DOCSLIB.ORG

Ca PLT Fampapavransaxrose

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Ca PLT Fampapavransaxrose California Plant Families Santa Barbara Botanic Garden Mary Carroll Vascular Plant Cladogram Flowering Plants Monosulcate pollen (1 groove)) Tricolpate pollen (3 furrows, different orientation than monosulcate) Platanaceae (flowers from Wikipedia) Flowers small, inconspicuous, hypogenous, regular, unisexual, receptacle short, smooth, hypogynous disk absent, perianth reduced, sepals number three to four, rarely eight, free or basally fused, shorter than the petals, triangular. Petals number three to four, rarely eight, truncated-spatulate or vestigial, scarious, frequently absent in the female flowers, male flowers with androecium haplostemonous, isostemonous, oppositisepal, with three to four, rarely eight, stamens, gynostemium short or vestigial, anthers basifixed, not versatile, dithecous, tetrasporangiate, elongated, connectivum apically widened, peltate, dehiscence along longitudinal valves; pistillidium sometimes present, female flowers with superior gynoecium carpels apocarpous in two or three whorls, imperfectly closed apically, surrounded by petals, linear stylodious, stigmas internal, decurrent in two ridges, more or less dry, two ovules per carpel but one nearly always aborts, orthotropous, bitegmic, crassinucellated, pendulous, apical to marginal placentation, three or four staminodes 3-4, no nectaries Ranunculaceae herbaceous, rarely woody (and then a vine) Ranunculaceae free perianth parts, radial symmetry in most Ranunculaceae free perianth parts, radial symmetry in most Ranunculaceae free perianth parts, bilateral symmetry in very few Ranunculaceae sepals somemes absent Ranunculaceae numerous free stamens Pisls = gynoecium –> 3 opons (unicarpellate, apocarpous, syncarpous) Ranunculaceae apocarpous Tricolpate pollen (3 furrows, different orientation than monosulcate) Platanaceae (flowers from Wikipedia) Flowers small, inconspicuous, hypogenous, regular, unisexual, receptacle short, smooth, hypogynous disk absent, perianth reduced, sepals number three to four, rarely eight, free or basally fused, shorter than the petals, triangular. Petals number three to four, rarely eight, truncated-spatulate or vestigial, scarious, frequently absent in the female flowers, male flowers with androecium haplostemonous, isostemonous, oppositisepal, with three to four, rarely eight, stamens, gynostemium short or vestigial, anthers basifixed, not versatile, dithecous, tetrasporangiate, elongated, connectivum apically widened, peltate, dehiscence along longitudinal valves; pistillidium sometimes present, female flowers with superior gynoecium carpels, apocarpous in two or three whorls, imperfectly closed apically, surrounded by petals, linear stylodious, stigmas internal, decurrent in two ridges, more or less dry, two ovules per carpel but one nearly always aborts, orthotropous, bitegmic, crassinucellated, pendulous, apical to marginal placentation, three or four staminodes 3-4, no nectaries Poppy Family Papaveraceae usually herbaceous, rarely woody Papaveraceae Papaveraceae usually half the # sepals as petals (4 or 6 petals, 2 or 3 sepals) Papaveraceae 4 petals Papaveraceae or 6 petals and numerous stamens Papaveraceae syncarpous, ovary superior, fruit a capsule Saxifragaceae herbaceous, with palmate leaves, oen basal Saxifragaceae inflorescence with reduced leaves Saxifragaceae receptacle a hypanthium Saxifragaceae gen. 5 sepals, 5 petals, 5-10 stamens Saxifragaceae gen 2-3 carpels fused at base, ovary superior to parally inferior Grossulariaceae Always woody Alternate palmate leaves Currants without nodal spines Gooseberries (usually) with nodal spines Grossulariaceae 4 – 5 sepals 4-5 petals (same number as sepals) 4-5 stamens (same number as sepals) Grossulariaceae 4 – 5 sepals 4-5 petals (same number as sepals) 4-5 stamens (same number as sepals) Grossulariaceae Hypanthium – cup to tube shaped Grossulariaceae Hypanthium – cup to tube-shaped Grossulariaceae Ovary inferior (pedicels often pendent) The Rose Family - Rosaceae Herbaceous to woody Hypanthium (expanded receptacle) 5 sepals, 5 petals – separate (free), radial symmetry Numerous stamens usually Ovary superior (unicarpellate, apocarpous) or inferior (syncarpous Rosaceae Numerous stamens Rosaceae Hypanthium Rosaceae Unicarpellate = ovary superior, one ovary, one style, one sgma, one chamber Rosaceae Apocarpous = ovaries superior, more than one separate carpel per flower Rosaceae Syncarpous = more than one fused carpel per flower, in this family, ovary always inferior .
Load more

Recommended publications
  • Auxin Regulation Involved in Gynoecium Morphogenesis of Papaya Flowers
    Zhou et al. Horticulture Research (2019) 6:119 Horticulture Research https://doi.org/10.1038/s41438-019-0205-8 www.nature.com/hortres ARTICLE Open Access Auxin regulation involved in gynoecium morphogenesis of papaya flowers Ping Zhou 1,2,MahparaFatima3,XinyiMa1,JuanLiu1 and Ray Ming 1,4 Abstract The morphogenesis of gynoecium is crucial for propagation and productivity of fruit crops. For trioecious papaya (Carica papaya), highly differentiated morphology of gynoecium in flowers of different sex types is controlled by gene networks and influenced by environmental factors, but the regulatory mechanism in gynoecium morphogenesis is unclear. Gynodioecious and dioecious papaya varieties were used for analysis of differentially expressed genes followed by experiments using auxin and an auxin transporter inhibitor. We first compared differential gene expression in functional and rudimentary gynoecium at early stage of their development and detected significant difference in phytohormone modulating and transduction processes, particularly auxin. Enhanced auxin signal transduction in rudimentary gynoecium was observed. To determine the role auxin plays in the papaya gynoecium, auxin transport inhibitor (N-1-Naphthylphthalamic acid, NPA) and synthetic auxin analogs with different concentrations gradient were sprayed to the trunk apex of male and female plants of dioecious papaya. Weakening of auxin transport by 10 mg/L NPA treatment resulted in female fertility restoration in male flowers, while female flowers did not show changes. NPA treatment with higher concentration (30 and 50 mg/L) caused deformed flowers in both male and female plants. We hypothesize that the occurrence of rudimentary gynoecium patterning might associate with auxin homeostasis alteration. Proper auxin concentration and auxin homeostasis might be crucial for functional gynoecium morphogenesis in papaya flowers.
    [Show full text]
  • BIOLOGY ASSIGNMENT CLASS IX UNIT II- FLOWERING PLANT Chap 3- the Flower
    BIOLOGY ASSIGNMENT CLASS IX UNIT II- FLOWERING PLANT Chap 3- The Flower Flower is the most attractive,brightly coloured and significant part of plant. It help in sexual reproduction and later forms fruit and seeds. Characterstic of flower : The flower possesss the following characterstic – 1. Flower is the highly modified and specialized shoot meant for Sexual Reproduction. 2.The nodes and internodes are highly condensed to form a flat thalamus or receptacles. 3. Thalamus is quiet short and usually borne at the end of the stalk called Pedicel. 4.The flower arise from the leaf like structure called bract. 5. The floral part of flower are borne on thalamus in the form of four whorls. 6. These four whorl includes from outer side – Calyx ( sepal ), Corolla( petal ) , Androecium (stamen ), Gynoecium ( carpel ). Parts of a Flower : Floweris attached with the stem or its branch with the help of a stalk called Pedicel. If the flower is without pedicel it is called sessile . The pedicel arises from the axil of green leaf like structure called Bract.The other swollen end of the pedicel is Thalamus or torus . On the thalamus usually four whorl of Floral structure are present These four whorl from outside to inside are : i. Calyx iii. Androecium ii. Corolla iv. Gynoecium I. CALYX It is the outermost and the lowermost whorl , green in colour which cover an unopened bud .The single unit of Calyx is called sepal .It may be fussed or free from each other. Additional floral whorl outside the Calyx is called Epicalyx. Function of Calyx : i.
    [Show full text]
  • Repeated Climate-Linked Host Shifts Have Promoted Diversification in a Temperate Clade of Leaf-Mining Flies
    Repeated climate-linked host shifts have promoted SPECIAL FEATURE diversification in a temperate clade of leaf-mining flies Isaac S. Winklera,b,1, Charles Mitterb, and Sonja J. Schefferc aDepartment of Entomology, North Carolina State University, Campus Box 7613, Raleigh, NC 27695-7613; bDepartment of Entomology, University of Maryland, 4112 Plant Sciences Building, College Park, MD 20742; and cSystematic Entomology Laboratory, Plant Science Institute, Agricultural Research Service, United States Department of Agriculture, 10300 Baltimore Avenue, Building 003, Room 231, BARC-West, Beltsville, MD 20705 Edited by Anurag A. Agrawal, Cornell University, Ithaca, NY, and accepted by the Editorial Board July 30, 2009 (received for review May 1, 2009) A central but little-tested prediction of ‘‘escape and radiation’’ ever, there is still little evidence on the degree to which changes coevolution is that colonization of novel, chemically defended host in either plant defense or insect ‘‘offense’’ promote diversifica- plant clades accelerates insect herbivore diversification. That the- tion (7). Progress on the insect side has come from several recent ory, in turn, exemplifies one side of a broader debate about the reports plausibly attributing an instance of significantly elevated relative influence on clade dynamics of intrinsic (biotic) vs. extrinsic insect diversity to a co-occurring shift to a new host taxon (5, 10, (physical-environmental) forces. Here, we use a fossil-calibrated 11). Any single instance of elevated diversification, however, molecular chronogram to compare the effects of a major biotic could reflect other influences that happen to be confounded factor (repeated shift to a chemically divergent host plant clade) with the host shift.
    [Show full text]
  • Plant Taxonomy Table
    COMMON AND LATIN NAMES OF IMPORTANT PLANT TAXA LATIN NAME* COMMON NAME Abies Fir Acer Maple Acer negundo Box elder Aesculus Buckeye; Horse Chestnut Alnus Alder Ambrosia Ragweed Apiaceae [Umbelliferae] Carrot or parsley family Artemisia Sagebrush; sage; wormwood Asteraceae [Compositae] Aster or Sunflower Family Betula Birch Boraginaceae Borage family Brassicaceae [Cruciferae} Mustard family Caryophyllaceae Pinks Castanea Chestnut Compositae (Asteraceae) Aster or Sunflower family Cornus Dogwood Corylus Filbert; hazelnut Cruciferae (Brassicaceae) Mustard family Cupressaceae Junipers, cypresses, "cedars", others Cyperaceae Sedge family Ericaceae Heath family Fabaceae [Leguminosae] Pea family Fagus Beech Fraxinus Ash Gramineae (Poaceae) Grass family Juglans Walnut; butternut Labiatae (Lamiaceae) Mint family Larix Larch; tamarack Leguminosae (Fabaceae) Pea family Liliaceae Lily family Liriodendron Tulip tree or yellow poplar Nuphar Water lily Onagraceae Evening primrose family Papaveraceae Poppy family Picea Spruce Pinus Pine Plantago Plantain Poaceae [Gramineae] Grass family Polemonium Jacob's ladder Polygonaceae Buckwheat family Populus Poplar; cottonwood; aspen Potamogeton Pondweed Primulaceae Primrose family Quercus Oak Ranunculaceae Buttercup family Rosaceae Rose family Rhus sumac, incl. poison ivy, etc. Salix Willow Saxifragaceae Saxifrage family Scrophulariaceae Snapdragon family Sparganium Bur reed Thalictrum Meadow rue Tilia Linden or basswood Tsuga Hemlock Typha Cattail Ulmus Elm Umbelliferae (Apiaceae) Carrot or parsley family * Names of genera are always italicized; family names are given in Roman characters. All proper plant family name ends in -aceae; family names above that don't have this ending are old names, and the proper modern name is included in parentheses. .
    [Show full text]
  • Outline of Angiosperm Phylogeny
    Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese
    [Show full text]
  • The Developmental and Genetic Bases of Apetaly in Bocconia Frutescens
    Arango‑Ocampo et al. EvoDevo (2016) 7:16 DOI 10.1186/s13227-016-0054-6 EvoDevo RESEARCH Open Access The developmental and genetic bases of apetaly in Bocconia frutescens (Chelidonieae: Papaveraceae) Cristina Arango‑Ocampo1, Favio González2, Juan Fernando Alzate3 and Natalia Pabón‑Mora1* Abstract Background: Bocconia and Macleaya are the only genera of the poppy family (Papaveraceae) lacking petals; how‑ ever, the developmental and genetic processes underlying such evolutionary shift have not yet been studied. Results: We studied floral development in two species of petal-less poppies Bocconia frutescens and Macleaya cordata as well as in the closely related petal-bearing Stylophorum diphyllum. We generated a floral transcriptome of B. frutescens to identify MADS-box ABCE floral organ identity genes expressed during early floral development. We performed phylogenetic analyses of these genes across Ranunculales as well as RT-PCR and qRT-PCR to assess loci- specific expression patterns. We found that petal-to-stamen homeosis in petal-less poppies occurs through distinct developmental pathways. Transcriptomic analyses of B. frutescens floral buds showed that homologs of all MADS-box genes are expressed except for the APETALA3-3 ortholog. Species-specific duplications of other ABCE genes inB. frute- scens have resulted in functional copies with expanded expression patterns than those predicted by the model. Conclusions: Petal loss in B. frutescens is likely associated with the lack of expression of AP3-3 and an expanded expression of AGAMOUS. The genetic basis of petal identity is conserved in Ranunculaceae and Papaveraceae although they have different number of AP3 paralogs and exhibit dissimilar floral groundplans.
    [Show full text]
  • Ranunculaceae) for Asian and North American Taxa
    Mosyakin, S.L. 2018. Further new combinations in Anemonastrum (Ranunculaceae) for Asian and North American taxa. Phytoneuron 2018-55: 1–11. Published 13 August 2018. ISSN 2153 733X FURTHER NEW COMBINATIONS IN ANEMONASTRUM (RANUNCULACEAE) FOR ASIAN AND NORTH AMERICAN TAXA SERGEI L. MOSYAKIN M.G. Kholodny Institute of Botany National Academy of Sciences of Ukraine 2 Tereshchenkivska Street Kiev (Kyiv), 01004 Ukraine [email protected] ABSTRACT Following the proposed re-circumscription of genera in the group of Anemone L. and related taxa of Ranunculaceae (Mosyakin 2016, Christenhusz et al. 2018) and based on recent molecular phylogenetic and partly morphological evidence, the genus Anemonastrum Holub is recognized here in an expanded circumscription (including Anemonidium (Spach) Holub, Arsenjevia Starod., Tamuria Starod., and Jurtsevia Á. Löve & D. Löve) covering members of the “Anemone ” clade with x=7, but excluding Hepatica Mill., a genus well outlined morphologically and forming a separate subclade (accepted by Hoot et al. (2012) as Anemone subg. Anemonidium (Spach) Juz. sect. Hepatica (Mill.) Spreng.) within the clade earlier recognized taxonomically as Anemone subg. Anemonidium (sensu Hoot et al. 2012). The following new combinations at the section and subsection ranks are validated: Anemonastrum Holub sect. Keiskea (Tamura) Mosyakin, comb. nov . ( Anemone sect. Keiskea Tamura); Anemonastrum [sect. Keiskea ] subsect. Keiskea (Tamura) Mosyakin, comb. nov .; Anemonastrum [sect. Keiskea ] subsect. Arsenjevia (Starod.) Mosyakin, comb. nov . ( Arsenjevia Starod.); and Anemonastrum [sect. Anemonastrum ] subsect. Himalayicae (Ulbr.) Mosyakin, comb. nov. ( Anemone ser. Himalayicae Ulbr.). The new nomenclatural combination Anemonastrum deltoideum (Hook.) Mosyakin, comb. nov . ( Anemone deltoidea Hook.) is validated for a North American species related to East Asian Anemonastrum keiskeanum (T.
    [Show full text]
  • Functional Analyses of Genetic Pathways Controlling Petal Specification in Poppy Sinéad Drea1, Lena C
    RESEARCH ARTICLE 4157 Development 134, 4157-4166 (2007) doi:10.1242/dev.013136 Functional analyses of genetic pathways controlling petal specification in poppy Sinéad Drea1, Lena C. Hileman1,*, Gemma de Martino1 and Vivian F. Irish1,2,† MADS-box genes are crucial regulators of floral development, yet how their functions have evolved to control different aspects of floral patterning is unclear. To understand the extent to which MADS-box gene functions are conserved or have diversified in different angiosperm lineages, we have exploited the capability for functional analyses in a new model system, Papaver somniferum (opium poppy). P. somniferum is a member of the order Ranunculales, and so represents a clade that is evolutionarily distant from those containing traditional model systems such as Arabidopsis, Petunia, maize or rice. We have identified and characterized the roles of several candidate MADS-box genes in petal specification in poppy. In Arabidopsis, the APETALA3 (AP3) MADS-box gene is required for both petal and stamen identity specification. By contrast, we show that the AP3 lineage has undergone gene duplication and subfunctionalization in poppy, with one gene copy required for petal development and the other responsible for stamen development. These differences in gene function are due to differences both in expression patterns and co- factor interactions. Furthermore, the genetic hierarchy controlling petal development in poppy has diverged as compared with that of Arabidopsis. As these are the first functional analyses of AP3 genes in this evolutionarily divergent clade, our results provide new information on the similarities and differences in petal developmental programs across angiosperms. Based on these observations, we discuss a model for how the petal developmental program has evolved.
    [Show full text]
  • Molecular Communication for Coordinated Seed and Fruit Development: What Can We Learn from Auxin and Sugars?
    International Journal of Molecular Sciences Review Molecular Communication for Coordinated Seed and Fruit Development: What Can We Learn from Auxin and Sugars? Hélène S. Robert Mendel Centre for Genomics and Proteomics of Plants Systems, CEITEC MU-Central European Institute of Technology, Masaryk University, 625 00 Brno, Czech Republic; [email protected]; Tel.: +420-549-49-8421 Received: 30 January 2019; Accepted: 19 February 2019; Published: 21 February 2019 Abstract: Seed development in flowering plants is a critical part of plant life for successful reproduction. The formation of viable seeds requires the synchronous growth and development of the fruit and the three seed structures: the embryo, the endosperm, the seed coat. Molecular communication between these tissues is crucial to coordinate these developmental processes. The phytohormone auxin is a significant player in embryo, seed and fruit development. Its regulated local biosynthesis and its cell-to-cell transport capacity make of auxin the perfect candidate as a signaling molecule to coordinate the growth and development of the embryo, endosperm, seed and fruit. Moreover, newly formed seeds need nutrients and form new carbon sink, generating high sugar flow from vegetative tissues to the seeds. This review will discuss how auxin and sugars may be considered as signaling molecules to coordinate seed and fruit development. Keywords: auxin; sucrose; embryo; embryo; endosperm; seed; fruit; molecular communication 1. Auxin in Seed Development Plant hormones are instrumental players for many aspects of plant development. In flowering plants, every step leading to seed formation requires crosstalk between various hormones: flower primordium development, floral organ development, including stamens, gynoecium patterning, ovule formation, ovule number, fertilization, seed formation, fruit initiation [1–8].
    [Show full text]
  • Ranunculus Parviflorus (Ranunculaceae) Naturalized in Kansas
    Singhurst, J.R., J.N. Mink, and W.C. Holmes. 2018. Ranunculus parviflorus (Ranunculaceae) naturalized in Kansas. Phytoneuron 2018-22: 1–3. Published 25 April 2018. ISSN 2153 733X RANUNCULUS PARVIFLORUS (RANUNCULACEAE) NATURALIZED IN KANSAS JASON R. SINGHURST Wildlife Diversity Program Texas Parks and Wildlife Department 4200 Smith School Road Austin, Texas 78744 [email protected] JEFFERY N. MINK 3229 Cole Avenue Waco, Texas 76707 [email protected] WALTER C. HOLMES Department of Biology Baylor University Waco, Texas 76798-7388 ABSTRACT Ranunculus parviflorus is documented here in the naturalized flora of southeast Kansas, where it was collected at two locations in the town of Neodesha. Ranunculus parviflorus L. (Ranunculaceae) (smallflower buttercup) (Figure 1) is a herbaceous annual native to Europe (Salisbury 1931; Fernald 1950; Fitzgerald 2002). The species is naturalized in subtropical and temperate areas of North and South America (Benson 1948; Hernandez 1993) and Australia and New Zealand (Bock 1979). Ranunculus parviflorus is now found from California north through Oregon and Washington (to British Columbia) and in the eastern USA from western Florida north to New York, west through the Midwest to central Missouri, south through eastern Oklahoma and eastern Texas, east to Florida, including Hawaii (Brouillet et al. 2006; Hickman 1993; Lowe 1921; Mohr 1901; Stone et al. 1992; Wunderlin et al. 1996). Early collections in the USA were made by Chickering at Ft. Monroe, Virginia, in 1878 (KANU) and Curtiss in Gadsden Co., Florida, in 1889 (USFH 6400) (Wunderlin et al. 2018). Ranunculus parviflorus has not been included in floras or checklists for Kansas (Brooks 1986; Haddock et al.
    [Show full text]
  • The Classification of Plants, Viii
    70 The Ohio Naturalist [Vol. XIII, No. 4, THE CLASSIFICATION OF PLANTS, VIII. JOHN H. SCHAFFNER. Below is presented a synopsis of the fifteen plant phyla given in the preceding paper of this series. The classification of the fungi follows with a key to the orders. The following changes should be made in the arrangement of the families of Anthophyta as presented in the sixth paper: Transfer the Parnassiaceae from Saxifragales to Ranales following the Ranunculaceae. Interchange the position of Loganiaceae and Oleaceae. Also interchange the position of Bromeliaceae and Dioscoreaceae. SYNOPSIS OF THE PLANT PHYLA. A. Plant body unicellular or filamentous, or if a solid aggregate through the ovary, when present, not an archegonium; never seed-producing; nonsexual, with a simple sexual life cycle, or with an alternation of generations. I. Cells typically with poorly differentiated nuclei and chromatophores, reproducing by fission; motile or nonmotile, colored or colorless, with or without chlorophyll but never with a pure chlorophyll- green color; resting spores commonly present. Phylum 1. SCHIZOPHYTA. II. Cells with well differentiated nuclei, and if holophytic usually with definite chromatophores; with or without chlorophyll; colorless, green, or variously tinted by coloring matters. (I.) Nonsexual, unicellular plants without chlorophyll having a plasmodium stage of more or less completely fused amoeboid cells from which complex .sporangium-like resting bodies are built up. Phylum 2. MYXOPHYTA. (II.) Plants not developing a plasmodium, but the cells normally covered with walls in the vegetative phase. 1. Unicellular or filamentous plants containing chlorophyll, either brown with silicious, two-valved walls or green with complex chromatophores, the walls not silicificd; conjugating cells not ciliated, isogamous.
    [Show full text]
  • The Discovery and Naming of Papaver Orientale Sl (Papaveraceae)
    The discovery and naming of Papaver orientale s.l. (Papaveraceae) with notes on its nomenclature and early cultivation H. Walter Lack Abstract LACK, H.W. (2019). The discovery and naming of Papaver orientale s.l. (Papaveraceae) with notes on its nomenclature and early cultivation. Candollea 74: 47 – 64. In English, English abstract. DOI: http://dx.doi.org/10.15553/c2019v741a7 Papaver sect. Macrantha Elkan is a widespread and common polyploid complex comprising the diploid Papaver bracteatum Lindl., the tetraploid Papaver orientale L. and the hexaploid Papaver pseudo-orientale (Fedde) Medw. All three species are restricted in their distribution to the Caucasus area in the wide sense including north-eastern Turkey and north-western Iran. Papaver orientale and Papaver pseudo-orientale were first collected by one of the members of the expedition headed by Joseph Pitton de Tournefort (1656 – 1708) and introduced into cultivation in the Jardin du Roi in Paris as early as 1702. From there living material of both taxa was quickly distributed to other botanical gardens in Amsterdam, Leiden and the Chelsea Physic Garden in London. For decades both species were known only from cultivated specimens. Due to hybridisation both in the field and in cultivation the relationships between the two taxa remained unclear and were further blurred by the introduction of Papaver bracteatum into cultivation which began in Gorenki near Moscow, Berlin and Chelsea around 1800. Based on ample evidence never studied before like unpublished illustrations kept in Paris and Vienna, Tournefort’s unpublished field book, and seed lists this paper unravels this complex historical and taxonomic story.
    [Show full text]