Insects in the Killbuck Marsh Wildlife Area: 1993 Survey1

R. N. WILLIAMS, M. S. ELLIS, AND D. S. FICKLE, Ohio Agricultural Research and Development Center, Ohio State University, 1680 Madison Avenue, Wooster, OH 44691

ABSTRACT. The Killbuck Marsh Wildlife Area was the focus of a seven month survey performed in 1993 to determine the diversity of selected . Primary emphasis was focused on three families of Coleoptera: ground , including tiger beetles (Carabidae); sap beetles (Nitidulidae); and carrion beetles (Silphidae). Rare or endangered within these families were of particular interest and constant vigilance was made to detect them. Five collection methods were used at five sites within the Killbuck Marsh. These included: ultraviolet (black light) traps, flight interception (window) traps, bait traps, carrion bait sampling, and aerial and aquatic sweep netting. In all, 68 ground , 30 , and seven carrion beetle species were identified. In addition to these families, beetles from 47 other families (372 species) of Coleoptera were collected and identified. Aside from Coleoptera, several dragonflies and damselflies (Odonata), caddisflies (Trichoptera), butterflies and moths (Lepidoptera), and mosquitoes and midges (Diptera) were also taken. Six species considered uncommon were encountered: Agonum cupripenne (Say), Agonum galvestonicum Casey, Chlaenius niger Randall, Oo'des americanum Dejean, discus (F.), and Stenocrepis cuprea (Chaudoir). One hister beetle (Histeridae), Anapleus marginatus LeConte, was also very uncommon for this area.

OHIO J. SCI. 95 (3): 226-232, 1995

INTRODUCTION MATERIALS AND METHODS The Killbuck Marsh Wildlife Area, the second largest Five sites representing different habitats were chosen wetland remaining in Ohio, was the focus of a seven in order to capture a broad range of insects. Site one was month survey in 1993 to determine its fauna. This a fairly isolated, narrow, vegetated island within the is the first survey of this magnitude of the Killbuck Marsh marsh, and it yielded most of the aquatic insects such as which basically lists all of the species we could get iden- predacious diving beetles (), water scavenger tified from several trapping methods. Such benchmark beetles (), and crawling water beetles information is necessary in order to measure/compare (Haliplidae). Site two was located on a bank facing the biodiversity in the future. Should this habitat come under marsh with access to woods and farmland. Site three was threat by natural or human conditions, this survey will a narrow tree line surrounded by grassland and open serve as a basis to assess the impact these conditions fields. A low-lying area on the Killbuck Creek's flood- might have on the local insect fauna. plain served as site four. Finally, site five was a dense Additional projects associated with the Killbuck upland woodlot adjacent to the marsh approximately Marsh Wildlife Area have recently been conducted to 5 m above the level of the river. A detailed map of the examine the Killbuck Creek's water quality (Ohio En- Killbuck Marsh Wildlife Area and any additional infor- vironmental Protection Agency 1986, unpublished) and mation about this habitat may be obtained from the Area the degrees of organic pollution on benthic macro- Manager in Shreve, OH, at (216) 567-3390. invertebrates (Sommer 1994). Primary collection emphasis was focused on Coleoptera, The Ohio Department of Natural Resources (ODNR) especially ground/tiger beetles (Carabidae), sap beetles began purchasing land for the Killbuck Marsh Wild- (Nitidulidae), and carrion beetles (Silphidae). Many other life Area in 1969, and it now encompasses approxi- orders and families of insects were collected and, with mately 5,500 acres maintained by the ODNR Division the aid of specialists, determined to species. Groups of of Wildlife. The marsh is located in a U-shaped glacial particular interest in this wetland setting were the dragon- outwash valley about 6.5 km southwest of Wooster in flies and damselflies (Odonata), caddisflies (Trichoptera), Wayne and Holmes counties between State Routes mosquitoes (Diptera), various families of aquatic beetles 226 and 83. About 60% of this wetland is flooded (Coleoptera), and butterflies and moths (Lepidoptera). during some portion of the season and produces Five collection methods were utilized in the study to diverse hydrophytic vegetation. Some of the com- optimize the diversity of insects encountered. All traps ponents included are cattail {Typha latifolid), button- were set on 7 May 1993, collected weekly unless other- bush (Cephalanthus occidentalism, and arrowhead wise noted, and removed on 14 October 1993. Four {Sagittaria latifolid). general purpose "black light" survey traps (O. B. Enter- prises, Oregon, WI) (two AC and two DC) were placed, one each, at sites one, two, three, and five. All of the light trap entrances were situated approximately 1 m 'Manuscript received 21 October 1994 and in revised form 6 March above ground. The four light traps were run 48 hours 1995 (#94-23). continuously per week. OHIO JOURNAL OF SCIENCE R. N. WILLIAMS ET AL. 227

A flight intercept (window) trap similar to the design TABLE 1 used in Peck and Davies (1980) was used at sites one, two, four, and five. The window pane of each trap was 1993 Killbuck Marsh Wildlife Area insect list. about 0.7 m2 with troughs made of PVC pipe on each side of the pane. The bottom of each trap was at ground level and the PVC troughs were filled with soapy Species water. Four nitidulid inventory technique (NIT) traps (Williams et al. 1993) were utilized, two with whole ODONATA (6 Families; 24 Species) wheat bread dough and two with fermenting brown I Dragonflies sugar, at each of the five sites. Each bait was used at Aeschniclae: Aeschna mutata Hagen two levels, ground level and 1 m above ground level, at Aeschna canadensis Walker every site. Anax longipes Hagen In order to survey for carrion insects, unmarketable Anax junius Dairy Gomphidae: turkeys and road kills, mostly squirrels, were placed at Gomphus lividus Selys random around the five sites, secured with chicken Gomphus villosipes Selys wire, and checked every two weeks. Three multiple-plate Libellulidae: aquatic macroinvertebrate samplers (Hester and Dendy Celithemis elisa Hagen Celithemis eponina Drury 1962) were used to capture (riffle beetles). Sweep Erythemis simplicicollis Say nets were used to survey insects on vegetation, and Libellula luctuosa Burmeister aquatic sweep nets were used weekly at all five sites. Libellula pulchella Drury Libellula lydia Drury During weekly collection of all traps, insects were Pachydiplax longipennis Burmeister either placed directly in 70% ethanol (NIT traps and Perithemis tenera Say sweep net material) or frozen and then placed in Sympetrum vicinum Hagen ethanol (light and window trap specimens) to preserve Tramea lacerata Hagen II Damselflies them for later identification. Coenagrionidae: In addition to the present paper, we have produced Enallagma aspersum Hagen a short video on the history and public use of the Enallagma civile Hagen Ischnura posita Hagen Killbuck Wetlands entitled "An Introduction to The Ischnura verticalis Say Killbuck Marsh Wildife Area of Northeast Ohio" which Lestidae: is available through OSU Extension Publications at Lestes congener Hagen (614) 292-1607. Lestes forcipatus Rambur Lestes rectangularis Say Calopterygidae: RESULTS AND DISCUSSION Calopteryx maculata Beauvois The total number of insects includes nearly 500 TRICHOPTERA (5 Families; 20 Species) species in 85 families belonging to 10 orders (Table 1). Hydropsychidae: Netspinning Caddisflies Of the five trapping methods used (window trap, light Ceratopsyche slossonae (Banks) trap, NIT trap, carrion bait, and sweep net sampling), Cheumatopsyche campyla Ross Cheumatopsyche pettiti (Banks) the light traps yielded the greatest number of species Hydropsyche betteni Ross comprising an estimated 70% of the total number of Hydropsyche dicantha Ross species caught. Several ground beetles, water scavenger Hydropsyche rossiVM., Vos., Par. beetles, and variegated mud-loving beetles (Heteroceri- Leptoceridae: Longhorned Caddisflies Ceraclea maculata (Banks) dae) were collected in these traps in large numbers. Most Ceraclea tarsipunctata (Vorhies) of the butterflies and moths, caddisflies, and predacious Leptocerus americana (Banks) diving beetles were also taken in the light traps. Oecetis inconspicua (Walker) Triaenodes injustus Milne All sap beetles, with three exceptions, were caught Triaenodes tardus (Hagen) in the NIT traps baited with fermenting brown sugar or Nectopsyche sp. wheat dough. The three exceptions were Prometopia Limnephilidae: Northern Caddisflies sexmaculata (Say), which was collected only in the Limnephilus indivisus Walker Limnephilus submonilifer Walker light traps; Epuraea rufa (Say), which was collected Pycnopsyche lepida (Hagen) frequently in the light and window traps; and Omosita Phryganeidae: Large Caddisflies colon (L.), which was lured to carrion in large numbers. Agrypnia vestita (Walker) Phryganea sayi Milne Connell (1956) reported rarely collecting E. rufa in light Ptilostomis postica (Walker) traps in Delaware. One specimen of Prometopia bisig- Rhyacophilidae: nata was collected at a Mercury light on 4 March 1962 Rhyacophila ledra Ross (Gillogly 1969). P. sexmaculata is normally associated LEPIDOFTERA (9 Families; 24 Species) with sap flows, and was among the principal species I Butterflies found associated with oak wilt in West Virginia (True Danaidae: Milkweed Butterflies et al. I960). Several ground beetles were also collected Danaus plexippus plexippus L. in the NIT traps. Lycaenidae: Blues & Hairstreaks Lycaena hyllus Cramer The carrion bait (rodent and turkey carcasses) attracted Nymphalidae: Brushfooted Butterflies many carrion beetles, trox beetles (Trogidae), and one Limenitis arthemis arthemis Drury 228 INSECTS OF THE KILLBUCK MARSH VOL. 95

TABLE 1 (Continued) TABLE 1 (Continued)

Species Species

Nymphalls antiopa antiopa L. Bembidion rapidum (LeConte) Papilioniclae: Swallowtails Bembidion versicolor (LeConte) Papilio glaucus glaucus L. Blernus discus (F.) Papilio troihis troilus L. Bradycellus atrimedius (Say) Pieridae: Whites and Sulphurs Calathus gregarius (Say) Colias eurytheme Boisduval Chlaenius emarginatus Say Colias philodice philodice Godart Chlaenius impunctifrons Say Satyridae: Satyrs & Wood Nymphs Chlaenius lithophilus Say Enodia anthedon A. H. Clark Chlaenius nemoralis Say II Moths Chlaenius niger Randall Noctuidae: Owlet Moths Chlaenius pennsylvanicus Say Acronicta hasta Guenee Cicindela punctulata Olivier Agriopodes fallax (Herr.-Schaff.) Cicindela sexguttata F. Bellura obliqua (Walker) Clivina americana Dejean Lacanobia subjuncta (Grote & Rob.) Clivina bipustulata (F.) Leucania lapidercea (Grote) Clivina impressefrons LeConte Leucania multilinea Walker Colliuris pensylvanica (L.) Nedra ramosula (Guenee) Dyschirius erythrocerus LeConte Ochropleura plecta (L.) Dyschirius haemorrhoidalis (Dej.) Pseudaletia unipuncta (Haworth) Elaphropus xanthopus (Dejean) Simyra henrici (Grote) Elaphrus californicus Mannerheim Saturniidae: Silkworm & Royal Moths Elaphrus ruscarius Say Actias luna L. Harpalus caliginosus (F.) Automeris io F. Harpalus pensylvanicus (DeGeer) Eacles imperialis Drury analis Dejean Hyalophora cecropia L. Lebia atriventris Say Sphingidae: Sphinx Moths Lebia fuscata Dejean Amphion floridensis B. P. Clark Lebia grandis Hentz Lebia solea Hentz COLEOPTERA (50 Families; 372 Species) Lebia tricolor Say Alleculidae: Comb-clawed Beetles Lebia viridis Say Mycetochara fraterna (Say) Leptotrachelus dorsalis (F.) Anobiidae: Deathw. & Drug. Beetles Oodes americanus Dejean Stegobium panicium (L.) Paratachys proximus (Say) Anthicidae: Antlike Flower Beetles Platynus cincticollis Say Anthicus cervinus LaFerte-Senect. Platynus hypolithos (Say) Notuxus monodon F. Poecilus chalcites (Say) Notoxus murinipennis LeConte Poecilus lucublandus (Say) Sapintus fulvipes (Laf.) Pterostichus corvinus (Dejean) : False Powder. Beetles Pterostichus commutabilis (Motsc.) Lichenophanes bicornis (Weber) Pterostichus stygicus (Say) Xylobiops basilare (Say) Schizogenius lineolatus (Say) Bruchidae: Seed Beetles Stenocrepis cuprea (Chaudoir) Amblycerus robiniae (F.) Stenolophus fuliginosus Dejean Cantharidae: Soldier Beetles Stenolophus ochropezus (Say) Cantharis bilineatus Say Badister (s. str.) sp. Cantharis impressus LeConte Dyschirius sp. 1 Cbauliognathus pennsylvanicus DeG. Dyschirius sp. 2 Podabrus modestus (Say) Cerambycidae: Longhorned Beetles Podabrus rugosulus LeConte Anelaphus villosus F. Carabidae: Ground and Tiger Beetles Eburia quadrigeminata (Say) Agonum cupripenne (Say) Elaphidion mucronatum (Say) Agonum extensicolle (Say) Eupogonus tomentosus (Halderman) Agonum galvestonicum (Casey) Gaurotes cyanipennis (Say) Agonum harrisii LeConte Graphisurus fisciatus (DeGeer) Agonum lutulentum (LeConte) Megacyllene robiniae (Forster) Agonum melanarium (Dejean) Micranoplium unicolor (Haldeman) Agonum placidum (Say) Orthosoma brunneum Forster Agonum tenue (LeConte) Prionus laticollis (Drury) Amara cupreolata Putzeys Smodicum cucujiforme (Say) Amara impuncticollis (Say) Xylotrechus colonus (F.) Amara pallipes Kirby Chrysomelidae: Leaf Beetles Anisodactylus discoideus Dejean Acalymma vittatum (F.) A nisodactylus furvus LeConte Altica litigata Fall Anisodactylus harrisii LeConte Calligrapha californica Brown Anisodactylus kirbyi Lindroth Chalepus dorsalis Thunberg Bembidion affine Say Chrysochus auratus (F.) Bembidion americanum Dejean Diabrotica longicornis barberi S&L Bembidion bifossulatum (LeConte.) Diabrotica undecimpunctata howardi Barber Bembidion impotens Casey Disonycha pennsylvanica (Illiger) Bembidion patruele Dejean Donacia emarginata Kirby OHIO JOURNAL OF SCIENCE R. N. WILLIAMS ET AL. 229

TABLE 1 (Continued) TABLE 1 (Continued)

Species Species

Donacia subtilis Kunze Dytiscus hybridus Aube Lema collaris Say Graphoderus liberus (Say) Leptinotarsa decemlineata (Say) Hydaticus modestus Sharp Paria canella (F.) Hydrobius consimilis LeConte Phyllotreta striolata (F.) Hydrocanthus tricolor Say Trirhabda virgata LeConte Ilybius biguttulus (Germar) Tymnes tricolor (F.) Ilybius oblitus Sharp Zygogramma suturalis (F.) Laccophilus fasciatus Aube Cleridae: Checkered Beetles Thermonectus basillaris Harris Cymatodera undulata Say Thermonectus nigrofaciatus Aube Enoclerus nigripes (Say) Tropisternus lateralis nibatus (LeConte) Placopterus tboracicus (Olivier) Uvarus lacustris (Say) Zenedosus sanguineus (Say) Agabus sp. Coccinellidae: Lady Beetles Celina sp. Adalia bipunctata (L.) Copelatus sp. Brachyacantha ursina (F.) Hydaticus sp. Calvia 14-guttata (L.) Hydroporus sp. 1 Coccinella novemnotata Herbst Hydroporus sp. 2 Coleomegilla fuscilabris Mulsant Hygrotus sp. Cycloneda sanguinea (L.) Laccophilus sp. 1 Diomus terminatus (Say) Laccophilus sp. 2 Hippodamia glacialis (F.) Rbantus sp. Neomysia pullata (Say) Uvarus sp. Psyllobora vigintimaculata (Say) Elateridae: Click Beetles : Silk. Fungus Beetles Aeolus dorsalis (Say) Telmatophilus americanus LeConte Ampedus linteus (Say) Atomaria (s. str.) pusilla Payk. Conoderus bellus (Say) Atomaria (s. str.) sp. Hemicrepidius memonius (Herbst) Atomaria (Anchicara) sp. 1 Agriotes sp. Atomaria (Anchicara) sp. 2 Elmidae: Riffle Beetles Atomaria (Anchicara) sp. 3 Ancronyx variegata Germar Cucujidae: Flat Bark Beetles Dubiraphia vittata (Melsheimer) Cucujus clavipes F. Macronychus glabratus (Say) Curculionidae: Weevils trivittatus (W. Brown) Anametis granulata (Say) Stenelmis crenata (Say) Calendra australis (Chittenden) Stenelmis vittipennis Zimm. Ceutorhynchus americanus Buch. Stenelmis sp. Conotrachelus anaglypticus (Say) Endomychidae: Handsome Fungus Beetles Conotrachelus geminatus LeConte Endomychus biguttatus Say Cyrtepistomus castaneus (Roelof.) : Pleasing Fungus Beetles Dorytomus vagenotatus Csy. Lschyrus quadripunctatus (Olivier) Hypera compta (Say) Megalodacne fasciata (F.) Listronotus appendiculatus (Bohe.) Gyrinidae: Whirligig Beetles Listronotus porcellus (Say) Dinuetus assimilis Aube Listronotus squamiger (Say) Dinuetus discolor Aube Lixus concavus Say Haliplidae: Crawling Water Beetles Lixus macer LeConte Haliplus borealis LeConte Odontopus calceatus (Say) Haliplus triopsis Say Pelenomus sulcicollis (F.) Peltodytes duodecimpunctatus (Say) Sitona cylindricollis F. Peltodytes edentulatus (LeConte) Spbenophorus australis australis (Chittenden) Peltodytes muticus (LeConte) Stenopelmus rufinasus Gyll. Helodidae: Tanysphyrus lemnae (F.) Scirtes tibialis Guerin-Meneville Tychius picirostris (F.) Cyphon sp. 1 Tyloderma aereum (Say) Cyphon sp. 2 Smicronyx sp. : Mud-loving Beetles Dermestidae: Dermestid Beetles Neoheterocerus pallidus Say Anthrenus verbasci (L.) pusillus (Say) Dermestes caninus Germar Heterocerus sp. Dryopidae: Long-toed Beetles Histeridae: Hister Beetles Helichus sp. Anapleus marginatus LeC. Dytiscidae: Predac. Diving Beetles Euspilotus assimilis (Paykull) Acilius fraternus Harris Geomysaprinus obsidianus (Casey) Acilius semisulcatus Aube Hister abbreviatus F. Acilius sylvanus Hilsenhoff Phelister subrotundus Say Bidessus affinis (Say) Platysoma lecontei Marseul Celina angustata Aube Saprinus lugens Erichson Celina hubbelli Young Hydrophylidae: Water Scavenger Beetles Coptotomus interrogatus (F.) Berosus pantherinus LeConte Coptotomus lenticus Hilsenhoff Berosus peregrinus (Herbst.) Cybister fimbriolatus Say Berosus striatus (Say) 230 INSECTS OF THE KILLBUCK MARSH VOL. 95

TABLE 1 (Continued) TABLE 1 (Continued)

Species Species

Cercyon haemorrhoidalis (F.) Carpophilus sayi Parsons Cercyon herceus Smet. Colopterus niger (Say) Cercyon mendax Smet. Colopterus semitectus (Say) Cercyon pubescens LeConte Conotelus obscurus Erichson Cercyon roseni Knisch Cryptarcha ampla Erichson Cryptopleurum minutum (F.) Cryptarcha concinna Melsheimer consors (LeConte) Cryptarcha strigatula Parsons Enochrus hamiltoni (Horn) Epuraea alternata Parsons Enochrus ochraceus (Melsh.) Epuraea peltoides Horn Enochrus sayi Gundersen Epuraea rufa (Say) Helocombus bifidus (LeConte) Epuraea rufida (Melsheimer) Hydrobius melaneus (Germar) fasciatus Olivier Hydrochara obtusata (Say) Glischrochilus quadrisignatus (Say) Hydrochara soror Smetana Glischrochilus sanguinolentus (Olivier) Hydrophilus triangularis Say Lobiopa undulata (Say) Sphaeridium lunatum F. Omosita colon (L.) Sphaeridium scarabaeoides (L.) Omosita sp. Tropisternus blatchleyi (Orchym.) Phenolia grossa (F.) Tropisternus lateralis (F.) Prometopia sexmaculata (Say) Tropisternus mexicanus LaPorte Stelidota geminata (Say) Tropisternus natator (Orchymont) Colopterus sp. Anacaena sp. Noteridae: Berosus sp. Suphisellus puncticollis Crotch Enochrus sp. Passandridae: (formerly Cucujidae) Helophorus sp. Catogenus rufus (F.) Hydrobius sp. Pselaphidae: Hydrochus sp. Brachygluta sp. Laccobius sp. Psephenidae: Water-penny Beetles Paracymus sp. Psephenus herricki DeKay Tropisternus sp. Pyrochroidae: Fire-colored Beetles Laemophloeidae: (formerly Cucujidae) Dendroides concolor Newman Laemophloeus biguttatus (Say) Neopyrochroa flabelata (F.) Lampyridae: Lightningbugs Scarabaeidae: Scarab Beetles Ellychnia corrusea (L.) Aphodius distinctus (MHller) Photinus pyralis (L.) Aphodius rusticola Melsh. Photuris pennsylvanica (DeGeer) Aphodius stercorosus Melsh. Languriidae: Lizard Beetles Ataenius gracilis (Melsh.) Acropteryx gracilis Newman Ataenius imbricatus (Melsh.) Languria mozardi Latrielle Ataenius platensis (Blanch.) Loberus impressus LeC. Ataenius spretulus (Haldeman) Toramus pulchellus (LeC.) Ataenius strigatus (Say) Lathridiidae: Min. Br. Scav. Beetles Cyclocephala immaculata (Olivier) Corticarina cavicollis (Mann.) Euphoria inda (L.) Melanophthalma sp. Popillia japonica Newman Corticaria sp. Pelidnota punctata L. Leiodidae: Serica sericea (Illiger) Colon megasetosum Peck & Step. ms. Trox foveicollis Harold Lucanidae: Stag Beetles Trox hamatus Robinson Pseudolucanus capreolus (L.) Trox scaber L. : Netwinged Beetles Phyllophaga sp. Caenia dimidiata (F.) Silphidae: Carrion Beetles Melandryidae: False Darkling Beetles Nicrophorus orbicollis Say Dircaea quadrimaculata (Say) Nicrophorus pustulatus Hersch. Synchroa punctata Newman Nicrophorus tomentosus Weber Meloidae: Blister Beetles Necrodes surinamensis (F.) Epicauta pennsylvanica (DeGeer) Necrophila americana L. Mycetophagidae: Hairy Fungus Beetles Oiceoptoma inaequalis F. Litargus balteatus LeC. Oiceoptoma noveboracensis (Forst.) Typhaea stercorea L. : (formerly Cucujidae) Mycetophagus punclatus Say Ahasverus advena (Waltl) Mycetophagus sp. planatus Germ. Nitidulidae: Sap Beetles Telephanus velox (Hald.) Amphicrossus ciliatus (Olivier) Staphylinidae: Rove Beetles Carpophilus antiquus (Melsheimer) Achenomorphus corticinus Grav. Carpophilus brachypterus (Say) Anotylus nitidulus Grav. Carpophilus corticinus Erichson Creophilus maxillosus (L.) Carpophilus freemani Dobson Homoetarsus bicolor (Grav.) Carpophilus hemipterus (L.) Hoplandria lateralis (Melsh.) Carpophilus lugubris Murray Lathrobium rubripenne (Csy.) Carpophilus marginatus Erichson Lithocharis ochracea (Grav.) Carpophilus marginellus Motsch. Neobisnius occidentoides Frank OHIO JOURNAL OF SCIENCE R. N. WILLIAMS ET AL.

TABU-: 1 (Continued) TABLE 1 (Continued)

Species Species

Neobisnius sobrinus (Er.) Hesperocorixa sp. Tachyporus nitidulus (F.) Gerridae: Water Striders Philonthus alumnus Er. Gerris sp. Philonthus cyanipennis (F.) Naucoridae: Creeping Water Bugs Aleocharinae sp. Pelocoris sp. Bledius sp. 1 Nepidae: Waterscorpions Bledius sp. 2 Nepa apiculata Uhler Caipelimus sp. 1 Rdnatra fusca Beauvois Carpelimus sp. 2 Notonectidae: Backswimmers Carpelimus sp. 3 Notonecta sp. Carpelimus sp. 4 Pleidae: Pygmy Backswimmers Carpelimus sp. 5 Plea striola Fieber Lathrobium sp. Saldidae: Shore Bugs Lobrathium sp. Sdldula sp. Neohypnus sp. Philonthus sp. INVERTEBRATES (4 Families; 4 Species) Tenebrionidae: Darkling Beetles Phylum—ANNELIDA Alphitobius diaperinus (Panzer) Class—CLITELLATA: Worms & Leeches Cynaeus angustus (LeConte) GNATHOBDELLIDA Diaperis maculata Olivier Hirudiniclae: Tenebrio molitor L. Macrobdella decora Say Hymenorus sp. (formerly Alleculidae) RHYNCHOBDELLIDA Throcsidae: Glossiphoniidae: Trixagus chevrolati (Beauv.) Helobdella stagnalis L. Trogositidae: Bark-gnawing Beetles Class—CRUSTACEA Tenebroides mauritanicus (L.) Subclass—MALACOSTRACA AMPHIPODA DIPTERA (4 Families; 36 Species) Gammaridae: Ceratopogonidae: Biting Midges, Punkies, or No-see-ums Cragonyx sp. 1 pupa ISOPODA Chaoboridae: Phantom Midges Asellidae: Corethrella brakeleyi (Coquillett) Asellus sp. Chironomidae: Midges Limnophyes sp. Zaureliella sp. Culicidae: Mosquitoes Aedes abserratus (Felt & Young) Aedes aurifer (Coquillett) species of sap beetle, Omosita colon. However, they did Aedes canadensis (Theobald) not yield any of the rare sap beetles in the Aedes cinereus Meigen Nitidula or the rare/endangered carrion beetle Nicro- Aedes grossbecki Dyar & Knab phorus americana L., which has not been encountered Aedes sticticus (Meigen) Aedes stimulans (Walker) group in Ohio since 1974 when it was taken in Hocking Aedes triseriatus (Say) County by David Harris. Aedes trivittatus (Coquillett) The macroinvertebrate samplers yielded many im- Aedes vexans (Meigen) Anopheles barberi Coquillett mature specimens including three Diptera, two Coleoptera, Anopheles crucians Wiedemann one Hemiptera, two Odonata, and several other Anopheles punctipennis (Say) invertebrates. However, no riffle beetles (Elmidae) were Anopheles quadrimaculatus Say collected from these traps. Anopheles walkeri Theobald Coquillettidia perturbans (Walker) Sweep net sampling of foliage accounted for most of Culex erraticus (Dyar & Knab) the leaf beetles (Chrysomelidae), checkered beetles Culex pipiens L. (Cleridae), and lizard beetles (Languriidae). Aquatic sweep- Culex restuans Theobald Culex salinarius Coquillett ing produced several additional predacious diving Culex territans Walker beetles, one large leech {Macrobdella decora Say), and Culiseta inornata (Williston) one immature Odonata (Pachydiplax longipennis Burm.). Culiseta melanura (Coquillett) Ground beetle collecting was quite successful. Six Culiseta minnesotae Barr Culiseta morsitans (Theobald) species of Carabidae uncommon in Ohio were en- Orthopodomyia alba Baker countered: Agonum cupripenne (Say), Agonum Orthopodomyia signifora (Coquil.) galvestonicum (Casey), Chlaenius niger Randall, Oodes Psorophora ciliata (F.) americanus Dejean, (F.), and Steno- Psorophora ferox (von Humboldt) Psorophora horrida (Dyar & Knab) crepis cuprea (Chaudoir). Psorophora mathesoni (Belk. & He.) Many of the species listed were made available by other Uranotaenia sapphirina (Ost.-Sac.) entomologists who made collections in the Killbuck HETEROPTERA (HEMBPTERA) (7 Families; 8 Species) Marsh Wildlife Area. These include Culicidae and Chao- Corixidae: Water Boatmen boridae collected in 1993, by Corinne Rickabaugh and 232 INSECTS OF THE KILLBUCK MARSH VOL. 95

Doris Troutman, and many of the Carabidae, Dytiscidae, ment Center, The Ohio State University. Manuscript number 148-94. Hydrophilidae, and Heteroptera collected by Foster Purrington in 1987. The Odonata list was provided by LITERATURE CITED Robert Glotzhober of the Ohio Historical Society from Connell, W. A. 1956 Nitidulidae of Delaware. Delaware Agric. Expt. collections made by Foster Purrington, also from 1987. Sta.Tech. Bui. 318. 67 pp. Gillogly, L. R. 1969 Nitidulidae (Coleoptera) collected by the Noona Dan Expedition in the Philippine and Bismarck Islands. Entomol. ACKNOWLEDGEMENTS. Gratitude is extended to those who assisted in Medd. 37: 207-224. species determinations. They include: Foster Purrington: Carabidae Hester, F. E. and J. S. Dendy 1962 A multiple-plate sampler for aquatic (Coleoptera); Keith Phillips: Cerambycidae (Coleoptera); Peter Kovarik: macro-invertebrates. Trans. Am. Fish. Soc. 91: 420-421. Histeridae (Coleoptera); Miguel Archangelsky: Dytiscidae and Peck, S. B. and A. E. Davies 1980 Collecting small beetles with large Hydrophilidae (Coleoptera); Dave Shetlar: Scarabaeidae (Coleoptera); area window traps. Coleopterists Bui. 34: 237-239- George Keeney: Silphidae (Coleoptera); Robert Glotzhober and Susan Sommer, S. E. 1994 A benthic macroinvertebrate assessment of Heady: Odonata; Charles Triplehorn: Coleoptera; Roy Rings: Lepidop- organic pollution in six Wayne County, Ohio streams. Independent tera; Brian Armitage: Trichoptera; Richard Berry and Robert Restifo: Study Thesis, Dept. of Biology, College of Wooster, Wooster, OH. Culicidae (Diptera); Sturgis McKeever: Chaoboridae (Diptera); Jeff 66 pp. DeShon and Mike Bolton: aquatic invertebrates. Additional thanks go True, R. P., H. L. Barnett, C. K. Dorsey, and J. G. Leach I960 Oak wilt to Karl Stephan who, on short notice, painstakingly sorted through the in West Virginia. Bui. 448T, West Virginia Univ., Agric. Exp. Stn., light trap residue and nearly doubled the list of Coleoptera. We es- Morgantown, WV. 119 pp. pecially thank the Division of Wildlife of the Ohio Department of Williams, R. N., D. S. Fickle, R. J. Bartelt, and P. F. Dowd 1993 Re- Natural Resources for funding this project on Ohio Wildlife Diversity. sponses by adult Nitidulidae (Coleoptera) to synthetic aggregation Salaries and research support were provided by State and Federal pheromones, a coattractant and effects of trap design and place- Funds appropriated to the Ohio Agricultural Research and Develop- ment. Eur. J. Entomol. 90: 287-294.

The 1994 Paper of the Year Award

was presented at the Annual Meeting of the OAS at Otterbein College on 29 April 1995 to:

Brian C. Reeder and Wendy R. Eisner

Department of Biological and Environmental Sciences Morehead State University Morehead, Kentucky and Byrd Polar Research Center The Ohio State University Columbus, Ohio

for their paper:

'Halocene Biogeochemical and Pollen History of a Lake Erie, Ohio, Coastal Wetland"

The Ohio Journal of Science 94: 87-93