = 8+m, CHN. China, Zhejiang, Longquan, Fengyangshan Feb 2005, R.-L. Zhu & al. 200502114 (HSNU). Nature Reserve, 1,530 m, 29 Jul 2006, R.-L. Zhu & al. 2006072910 (HSNU). Porella densifolia (Steph.) S. Hatt. n = 8, CHN. China, Zhejiang, Linan, Tianmushan Nature Re- LEPIDOLAENACEAE serve, 1,000 m, 26 Apr 2006, R.-L. Zhu & al. 2006042606 (HSNU). Trichocoleopsis sacculata (Mitt.) S. Okamura ▼n = 9, CHN. China, Zhejiang, Longquan, Fengyangshan Porella obtusata var. macroloba (Steph.) S. Hatt. & M.X. Nature Reserve, 1,510 m, 30 Jul 2006, R.-L. Zhu & al. 2006073007 Zhang (Fig. 1L) (HSNU). ▼n = 8, CHN. China, Zhejiang, Longquan, Fengyangshan Nature Reserve, 1,140 m, 28 Jul 2006, R.-L. Zhu & al. 2006072814 LEPIDOZIACEAE (HSNU). Bazzania tridens (Reinw. & al.) Trevis. (Fig. 1J) **n = 10, CHN. China, Zhejiang, Linan, Tianmushan Na- Porella ulophylla (Steph.) S. Hatt. ture Reserve, 900 m, 12 May 2004, R.-L. Zhu & al. 2004051312 n = 8, CHN. China, Zhejiang, Linan, Tianmushan Nature Re- (HSNU); China, Zhejiang, Linan, Tianmushan Nature Reserve, serve, 900 m, 25 Apr 2006, R.-L. Zhu & al. 2006042503 (HSNU); 1,000 m, 13 May 2004, R.-L. Zhu & al. 2004051313 (HSNU); China, Zhejiang, Linan, Tianmushan Nature Reserve, 950 m, 28 China, Hainan, Changjiang, Bawangling Nature Reserve, 1,150 m, Apr 2006, R.-L. Zhu & al. 2005042801 (HSNU). 27 Nov 2003, R.-L. Zhu & al. 2003112701 (HSNU). Literature cited LOPHOZIACEAE Zheng, M. & Zhu, R.-L. 2008. Cytological observation on Plicanthus birmensis (Steph.) R.M. Schust. (Fig. 1K) some Chinese species of Plagiochila (Plagiochilaceae, n = 9, CHN. China, Guangxi, Xingan, Maoershan Nature Marchantiophyta). Cryptog. Bryol. 29: 39–47. Reserve, 2,100 m, 3 Sep 2004, R.-L. Zhu & al. 20049307 (HSNU). Zheng, M. & Zhu, R.-L. 2009. Karyological studies on some species of Radula (Radulaceae, Jungermanniopsida, PORELLACEAE Marchantiophyta). Nova Hedwigia 88: 229–244. Porella chinensis (Steph.) S. Hatt.

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