Middle and Upper Ordovician Nautiloid Cephalopods of the Cincinnati Arch Region of Kentucky, Indiana, and Ohio
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Middle and Upper Ordovician Nautiloid Cephalopods of the Cincinnati Arch Region of Kentucky, Indiana, and Ohio U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1066-P Prepared in cooperation with the Commonwealth ofKentucky, University ofKentucky, and Kentucky Geological Survey AVAILABILITY OF BOOKS AND MAPS OF THE U.S. GEOLOGICAL SURVEY Instructions on ordering publications of the U.S. Geological Survey, along with prices of the last offerings, are given in the current- year issues of the monthly catalog "New Publications of the U.S. Geological Survey." Prices of available U.S. Geological Survey publica tions released prior to the current year are listed in the most recent annual "Price and Availability List." Publications that may be listed in various U.S. Geological Survey catalogs (see back inside cover) but not listed in the most recent annual "Price and Availability List" may be no longer available. 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GEOLOGICAL SURVEY PROFESSIONAL PA PER 1066-P Prepared in cooperation with the Commonwealth of Kentucky, University ofKentucky, and Kentucky Geological Survey Stratigraphic distribution, pakoecology, biogeography, and systematic paleontology of 50 species of Ordovician nautiloids from the midcontinent UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1995 U.S. DEPARTMENT OF THE INTERIOR BRUCE BABBITT, Secretary U.S. GEOLOGICAL SURVEY GORDON P. EATON, Director Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply endorsement by the U.S. Government. Library of Congress Cataloging in Publication Data Frey, Robert C. Middle and Upper Ordovician nautiloid cephalopods of the Cincinnati arch region of Kentucky, Indiana, and Ohio / by Robert C. Frey. p. cm. — (U.S. Geological Survey professional paper; 1066-P) (Contributions to the Ordovician paleontology of Kentucky and nearby states) Includes bibliographical references and index. Supt. of Docs, no.: 119.16:1066-P 1. Nautiloidea, Fossil—Kentucky. 2. Nautiloidea, Fossil—Indiana. 3. Nautiloidea, Fossil—Ohio. 4. Paleontology—Ordovician. 5. Animals, Fossil—Kentucky. 6. Animals, Fossil—Indiana. 7. Animals, Fossil—Ohio. I. Title. II. Series. III. Series: Contributions to the Ordovician paleontology of Kentucky and nearby states. QE807.N4F74 1995 564'.52'0977—dc20 94-27353 CIP For sale by U.S. Geological Survey, Information Services Box 25286, Federal Center, Denver, CO 80225 CONTENTS Page Page Abstract............................................................................................. PI Distribution and Fauna! Affinities of Ordovician Nautiloids in Introduction...................................................................................... 2 the Cincinnati Arch Region—Continued Acknowledgments ........................................................................... 3 Nautiloid Fauna of the Cincinnatian Provincial Series ......... P16 Regional Ordovician Paleogeographic Setting............................. 3 Edenian Nautiloid Fauna................................................. 16 Ordovician Lithostratigraphy in the Cincinnati Arch Region.... 4 Maysvillian and Early Richmondian Nautiloid Fauna. 17 Middle Ordovician Champlainian Provincial Series Nautiloid Fauna of the Leipers Limestone.................... 19 Lithostratigraphy in Kentucky ........................................ 6 Middle and Late Richmondian Nautiloid Faunas......... 19 High Bridge Group................................................................. 6 Late Ordovician Extinction Event............................................ 20 Lexington Limestone..........................................'................... 6 Nautiloid Taphonomy........................................................................... 21 Upper Ordovician Cincinnatian Provincial Series Nautiloid Biostratinomy............................................................ 21 Lithostratigraphy ............................................................. Silicified Nautiloids.................................................................... 22 Edenian Stage......................................................................... Nonsilicified Nautiloids............................................................. 22 Maysvillian Stage................................................................... 9 Nautiloid Classification....................................................................... 23 Richmondian Stage................................................................ 9 Systematic Descriptions...................................................................... 24 Distribution and Fauna! Affinities of Ordovician Nautiloids in Systematic Paleontology..................................................................... 27 the Cincinnati Arch Region......................... 10 References Cited................................................................................... 113 Nautiloid Fauna of the Tyrone Limestone...... 10 Appendix: Locality Register................................................................ 120 Rocklandian Nautiloid Extinction.................... 12 Index...................................................................................................... 125 Nautiloid Fauna of the Lexington Limestone., 15 ILLUSTRATIONS [Plates follow index) PLATE 1. Cartersoceras and Murrayoceras 2. Pojetoceras and Pleurorthoceras 3. Ordogeisonoceras and Geisonoceras 4. Proteoceras and Treptoceras 5-8. Treptoceras 9. Monomuchites 10. Isorthoceras 11. Polygrammocerast 12. Polygrammoceras, Anaspyroceras, and Gorbyoceras 13. Richmondoceras 14. Cameroceras 15. Vaginoceras 16. Triendocerast and Actinoceras 17. Actinoceras 18. Ormoceras, Deiroceras, and Orthonybyoceras 19. Plectoceras, Trocholites, and Oncoceras 20. Oncoceras, Bdoitoceras, Maelonoceras, Laphamoceras, Allumettoceras, Augustoceras, and Manitoulinoceras 21. Diestoceras and Beloitoceras 22. Diestoceras and Beloitoceras III IV CONTENTS Page FIGURE 1. Map showing Middle Ordovician paleogeography in Eastern North America............................................................................................ P4 2,3. Charts showing: 2. Ordovician lithostratigraphy in the Cincinnati arch region............................................................................................................. 5 3. Lexington Limestone lithostratigraphy............................................................................................................................................... 7 4,5. Diagrams showing: 4. External features of nautiloids............................................................................................................................................................ 25 5. Internal features of nautiloids.............................................................................................................................................................. 26 6. Drawing showing internal features of Cartersoceras shideleri.................................................................................................................... 29 7. Bivariate plot of measurements made on specimens of Pojetoceras floweri and Orthoceras regularis .........................................