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A New Sandy-Desert Subspecies of Megathymus Coloradensis (Megathymidae) from Extreme Northern Arizona
BULLETIN OF THE ALLYN MUSEUM Published by THE ALLYN MUSEUM OF ENTOMOLOGY Sarasota, Florida Number -- 17 12 APRIL 1974 A NEW SANDY-DESERT SUBSPECIES OF MEGATHYMUS COLORADENSIS (MEGATHYMIDAE) FROM EXTREME NORTHERN ARIZONA Ronald S.' and Dale Wielgus 3434 West Augusta Avenue, Phoenix, Arizona 85021 Recent descriptions in the genus Cercyonis (Satyridae) by Emmel & Emmel (1969; 1971) and Emmel & Mattoon (1972) have shown the existence of distinctive whitish races adapted to arid alkaline flats of the western deserts of the United States. Such environmental adaptation appears to extend to a population of Megathymus coloradensis Riley inhabiting a sandy-desert area of extreme northern Arizona. The discovery of the subspecies described herein was a result of our initial investigations into the biology of Megathymus streckeri (Skinner). The purpose of this paper is to describe the new population and to note its life history. This study considered the following subspecies: Megathymus coloradensis navajo Skinner, Megathymus coloradensis arizonae Tinkham, Megathymus coloradensis reubeni Stallings, Turner & Stallings, Megathymus coloradensis coloradensis Riley and Megathymus coloradensis browni Stallings & Turner. Megathymus coloradensis albasuffusa Ronald S. and Dale Wielgus, new subspecies. Female. Head: vertex grey, Palpus: white, with some black-tipped scales. Antenna: shaft white distad, ringed with black basad; club black. Thorax: grey dorsally with long greenish-brown hairs caudad, blackish ventrally. Abdomen: black dorsally, grey ventrally. -
Courtship and Oviposition Patterns of Two Agathymus (Megathymidae)
Journal of the Lepidopterists' Society 39(3). 1985. 171-176 COURTSHIP AND OVIPOSITION PATTERNS OF TWO AGATHYMUS (MEGATHYMIDAE) DON B. STALLINGS AND VIOLA N. T. STALLINGS P.O. Box 106, 616 W. Central, Caldwell, Kansas 67022 AND J. R. TURNER AND BEULAH R. TURNER 2 South Boyd, Caldwell, Kansas 67022 ABSTRACT. Males of Agathymus estelleae take courtship sentry positions near ten eral virgin females long before the females are ready to mate. Males of Agathymus mariae are territorial and pursue virgin females that approach their territories. Ovipo sition patterns of the two species are very similar. Females alight on or near the plants to oviposit and do not drop ova in flight. Few detailed observations of the courtship and oviposition of the skipper butterflies in natural environments have been published. For the family Megathymidae Freeman (1951), Roever (1965) (and see Toliver, 1968) described mating and oviposition of some Southwestern U.S. Agathymus, and over a hundred years ago (1876) Riley published an excellent paper on the life history of Megathymus yuccae (Bois duval & LeConte) which included data on oviposition of the female; otherwise, only the scantiest comments have been made. C. L. Rem ington (pers. comm.) and others tell us that there is a significant pos sibility that the Hesperioidea are less closely related to the true but terflies (Papilionoidea) than to certain other Lepidoptera and even that the Megathymidae may not be phylogenetically linked to the Hesper iidae. For several years we have been making on-the-scene studies of these two aspects of megathymid behavior, both for their interest in understanding the whole ecology of these insects and for their possible reflection on higher relationships. -
Louisiana's Animal Species of Greatest Conservation Need (SGCN)
Louisiana's Animal Species of Greatest Conservation Need (SGCN) ‐ Rare, Threatened, and Endangered Animals ‐ 2020 MOLLUSKS Common Name Scientific Name G‐Rank S‐Rank Federal Status State Status Mucket Actinonaias ligamentina G5 S1 Rayed Creekshell Anodontoides radiatus G3 S2 Western Fanshell Cyprogenia aberti G2G3Q SH Butterfly Ellipsaria lineolata G4G5 S1 Elephant‐ear Elliptio crassidens G5 S3 Spike Elliptio dilatata G5 S2S3 Texas Pigtoe Fusconaia askewi G2G3 S3 Ebonyshell Fusconaia ebena G4G5 S3 Round Pearlshell Glebula rotundata G4G5 S4 Pink Mucket Lampsilis abrupta G2 S1 Endangered Endangered Plain Pocketbook Lampsilis cardium G5 S1 Southern Pocketbook Lampsilis ornata G5 S3 Sandbank Pocketbook Lampsilis satura G2 S2 Fatmucket Lampsilis siliquoidea G5 S2 White Heelsplitter Lasmigona complanata G5 S1 Black Sandshell Ligumia recta G4G5 S1 Louisiana Pearlshell Margaritifera hembeli G1 S1 Threatened Threatened Southern Hickorynut Obovaria jacksoniana G2 S1S2 Hickorynut Obovaria olivaria G4 S1 Alabama Hickorynut Obovaria unicolor G3 S1 Mississippi Pigtoe Pleurobema beadleianum G3 S2 Louisiana Pigtoe Pleurobema riddellii G1G2 S1S2 Pyramid Pigtoe Pleurobema rubrum G2G3 S2 Texas Heelsplitter Potamilus amphichaenus G1G2 SH Fat Pocketbook Potamilus capax G2 S1 Endangered Endangered Inflated Heelsplitter Potamilus inflatus G1G2Q S1 Threatened Threatened Ouachita Kidneyshell Ptychobranchus occidentalis G3G4 S1 Rabbitsfoot Quadrula cylindrica G3G4 S1 Threatened Threatened Monkeyface Quadrula metanevra G4 S1 Southern Creekmussel Strophitus subvexus -
Butterflies and Moths of Pinal County, Arizona, United States
Heliothis ononis Flax Bollworm Moth Coptotriche aenea Blackberry Leafminer Argyresthia canadensis Apyrrothrix araxes Dull Firetip Phocides pigmalion Mangrove Skipper Phocides belus Belus Skipper Phocides palemon Guava Skipper Phocides urania Urania skipper Proteides mercurius Mercurial Skipper Epargyreus zestos Zestos Skipper Epargyreus clarus Silver-spotted Skipper Epargyreus spanna Hispaniolan Silverdrop Epargyreus exadeus Broken Silverdrop Polygonus leo Hammock Skipper Polygonus savigny Manuel's Skipper Chioides albofasciatus White-striped Longtail Chioides zilpa Zilpa Longtail Chioides ixion Hispaniolan Longtail Aguna asander Gold-spotted Aguna Aguna claxon Emerald Aguna Aguna metophis Tailed Aguna Typhedanus undulatus Mottled Longtail Typhedanus ampyx Gold-tufted Skipper Polythrix octomaculata Eight-spotted Longtail Polythrix mexicanus Mexican Longtail Polythrix asine Asine Longtail Polythrix caunus (Herrich-Schäffer, 1869) Zestusa dorus Short-tailed Skipper Codatractus carlos Carlos' Mottled-Skipper Codatractus alcaeus White-crescent Longtail Codatractus yucatanus Yucatan Mottled-Skipper Codatractus arizonensis Arizona Skipper Codatractus valeriana Valeriana Skipper Urbanus proteus Long-tailed Skipper Urbanus viterboana Bluish Longtail Urbanus belli Double-striped Longtail Urbanus pronus Pronus Longtail Urbanus esmeraldus Esmeralda Longtail Urbanus evona Turquoise Longtail Urbanus dorantes Dorantes Longtail Urbanus teleus Teleus Longtail Urbanus tanna Tanna Longtail Urbanus simplicius Plain Longtail Urbanus procne Brown Longtail -
Butterflies and Moths of El Paso County, Texas, United States
Heliothis ononis Flax Bollworm Moth Coptotriche aenea Blackberry Leafminer Argyresthia canadensis Apyrrothrix araxes Dull Firetip Phocides pigmalion Mangrove Skipper Phocides belus Belus Skipper Phocides palemon Guava Skipper Phocides urania Urania skipper Proteides mercurius Mercurial Skipper Epargyreus zestos Zestos Skipper Epargyreus clarus Silver-spotted Skipper Epargyreus spanna Hispaniolan Silverdrop Epargyreus exadeus Broken Silverdrop Polygonus leo Hammock Skipper Polygonus savigny Manuel's Skipper Chioides albofasciatus White-striped Longtail Chioides zilpa Zilpa Longtail Chioides ixion Hispaniolan Longtail Aguna asander Gold-spotted Aguna Aguna claxon Emerald Aguna Aguna metophis Tailed Aguna Typhedanus undulatus Mottled Longtail Typhedanus ampyx Gold-tufted Skipper Polythrix octomaculata Eight-spotted Longtail Polythrix mexicanus Mexican Longtail Polythrix asine Asine Longtail Polythrix caunus (Herrich-Schäffer, 1869) Zestusa dorus Short-tailed Skipper Codatractus carlos Carlos' Mottled-Skipper Codatractus alcaeus White-crescent Longtail Codatractus yucatanus Yucatan Mottled-Skipper Codatractus arizonensis Arizona Skipper Codatractus valeriana Valeriana Skipper Urbanus proteus Long-tailed Skipper Urbanus viterboana Bluish Longtail Urbanus belli Double-striped Longtail Urbanus pronus Pronus Longtail Urbanus esmeraldus Esmeralda Longtail Urbanus evona Turquoise Longtail Urbanus dorantes Dorantes Longtail Urbanus teleus Teleus Longtail Urbanus tanna Tanna Longtail Urbanus simplicius Plain Longtail Urbanus procne Brown Longtail -
Four New Species of Agathymus from Texas (Megathymidae)
1964 Journal of the Lepidopterists' Society 171 FOUR NEW SPECIES OF AGATHYMUS FROM TEXAS (MEGATHYMIDAE) by H. A. FREEMAN 1 160.5 Lewis, Garland, Texas I, as well as Stallings & Turner, have know for several years that there were several species mixed in material we have been calling Agath,ymus mariae (B. & B.) in Texas. During the past six years [ have been making a detailed study of that particular species complex. In working with this problem I used 32 locations in Texas and one in New Mexico and these constitute the known range of this species complex in the United States. As Agave lecheguilla Torr. is the known food plant of members of this complex their range can be followed by checking the range of the food plant. In making a study of the various habitats, the following information cn each was carefully noted: location, date, plant associates, type of soil, pH of the soil at the feeding level of the plant, elevation, average annual rainfall and the presence or absence of radiation. One of the most significant factors was the isolation of certain areas from the main gene pool of mariae. Another appeared to be the pH factor which seemed to have an influence on the presence or absence of various species in a given habitat. Typical mariae is distinctly associated with alkaline soil where the average is just below 8. In the area around Del Rio, Juno and Bracketville in Texas the reading L<: 7 or just slightly above, indicating near neutral or neutral soil. In the Chinati mountains the pH was around 7.4. -
SW Region USFS Sensitive Species List (7/21/99
SW Region USFS Sensitive species list Fed Global / (7/21/99 as corrected 2/23/00 ) Statu Sub-Spp State Forest of likely Scientific Name Common Name s Rank Rank State occurrence Mammals SOREX ARIZONAE ARIZONA SHREW S G3 S2S3 BOTH COR CYNOMYS LUDOVICIANUS BLACK-TAILED PRAIRIE DOG <not on 7/99 list?> Cand ALL KI,RB,BK PEROMYSCUS EREMICUS PAPAGENSIS CACTUS MOUSE S G5T1T2 SYN AZ THOMOMYS BOTTAE PAGUATAE CEBOLLETA POCKET GOPHER S G5T2 S2 NM SCIURUS NAYARITENSIS CHIRACAHUAE CHIRICAHUA FOX SQUIRREL S G5T1T2 S1S2 AZ COR EUTAMIAS MINIMUS CHUSKAENSIS CHUSKAEN LEAST CHIPMUNK S G5T1 S1 NM OVIS CANADENSIS MEXICANA DESERT BIGHORN SHEEP S G4G5T3 S3S4 BOTH COR,K,T OCHOTONA PRINCEPS NIGRESCENS GOAT PEAK PIKA S G5T1 S1? NM SFE THOMOMYS UMBRINUS GRAHAMENSIS GRAHAM MOUNTAINS POCKET GOPHER S G5T3Q SSYN AZ COR THOMOMYS BOTTAE GUADALUPENSIS GUADALUPE POCKET GOPHER S G5T2 S1 NM THOMOMYS UMBRINUS INTERMEDIUS HAUCHUCA MOUNTAINS POCKET GOPHER S G5T3 S3 AZ MICROTUS MEXICANUS NAVAHO NAVAJO MEXICAN VOLE S G5T2Q S1 AZ ZAPUS HUDSONICUS LUTEUS NEW MEXICAN (MEADOW) JUMPING MOUSE S G5T3 S2 BOTH A/S,CAR,LIN,SFE,GILA EUTAMIAS QUADRIVITTATUS AUSTRALIS ORGAN MOUNTAINS CHIPMUNK S G5T1 S1 NM EUTAMIAS QUADRIVITTATUS OSCURAENSIS OSCURA MOUNTAINS CHIPMUNK S G5T1 S1 NM EUTAMIAS MINIMUS ATRISTRIATUS PENASCO CHIPMUNK S G5T1 S1 NM LIN AMMOSPERMOPHILUS LEUCURUS TERSUS PROSPECT VALLEY WHITE-TAIL ANTELOPE SQRL S G5T1Q S1 AZ ANTILOCAPRA AMERICANA SONORIENSIS SONORAN PRONGHORN <remove to E list> E G5T1 S1 AZ THOMOMYS UMBRINUS QUERCINUS SOUTHERN POCKET GOPHER SSP S G5T3 SSYN AZ COR THOMOMYS -
Section IV – Guideline for the Texas Priority Species List
Section IV – Guideline for the Texas Priority Species List Associated Tables The Texas Priority Species List……………..733 Introduction For many years the management and conservation of wildlife species has focused on the individual animal or population of interest. Many times, directing research and conservation plans toward individual species also benefits incidental species; sometimes entire ecosystems. Unfortunately, there are times when highly focused research and conservation of particular species can also harm peripheral species and their habitats. Management that is focused on entire habitats or communities would decrease the possibility of harming those incidental species or their habitats. A holistic management approach would potentially allow species within a community to take care of themselves (Savory 1988); however, the study of particular species of concern is still necessary due to the smaller scale at which individuals are studied. Until we understand all of the parts that make up the whole can we then focus more on the habitat management approach to conservation. Species Conservation In terms of species diversity, Texas is considered the second most diverse state in the Union. Texas has the highest number of bird and reptile taxon and is second in number of plants and mammals in the United States (NatureServe 2002). There have been over 600 species of bird that have been identified within the borders of Texas and 184 known species of mammal, including marine species that inhabit Texas’ coastal waters (Schmidly 2004). It is estimated that approximately 29,000 species of insect in Texas take up residence in every conceivable habitat, including rocky outcroppings, pitcher plant bogs, and on individual species of plants (Riley in publication). -
89 Megathymus Yuccae in Texas, with the Description
1963 Journal of the Lepidopterists' Society 89 MEGATHYMUS YUCCAE IN TEXAS, WITH THE DESCRIPTION OF TWO NEW SUBSPECIES by H. A. FREEMAN After many hours of thought and discussion DON B. STALLINGS and I have reached the following conclusions concerning the arrangement of the various subspecies of Megathymus yuccae (Bdv. & Lec.) in the state of Texas. We have been working with this group for nearly twenty years, and there was a time when we had a few specimens of yuccae from isolated locations over the state and these clearly told us the complete story of yuccae in this area. Now that we have nearly five thousand specimens of Megathymidae in our separate collections the clarity of the picture dims. Since receiving a research grant from the National Science Foundation I have been making every effort to approach this problem from a strictly scientific viewpoint. There are so many factors that seem to have brought about the production of subspeciation in this group that I have been trying to associate these factors together so as to arrive at some definite conclusions. In the various habitats that I have been studying over the state such things as plant associates, type of soil, pH of soil, elevation, average rainfall, and whether or not there is any indication of radiation present, have been checked and recorded. These factors certainly seem to have had some influence upon the particular subspecies found in any given area. Genetically speaking there appear to be two major influences affecting the Texas populations, from colioradensis to the west and from yuccae to the east, while from the south comes a minor influence of wilsonorum. -
Checklist of Texas Lepidoptera Knudson & Bordelon, Jan 2018 Texas Lepidoptera Survey
1 Checklist of Texas Lepidoptera Knudson & Bordelon, Jan 2018 Texas Lepidoptera Survey ERIOCRANIOIDEA TISCHERIOIDEA ERIOCRANIIDAE TISCHERIIDAE Dyseriocrania griseocapitella (Wlsm.) Eriocraniella mediabulla Davis Coptotriche citripennella (Clem.) Eriocraniella platyptera Davis Coptotriche concolor (Zell.) Coptotriche purinosella (Cham.) Coptotriche clemensella (Cham). Coptotriche sulphurea (F&B) NEPTICULOIDEA Coptotriche zelleriella (Clem.) Tischeria quercitella Clem. NEPTICULIDAE Coptotriche malifoliella (Clem.) Coptotriche crataegifoliae (Braun) Ectoedemia platanella (Clem.) Coptotriche roseticola (F&B) Ectoedemia rubifoliella (Clem.) Coptotriche aenea (F&B) Ectoedemia ulmella (Braun) Asterotriche solidaginifoliella (Clem.) Ectoedemia obrutella (Zell.) Asterotriche heliopsisella (Cham.) Ectoedemia grandisella (Cham.) Asterotriche ambrosiaeella (Cham.) Nepticula macrocarpae Free. Asterotriche helianthi (F&B) Stigmella scintillans (Braun) Asterotriche heteroterae (F&B) Stigmella rhoifoliella (Braun) Asterotriche longeciliata (F&B) Stigmella rhamnicola (Braun) Asterotriche omissa (Braun) Stigmella villosella (Clem.) Asterotriche pulvella (Cham.) Stigmella apicialbella (Cham.) Stigmella populetorum (F&B) Stigmella saginella (Clem.) INCURVARIOIDEA Stigmella nigriverticella (Cham.) Stigmella flavipedella (Braun) PRODOXIDAE Stigmella ostryaefoliella (Clem.) Stigmella myricafoliella (Busck) Tegeticula yuccasella (Riley) Stigmella juglandifoliella (Clem.) Tegeticula baccatella Pellmyr Stigmella unifasciella (Cham.) Tegeticula carnerosanella Pellmyr -
Arizona Wildlife Notebook
ARIZONA WILDLIFE CONSERVATION ARIZONA WILDLIFE NOTEBOOK GARRY ROGERS Praise for Arizona Wildlife Notebook “Arizona Wildlife Notebook” by Garry Rogers is a comprehensive checklist of wildlife species existing in the State of Arizona. This notebook provides a brief description for each of eleven (11) groups of wildlife, conservation status of all extant species within that group in Arizona, alphabetical listing of species by common name, scientific names, and room for notes. “The Notebook is a statewide checklist, intended for use by wildlife watchers all over the state. As various individuals keep track of their personal observations of wildlife in their specific locality, the result will be a more selective checklist specific to that locale. Such information would be vitally useful to the State Wildlife Conservation Department, as well as to other local agencies and private wildlife watching groups. “This is a very well-documented snapshot of the status of wildlife species – from bugs to bats – in the State of Arizona. Much of it should be relevant to neighboring states, as well, with a bit of fine-tuning to accommodate additions and deletions to the list. “As a retired Wildlife Biologist, I have to say Rogers’ book is perhaps the simplest to understand, yet most comprehensive in terms of factual information, that I have ever had occasion to peruse. This book should become the default checklist for Arizona’s various state, federal and local conservation agencies, and the basis for developing accurate local inventories by private enthusiasts as well as public agencies. "Arizona Wildlife Notebook" provides a superb starting point for neighboring states who may wish to emulate Garry Rogers’ excellent handiwork. -
Genomics of a Complete Butterfly Continent
bioRxiv preprint doi: https://doi.org/10.1101/829887; this version posted November 4, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. Genomics of a complete butterfly continent Jing Zhang2,*, Qian Cong3,*, Jinhui Shen2, Paul A. Opler4 and Nick V. Grishin1,2,# 1Howard Hughes Medical Institute and 2Departments of Biophysics and Biochemistry, University of Texas Southwestern Medical Center, Dallas, TX, 75390, USA; 3Institute for Protein Design and Department of Biochemistry, University of Washington, Seattle, WA, 98195, USA; 4Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO, 80523, USA. *These authors contributed equally to this work, #Corresponding author: [email protected] Never before have we had the luxury of choosing a continent, picking a large phylogenetic group of animals, and obtaining genomic data for its every species. Here, we sequence all 845 species of butterflies recorded from North America north of Mexico. Our comprehensive approach reveals the pattern of diversification and adaptation occurring in this phylogenetic lineage as it has spread over the continent, which cannot be seen on a sample of selected species. We observe bursts of diversification that generated taxonomic ranks: subfamily, tribe, subtribe, genus, and species. The older burst around 70 Mya resulted in the butterfly subfamilies, with the major evolutionary inventions being unique phenotypic traits shaped by high positive selection and gene duplications. The recent burst around 5 Mya is caused by explosive radiation in diverse butterfly groups associated with diversification in transcription and mRNA regulation, morphogenesis, and mate selection.