The Silurian of the Pyrenees
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Brachiopod Phylogeny in the Cambrian Guliforms, Obolellates and Rhynchonelliforms (E.G., Zhang Et Al., 2009, 2014, 2015; Holmer Et Al., 2018A)
Permophiles Issue #66 Supplement 1 pods which may further address these questions. Glenn A. Brock The world’s oceans are changing. IPCC (2013) predictions Department of Biological Sciences, Macquarie University, NSW suggest that by the end of the current century our seas will be 2109, Australia ZDUPHUDQGWKHLUS+VLJQL¿FDQWO\ORZHU$OWKRXJKWKLVZLOOEH a challenge to all organisms, how will it impact brachiopods? Leonid E. Popov Given that they have a higher proportion of mineralised tissue 'HSDUWPHQWRI*HRORJ\1DWLRQDO0XVHXPRI:DOHV&DUGL൵ than virtually any other invertebrate group what will be the CF10 3NP, UK WKUHDWWRWKHPHLWKHULQWHUPVRIJURZLQJWKHLUVKHOOVLQWKH¿UVW Brachiopods are richly represented in the rock record and instance or repairing and maintaining that shell once it is made? as early as the Cambrian, where they show an impressive diver- In this talk I will review a series of experiments and historical sity of form and in shell morphology (e.g., Harper et al., 2017). studies undertaken with Emma Cross and Lloyd Peck (Cross et 3UHVHQWO\ WKH JURXS LV ¿UPO\ URRWHG ZLWKLQ WKH ORSKRWURFKR- al., 2015, 2016, 2018) that seek to explore the answers to these zoan branch of the bilaterian tree based on molecular data. Our questions. UHVHDUFKKDVLGHQWL¿HGVRPHPHPEHUVRIWKHHQLJPDWLF(DUO\ References Cambrian organophosphatic tommotiids as belonging to the &URVV(/3HFN/6 +DUSHU(02FHDQDFLGL¿FD- brachiopod stem (e.g., Holmer et al., 2002). Subsequent discov- tion does not impact shell growth or repair of the Antarctic HULHVRIWKH¿UVWHYHUDUWLFXODWHGVFOHULWRPHVRIEccentrotheca, brachiopod Liothyrella uva (Broderip, 1833). Journal of Paterimitra, and the inferred bivalved scleritome of Micrina f rom Experimental Marine Biology and Ecology, 462, 29–35. the lower Cambrian of South Australia reveals these three tom- Cross, E.L., Peck, L.S., Lamare, M.D. -
SILURIAN TIMES NEWSLETTER of the INTERNATIONAL SUBCOMMISSION on SILURIAN STRATIGRAPHY (ISSS) (INTERNATIONAL COMMISSION on STRATIGRAPHY, ICS) No
SILURIAN TIMES NEWSLETTER OF THE INTERNATIONAL SUBCOMMISSION ON SILURIAN STRATIGRAPHY (ISSS) (INTERNATIONAL COMMISSION ON STRATIGRAPHY, ICS) No. 27 (for 2019) Edited by ZHAN Renbin INTERNATIONAL UNION OF GEOLOGICAL SCIENCES President: CHENG Qiuming (Canada) Vice-Presidents: Kristine ASCH (Germany) William CAVAZZA (Italy) Secretary General: Stanley C. FINNEY (USA) Treasurer: Hiroshi KITAZATO (Japan) INTERNATIONAL COMMISSION ON STRATIGRAPHY Chairman: David A.T. HARPER (UK) Vice-Chairman: Brian T. HUBER (USA) Secretary General: Philip GIBBARD (UK) SUBCOMMISSION ON SILURIAN STRATIGRAPHY Chairman: Petr ŠTORCH (Czech Republic) Vice-Chairman: Carlo CORRADINI (Italy) Secretary: ZHAN Renbin (China) Other titular members: Anna ANTOSHKINA (Russia) Carlton E. BRETT (USA) Bradley CRAMER (USA) David HOLLOWAY (Australia) Jisuo JIN (Canada) Anna KOZŁOWSKA (Poland) Jiří KŘÍŽ (Czech Republic) David K. LOYDELL (UK) Peep MÄNNIK (Estonia) Michael J. MELCHIN (Canada) Axel MUNNECKE (Germany) Silvio PERALTA (Argentina) Thijs VANDENBROUCKE (Belgium) WANG Yi (China) Živilė ŽIGAITĖ (Lithuania) Silurian Subcommission website: http://silurian.stratigraphy.org 1 CONTENTS CHAIRMAN’S CORNER 3 ANNUAL REPORT OF SILURIAN SUBCOMMISSION FOR 2019 7 INTERNATIONAL COMMISSION ON STRATGRAPHY STATUTES 15 REPORTS OF ACTIVITIES IN 2019 25 1. Report on the ISSS business meeting 2019 25 2. Report on the 15th International Symposium on Early/Lower Vertebrates 28 3. Report on the 13th International Symposium on the Ordovician System in conjunction with the 3rd Annual Meeting of IGCP 653 32 GUIDELINES FOR THE ISSS AWARD: KOREN' AWARD 33 ANNOUNCEMENTS OF MEETINGS and ACTIVITIES 34 1. Lithological Meeting: GEOLOGY OF REEFS 34 SILURIAN RESEARCH 2019: NEWS FROM THE MEMBERS 36 RECENT PUBLICATIONS ON THE SILURIAN RESEARCH 67 MEMBERSHIP NEWS 77 1. List of all Silurian workers and interested colleagues 77 2. -
Revised Correlation of Silurian Provincial Series of North America with Global and Regional Chronostratigraphic Units 13 and D Ccarb Chemostratigraphy
Revised correlation of Silurian Provincial Series of North America with global and regional chronostratigraphic units 13 and d Ccarb chemostratigraphy BRADLEY D. CRAMER, CARLTON E. BRETT, MICHAEL J. MELCHIN, PEEP MA¨ NNIK, MARK A. KLEFF- NER, PATRICK I. MCLAUGHLIN, DAVID K. LOYDELL, AXEL MUNNECKE, LENNART JEPPSSON, CARLO CORRADINI, FRANK R. BRUNTON AND MATTHEW R. SALTZMAN Cramer, B.D., Brett, C.E., Melchin, M.J., Ma¨nnik, P., Kleffner, M.A., McLaughlin, P.I., Loydell, D.K., Munnecke, A., Jeppsson, L., Corradini, C., Brunton, F.R. & Saltzman, M.R. 2011: Revised correlation of Silurian Provincial Series of North America with global 13 and regional chronostratigraphic units and d Ccarb chemostratigraphy. Lethaia,Vol.44, pp. 185–202. Recent revisions to the biostratigraphic and chronostratigraphic assignment of strata from the type area of the Niagaran Provincial Series (a regional chronostratigraphic unit) have demonstrated the need to revise the chronostratigraphic correlation of the Silurian System of North America. Recently, the working group to restudy the base of the Wen- lock Series has developed an extremely high-resolution global chronostratigraphy for the Telychian and Sheinwoodian stages by integrating graptolite and conodont biostratigra- 13 phy with carbonate carbon isotope (d Ccarb) chemostratigraphy. This improved global chronostratigraphy has required such significant chronostratigraphic revisions to the North American succession that much of the Silurian System in North America is cur- rently in a state of flux and needs further refinement. This report serves as an update of the progress on recalibrating the global chronostratigraphic correlation of North Ameri- can Provincial Series and Stage boundaries in their type area. -
University of Birmingham Carbon Isotope (13Ccarb) and Facies
University of Birmingham Carbon isotope (13Ccarb) and facies variability at the Wenlock-Ludlow boundary (Silurian) of the Midland Platform, UK Blain, John Allan; Wheeley, James; Ray, David DOI: 10.1139/cjes-2015-0194 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Blain, JA, Wheeley, J & Ray, D 2016, 'Carbon isotope (13Ccarb) and facies variability at the Wenlock-Ludlow boundary (Silurian) of the Midland Platform, UK', Canadian Journal of Earth Science. https://doi.org/10.1139/cjes-2015-0194 Link to publication on Research at Birmingham portal Publisher Rights Statement: Publisher Version of Record available at: http://dx.doi.org/10.1139/cjes-2015-0194 Validated Feb 2016 General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document. -
(Silurian) Anoxic Palaeo-Depressions at the Western Margin of the Murzuq Basin (Southwest Libya), Based on Gamma-Ray Spectrometry in Surface Exposures
GeoArabia, Vol. 11, No. 3, 2006 Gulf PetroLink, Bahrain Identification of early Llandovery (Silurian) anoxic palaeo-depressions at the western margin of the Murzuq Basin (southwest Libya), based on gamma-ray spectrometry in surface exposures Nuri Fello, Sebastian Lüning, Petr Štorch and Jonathan Redfern ABSTRACT Following the melting of the Gondwanan icecap and the resulting postglacial sea- level rise, organic-rich shales were deposited in shelfal palaeo-depressions across North Africa and Arabia during the latest Ordovician to earliest Silurian. The unit is absent on palaeohighs that were flooded only later when the anoxic event had already ended. The regional distribution of the Silurian black shale is now well-known for the subsurface of the central parts of the Murzuq Basin, in Libya, where many exploration wells have been drilled and where the shale represents the main hydrocarbon source rock. On well logs, the Silurian black shale is easily recognisable due to increased uranium concentrations and, therefore, elevated gamma-ray values. The uranium in the shales “precipitated” under oxygen- reduced conditions and generally a linear relationship between uranium and organic content is developed. The distribution of the Silurian organic-rich shales in the outcrop belts surrounding the Murzuq Basin has been long unknown because Saharan surface weathering has commonly destroyed the organic matter and black colour of the shales, making it complicated to identify the previously organic-rich unit in the field. In an attempt to distinguish (previously) organic-rich from organically lean shales at outcrop, seven sections that straddle the Ordovician-Silurian boundary were measured by portable gamma-ray spectrometer along the outcrops of the western margin of the Murzuq Basin. -
Redox Changes in the Deep Shelf of the East Baltic Basin in the Aeronian and Early Telychian (Early Silurian)
Proc. Estonian Acad. Sci. Geol., 2004, 53, 2, 94–124 Redox changes in the deep shelf of the East Baltic Basin in the Aeronian and early Telychian (early Silurian) Enli Kiipli Institute of Geology at Tallinn University of Technology, Estonia pst. 7, 10143 Tallinn, Estonia; [email protected] Received 27 March 2003, in revised form 11 February 2004 Abstract. Aeronian black shales and overlying Telychian greenish-grey and red claystones of the deep shelf of the East Baltic Basin indicate different synsedimentary redox conditions of the bottom water of the sea. In the Aeronian, the primary bioproductivity rise caused accumulation of organic- rich black shale in deep shelf and formation of microcrystalline limestone with chertification, chert nodules, and barite in shoreward areas. In the early Telychian, the bioproductivity decreased, as concluded from the absence of indicators of high primary bioproductivity in the Aeronian. The suggested mechanism regulating primary bioproductivity and oxygen content of bottom waters of deep shelf was a change from wind-induced upwelling in the Aeronian to downwelling in the Telychian. The sedimentation rate did not influence the change in the redox regime of the deep shelf sediment, as it was low for both the Aeronian and Telychian. Key words: Aeronian, Telychian, East Baltic deep shelf, redox conditions, up- and downwellings. INTRODUCTION Differences in lithology, mineralogy, and rock chemistry evidence changes at the Aeronian–Telychian boundary. The changes concern sea water chemistry and early diagenesis of sediments with implication on hydrodynamic and atmospheric circulation, and palaeogeography. The area of investigation is southwest Estonia and west Latvia, which form a central, deeper part of the Palaeozoic East Baltic basin, treated here as the deep shelf. -
The Classic Upper Ordovician Stratigraphy and Paleontology of the Eastern Cincinnati Arch
International Geoscience Programme Project 653 Third Annual Meeting - Athens, Ohio, USA Field Trip Guidebook THE CLASSIC UPPER ORDOVICIAN STRATIGRAPHY AND PALEONTOLOGY OF THE EASTERN CINCINNATI ARCH Carlton E. Brett – Kyle R. Hartshorn – Allison L. Young – Cameron E. Schwalbach – Alycia L. Stigall International Geoscience Programme (IGCP) Project 653 Third Annual Meeting - 2018 - Athens, Ohio, USA Field Trip Guidebook THE CLASSIC UPPER ORDOVICIAN STRATIGRAPHY AND PALEONTOLOGY OF THE EASTERN CINCINNATI ARCH Carlton E. Brett Department of Geology, University of Cincinnati, 2624 Clifton Avenue, Cincinnati, Ohio 45221, USA ([email protected]) Kyle R. Hartshorn Dry Dredgers, 6473 Jayfield Drive, Hamilton, Ohio 45011, USA ([email protected]) Allison L. Young Department of Geology, University of Cincinnati, 2624 Clifton Avenue, Cincinnati, Ohio 45221, USA ([email protected]) Cameron E. Schwalbach 1099 Clough Pike, Batavia, OH 45103, USA ([email protected]) Alycia L. Stigall Department of Geological Sciences and OHIO Center for Ecology and Evolutionary Studies, Ohio University, 316 Clippinger Lab, Athens, Ohio 45701, USA ([email protected]) ACKNOWLEDGMENTS We extend our thanks to the many colleagues and students who have aided us in our field work, discussions, and publications, including Chris Aucoin, Ben Dattilo, Brad Deline, Rebecca Freeman, Steve Holland, T.J. Malgieri, Pat McLaughlin, Charles Mitchell, Tim Paton, Alex Ries, Tom Schramm, and James Thomka. No less gratitude goes to the many local collectors, amateurs in name only: Jack Kallmeyer, Tom Bantel, Don Bissett, Dan Cooper, Stephen Felton, Ron Fine, Rich Fuchs, Bill Heimbrock, Jerry Rush, and dozens of other Dry Dredgers. We are also grateful to David Meyer and Arnie Miller for insightful discussions of the Cincinnatian, and to Richard A. -
International Stratigraphic Chart
INTERNATIONAL STRATIGRAPHIC CHART ICS International Commission on Stratigraphy Ma Ma Ma Ma Era Era Era Era Age Age Age Age Age Eon Eon Age Eon Age Eon Stage Stage Stage GSSP GSSP GSSA GSSP GSSP Series Epoch Series Epoch Series Epoch Period Period Period Period System System System System Erathem Erathem Erathem Erathem Eonothem Eonothem Eonothem 145.5 ±4.0 Eonothem 359.2 ±2.5 542 Holocene Tithonian Famennian Ediacaran 0.0117 150.8 ±4.0 Upper 374.5 ±2.6 Neo- ~635 Upper Upper Kimmeridgian Frasnian Cryogenian 0.126 ~ 155.6 385.3 ±2.6 proterozoic 850 “Ionian” Oxfordian Givetian Tonian Pleistocene 0.781 161.2 ±4.0 Middle 391.8 ±2.7 1000 Calabrian Callovian Eifelian Stenian Quaternary 1.806 164.7 ±4.0 397.5 ±2.7 Meso- 1200 Gelasian Bathonian Emsian Ectasian proterozoic 2.588 Middle 167.7 ±3.5 Devonian 407.0 ±2.8 1400 Piacenzian Bajocian Lower Pragian Calymmian Pliocene 3.600 171.6 ±3.0 411.2 ±2.8 1600 Zanclean Aalenian Lochkovian Statherian Jurassic 5.332 175.6 ±2.0 416.0 ±2.8 Proterozoic 1800 Messinian Toarcian Pridoli Paleo- Orosirian 7.246 183.0 ±1.5 418.7 ±2.7 2050 Tortonian Pliensbachian Ludfordian proterozoic Rhyacian 11.608 Lower 189.6 ±1.5 Ludlow 421.3 ±2.6 2300 Serravallian Sinemurian Gorstian Siderian Miocene 13.82 196.5 ±1.0 422.9 ±2.5 2500 Neogene Langhian Hettangian Homerian 15.97 199.6 ±0.6 Wenlock 426.2 ±2.4 Neoarchean Burdigalian M e s o z i c Rhaetian Sheinwoodian 20.43 203.6 ±1.5 428.2 ±2.3 2800 Aquitanian Upper Norian Silurian Telychian 23.03 216.5 ±2.0 436.0 ±1.9 P r e c a m b i n P Mesoarchean C e n o z i c Chattian Carnian -
GEOLOGIC TIME SCALE V
GSA GEOLOGIC TIME SCALE v. 4.0 CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST HIST. ANOM. (Ma) ANOM. CHRON. CHRO HOLOCENE 1 C1 QUATER- 0.01 30 C30 66.0 541 CALABRIAN NARY PLEISTOCENE* 1.8 31 C31 MAASTRICHTIAN 252 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 Lopin- 254 32 C32 72.1 635 2A C2A PIACENZIAN WUCHIAPINGIAN PLIOCENE 3.6 gian 33 260 260 3 ZANCLEAN CAPITANIAN NEOPRO- 5 C3 CAMPANIAN Guada- 265 750 CRYOGENIAN 5.3 80 C33 WORDIAN TEROZOIC 3A MESSINIAN LATE lupian 269 C3A 83.6 ROADIAN 272 850 7.2 SANTONIAN 4 KUNGURIAN C4 86.3 279 TONIAN CONIACIAN 280 4A Cisura- C4A TORTONIAN 90 89.8 1000 1000 PERMIAN ARTINSKIAN 10 5 TURONIAN lian C5 93.9 290 SAKMARIAN STENIAN 11.6 CENOMANIAN 296 SERRAVALLIAN 34 C34 ASSELIAN 299 5A 100 100 300 GZHELIAN 1200 C5A 13.8 LATE 304 KASIMOVIAN 307 1250 MESOPRO- 15 LANGHIAN ECTASIAN 5B C5B ALBIAN MIDDLE MOSCOVIAN 16.0 TEROZOIC 5C C5C 110 VANIAN 315 PENNSYL- 1400 EARLY 5D C5D MIOCENE 113 320 BASHKIRIAN 323 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 1500 CALYMMIAN 6 C6 APTIAN LATE 20 120 331 6A C6A 20.4 EARLY 1600 M0r 126 6B C6B AQUITANIAN M1 340 MIDDLE VISEAN MISSIS- M3 BARREMIAN SIPPIAN STATHERIAN C6C 23.0 6C 130 M5 CRETACEOUS 131 347 1750 HAUTERIVIAN 7 C7 CARBONIFEROUS EARLY TOURNAISIAN 1800 M10 134 25 7A C7A 359 8 C8 CHATTIAN VALANGINIAN M12 360 140 M14 139 FAMENNIAN OROSIRIAN 9 C9 M16 28.1 M18 BERRIASIAN 2000 PROTEROZOIC 10 C10 LATE -
The Early Ludfordian Leintwardinensis Graptolite Event and the Gorstian–Ludfordian Boundary in Bohemia (Silurian, Czech Republic)
This is the peer reviewed version of the following article: Štorch, P., Manda, Š., Loydell, D. K. (2014), The early Ludfordian leintwardinensis graptolite Event and the Gorstian–Ludfordian boundary in Bohemia (Silurian, Czech Republic). Palaeontology, 57: 1003–1043. doi: 10.1111/pala.12099, which has been published in final form at 10.1111/pala.12099. This article may be used for non-commercial purposes in accordance with Wiley Terms and Conditions for Self-Archiving. The early Ludfordian leintwardinensis graptolite Event and the Gorstian– Ludfordian boundary in Bohemia (Silurian, Czech Republic) Petr Štorch, Štěpán Manda, David K. Loydell Abstract The late Gorstian to early Ludfordian hemipelagic succession of the south-eastern part of the Prague Synform preserves a rich fossil record dominated by 28 species of planktic graptoloids associated with pelagic myodocopid ostracods, pelagic and nektobenthic orthocerid cephalopods, epibyssate bivalves, nektonic phyllocarids, rare dendroid graptolites, brachiopods, crinoids, trilobites, sponges and macroalgae. Faunal dynamics have been studied with particular reference to graptolites. The early Ludfordian leintwardinensis graptolite extinction Event manifests itself as a stepwise turnover of a moderate diversity graptolite fauna rather than an abrupt destruction of a flourishing biota. The simultaneous extinction of the spinose saetograptids Saetograptus clavulus, Saetograptus leintwardinensis and the rare S. sp. B. at the top of the S. leintwardinensisZone was preceded by a short- term acme of S. clavulus. Cucullograptus cf. aversus and C. rostratusvanished from the fossil record in the lower part of the Bohemograptus tenuis Biozone. No mass proliferation of Bohemograptus has been observed in the postextinction interval. Limited indigenous speciation gave rise to Pseudomonoclimacis kosoviensis and Pseudomonoclimacis cf.dalejensis. -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2018/08 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age GSSP GSSP GSSP GSSP EonothemErathem / Eon System / Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon Erathem / Era System / Period GSSA age (Ma) present ~ 145.0 358.9 ± 0.4 541.0 ±1.0 U/L Meghalayan 0.0042 Holocene M Northgrippian 0.0082 Tithonian Ediacaran L/E Greenlandian 152.1 ±0.9 ~ 635 Upper 0.0117 Famennian Neo- 0.126 Upper Kimmeridgian Cryogenian Middle 157.3 ±1.0 Upper proterozoic ~ 720 Pleistocene 0.781 372.2 ±1.6 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian Callovian 1000 Quaternary Gelasian 166.1 ±1.2 2.58 Bathonian 382.7 ±1.6 Stenian Middle 168.3 ±1.3 Piacenzian Bajocian 170.3 ±1.4 Givetian 1200 Pliocene 3.600 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Devonian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.63 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Proterozoic Neogene Sinemurian Langhian 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- 2050 Burdigalian Hettangian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 27.82 Gorstian -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2014/02 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age Erathem / Era System / Period GSSP GSSP age (Ma) GSSP GSSA EonothemErathem / Eon System / Era / Period EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon GSSP age (Ma) present ~ 145.0 358.9 ± 0.4 ~ 541.0 ±1.0 Holocene Ediacaran 0.0117 Tithonian Upper 152.1 ±0.9 Famennian ~ 635 0.126 Upper Kimmeridgian Neo- Cryogenian Middle 157.3 ±1.0 Upper proterozoic Pleistocene 0.781 372.2 ±1.6 850 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian 1000 Callovian 166.1 ±1.2 Quaternary Gelasian 2.58 382.7 ±1.6 Stenian Bathonian 168.3 ±1.3 Piacenzian Middle Bajocian Givetian 1200 Pliocene 3.600 170.3 ±1.4 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.62 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Langhian Sinemurian Proterozoic Neogene 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- Hettangian 2050 Burdigalian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 28.1 Gorstian Oligocene Upper Norian 427.4 ±0.5 2500 Rupelian Wenlock Homerian