Grandifrons and F Geijskesii

Total Page:16

File Type:pdf, Size:1020Kb

Grandifrons and F Geijskesii J Hattori Bot. lab. No. 75: 15- 22 (Feb. 1994) A RE-EVALUATION OF FISSJDENS SUBGENUS PACHYFISSJDENS, WITH A DETAILED DISCUSSION OF FISSIDENS GRANDIFRONS AND F GEIJSKESII 1 2 RONALD A. PURSELL AND BRUCE H. ALLEN ABSTRACT . Subgenus Pachyfissidens, lectotypified by Fissidens grandifrons Brid., is based essen­ tially on plants with stiff leaves and pluristratose laminal cells. It is an unnatural and taxonomically superfluous taxon. Species assigned to this subgenus are placed in subgenus Fissidens section Aloma C. Miill. (Fissidens sedgwickii Broth. & Dix.), section Amblyothallia C. Miill. (Fissidens grandifrons Brid.), section Crispidium C. Miill. (Fissidens strictus Hook. f. & Wits.), and section Fissidens (Fissi­ dens rochensis Broth. and F. ventricosus Lesq.), subgenus Serridium (C. Miill.) lwats. (Fissidens boninensis lwats., F. jaiorum Whitt. & Mill., and F. perdecurrens Besch.), and subgenus Sarawakia (C. Miill.) lwats. (Fissidens geijskesii Florsch.). Pachyfissidens, established as a section of Fissidens Hedw. by Muller (l 848), was rec­ ognized as a subgenus by Kindberg ( 1897) and later accepted at this level by Brotherus ( 1901, 1924) in his widely adopted system of classfication. The taxon is distinguished by its rigid leaves, pluristratose laminal cells, an absence of a central strand in the stem, and estomate capsules. Pachyfissidens is here lectotypified with Fissidens grandifrons Brid., Sp. Muse. I : 170. 1806. (Lectotype. In Nova Anglia habitat, without collector, B). Bruggeman-Nannenga (1974) demonstrated that the pluristratose lamina! cells and lack of a central strand are also found in species that unquestionably belong to subgenus Fissidens. Furthermore, estomate capsules are found in the subgenera Octodiceras (Brid.) Broth. and Sarawakia (C. Miill.) lwats., with both stomate and estomate capsules occuring in some members of the latter subgenus. In Sarawakia the estomate condition was consid­ ered a feature influenced by habitat (Pursell et al. 1988). Nevertheless, despite its lack of any uniquely defining characters, Bruggeman-Nannenga maintained Pachyfissidens as a section in subgenus Fissidens. Recent data derived from SEM studies on the p~ristome (Allen 1980, Bruggeman­ Nannenga & Berendsen 1990) and light-microscope studies on the costa (Bruggeman-Nan­ nenga 1990, Stone 1990) have been shown to be valuable in understanding the infra-gener­ ic relationships of Fissidens. These two features frequently appear coordinated throughout Fissidens and are here used in a re-examinations of Pachyfissidens. Pachyfissidens throughout its history has been a depository for species that share a similar type of habitat, i.e., submersion in rapidly running water. The principal features of this group appear to represent a cluster of convergent characters rather than a suite of coor­ dinated characters indicative of a natural relationship. Nine species of Fissidens are 1 Department of Biology, 208 Mueller Laboratory, The Pennsylvania State University, University Park, Pennsylvania 16802- 530 I, U.S.A. 2 Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166-0299, U.S.A. 16 J. Hattori Bot. Lab. No. 75 I 9 9 4 presently placed in Pachyfissidens, i.e., F boninensis Iwats., F geijskesii Florsch., F gran­ difrons Brid., F jaiorum Whitt. & Mill., F rochensis Broth., F perdecurrens Besch., F sedgwickii Broth. & Dix., F strictus Hook. f. and Wils., and F ventricosus Lesq. The pur­ pose of this paper is to discuss these species and their taxonomic positions. Fissidens grandifrons is a widely dispersed aquatic species in the Northern Hemi­ sphere. Although gametophytically specialized for an aquatic habitat, e.g., the long-stalked archegonia (lwatsuki & Suzuki 1982) and thick imbricate leaves, the species has retained a terrestrial sporophyte, i.e., one having a long exserted seta. Such a sporophyte usually has stomata in the capsule. However, we have not been able to demonstrate the presence of stomata in the few sporophytes of F. grandifrons available for study. But, this combination of terrestrial and aquatic sporophytic features is not unique. The same combination of fea­ tures can be found in Dichelyma Myr., a genus in the Fontinalaceae with setae up to 20 mm in length and estomate capsules. The peristome of F. grandifrons has been described as belonging to the taxifolius-type (Bruggeman-Nannenga & Berendsen 1990). This type is commonly found in section Ser­ ridium C. Miill. [=subgenus Serridium (C. Miill.) Iwats.], but it also occurs in section Am­ blyothallia C. Miill. There is, however, no concensus on the peristome types in these two sections. Allen (1980) concluded that species of both sections had the same basic type, i.e. the taxifolius-type. Bruggeman-Nannenga and Berendsen (1990), on the other hand, recog­ nized two basic types of peristomes in these two sections, the taxifolius-type and the sim­ iliretis-type, with the former having three variations. They considered the taxifolius-type characteristic of Serridium while the similiretis-type was found mainly in Amblyothallia. However, these types of peristome were not restricted to one section or the other. To the contrary, some species of Serridium were found to possess a similiretis-type peristome and species of Amblyothallia were found with a taxifolius-type peristome. Recently, Stone (1990), upon examining some Australian species, thought that only one type of peristome was represented in Serridium and Amblyothallia. The lanceolate leaves of F. grandifrons (Fig.1) are usually tightly imbricate and stiff. A recent examinations of specimens from sites throughout the range of the species, however, has revealed that there is considerable variation in leaf thickness and costal structure. Mar­ ginal lamina! cells in the distal part of the leaf can be unistratose (Figs. 2, 3, 8, 12, 14), bis­ tratose (Figs. 6, 10), and even quadristratose (Fig. 5). The interior lamina! cells can be bis­ tratose (Fig. 14), pentastratose (Fig. I 0), hexastratose (Fig. 12) to pluristratose (Fig. 5). Moreover, the subsurface lamina! cells are generally larger than the surface cells. The basic structure of the costa includes three stereid bands in the proximal part of the leaf. A ventral and two lateral stereid bands are evident in Figs. 3, 4, and 13 (the ventral band in this last figure is interrupted) and also in the illustration in lwatsuki and Suzuki (1982, Plate LIII). The ventral band can be reduced to a single file of stereid cells (Figs. 7, 15) or it can be missing (Figs. 9, 11). There are basically two stereid bands in the distal part of the leaf (Figs. 6, 8, 12), although one can be lost (Figs. 10, 14), or both can be eliminated (Fig. 5), in which case the costa consists of more or less homogeneous cells. In both the proximal and distal parts of the leaf the lateral stereid bands may fuse (Figs. 2, 11 ; see also Fig. 45F R. A. PURSELL & B. H. AL LEN: A re-evaluation of Fissidens subg. Pachyjissidens 17 Figs. 1- 5. Fissidens grandifrons Brid. 1. Habit sketch of a leaf. 2, 5. Transverse sec­ tions of distal part of leaf. 3- 5. Transverse sections of proximal part of leaf. [1 - 3 from Pursell, Allen & Magill 10984, PAC; 4-5 from Frye: Moss Exsiccati 25, PAC] 18 J. Hattori Bot. Lab. No. 75 1 9 9 4 0 s~. ?o~ § 9 gO•Oo' • o' · ·~ ~·• •• o :·o~ o ~~a ~0r- 6 15 a8Q'.j 0 JJfm 14 ~D.o~··@ Q ~o o ~ 'J/013 12 Figs. 6-15. Fissidens grandifrons Brid. 6, 8, 10, 12, 14. Transverse sections of distal part of leaf. 7, 9, 11, 13, 15. Transverse sections of proximal part of leaf. [6-7 from Wilson 1909, PAC; 8- 9 from Sharp, Clebsch & Thornburgh 2230, PAC; 10- 11 from Bruggeman­ Nannenga 406, PAC; 12- 13 from Lin 12831 , HIRO; 14- 15 from Higuchi 20162, HIRO] R. A . PURSELL & B. H. A t.,LEN : A re-evaluation of Fissidens subg. Pachyfissidens 19 in Crum & Anderson 1981 ). The type of costa that is found in F grandifrons has been shown (Bruggeman-Nannen­ ga 1990, Stone 1990) to be a unique feature of section Amblyotha/lia. Most species of Fis­ sidens have a costa that has two stereid bands throughout its length, but in Amblyotha/lia there are three stereid bands in the proximal half of the leaf, two lateral and one ventral, separated usually by more than three enlarged cells (we include in this term 'guide cells,' 'deuters,' 'conjunctivae,' and 'conductivae' of authors). Therefore, in view of its costal structure, F grandifrons is hereby transferred to section Amblyotha/lia. Specimens examined. Taiwan. Pingtung Co. : Lai 9850 (NY); Taichung Co.: Lai 8699, c.fr. (NY); Taitung Co.: Lai 9651 (NY), Lin 12831, c.fr. (HIRO). Pakistan. Northwest Frontier Prov.: Haghan Valley, Higuchi 20162, c.fr. (HIRO). France. Hautes-Pyrenees: Bruggeman-Nannenga 406 (PAC). U.S.A. California. Mendocino Co. : Wilson 1909 (PAC). Idaho. Fremont Co. (?):Frye: Moss Exsiccati 25 (PAC). Missouri. St. Frarn;ois Co.: Pursell, Allen & Magi/110984 (PAC). Mexico.: Oax­ aca: Sharp, Clebsch & Thornburgh 2230 (PAC). Fissidens geijskesii is known from only the type collections made in Suriname, north­ ern South America. Its leaves (Fig. 16) are similar in shape, arrangement, and texture to those of F. grandifrons. The costa of F. geijskesii in transverse section, however, reveals two stereid bands throughout its length (Figs. 17, 18), separated distally by two enlarged cells and proximally by three or more enlarged cells. The costa is similar to the bryoides-type (Bruggeman-Nannenga 1990) which is found in all groups of Fissidens except sections Amblyothallia and Crispidium C. Miill., and subgenus Serridium. The sporophyte of F. geijskesii has an immersed estomate capsule and a very short seta (0.4 mm long), a feature found only in the aquatic subgenera Octodiceras and Sarawakia. The peristome of F geijskesii is fragile and missing from the holotype, paratypes, and most isotypes. There is, however, an isotype in LAF in which the peristomes are intact. The teeth are reduced, i.e., undivided or irregularly divided, and are papillose throughout their length.
Recommended publications
  • About the Book the Format Acknowledgments
    About the Book For more than ten years I have been working on a book on bryophyte ecology and was joined by Heinjo During, who has been very helpful in critiquing multiple versions of the chapters. But as the book progressed, the field of bryophyte ecology progressed faster. No chapter ever seemed to stay finished, hence the decision to publish online. Furthermore, rather than being a textbook, it is evolving into an encyclopedia that would be at least three volumes. Having reached the age when I could retire whenever I wanted to, I no longer needed be so concerned with the publish or perish paradigm. In keeping with the sharing nature of bryologists, and the need to educate the non-bryologists about the nature and role of bryophytes in the ecosystem, it seemed my personal goals could best be accomplished by publishing online. This has several advantages for me. I can choose the format I want, I can include lots of color images, and I can post chapters or parts of chapters as I complete them and update later if I find it important. Throughout the book I have posed questions. I have even attempt to offer hypotheses for many of these. It is my hope that these questions and hypotheses will inspire students of all ages to attempt to answer these. Some are simple and could even be done by elementary school children. Others are suitable for undergraduate projects. And some will take lifelong work or a large team of researchers around the world. Have fun with them! The Format The decision to publish Bryophyte Ecology as an ebook occurred after I had a publisher, and I am sure I have not thought of all the complexities of publishing as I complete things, rather than in the order of the planned organization.
    [Show full text]
  • Part 2 – Fruticose Species
    Appendix 5.2-1 Vegetation Technical Appendix APPENDIX 5.2‐1 Vegetation Technical Appendix Contents Section Page Ecological Land Classification ............................................................................................................ A5.2‐1‐1 Geodatabase Development .............................................................................................. A5.2‐1‐1 Vegetation Community Mapping ..................................................................................... A5.2‐1‐1 Quality Assurance and Quality Control ............................................................................ A5.2‐1‐3 Limitations of Ecological Land Classification .................................................................... A5.2‐1‐3 Field Data Collection ......................................................................................................... A5.2‐1‐3 Supplementary Results ..................................................................................................... A5.2‐1‐4 Rare Vegetation Species and Rare Ecological Communities ........................................................... A5.2‐1‐10 Supplementary Desktop Results ..................................................................................... A5.2‐1‐10 Field Methods ................................................................................................................. A5.2‐1‐16 Supplementary Results ................................................................................................... A5.2‐1‐17 Weed Species
    [Show full text]
  • Bryophyte Ecology Table of Contents
    Glime, J. M. 2020. Table of Contents. Bryophyte Ecology. Ebook sponsored by Michigan Technological University 1 and the International Association of Bryologists. Last updated 15 July 2020 and available at <https://digitalcommons.mtu.edu/bryophyte-ecology/>. This file will contain all the volumes, chapters, and headings within chapters to help you find what you want in the book. Once you enter a chapter, there will be a table of contents with clickable page numbers. To search the list, check the upper screen of your pdf reader for a search window or magnifying glass. If there is none, try Ctrl G to open one. TABLE OF CONTENTS BRYOPHYTE ECOLOGY VOLUME 1: PHYSIOLOGICAL ECOLOGY Chapter in Volume 1 1 INTRODUCTION Thinking on a New Scale Adaptations to Land Minimum Size Do Bryophytes Lack Diversity? The "Moss" What's in a Name? Phyla/Divisions Role of Bryology 2 LIFE CYCLES AND MORPHOLOGY 2-1: Meet the Bryophytes Definition of Bryophyte Nomenclature What Makes Bryophytes Unique Who are the Relatives? Two Branches Limitations of Scale Limited by Scale – and No Lignin Limited by Scale – Forced to Be Simple Limited by Scale – Needing to Swim Limited by Scale – and Housing an Embryo Higher Classifications and New Meanings New Meanings for the Term Bryophyte Differences within Bryobiotina 2-2: Life Cycles: Surviving Change The General Bryobiotina Life Cycle Dominant Generation The Life Cycle Life Cycle Controls Generation Time Importance Longevity and Totipotency 2-3: Marchantiophyta Distinguishing Marchantiophyta Elaters Leafy or Thallose? Class
    [Show full text]
  • Article ISSN 2381-9685 (Online Edition)
    Bry. Div. Evo. 39 (1): 075–093 ISSN 2381-9677 (print edition) DIVERSITY & http://www.mapress.com/j/bde BRYOPHYTE EVOLUTION Copyright © 2017 Magnolia Press Article ISSN 2381-9685 (online edition) https://doi.org/10.11646/bde.39.1.12 Diversity of the rheophytic condition in bryophytes: field observations from multiple continents JAMES R. SHEVOCK1, WEN-ZHANG MA2 & HIROYUKI AKIYAMA3 1Department of Botany, California Academy of Sciences, 55 Music Concourse Dr., Golden Gate Park, San Francisco, California 94118, U.S.A. 2Herbarium, Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sci- ences, Kunming, Yunnan 650201, China 3Museum of Nature & Human Activities, Hyogo, Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka-6, Sandi-shi, Hyogo 669-1546, Japan Abstract Bryophytes occurring in riparian systems where they are seasonally submerged or inundated are poorly documented in many parts of the world. The actual number of rheophytic bryophytes remains speculative but we believe the number could easily exceed 500 taxa. Rheophytic bryophytes generally display highly disjunct populations and adjacent rivers and streams can have considerably different species composition. Water management in the form of flood control, dams, and hydroelectric development can adversely impact many rheophytic bryophyte species and communities due to changes in river ecology, timing of water flow, and water temperature. Specimens of rheophytic bryophytes are underrepresented in herbaria and la- bels rarely indicate the actual micro-habitat and ecological attributes for bryophytes collected within riparian systems. Many rheophytes are morphological anomalies compared to their terrestrial relatives and the evolution of the rheophytic condition has occurred repeatedly in many bryophyte lineages.
    [Show full text]
  • TWO NEW Fissidens SPECIES (FISSIDENTACEAE) FROM
    Polish Botanical Journal 58(1): 117–125, 2013 DOI: 10.2478/pbj-2013-0013 TWO NEW FISSIDENS SPECIES (FISSIDENTACEAE) FROM TROPICAL EASTERN AfRICA AND FISSIDENS BESSOUENSIS CORB., A REMARKABLE SPECIES WITH FIMBRIATE LEAVES AND MULTICELLULAR SPORES1 MARIA ALIDA BRUGGE M AN -NANNENGA Abstract. Fissidens acrophilus Bruggeman-Nannenga, sp. nov. (subgenus Fissidens) from Kenya and Uganda and F. obscuri- frons Bruggeman-Nannenga, sp. nov. (subgenus Fissidens?) from Tanzania are newly described and figured. F. obscurifrons is compared to the little known Madeiran F. nobreganus Luisier & P. de la Varde. A revision of F. bessouensis Corb. (subgenus Octodiceras) with description, diagnosis and figures and a discussion including adaptations to an aquatic habitat is presented. Multicellular spores and fimbriate leaf margins are reported in the Fissidentaceae for the first time. Rectification: F. anguste- limbatus Mitt. is eliminated from the African flora. Key words: Taxonomy, Africa, Fissidens acrophilus spec. nov., Fissidens obscurifrons spec. nov., Fissidens nobreganus, Fis- sidens bessouensis, fimbriate leaves and multicellular spores in Fissidens Maria Alida Bruggeman-Nannenga, Griffensteijnseplein 23, 3703 BE Zeist, The Netherlands While preparing a paper on subgenus Fissidens 3960 m. 17 January 1992. Kungu 9222/AD (HOLO T YPE : in tropical eastern Africa two new species were L; ISO T YPE : EGR). discovered. Both are from wet habitats. Fissidens DESCRIP T ION (perigonial data from paratype). acrophilus Bruggeman-Nannenga, sp. nov. is a lim- Growing in mats; stem with central strand of bate species with smooth cells found on Mt. Elgon ± 10 cells, with 1–3 tiers of inner and 1–3 tiers (Kenya and Uganda) between 3250 and 3960 m.
    [Show full text]
  • Mosses of Bhutan II*. a Checklist of the Mosses of Bhutan
    Journal of Bryology ISSN: 0373-6687 (Print) 1743-2820 (Online) Journal homepage: http://www.tandfonline.com/loi/yjbr20 Mosses of Bhutan II*. A checklist of the mosses of Bhutan D. G. Long To cite this article: D. G. Long (1994) Mosses of Bhutan II*. A checklist of the mosses of Bhutan, Journal of Bryology, 18:2, 339-364, DOI: 10.1179/jbr.1994.18.2.339 To link to this article: http://dx.doi.org/10.1179/jbr.1994.18.2.339 Published online: 18 Jul 2013. Submit your article to this journal Article views: 20 View related articles Citing articles: 5 View citing articles Full Terms & Conditions of access and use can be found at http://www.tandfonline.com/action/journalInformation?journalCode=yjbr20 Download by: [British Bryological Society] Date: 09 May 2017, At: 04:09 Journal of Bryology (1994) 18: 339-364 Mosses of Bhutan 11*. A checklist of the mosses of Bhutan DAVID G. LONG Royal Botanic Garden, Edinburgh, U.K. SUMMARY A checklist is given of the mosses of Bhutan based on published reports. Totals of 156 genera and 282, species are recorded. The main past collectors of mosses in Bhutan are listed, together with sources of erroneous reports; 44 erroneous and doubtful taxa are excluded. The name Breutelia setschwanica is applied to the only Himalayan mem- ber of the genus. The name Barbula [lavicans is proposed as an avowed substitute for the illegitimate Tortula [lavescens Hook. & Grev. (Barbula fuscescens C. Mull. nom. inval.) KEYWORDS: Mosses, checklist, Bhutan, Himalaya, Breutelia setschwanica, Barbula [lavicans. INTRODUCTION Of all the countries and states in the Himalayan region, Bhutan is probably one of the richest bryologically yet one of the least-known.
    [Show full text]
  • An Updated Checklist of Mosses of Xinjiang, China1 Новый Список Мхов Синьцзяня, Китай1
    Arctoa (1995) 4: 1-14 AN UPDATED CHECKLIST OF MOSSES OF XINJIANG, CHINA1 НОВЫЙ СПИСОК МХОВ СИНЬЦЗЯНЯ, КИТАЙ1 BENITO C. TAN2, JIAN-CHENG ZHAO3 AND REN-LIANG HU4 Abstract An updated checklist of Xinjiang mosses based on past publications and new collec- tions made in 1993 showed a total of 130 genera, 339 species and 8 infraspecific taxa. Twelve species and two varieties are reported here as new to China. They are Amblyodon dealbatus, Bryum schleicheri var. latifolium, Conardia compacta, Cynodontium schistii, Dicranella schreberiana, Orthotrichum cupulatum, O. hallii, O. pallens, O. speciosum var. elegans, Physcomitrium pyriforme, Pottia bryoides, P. heimii, Sphagnum warnstorfii and Tortella nitida. Amblyodon and Conardia are two new moss generic records for China. Резюме Новый список листостебельных мхов Синьцзян-Уйгурского автономного округа (северо-западный Китай) основан на последних публикациях и сборах авторов в 1993 году. Список включает 339 видов и 8 внутривидовых таксонов из 130 родов. Двенадцать видов и две разновидности приводятся впервые для Китая: Amblyodon dealbatus, Bryum schleicheri var. latifolium, Conardia compacta, Cynodontium schistii, Dicranella schreberiana, Orthotrichum cupulatum, O. hallii, O. pallens, O. speciosum var. elegans, Physcomitrium pyriforme, Pottia bryoides, P. heimii, Sphagnum warnstorfii и Tortella nitida. Два рода, Amblyodon и Conardia, найдены в Китае впервые. INTRODUCTION Himalayan Range, moist air currents from the The study area, Xinjiang Uygur Autonomous Indian Ocean fail to reach Xinjiang, resulting in Region, is the largest province of China, repre- the vast expanse of arid terrain. Hydrologically, senting roughly 1/6 of the national territory. It there is a progressive increase of precipitation is situated in the northwestern corner of the coun- from south to north, with the Altai region re- try (34° 15'-49° 11' N, 73° 21'-96° 25' E) with a ceiving the highest annual precipitation.
    [Show full text]
  • An Updated List of Mosses of Korea
    Journal of Species Research 9(4):377-412, 2020 An updated list of mosses of Korea Wonhee Kim1,*, Masanobu Higuchi2 and Tomio Yamaguchi3 1National Institute of Biological Resources, 42 Hwangyeong-ro, Seo-gu, Incheon 22689 Republic of Korea 2Department of Botany, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba 305-0005 Japan 3Program of Basci Biology, Graduate School of Integrated Science for Life, Hiroshima University, 1-3-1 Kagamiyama, Higashi-hiroshima-shi 739-8526 Japan *Correspondent: [email protected] Cardot (1904) first reported 98 Korean mosses, which were collected from Busan, Gangwon Province, Mokpo, Seoul, Wonsan and Pyongyang by Father Faurie in 1901. Thirty-four of these species were new species to the world. However, eight of these species have been not listed to the moss checklist of Korea before this study. Thus, this study complies the literature including Korean mosses, and lists all the species there. As the result, the moss list of Korea is updated as including 775 taxa (728 species, 7 subspecies, 38 varieties, 2 forma) arranged into 56 families and 250 genera. This list include species that have been newly recorded since 1980. Brachythecium is the largest genus in Korea, and Fissidens, Sphagnum, Dicranum and Entodon are relatively large. Additionally, this study cites specimens collected from Jeju Island, Samcheok, Gangwon Province, and Socheong Island, and it is possible to confirm the distribution of 338 species in Korea. Keywords: bryophytes, checklist, Korea, mosses, updated‌ Ⓒ 2020 National Institute of Biological Resources DOI:10.12651/JSR.2020.9.4.377 INTRODUCTION Choi (1980), Park and Choi (2007) reported a “New List of Bryophytes in Korea” by presenting an overview of The first study on Korean bryophytes was published by bryophytes surveyed in Mt.
    [Show full text]
  • Prepared For
    Trans Mountain Pipeline ULC Volume 5C, ESA - Biophysical Technical Reports Trans Mountain Expansion Project Vegetation Technical Report APPENDIX B POTENTIALS TABLES 7894/December 2013 REP-NEB-TERA-00008 Page B-1 TABLE B1 POTENTIAL RARE PLANT AND LICHEN SPECIES IN THE BOREAL FOREST – CENTRAL MIXEDWOOD AND DRY MIXEDWOOD, FOOTHILLS – LOWER FOOTHILLS, PARKLAND – CENTRAL PARKLAND NATURAL, ROCKY MOUNTAIN – MONTANE SUBREGIONS OF THE PROPOSED PIPELINE PROJECT Right-of-Way Preferred Habitat Within Known on Proposed Provincial Federal/Global Scientific Name Common Name Habitat Species Range Right-of-Way Designations Designations VASCULAR PLANTS Adenocaulon bicolor pathfinder Moist woods and thickets. Flowering from June to September. Yes Yes S21 -- Adiantum aleuticum western maidenhair Moist forests, rocks, rocky scree and banks. Sporulating from summer to fall. Yes Yes S21 -- fern Agrostis exarata spike redtop Moist slopes and open areas; usually areas that hold snow late in the growing -- Yes S21 -- season. Flowering from late June to August. Agrostis humilis low bent grass Moist slopes and meadows. Flowering from August to September. Yes Yes S21 -- Agrostis mertensii northern bent-grass Moist slopes. Flowering from July to August. Yes Yes S21 -- Allium geyeri Geyer's onion Wet meadows and streambanks. Flowering from June to July. -- Yes S21 -- Almutaster pauciflorus few-flowered aster Saline soils, saline shores and depressions. Flowering from spring to fall. Yes -- S2S31 -- Alopecurus alpinus alpine foxtail Shores and open woodland. Flowering from June to August. -- Yes S2?1 -- Anemone quinquefolia wood anemone Moist woods. Flowering in July. Yes Yes S11 -- Antennaria aromatica scented everlasting Limestone talus. Flowering from July to early August. -- -- S21 -- 1 Page B Page Antennaria corymbosa corymbose Open woods and meadows.
    [Show full text]
  • Checklist and Country Status of European Bryophytes – Towards a New Red List for Europe
    ISSN 1393 – 6670 Checklist and country status of European bryophytes – towards a new Red List for Europe Cover image, outlined in Department Green Irish Wildlife Manuals No. 84 Checklist and country status of European bryophytes – towards a new Red List for Europe N.G. Hodgetts Citation: Hodgetts, N.G. (2015) Checklist and country status of European bryophytes – towards a new Red List for Europe. Irish Wildlife Manuals, No. 84. National Parks and Wildlife Service, Department of Arts, Heritage and the Gaeltacht, Ireland. Keywords: Bryophytes, mosses, liverworts, checklist, threat status, Red List, Europe, ECCB, IUCN Swedish Speices Information Centre Cover photograph: Hepatic mat bryophytes, Mayo, Ireland © Neil Lockhart The NPWS Project Officer for this report was: [email protected] Irish Wildlife Manuals Series Editors: F. Marnell & R. Jeffrey © National Parks and Wildlife Service 2015 Contents (this will automatically update) PrefaceContents ......................................................................................................................................................... 1 1 ExecutivePreface ................................ Summary ............................................................................................................................ 2 2 Acknowledgements 2 Executive Summary ....................................................................................................................................... 3 Introduction 3 Acknowledgements ......................................................................................................................................
    [Show full text]
  • Rare, Threatened and Endangered Species of Oregon
    RARE, THREATENED AND ENDANGERED SPECIES OF OREGON OREGON NATURAL HERITAGE INFORMATION CENTER May 2004 Oregon Natural Heritage Information Center Institute for Natural Resources Oregon State University 1322 SE Morrison Street Portland, OR 97214-2531 (503) 731-3070 http://oregonstate.edu/ornhic/ With assistance from: Native Plant Society of Oregon The Nature Conservancy Oregon Department of Agriculture Oregon Department of Fish and Wildlife Oregon Department of State Lands Oregon Natural Heritage Advisory Council U.S. Fish and Wildlife Service U.S. Forest Service Bureau of Land Management Compiled and Published by the staff of the Oregon Natural Heritage Information Center: Jimmy Kagan, Director/Ecologist Eric Scheuering, Zoology Data Manager Sue Vrilakas, Botany Data Manager John Christy, Wetlands Ecologist, Bryologist Eleanor Gaines, Zoologist Jon Hak, GIS Program Manager Cliff Alton, Data Output/IS Manager Claudine Tobalske, GIS Analyst Fern McArthur, Botany Data Handler Annie Weiland, Zoology Data Assistant Kuuipo Walsh, GIS Analyst Theresa Koloszar, Office Manager/Grants Specialist Cover Illustration: Pink sand-verbena (Abronia umbellata ssp. breviflora) by Diane Bland. Bibliographic reference to this publication should read: Oregon Natural Heritage Information Center. 2004. Rare, Threatened and Endangered Species of Oregon. Oregon Natural Heritage Information Center, Oregon State University, Portland, Oregon. 105 pp. CONTENTS Introduction............................................................................................................................................................1
    [Show full text]
  • A Miniature World in Decline: European Red List of Mosses, Liverworts and Hornworts
    A miniature world in decline European Red List of Mosses, Liverworts and Hornworts Nick Hodgetts, Marta Cálix, Eve Englefield, Nicholas Fettes, Mariana García Criado, Lea Patin, Ana Nieto, Ariel Bergamini, Irene Bisang, Elvira Baisheva, Patrizia Campisi, Annalena Cogoni, Tomas Hallingbäck, Nadya Konstantinova, Neil Lockhart, Marko Sabovljevic, Norbert Schnyder, Christian Schröck, Cecilia Sérgio, Manuela Sim Sim, Jan Vrba, Catarina C. Ferreira, Olga Afonina, Tom Blockeel, Hans Blom, Steffen Caspari, Rosalina Gabriel, César Garcia, Ricardo Garilleti, Juana González Mancebo, Irina Goldberg, Lars Hedenäs, David Holyoak, Vincent Hugonnot, Sanna Huttunen, Mikhail Ignatov, Elena Ignatova, Marta Infante, Riikka Juutinen, Thomas Kiebacher, Heribert Köckinger, Jan Kučera, Niklas Lönnell, Michael Lüth, Anabela Martins, Oleg Maslovsky, Beáta Papp, Ron Porley, Gordon Rothero, Lars Söderström, Sorin Ştefǎnuţ, Kimmo Syrjänen, Alain Untereiner, Jiri Váňa Ɨ, Alain Vanderpoorten, Kai Vellak, Michele Aleffi, Jeff Bates, Neil Bell, Monserrat Brugués, Nils Cronberg, Jo Denyer, Jeff Duckett, H.J. During, Johannes Enroth, Vladimir Fedosov, Kjell-Ivar Flatberg, Anna Ganeva, Piotr Gorski, Urban Gunnarsson, Kristian Hassel, Helena Hespanhol, Mark Hill, Rory Hodd, Kristofer Hylander, Nele Ingerpuu, Sanna Laaka-Lindberg, Francisco Lara, Vicente Mazimpaka, Anna Mežaka, Frank Müller, Jose David Orgaz, Jairo Patiño, Sharon Pilkington, Felisa Puche, Rosa M. Ros, Fred Rumsey, J.G. Segarra-Moragues, Ana Seneca, Adam Stebel, Risto Virtanen, Henrik Weibull, Jo Wilbraham and Jan Żarnowiec About IUCN Created in 1948, IUCN has evolved into the world’s largest and most diverse environmental network. It harnesses the experience, resources and reach of its more than 1,300 Member organisations and the input of over 10,000 experts. IUCN is the global authority on the status of the natural world and the measures needed to safeguard it.
    [Show full text]