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March 2001

Curculionoidea (Insecta: Coleoptera) from the state of ,

M. D. Salas-Araiza Instituto de Ciencias Agricolas

C. W. O'Brien Florida A&M University, Tallahassee, FL

J. Romero-Napoles Colegio de Postgraduados, Texcoco, Mex.

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Salas-Araiza, M. D.; O'Brien, C. W.; and Romero-Napoles, J., "Curculionoidea (Insecta: Coleoptera) from the state of Guanajuato, Mexico" (2001). Insecta Mundi. 181. https://digitalcommons.unl.edu/insectamundi/181

This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI. Vol. 15. No. 1. March. 2001 45

Curculionoidea (Insecta: Coleoptera) from the state of Guanajuato, Mexico

M.D. Salas-Araiza Instituto de Ciencias Agricolas Apartado Postal 311 36500, Gto [email protected] C.W. O'Brien Center for Biological Control College of Engineering Sciences, Technology and Agriculture Florida A&M University, Tallahassee, FL 3230'7-4100 [email protected].

J. Romero-NApoles Instituto de Fitosanidad Colegio de Postgraduados Texcoco, Milx. [email protected]

Resumen. Este trabajo contribuye a1 conocimiento de las especies de curculionoideos preseiltes en el estado de Guanajuato, MBxico. Se determinaron 73 g6neros y 125 especies; se destacan, por el nGmero de Qstas,las subfamilias . y con 24, 24 y 23 especies, respectivamente. Los gQneroscoil mayor cantidad de especies fueron Geraer~sy Ai~tlioizoi~i~~scon siete, y Coizotmchelr~scoil seis. Se obtuvieron 18 nuevos registros para la Repitblica Mexicana. El 53.7% de las especies de c~~rculionoideosdel estado de Guanajuato son de afinidad neotropical, el 26% son de afinidad neArtica, el 18.4% son de distribuci6n cosmopolita, y el resto (1.9%) antes solo se han registrado en las Antillas.

Summary. This study is a contributioil to the knowledge of the species present in the State of Guanajuato, Mexico. During this study, 73 genera and 125 species were identified. The subfamilies best represented were Baridinae, Curculioninae and Entimilme with 24, 24 and 23 species, respectively. The genera best represented were Geraer~sand A~L~~ZOILOIIZZLSwith seven species and Coizotrachelz~swith six. Of the 125 species identified, 18 represent new country records. Of the weevil species found in Guanajuato, 53.7% are Neotropical, 26% are Nearctic, 18.4% are cosmopolitan, and the remainder (1.9%) previously were known only from the West Indies.

Introduction (1983) indicated that 36 of the then recognized 52 Neotropical subfamilies had been recorded for Mex- The are the family of living organisms ico, which constituted 69.23% of the total number of with the largest number of species lrnown. They subfamilies of Curculionoidea. can be found in all terrestrial habitats, and some The endemism of the Mexican weevil genera is species are aquatic. Larvae of many species feed on 6.5% and the endemism of the species is 40.5%. In 'oots, stalks, branches, trunlis, leaves, flowers, the United States and Canada. endemism is much fruits and seeds of grasses, shrubs and trees; some higher (31.2% of the genera and 76.6% of the feed on mosses, and others on fungi (Zimmerman, species): this fact is due to the geographical location 1994). of Mexico in the center of the continent, which O'Brien & Wibiner (1978) mentioned that favors the faunal exchange with the northern and through 1971 there were 4,237 genera and 44:883 southern regions (Anderson & O'Brien, 1996). In species of se7~sz~lato recognized as the central area of Mexico, particularly in the State valid in the world. Also they pointed out that the of Guanajuato, knowledge of the Curculionoidea is Neotropical region contains the largest number of poor. This is the first study toward the contribution described species, with a total of 12,962. Luna 4 6 Volume 15, No. 1, March, 2001, INSECTA MUNDI to taxonomic knowledge of the superfamily Curcu- Results and discussion lionoidea in Guanajuato. Weevil fauna Methods and materials During this study, 1,787 specimens were col- lected, belonging to five families, 17 subfamilies, 42 Location of the study area tribes, 73 genera and 125 species. The subfamilies This study was conducted in the State of Gua- best represented are Baridinae (15 genera and 24 najuato, which is located near the center of Mexico, species), Curculioninae (10 genera and 24 species) between 19" 55' 08" and 21" 52' 09North latitude and Entiminae (16 genera and 23 species). The and between 99" 36' 06" and 102" 05' 07" West lon- genera represented by the most species are Gerae- gitude. LLSand A7~tkorzoilzuswith seven each, and Conotra- chelus with six. Eighteen of the species represent Collection, preparation and identification of new country records (Table 1). Of special signifi- the material cance are Sitolza hispidulus (Fabricius)(Entiminae) Between January 1996 and December 1997,20 and Hypera postica (Gyllenhal)(Hyperinae), which collecting trips were carried out in different areas were collected in large numbers in alfalfa fields and representative of the State. These areas correspond have been reported to be important pests of that to an arid region in the northern municipalities, the crop in the United States and Europe. mountainous country of central Guanajuato, an agricultural area of the Bajio, and the hotter and humid area of the south. Weevils were collected Weevil species from the state of from trees by placing a mat underneath and shak- Guanajuato ing the foliage; a beating sheet was used for herba- ceous and shrubby vegetation; some weevils with Family APIONIDAE nocturnal habits were captured using an ultravio- let trap; some apterous species were obtained using Subfamily APIONINAE pitfall traps; active fliers were captured using flight intercept traps in addition to yellow pan traps filled Tribe APIONINI with water. Seeds and branches with larval dam- age were placed in rearing chambers in order to Apion spp. Sierra de Lobos, Le6n. 20lIXl95, 2OIIIII obtain adults. 96, 20NIIl96. Acacia spp.; Arnara,rzthus spp.; The main reference used for the identification Mimosa spp.; Quercz~sspp. (10 specimens). of the species was the Biologia Centrali-Americana Apion nr. tenuirostrum Smith. 9mi. N.E. Guade- (Champion, 1902-1909; Sharp, 1889-191 1). The keys lupe Victoria 8000 feet, VIII/16174, (3 speci- in Kissinger (1964), Hatch (1971), Dillon & Dillon mens); 1mi. N. Sauceda 7000 Feet, VIl2911971. (1972), Arnett (1973), Whitehead (1980), Arnett (lspecimen). (1985), and Bright (1994) were used as well. The Apion nr. varicorne Smith. 1 mi. N. Sauceda names and higher classification follow O'Brien & 7000 Feet, V112911971. (4 specimens) Wibmer (1982 & 1984), Wibmer & O'Brien (1986 & 1989) and Alonso-Zarazaga & Lyal (2000). Many Tribe PIEZOTRACHELINI species were identified through comparisons with specimens from the following collections: Charles Chrysapion auctum (Sharp). El Copal, Irapua- W. O'Brien Collection (Tallahassee, Florida); Na- to. 4NI.1196. Maleza. (1 specimen). tional Collection of INIFAP (, Guana- juato), and IFIT-CP Collection (Montecillos, Mexi- Family co). Reference material is deposited at the Entomo- logical Collection "Leopoldo ~inocoCorona" of ICA- Subfamily ATTELABINAE UGTO (Irapuato, Guanajuato). Tribe ATTELABINI

Himatolabus umbosis Hamilton. 3 km S XichG. 3NIIIl96. Maleza. (2 specimens). INSECTA MUNDI, Vol. 15, No. 1, March, 2001

Tribe PILOLABINI apicalis (Boheman). 1.5 lim N La Sauceda; El Copal, Irapuato. 29NI/71,281W96. Pilolabus klugii (Gyllenhal). Manzanar, San Maleza. (2 specimens). Luis de la Paz. 3NIIIl96. Maleza. (1 specimen). Trepobarisperlonga Champion. Gervasio Men- doza, Salvatierra; El Copal, Irapuato; Cd. Man- Family uel Doblado. 30NIII195, 31NII196, 811x196, 111 1x196, 28NII197. Ager~iataliscar~a Wats.; Bac- Subfamily RHYNCHITINAE charis spp.; Gramineae; Zea rlzays L. (4 speci- mens). Tribe RHYNCHITINI Ti-ichobaris championi Barber. 1.5 km N La Sauceda; Cd. Manuel Doblado; El Copal, Ira- Eugnamptus spp. 1 km Rancho de Enmedio car- puato. 29NI171, 10NIIIl89, 3lIVl95, 10NIIl95, retera Dolores-Guanajuato; Cuenca de la Espe- 251IW95, 25lIVl96, 4NII197, 25lIW97. Clzeizop- ranza, Guanajuato; km 22 carretera Lecin-San odiu~nspp.; Plzysalis spp.; Sorghurn vulgare Felipe; Manzanar, , 4NIIl96, (L.). (12 specimens). 6NII196, 3NIII196, 31NIII196, 28NIl97, Quer- major Barber. 1.5 km N La Sauce- cus spp., maleza, (7 specimens). da, Guanajuato. 29NI171. (1 specimen). Haplorhynchites mexicanus (Gyllenhal). El Trichobaris pellicea (Boheman). km 11 car- Copal, Irapuato; Las Antenas, Irapuato. 7NIII retera Le6n-Ocampo. 28NIl97. Maleza. (1 spec- 98, 111x198, 24lW98. Heliar~thusspp.; maleza; imen). Sirnsia ar,zplexicaulis (Cav.) Pers. (31 speci- mens). Tribe Haplorhynchites quadripennis (Fall). Las An- tenas, Irapuato. 241W98. Sirnsia a~nplexicaz~lis ambigua Casey. 4 lim S San Luis (Cav.) Pers. (4 specimens). de la Paz. 13NIII198. Gramineae. (1 specimen). Haplorhynchites subauratus (Sharp). Santa Stictobaris ornatella Casey. San Pedro, Comon- Rosalba, . 811x196. Biderzs pilosa L. (1 fort; 1.5 km N La Sauceda; El Copal, Irapuato; specimen). Las Antenas, Irapuato; km 5 carretera Sala- Involvulus mollis (Sharp). 1 km E Rancho de manca-Irapuato. 20NI171, 20NIIl89, 10lIW90, Enmedio, Guanajuato; km 22 carretera Lecin- 5NIIIl94, 3NIIIl95, 20NIIl96; 23NIIIl96, 211 San Felipe. 16NII196,28NI/97. Quercus spp. (3 W97, 311111198, 21lIVl98, 4NII198, 19NIIl98, 51 specimens). VIIIl98. Amarai~thz~sspp.; en trampa amarilla. (30 specimens). Family CURCULIONIDAE Tribe MADOPTERINI Subfamily BARIDINAE Apinocis angustus (Casey). Los Esp&rragos,Si- Tribe lao. 261x1196. Cylzodoi~dactyl or^ (L.). (4 speci- mens). corrusca (Boheman). El Copal, Irapuato. Barilepton quadricolle LeConte. El Copal, Ira- 2/11/91, 28lW96. Baccharis spp. (3 specimens). puato. 121x1196, 13/111/97,9/1V/97.Cyperaceae; Baris strenua (LeConte). El Copal, Irapuato; Gramineae. (68 specimens). Las Palomas, Guanajuato; carretera Morole6n- Barinus spp. El Copal, Irapuato. 5111190. En tram- Morelia camino a1 Aguacate; Frias, Cd. Manuel pa amarilla. (3 specimens). Doblado; km 5 carretera Salamanca-Irapuato. Centrinopus mistus Champion. El Copal, Ira- 3~111/94,5N111/94,6N11195,12~11/95,31N111 puato. 6NIIl95. Arte~nisiasp. (1 specimen). 95, 31lW95, 9NIIl96, 3NIII196, 31NIII196, 81 Geraeus candidus Champion. El Copal, Irapua- IW96, 15lIW97, 7NIIl98, 27NIIl98, 1lIW98. to. 8/VII194,28/VII/98.Asteraceae; Mirnosa spp. Arnara7zthus spp.; Cosrnos bipirz7zatus Cov.; (3 specimens). Heliar~thusa7zltuus L.; Prosopis spp.; Sirnsia Geraeus dugesi Champion. ; 16 km arnplexicaulis (Cov.) Pers.; Sorghum vulgare S Cd. Manuel Doblado. 17NII196; 9NIIIl98. (L.). (26 specimens). Baccharis spp.; Gramineae. (3 specimens). 48 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Geraeus hospes (Casey). El Copal, Irapuato. 31 ca. 5NIIl91, 131W93, 30lIIIl96, 15Nl96, 23Nl VIIIl94, 4lIW96, 10NIIl97, 2511x197. Alnaran- 96,31N/96,14NI/96,20N1/96,26NI/96,28NII thr~ssp.; Arte~nisialudoviciarza Nutt.; Coriau- 96, 7lVIIl96, 9NIIl96, 23NIII196, 28/11/98. drunz sativu~nL.; Prosopis sp. (4 specimens). Gramineae; Medicago sativa L.; Partlze7ziurn Geraeus perscitus (Herbst). El Copal, Irapuato. lzystel-ophorus L. (19 specimens). 14NIIIl90, 51IW94, 26lIW95, 28lIW95, 28NIl 96, 281x196, 13lIIIl97, 111IW97, 25lIW97, 301 1x197. Ainar.arzthus spp.; Col-iarzdn~nzsativur7z Tribe PHYTOBIINI L.; Gramineae; Malva pal-uiflora L. (14 speci- me ns) . pyrrhopus Boheman. El Copal, Ira- Geraeus senilis (Gyllenhal). El Copal, Irapuato. puato. 161x197. Polygoizu~~zpr~izctatz~irL Ell. (40 611x190, 1NIIl93, 311x193. Zea nays L. (4 spec- specimens). imens). Geraeus tonsus Champion. El Copal, Irapuato; AcBmbaro; 15 km NE . 311W93, Subfamily (formerly 16NIIIl94, 17NIIl96. Maleza; Zea nzays L. (2 ZYGOPINAE) specimens). Geraeus varius Champion. 3 km S San Josh Tribe LECHRIOPINI Iturbide. 2NIIIl96. Maleza. (1 specimen). Limnobaris discreta Champion. Rancho de Lechriops auritus (Schonherr). El Copal, Ira- Enmedio, Guanajuato. 16lVIIl96, 24lVIl97. puato. 2911x195, 1811x196. Mar*rubiu7rzvulgare Quel-cus spp. (2 specimens). L. (3 specimens). Nicentrus femoralis Champion. El Copal, Ira- puato. 7NIIl90, 27A7III190, 31NIIIl90, 14IIW Tribe PIAZURINI 90, 191x190, lONIIIl93, 26NIIIl93, 71IW93, 31 VII195, 10NIIIl95, 23NIIIl95, 6NIIIl97, 121 Cratosomus punctulatus Gyllenhal. Irapuato; VIIIl97, lllIW97, 3011x197, 12NIIIl98. Citrr~s Le6n. 2NI172, 18NIIIl91, 25lIXl91. Maleza; sirzer~sis(L.) Osb.; Medicago sativa L.; Sor- Plzysalis spp. (3 specimens). ghum vrclgare (L.); Zea nzays L. (20 specimens). Pseudocentrinus ochraceus (Boheman). El Tribe ZYGOPINI Copal, Irapuato. 29lIW94, 23NIIIl96, 27NIIIl 96, 411x196, 611x196, 11lIXl96, 8/IX/98. Baccka- adspersus (LeConte). El ris salicifolia (Ruiz & Pav6n); Gramineae; Mar- Copal, Irapuato. 6Nl96, 10NIIl97. Arter~zisia rubiul?z vulgare L. (32 specimens). lz~douici~~~aNutt. (4 specimens). Cylindrocopturus biradiatus Champion. El Tribe OPTATINI Copal, hapuato; Salamanca. 8lIIIl98, 18lIIIl98, 31lIIIl98, 2lIVl98, 8lIV198, 15lIVl98, 211IV198, Optatus palmaris (Pascoe). Salamanca; Pknja- 5Nl98, 7NIIl98. Opurztia spp. (34 specimens). mo. 30NIl52, 8NIIl89. Alzlzolza, cherilnola Mill. Cylindt-ocopturusganglbaueri (Heller). MBrqu- (9 specimens). ez, Irapuato. 4NII198. Opui~tiaspp. (8 speci- mens). Subfamily

Tribe CEUTORHYNCHINI Subfamily COSSONINAE

Ceutorhynchus truquii (Champion). El Puerto Tribe COSSONINI del Milagro: XichG. 3NIIl96. Brassica carnpes- tris L., B. rapa L.; Quercu,s spp. (38 specimens). Cossonus convexirostris Champion. El Copal, Irapuato; Santiago Maravatio; Le6n. 10NIII94, Tribe 3/11/96, 7NI1196, 31NIIIl96. Sobre suelo. (4 specimens). aspel- (LeConte). El Copal, Irapuato; La Providei~cia,Abasolo; La Ordeiia, Salaman- INSECTA MUNDI, Vol. 15, No. 1, March, 2001

Tribe DRYOTRIBINI (formerly Ai?za,rartthusspp.; Medicago sativa L. (8 speci- COTASTERINI) mens). Antlzonomus texanus Dietz. 1.6 lim N La Sauce- Caulophilus oryzae (Gyllenhal). Irapuato. 301 da, Guanajuato; El Copal, Irapuato; San Felipe. VIIl98. En semilla de Persea arlzericar~aMill. en 29NI174, 281IW95, 22lIW96, 161x197. Sphaer- suelo. (30 specimens). alcia ar-~gustifolia(Cov.) G. Don. (9 specimens). Anthonomus thyasocnemoides Hustache. El Subfamily Copal, Irapuato. 21x195, 71111196, 81x196, 7NIII 97, 101IW97. Acacia spp.; Arteinisia spp.; Ma,r- Tribe CRYPTORHYNCHINI I'ZL~~UI?~urclgare L. (16 specimens). Pseudanthonomus gluon Clark. El Copal, Ira- Gerstaeckeria mutillaria (Gerstaecker). Vic- puato. 28/IX/95,25/X/96,28/W96.Iponzoea spp. toria. 20Nl96. Larva alimentandose de Main- (8 specimens). rnillaria e~.ythrosperr?zaBoed. (2 specimens). Smicraulax tuber-culatus Pierce. El Copal, Ira- Gerstaeckeria unicolor Fisher. Barro Colorado, puato. 28lIW95. Psittaca~~thrcscalyculatus G. San Luis de la Paz; Manzanar, San Luis de la Don. (1 specimen). Paz; 3 l.;m N San Josit Iturbide. 2NIIIl96, 31 VIIIl96, 13NIIIl98. Cylir~dropz~r~tiair?zbrica.ta (Haw.) Knuth. (37 specimens). Tribe CURCULIONINI Maemactes punctatus Kissinger. El Copal, Ira- puato. 81W96, 171x1198, 241x1198. En trampa Curculio ortegai Gibson. 1 km E de Rancho de pitfall. (5 specimens). Enmedio, Guanajuato. 16NIIl96. Quercus spp. Pseudomopsis similis Champion. El Copal, Ira- (1 specimen). puato. 10NII/97. Aca,cia sp. (1 specimen). Tribe DERELOMINI Subfamily CURCULIONINAE Phyllotrox suturalis (Boheman). La Providen- Tribe cia, Abasolo; Las Palomas, Guanajuato; El Co- pal, Irapuato. 20/1/96,30/111/96,6/1V/96.Acacia Anthonomus deliqulus Clark & Burke. El Co- spp.; Bacchal-is spp; Medicago sativa L. (9 spec- pal, Irapuato; 25 km N Santa Cruz de Juventi- imens). no Rosas. 23/X/96,17/VII/97. Ipornoeasp.; Quer- CLLS sp. (2 specimens). Tribe OTIDOCEPHALINI Anthonomus eugenii Cano. El Copal, Irapuato; Gervasio Mendoza, Salvatierra; San Vicente, Myrmex spp. 1 km S ; 25 km N Juventino Salamanca; Granja Guadalupe, Irapuato. 41 Rosas. 111x196, 17NIIl97. Maleza; Quercus sp. VIIl89, 15lX190, 411x193, 1NII194, 6NII195, 311 (2 specimens). VIIIl96, 2511x197, 3011x197. Arternisia spp.; Capsicur7-Larzr~urn L.; Medicago sativa L.; Pisur~z Tribe PRIONOBRACHIINI sativurn L.; Sola~zu~?z7-ostratz~nz Duval. (16 spec- imens). Prionobrachium schoenhel-ri Faust. San Anto- Anthonomus mexicanus Boheman. Irapuato; nio, Salvatierra; El Copal, Irapuato. 17NIII96, Gervasio Mendoza, Salvatierra. 281IW95, 311 231x196, 3lIVl97, 24NIl97, 28NIIl97. Age~zia VIII196, lOlIXl96, 1llIW96, 18lIW96, 8lW96, ta,lisca,iza Wats; Malvaceae. (9 specimens). 221x196, 251x196, 10lIXl97, 161W97. Marrubi- urn uz~lgal-eL. (57 specimens). Tribe SMICRONYCHINI Anthonomus squamans Champion. El Copal, Irapuato. 4NII196, 10lIW97, lllIW97, 161XJ97. Smicronyx albonotatus Anderson. Km 23 car- Cor~izacai~adier~sis (L.) Cronq.; Cyi~odoizdac- retera Dolores-Guanajuato; km 56 carretera tylor~(L.); Marrubiurn vu1gai.e L.; Sphael-alcia Le6n-Ocampo. 4NIIIl96, 28NIl97. Maleza. (3 a~~gustifolia(Cov.) G. Don. (9 specimens). specimens). Anthonomus suturalis LeConte. El Copal, Ira- Smicr-onyx corniculutus (Fbhraeus). El Copal, puato. 6NII/95,20/IW95, 7NII197. Acacia spp.; Irapuato; 8 km S San Luis de la Paz; Apaseo El lume 15, No. 1, March, 2001, INSECTA MUNDI

Grande. 26lIVl96, 16NII/97,16NIII/97,13NIII/ Listl-onotusgt-ypidioides (Dietz). Cruce Rio Laja 98. Acacia spp.; Arteinisia ludoviciai~aNutt. y carretera San Felipe-Guanajuato. 16NIIl96.

, (56specimens). Maleza. (2 specimens). Smicronyx quadrifer Casey. El Copal, Irapuato. Listt-onotus punctiger LeConte. El Copal, Ira- 20/III/96. Acacia sp. (1 specimen). puato. 18lIXl98. Maleza. (1 specimen). Smicronyx spi-etus Dietz. Km 39 carretera Le6n- Ocampo. 28NIl97. Qrcercz~ssp. (1 specimen). Subfamily ENTIMINAE Smict-onyx tenuirostris Champion. El Copal, Irapuato. 9NIl98. Maleza. (1 specimen). Tribe BRACHYDERINI

Tribe Ameladus inornatus Sharp. El Copal, Irapuato. 29/IX/95. Maleza. (1 specimen). aulacis Clark. Las Palomas, Guanajua- Chaetopantus sp. Km 13 carretera J. Rosas-Ce- to. 21x1198. Milnosa spp. (47 specimens). laya; 8 lim E San Luis de la Paz; Santa Catar- Sibinia candidata Champion. El Copal, Irapua- ina. 2NIIIl96, 16NIII/97, 13NIIIl98. Gramine- to. 6NIIl90, 9NII196, 5NIl97, 30NIl98. Sor- ae. (3 specimens). glzz~muu1gai.e (L.). (8 specimens). Mitostylus setosus (Sharp). Km 70 carretera Sibinia inermis (Casey). Carretera Juventino Guanajuato-Querbtaro; Arandas, Irapuato; km Rosas-Guanajuato; El Copal, Irapuato; km 3 56 carretera Le6n-Ocampo; El Bobo, San Luis carretera San Josi! Iturbide-Guanajuato; San de la Paz; Santa Catarina; Camino a la Gaurida, Felipe; El Milagro, carretera Guanajuato-Ju- Irapuato; El Copal, Irapuato. 10lIW82, ION111 ventino Rosas. 9/XI/94, 221111195, 91x1195, 29/11 94, 12NIIl95, 6NIIl96, 2NIIIl96, 6lIW96, 281 96, 31/1/96, 71111196, 11/III/96, 12/III/96, 2OIIIII VIl97, 7NIIl98, 27NIIl98, 28NIIl98, 13NIIII 96, 27/111/96, 1IIVl96, 22/IV/96, 9Nl96, 16NIII 98, 29NIIIl98, 11IW98. Ainarantlzz~sspp.; Arte- 96, 2NIIIl96, 711x196, 9N/97, 7NIIl97. Acacia ilzisia ludoviciaiza Nutt.; Capsicz~nzarzizuin L.; spp.; Bougal~uilleaspp.; Sornghunz vulgare (L.). Cucuinis sativz~sL.; Gramineae; Medicago sa- (123 specimens). tiva L.; Miinosa spp.; Verbesir~auirgata Cov. Sibinia setosa (LeConte). 30 km NE Dolores (46 specimens). Hidalgo; El Copal, Irapuato; La Aldea, . 16NIIIl71, 3NIIIl94, 5/X/94, 1/11/95, 22/11/95, Tribe 1ONl95, 17Nl95, 24Nl95, 31Nl95, 7NIl95, 311 111196, 26lIVl96. Acacia spp.; Pi-osopis spp. (56 Bradyrhynchoides rugicollis (Sharp). Las An- specimens). tenas, Irapuato. 7NIIl98, 9NIIl98, 27NIIl98. Sibinia suturalis (Schaeffer). El Copal, Irapua- Alnarai~tlzusspp.; Polygonaceae. (6 specimens). to; Manzanar, San Luis de la Paz; Camino a la Epicaerus costatus Sharp. El Cubilete, Silao. 31 Garrida, Irapuato. 28/11/94, 20NIIl94, 3NIIIl 11/86. Male~a.(1 specimen). 94, 31NIII/94, 141IXl94, 7NI/95, 22NIl95, 311 Epicaerus squalidus Sharp. Santa Catarina. 21 VIIIl96, 7NIIl97, 10NII/97, 3NIIIl98, 29NIIIl VIIIl96. Maleza. (2 specimens). 98. Acacia spp.; Arterlzisia ludoviciarza Nutt.; Naupactopsis delumbis Champion. Rancho de Mii~zosaspp.; Prosopis spp. (34 specimens). Enmedio, Guanajuato. 16NIIl96. Qzcercus sp. (1 specimen). Incertae sedis Tribe NAUPACTINI Macrorhoptus sphaeralciae Pierce. El Copal, Irapuato. 81x196. Ma,rrzcbiuln vulgare L. (3 spec- Megalostylus albicans (Lacordaire). El Car- imens). men, San Francisco del Rinc6n. 811x196. Male- za. (1 specimen). Subfamily CYCLOMINAE (formerly Pantomorus albosignatus Boheman. 9 liln N RHYTIRRHININAE) Dolores Hidalgo; Santa Catarina; El Principe, San Diego de la Uni6n; Manzanar, San Luis de Tribe RHYTIRRHININI (formerly la Paz; San Jerbnirno, ; cruce Rio LISTRODERINI) Lajas y cametera San Felipe; krn 25 carretera Dolores Hidalgo-Guanajuato; 3 km S San Josk INSECTA MUNDI, Vol. 15, No. 1, March, 2001

Iturbide; Chamhcuaro, Celaya; El Copal, Ira- Pandeleteius ciliatipennis Champion. Rancho puato; Rio Blanco, Atarjea. 16NIIIl71, 8NIIl de Enmedio, Guanajuato; 25 km N Juventino 94, 28NIIl94, 10NIIIl94, 7NIl95, 25NIl95, 121 Rosas. 16NIIl96, 17NIIl97. Quercus spp. (4 VIIl95, 2NIIl96, 16NIIl96, 17NIIl96, 20NIIl specimens). 96, 2NIIIl96, 3NIIIl96, 4NII1/96,24NI/97, 251 Pandeleteius tibialis Boheman. Rancho de En- VIl97, 3NIIl97. Acacia. spp.; Cylindropurztia medio, Guanajuato; 29 km N Juventino Rosas; irnbricata (Haw.) Knuth.; Mimosa. spp.; Proso- Santa Rosa, Guanajuato; 25 km N LeBn; Cuen- pis spp.; Sclzir~us~nolle L.; Sorghum vulgare ca de la Esperanza, Guanajuato. 16NIIl96, 41 (L.). (93 specimens). VIIIl96, 12lXl96, 28NIl97, 7NIIl97, 17NII197. Pantomorus cervinus (Boheman). El Copal, Ira- Arbutus spp.; Quercus spp. (21 specimens). puato. 15lIW97. En follaje de Persea arnericana Pandeleteius viridiventris Champion. El Du- Mill. (5 specimens). razno, Atarjea. 14Nl94. Pinus ce~nbroidesZucc. Pantomorus uniformis Sharp. 14 km N Le6n; El (1 specimen). Copal, Irapuato; Camino a la Garrida, Irapua- Pandeleteius vitticollis Champion. Santa Rosa, to. 16/VIII/74, 81x196, lllIXl97, 29NIIIl98. Guanajuato. 12lW96. Quercus spp. (2 speci- Marr~~bir~mvulga.re L.; Zaluzaizia augusta (Lay) mens). Sch. (3 specimens). Tanymecus confusus Say. El Copal, Irapuato; Las Antenas, Irapuato. 22NIIl93, 1NIII/93, Tribe 10NIIl95, 12NIIl95, 41IW96, 2511x197. Alna- rarzthz~sspp.; Asparagus officinalisL.; Chelzop- cinereus FAhraeus. El Copal, Ira- 0diu7n spp.; Sorghum vulg.are (L.). (9 speci- puato. 28/11/94, 191W94, 81x196, 28lW96. Ipo- mens). rnoea heredaceae Sess6 & Moc.; Malvaparviflo- ra L.; Mirnosa sp.; Prosopis sp. (4 specimens). Subfamily ERIRHININAE Ophryastes sordidus LeConte. El Copal, Ira- puato. 241x1198, 21x11198, 17lXIIl98. Trampa Tribe TANYSPHYRINI pitfall en pastizal. (13 specimens). Tanysphyrus lemnae (Fabricius). El Copal, Ira- Tribe POLYDRUSINI puato. 28lIW90. En follaje de Zea nzays L. (1 specimen). Pythis amplicollis Champion. El Copal, Irapua- to. 15Nl97. Medicago sativa L. (1 specimen). Subfamily HYPERINAE

Tribe SITONINI Tribe HYPERINI

Sitona hispidulus (Fabricius). Apaseo el Grande; Hypera postica (Gyllenhal). El Copal, Irapuato. Arandas, Irapuato; El Copal, Irapuato. 6NIII 12/III/96, 13lIIIl96, 41IW96, 71111197, 18lIVl97, 96, 16NIIIl97. Gramineae; Medicago sativa L. 2Nl97, 15Nl97, 1llIW97, 30lIW97, 27NII198. (5 specimens). Medicago sativa L. (57 specimens).

Tribe TANYMECINI Subfamily LIXINAE (formerly CLEONINAE)

Isodacrys geminatum Howden. Ojo de Calvillo, Tribe LIXINI Dolores Hidalgo. 16NIIl96. Gramineae. (1 spec- imen). Lixus inermis Champion. San Antonio, Salvati- Isodrusus debilis Sharp. 25 km NE Guanajuato; erra. 17NIIl96. Maleza. (8 specimens). Santa Rosa, Guanajuato; Rancho de Enmedio Lixus merula Suffrian. El Copal, Irapuato. 161x1 Guanajuato; Las Palomas, Guanajuato; 22 km 97. Polygo~zu~npurzctatuin Ell. (1 specimen). N de Guanajuato. 16NIIIl74, 16NII/96, 121x1 Microlarinus lypriformis (VVollaston). El Co- 96, 3NIII197, 21x1198. Quercus spp. (21 speci- pal, Irapuato. 28lIW96, 22lW96, 18lIVl97, 151 mens). Vl97, 17/XII/98, 19/1/99, 22/1/99, 2/11/99. Bac- Pandeleteinus subcancer Howden. 30 km NE clzaris sp.; Hyptis ~nutabilis(Rich.) Briq.; Med- Dolores Hidalgo. 16NIIIl74. (2 specimens). 52 Volume 15, No. 1, March, 2001, INSECTA MUNDI

icag-o sativa L.; trainpa pitfall en pastizal. (14 dimidiatus Champion. XichG. specimens). 3NIIIl96. En follaje de Martgifera iltdica L. (2 specimens). Subfamily MESOPTILIINAE (formerly Conotrachelus flavangulus Champion. El Co- MAGDALIDINAE) pal, Irapuato. 411x192, 4NIJ.196, 18lIVl97, 5NIl 97, llNIIl97. Trampa de luz. (5 specimens). Tribe LAEMOSACCINI Conotrachelus leucophaeatus FAhraeus. El Copal, Irapuato. 2NIl93, 3NIIl95, 1NII196, 81 exsculpt us Champion. Cruce Rio V/97,29Nl97,5/VI/97,4MI/97,1lNII197.Aster- Laja y carretera San Felipe-Guanajuato. 161 aceae; Rosa spp.; en trampa de luz. (15 speci- VIIl96. Maleza. (1 specimen). mens). Laemosaccus ruficornis Champion. Sierra de Conotrachelus lobatus Champion. Irapuato. Lobos, Le6n. 20NIIl96. Quercz~ssp. (1 speci- 19NIIl93. Trampa de luz. (1 specimen). men). Pheloconus rubicundulus (Boheman). El Co- pal, Irapuato. 281IW95, 9NIIl96, 81W96, 22lW Subfamily 96, 251x196, 10lIXl97, lllIXl97. Gramineae; Malva parviflora L.; Mal-rubiz~muulgare L. (98 Tribe CLEOGONINI specimens).

Rhyssomatus nigerrimus FAhraeus. El Copal, Family DRYOPHTHORIDAE Irapuato; Camino a la Garrida, Irapuato; Man- zanar, San Luis de la Paz; km 15 carretera Subfamily RHYNCHOPHORINAE Victoria-XichG; San JesGs del Monte, San Fran- cisco del Rinc6n; Cuenca de la Esperanza, Gua- Tribe LITOSOMINI (formerly SITOPHILINI) najuato; El Carmen, Purisima del Rincbn. 151 VI191, 20NIIl91, 4lIXl91, 8NIIl94, 18NII194, Sitophilusgranarius (Linnaeus). El Copal, Ira- 20NIIl94, 29NIIl94, 3NIII194, 10NIIIl94, 171 puato. 23/1/96. Zea rnays L. (1 specimen). VIIIl94, 23NIIIl94, 31NIIIl94, 7lIW94, 14lIW Sitophilus linearis (Herbst). Lebn. 3NII195. 94, 5lW94, 21IXI95, 3NIII196, 4NIII196, 231 Sobre suelo. (1 specimen). VIIIl96, 3 1NIII196, 8lIW96, 1llIW96, 181x196, Sitophilus oryzae (Linnaeus). El Copal, Irapua- 10NII/97,27NII/98,28NII/98,29NIII/98.Aca- to. 7lVIIl89, 121x195, 7lVIIl97. Gramineae; cia spp.; Arnarartthus spp.; Baccharis salicifo- Partlze~~iurnlzysteroplzorz~s L.; Zea mays L. (5 lia (Ruiz & Pav6n); Ficus spp.; Gramineae; specimens). Ipomoea spp., I. heredaceae Sess6 & Moc.; Mi- Sitophilus zeamais Motschulsky. El Copal, Ira- rnosa spp.; Prosopis spp.; P~~LLILUSpersicae L. puato. Zea rnays L. (22 specimens). (88 specimens). Rhyssomatus sculpturatus Champion. Valle Tribe SPHENOPHORINI de Santiago; El Copal, Irapuato. 141Xl90, 4lIW 91, 22NIIl93, 27lIW95, 2811x195, 21W95, 8lW Metamasius spinolae (Gyllenhal). 1 km E de 95, 4lIXl96, lllIXl96, 1811x196, 8lW96, 221x11 Rancho de Enmedio, Guanajuato; El Copal, 96. Baccharis spp.; Biderts spp.; Cupressus spp.; hapuato; San Antonio el Rico, Irapuato; km 39 Ipornoea spp., I. heredacea Sessb & Moc.; Mar- carretera Lebn-Ocampo; La Garrida, Irapuato; rz~biurnuulgare L. (37 specimens). Manzanar, San Luis de la Paz. 18NIl90, 201 VIIIl90, 9NIIIl94, 5NIIl95, llNIIl95, 28NIl Tribe CONOTRACHELINI 96, lNIIIl96, 3NIII196, 23NIl97, 28NIl97, 81 VII/97,28/VII/98:29NIIl98. Opurttia spp.; Qz~er- Conotrachelus aguacatae Barber. Las Ante- cus spp. (38 specimens). nas, Irapuato. 1511x197. En follaje de Persea Rhodobaenus auctus Chevrolat. Las Antenas, arnericana Mill. (21 specimens). Irapuato; El Copal, Irapuato. 9NIIl90, 16lXIl Conotrachelus cinerascens Champion. Las 91, lllXIl92, 22NII193, 6NIIl95, 12NII195, 91 Antenas, Irapuato. 29lIW97. En flor de Titho- VIIl96, 241x196, 4NIIl97. Arnarartthus spp.; nia tubaefol-rnis (Jacq.) Cass. (1 specimen). Aspa.ragus officirtalis L.; Gramineae; Plzysa.lis pubescens L.; Populus spp.; Sorghu~nhalap- INSECTA MUNDI, Vol. 15, No. 1, March, 2001

erzse (L.) Pers.; Tithortia tubaefornzis (Jacq.) species of Baridinae. During this study, 15 genera Cass.; Triticuln aestivurrz L. (11 specimens). and 24 species of this subfamily were identified. Rhodobaenus mundus (Champion). San Anto- The subfamily Entiminae is also one of the largest, nio, Salvatierra. 17NIIl96. Maleza. (1 speci- with 107 genera and 899 species; 16 genera and 23 men). species were identified in the material collected in Scyphophorus acupunctatus Gyllenhal. Puen- Guanajuato. The subfamily Tychiinae, which is tecillas, Guanajuato; Hacienda Silva, Romita. predominantly Nearctic, contains eight genera and 12/IV/00, 12Nl00. Aga,ve spp. (12 specimens). 58 species in the United States; five species of Sphenophorus cicatristriatus FBhraeus. Vil- Sibirzia were identified in this study. lagrhn. 10/XI/92. Maleza. (1 specimen). Sphenophorus phoeniciensis Chittenden. El Conclusions Dorado, Irapuato. 12/XII/93, 16/IX/97, 13NIl 98. Cyrzodorz dactylolz (L.). (4 specimens). Only a handful of major publications deal with Sphenophorus venatus (Say). El Dorado, Ira- Mexican Curculionoidea; to date, the only taxo- puato. 10/IV/97. Cyrzodorz dactylorz (L.). (1 spec- nomic reference work for all groups of Mexican imen). weevils is the Biologia Centrali-Americana (Sharp, 1889-1911; Champion, 1902-1909). There are some Faunal distribution studies corresponding to limited areas, such as The State of Guanajuato has 2.6% of the weevil Kissinger (1962) for the Yucatan Peninsula, as well genera and 0.68% of the weevil species recorded for as some taxonomic revisions including Mexican the American Continent, and 15.8% of the genera species (e.g., Barber, 1935; Clark,1982, 1983; Clarlr and 5.2% of the species recorded for Mexico. & Burke,1985, 1986, 1987; O'Brien, 1977, 1984, The geographic location of Central America, 1989; Vaurie, 1951, 1954, 1967, 1970, 1981). The including Mexico, allows an extensive faunal ex- present study contributes to knowledge of the dis- change, evidenced by the low endemism at the tribution of Mexican weevils and we believe that generic level. Only 29 genera of Curculionoidea are the 125 species identified represent less that half of endemic to Mexico and 57 to Central America; in the species present in the State of Guanajuato. addition, most endemic genera are monotypic (25 of The subfamilies Baridinae, Curculioninae and the 29 genera present in Mexico and 49 of the 57 Entiminae are represented by the largest number genera present in Central America), which sug- of genera. The former is one of the largest weevil gests that they are members of a highly evolved subfamilies but it is in need of revision. Many taxon (Anderson & O'Brien, 1996). It is probable species were collected that could not be identified. that the number of species listed in this work Eighteen percent of the species collected during represents less than half of the species present in this study are known to be pests of crops. This Guanajuato, since the irregular nature of the ter- percentage may increase later given the large num- rain provides various microhabitats that favor iso- ber of weevil species able to feed on cultivated lation of populations leading to speciation. plants. A comparison of the weevil fauna of Guanajua- to with the weevil fauna of other regions shows that Acknowledgements 53.7% of the species (e.g., Partdeleteius and Conotra,- chelus) are Neotropical, 26% (e.g., Slnicrorzyx) are We gratefully acknowledge the invaluable as- Nearctic, 18.4% (e.g., Sitophilus) are cosmopolitan, sistance of Dr. Guillermo J. Wibmer, who translat- and 1.9% previously were known only from the ed the text from Spanish to English and who made West Indies. Most of the Nearctic species are dis- numerous helpful suggestions and corrections. The tributed in the southwestern United States, and it research of C. W. O'Brien was supported in part by is possible that the corridor between the Sierra a grant FLAX 97007 from CSREES (Cooperative Madre Oriental and the Rocky Mountains has State Research, Education, and Extension Service, contributed to their spread. Some species are present USDA). in the southern United States, because the Mexi- can Plateau is an extension of the Chihuahuan Literature Cited desert. Three subfamilies (Baridinae, Entiminae and Molytinae) are primarily Neotropical. O'Brien Alonso-Zarazaga, M.A. and C.H.C. Lyal. 2000. & Wibmer (1978) record 411 genera and 2,968 A World Catalogue of Families and Genera of 54 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Curculionoidea (Insecta: Coleoptera) (Except- ic Publications. South Lancaster, Massachu- ing Scolytidae and Platypodidae). Entomoprax- setts. 141 pp. is. Barcelona. 315 pp. Luna, K. 1983. Contribuci6n a1 estudio de 10s Anderson, R.S. and C.W. O'Brien. 1996. Curcu- Colebptera, Curculionidae, Baridinae asocia- lionidae (Coleoptera). 62: Llorente B.J., A. Garcia dos a las Compositae de la Caiiada de Otongo, A., and E. GonzBlez S. (Eds.). Biodiversidad, Hgo. Tesis de Licenciatura. Facultad de Cien- Taxonomia y Biogeografia de Artr6podos de cias. UNAM. 15 pp. M6xico. Instituto de Biologia. UNAM. Mdxico, O'Brien, C.W. and G.J. Wibmer. 1978. Numbers D.F. 500 pp. of genera and species of Curculionidae (Co- Arnett, R.H. 1973. The of the United leoptera). Entomol. News, 89(3-4):89-92. States. The American Entomological Institute. O'Brien, C.W. and G.J. Wibmer. 1982. Annotat- New York. 1112 pp. ed checklist of the weevils (Curculionidae serzsu Arnett, R.H. 1985. American . A Handbook lato) of North America, Central America, and of the Insects of America North of Mexico. Van the West Indies (Coleoptera: Curculionoidea). Nostrand Reinhold Company. New York. 850 Mem. Am. Entomol. Inst. 34. The American PP. Entomological Institute. Ann Arbor, Michigan. Barber, H.S. 1935. The tobacco and solanum wee- 382 pp. vils of the Trichobaris. United States Sharp, D. 1889-1911. Biologia Centrali-America- Department ofAgriculture. Miscellaneous Pub- na. Insecta. Coleoptera. Rhynchophora. Curcu- lications, 226, pp. 1-28. lionidae. Attelabinae, Pterocolinae, Allocoryn- Bright, D.E. 1994. Revision of the genus Sitorza inae, Apioninae, Thecesterninae, Otiorhynchi- (Coleoptera: Curculionidae) of North America. nae (part, "Apterae"), vol. 4, pt. 3, pp. 1-40 Ann. Entomol. Soc. Am. 87(3):277-306. (1889); 41-80 (1890); 81-168 (1891); 169-178 Champion, G.C. 1902-1909. Biologia Centrali- (1911). Americana. Insecta. Coleoptera. Rhynchopho- Vaurie, P. 1951. Revision of the genus Calerzdra ra. Curculionidae. C~~culioninae(part), vol. 4, (formerly Spherzophorus) in the United States pt. 4, pp. I-VIII, 1-144 (1902); 145-312 (1903); and Mexico (Coleoptera, Curculionidae). Bull. 313-440 (1904); 441-600 (1905); 601-750 (1906); Am. Mus. Nat. Hist., 98(2):31-186. pt. 5, pp. I-VIII, 1-136 (1906); 137-240 (1907); Vaurie, P. 1954. New species of Calendra from 241-400 (1908); 401-513 (1909). Mexico, with notes on others (Coleoptera, Cur- Dillon, E.S. and L.S. Dillon. 1972. A Manual of culionidae). Am. Mus. Nov., 1681, pp. 1-8. Common Beetles of Eastern North America. Whitehead, D.R. 1980. The weevil Apiorz (Perap- Vol. 2. Dover Pub. New York. 894 pp. ion.) curtirostre Germar in North America, and Hatch, M.H. 1971. The Beetles ofthe Pacific North- the systematic position of Perapiorz Wagner west. University of Washington Press. Seattle. with a review of the North American species 662 pp. (Coleoptera: Curculionoidea: Apionidae). Co- Kissinger, D.G. 1962. The curculionid beetles leopt. Bull., 34(4):397-400. collected by the Explorers Club. American Wibmer, G.J. and C.W.O'Brien. 1986. Annotat- Museum of Natural History Entomological ed checklist of the weevils (Curculionidae ser~sz~ Expedition to Yucatan, Mexico, in 1952 (Co- lato) of South America (Coleoptera: Curculion- leoptera, Curculionidae). Am. Mus. Nov., 2086, oidea). Mem. Am. Entomol. Inst. Number 39. pp. 1-28. The American Entomological Institute. Gaines- Kissinger, D.G. 1964. Curculionidae of America ville, Florida. 563 pp. North of Mexico. A key to the genera. Taxonom- Zirnmerman, E. 1994. Australian Weevils. Vol. 1. CSIRO. Australia. 741 pp. INSECTA MUNDI, Vol. 15, No. 1, March, 2001 55

Table 1. Families, subfamilies, tribes, genera and species of Curcullonoidea from the State of Guanajuato, Mexico

FAMILIES SUBFARIILIES TRIBES SPECIES (* = new for Mexico) Apionidae Apioninae Apionini Apio~zspp. Apion nr. te7~zcirostr~c~nSmith Apion nr. varicor~~eSmith Piezotrachelini Clzrysapion arect~e~n(Sharp) Attelabidae Attelabinae Attelabini Hi~natolabuszclnbosis Hamilton Pilolabini Pilolabus lzlrcgii (Gyllenhal) Rhynchitidae Rhynchitinae Rhynchitini Eug~ia~nptusspp. Haplorlzynclzites 7nesica7iz~s(Gyllenhal) " Haplorhynclzites quadripe~zizis(Fall) Haplorlzynchites subaz~ratzcs(Sharp) Involvzelr~s~nollis (Sharp) Curculionidae Baridinae Baridini Baris corr~csca(Boheinan) Baris strenua (LeConte) Pseudobaris apicalis (Boheman) Trepobaris perlo~~gaChampion Triclzobaris clzarnpiorii Barber Trichobaris rnajor Barber T7-iclzobaris pellicea (Boheman) Madarini " Onychobaris arrzbigua Casey Stictobaris ornatella Casey Madopterini " Apinocis angustus (Casey) " Barilepto7i qrcadricolle LeConte Barinz~sspp. Centri7zoprcs rnistzcs Champion Geraeus candidus Champion Geraeus dugesi Champion Geraeus lzospes (Casey) Geraeus perscitus (Herbst) Geraeus senilis (Gyllenhal) Gerae~cstonsus Champion Geraeus uarius Champion Li~nnobarisdiscreta Champion * Nicentrus fe~noralisChampion Psezcdoce7ztrinres ochracer~s(Boheman) Optatini Optatus pallnaris (Pascoe)

Ceutorhynchinae Ceutorhynchini Ceretorhynclzzes trzcquii (Champion) Cnemogonini * Azcleutes asper (LeConte) Phytobiini " Rlzi7zoncus pyrrhopus Boheman

Conoderinae Lechriopini Lechriops auritus (Schonherr) Piazurini Cratoso~nuspunctulatus Gyllenhal Zygopini Cyliizdrocopturus adspersus (LeConte) Cylindrocopturzcs biradiatus Champion C~~lindrocopturusga7zglbaueri (Heller)

Cossoninae Cossonini Cossonus co~zuezirostrisChampion Dryotribini Cazelophilus oryzae (Gyllenhal) Cryptorhynchinae Cryptorhynchini Gerstaeclzeria mutillaria (Gerstaecker) Gerstaeckeria u7zicolor Fisher Maernactes punctatus Kssinger Pseudol~zopsissi~nilis Champion

Curculioninae Anthonomini A7ztho1to1nresdeliqulus Clark & Burke A7zthonornus eugeliii Cano 56 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Table 1. Continued. FAMILIES SUBFAMILIES TRIBES SPECIES (* = new for Mexico) A7~tho7zo~nz~s~nericaizz~s Bohelnan Ai~tJzo7~07lzz~sS~LL~T~L~IZS Chainpion * A~L~~LO~LO~~LLSsz~tz~ralisLeConte A7ztlzoizo7~zz~~tera7zzcs Dietz " A~L~~LOILO~?~ZLStlzyasoc7~e1noides Hustache Pser~dai~tlzoi~o7~z~~sglz~oi~Clark "S~~zicrazclartr~berczslatz~s Pierce Curculionini Cz~rcz~lioortegai Gibson Derelonlini Plzyllotror sz~tz~ralis(Boheman) Otidocephalilli Myr17ze.u spp. Prionobrachiini Prio~zobraclziuilzsclzoe7tl~erri Faust Smicronychiili " S1lzicro7yr albo7~otatz~sAnderson " Sinicr-onyx coi.lzicz~latz~s(FAhraeus) " S17zict-oi~yrqz~,adt-ifer Casey " S1rricro7yr spretzss Dietz S~?zici*oi~yrte7~z~irostris Chalnpion Tychiini Sibitzia azclacis Clarlr Sibinia caizdidata Champion Sibizia iizei-i~zis(Casey) Sibitria setosa (LeConte) Sibi~riasuturalis (Schaeffer) (lircel-tae sedis) Macror1zoptz1,ssplzaeralciae Pierce

Rhytirrhiniili Listro~rotl~sgrypidioides (Dietz) List~-oizotz~sp~~i~ctigel. LeConte

A~neladz~siizoriaatzi,~ Sharp Clzaetopa7ztz~ssp. Mitostylz~ssetosz~s (Sharp) Geoneinini Bradyrl~yylzclzoidesrz~gicollis (Sharp) Epicae7-zrs costatzss Sharp Epicaerzcs sqr~alidz~sSharp A7az1,pactopsis delz~nzbisChainpioil Megalostylr~salbicans (Lacordaire) Pa7zto1~zor.r~~albosig1~atzi.s Boheinan Pa7ztorizorr~scervii~zcs (Boheman) Pa7rtorrl,orns LLIL~~OTIIZ~SSharp Ophryastini Oplzryastes ciizererrs FAhraeus Ophryastes sordidrcs LeConte Polydrusini Pytlzis arlzplicollis Champion Sitonini " Sitorza lzispidulz~s(Fabricius) Tanyinecini Isodacrys ge1rzi7~atr~1lzHowden Isodrzssz~sdebilis Sharp Pa7zdeletei~zr~ssz~bcaizcer Howden Pai~deleteiz~sciliatipei~~~is Champion Pa~zdeleteiustibialis Boheman Pal~deleteiusviridive7ztris Chainpion Paizdeleteir~svitticollis Champion Tar~y~necz~sco~zfz~sus Say

Erirhininae Tanysphyrini " Taizysp1zy.yrzcs lerlz7zae (Fabricius)

Hyperinae Hyperini " H3per-a postica (Gyllenhal)

Lixiilae Lixini Lirus i7eernzis Champion " Lirz~s~~zerz~la SufGiall INSECTA MUNDI, Vol. 159No. 1, March, 2001 57

Table 1. Continued.

FAMILIES SUBFAMILIES TRIBES SPECIES ("" = new for Mexico) " Microlarii~uslyprifori7zis (Wollaston)

Mesoptiliinae Laemosaccini Lae~~zosaccz~sexsculptz~s Chainpion Lae~~zosaccz~srzsficorizis Champion

Molytinae Cleogonini Rlz~~sso~~zatz~s1zige7-ri77zz~s FBhraeus Rhyssol7zatz~ssculptzrratz~s Champion Conotrachelini Co~zotraclzelz~saguacatae Barber Coicotraclzelz~sciizerascei~s Champlon Co7~otrachelz~sdilrzidratz~s Chalnpion Co7zotraclzelr~sflaua7zgr~lz~s Champlon Coizotraclzelr~sle~~coplzaeatus FBhraeus Colzoti~achelr~slobatz~s Champion Plzelocol~zis1.z~bi~z~7zdz~lz~s (Boheman)

Dryophtlloridae Rhynchophorinae Litosolnini Sitophilz~sgra7zariz~s (Linnaeus) Sitophilus li7zearis (Herbst) Sitophilz~soryzae (Linnaeus) Sitophilz~szealnuis Motschulsliy Sphenophorini Metal~lasiz~sspi7zolae (Gyllenhal) Rlzodobae7zz~sazrctz~s Chevrolat Rlzodobae~~z~s112 ZIIL~ZLS (Champion) Scyphophorus acz~pz~i~ctatz~sGyllenhal Sphei~ophorrrscicatristriatz~s FBhraeus Sphe7~oplzo1.u~plzoe7zicier~sis Chittenden Splre~~oplzorz~sue7~atz1,s (Say)