INTERNATIONAL UNION OF BIOLOGICAL SCIENCES SECTION FOR PALAEOBOTANY

International Organisation of Palaeobotany

IOP NEWSLETTER 110

August 2016

CONTENTS

FROM THE SECRETARY/TREASURER IPC XIV/IOPC X 2016 IOPC 2020 IOP MEMBERSHIP IOP EXECUTIVE COMMITTEE ELECTIONS IOP WEBMASTER POSITION OBITUARY: Dr. Richard D. Dayvault WHAT HAPPENED TO THE OUPH COLLECTIONS? THE PALAEOBOTANY OF ITALY UPCOMING MEETINGS CALL FOR NEWS and NOTES

The views expressed in the newsletter are those of its correspondents, and do not necessarily reflect the policy of IOP. Please send us your contributions for the next edition of our newsletter (June 2016) by May 30th, 2016.

President: Johanna Eder-Kovar (Germany) Vice Presidents: Bob Spicer (Great Britain), Harufumi Nishida (Japan), Mihai Popa (Romania) Members at Large: Jun Wang (China), Hans Kerp (Germany), Alexej Herman (Russia) Secretary/Treasurer/Newsletter editor: Mike Dunn (USA) Conference/Congress Chair: Francisco de Assis Ribeiro dos Santos

IOP Logo: The evolution of architecture (© by A. R. Hemsley) IOP 110 2 August 2016

included in that newsletter to the address at the end of this newsletter. FROM THE In addition, please send any issues that you SECRETARY/TREASURER think need to be addressed at the Business meeting. I will add those to the Agenda. Dear IOP Members, Respectfully, I am happy to report, that IOP seems to be on track and ready for a new Executive Mike Council to take over. The elections are progressing nicely and I will report the results in the September/October Newsletter. IPC XIV/IOPC X 2016 The one area that is still problematic is the webmaster position. We really to talk amongst ourselves, and find someone who is willing and able to do the job. I discuss the position in more detail below.

Everything seems to be falling into place for IOPC X at Salvador Brazil. The Olympics seem to have gone well, so the Brazilians are all practiced as great hosts, so our meeting should be great fun. I look forward to seeing all of you there.

As always I am eternally grateful for submissions to the newsletter, even if the The 2016 joint meeting of the International news is bad. Carole Gee reports on the bad Palynological Congress and the International news of the passing of Richard Dayvault, Organization of Palaeobotanists will be held and well as the good news of the publication 23-28 October 2016 at Salvador, Bahia, of ‘The Palaeobotany of Italy’. And Gar Brazil. The venue is The Bahia Othon Rothwell provided an important update on Palace. the status of the Ohio University Paleobotanical collections since his The homepage with registration details is up retirement from that institution. and running.

I will publish one more newsletter in late The Website is: September-Early October to report on the results of the elections, the status of IOP, http://www.ipciopcbrazil.com/ and the upcoming IOPC 2016 meetings. The site is very easy to use, but if your As always, please send anything you want Institution has a robust Firewall the site may IOP 110 3 August 2016 be blocked as it was for me. I simply IOP EXECUTIVE registered from home and had no problem. COMMITTEE ELECTIONS Time is running out, but do please remember to double check the requirements for The IOP Executive Committee elections are entering Brazil. For example, US citizens underway and will be open until 30 August. need a visa, but every country is different. To date 73 members have voted for a participation rate of 26%. That is an even worse voting rate than American elections, and I know that you don’t want to make us IOPC 2020 Ugly Americans look good. So if you haven’t already, please vote. If you need At this time we have no official bids to host another ballot, please just let me know. IOPC2020. IOP Webmaster IOP MEMBERSHIP I was very sorry to accept the resignation of Thank you for the great response to the Alan Spencer as our Webmaster. Alan has recent call for membership updates. I did a done a great job for us for more than 10 hard cull to clean up the rolls for the years now, but he has decided that he needs incoming Secretary/Treasurer and so we to focus on other things. I would personally now have 279 members. I deleted any like to thank him publically, and member who was more than 2 years in acknowledge how much he has helped me arrears. I used the 2 year cut-off because the these past years. His help was invaluable. I current system won’t allow anyone to pay hope all of you also take the time to thank dues until after their membership has him for his service to IOP. expired, so it is easy to forget to pay when one’s membership expires on January 1. But it does mean that we need someone to take on that position. Alan has generously In addition, some members have a bias offered to be there to assist in the transition, against using Paypal. If there are issues with so things should go smoothly for our next Paypal, please let me know what they are, webmaster. If you have an interest in the and we can discuss them at the Business position, or know someone who would like Meeting. That system makes life very easy to serve IOP as Webmaster, please let me for the Secretary/Treasurer but if there are know as soon as possible. This position is problems we need to know what they are so not voted on, and you can serve as long or as we can address them. Sending checks in short of time as you would like. anything except US dollars is not helpful because the exchange costs are too Our organisation can’t function properly prohibitive. without a webmaster. Alan and our previous webmasters have done a great job of setting up and maintaining the site, so it just needs someone with the time and skills to keep it IOP 110 4 August 2016 running. Please consider this position, if it toxic uranium mine waste sites in western fits your agenda. Colorado, but on the weekends, he and his wife Jalena became avid hunters of OBITUARY . In fact, Dick had collected fossil plants of all sorts for a number of decades; his email moniker was “geocycad.” Dr. Richard D. Dayvault 6 May 6 1948 – 29 August 2015 However, it was through the fossil cones, of which he had amassed hundreds of specimens from all over Utah, that we met. In 2007, Dick and Steven Hatch published an article on “ Cones from the and Rocks of Eastern Utah” in which they appealed to the paleobotanical world to have a closer look at these cones. Reading this article, I could not believe the wealth of three-dimensionally preserved cones that had been gathered together by Dick through the amalgamation of older private collections and his own dedicated collecting. By heeding his call in the article, Dick and I became friends, colleagues, and research partners in 2008 on conifer cones from the Upper Jurassic Morrison Formation from Utah and Dick Dayvault with wife Jalena in the Wyoming. Upper Jurassic Morrison Formation of NE Utah Hence, we didn’t know each other very The last year has been an extremely sad one long—three years short of a decade—but we for all of us with the passing of so many would go prospecting for cones in Utah beloved paleobotanists and palynologists. pretty much every single year and As we mourn the loss of our colleagues, we collaborated intensively for an overview also reflect on their contribution to our field paper entitled "Palaeobiodiversity of conifer of science and personal enrichment to our seed cones in the Upper Jurassic Morrison lives. I’d like to tell you about my research Formation of Utah, USA,” which was partner and friend of mine, Dr. Richard D. published in Palaeobiodiversity and Dayvault, who died about a year ago. Palaeoenvironments just last year (June 2014). Dick was a professional geologist and mineralogist who worked on environmental Born and raised in North Carolina, Dick clean-up for and with the Department of spent his early years collecting rocks, Energy in Grand Junction, Colorado, for 35 minerals, and crystals. He took his interests years. Weekdays on the job, he cleaned up into college, where he received his B.S. and IOP 110 5 August 2016

M.S. degrees in Geology from East Carolina and hydrologist, by his mother Anna Lee, University in Greenville, N.C. Afterwards, brother Steve (and wife Paula) of North Dick moved on to the University of South Carolina, and honorary sister, K.D. Carolina in Columbia, where he was Curator Ashbrook of Grand Junction. at the L.L. Smith Geology Museum and taught physical and historical geology Yes, Dick and I had only seven years of classes. He then continued to teach these fieldwork and research together, and he has courses, plus igneous petrology labs, at East left us all much too soon, but Dick’s Carolina University. contributions to Jurassic of North America and to the Morrison Thus, Dick was on the academic track, when Formation will stand forever. the call for a geoscientist drew him westward to the Department of Energy in Grand Junction, Colorado. Dick told me Carole Gee that this first road trip west—replete with University of Bonn gorgeous arid landscapes, breathtaking Germany geology, and wide open spaces—hooked him, like they continue to captivate me. Many of our long drives to fossil cone localities were spent chatting and taking in WHAT HAS HAPPENED TO the strikingly beautiful rock formations of THE OHIO UNIVERSITY eastern Utah. PALEOBOTANICAL Paleontology came relatively late in Dick’s COLLECTIONS? career, but like all of his interests, he embraced it with enthusiasm, vigor, as well As many of you know, Gar Rothwell has as a scholarly approach, for Dick was not retired from Ohio University, and with that, just a “rockhound.” In addition to collecting the Ohio University Paleobotanical and minerals in his spare time, for the Collections are now dispersed to a number past few decades, Dick wrote and published of museums and collections. Because the extensively, for example, Ancient Forests: A ability to reference and re-examine Closer Look at Fossil Wood, a aesthetically published specimens is an integral part of well-illustrated book couthored with Frank paleobotanical research, Gar has submitted Daniels, and a number of scientific articles the following update on where we can find on fossil plants, silicified wood, geodes, and the specimens we might need for our minerals. Dick was a consulting editor for research. the magazine Rocks and Minerals from 1976 to 2015, as well as the recipient of several Publications with OUPH specimen awards by geological societies in Colorado. content: Dick passed away at home at the end of August 2015 after a brief illness. He is The following list of publications, survived by his wife Jalena, a civil engineer although probably incomplete, includes nearly all of the studies that are based on IOP 110 6 August 2016 specimens that until recently have been Society of Edinburgh (Earth housed at Ohio University. Specimens that Sciences) 81: 127-159. support this research, and that were housed Beeler, H. E. 1983. Anatomy and frond in the Ohio University Paleobotanical architecture of ovata and Herbarium, have been transferred to the N. scheuchzeri from the Upper Field Museum of Natural History, Chicago, of the Appalachian and the Natural History Museum, University Basin. Canadian Journal of of Kansas, Lawrence. The collections have 61: 2352-2368. been relocated by general topic as follows: Dunn, M. T., M. Krings, G. Mapes, G. W. Rothwell, R. H. Mapes, K-Q Sun. Coal Ball material – Field Museum 2003. Medullosa steinii sp. nov., a /Lower seed seed vine from the Late plants – University of Kansas . Review of Psaronius fern trunks – Field Museum Palaeobotany and Palynology 124: Paleozoic – University of Kansas 307-324. Anatomical sections of living plants – Dunn, M. T., G. Mapes and G. W. Rothwell. Field Museum 2001. On Paleozoic plants from Mississippian (Weddington Delta, marine strata: Hexaloba finisensia Arkansas) fossils – University of Kansas new genus and , a Miscellaneous collections – Field Museum trigonocarpalean from the Virgilian () Finis Shale of Texas. Journal of Paleontology 76: Publications 173-180. Dunn, M. and G. W. Rothwell. 2012. Basinger, J. F., G. W. Rothwell, and W. N. Phenotypic plasticity of the Stewart. 1974. Cauline vasculature hydrasperman seed fern Tetrastichia and trace production in bupatides Gordon. International medullosan pteridosperms. American Journal of Plant Sciences 173: 823- Journal of Botany 61: 1002-1015. 834. Bateman, R. M., P. Kenrick, and G. W. Dunn, M. T., G. W. Rothwell, and G. Rothwell. 2007. Do eli ghuelrabtaecous Mapes. 2003. On Paleozoic plants lycopsids occur in Palaeozoic strata? from marine strata: Trivena Hestia eremosa gen. et sp. nov. from arkansana gen. et sp. nov., a the Dinantian of Oxroad Bay, East lyginopterid from the Fayetteville Lothian, Scotland. Review of Formation (middle Chesterian/Upper Palaeobotany and Palynology 144: Mississippian) of Arkansas, USA. 323-335. American Journal of Botany 90: Bateman, R. M., and G. W. Rothwell. 1990. 1239-1252. A reappraisal of the Dinantian floras Dunn, M. T., G. W. Rothwell, and G. at Oxroad Bay, East Lothian, Mapes. 2006. The Fayetteville Flora Scotland, I. Floristics and the of Arkansas (USA): A Snapshot of development of whole plant Terrestrial Vegetation Patterns concepts. Transactions of the Royal within a Clastic Swamp at Upper IOP 110 7 August 2016

Mississippian Time. P. 127-137. In: American Journal of Botany 67: DiMichele, W.A, and S. Greb, eds. 1051-1058. Wetland paleoecology through time. Hamer, J. J., and G. W. Rothwell. 1983. Geological Society of America, Phillipopteris gen. nov.-- Special Paper 139. anatomically preserved sporangial Erwin, D. M., and G. W. Rothwell. 1989. fructifications from the Upper Gillespia randolphensis sp. nov. Pennsylvanian of the Appalachian (Stauropteridales), from the Upper Basin. American Journal of Botany Devonian of West Virginia. 70: 1378-1385. Canadian Journal of Botany 67: Hamer, J. J., and G. W. Rothwell. 1988. The 3063-3077. vegetative structure of Medullosa Feng, B. C., and G. W. Rothwell. 1989. endocentrica Baxter Microsporangiate cones of (Pteridospermopsida). Canadian Mazocarpon bensonii from the Journal of Botany 66: 375-387. Upper Pennsylvanian of the Hernandez-Castillo, G. R., G. W. Rothwell Appalachian Basin. Review of and G. Mapes. 2001. Compound Palaeobotany and Palynology 57: pollen cones in Paleozoic conifers. 289-297. American Journal of Botany 88: Frankenberg, J. M. and Eggert, D. A., 1969. 1139-1142. Petrified Stigrnaria from North Hernandez-Castillo, G. R., G. W. Rothwell America, I. Stigrnaria ficoides, the and G. Mapes. 2001. Thucydiaceae underground portions of fam. nov., with a review and Lepidodendraceae. reevaluation of Paleozoic walchian Palaeontographica 128B: 1-47. conifers. International Journal of Gillespie, W. H., G. W. Rothwell, and S. E. Plant Sciences 162: 1155-1185. Scheckler. 1981. The earliest seeds. Hernandez-Castillo, G. R., G. W. Rothwell, Nature 293: 462-464. R. A. Stockey and G. Mapes. 2003. Good, C. S., and G. W. Rothwell. 1988. A Growth architecture of Thucydia reinterpretation of the Paleozoic fern mahoningensis, a model for Norwoodia angustum. Review of primitive walchian conifer plants. Palaeobotany and Palynology 56: International Journal of Plant 199-204. Sciences 164: 443-452. Good, C. W., G. W. Rothwell, and T. N. Hernandez-Castillo, G. R., G. W. Rothwell Taylor. 1982. A new and R. A. Stockey. 2009. The role of Stephanospermum from the multivariate analysis in Appalachian Basin of North reconstructing fossil conifers. Pages America. Review of Palaeobotany 21-28. In: Bieleski, R.L. and M.D. and Palynology 36: 231-240. Wilcon, eds., , Grove, G. G., and G. W. Rothwell. 1980. Proceedings of the 2002 Mitrospermum vinculum sp. nov., a Araucariaceae Symposium. cardiocarpalean ovule from the International Dendrology Society, Upper Pennsylvanian of Ohio. Kunedin, New Zealand. IOP 110 8 August 2016

Hernandez-Castillo, G. R., R. A. Stockey, G. development of L. III. The W. Rothwell and G. Mapes. 2009. sequence of root initiation in three- Reconstruction of the Pennsylvanian- and four-lobed plants of I. age walchian conifer Emporia tuckermanii A. Br. and I. nuttallii A. cryptica sp. nov. (Emporiaceae, Br. Botanical Gazette 139:271-283. Voltziales). Review of Knaus, M. J. 1995. The Species of the Early Palaeobotany and Palynology 157: Carboniferous Fossil Plant Genus 218-237. Genselia. International Journal of Hernandez-Castillo, G. R., R. A. Stockey, G. Plant Sciences 156: 61-92. W. Rothwell and G. Mapes. 2009. A Knaus, M. J., G. R. Upchurch Jr. and W. H. new voltzialean conifer, Emporia Gillespie. 2000. Charbeckia royalii sp nov. (Emporiaceae), from macrophylla gen. et sp. nov. from the the Hamilton Quarry, Kansas. Lower Mississippian Price (Pocono) International Journal of Plant Formation of southeastern West Sciences 170: 1201-1227. Virginia. Review of Palaeobotany Hernandez-Castillo, G. R., R. A. Stockey, G. and Palynology 111: 71–92. W. Rothwell and G. Mapes. 2009. Mapes, G., and G. W. Rothwell. 1980. Whole plant reconstruction of Quaestora amplecta gen. et sp. nov., Emporia lockardii (Emporiaceae) a structurally simple medullosan Voltziales and initial thoughts on stem from the Upper Mississippian Paleozoic conifer ecology. of Arkansas. American Journal of International Journal of Plant Botany 67: 636-647. Sciences 170: 1056-1074. Mapes, G., and G. W. Rothwell. 1988. Jennings, J. E., E. E. Karrfalt, and G. W. Diversity among Hamilton conifers. Rothwell. 1983. Structure and In: G. Mapes and R. H. Mapes, eds., affinities of Protostigmaria Regional geology and paleontology eggertiana. American Journal of of upper Paleozoic Hamilton quarry Botany 70: 963-974. area in southeastern Kansas, Kansas Karrfalt, E. E., and D. A. Eggert. 1977a. The Geological Survey Guidebook Series comparative morphology and 6: 225-244. development of Isoetes L. I. Lobe Mapes, G., and G. W. Rothwell. 1991. and furrow development in I. Structure and relationships of tuckermanii A. Br. Botanical Gazette primitive conifers. Neus Jahrbuch 138:236-247. fur Geol. Palaontol., Stuttgart. 183: Karrfalt, E. E., and D. A. Eggert. 1977b. The 269-287. comparative morphology and Mapes, G., and G. W. Rothwell. 1984. development of Isoetes L. II. Permineralized ovulate cones of Branching of the base of the in Lebachia from Late Paleozoic I. tuckermanii A. Br. and I. nuttallii limestones of Kansas. Palaeontology A. Br. Botanical Gazette 138:357- 27: 69-94. 368. Mapes, G., and G. W. Rothwell. 1998. Karrfalt, E. E., and D. A. Eggert. 1978. The Pollen cone structure of the Late comparative morphology and Pennsylvanian (Stephanian) conifer IOP 110 9 August 2016

Emporia. Journal of Paleontology Pigg, K. B., and G. W. Rothwell. 1979. 72: 571-576. Stem-root transition of an Upper Mapes, G. and G. W. Rothwell. 2003. Pennsylvanian woody lycopsid. Validation of the names American Journal of Botany 66: 914- Emporiaceae, Emporia, and Emporia 924. lockardii. Taxon 52: 327-328. Pigg, K. B., and G. W. Rothwell. 1983. Mapes, G., G. W. Rothwell, and M. T. Chaloneria gen. nov., heterosporous Haworth. 1989. Evolution of seed from the Pennsylvanian dormancy. Nature 337: 645-646. of North America. Botanical Gazette Mickle, J. E. 1983. Psaronius, tree fern of 144: 132-147. the coal age. The Living Museum. Pigg, K. B., and G. W. Rothwell. 1983. 45: 12-14. Megagametophyte development in Mickle, J. E. 1984. Aspects of growth and the Chaloneriaceae fam. nov., development in the Pennsylvanian permineralized Paleozoic Isoetales age marattialean fern Psaronius. (Lycopsida). Botanical Gazette 144: Botanical Gazette 145: 407-419. 297-302. Mickle, J. E. 1984. of specimens Pigg, K. B., and G. W. Rothwell. 1985. of the Pennsylvanian-age Cortical development in Chaloneria marattialean fern Psaronius from cormosa (Isoetales), and the Ohio and Illinois. Illinois State biological derivation of compressed Museum scientific paper No. XIX. decortication taxa. Springfield, Illinois, 64 pp. Palaeontology 28: 545-553. Mickle, J. E., and G. W. Rothwell. 1982. Ramanujam, C. G. K., G. W. Rothwell, and from the Upper W. N. Stewart. 1974. Probable Pennsylvanian of Ohio and Illinois. attachment of the Dolerotheca Journal of Paleontology 36: 392-402. campanulum to a Myeloxylon- Mickle, J. E., and G. W. Rothwell. 1986. Alethopteris type frond. American Vegetative and fertile structures of Journal of Botany 61: 1057-1066. Cyathotheca ventilaria from the Rice, J., G. W. Rothwell, G. Mapes and R. Upper Pennsylvanian of the H. Mapes. 1996. Suavitas imbricata Appalachian Basin. American gen. et sp. nov., an anatomically Journal of Botany 73: 1474-1485. preserved seed analogue from Upper Millay, M. A. and G. W. Rothwell. 1983. Pennsylvanian marine deposits of Fertile pinnae of Biscalitheca midcontinent North America. () from the Upper American Journal of Botany 83: Pennsylvanian of the Appalachian 1083-1090. Basin. Botanical Gazette 144: 589- Riggs, S. D., and G. W. Rothwell. 1985. 599. Sentistrobus goodii n. sp., a Millay, M. A., G. W. Rothwell, and D. A. permineralized sphenophyllalean Eggert. 1980. Ultrastructure of cone from the Upper Pennsylvanian Stewartiotheca (Medullosaceae) of the Appalachian Basin. Journal of prepollen. American Journal of Paleontology 59: 1194-1202. Botany 67: 223-227. IOP 110 10 August 2016

Rothwell, G. W. 1971. Additional Palaeontology 19: 307-315. observations on Conostoma anglo- Rothwell, G. W. 1976. Primary vasculature germanicum and C. oblongum from and systematics. the Lower Pennsylvanian of North Review of Palaeobotany and America. Palaeontographica 131B: Palynology 22: 193-206. 167-178. Rothwell, G. W. 1977. Evidence for a Rothwell, G. W. 1971. Ontogeny of the pollination drop mechanism in Paleozoic ovule: Callospermarion Paleozoic pteridosperms. Science pusillum. American Journal of 198: 1251-1252. Botany 58: 706-715. Rothwell, G. W. 1977. The primary Rothwell, G. W., and T. N. Taylor. 1971. vasculature of Cordaianthus Studies of Paleozoic calamitean concinnus. American Journal of cones: Weissia kentuckiense gen., et Botany 64: 1235-1241. sp. nov. Botanical Gazette 132: 215- Rothwell, G. W. 1978. Doneggia complura 224. gen. et sp. nov., a filicalean fern from Rothwell, G. W. 1972. Palaeosclerotium the Upper Pennsylvanian of Ohio. pusillum gen., et sp. nov., a fossil Canadian Journal of Botany 56: eumycete from the Pennsylvanian of 3096-3104. Illinois. Canadian Journal of Botany Rothwell, G. W. 1980. The 50: 2353-2356. Rothwell, G. W. 1972. Pollen organs of the (Pteridospermopsida): II. Pennsylvanian Callistophytaceae Reproductive Features. (Pteridospermopsida). American Palaeontographica 175B: 85-106. Journal of Botany 59: 993-999. Rothwell, G. W. 1981. The Callistophytales Rothwell, G. W. 1972. Evidence of pollen (Pteridospermopsida); reproductively tubes in Paleozoic pteridosperms. sophisticated Paleozoic Science 175: 772-774. . Review of Rothwell, G. W. 1973. Vegetative structures Palaeobotany and Palynology 32: of the Callistophytaceae 103-121. (Pteridospermopsida). Ph.D. Rothwell, G. W. 1981. Cordaianthus dissertation, Univ. Alberta, 126 pp. duquesnensis sp. nov., anatomically Rothwell, G. W. 1975. The preserved ovulate cones from the Callistophytaceae Upper Pennsylvanian of Ohio. (Pteridospermopsida): I. vegetative American Journal of Botany 68: 239- structures. Palaeontographica 151B: 247. 171-196. Rothwell, G. W. 1982. New interpretations Rothwell, G. W. 1976. Petrified of the earliest conifers. Review of Pennsylvanian age plants of eastern Palaeobotany and Palynology 37: 7- Ohio. Ohio Journal of Science 76: 28. 128-132. Rothwell, G. W. 1984. The apex of Rothwell, G. W. 1976. A new pteropsid Stigmaria (Lycopsida), rooting organ fructification from the Middle of Lepidodendrales. American Pennsylvanian of Kansas. Journal of Botany 71: 1031-1034. IOP 110 11 August 2016

Rothwell, G. W. 1986. Classifying the Tidwell, W. D., Wild, V. and earliest gymnosperms. In: Spicer, R. Wright, W.W. Secrets of petrified A., and B. A. Thomas eds., plants. D=ORO Verlag, Systematic and taxonomic Heppenheim, Germany. approaches in Paleobotany. The Rothwell, G. W., and A. H. Blickle. 1982. Systematics Association Special Psaronius magnificus n. comb., a Volume No. 31, Linnean Society of marattialean fern from the Upper London, Academic Press, London, Pennsylvanian of North America. pp. 137-161. Journal of Paleontology 36: 259-268. Rothwell, G. W. 1987. The role of Rothwell, G. W., and D. A. Eggert. 1970. A development in plant phylogeny: A Conostoma with a tentacular paleobotanical perspective. Review sarcotesta from the Upper of Palaeobotany and Palynology 50: Pennsylvanian of Illinois. Botanical 97-114. Gazette 131: 359-366. Rothwell, G. W. 1987. Complex Paleozoic Rothwell, G. W., and D. A. Eggert. 1982. Filicales in the evolutionary radiation What is the vascular architecture of of . American Journal of Botany complex medullosan pollen organs? 74: 458-461. American Journal of Botany 69: 641- Rothwell, G. W. Cordaitales. 1988. In: C.B. 643. Beck (ed.), Origin and Evolution of Rothwell, G. W., and D. A. Eggert. 1986. A Gymnosperms. Columbia University monograph of Dolerotheca Halle, Press. pp. 273-297. and related complex permineralized Rothwell, G. W. 1988. Upper Pennsylvanian medullosan pollen organs. Steubenville coal-ball flora. Ohio Transactions of the Royal Society of Journal of Science 88: 61-64. Edinburgh, Earth Sciences 77: 47-79. Rothwell, G. W. 1990. Botryopteris forensis Rothwell, G. W., and D. M. Erwin. 1985. (Botryopteridaceae), a trunk epiphyte The rhizomorph apex of of the tree fern Psaronius. American Paurodendron; implications for Journal of Botany 78: 782-788. homologies among the rooting Rothwell, G. W. 1993. Cordaixylon organs of Lycopsida. American dumusum (Cordaitales). II. Journal of Botany 72: 86-98. Reproductive biology, phenology Rothwell, G. W., and D. M. Erwin. 1987. and growth ecology. International Origin of seed plants: an Journal of Plant Science 154: 572- aneurophyte-seed fern link 586. elaborated. American Journal of Rothwell, G. W. 2002. Coal Balls: Botany 74: 970-973. remarkable evidence for Paleozoic Rothwell, G. W. and C. W. Good. 2000. fossil plants and the communities in Reconstruction of the Pennsylvanian which they grew. P. 47-63. In: age filicalean fern Botryopteris Barthel, M., Dernbach, U., Galtier, tridentata. International Journal of J., Jung, W., Kerp, H., Noll, R., Plant Sciences 161: 495-507. Rößler, R., Rothwell, G.W., Rothwell, G. W. and E. E. Karrfalt. 2008. Selmeier, A., Stockey, R. A., Growth, development and IOP 110 12 August 2016

systematics of ferns: Does Rothwell, G. W., G. Mapes and R. H. Botrychium s.l. (Ophioglossales) Mapes. 1997. Late Paleozoic really produce secondary xylem? conifers of North America; structure, American Journal of Botany 95: 414- diversity and occurrences. Review 423. of Palaeobotany and Palynology 95: Rothwell, G. W. and S. Lev-Yadun. 2005. 95-113. 375 million-year-old fossil evidence Rothwell, G. W., and J. E. Mickle. 1982. of polar auxin flow. American Rhetinotheca patens n. sp., a Journal of Botany 92: 903-906. medullosan pollen organ from the Rothwell, G. W., and G. Mapes. 1988. Upper Pennsylvanian of North Vegetation of a Paleozoic conifer America. Review of Palaeobotany community. In: G. Mapes and R. H. and Palynology 36: 361-374. Mapes, eds., Regional geology and Rothwell, G. W., and J. M. Pryor. 1991. paleontology of upper Paleozoic Developmental dynamics of Hamilton quarry area in southeastern arborescent lycophytes; Capical and Kansas, Kansas Geological Survey lateral growth in Stigmaria ficoides. Guidebook Series 6: 213-224. American Journal of Botany 78: Rothwell, G. W. and G. Mapes. 2001. 1740-1745. Barthelia furcata gen. et sp. nov., Rothwell, G. W., H. Sanders, S. E. Wyatt with a review of Paleozoic and S. Lev-Yadun 2008. A fossil coniferophytes and a discussion of record for growth regulation; the role coniferophyte systematics. of auxin in wood evolution. Annals International Journal of Plant of the Missouri Botanical Garden 95: Sciences 162: 637-667. 121-134. Rothwell, G. W. and G. Mapes. 2003. Rothwell, G. W., and S. E. Scheckler. 1988. Validation of the names Biology of ancestral gymnosperms. Utrechtiaceae, Utrechtia, and In: C.B. Beck, Ed. Origin and Utrechtia floriniformis. Taxon 52: Evolution of Gymnosperms. 329-330. Columbia University Press. pp. 85- Rothwell, G. W., G. Mapes, and G.R. 134. Hernandez-Castillo. 2005. Rothwell, G. W., S. E. Scheckler, and W. H. Hanskerpia gen. nov. and Gillespie. 1989. Elkinsia gen. nov., a phylogenetic relationships among the Late Devonian gymnosperms with most ancient conifers (Voltziales) cupulate . Botanical Gazette Taxon 54: 733-750. 150: 170-189. Rothwell, G. W., G. Mapes and R. H. Rothwell, G. W., and A. C. Scott. 1983. Mapes. 1996. Anatomically Coprolites within marattiaceous fern preserved vojnovskyalean seed stems (Psaronius) from the plants in Upper Pennsylvanian Pennsylvanian of the Appalachian (Stephanian) marine shales of North Basin, U.S.A. Palaeogeography, America. Journal of Paleontology Palaeoclimatology, Palaeoecology 70: 1067-1079. 41: 227-232. IOP 110 13 August 2016

Rothwell, G. W., and A. C. Scott. 1988. structures. Botanical Gazette 145: Heterotheca Benson; Lyginopterid 275-291. pollen organs or coprolites? Bulletin Rothwell, G. W., and K. L. Whiteside. 1974. of the British Museum (N.H.), Rooting structures of the Geology 44: 41-43. Carboniferous medullosan Rothwell, G. W., and R. Serbert. 1991. pteridosperms. Canadian Journal of Pollination biology of Elkinsia Botany 52: 97-102. polymorpha, implications for the Rothwell, G. W., S. W. Wyatt and A. M. F. origin of gymnosperms. In: F. Tomescu. 2014. Plant evolution at Schaarschmidt, ed. Special issue: the interface of paleontology and International symposium on developmental biology: an organism palaeobotany; anatomical centered paradigm. American investigations of fossil plants. Cour. Journal of Botany 101: 899-913. Forschungsinst. Senckenb. 147: 225- Sanders, H., G. W. Rothwell and S. E. 231. Wyatt. 2007. A paleontological Rothwell, G. W. and R. A. Stockey. 2008. context for the developmental Phylogeny and evolution of ferns: a mechanisms of evolution. paleontological perspective. P. 332- International Journal of Plant 366. In: T.A. Ranker and Haufler, Sciences 178: 719-728. C.H., eds. The biology and evolution Sanders, H., G. W. Rothwell and S. E. of ferns and lycophytes. Cambridge Wyatt. 2009. Key morphological University Press. alterations in the evolution of . Rothwell, G. W., and T. N. Taylor. 1971. International Journal of Plant Weissistachys kentuckiensis: a new Sciences 170: 860-868. name for Weissia kentuckiense Sanders. H., G. W. Rothwell and S. W. Rothwell and Taylor. Botanical Wyatt. 2011. Parallel evolution of Gazette 132: 371-372. auxin regulation in rooting systems. Rothwell, G. W., and T. N. Taylor. 1972. Plant Systematics and Evolution 291: Carboniferous pteridosperm studies: 221-225. morphology and anatomy of Serbet, R. and G. W. Rothwell. 1992. Schopfiastrum decussatum. Characterizing the most primitive Canadian Journal of Botany 50: seed ferns. I. A reconstruction of 2649-2658. Elkinsia polymorpha. International Rothwell, G. W., T. N. Taylor, and C. Journal of Plant Science 153: 602- Clarkson. 1979. On the structural 621. similarity of the Paleozoic ovules Serbet, R., and G. W. Rothwell. 1995. Conostoma platyspermum and C. Functional morphology and leptospermum. Journal of homologies of gymnospermous Paleontology 53: 49-54. ovules: Evidence from a new species Rothwell, G. W., and S. Warner. 1984. of Stephanospermum (). Cordaixylon dumusum n. sp. Canadian Journal of Botany 73: 650- (Cordaitales). I. vegetative 661. Stubblefield, S. P. 1984. Taxonimic IOP 110 14 August 2016

delimitation among Pennsylvanian Monoletes (Medullosales). American marattialean fructifications. Journal Journal of Botany 69: 570-578. of Paleontology 58: 793-803. Tomescu, A. M. F., R. Honegger and G. W. Stubblefield, S. P., and G. W. Rothwell. Rothwell. 2008. Earliest fossil record 1980. Conostoma chappelicum n. of bacterial-cyanobacterial mat sp., lagenostomalean ovules from consortia: the early Passage Kentucky. Journal of Paleontology Creek biota (440 Ma, Virginia, 54: 1012-1016. USA). Geobiology 6: 210-124. Stubblefield, S. P., and G. W. Rothwell. Tomescu, A. M. F., L. Pratt, G. W. 1981. Embryogeny and reproductive Rothwell, P. K. Strother, G. C. biology of Bothrodendrostrobus Nadon. 2009. Carbon isotopes mundus (Lycopsida). American support the presence of extensive Journal of Botany 68: 625-634. land floras pre-dating the origin of Stubblefield, S. P. and G. W. Rothwell. vascular plants. Palaeogeography, 1989. Cecropsis luculentum gen. Palaeoclimatology, Palaeoecology nov., evidence for heterosporous 283: 46-59. in the Upper Tomescu, A. M. F. and G. W. Rothwell. Pennsylvanian of North America. 2006. Wetlands before American Journal of Botany 76: tracheophytes: Thalloid terrestrial 1,415-1,428. communities of the Early Silurian Stubblefield, S. P., G. W. Rothwell, and T. Passage Creek biota (Virginia). P. N. Taylor. 1984. Conostoma 41-56. In: DiMichele, W.A, and S. williamsonii n. sp., a Greb, eds. Wetland paleoecology lagenostomalean ovule from the through time. Geological Society of Pennsylvanian of the Illinois Basin. America, Special Paper 139. Canadian Journal of Botany 62: 96- Tomescu, A. M. F., G. W. Rothwell and R. 101. Honegger. 2006. Cyanobacterial Stubblefield, S. P., and T. N. Taylor. 1983. macrophytes in an Early Silurian Studies of Paleozoic fungi. 1. The (Llandovery) continental biota: structure and organization of Passage Creek, lower Massanutten Traquaria (Ascomycota). Ameican Sandstone, Virginia, USA. Lethaia Journal of Botany 70: 387-399. 39: 329-338. Stubblefield, S. P., T. N. Taylor, C. I. Miller, Tomescu, A. M. F., G. W. Rothwell and R. and G. T. Cole. 1983. Studies of Honegger. 2009. A new genus and Carboniferous fungi. II. The structure species of filamentous microfossil of and organization of Mycocarpon, cyanobacterial affinity from Early Sporocarpon, Dubiocarpon, and Silurian fluvial environments (lower Coleocarpon (Ascomycotina). Massanutten Sandstone, Virginia, American Journal of Botany 70: USA). Botanical Journal of the 1482-1498. Linnean Society 160: 284-289. Taylor, T. N., and G. W. Rothwell. 1982. Tomescu, A. M. F., G. W. Rothwell and G. Studies of seed fern pollen: Mapes. 2001. Lyginopteris royali sp. Development of the exine in nov. from the Upper Mississippian of IOP 110 15 August 2016

North America. Review of fern. Evolutionary Trends in Plants Palaeobotany and Palynology 116: 2: 25-29. 159-173. Trivett, M. L., and G. W. Rothwell. 1991. Tomescu, A. M. F., G. W. Rothwell and M. Diversity among Paleozoic L. Trivett. 2006. Kaplanopteridaceae Cordaitales. Neus Jahrbuch fur Fam. Nov., additional diversity in the Geol. Palaontol., Stuttgart. 183: 289- initial radiation of filicalean ferns. 305. International Journal of Plant Sciences 167:615-630. Tomescu, A. M. F., G. W. Rothwell and M. PALAEOBOTANY OF ITALY L. Trivett. 2008. Reiterative growth in the complex adaptive architecture of the Paleozoic (Pennsylvanian) filicalean fern Kaplanopteris clavata. Plant Systematics and Evolution 270: 209-216. Trivett, M. L. 1992. Growth architecture, strufture, and relationships of Cordaixylon iowensis nov. comb. (Cordaiatales). International Journal of Plant Sciences 153: 273-287. Trivett, M. L. 1993. An architectural analysis of Archaeopteris, a fossil tree with pseudomonopodial and opportunistic agdventitious growth. Botanical Journal of the Linnean Society 111: 301-329. Trivett, M. L., and G. W. Rothwell. 1985. Morphology, systematics, and paleocology of Paleozoic fossil The volume „Palaeobotany of Italy“ is plants: Mesoxylon priapi, sp. nov. edited by the organizers of the 9th European (Cordaitales). Systemic Botany. 10: Palaeobotany and Palynology Congress in 205-223. Padova, Italy (EPPC2014). Italy’s Trivett, M. L., and G. W. Rothwell. 1988. palaeobotanical record is extensive. Diversity among Paleozoic However, this heritage has largely been Cordaitales: The vascular forgotten. More than 300 million years lie architecture of Mesoxylon birame between the oldest plant fossils discovered Baxter. Botanical Gazette 149: 116- in Italy and Quaternary plant remains found 125. in archaeological excavations. Fossil floras Trivett, M. L., and G. W. Rothwell. 1988. throughout Italy are remarkable in that they Modeling the growth architecture of show a surprising abundance and diversity fossil plants: A Paleozoic Filicalean over the millions of years. The Italian palaeobotanical heritage represents an IOP 110 16 August 2016 important source of new information on the evolution of plants and the ecosystems in which they lived, but can also help in predicting future environmental scenarios. The present volume is the first general book on fossil floras of Italy; it is bilingual with an Italian and English text. The book is divided in a detailed overview on the history of palaeobotany in Italy, a general part on materials and methods in palaeo- and archeobotany and nine chronological chapters (from the Carboniferous to Archeobotany). Each chapter is divided in geographic areas as well as in macro- and microfloras. Additional information are The 10th European Palaeobotany and given on the institutions different collections Palynology Conferencewill be held in are stored in. This book provides a state of Dublin on the 12th to 19th of August 2018 the art of the different floras with an at University College Dublin. Partner exhaustive reference list. organizations include Trinity College Dublin Since the first volume was sold out within and the National Museum of Ireland. We two years, a second, slightly corrected look forward to welcoming you in 2018 version has been re-printed. Jennifer McElwain, Earth Institute, Bibliographic information: University College Dublin Chair of EPPC organizational committee La storia delle piante fossili in Italia, seconda edizione - Paleobotany of Italy, second edition Evelyn Kustatscher, Guido Roghi, Adele 25th International Workshop on Plant Bertini & Antonella Miola (eds.) Taphonomy 2015, November 25 - 26, Veröffentlichungen des Naturmuseums 2015, Bonn, Germany Südtirol, volume 9 pp. 400, 20.6 x 27.6 cm, Euro 25,00 Bozen/Bolzano, Naturmuseum The 25th International Workshop on Plant Südtirol/Museo di Scienze Naturali dell’Alto Taphonomy will be held on the last weekend Adige 2016 in November in Bonn, from Friday, ISBN: 978-88-87108-09-5 November 25 to Saturday, November 26, 2016. The date provides terrific UPCOMING MEETINGS opportunities for stimulating discussions over Glühwein at the Bonn Christmas The 10th European Palaeobotany and Market, so please mark it on your calendar! Palynology Conference The 2016 workshop will feature a keynote lecture from Robert Gastaldo (Colby IOP 110 17 August 2016

College, Maine, USA) on “Time, Space, Sedimentary Systems, and Some Thoughts on Geochemistry of Why, or Not, We Have A Fossil Plant Record,” as well as a round-table discussion on the taphonomy of leaf cuticle led by Lutz Kunzmann (Senckenberg Naturhistorische Sammlungen Dresden, Germany). Contributed talks and posters on all aspects of plant taphonomy are welcome.

Although there will be formal presentations, we hope that the meeting will retain its informal, discussion-oriented, workshop flavor, and students and new members wanting to check out plant taphonomy are especially wanted. The deadline for pre-registration and abstracts is October 1, and more information will be contained in a second mailing at the end of summer. If you are interested in receiving this second circular or if you have any questions, please send an email to Carole Gee ([email protected]).

Carole Gee and Thomas Litt University of Bonn Germany

CALL FOR NEWS AND NOTES

Please send submissions for the next news letter by 15 September 2016 to:

Mike Dunn Department of Biological Sciences Cameron University Lawton, Oklahoma 73505 Ph.: 580-581-2287 email: [email protected]