Index

cases code) from numbers The hyphen separates the names of taxa at the left side (in most followed by a

and the side. Names of followed code of letters of pages, figures plates at right accepted species are by a

with their in this A B: Ch: and numbers corresponding sequence paper. means , Bracisepalum,

Chelonistele, E: Entomophobia, G: Geesinkorchis, and N: Nabaluia. Synonyms are followed by '='and

An 'Excl.' excluded - subsequently by the code of the species to which they belong. refers to species.

asterisk indicates the where the is 'fig.' refers to a line-drawing; to a coloured page taxon described; 'plate'

Two unnumbered with their number. plate. maps are given page

Acriopsidinae Dressier — 1 Chelonanthera Blume - 23, 24

Acriopsis Reinw. ex Blume - 1, 2, 3*, 4 map clypeata (Lindley) Pfitzer - 23

= — annamensis Guill. ?A5c 16 gibbosa Blume - 23

= — = annamica Finet ?A5c 15, 16 sulphurea Blume ChlOa - 23, 37

= - borneensis Ridley A2b 7 ventricosa Blume - 23

carrii Holttum A1 - 2, 3, 5*, fig. 2 Chelonistele Pfitzer - 17, 23, 24*, 25 map, 43, 47

densiflora Lindley A2 — 2, 3, 5, 6*, 7 amplissima (Ames & Schweinf.) Carr Chi -

var. borneensis (Ridley) Minderhoud & 26*, fig. 10, plate 2d

de Vogel A2b - 6, 7*, fig. 3f-g apiculata (Lindley) Pfitzer Excl. — 23, 40

var. densiflora A2a — 6*, fig. 3a-e biflora (Par. & Rchb. f.) Pfitzer Excl. - 23, 40

= floribunda Ames A5b - 13 brevilamellata (J.J. Smith) Carr Ch2 - 24, 26,

gracilis Minderhoud & de Vogel A3 — 1, 3, 5, 28*, fig. 11

= 7*, fig. 4, plate la clemensii (Ames) Carr N2 - 47, 49

= = griffithii Rchb. f. A5a - 11 crassifolia Carr Chi Ob — 39

= = harai Tuyama ?A5a — 12, 13 cuneata (J.J. Smith) Carr ChlOa - 38, 39

- indica Wight A4 3, 5, 8*, fig. 5, plate lb dentifera de Vogel Ch3 - 26, 29*, fig. 12

= insulari-sylvatica Fukuyama ?A5a - 12, 13 ingloria (J.J.Smith) Carr Ch4 - 18, 26, 30*,

javanica Reinw. ex Blume A5 — 3,5,9*, 13,15, fig. 13, plate 3a

= 16 keithiana W.W. Smith N2 - 40, 47, 49

var. auriculata Minderhoud & de Vogel kinabaluensis (Rolfe) de Vogel Ch5 — 26, 31*,

A5c - 1, 3, 10, 14*, 16, fig. 6 fig. 14

= var. floribunda (Ames) Minderhoud & de kutaiensis (J.J. Smith) Carr ChlOa — 38, 39

Vogel A5b - 10, 13*, fig. ld-e (p. 11) lamellulifera Carr Ch6 — 26, 32*, 47, fig. 15

var. javanica A5a — 3, 10*, 13, fig. la—c, lurida (L. Linden & Cogn.) Pfitzer Ch7 - 26,

f-h (p. 11), plate lc, Id 33*

= var. nelsoniana (Bailey) J.J. Smith A5a - 11 var. grandiflora de Vogel Ch7b - 34*, fig.

= javanica (non Reinw. ex Bl.) Blume A5c — 14 16j-k, plate 3c

= - javanica (non Reinw. ex Bl.) Schltr. A5b - 13 var. lurida Ch7a 34*, fig. 16a-i, plate 3b

— = latifolia Rolfe Excl. 16 perakensis (Rolfe) Ridley ChlOa - 23,38,39,

= nelsoniana Bailey A5a — 11 43

= = var. pallidifloraSchltr. A5a - 12 pinniloba (J.J. Smith) Carr Chl0a-38, 39

= - = papuana Kraenzlin A5a 12 pusilla (Ridley) Ridley ChlOa - 37, 39

= — philippinensis Ames A5a 12 ramentacea J.J.Wood Ch8 - 26, 35*, fig. 17

= ?A5a - picta Lindley 11 richardsii Cart Ch9 - 26, 35*, fig. 18

=

- purpurea Ridley A2a 7 sulphurea (Blume) Pfitzer ChlO - 24, 36*, 43

- ridleyi Hooker f. A6 2, 3, 5, 15*, fig. 7, plate var. crassifolia (Carr) de Vogel Chi0b - 18,

2a, 2b 26, 39*, fig. 19e-g, fig. 20e

=

sumatrana Schltr. ?A5a -11,13 var. sulphurea ChlOa — 18, 26, 37*, fig.

19a-d, fig. 20a-d, f, g, plate 3d,

Basigyne J.J. Smith - 19 cover

Smith — Bracisepalum J.J. 17, 19* tenuiflora (Ridley) Pfitzer Excl. - 40

densiflorum de Vogel Bl - 19, 20*, fig. 8, unguiculata Carr Chll - 26, 39*, fig. 21

plate 2c vermicularis (J.J. Smith) Kraenzlin Excl. — 40

selebicum J.J.Smith B2 — 18, 19, 20*, fig. 9 Chelonostele auct. non Pfitzer: Ridley - 43, 44

52 Orchid Monographs 1 (1986) — — 47 Geesinkorchis de 43* Coelogyne Lindley 19, 23, 24,43, Vogel 17,

- - 44*, 23, sect. Chelonistele J.J. Smith 23, 24 alaticallosa de Vogel G1 43, fig.

sect. Longifoliae Pfitzer - 43 plate 4a, 4b

G2 — subgen. Chelonistele (Pfitzer) Butzin - 24 phaiostele (Ridley) de Vogel 18, 40, 43,

- 4c amplissima Ames & Schweinf. Chi — 26, 28 44, 45*, fig. 24, plate

var. schweinfurthiana J.J. Smith = Chi —

Nabaluia Ames — 47* 26, 28 17, 23, 43,

- 39 de N1 — 49, fig. 25, beyrodtianaSchltr. = ChlOa 23, 38, angustifolia Vogel 47, 48*,

biflora Par. & Rchb. f. — 23 plate 4d

- 26 bilamellata Lindley - 47 clemensii Ames N2 40,47,48, 49*, fig. - = non Ames 48 brevilamellata J.J. Smith Ch2 — 23, 28 clemensii auct. Ames:

= de - 27 crassifolia (Carr) Masamune ChlOb - 39 exaltata Vogel N3 48, 50*, fig.

= croockewitii Teysm. & Binnend. ChlOa —

— 23 23,38,39 Panisea Lindley

= - 23 cuneata J.J. Smith ChlOa — 38 apiculata Lindley

= - 40 decipiens Sander ?Chl0a - 38 uniflora (Lindley) Lindley

- 47 gibbosa(Blume) Rchb. f. — 23 Pholidota W.J. Hooker 17, 23, 41,43,

= de Vriese - 43 ingloria J.J. Smith Ch4 — 30 gibbosa (Blume)

= J. Hooker — 43 kutaiensis J.J. Smith ChlOa - 38 imbricata W.

= = - kinabaluensis Ames El - 41 lamellulifera (Carr) Masamune Ch6 32

— = - Rchb. f. 47 lurida L. Linden & Cogn. Ch7 23, 34 nervosa (Blume) 41, 43,

= - 38 = perakensis Rolfe ChlOa - 23, 38 pusilla (Ridley) Kraenzlin ChlOa

= Smith Ch5 - 31 = phaiostele G2 - 43,45 sigmatochilus(Rolfe) J.J.

= - pinnilobaJ.J. Smith ChlOa — 38 sulcata J.J. Smith 43

= = - pusilla Ridley ChlOa - 23, 37 triloba J.J. Smith G2 45

= - 43 richardsii (Carr) Masamune Ch9 - 35 ventricosa (Blume) Rchb. f.

— 23 = ridleyana Schltr. G2 — 43,45 Pleione D. Don

= = & O. Kuntze sarawakensis Schltr. Ch7a - 34 croockewitii (Teysm. Binnend.)

= - ChlOa - 38 sulphurea (Blume) Rchb. f. ChlOa 23, 37 23,

= O. Kuntze ChlOa — 37 tenuiflora Ridley - 23 sulphurea (Blume) 23,

= unguiculata(Carr) Masamune Chll — 39

- - Rolfe 24 ventricosa (Blume) Rchb. f. 23 Sigmatochilus

= kinabaluensis Rolfe Ch5 — 31 — 1 24,

=

- Cyrtopodiinae — 1, 2 Spathoglottis ?trivalvis Lindley A5a 11

Dendrochilum Blume - 19 Thecopus —1,2

Thecostele — 1,2

— Entomophobia de Vogel - 17, 41* alata (Roxb.) Par. & Rchb. f. 16

kinabaluensis (Ames) de Vogel El - 41*, Thecostelinae -1,2

fig. 22

- Epidendreae — 1 Vandeae 1

— - 1 Vandoideae 1

Orchid Monographs 1 (1986) 53 Legends of the figures

For Fig. 1, see page 11.

Holttum. b. flower d. and column Fig. 2. Acriopsis carrii a. Habit; flower; c. analysis; lip (a.

Carr 135; b-d. SF 10254).

flower Fig. 3. Acriopsis densiflora Iindley var. densiflora. a. Habit; b. flower; c. analysis; d. lip and column; e. lip. — Acriopsis densiflora Lindley var. bomeensis (Ridley) Minderhoud&

de f. Habit; and column Niel b-d. Herb. Paris e. van der Est Vogel. g. lip (a. van 3718, s.n.;

s.n.; f-g. Collenette 29/79).

Fig. 4. Acriopsis gracilis Minderhoud & de Vogel. a. Flower; b. flower analysis; c. lip and column (a—d. Giles 620).

indica b. flower column Fig. 5. Acriopsis Wight, a. Habit; analysis; c. flower; d. lip and

Weber b—d. Seidenfaden& Smitinand (a. s.n.; 8345).

Blume auriculata Minderhoud de Fig. 6. Acriopsis javanica Reinw. ex var. & Vogel. a. Habit; b. flower; c. flower analysis; d. lip and column; e. cross-section through lip and column (a—d.

Leiden, cult. Hort. (Roos & Franken) 20788; e. Mason 546).

Fig. 7. Acriopsis ridleyi Hooker f. a. Habit; b. flower; c. flower analysis; d. lip and column; e.

fruit (a. Burkill 3331;b-e. Bridson 353).

Fig. 8. Bracisepalum densiflorum de Vogel. a. Habit; b. bract; c. flower; d. flower analysis; e. lip and column; f. column (a—f. Leiden, cult. Hort. (de Vogel) 20313).

Smith, flower Fig. 9. Bracisepalum selebicum J.J. a. Habit; b. bract; c. flower; d. analysis; e. lip and column; f. colum (a—f. Leiden, cult. Hort. (de Vogel) 20446).

flower Fig. 10. Chelonisteleamplissima (Ames & Schweinf.) Carr. a. Habit; b. flower; c. anal-

outline Chai ysis; d. column; e. cross-section through ovary (a. & c—e. Nooteboom& 2247;

b. Singapore, cult. Hort. 11.111.66 (Kew ale. 31926)).

Fig. 11. Chelonistele brevilamellata(J.J. Smith) Carr. a. Habit; b. flower analysis; c. backside

of the lip; d. column (a—d. Endert 4427).

Fig. 12. Chelonistele dentifera de Vogel. a. Habit; b. flower analysis; c. lip and column; d.

column; e. lip (a. & c—e. S 21954; b. Burtt B. 11375).

Carr. b. flower d. col- Fig. 13. Chelonistele ingloria (J.J.Smith) a. Habit; flower; c. analysis;

cult. Hort. Martin umn (a. & c-d. Endert 3972; b. Leiden, (Burtt & 5215) 22064).

de b. flower Fig. 14. Chelonistele kinabaluensis (Rolfe) Vogel. a. Habit; flower; c. analysis;

the lobes Clemens e—f. Clemens d. lip and column; e. column; f. column, spread (a—d. 51192;

50658).

Carr. flower d. col- Fig. 15. Chelonistele lamellulifera a. Habit; b. flower; c. analysis; lip; e.

umn,left lobes spread (a. & d. Synge S. 521; b-c. & e. Burtt & Martin B. 5405).

54 Orchid Monographs 1 (1986) Fig. 2. Holttum.

55 Orchid Monographs 1 (1986) 16. Chelonistele lurida Linden & Pfltzer Fig. (L. Cogn.) var. lurida. a. Habit; b. flower; c.

flower analysis; d. column, lobes spread; e. anther; f. pollinia; g. flower analysis; h. lip and

column; i. column, lobes spread. — Chelonistele lurida var. grandiflora de Vogel. j. Flower

analysis; k. column, lobes spread (a. Clemens s.n., 7-1-1932; b-f. SAN (Sinanggul) 38318;

g—i. Bruxelles, cult. Hort. (Herb. Cogniaux); j-k. Burtt & Woods B. 2169).

Fig. 17. Chelonistele ramentacea J.J.Wood, a. Habit; b. flower analysis; c. column (a—c.

Burtt & Woods B. 2192).

Fig. 18. Chelonistele richardsii Carr. a. Habit; b. flower; c. flower analysis; d. lip and column;

Richards e. column, lobes spread (a-e. S. 59).

Pfltzer Fig. 19. Chelonistele sulphurea (Blume) var. sulphurea. a. Habit; b. flower; c. flower

- Chelonistele analysis; d. lip and column. sulphurea var. crassifolia (Carr) de Vogel. e. Habit;

f. flower & Blume H.L.B. b. & d. cult. flower; g. analysis (a. c. s.n., 902,322-1084; Leiden, Hort. (Roos& Franken) 21133;e-g. Clemens 27169).

Fig. 20. Variation in shape of lip and column, in different specimens of Chelonistele sul- Pfltzer phurea (Blume) var. sulphurea. a. Leiden, cult. Hort. (Roos & Franken) 21133, Java

cult. (‘sulphurea’) b. Kew, Hort. s.n. (IX-1904), Malay Peninsula (‘pusilla’);I; c. Bogor, cult.

Hort. (Kuyper) 129; d. Endert 4524, (‘pinniloba’), type); f. Endert 4429, Borneo

Endert - (‘kutaiensis’ ', type); g. 3879, Borneo {'cuneata ', type). Chelonistelesulphurea var.

crassifolia (Carr) de Vogel. e. Clemens 27169, Borneo ( ‘crassifolia’'■'< type).

Chelonistele Fig. 21. unguiculata (Carr) Masamune. a. Habit; b. bract; c. flower; d. flower

and f. base of lobes analysis; e. hp column; Up; g. column, spread (a-g. Synge S. 399).

Fig. 22 Entomophobia kinabaluensis (Ames) de Vogel. a. Habit; b. infructescence; c. bract;

flower f. from d. flower; e. analysis; lip above; g. column; h. lip and column; i. lip and col-

umn, part of the lip removed to show the callus; j. anther; k. pollinia (a—e. Clemens 30607;

f-k. Geesink 9039).

Fig. 23. Geesinkorchis alaticallosa de Vogel. a. Habit; b. bract; c. & d. flower; e. flower anal-

f. detail of of ysis; lip and column; g. backside lip; h. column; i. anther;j. pollinarium, two

pollinia removed; k. seed (a—k. Geesink 8965).

Fig. 24. Geesinkorchis phaiostele (Ridley) de Vogel. a. Habit; b. bract; c. & d. flower; e. fruit;

f. flower analysis; g. lip and column; h. column; i. anther; j. pollinarium, two pollinia re-

moved Hewitt b. Clemens (a. s.n., Mt Poe; 20398; c—j. Leiden, cult. Hort. (Burtt & Woods

B. 2667) 22090).

Fig. 25. Nabaluia angustifolia de Vogel. a. Habit; b. flower; c. flower analysis; d. Up and col-

detail of backside of f. & h. column & Clemens e-f. umn; e. & g. Up; (a-d. g-h. 27159;

Fuchs & CoUenette 21381).

Fig. 26. Nabaluia clemensii Ames. a. Habit; b. flower analysis; c. lip and column; d. detail of backside of Up; e. column (a-e. Moulton 103).

27.. Fig. Nabaluia exaltata de Vogel. a. Habit, detached young shoot and inflorescence; b. bract; c. flower; d. flower analysis; e. lip and column; f. detail of backside of lip; g. column

(a—g. Burtt & Martin B. 5259).

Orchid 1 Monographs (1986) 56 de & var. (Ridley) Minderhoud & Vogel. Fig. 3. Lindley var.

57 Orchid Monographs 1 (1986) Fig. 4. Minderhoud & de Vogel.

58 Orchid Monographs 1 (1986) Fig. 5. Wight.

59 Orchid Monographs 1 (1986) Fig. 6. Reinw. ex Blume vai. Minderhoud & de Vogel.

60 Orchid Monographs 1 (1986) f. Fig. 7 Hooker

61 Orchid Monographs 1 (1986) Fig. 8. de Vogel.

62 Orchid Monographs 1 (1986) Fig. 9. J. J. Smith.

63 Orchid Monographs 1 (1986) Fig. 10. (Ames & Schweinf.) Can.

64 Orchid Monographs 1 (1986) Fig. 11. (J.J.Smith) Carr.

Orchid Monographs 1 (1986) 65 Fig. 12. de Vogel

66 Orchid Monographs 1 (1986) Fig. 13. (J.J. Smith) Cair.

67 Orchid Monographs 1 (1986) Fig. 14. (Rolfe) de Vogel

68 Orchid Monographs 1 (1986) Fig. 15. Carr.

69 Orchid Monographs 1 (1986) Fig. 16. (L. Linden & Cogn.) Pfitzer vax. & var. de Vogel.

70 Orchid Monographs 1 (1986) Fig. 17. J.J.Wood.

71 Orchid Monographs 1 (1986) Fig. 18. Carr.

72 Orchid Monographs 1 (1986) var. (Can) de Vogel. Fig. 19. (Blume) Pfitzer var. &

73 Orchid Monographs 1 (1986) Fig. 20. Variation in shape of lip and column, in & var.

74 Orchid Monographs 1 (1986) Masamune. Fig. 21. (Carr)

75 Orchid Monographs 1 (1986) Fig. 22. (Ames) de Vogel.

76 Orchid Monographs 1 (1986) de Fig. 23. Vogel

77 OrcWd Monographs 1 (1986) Fig. 24. (Ridley) de Vogel.

78 Orchid Monographs 1 (1986) de Fig. 25. Vogel.

79 Orchid Monographs 1 (1986) Fig. 26. Ames.

80 Orchid Monographs 1 (1986) Fig. 27. de Vogel.

Orchid Monographs 1 (1986) 81 Legends of the plates

Plate 1 Plate 2

a. Acriopsis gracilis Minderhoud& de Vogel a. Acriopsis ridleyi Hooker f.

From Borneo, Sabah Singapore, cult. Hort., possibly from

Photograph A. Lamb or Borneo

Photograph J. Comber

b. Acriopsis indica Wight

From Java b. Acriopsis ridleyi Hooker f.

Photograph J. Comber Leiden, cult. Hort. (Roos & Franken)

20761, from Borneo, Kalimantan

Blume B. N. Kieft c. Acriopsis javanica Reinw. ex Photograph

var. javanica (collection Rijksherbarium, Leiden)

Leiden, cult. Hort. (Roos & Franken)

20741, from Sumatra c. Bracisepalum densiflorum de Vogel

Photograph B. N. Kieft Leiden, cult. Hort. (de Vogel) 20313,

(collection Rijksherbarium, Leiden) from Celebes

Photograph C.G.J.M. van Vliet

Blume d. Acriopsis javanica Reinw. ex (collection Rijksherbarium, Leiden)

var. javanica

Leiden, cult. Hort. 21370, from Aus- d. Chelonistele amplissima (Ames &

tralia Schweinfurth) Carr

Photograph B.N. Kieft From Borneo, Sabah

(collection Rijksherbarium, Leiden) Photograph J. Comber

Plate 3 Plate 4

a. Chelonistele ingloria (J.J. Smith) Carr a. Geesinkorchis alaticallosa de Vogel

Leiden, cult. Hort. (Burtt & Martin Bogor, cult. Hort. 981.VIII.62 (Darnaedi

B. 5215) 22064, from Borneo, & Waluyo), from Borneo, Kalimantan

Sarawak Photograph B. N. Kieft

Photograph B. N. Kieft (collection Rijksherbarium, Leiden)

(collection Rijksherbarium, Leiden)

b. Geesinkorchis alaticallosa de Vogel b. Chelonistele lurida (L. Linden & Cogn.) From Borneo, Sabah

Pfitzer var. lurida Photograph J. Comber

From Borneo, Sabah

Photograph J. Comber c. Geesinkorchis phaiostele (Ridley)

de Vogel c. Chelonistele lurida (L. Linden & Cogn.) Leiden, cult. Hort. (Burtt & Woods

Pfitzer var. grandiflora de Vogel B. 2667) 22090, from Borneo, Sabah

Lewis 336, from Borneo, Sarawak Photograph B. N. Kieft

Photograph G.P. Lewis (collection Kew) (collection Rijksherbarium, Leiden)

d. Chelonistele sulphurea (Blume) Pfitzer d. Nabaluia angustifolia de Vogel

var. sulphurea Kew, cult. Hort. (Brentnall 154), from

From Sumatra Borneo, Sabah

Photograph J. Comber Photograph W. Rossi (collection Kew)

82 Orchid Monographs 1 (1986) Plate 1

a. Minderhoud & de Vogel b. Wight

d. Reinw. Blume c. Reinw. ex Blume var ex var

Orchid Monographs 1 (1986) 83\1 Plate 2

a. Hooker f. b. Hooker f.

c. de Vogel d. (Ames& Schweinf.) Carr

84\2 Orchid Monographs 1 (1986J Plate 3

Carr b. Linden & Cogn.) Pfitzer a. (J.J. Smith) (L.

& Pfitzer d. (Bl.) Pfitzer var. c. (L. Linden Cogn.)

85\3 Orchid Monographs 1 (1986) Plate 4

a. de Vogel b. de Vogel

c. (Ridley) de Vogel d. de Vogel

86\4 Orchid Monographs 1 (1986)