S ENCKENBERGIANA BIOLOGI CA Wissenschaftliche Mitteilungen Der Senckenbergischen N Aturforschenden Gesellschaft

S ENCKENBERGIANA BIOLOGI CA Wissenschaftliche Mitteilungen der Senckenbergischen N aturforschenden Gesellschaft

Schriftleitung Dr. DIETER KOCK, Dr. HEINZ SU-IRODER und Dr. RICHARD ZUR STRASS EN

Senckenbergiana bioI. 170 (1 98 9) 1 (1/3) \ - 13 Frankfurt am M ain, 2. S. 1990

Eptesicus (Eptesicus) bottae (PETERS 1869) in Saudi Arab ia with notes on its subspecies and distribution (Mammalia: Chiroptera : Vespertilionidae). By

hAD A. NA DER, & DI ET ER KOCK,

Riyadh , FrJtlkiun ;t. M

\,\Ii t h 1 map and 1 table.

A b s t r act : Eptesicli s boltae is an additio nal species to t he bat fauna of Saudi Arabia and data fo r the nominate su bspecies are given for the first time since its original description Addit ional measurements and n otes are given for recognition o f t he known subspecies innesi, hingstol1l, ana lOiL clis (new mater ial), taJianimontis, and ognevi (new material); jists of synonyms are attempted for th ese subspecies and their distribution is mapped. Data on reproductive activity are included. - E. hingstoni co llected at Shanna/SE-Arabia actually represents E. nasulus; E. nilsson i recorded fo r Iraq had been re-i dentifi ed as E. ba ltae.

Eptmcus (Epteslcus) bottae (P ETERS 1869) was collected recently In SW-Arabia. It was not Included preVi ously in the bat fauna of Saud i Arabia; the total number of bat species thus raises to 22 for this country.

A c know 1e d g e m e n t s: For the loan of specimens, we thank Dr. H. VI ERHA US (Bad Sass endorf-Lohne, F RG ), and gratefully recogn ize the deposit of refere nce speci mens in (he SMF collec(ion by Prof. V. HA NA K (Prague, CSSR), (he re-examinat ion of the bOllae h olot ype - by Dr. F. PET TER , M usee N ational d'Histo ire Naturell e, Paris (M H N P), the re-examinat ion of an u nsuffi ciently documented specimen o f "E. hingstoni" by D r. J. E . HILL, Bri(ish M useum of N atu ral H is rory, Lo ndon (BM), and t he valuable transla(ions made fo r us by Mrs. K. JENTOCH (SMF) and Dr. S. I. GOLOVATCH , Moscow. - J 2

Tab.!. Body and sku Jl measurements of sub Eptesicus (Eptesicus) bottae bottae (PETERS 1869).

1869 lIesperus bottae PETERS, Mber. preuss. Akad. \XIiss. Berlin, 1869: 406; terra typ. : Arabi a omaner [= S\XI-Yem en], holotype ad. 9; leg. [Po E.] BO TTA 1837 [I.! S37 = date of deposit in bottae MNHP]. PETERS HARRISON this paper HARRIS! 187 8 Vesperugo serotinus, - DOB SON, Cat. Chiropt. coi l. Brit. Mus.,: 191; n. comb., bottae IB69 1964 Type d a synonym. Type ~ 400 - 114 91 1897 [Vespemgo (Vespe rus) ] serolinus, - TROUESSART, Cat. Mamm. vivo quam foss., 1: 107; 100 102 Total - 50 40 BOltac a syn., terra typ.: N-Africa [sic l ]. 43.5 41 Tail - 10 7.8 B.O 7 1941 Eptesicus serolinus, - RODE , Bull. Mus. na tn. Hist. nat. , (2) 13 (4): 247; 0' [= 9] HF 17 - 17 14.2 E 14 holotype of bOllae, terra typ.: Arabia; n. comb. H+B 41 - 44 39 1942 Eptesicu5 (Eptesicus) bOHae, - TATE, Bull. Amer. Mus. nat. H ist., 80 (7): 275,277; loco FA 40 15 . 7 17.0 16. 4- 17.0 typ.: Botta [s ic!], Arabi a. Gl 15 . 6- 16.1 14.9 CBl 10 .4- 10.7 10.3 1951 Eptesicus isabellinus bOllae, - ELLERM ANN & MORRI SO N-SCOTT, Checklist Palaearct. ZB 11.0 7.4 B.O 8 .0- 8.2 lndian Mamm.,: 156 ; [restricted] terra typ.: Yeme n; n. comb. BB 3.7- 3.8 3.6 IC 4.0 5.4 1964 Eptesicus bottae bottae, - H ARRISON, Mamm. Arabia, 1: 140; terra typ.: Yemen; 0' [= 6.0 5.9- 6.0 C-H3/ 6 .2 7.0 6.6- 6.7 9] nolotype, sku ll extracted. C-H/3 11.9- 12.3 11.1 H 12.0 Ma t e rial: Sa u d i Arab i a: Misfe ra, 40 km (by road) N Abha, 9.Y., 5. & 8VI.1987, 30'0' ad. (3 sk ull s, 3 skins), ColI. IA N 1131, 1153-54; llVI.198 7, J 0' ad. (skull, skin), SMF 73811, leg. AWADH AL AHMERY. M easurem ent s: Se e Tab. 1. anatoJicus HARRISON 1964, HARRISON H o I 0 t y p e: According to the origin al description, the holotype is an ad. 9 FELTEN DeBlASE 1980, this pape in alcohol; the skull since then has been extracted and its measurements were give n 1971 NADER & KDCK 1983 4dd by HARRISON (1964) for the first time. RODE (1941) and HARRISON (1964) stated Type ~ 6~~ 107 -126.5 105 -120 that the type is a 0'. The holotype (MHNP 212) was kindly re-exa mined by Dr. Total 42.0- 54.0 40 .0- 52 Tail F. PITTER and confirmed that it is indeed a 9. 9.0- 10.0 7 .B- ~ HF 16 .4- 11 14 .0- 18.0 E 0 H+B T y pel c a lit y : E. bottae was described (P ETT ERS 18 69) witho ut exact locality 46.2- 49.4 ~.~ 5 FA 45.0 fro m Arabia. 1.LLERMANN & MOR RISON -SCOTT (1951) restricted the terra typica to 17.2- 18. 8 U.~~ Gl 17 .9 W .~l 17 _7 16.5- 17.B Yemen (= North Yemen of today; South Yem en of today at that time consisted C8L 12.~1 12.9 12.4- 12.8 05 Z8 7 .f.- 9 .1 02 B.~ of several Sultanates) without giving any elaboration on their action. - A similar BB 3. 4- 4.2 3.~ IC 4 .2 hi story is attache d to other items coll ected by P. E. BOTTA: the stone chat, Oenanthe 6.0- 6.8 6.~ C- H3/ 6.6 6.8- 7 . 9 6.7 botlae (BONAPARTE 1854), was described without a type locality, later restricted to C-H/3 12.6- 14 . 1 12.7-; "Yemen" (ME INERTZH AGEN 1954); the chameleon, ChamaeLeo caLyptratus D UMERIL H & DUME RIL 1851, described from the Nile Region , was restricted to a type locality in Yemen (HILLEN IUS & GASPERETTl 1984), as its collector journeyed in Yeme n from Hodeida (Hudaydah) in IX. 183 6 to Hays, Ta'izz, back to Hays then to Al Mukha from where he departed (BOTTA 1841). This travel rome restricts the terra ogn eV'i typica to SW -Yemen). Oe8lASE 1980 STR ElKOV et a1. H 27ZZ 3dd 1066 Ta x 0 nom y : HARRISON (1964) while characterizing the different subspecies 93 - 101 02 Total 42.0-50.0 41.0-! of E. bottae had only the alcohol preserved holotype of boltae ava ilable to him. Tail 44 - 45 02 9 - 10 02 He suggested then that the nominate form may be synonymous wi t h the closely HF 16.5-1B.2 17 .0 E 15 - 19 57.2 related innesi (LATAST E 1887) . The newly collected material from Saudi Ar~hia is 53.5-69.0 H+B - 43.5 38 .0 FA 42.4- 45.8 16.0 assigned to b. bottae due to the geographic proximity to the terra typica "SW 17.1- 17.5 15.5- 17.0 GL 15 .7-16.B 16.1 Yemen", and gives the opportunity to com pare it with znnesi. E. b. bottae is darker CBl 1B.B- 16.9 10. 6 10 .9- 12.2 02 10 .5-11.1 ZB 7.6- 8.3 7.6· in color of the upper parts, being brownish ye ll ow and the lower parts paler. The 8.8- 9.4 BB 3.6- 4.2 3.7· 3.B- 4.1 base of the hair is blackish. The wing membranes and the ears are dark brown close IC 6 .0- 6.3 5.6· 6.0- 6. 5 C-H3/ 6.5- 7.1 6.1 to blackish. In size it is larger in greatest length of skull, zygo matic breadth, 6.7- 7.2 C- H/3 11.7-12.3 11.4 breadth of braincase, maxillary and mandibular toothrows, and mandible length. H 12.9- 13.0 3

bottae (PETERS 1869). Tab. 1. Body and sk ull measurements of subspecies currently recogni zed of Eplesicus bollae. \'(fiss. Berlin, 1869: 406; terra typ.: Ara bia I BOTTA 1837 [1.l837 = dare of d . ep OSH In bottae omanensis innesi WASSIF 1962, )t. coil. Brit. Mus ' 191· 0 co mb b ')' ,. 'J OU ae PETERS HARRISON HARRISON 1963 HARRI SON 1963, 1869 1964 this paper HARRISON 1976 NADER & KOCK 1983 Lectotype RT, Cat. Mamm. vivo qu am (oss ., 1: 107; Type ~ 400 Type 0 2M o 2Z~ 300 Total 100 102 -114 91 92 90 99.3 98 99.5 97 n1 n. Hist. nat. , (2) 13 (4): 247; 0' [= 9) Tail 43.5 41 - 50 40 43 41 42.6 44 40.3 39.3 - 42 .0 n2 >mb. HF 8.0 7 - 10 7.8 7.8, 7.8 8.6 7.7, 8.9 7 .3 - 8 . 3 E 14 17 - 17 14.2 15.2, 14.7 14.5 13.6, 16.5 13.4 - 15 .0 ler. Mus. nat. Hist., 80 (7): 275, 277; loco H+8 FA 40 41 - 44 39 40.8, 39.3 41.7 40.3, 42. 3 40.3 n2 GL 17.0 16 . 4- 17.0 15.7 15.3, 14.9 16.3 16.6 15.6 - 16 . 1 n2 : MORRI SON ,SCOTT , Checkl ist Palaea rct. CBL 15.6- 16.1 14.9 14.4 15.6 16.0 15.2 - 15 .3 Yemen; n. comb . 28 11.0 10.4- 10.7 10.3 10.0. 9.7 10.4 10.4 10.35- 11 .0 n2 8.0 8.0 7.4 7.7 7.2 Arabia, 1: 14 0 v 8B 8.2 7.2, 7.0 7. 2 - B.1 ; rerra typ.: l emen; 0' [= IC 4.0 3.7 3.8 3.6 3.4, 3.5 3.8 3. 3 3.8 - 3.9 C-/13/ 6.0 5.9 6.0 5.4 5.3, 5.2 5.6 5.7 5.3 - 6.8 C- M/3 7.0 6.6 6.7 6.2 6.0, 5.7 6.1 6.3 5.9 - 7.4 (by road) N Abha, 9,V, S. & 8V1.l 98 7, M 12.0 11.9- 12.3 11.1 11.0, 10.5 11.6 12.0 11. 0 - 11. 7 n2 )4; I1VU987, 10' ad. (skull, ski n), SMF

scription, the holotype is an ad. 9 anato[icus hingstoni HARRISON 1964, ted and its measurements were gi ven FEL TEN OeBLASE 1980, HARRISON 1964, HARRISON 1964 1971 NADER & KOCK 1983 this paper ,(1941) and HARRISON (1964) stated Type .12) was klOdly re-examined by D Type ~ &I'~ 400 sex? 7~~ 500 9. ~ Total 107 -126.5 105 -120.7 106 - 114 102 -110 .5 Tail 42 .0 54.0 40.0- 52.3 43 - 52 44 - 51.5 HF 9.0- 10.0 7.8 9.4 n3 9 . 3 7.5 10.0 9 - 10 .1 )ETTERS 1869) without exact locality E 14 .0 18.0 16 . 4- 18.0 14 - 17 14 - 18 H+B (1951) restncted the terra typica to FA 45.0 46.2- 49.4 46.0- 51.7 45 .0 45.9- 50.0 n5 43.0- 46 .3 len of today at that time consisted GL 17.9 17.2- 18.8 17 .1- 18.3 16 .6 17 .3 IB.O n4 16.7- 17.4 n3 C8L 17.7 16.5- 17.8 16.6-17.8 15.9 16 .8 17.4 n4 16.1- 16.8 n3 ratIon on their ac ti on. - A simi lar ZB 12.9 12.4- 12.8 n5 12 .0 12.3 n2 11.2 11 . 0- 11.8 n2 10.9 11.6 n2 E. BOTTA: the Stone chat, Oenanthe BB 7.f 9.1 n2 B.O 8.6 7.9 7.6 B n4 7.5 7.B n3 Ie 4.2 3.4 4.2 3 . 5 3.8 3 . 6 3 . 3 3.9 n4 3.4 3 . 7 n3 l[ a type locality, later restricte d to C-M3/ 6.6 6.0 6.8 6.0 6.2 5.8 5.9 n4 5.7 6.1 n3 on, Chamaeleo calyptratus DUMERIL C-H/3 6.8 7.9 6.7 6.9 6.7 6 .5 6.8 n4 6.6 6 . 8 n3 M 12.6- 14.1 12.7 13.2 12 . 0 11 . 6- 13.0 n4 12.2- 12 . 8 n3 .on, was restricted to a type locality > Its collector Journeyed in Yemen ;, Ta 'izz, back to Hays then to Al ThIS travel route restricts the terra ognevi taftanimontis

O.BLASE 1980 STRELKOV et al. 1978 ROGUIN 198B 'acterizing the different subspecies 300 1000 27~Z Type ~ 3n ,lotype of bottae available to him. Total 93 -101 n2 be synonymous with the closely Tail 44 - 45 n2 42.0-50 .0 41.0 53.0 HF 9 - 10 n2 8.5 ted matenal from Saudi Arabia is E 15 - 19 16.5-18.2 17.0-20.4 14 )Xlmlty to the terra typica "SW H+8 53.5-69.0 57.2-68.5 57 FA 42.4- 45. 8 - 43.5 38.0-45.3 42. 85 41.2 -44.65 t with mnesi. E. b. bottae is darker GL 17.1- 17.5 15.5-17.0 16.0-17.5 16.75 15.95-17.25 ow and the lower pans paler. The C8L 1B.8- 16.9 15.7-16.8 16.1-17.1 16.4 15.45-17 . 0 ZB 10.9- 12.2 n2 10.5-11.1 10.6-11.7 10.95 10.45-11.30 ; and the ears are dark brown close 88 8.8 9.4 7.6- 8.3 7.6- 8.2 IC 3.8 4.1 3. 6- 4.2 3.7- 4.2 3.7 3.7 - 3.9 gth of skull, zygomatic breadth C-1>I3/ 6.0 6.5 6.0- 6.3 5.6- 6.5 6.0 5.85- 6 . 35 toothrows, and mandible length ' C- M/3 6.7 7.2 6.5- 7.1 6.1- 7.1 6.5 6.3 - 6.95 M 12.9- 13 . 0 11.7-12.3 11.4-12 .8 12.65 12.15-12 . 9 4

d" H . ST J. P HlLBY, 22. 11.1932) was examined v measurements are smaller than bottae, he not defi ni tel y allocated. Dr. J. E. H ILL ki n o measured, an d id entified as Eptesicu s naS 0 0 ~ ,~~ ~~ o <7 Eptesicus (Eptesicus) oo oj ___.: __ _ .0 0,' A sy nonymy for this su b,pecies o o ~, Mate ri al : See N.\ DER & KoCK (E -- 0 19 .. ~ / '''''. M ea s ur e m e n t s : See Tab. 1. o 0 ~ 8 ..' , .': ,." ~ 09-_q-~. " . og ~-'>----'?~~ .. '- O?..'-:r ::-"-';. .' _ . ~___ ''''" The known localities (Map 1) f 0 .., rnari zed by NA DER & KOC K (1983). 0 "- , " , ' . . . spec imen from Abu Rawash as did :) , ' - -,. ',: ' / ~:- Geddi. BATES & HARRISON (1989) I Jorda n (nr. Ru m; too late for inclu

-- -~ ...

~ '\ Eptesicus (Eptesicus) I l ~ '-. "'--..J :_. \ 1913 WETTSTE IN, Eptesicu5 seYocinus, - ~ spec ime ns from Iraq [see SPITZEl' 1919 Ep tesicus hingston i THOMAS, J. Bon .. 1942 Eptesicus (Eptesicus) hingstoni, - " 1951 Eptesicus sodalis hingstoni, - ELI Indian Mamm.,: 156; n. comb. o I A ~ .... ~ ~. 195 6 Eptesicus hings lOni, - H ARRISON, • bottae Proc. zool. So c., 127: 452. I .:' ... Orlwnensis 1959 Epresicus nilssoni nilssoni, - HAT " InneS I Ll. hlngs toni 31; specimen fro m Baghdad [HA I • an atohcus 1963 EpresiCIIs innes i, - H ARRtSON, Z ,' ~ o CKjI)(lVI 'Y. Q 250 500 km .. taftanimont is 1964 Epresicl{s botlae hmgstoni, - H A 1966 Epres lcu5 serot in ll s, - AL ROB AAE Map 1. Known di stribution of Eptesicus botta e; type localities are prese nted by sy mbols w ith SMF [see FELTEN 1971: 372). double contours; localities not known exactl y are indi ca ted by ques ti ons mark . - Drawin g: Materi al: See FELT EN (197 1) fo Mus. Se nck. (JULIA ALTM ANN) . Suleimania, 15.XI.196 1, 3a a 5 99 E ROB AAE . - I r a q: Al Mahmudiya, 24.1 2 skeletons), Coil. IAN 247-8,256- 67 leg. I. A. NADE R. - Baqubah, 10.X.!' T he range of I1leasurements now es tablish ed for b. bottae in gen er al is bel ow (now at BRCB) , leg . I A. NADE R. that of hingstoni and anatolicus; these twO subspecies geographically separate b. bot Mea s u r e m e n t s : See Tab. 1. tae fr om the rather si mil ar sized ognevi, which is howe ver clearly lighter in colora tion. On th e ot her hand b. bottae is larger (han omanensis in forearm, skull and The occurrence of E. b. hingsto toothrow lengths, and is brow nish instead of greyish. by the records co mpiled in HARI record from 35 km N An-Naja£. E Di s t rib u ti 0 n (Map 1): The no mInate subspecies has been co ll ec ted on ly ting sites, FELTEN (197 1) gave new twice (SW-Yem en; Misfe ra/Saudi Arabia, see above). dealt with by AL ROBAA E (196( Notc: DOllMAN (1933) recorded an Epleslcu5 sp. from Shanna, which POCO CK (19 35) reported by WETTSTEIN (1913) as 1 identified as E. hmgstoni ( = E. bottae hingstoni). This locality (18.58. N .-51.05.E.) is close and H ARR ISO N(1964), were re-ideJ to the undefined borders of Saudi Arabia, South Yemen and O man. H AR RISON ( 1964) stated adding Babylon as a further loca that this specimen (skull not found) may belong to £. bottae, but gave no external HATT (1959: 31) listed E sodail mea suremcnts. T he spec imen (BM 34.8.8.1, <;> in ale ., sku ll extracted , but nor located, leg . records, but the only specimens 5

H. ST. J. PHI LBY , 22 .U.1932) was ex am ined for the present study (IAN, Il.1 989). T he external measurements are sma lle r tha n bo ltae, however, without its skull, the specific identity was not definitely all ocated . Dr. J. E. H JLL kindly informed us (in litt.) that the skull was located, measured, and ide nti fied as Eptesicus nasullis (DOBSON 1877).

Eptesicus (Eptesicus) bottae innesi (LATASTE 1887).

A syno nymy for this subspec ies wa s given by N ADER & KO C K (1983). M a t erial: See NAD ER & KO CK (198 3) . Nl e a s ur e m en t s : See Tab. 1. Th e k nown local ities (Map 1) fo r mnesi in Egypt and Palest ine were sum marized by NAD ER & KO CK (1983). WA SSIF et a!. (1984) reponed agai n a known \ ~ - I specimen from Abu Rawash as did QUM SIY EH (1985) on a specim en from Ei n Geddi. BATES & H ARtUSO N (1989) record this subspecies for the first time fro m ' . ,~ - ~ - Jordan (nr. Rum; roo la te for inclu sion in Map 1). l, ~ ., -_~- " - '--' , ~ , Eptesicus (Eptes icus) bottae hingstoni THOMA S J919. 1913 Eptesieus serotinus, - WETTSTEI N, Ann. k. k. naturhist. Hofm us . \XIien, 27 (1 9): 466 ; spec imens from Iraq [see SPITZ EN BERGER 1979: 455 ]. 191 9 Eptesieus hingst on i THO MAS, J. Bo mbay nat. Hist. Soc., 26: 745; loco typ.: Baghdad, Iraq. 194 2 Eptesieus (I:.ptesicus) hingstoni, - TATE , Bull. Am er. Mus. nat. Hist. , 80 (7): 27 5, 277. 1951 Eptesicus soda lis hlngstoni, - ELLERMANN & MOR RI SON -S COTT, Checklist Palaearct. fndian Ma mm ., : 156; n co mb. 1956 Eptesleus hingsto nl, - HARR ISON , Bonn. zooL Bei tr., 7 (1/3): 2; Mamm ., 37 (2): 262 ; • bottae Proe. zoo!. Soc., 127: 452. • om(,)nensis • Innes. 1959 Ep leS icus nilssol1i ni!ssoni, - H ,\T T, Mi sc . PubL Mus. Zo o1. Univ. Michi ga n, DO 106: llo hlO9S10l1i • ana1 0hcus 31; specimen from Baghdad [HARR ISON 1972: 627]. ===> o ognev. , SOO I(m 1963 Eptesicus innesi, - H AR RI SON, Z. Sa ugetierkde., 28 (2): 11 0; hingstoni a sy nonym.

Material : See N ADER & KOCK (1983). himself were id entified as a [first] record of Eptesicus n. nilssoni (KEYSERLlNG & Adana, SVlIL1976, 1() 19 subad. (2 skulls, 2 BLASIUS 18 39) for Iraq (from Baghdad and another undocumented locality) and al so KUMER LOEVE 1982) . incidentally for the whole Arabian peninsula. However, HARRISON (1964) did not Mea sur e men t s: See Tab. 1. Include E. nilssoni as a member of the Arabian bat fauna; als o he did not comment The range of skull measurements now upon such a di stributionally exceptional record. Only later the Baghdad specimen approach to and overlap with E. sodalis Bp of H ATT (1959) was examined by HARRISON (1972: 627) and its identification of upper toOthrow (BAUER 1968: tab. 1). corrected as E. bottae. remains to be studied more closely, de~ E. nilssoni was also li sted for Iraq by AL ROBAA E(1966), based on HATT (1959), not noting soda/is in E. serotinus. The occurrence of anatoliCus in Turl however th e deletion by HARRISON (1964). This species is in cluded in the chec k-l ist of MAl-WI & GEO RG (1969) tOo. A more modern lis tin g of Iraqi mam mals (KADH IM et al. 1977) (1983), for N-Iraq by HARRISON (1964: 2 does not mention E nilssoni among th e 15 bat species (14 named) of the country, apparently SW-Iran by DEBLASE (1980). Iraqi record! being aware of HARRISON 'S (1972) correction. However, recently H.'NAK & HORACEK (1987) ar e based o n HARRISON (19:>6) which 1; referred to HATT 'S (1959) reco rd aga in in establishing rhe sourhern limit of distribution for RISON 1964) . E. niLssoni. A r ecord from Sulelmama (FELTEN 1971), being outside the Iraqi lowlands, raised the question whether the Iraqi population assembled by FELTEN (1971) in measurement data of hingstoni, might in reality comprise further larger sized b. Eptes icus (Eptesicus) botta anatolicus. The measurements of the Suleimania series show better agreement with 1976 Eptesicus bo ttae omanensis H ARR ISO N, I lowland hingstom (FA 3c/c/ 44-45, 599 46-47' 5; greatest length of skull Jeb el al Akhdar, 1830 m, Oman. [unsexed]: 16'6-17'4; condylobasal length 16'4-17'0; zygomatIc breadth 11'1-11 '4; upper toothrow 5'9-6'2; lower toothrow 6'6-6'8) than with anatolicus Measur e m e nts: See Tab. 1. (see Tab. 1). Three specimens of omanensis are I 1 Birkat Sharaf als Wadi Sahtan, in t Ep tesicus (Eptesicus) bottae anatolicus FELTEN 1971. HARRISON 1976, 1977, 1983).

1905 Vesperlilio sp., near V serOlinus, - THOMAS, Proc. zooJ. Soc., 1905 (2): 520; sp ecimens fro m Mala-i-Mir [= Izeh] and Telespid, Khuzi stan , [ran [s ee D EBLA SE 1980: 192]. 1919 E [p lesicusj. serotinus luyeomanu5, - THOMA S, J. Bombay nat. Hist. Soc., 26 (3): 745; BM specimens from (Izeh, Tel es pid and Basht] [ran [se e HARRISON 1964: 143, D EBLAS E Eptesio/s (Eptesicus) botta 1980: 194]. 1921 Eplesicus mirza, - CHEESMAN, J. Bombay nat. Hist. Soc., 27 (3): 574; specimens from Epcesicus bOllae taftanimonlis DE F 1988 Telespid, Iran [THOMAS 1905: 520] and from Bas ht, Khuzistan, Iran [see DEBLA SE 1980: Ku sheh, W-side of Kuh -eTaftan, lr 192, 194]. Eptesicus bottae taflahimontis [sic\], 1988 1956 Eplesicus serolinus luYCOmant/s, - HARRISON, J. Mammal., 37 (2): 262; specimens from See Tab. 1. Ser'Amadia and Kanl Mase, Kurdistan, Iraq [see HARRISON 1964: 139 , 143] and BM l\lcasurem ent s : speci mens from Iran [THOMAS 1919]; [see DEBLASE 1980: 194]. Measuremencs given for the recer 1964 Eptesicus boltae subsp. indet. , HARRI SO N, Mamm. Arabia, 1: 139; spec im ens of t he range of £. b. ognevt (see DE Roc H ARRI SON (19 56: 139) and [fid e HARRISON 1964: 143, HARRTSON 1976: 417, DEBLA SE et al. 1978: tab. 9). However, the h; 1980] of THO MAS (1919). ured 1967 Eptesicus boltae, - LAY, fieldiana Zool. , 54: 233; sp eci mens from Izeh, Telespid and being described as not bicolo , ' Bas ht [see DEBLASE 1980: 194]. stated by GAISLER (1 970), HAN,\K 1970 Eptesicus serotinus schi1"i12iensis [sic I], - GA ISLER, Acta Sci. nat. Brno, 4 (6) : 30, tab. (1987). 18; 32, tab. 19; specimens from Basht, Telespid and Mala-i-Mir, SW-[ran [see DEBLASE While the size of ta/tanimontis 1980: 202]' range, further characters like sh ap 1971 Eptesicus anatolicus FELTEN, Senckenbergiana bioI., 52 (6): 371; loc. typ.: Alanya, coast reduction of the third commissur, of S-Anatolia, ViI. Antalya, Turkey. identity w ith bottae (see HANAK I 1976 Eplestcus botlae analOlicus, - HARRISON , Mammalia, 39 (3): 417; n. comb. E. b. ta/tanimontis is present 1982 Eptesicus analolicus, - KUM ERLOEV E, Sauget ierkdl. Mitt., 30 (1): 27. ROGulN 1988; Map 1). The above synonymy is extended from a previous one given by NADER & KO CK (19 83). 7

~cord of Eptesicus 'I . ld and h n. nz ssonz (KnsERLlNG & anot Mat e r"l: See N ADER & KOCK (19 83 ). - Turke y : Toprakkala, nr. Ceyhan , ViI. :ninsula er undocumented locality) and Adana, SVII1.l976, 10- 19 subad. (2 skulls, 2 skins) , leg. & Coil . H. VIERHAUS 911 -2 (see A. b' . Hb owever, HARRISON (1964) did also KUM ERLOEVE 1982) . r a Ian at fauna; al so he d'd not Mea su r erne n [ s : See Ta b. 1. lal record Only I h 1 not comment . ater t e Baghd d . ARRISON (1972: 627) d' . a specImen The range of skull measurements now available for thi s subspecies shows certain an Its ldentlfIcation approach to and overlap with E. sodalis BARRETT-HAMILTON 1910, except for length of upper tooth row (BAUER 1968 : tab. 1). The relationship between these two taxa ROBAAE (1966), based on HATT (19 . remains to be studied more closely, despite HILL & H ARRISON (1987) Included ). This species is included 59), not noting rn listing of Ira . In the ch eck·IIst of so daLls in E. serotinus. Jat species (14 n;mm~)mmfalhs (KADHIM et.1. 1977) The occurrence of anatolicus in Turkey was summ arized by N ADER & KOCK e 0 [ e cOuntry I n. HOwever recently H' ' apparent y (1983), for N-Iraq by H ARRISON (1964: as hingstoni) and DEBLA SE (1980), and for .' ANA K&H 'c rabl,shing th e so uth j"' f ORA 'EK (1987) SW-Iran by DEBLA SE (1980). Iraqi records of E. serotinus mapped by OSBORN (1963) ern Imlt 0 distriburion for are based on H ARRI SON (1956) whicrl later were reid entified as anatolicus (HAR RISON 1964). 1971), being ou tside the I . lopulation assembled b F raql lowlands, . Y ELTEN (1971) . n realIty comprise fu th I . In '. r er arger SIzed b ~ Im an la series sh b . 599 46-47' . ow etter agreemem with Eptesicus (Eptesicus) bottae omanensis HARRI SO N 1976. 5, greatest length of sk 11 1engt h 16'4-17'0. . u 1976 Eptesicus bonae omanemis HARRI SON , Mammalia, 39 (3): 415; loc. typ.: Masj id Ma'illah, ~r toothrow 6'6-6'8) t¥gomathlc breadth an W It analOizcus Jebel al Akhdar, 1830 m, Oman. Measurements: See Tab. 1. Three specime ns of omanensis are presently known from the type locality and 'e anatolicus FELTEN 1971. Birkat Sharaf als Wadi Sahtan, in the Jebel al Akhdar of N-Oman (Map 1; lAS, Proc. zooL Soc., 1905 (2). '-, . HARRISON 1976, 1977, 1983). Khuzistan r . ~20, SpeCim ens J ] , ran [se e D£BLASE 1980 : 192] MAS , . Bombay nat. Hist 5 . :ashtJ Iran [s ee HARRI SON 1·96~C.;/36D(3 )B: 745; . , E LA SE , nat. Hist. Soc., 27 (3)' 5. . Eptesicus (Eptesicus) bottae taftanimontis DE ROGUIN 1988. '01 Basht Kh ' . 74, speCImens from , UZ lstan, Iran [see DEBlAS 1980: E 1988 Eptesicus bonae ta/Lanimontis DE ROGUIN , Re v. sui sse ZooL , 9S (2 ): 598; loc. typ.: )N,]. Mamm al., 37 (2)· 26. . Kusheh, W-side of Kuh-e-Taftan, Iranian Sis tan and Baluchistan. 'ag [H . 2, specImens from 1988 Eptesicus bottae ta/tahimontis [sic!], - SE XSON & PUR DY, Wildlife Review, 211: 60, 630. > see ARI\ISON 1964: 139. 143] a d B , DEBlASE 1980: 194]. . n M Mea s ur e m e n t s: See Tab. 1. ~, Mamm. Arabia 1· 139 . , 1964· 143 H ,. ; specimens of . , ARRIS ON 1976: 417, DEBLA SE Measurements given for the recendy de sc ribed ta/tanimontis seem to fall within the range of E. b. ognevi (see DE ROGUI~.; 1988, tab.: 599, compared with STRELKOV :4: 233; spec imens from Izeh "r J 'd ' Ie espl and et at 1978: tab. 9). However, the hair of tafianimontls is an interesting character, being described as not bicoloured, an exception of the specific hair colourati on as AISlER, Acta Sci. nat B llO >pid and Mala-i-Mir swrr , 4 (6): 30, tab. stated by GAI SL ER (1970), HANAK & GAISLER (1971) and HANAK & HORA CEK , - ran [see DEBlA s£ (1987). a bioi., 52 (6): 371; loc. typ.: AJanya, coast While the size of tafianimontis agrees with E. bonae in that part of the species l 2 range, further characters like shape and proportions of r . and PM 2, as well as lam malia, 39 (3): 417· b J . kd ' n. com reduction of the third commissure of M need to be studied to confirm sp ecies tIer I. Mi tt., 30 (1): 27. . identity with bOtlae (see H ANAK & HOR AC EK 1987). ous one given b), N E. b. ta/tanimontis is presently restricted to the Kuh-e-Taftan, SE-Iran (DE ADER & KocK. (1983). RO GU IN 1988 ; Map 1). 8

Eptesicus (Eptesicus) bottae ognevi BOBR IN SKI I 1918. sic !: 22) in the bat fauna of E-Afghan paper of MEYER-OEHME (1965). GAISLE 18 65 V(espertilioj. (U)'comanus, - DE FILIP PI, N ote viaggio in Persia nel ] 862,: 342; speci me n HA NAK & GAI SLER (1971) considered MZST 903 fro m Zanjan, N W-lran [see D EBLASE ]980: ]92 , 196, 20], 205]. N IET HA MMER (1983) Kabul appears as a ]918 EptesiCtls ognevi BO BRINS KI I, [Fauna & Flora of Russia], 15: ]2; loc. typ. : Sokhta (1982) had included E. (as Chinar, G issaro-Darvaz, W-Tadjikistan [from KUZYAK IN 1944, 1950, bu t modern isabellinus geographic names]. Afghanistan for their zoogeographical 1925 Vespertilia (EptesiCtls) soda/is ogn evi, - BOBRI NSK ll, Bull. Mosk. O bstsh. Ispit. Prirod., O EHME'S (1965) specimen in question 34: 35 4; n. stat. (1971) as E. niLssoni kashgaTicus BOB. 1927 Eptesicus sodahs ognevi, - OC NE V, J. Mamm., 8: 140-157; n. stat. BOBRINSKlJ 1926, vide HILL & H ARR IS( 1942 Eptesicus (Eptesicus) soda lis ognevi, - TATE, Bull. Ame r. Mus . nat. H ist. , 40 (7): 275 , 277 . inform ation w accept the occurrence 194 4 Vespertilio ognev i, - KUZYAKIN, Oprede l. Ml ekopit. ,: 96, 103; n. comb. E. b. ognevi was recorded for Pak 1951 Eplesicui sodalis ognevi, - ELLE RM AN & MORRI SON-S COTT, Chec klist Palaearct. India n Aswr, Azad Kashmir, based on a BM s Mamm.,: 156; te rr. typ.: Bokhara D istr., Turkestan [nee Buk hara, Uzbekistan]. it from his later check-list (SIDDIQI l' 1956 Eptesicus serotinus, - MI SONNE, Mem . Inst. r. Sci. nat., (2) 59: 26 [partim, literature specimen (Shenkagarh, Astor Distr., G reco rd for Zanjan, N W-I ran]. 1963 Eptesicus (Eptesicus) ognevi, - GROMOV et aI., Mle kopit. Fau na SS SR. , 20] . which is more likely w be referred t( 1967 Eptesicus seratinlls, - LAY, Fiel diana Zool. , 54: 145 [parti m, li te rat ure record for Zanjan, The occurrence of E. soda/is ognevi NW-Iran]. BOBRIN SKII (1931), based on one speci 1971 Eptcsicus bOllae ognevi, - HAN ,\K & G AISLER, Vest. ces koslov. Spol. zoo!., 35 (1): 20; one from SW- Mongolia (K VZYAKI N 1 n. com b. although repeate d in literature (e. g. 1972 Epresicus ognev [s ic !], - GR OM OV & j ANuS HEVIC H, Mlekopit. Ki rg isi i,: 88. JANVSH EVICH 1972). BA NN IKOv (1954: 7 (1968: 25, sub E. sodalis) identify this The above synonymy may neither be complete nor correct, due to our poor Chinese terriwry (Transaltai Gobi, Air command of Russ ian and difficulty of access to relevant literature published in this occurrence of E. s. ognevi in NW-M language. omiting the reference!), but do not inc M ateria l: See N ADER & KO CK (1983). - U SS R: Gobustan, 60 km S Baku , Azerbaid (: 112). SOKOLOV & aRLOV (1980: 67) zh an SSR, 20. & 21Vl.I979, 10" 2 99 (3 skulls, 3 skins), SM F 73493-5, leg. v. H ANAK . - ognevi in Mongolia, but give no expl Osh region, Kirgizian SSR, VII .-VIII.1 988 , 9 (skull, ale.), SMF 73942, vend. H . BURDA. In summary, the wtal distribution M e a su r e men t s : See Ta b. 1; this gi ves a sel ect ion of larger mate ri al as cited, other dat a, time, characterizes a bat which occurs e.g. from Syr Darya area give FA 46'5 (BO BRINS KII 1931). is not recorded from within the Sah. E. b. ognevl is the third subspeci es occurring in Iran. Records from Iran ian Mediterranean or mountainous habit; Azerbaidzhan and Khorassa n we re reviewed by DEBL ASE (1980). T he di stribution B i 0 log y : Hardly any informati of ognevi in the southern regions of the USSR (Map 1) is based on BOBRINSKIJ available. Eight 9 9 from Baghdad we et a1. (1965) with addition of the above materi al from Azerbaidzhan; on STRELKOV were found in Iraq in mid-April (AL R & SHAJ-MARDANOv (1983) for Kazachstan wi th additions given by NAD ER & KOCK Yotva ta / Palestine, collected on 22. Apr (1983); on GROMOV & ]ANV SHEVI CH (1972) fo r Kirgizia with addition of ab ove (HARR ISON 1964). One of two 99 material ; on ST RELKOV et al. (1978) for Turkmenia with an addition given by Baku l Azerbaidzhan had also twin en H ARR ISON (1976: Meshed-M esrian). the six specimens from Kusheh/SE-Ir; The records of this subspec ies in Afghanistan need a thorough rev iew: ROGVIN 1988) and also one of d KV ZYAKIN (1944,1950) in desc rib ing the range of ognevi in Soviet Central Asia lists Ceyhan/Turkey. At Al Mahm udiyah a locality "Feisabad" in upper Bokhara (= Tadjikistan), but hi s maps show a April and 5. May 1965 and again on 17 locality in N -Afghan istan, which co ul d be identified as Fai sa bad, Badakhshan animals were collected: 19 carrying t Prov./N E-Afghanistan, as was done by AELLE N (1959). This locality beca me an yo ungs and 1 9 carrying one young a acc epted Afghan record by GAISLER (1971) and N IET HAMME R (1983), but not by length of the 40'0' from SW-Saudi Al N ERONOV & ARSE NIEVA (1982) an d wa s co nsidered as "more or less uncertain" by length was about 5 mm. - Body wei H ANAK & GATSLE R (1971: 18). - ViEYER-OEHM E (1965) collected an Eptesicus sp. g and that of the holotype of tafiam in Kabul, which had smaller forearm length than b. ognevi and which the author It is noteworthy that all four specir left unidentified, although he cites a somewhat Jarger forearm length of an E. collected on four different dates are C isabellinus bottae fr om Kas hmir. KU LLMAN N (1970) included E. bottae (= ogne'W l, old buildung betwee n 9-11 p.m., evil 9

ognevi BOB RI NSK JI 1918. sic!: 22) In the bat fauna of E-Afghani stan; the basis for this could only be the paper of MEY ER-OEH ME (1965). Gt\I SLE R (1971) li sted Kabul with a question mark, ote viaggio in Persia ne l 1862,: 342; specimen HANAK & G AISL ER (1971) considered this as " more or less uncertain", but in )EB LAsE 1980: 192 , 196 , 201 , 205 ). Flora of Russia], 15: 12; loe. typ. : So kh ta N IETHAMMER (198 3) Kabul appears as an accepted locality. NERO NOV & ARS EN JEVA [fro m KU ZYAKIN 1944, 1950, but modern (1982) had included E. Isa bellinu5 (as a species separate from E. serotznus!) in E Afghanistan for therr zoogeographical analysIs of t he country's bat fauna. ME YER IB RINSK II, Bu ll. Mosk . Obstsh . lspit. Prirod. , OEHME 'S (1965) specimen in questi o n from Kabul had been identified by FELTEN (1971) as E. nilssonl kashgaricus BOBRI NS KII 1926 (= synonym of E. gobiensls Imm. , 8: 140-157; n. stat. BOBRINSKII 1926, vide HILL & HARRISON 1987). We did not find any other reliable Bull. Am er. Mus . nat. H ist. , 40 (7): 275 , 277. information to accept the occurrence of E. bottae in Afghanistan . . Mlekopit., : 96, 103; n. comb. E. b. ognev i was recorded for Pakistan by SIDDIQI (1961) from Shenkagarh, ;l OR RISO N-SCOTT, Checklist Pala earct. Indian Asto r, Azad Kashmir, based on a BM specimen (E. isabellinus bottae), but omitted Turkesta n [n ee Bukhara , Uzbek istan). st. r. Sci. nat. , (2) 59: 26 [part im , li terature it from his later check-list (SI DD IQI 1969). ROBERTS (1 977) r efe rs to an identical specimen (Shenkagarh, Astor D istr., Gilgit, 3200 m, labelled E. isabellznus bottae), tal., Ml ekopit. Fauna SSS R.,: 20 1. which IS more li ke ly to be refe rred to E. bottae ognevi. 54 : 145 [part im, literature re cord for Zanjan , The occurrence of E. sodalis ognevl in regions further north was mentioned by BOBRINSKJI (1931), based on one specimen from N W- Mongolia. This record and ,L ER, Vest. ceskoslov. Spol. zo o!. , 35 (1): 20; one from SW-Mongolia (KUZYAK IN 1944: Eczin) have not been mapped here, although repeated in literature (e.g. HA NAK & GA IS LE R 1971, G ROMOV & USHEVIC H, Mlekopit. Kirg is il,: 88 . JA NUS HEVICH 1972). BANN IKOV (1954: 77, sub V ognevi) and STUBBE & C HOTOLCHU complete nor correct, due to our poor (1968: 25, sub E. sodalis) identify this locality as Edsin-gol, or Echin-gol Oasis, in Chinese territory (Transaltai Gobi, Aimak S-Gobi). The la tter authors also cite the :s to relevant literature published rn this occurrence of E. s. ognevl in N W-Mongol ia from BO BRI NS KII (1931) (however omiting the r eference!), but do not include the spec ies in their Mongolia check-list j S S R: Gobustan , 60 k m 5 Baku , Azerbaid (: 112). SOKOLOV & ORLOV (1980: 67) would not exclude the occurrence of E. b. ;, 3 skin s), SMF 73493- 5, leg. V. H ANA K. ognevi in Mongolia, but give no explanation. (skull, ale.) , SMF 73942, vend . H. BURD A. In summary, the total di stribution of the species as outlined here for the first lect ion of la rger material as cited , other dat a, time, characterizes a bat which occur:; in the south of the Palaearctic Region, but 15 KII 193 1). is not recorded from within the Saharo-Sindian Subregion except marginally in cUfring in Iran . Records from Iranian Mediterranean or mountaino us habitats. d by D EBL AsE (1980) . T he distribution B i 0 log y: Hardly any information on the reproduction of this species is USSR (Map 1) is based on BOB RINSKII available. Eight Q Q from Baghdad we re inseminated on 26. Dec. and gravid Q Q te rial from Azerbaidzhan; on STRE LKOV were found in Iraq in mid-April (AL RO BAAE 1966: sub E. serotinus). One Q from vith additions given by N ADER & KoCK Yotvata/Pales tine, collected on 22. A pri l had two embryos measuring about 14 mm 2) for Kirgizia with addition of above (HARRISON 1964). One of two Q Q collected on 20. and 21. June south of I'urkmenia with an add iti on give n by Baku/Azerbaidzhan had also twin em bryos, the other had one embryo. Two of the six specimens from Kusheh/SE- Iran co llected on 14. June were subadults (DE Jghanistan need a thorough review: RO GUIN 1988) and also one of the two collected on 5. Aug. from near Ige of ognevl In SOVIet Central Asia lists Ceyhan/Turkey. At Al MahrnudiyahlIraq preg nant Q Q were collected on 28. := Tadjikistan), but hi s maps show a April and 5. May 1965 and again on 17. Ap ril and 15. May 1968. Also, the following )e identified as Faisabad, Badakhshan animals were collected: 1 Q carrying two youngs on 28. pril; 4 Q Q carrying two EL LEN (1959). This locality became an youngs and 1 Q carrying one young and 2 Q Q were lactating on 5. May. - Testes ) and N IETHAM MER (1983), but not by length of the 4u u from SW-Saudi Arabia ranged from 4-5 mm and their p hallar Is idered as "more or Jess u ncertain" by length was abou t 5 mm. -Body weight of these specimes ranged between 7-10 ) EHME (1965) collected an Eptesicus sp. g and that of the holotype of taJtanimontis was 14 g. h than b. ognevl and which the author It is noteworthy that all four specimens from Misfera/Saudi Arabia which were ew hat larger forearm length of an E. collected on four different dates are U U , but were taken at ni ght from the same N (1970) lI1cluded E. bottae (= ognew i, old buJldung between 9-11 p.m., evidently upon their return from their feeding 10 excursions. A search by the senior author fo r their day roost was unsuccessfu l. G AISLER , J. (1970): The bats (ChiroF HARRISON (1956) records roosting si te s in Iraq as under corru gated iron roofi ng. expeditions of 1965-1967 . . No ectoparasites were o bserved o n the specimens examined. Although losses __ _ (1971): Systematic review a after capture, by preparation and transport may have been effective, this may be hitherto recorded from Af! due to specific ro osting sites and habits of a hos t speci es which is unfavourable for GROMOV, I. M. & JANU SHEVI CH , A. J the development of som e ectoparasites , e.g. fleas and barfl ies. (lsdatel. " Him" ) [in Ru ss iaI HANAK, V. & GAISLER, ]. (197 1): T remarks on some other sper Zu s amm e nfas s un g. spol. zoo1., 3S (1): 11-24; : Eptesiws bOllae wird erst mals fur Sau di Arabien belegt. Daren fur die N ominatform sind HANAK, v. & HORACEK, I. (1987): die ersten Erganzungen seit ihrer Besc hreibung. Weitere Ma{l,angaben und Bemerkungen, (Chiroptera: Vesperti lionic ein schlie{l,]i ch Syno ny mie-Li sten, zu den bisher bekannten Unterarren mneSI , hingstoni, 377-38 8; Wi en [for 1984/8 anatolicus, cafian im ontis und ognevi werden ge geben und di e Gesamtverbreitung kart iert. H ARR ISON, D. L. (1956): Notes on sc Daten zum Fo rtpflanzu ngsgesc he he n werden mitgete ilt. Beirr., 7 (1/3): 1-7 ; Bo nn. (1963): A new bat for Israe on the affi ni ties of the sper R efe r e n ces. Berlin . (1964): The mam mals of AI AUL EN, V. (1959): Contribution a l'etude de la faune d'A fghanistan . 9. C hriopceres. - Rev. pp.; London (E. Benn). suisse Zool., 66 (21): 353- 386; Genc ve. (1972): The mammals of j AL RO BA AE, KH. (1 966) : U ntersuchungen der Lebensweise irakisc her FledeI-mause. - pp.; Lon do n (E. Ben n). Sauge ti erkdl. Mitt., 14 (3): 177-211 ; Mti nc hen. (1976): Scientific results of BANNI KOv, A. G. (1954): Mlekopi tajus hchiche Mon go lskoi narodnnoje respu bli ki [Mam mal s of a new subspecies of Bolt of thc Mongolian Peoples ' Republic). - 649 pp.; Moskva (Isdatel. Akad. Nauk Vespertilionid ae) from Orr SSSR) [in Ru ss ian] BATES, P. J. J. & H ARR ISON , D . L. (1989 ): New records of small mammals from Jordan. (1977): Ma mmals obtained Bonn. zool. Beirr., 40 (3 / 4): 223 -226 ; Bonn. the Sultanate of Oman. BAUER, K. (1968): Ein dritter Fund von Eptesicus sodalis (Chi roptera, Ves perrilionida e). (1983): The mammal faun ; Ann. naturhist. Mu s. \XIi en, 72 : 29-32; \XIie n. the Oman Flora and Faun BO BRINSKII, N. A. (1925) : [M aterials on che fauna of bats of Turkestan (Wescern Tur kestan HATT, R. T. (1959 ): The mammals 0 wit h the regions of Semiretchie and of Transkas pia , Khiva and Bucha I' a). - Bull. 106: 1-113; Ann Arbor. Mosk . Obm h. Ispit. Prirod., 34: 330-374; Moskva [in Ru ssia n). HI LL, J. E . & HA RR ISON, D. L. (1 91 - -- (1931): N eue Angaben tiber die geographisc he Verbreitung der Fledermause Vespenili on id ae) wi th a sy (Chi roptera) Rulllands. - Zool . Anzeiger, 96 (9/ 10) : 265 - 268; Le ipzig. and the description of ne (Zool.), 52 (7): 225- 305; I BOBR IN SK II, N. A., KU SNE ZO V, B. A . & KUZjA KI N, A. P. (1 96 5): O predelitelje ml ekopi taju shchich SS SR. - 384 pp.; Moskva (Isdatel. Prosves henje) [i n Ru ssi an ). HILLENIUS, D. & GASPERETTI, J. (19 Arabia, 6: 513 - 527; Jedda BOTTA, P. E. (1 841): Relaci on d'un voyage dans Ie Yemen entrepris en 1837. - 148 pp ; Paris (B. Duprat) [no vid.). KADHIM, A . H ., N ADAC HOVSK I, A. know led ge of Iraqi mamn DEBLASE, A. F. (1980): The bats of Iran : sys te macics, di stributi on, ecol ogy. - Field ia na Zool. , (NS) 4: i-xvii, 1-424; Chicago. KUL LMANN, E. (1970): Die Tierwelt - Freunde Kaln . Zoo, D OLLM i\N , J. G. (1933): Mammals of the Rub' 31 Kh al i, 39 4. - In : PHILB¥, H. ST. J. B.: 1: The empt), quarter, bein g a description of t he great south dese rt of Arabia known KUMERLOEVE, H. (1982): Zur Erfor as Rub' al Khali. - xxi v + 433 pp.; Lond on (Constable & Co. ). Saugetierkdl. Mitt., 30 (1 ELLER MAN, J. R. & MORRI SON-SCOTT, T. C. S. (1951): Chec kli st of Palaearctic and Indian KU ZYAKIN, A. P. (1944): Ordo C hin mammals 175 8- 1946. - 810 pp. ; Lon don (Trus!. Bric. Mus. N~t. 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TH OMAS, O. (19 05): O n a collection c MA HDI, N . & G EORG , P. V. (1969): A systemat ic list of the vertebrates of Iraq. - Iraq nat. British Museu m by Col. A H ist. Mus. Pu bl., no. 26: i-iv, 1- 104; Baghdad. Lo ndon. MEINERTZHAGEN, R. (1954): Birds of Arabia. - xiii + 62 4 pp.; Edinburgh -London (Oliver _ __ (J 919):Somenew mammal & Boyd ). 745- 749 ; Bombay. MEYER-OEHME , D. ( 1965): Die Sauget ie re Afghanisrans. Teil III: Chiro ptera. - Science, 1965: WA SS IF, K. (1962): The serotine bat, E 42-58; Kab ul. zooL Soc. Egy pt, 17 : 108 NADER, I. A. (1969): Animal re mai ns in pellets of the barn ow l, fro m the vicinity Tyto alba, \XiA SS IF, K., MADKOUR, G. & SO LIMA N of An-N ajaf, Iraq . - Bull. Iraq nat. Hist. Mus., 4 (1): 1-7; Ba ghdad. of bats from Egy pt. - 36 NADE R, I. A. & KOC K, D. (1983): Notes on some bats fro m the Near East (Mammalia : Egypt) . Chiroptera). - Z. Sau getierkde., 48 (1): 1-9; H amburg-Berlin. WETTSTE IN , O. VO N (1913): Wi sse nsc h: NE RONOV, V. M. & ARSENI EVA, L. P. (1982): Regio nal peculi arities of the fa una of bat s of 1910. Die Ch iropterenaust Afghanista n. - Zool. Zh., 61 (4) : 585- 592; Moskva [in Ru ssi an]. 465-471; Wien . NIET HAMM ER, j. (1983): Die Sa uge tiere Afghan istans. - Mitt. dtsch. Orient-Inst., no. 22: 211 - 228; Hamburg . OSBORN , D. j. (1963): N ew di stributional records of bats fro m Turkey. - Mam mal ia, 27 (2): 21 0-217; Paris. PETERS , W. (1869): Bemerkungen liber neue oder wenige r bekannte Fl edenhiere, bes onders des Pariser Mu se ums. - Monatsbe r. kgl. preuss. Akad. Wiss ., 1869 : 391 - 406; Authors: Dr. IY AD A. NA DER, Nation Berlin . mel1t (NCWC D), P.O.Box 61681 , Forschungs institut Senckenberg, Sen PO COC K, R. I. (1935): The mammals coll ected in S. E. Arabia by Mr. BERTR AM THO MAS and Mr. H . St. J. PHIL BY. - An n. Mag. nat. Hist., (10) 15 : 441-467; London. QUM SIYEH, M. B. (1985): The bats of Egypt. - Spec. Publ. Mu s. Texas Tec h. Univ., no. 23: 1-102; Lubbock ROB ERTS, T. J. (19 77): The mamm als of Paki stan. - xx vi + 361 pp.; London-Tonbridge (E. Benn). ROD E, P. (1941): Catalogue des types de ma mmiferes du Museum National d'H istoire N aru rell e. Ordre des chiropteres . - Bull. Mus. natn. Hist. nat. , (2 ) 13 (4) 227- 252; Paris. ROGUIN, L. DE (1988) : N otes su r quelques mammiferes du Baluchistan iran ien. - Rev. suisse Zoo l , 95 (2) 595-606; Geneve. SID DIQI, M. S. U. (196 1): Checklist of mammals of Pakistan with pa rti cular reference to the mammalian coll ection in the Bri tish Mus eum (Natul'al History), Lon don. Bi ologia, 7 (1 /2) : 93 -225; Lahore. - - - (1969): Faun a of Paki sta n. - 160 pp.; Karach i (Agric. Res. Co uncil) . SOKOLOV, V. E. & ORLOV, V. N . (1980) : [Key to the mammals of the Mongol ian People's Republic.] - 352 pp.; Moskva (Ak ad. Nauk) [in Russ ian]. SP IT ZENBE RGER, F. (1979): Die Saugetierfauna Zyperns. Teil II : Chi roptera, Lago morpha, Carn ivora and Artiodac tyla. - Ann. naturhis t. Mus. Wien, 83: 43 9- 465; \Vien . STREL KOV, P. P. & SHAJM AR DANOV, R. T. (1983): [The new da ta on tfle di st ribution of bats (Chiroptera) in Kazak hstan]. - Fau na, sys temat ik a i biologij a mlekopitajushch ic h. Trudi zo ol. Inst., Akad. Nau k SSSR, 119: 3-37; Leningrad (Zool. lnst. Akad . N auk SSSR) [i n Russian ]. STRELKOV, P. P., SOSNOVT ZE VA, V. P & BABAEV, H. B. (1978): [Bats (Chiroptera) of Turkmen ia. ] - Funct ion al morphology and systematic of mammals. Trudi zoo l. Inst. , Akad. Nauk SSSR, 79: 3-71; Leningrad [in Russian]. ST UBB E, M. & CH OTO LCH lJ, N . (1968): Zur Saugetie rfa una der Mongo lei. - Mitt. zool Mus. Berl. , 44 (1): 5-1 2 1; Berl in. 13

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