DOCSLIB.ORG

Elevational Studies of Silphidae (Insecta: Coleoptera) in Southeast Tennessee

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Elevational Studies of Silphidae (Insecta: Coleoptera) in Southeast Tennessee University of Tennessee, Knoxville TRACE: Tennessee Research and Creative Exchange Masters Theses Graduate School 6-1971 Elevational Studies of Silphidae (Insecta: Coleoptera) in Southeast Tennessee Barry Charles Lumpkin University of Tennessee, Knoxville Follow this and additional works at: https://trace.tennessee.edu/utk_gradthes Part of the Zoology Commons Recommended Citation Lumpkin, Barry Charles, "Elevational Studies of Silphidae (Insecta: Coleoptera) in Southeast Tennessee. " Master's Thesis, University of Tennessee, 1971. https://trace.tennessee.edu/utk_gradthes/4237 This Thesis is brought to you for free and open access by the Graduate School at TRACE: Tennessee Research and Creative Exchange. It has been accepted for inclusion in Masters Theses by an authorized administrator of TRACE: Tennessee Research and Creative Exchange. For more information, please contact [email protected]. To the Graduate Council: I am submitting herewith a thesis written by Barry Charles Lumpkin entitled "Elevational Studies of Silphidae (Insecta: Coleoptera) in Southeast Tennessee." I have examined the final electronic copy of this thesis for form and content and recommend that it be accepted in partial fulfillment of the requirements for the degree of Master of Science, with a major in Wildlife and Fisheries Science. Arthur C. Cole, Major Professor We have read this thesis and recommend its acceptance: J.N. Liles, R.R. Schmoller Accepted for the Council: Carolyn R. Hodges Vice Provost and Dean of the Graduate School (Original signatures are on file with official studentecor r ds.) /,/ May 4, 1971 To the Graduate Council: I am submitting herewith a thesis written by Barry Charles Lumpkin entitled 11Elevational Studies of Silphidae (Insecta: Coleoptera) in Southeast Tennessee." I recommend that it be accepted for nine quarter hours of credit in partial fulfillment of the requirements for the de­ gree of Master of Science, with a major in.Zoology. We have read this thesis and recommend its acceptance: Accepted for the Council: Graduate Studies and Research ELEVATIONAL STUDIES OF SILPHIDAE (INSECTA: COLEOPrERA) IN SOUTHEAST TENNESSEE A Thesis Presented to the Graduate Council of The University of Tennessee In Partial Fulfillment of the Requirements for the Degree Master of Science by Barry Charles Lumpkin June 1971 ACKNOWLEDGEMENTS The author wishes to express his sincere appreciation to his major professor, Dr. Arthur C. Cele, for his patient guidance and scholarly criticism in the preparation of this manuscript. Thanks are also given to the other members ef the author's advisory committee, Dr. J. N. Liles and Dr. R. R. Schmoller, for their editorial advice and technical assistance. Additional thanks is extended to the Great Smoky Mountains National Park and staff for its valuable assistance in carrying out this project, and for the use of the park insect collection as an aid in identification. Also thanks to the Department of Zoology and Ento­ mology, The University of Tennessee, Knoxville, for providing a re­ search vehicle. ii 97�663 ABSTRACT During the period of June 17-December 2, 1970 baited pit-fall traps were used to collect 3467 specimens of Silphidae and related car­ rion frequenting Coleoptera at varying altitudes (800-6500 ft. ) in south­ east Tennessee. Ten species of Silphidae and representatives of five other families of beetles (Carabidae, Staphylinidae, Histeridae, Scara­ baeidae, and Trogidae) were consistently collected from the traps during the study period, The genus Nicrophorous appeared to occupy the higher elevations in the study area while the genus Silphidae occupied the lower elevations. !· marginatus and!· hydrophiloides appeared to have their elevational centers at higher altitudes than those found in the study area. The population peaks for!· vespilloides, !• orbicouis, li• tomentosus, and�. americana were reached in late summer. !· margin­ atus, !· pustulatus, !• surinamensis, �. noveboracensis, and�. inegualis appear to have reached their population peaks before this study began. The Carabidae and Staphylinidae exhibited three centers of elevational distribution (800, 2000, and 5000 ft.). The Histeridae exhibited two elevational peaks (800 and 1000 ft.), while both the Scarabaeidae and Trogidae only exhibited one peak each (900 and 1000 ft.). The Histeridae, Scarabaeidae, and Trogidae appear to reach their population peaks in late summer, while Carabidae and Staphylinidae appear to have reached their population peaks before this study began. iii TABLE OF CONTENTS CHAPTER PAGE I. INTRODUCTION. .. 1 II. PROCEDURE . 3 Location of Study Area. 3 Materials and Methods . 12 Microclimatic Conditions of the Study Area. 16 III. RESUL_TS • 23 Identification. 23 Elevational Centers of Distribution of the Silphidae. 26 Seasonal Succession of the Silphidae. 26 Notes on Related Coleoptera . 30 DI. DISCUSSION. 38 v. SUMMA.RY . 44 LITERATURE CITED. 46 VITA. 50 iv LIST OF TABLES TABLE PAGE I. Altitude and County-State Locatien of Trap Sites •• . 6 II. Means (Degrees F.) of Ground Temperatures for Indi- vidual Traps and Trap Sites June 17 Through December 2 •• 18 III. Means (Degrees F.) of Air Temperatures for Individual Traps and Trap Sites June 17 Through December 2 ••• 21 rJ. Total Number of Coleoptera Cellected in Carrion Baited Traps June 17 Through December 2 . • . 24 v. Total Number of Silphidae Collected in Carrion Baited Traps June 17 Thr0Ugh December 2 . • • � . 24 VI. Numbers of Specimens of Silphidae Collected from A and B Traps at Each Trap Site June 17 Through December 2 . - . • 27 VII. Centers of Elevational Distribution for the Silphidae. • 0 29 VIII. Comparison of the Numbers of Silphidae Collected During Three Time Periods 0 • 0 e e • • e • . 31 IX. Numbers of Specimens of Related Coleoptera Collected from A and B Traps at Each Trap Site June 17 Through December 2 •••••• i •••••• . 33 X. Comparison of the Numbers of Related Coleoptera Collected During Three Time Periods., • • • • • Q • • • • • • • • • • 36 V LIST OF FIGURES FIGURE PAGE 1. Map of Study Area Showing Numbered Trap Sites Represented by X Marks. Small Insert Map Shows Study Area in Rela- tion to Tennessee•• v •••••••••••• . 5 2. Average Drop in Mean Ground Temperature (Degrees F.) Per Trap Site as Elevation is Increased ••••• •• • . 20 3. Average Drop in Mean Air Temperature (Degrees F.) Per Trap Site as Elevation is Increased. • . • . • . • . • 22 4. Elevational Data for N. vesJ2illoides (-) and S. americana (- ---) • . • • . • • • . • . • • . • 28 5. Elevational Data for!• orbicouis (-)and!• tomentosus E- --�) • e • • e • e • • • 0 • 0 0 • • • t • • . 28 6. Elevational Data for !•margina tus (-) and !• J2UStulatus t'-- � . 0 • ' . 28 7. Elevational Data for N. surinam.ensis (-)and!•hydro­ philoides � ---) • • • • D f 8 e e O e • e • 0 0 e • e • • • 28 . ' 8. · Seasonal Population Trends of Silphidae for Study Period June 17 Through December 2, !i� vespilloides (----�, !• orbicouis E-· -·), !• tom.entosus (-··-�, !• marginatus <--L !· pustu1atus (...... ), [9 americana �---L !· surinam.ensis (-•.-), and !• hydrophiloides (- - • �. • . 32 vi vii FIGURE PAGE 9. Elevational Data for Carabidae (-), Staphylinidae E- - --L and the Histeridae (• ••.•) • • ' • 0 • 0 • • • 0 • • . 34 10. Elevational Data for Scarabaeidae (--) and Trogidae �--� 34 11. Seasonal Population Trends of Related Coleoptera for Study Period June 17 Through December 2, Carabidae (-), Staphylinidae (----), Histeridae (• • • -�, Scarabaeidae E-·-. -, , and Trogidae (-.. -�• . • • • 41 37 12. Centers of Elevational Distribution for Silphidae••••••• 40 CHAPTER I INTRODUCTION The Silphidae (Coleoptera) have been universally recognized by researchers as carrion related insects, Many superstitions and false ideas have been passed down from generation to generation concerning these active little insects (Abbott, 1927a, Fabre, 1899) , Enormous amounts of information on these insects has appeared in scientific journals for quite some time. One of the earlier papers by C. U. Clark (1895) noted the feeding habits of these beetles, Further literature review reveals that almost all of the research that has been, and is being done on the Silphidae concerns their food locating and feeding habits (Steele, 1927; Walsh, 1933; Cole, 1942; Shubeck, 1968). Al­ though this seems to be the more important aspect of the relationship of silphids to man, other ecological factors need to be carefully studied in order to better understand these insects. One such factor is the elevational relationships of the Silphidae. It is known that there are many changes in environmentai infiuences (weather, soii, vegetation, etc.) as elevation increases or decreases, Likewise, the insect types should change along with these environmentai influences. Research on summer foliage insects of the Smoky Mountains of Tennessee by R. H. Whittaker in 1952, showed definite distributional patterns with relation to elevation in many groups of insects. Although ranges of most of the insects studied during this project overlapped tre­ mendously, centers of species distribution seemed to appear at various 1 2 elevations. Walker (1957) studied arthropods associated with decaying materials in our habitats of Middle Tennessee, but altitude was not a factor studied. Walker used different types of bait, and found that the degree of difference in collections from var ious habitats was quite large. G. F. Bornemissza (1957) studied the arthropod succession of carrion in Western Australia. Five different stages of carcass decompo­ sition were
Load more

Recommended publications
  • The Phylogeny of Ptiliidae (Coleoptera: Staphylinoidea) – the Smallest Beetles and Their Evolutionary Transformations
    77 (3): 433 – 455 2019 © Senckenberg Gesellschaft für Naturforschung, 2019. The phylogeny of Ptiliidae (Coleoptera: Staphylinoidea) – the smallest beetles and their evolutionary transformations ,1, 2 3 4 Alexey A. Polilov* , Ignacio Ribera , Margarita I. Yavorskaya , Anabela Cardoso 3, Vasily V. Grebennikov 5 & Rolf G. Beutel 4 1 Department of Entomology, Biological Faculty, Lomonosov Moscow State University, Moscow, Russia; Alexey A. Polilov * [polilov@gmail. com] — 2 Joint Russian-Vietnamese Tropical Research and Technological Center, Hanoi, Vietnam — 3 Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Barcelona, Spain; Ignacio Ribera [[email protected]]; Anabela Cardoso [[email protected]] — 4 Institut für Zoologie und Evolutionsforschung, FSU Jena, Jena, Germany; Margarita I. Yavorskaya [[email protected]]; Rolf G. Beutel [[email protected]] — 5 Canadian Food Inspection Agency, Ottawa, Canada; Vasily V. Grebennikov [[email protected]] — * Cor- responding author Accepted on November 13, 2019. Published online at www.senckenberg.de/arthropod-systematics on December 06, 2019. Published in print on December 20, 2019. Editors in charge: Martin Fikáček & Klaus-Dieter Klass. Abstract. The smallest beetles and the smallest non-parasitic insects belong to the staphylinoid family Ptiliidae. Their adult body length can be as small as 0.325 mm and is generally smaller than 1 mm. Here we address the phylogenetic relationships within the family using formal analyses of adult morphological characters and molecular data, and also a combination of both for the frst time. Strongly supported clades are Ptiliidae + Hydraenidae, Ptiliidae, Ptiliidae excl. Nossidium, Motschulskium and Sindosium, Nanosellini, and a clade comprising Acrotrichis, Smicrus, Nephanes and Baeocrara. A group comprising Actidium, Oligella and Micridium + Ptilium is also likely monophy- letic.
    [Show full text]
  • The Hoosier- Shawnee Ecological Assessment Area
    United States Department of Agriculture The Hoosier- Forest Service Shawnee Ecological North Central Assessment Research Station General Frank R. Thompson, III, Editor Technical Report NC-244 Thompson, Frank R., III, ed 2004. The Hoosier-Shawnee Ecological Assessment. Gen. Tech. Rep. NC-244. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Research Station. 267 p. This report is a scientific assessment of the characteristic composition, structure, and processes of ecosystems in the southern one-third of Illinois and Indiana and a small part of western Kentucky. It includes chapters on ecological sections and soils, water resources, forest, plants and communities, aquatic animals, terrestrial animals, forest diseases and pests, and exotic animals. The information presented provides a context for land and resource management planning on the Hoosier and Shawnee National Forests. ––––––––––––––––––––––––––– Key Words: crayfish, current conditions, communities, exotics, fish, forests, Hoosier National Forest, mussels, plants, Shawnee National Forest, soils, water resources, wildlife. Cover photograph: Camel Rock in Garden of the Gods Recreation Area, with Shawnee Hills and Garden of the Gods Wilderness in the back- ground, Shawnee National Forest, Illinois. Contents Preface....................................................................................................................... II North Central Research Station USDA Forest Service Acknowledgments ...................................................................................................
    [Show full text]
  • Ofcanada Part13
    THE INSECTS ANDARAOHNIDS OFCANADA PART13 The ofca,.m'ffitrslP; Coleo r* SgHHy'" THE INSECTS ANDARACHNIDS OFCANADA t%RT13 The Carrion Beetles of Canada and Alaska Coleoptera Silphidae and Agyrtidae Robert S. Andersonl and Stewart B. Peck2 Biosystematics Research Institute Ottawa, Ontario Research Branch Agriculture Canada Publication 1778 1985 rUniyersity of Alberta, Edmonton, Alberta 2Carleton University, Ottawa, Ontario oMinister of Supply and Services Canada 1985 Available in Canada through Authorized Bookstore Agents and other bookstores or by mail from Canadian Government Publishing Centre Supply and Services Canada Ottawa, Canada KIA 0S9 Catalogue No. A42-42,21985-l3E Canada: $7.00 ISBN 0-662-11752-5 Other Countries: $8.40 Price subject to change without notice Canadian Cataloguing in Publication Data Anderson, Robert Samuel The carrion beetles of Canada and Alaska (Coleoptera: Silphidae and Agyrtidae) (The Insects and arachnids of Canada, ISSN 0706-7313 ; pt. 13) (Publication ;1778) Includes bibliographical references and index. l. Silphidae. 2. Beetles - Canada. 3. Beetles -- Alaska. I. Peck, Stewart B. II. Canada. Agricul- ture Canada. Research Branch. III. Title. IV. Series. V. Series: Publication (Canada. Agri- culture Canada). English ; 1778. QL596.S5A5 1985 595.76 C85-097200-0 The Insects and Arachnids of Canada Part l. Collecting, Preparing, and Preserving Insects, Mites, and Spiders, compiled by J. E. H. Martin, Biosystematics Research Institute, Ottawa, 1977. 182 p. Price: Canada $3.50, other countries $4.20 (Canadian funds). Cat. No. A42-42/1977 -1. Partie 1. R6colte, prdparation et conservation des Insectes, des Acariens et des Araign6es, compil6 par J.E.H. Martin, Institut de recherches biosyst6- matiques, Ottawa, 1983.
    [Show full text]
  • A New Genus and Species Record for Mongolia and Republic of Korea, Alliphis Necrophilus Christie, 1983 (Parasitiformes: Eviphididae)
    Journal148 of Species Research 6(Special Edition):148-151,JOURNAL OF2017 SPECIES RESEARCH Vol. 6, Special Edition A new genus and species record for Mongolia and Republic of Korea, Alliphis necrophilus Christie, 1983 (Parasitiformes: Eviphididae) Eunsun Keum1, Badamdorj Bayartogtokh2 and Chuleui Jung3,* 1Research Institute of Agricultural Science and Technology, Andong National University, Andong 36729, Republic of Korea 2Department of Zoology, School of Biology and Biotechnology, National University of Mongolia, Ulaanbaatar 210646, Mongolia 3Department of Plant Medicine, Andong National University, Andong 36729, Republic of Korea *Correspondent: [email protected] Assocations of wingless mites with winged arthropods highlight important life history strategies and ecological interactions. Here we report a mesostigmatan mite species association with an earth-boring dung beetle. Mite spciemens were collected from the soil of organic apple orchards in the Province of Gyeongsangbuk-do, Republic of Korea, and cattle dung was collected from grassland in Mongolia. The species was identified as Alliphis necrophilus Christie, 1983 (Acari: Eviphididae). This is the first report of any species in the genus Alliphis in Korea. Some descriptive details and ecological remarks are also provided. Keywords: association, Chromogeotrupes auratus, dung beetle, Geotrupes amoenus, Geotrupidae, phoresy Ⓒ 2017 National Institute of Biological Resources DOI:10.12651/JSR.2017.6(S).148 INTRODUCTION the Eviphididae. It currently includes about 20 species mostly associated with scarab beetles and is primarily Mites in the superfamily Eviphidoidea are primarily distributed in the Palaearctic region, although some spe- humus and litter-inhabiting predators. Many mites in cies are found in Africa and Australia (Mašán and Halli- the Eviphidoidae have adapted to transient habitats such day, 2010).
    [Show full text]
  • View Preprint
    A peer-reviewed version of this preprint was published in PeerJ on 15 March 2017. View the peer-reviewed version (peerj.com/articles/3088), which is the preferred citable publication unless you specifically need to cite this preprint. Fusco NA, Zhao A, Munshi-South J. 2017. Urban forests sustain diverse carrion beetle assemblages in the New York City metropolitan area. PeerJ 5:e3088 https://doi.org/10.7717/peerj.3088 Urban forests sustain diverse carrion beetle assemblages in the New York City metropolitan area Nicole A. Fusco1, Anthony Zhao2 and Jason Munshi-South1,* 1Louis Calder Center – Biological Field Station, Fordham University, 31 Whippoorwill Road, Armonk, NY 10504, USA. 2Department of Entomology, University of Maryland, College Park, MD 20742, USA. *Corresponding author: Jason Munshi-South Louis Calder Center, Fordham University, 31 Whippoorwill Road, Armonk, NY 10504 USA [email protected] Phone: (914) 273-3078 x20 1 PeerJ Preprints | https://doi.org/10.7287/peerj.preprints.2481v2 | CC BY 4.0 Open Access | rec: 1 Feb 2017, publ: 1 Feb 2017 1 Abstract 2 Urbanization is an increasingly pervasive form of land transformation that reduces biodiversity 3 of many taxonomic groups. Beetles exhibit a broad range of responses to urbanization, likely due 4 to the high functional diversity in this order. Carrion beetles (Order: Coleoptera, Family: 5 Silphidae) provide an important ecosystem service by promoting decomposition of small-bodied 6 carcasses, and have previously been found to decline due to forest fragmentation caused by 7 urbanization. However, New York City (NYC) and many other cities have fairly large 8 continuous forest patches that support dense populations of small mammals, and thus may harbor 9 relatively robust carrion beetle communities in city parks.
    [Show full text]
  • Good-Bye Scydmaenidae, Or Why the Ant-Like Stone Beetles Should Become Megadiverse Staphylinidae Sensu Latissimo (Coleoptera)
    Eur. J. Entomol. 106: 275–301, 2009 http://www.eje.cz/scripts/viewabstract.php?abstract=1451 ISSN 1210-5759 (print), 1802-8829 (online) Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera) VASILY V. GREBENNIKOV1,2 and ALFRED F. NEWTON3 1Entomology Research Laboratory, Ottawa Plant and Seed Laboratories, Canadian Food Inspection Agency, K.W. Neatby Bldg., 960 Carling Avenue, Canadian National Collection of Insects, Arachnids and Nematodes, Ottawa, Ontario K1A 0C6, Canada; e-mail: [email protected] 2Entomology Group, Institut für Spezielle Zoologie und Evolutionsbiologie, Friedrich-Schiller-Universität Jena, Erbertstraße 1, 07743 Jena, Germany 3Field Museum of Natural History, 1400 South Lake Shore Drive, Chicago IL 60605-2496, USA; e-mail: [email protected] Key words. Coleoptera, Staphylinidae sensu latissimo, Staphylinine Group, Scydmaenidae, taxonomy, phylogeny, classification, morphology, larvae, 18S rDNA, parsimony, Bayesian, neighbour joining Abstract. Ant-like stone beetles (Coleoptera: Scydmaenidae) include more than 4,850 described species in about 90 genera main- tained as a separate cosmopolitan family since 1815. Recent authors have hypothesised that Scydmaenidae might be rooted deep inside rove-beetles (Staphylinidae). To test this hypothesis we analysed 206 parsimoniously informative larval and adult morpho- logical characters scored for 38 taxa. Strict consensus topologies from the shortest trees in all 12 analyses consistently placed Scyd- maenidae as sister to (Steninae + Euaesthetinae) in a monophyletic Staphylinine Group (with or without Oxyporinae). The single fully resolved and most consistently supported topology maintains a monophyletic Staphylinine Group consisting of Oxyporinae + (Megalopsidiinae + ((“Scydmaenidae” + (Steninae + Euaesthetinae)) + (Leptotyphlinae + (Pseudopsinae + (Paederinae + Staphylini- nae))))); Solierius lacks larval data and is ambiguously placed within the Group.
    [Show full text]
  • NATURAL RESOURCES COMMISSION Information Bulletin #2
    Indiana Register NATURAL RESOURCES COMMISSION Information Bulletin #2 (Eighth Amendment) December 1, 2017 SUBJECT: Roster of Indiana Animals, Insects, and Plants that are Extirpated, Endangered, Threatened, or Rare (also described as Special Concern). I. HISTORY The initial roster was published February 1, 1992 (15 IR 848), republished April 1, 1991 (15 IR 1312); and subsequently amended to include additional species and published on February 1, 2005 (28 IR 1581). Since 2005, revisions have been made to several of the endangered species lists. The term "special concern" replaced the references to "rare" as it relates to wild animals and is expanded to include species in a legal status transition. Federal funding is available for species that are endangered or of special concern. In the fourth amendment, posted at 20070815-IR-312070469NRA on August 15, 2007, "rare", "threatened", and "extirpated" classifications for insect species were retained. Classification definitions were added for vascular plant species. In this document, species are reclassified and scientific names were modified. In the fifth amendment, posted at 20120125-IR-312120047NRA on January 25, 2012, the listing for endangered reptiles and amphibians was modified. Notable was removal of the American bald eagle from the endangered species list. Some species removed from the endangered list were redesignated as "special concern". Amendments were made to use scientific names that are consistent with those for species of animals listed in 312 IAC 9. In the sixth amendment, posted at 20140129-IR-312140023NRA on January 29, 2014, the listing for endangered birds, mollusks, insects, and vascular plants was modified. Notable was the removal of the peregrine falcon from the endangered species list.
    [Show full text]
  • An Annotated List of Insects and Other Arthropods
    This file was created by scanning the printed publication. Text errors identified by the software have been corrected; however, some errors may remain. Invertebrates of the H.J. Andrews Experimental Forest, Western Cascade Range, Oregon. V: An Annotated List of Insects and Other Arthropods Gary L Parsons Gerasimos Cassis Andrew R. Moldenke John D. Lattin Norman H. Anderson Jeffrey C. Miller Paul Hammond Timothy D. Schowalter U.S. Department of Agriculture Forest Service Pacific Northwest Research Station Portland, Oregon November 1991 Parson, Gary L.; Cassis, Gerasimos; Moldenke, Andrew R.; Lattin, John D.; Anderson, Norman H.; Miller, Jeffrey C; Hammond, Paul; Schowalter, Timothy D. 1991. Invertebrates of the H.J. Andrews Experimental Forest, western Cascade Range, Oregon. V: An annotated list of insects and other arthropods. Gen. Tech. Rep. PNW-GTR-290. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 168 p. An annotated list of species of insects and other arthropods that have been col- lected and studies on the H.J. Andrews Experimental forest, western Cascade Range, Oregon. The list includes 459 families, 2,096 genera, and 3,402 species. All species have been authoritatively identified by more than 100 specialists. In- formation is included on habitat type, functional group, plant or animal host, relative abundances, collection information, and literature references where available. There is a brief discussion of the Andrews Forest as habitat for arthropods with photo- graphs of representative habitats within the Forest. Illustrations of selected ar- thropods are included as is a bibliography. Keywords: Invertebrates, insects, H.J. Andrews Experimental forest, arthropods, annotated list, forest ecosystem, old-growth forests.
    [Show full text]
  • Insect Succession on Carrion in Three Biogeoclimatic Zones of British
    INSECT SUCCESSION ON CARRION IN THREE BlOGEOCLlMATlC ZONES OF BRITISH.COLUMBIA a . Leigh Celerte Dillon B.Sc., University of Waterloo, 1993 THESIS SUBMITTED IN PARTIAL FULFILMENT OF i THE REQUIREMENTS FOR THE DEGR& OF -9 MASTER OF SCIENCE in the Deparbnent of Biological Sciences O Leigh Celeste Dillon, 1997 SIMON FRASER UNIVERSITY April 1997 All rights reserved. This work may not be reproduced in whole or in part, by photocopy or other means, without permiasion of the author. b , APPROVAL 3 Name: Leigh C. Dillon Degree: Master of Science i p Title of Thesis: 4 INSECT SUCC~%SIONON CARRION IN THREE BIOGEOCLIMATIC ZONES OF BRITISH COLUMBIA. Examining Committee: t Ghair: Dr. F. Law, Professor D . ~;~kndersdn,'AssistantProfessor, Senior Supervisor Zepartment of Biological Sciences, SFU -~ Dr. 4. Borden, Professor Department of Biological Sciences, SFU Dr.A. Harestad, Associate Professor. Department of Biological Sciences, SFU - Dr. P. Belton, ~djunct-~rofessor Department of Biological Sciences, SFU Public Examiner National Library Bibliotheque nationale qf Canada du Canada Acquisitions and Acquisitions et Bibliographic Services services bibliographiques 395 Wellington Street 395, rue Wellington Ottawa ON K1A ON4 Ottawa ON KlA ON4 Canada Canada ~ Your hle Votre reference Our h& Notre rekrence The author has granted a non- L'auteur a accorde une licence non + exclusive licence allowing the exclusive pennettant a la National Library of Canada to Bibliotheque nationale du Canada de reproduce, loan, distribute or sell reproduire, prGter, distribuer ou copies of th~sthesis in microform, vendre des copies de cette these sous paper or electronic formats. - la forme de microfiche/film, de reproduction sur papier ou sur format electronique.
    [Show full text]
  • Short-Term Response of Ground-Dwelling Macroarthropods to Shelterwood Harvests in a Productive Southern Appalachian Upland Hardwood Forest
    Short-term Response of Ground-dwelling Macroarthropods to Shelterwood Harvests in a Productive Southern Appalachian Upland Hardwood Forest John Westby-Gibson, Jr., Cathryn H. Greenberg, Christopher E. Moorman, T.G. Forrest, Tara L. Keyser, Dean M. Simon, and Gordon S. Warburton Forest Service Southern Research Station e-Research Paper RP-SRS-59 December 2017 AUTHORS: John Westby-Gibson, Jr., Student, University of North Carolina at Asheville, Department of Biology, One University Heights, Asheville, NC 28804; Cathryn H. Greenberg, Research Ecologist, U.S. Department of Agriculture, Forest Service, Southern Research Station, Bent Creek Experimental Forest, Asheville, NC 28806; Christopher E. Moorman, Professor, North Carolina State University, Fisheries, Wildlife, and Conservation Biology Program, Department of Forestry and Environmental Resources, Raleigh, NC 27695; T.G. Forrest, Professor, University of North Carolina at Asheville, Department of Biology, One University Heights, Asheville, NC 28804; Tara L. Keyser, Research Forester, U.S. Department of Agriculture, Forest Service, Southern Research Station, Bent Creek Experimental Forest, Asheville, NC 28806; Dean M. Simon, Wildlife Forester (retired), North Carolina Wildlife Resources Commission, 8676 Will Hudson Road, Lawndale, NC, 28090; and Gordon S. Warburton, Director of Western North Carolina Region (retired), North Carolina Wildlife Resources Commission, 783 Deepwoods Dr., Marion, NC 28752. Corresponding author: Cathryn H. Greenberg, Research Ecologist, U.S. Department of Agriculture, Forest Service, Southern Research Station, Bent Creek Experimental Forest, Asheville, NC 28806; [email protected]. Product Disclaimer The use of trade, firm, or model names in this publication is for reader information and does not imply endorsement by the U.S. Department of Agriculture of any product or service.
    [Show full text]
  • A CATALOG of the COLEÓPTERA of AMERICA NORTH of MEXICO FAMILY: SILPHIDAE R
    ÑaHPtk as A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO FAMILY: SILPHIDAE r: CO ;~~ in NAL Digitizing ProjectEll ah52928 ^^i. UNITED STATES AGRICULTURE PREPARED BY ñka DEPARTMENT OF HANDBOOK AGRICULTURAL ^S&?f AGRICULTURE NUMBER 529-28 RESEARCH ^^^^ SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO Fascicle' Family Year issued Fascicle' Family Year issued Fascicle' Family Year issued 1 Cupedidae 1979 46 Callirhipidae 102 Biphyllidae 2 Micromalthidae 1982 47 Heteroceridae 1978 103 Byturidae 1991 3 Carabidae 48 Limnichidae 1986 104 Mycetophagidae 4 Rhysodidae 1985 49 Dryopidae 1983 105 Ciidae ^. 1982 5 Amphizoidae 1984 50 Elmidae 1983 107 Prostomidae 6 Haliplidae 51 Buprestidae 109 Colydiidae 8 Noteridae 52 Cebrionidae 110 Monommatidae 9 Dytiscidae 53 Elateridae 111 Cephaloidae 10 Gyrinidae 54 Throscidae 112 Zopheridae 13 Sphaeriidae 55 Cerophytidae 115 Tenebrionidae 14 Hydroscaphidae 56 Perothopidae 116 Alleculidae 15 Hydraenidae 57 Eucnemidae 117 Lagriidae 16 Hydrophilidae 58 Telegeusidae 118 Salpingidae 17 Georyssidae 61 Phengodidae 119 Mycteridae 18 Sphaeritidae 62 Lampyridae 120 Pyrochroidae 1983 20 Histeridae 63 Cantharidae 121 Othniidae 21 Ptiliidae 64 Lycidae 122 Inopeplidae 22 Limulodidae 65 Derodontidae 1989 123 Oedemeridae 23 Dasyceridae 66 Nosodendridae 124 Melandryidae 24 Micropeplidae 1984 67 Dermestidae 125 Mordellidae 1986 25 Leptinidae 69 Ptinidae 126 Rhipiphoridae 26 Leiodidae 70 Anobiidae 1982 127 Meloidae 27 Scydmaenidae 71 Bostrichidae 128 Anthicidae 28 Silphidae 1993 72 Lyctidae 129 Pedilidae 29 Scaphidiidae
    [Show full text]
  • A Faunal Study of Illinois Silphidae (Coleoptera)
    Eastern Illinois University The Keep Masters Theses Student Theses & Publications 1975 A Faunal Study of Illinois Silphidae (Coleoptera) Brian Baldwin Eastern Illinois University This research is a product of the graduate program in Zoology at Eastern Illinois University. Find out more about the program. Recommended Citation Baldwin, Brian, "A Faunal Study of Illinois Silphidae (Coleoptera)" (1975). Masters Theses. 2487. https://thekeep.eiu.edu/theses/2487 This is brought to you for free and open access by the Student Theses & Publications at The Keep. It has been accepted for inclusion in Masters Theses by an authorized administrator of The Keep. For more information, please contact [email protected]. B A Fauna] Study of Ill ino1 {CoJeoptera) S1Jphidae (TITLE) BY Brian Baldwin Ed., 1971 B.S. in Eastern Illinois University, THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF .., ' ' Master of Science in Zoology IN THE GRADUATE SCHOOL, EASTERN ILLINOIS UNIVERSITY CHARLESTON, ILLINOIS 1975YEAR I HEREBY RECOMMEND THIS THESIS BE ACCEPTED AS FULFILLING THIS PART OF THE GRADUATE DEGREE CITED ABOVE ADVISER f<l?S- , , t)Jt(J - 1JJ14q /f7-5 DATE A Faunal of Illinois Study (Coleoptera) Silphidae By Brian Baldwin Ed., Eastern Illinois University, 1971 B.S. in ABSTRACT OF A THESIS Submitted fulfillment of the requirements for the in partial degree of of at the Graduate School of Master Science Eastern Illinois University CHARLESTON, ILLINOIS 1975 of the family Silphidae Illinois was A faunal study in made through examination of over 1400 A brief specimens. history of the taxonomy of the family a description of silphid and ecology was presented.
    [Show full text]