Observations on Coleoptera 01 Mallorca, Balearic Islands

P.F. WHITEHEAD

Whitehead, P.F., 1993. Observations on Coleoptera 01 Mal/orca, Balearic Islands. SHNB BolI. Soco Hist. Nat. Ba/ears, 36:45-56 ISSN 0212-260X. Palma de Mal/orca. In the spring 01 1987 and autumn 01 1990 the writer amassed a large databank on the Coleoptera of Mal/orca, as a result 01 intensive lieldwork. The particular concern of the author lies with the conservation and historical development 01 in­ vertebrate faunas, understanding 01 which is largely dependant on the exact taxo­ nomic knowledge 01 the specialist entomologist. The lindings presented herewith @ are significant in a number 01 ways. Not only do they underline the potential bilo­ gical richness 01 Mal/orca, but they also begin to portray the quality of the launa. SOCIETAT D'HISTORIA Records lor hygrophilous, amphibious, and aquatic general such as Paragaeus, NATURAL DE LES BALEARS Oodes, , Cercyon, Paracymus, Limnoxenus and Cymbiodyta derñonstrate the advanced state 01 development of the launa of the north coast wetlands. This wil/ undoubtedly be confirmed by further work, and the evidence from other authors, as yet unpublished. New records provided here support launistic links with both and Alrica (Zonitis fernancastroi Pardo, Psylliodes laticollis Kutschera). Corsica and Sardinia (Crioceris asparagi var. campes tris Linnaeus) and Macarone­ sia (Dactylotrypes longicollis (Wol/aston)), although scolyfids such as Dactylotrypes are prime contenders lor assuming dissemination by mano Keywords: Mallorca, Coleoptera, conservation.

OBSERVACIONS SOBRE ELS COLEOPTERS DE MALLORCA, ILLES BALEARS. En el decurs de la primavera de 1987 i la tardor de 1990, com a resultat d'una intensa tasca de camp, es va reunir una considerable base de dades sobre coleopters de Mal/orca. L'autor s'interessa especialment per la conservació i desenvolupament historie de les faunes invertebrades, el qual ha de basar-se en un coneixement taxonomic precís i acurat per part deIs entomolegs. Les trobal/es presentad es en aquest trebal/ són signilicativament importants. A més a més de destacar la riquesa biologica potencial de Mal/orca, es comem;a a posar de rel/eu la qualitat d'aquesta fauna. Les trobal/es de generes higrofils, amfibis i aquatics com: Panagaeus, Oodes, Polistichus, Cercyon, Paracymus, Limnoxenus i Cymbio­ dyta demostren J'elevat grau de desenvolupament de les faunes de zones humides de la costa nord de Mallorca. Aixo sera indubtablement conlirmat per posteriors estudis. Les noves citacisus presentades recolzen els I/iganus launistics entre África i Europa (Zonitis fernoncastroi Pardo, Psylliodes láticollis Kustchera). així 46 BolI. Soco Híst. Nat. Balears, 36 (1993)

com amb Corsega i Sardenya (Crioceris) i amb Macaronésia (Doctylotrypes locjicollis (Wollas Toug). malgrat aquest sigui un cas d'antropocoria. Keywords: Mallorca, Coleoptera, Conservació.

P.F. WHITEHEAD, Moor Leys, Little Coberton, Pershore, Worcestershire, WR10 3eh. England.

Recepció del manuscrit, 03-feb-93. Revisió acceptada, 01-juny-93

Introduction lago received the "Tourism for Tomor­ row" award in direct recognition of new Although the Coleoptera of Mallorca conservation legislation. and the Balearic archipelago has been My visit of 1987 acquainted me relatively well documented (Jordá 1922, with the biotopes of Mallorca and its 1923, 1923a, 1927; Español 1951, entomofauna, although throughout there 1954, 1955, 1959, 1976, 1977; Compte, is bias towards the north coast of the 1953, 1963, 1966, 1968; Joliver 1953; island. In 1990, our arrival on Mallorca Bias, 1977, 1981; Jeanne & Zaballos, coincided with rather dramatic events; 1986; Montes & Soler 1986; Garcia­ during 5 days, mostly on the 8th-10th Aviles, 1990; Valladares & Montes October, up to 200 litres of rain per m2 1991), no faunal overview appears to fell in the north of the island, creating a have be en published since that of Te­ disaster area, especially in the settle­ nenbaum (1915). Because of the large ments Llubí, Muro, and Sa Pobla (Ba­ amount of new data yet to be uncover­ día d'Alcúdía, 28 oct. 1990; Mallorca ed, this may well remain sorne way off. Daíly Bulletin, 816, 27-10-1990. The This account utilises muy own ori­ issue of Badía d'Alcúdía makes illumi­ ginal observations, and provides limited nating reading on the anthropogenic discussion on important biocoenoses, in contribution to the subsequent flooding). the hope that they will contirbute, albeit This rain had a tremendous scour­ in a rather minor sense, to such a ing and dispersive effect on the wetland future endeavour. The general approach and riparian entomofaunas, large com­ to conservation on Mallorca, has how­ munities of which were swept into the ever, progressed with encouraging sea in the northern bays. We found speed. Between my visot of 1987 and Nepa cínerea Linnaeus (Hemiptera, Ne­ 1990 the S'albufera de Mallorca was pidae) surviving on the edge of the sea formally declared a "Pare Natural" and in the Badia de Pollensa up to 7 after the Govern Balear through its Díreccíó the flood. The October visit was there­ General del Medí Ambíent has declared fore dominated by study of these flood­ a number of new Area Natural d'Espe­ dispersed , and this is reflected cíal Interes, backed up by informative in the following lists. brochures. In 1992 the whole archipe- P.F. Whitehead, Observations on Coleoptera of Mallorca 47

Selected observations on Coleop­ consumed a fully-grown Eobania vermi­ tera culata (Müller), whilst living inside the shell for a number of days. The habits Although species new to the Ba­ of M. morbillosus (Fabricius) were docu­ learic Archipelago should have promi­ mented by the malacologist Boettger nence in this list, it also contains infor­ (1921). mation of biological interest as well as Dyschirius chalybaeus Putzeys. observations of a more routine nature. Salines de S'Albufera, 19-4-1987. Because there is no up-to-date over­ Bem-bidion laetum Brullé. One view of the Coleoptera of Mallorca, I record; on evaporite-rich mud near lake have experienced difficulty in assessing at S'Albufereta, 15-4-1987. the status of a number of species on Tachys (Porotachys) bisulcatus (Ni­ the island. Examples of these species colai). 6 amongst flood debris on beach include Myrmecopora la esa (Erichson) at mouth Torrent de S'Albufereta, 25-10- (found in numbers on the littoral of the 1990. Not listed for Mallorca by Jeanne Badia de Pollen~a), Otiorhynchus juven­ & Zaballos (1986). cus Gyllenhal (in flood drift, mouth of Tachys dimidia tus (Motschulsky). Torrent de S'Albufereta, October 1990) Originally described by Motschulsky as perhaps not seen since the early years a variety of Tachys scutellaris Ste­ of this century on Mallorca and princi­ phens, not specifically cited for Balea­ pally a French species (M.L. Cox, in rics by Jeanne & Zaballos (1986). 5, litt.) and Otiorhynchus trophonius Reitter with 160 Tachys scutellaris Stephens, (a population in the general area of amongst flood debris, mouth of Torrent Alcudia evidently somewhat "culture-fa­ de S'Albufera, October 1990. voured") with a holo-mediterranean dis­ Olisthopus elongatus Wollaston and tribution. None of these were cited by Olisthopus fusca tus Dejean. Although Tenenbaum (1915); I request the tole­ these two species are sometimes con­ rance of readers for apparent short­ fused, the evidence on Mallorca is that comings in this regard. they occupy distinct biotopes. O. elon­ Many record are omitted from this gatus occurs from sea-Ievel to at least account. I surveyed many of salines of 740m. altitude (breeding populations on Mallorca in detail, observing extensive limestone (pupae), Fornalutx, April faunas of halophilous species generally 1987), whilst O. fusca tus is evidently distributed in the Mediterranean Sea halophilous; under seaweed on beach area. Badia de Pollen~a, October 1990, sali­ nes de S'Albufera, April 1987. Zabrus ignavus Cziki. Fairly com­ Carabidae mon granivorous species of cultivated areas in the north of the island. One, Macrothorax morbillosus balearicus under seaweed on shingle, Badia de (Lapong). Widespread malacophagous Pollen~a, 22-10-1990. species. Adult eating Rumina decollata Harpalus fulvus Dejean. On ce­ (Linnaeus), Vall de Bóquer, 17-4-1987. ment-waste tip, El Tucan, near Alcúdia, During April 1987, a last instar larva April 1987, normally psammophilous. 48 BolI. Soco Hist. Nat. Balears, 36 (1993)

Anisodactylus virens (Dejean). April coexist at a single site, provides a 1987: Salines de S'Albufera and S'Albu­ strong optimistic assessment of its eco­ fe reta, saline mudo Halophilous species logical vigour. P. connexus flies at night recently found in large numbers, Parc and can cover long distan ces in optimal de S'Albufera (Palmer & Vives, 1993). conditions (Whitehead, 1992). Scybalicus oblongiusculus (Dejean). One site, males under seaweed, Badia de Pollenga, 22-10-1990. Not specified Dytiscidae for Balearic archipelago by Jeanne & Zaballos (1986). Principally a West Pa­ Potamonectes ceresyi (Aubé). Very laearctic species, reaching Africa. localised on Mallorca, presently known Acupalpus brunnipes (Sturm). One from two sites (Garcia-Aviles, 1990). In under flood debris on beach by Torrent saline drains, Salines de L1evant, 22-4- de S'Albufereta, 21-10-1990, with Acu­ 1987. palpus elegans (Dejean). Not cited for Copelatus haemorrhoidalis (Fabri­ Mallorca by Jeanne & Zaballos (1986). cius). One in flood debris, Parc de S'AI­ Panagaeus cruxmajor (Linnaeus). bufera, 23-10-1990. An important species with a high con­ Dytiscus circumflexus Fabricius. Ra­ servation value, declining in many parts re in the archipelago, known only from of its range. A primary wetland indica­ north-east Mallorca in clean, unpolluted tor, usually amongst helophytes. 3 in rivers and streams (Garcia-Aviles, 1990). flood debris, Parc de S'Albufera, 23-10- Dead on road, Alcúdia, 15-4-1987. 1990. Oodes gracilis Villa. An important species with a high conservation value, Hydrophilidae in established freshwater wetlands. Pair in flood debris, Parc de S'Albufera, 23- Cercyon sp. A Cercyon of the 10-1990. New to the Balearic archipela­ aquatic convexiusculus-group was found go and very localised in Spain. J. Vives in flood debris at the Parc de S'Albufe­ (Barcelona) is aware of records from ra on 23-10-1990. The specimen is ex­ Valencia, Teruel, Zaragoza and Prat de tremely like Cercyon sternalis Sharp, but L1obregat, Barcelona (Vives, in litt., 25- with slightly les$ strongly micros­ 2-1991). Jeanne & Zaballos (1986) re­ culptured elytra. It may prove to be an cord O. gracilis only from Gerona. This underscribed sibling. species highlights the importance of the Paracymus aeneus (Germar). Parc freshwater habitat in the Mediterranean de S'Albufera, 25-10-1990. basin. Limnoxenus niger (Zschach). Parc Polistichus connexus (Fourcroy). de S'Albufera, 23-10-1990. Generally widespread and sporadic in Cymbiodyta marginella (Fabricius). the Iberian peninsula (Vives & Vives, Parc de S'Albufera, 23-10-1990. AI­ 1978), with previous recors for Mallorca. though the Cercyon has so far resisted One in flood debris, Parc de S'AI­ attempts to identify it, these four hydro­ bufera, 23-10-1990. The fact that P. philids make a strong positive conser­ cruxmajor,O. gracilis and P. connexus vation statement, admirably underlining P.F. Whitehead, Observations on Coleoptera of Mallorca 49

the biological value of rich fen in warm­ One, flood debris on beach by Torrent temperate climates. de S'Albufereta, 25-10-1990. Ochthephilum jacquelini (Boieldieu). 9, flood debris on beach by Torrent de Histeridae S'Albufereta, 25-10-1990. Staphylinus ater Gravenhorst. Litto­ Kissister minimus (Aubé). One, un­ ral; Badia de POllen¡;a, Badia d'Alcúdia. der seaweed on fine beach-gravel, Ba­ Frequent, Parc de S'Albufera. dia de POllen¡;a, 23-10-1990. Whitehead Mycetoporus angularis Mulsant & (1990) recorded K. minimus breeding in Rey. In flight, Port d'Alcúdia, 28-10- large numbers in Somerset, England, in 1990. a similar situation. Sepedophilus pedicularius (Graven­ horst). Wetland species. Lago Esperan­ za, Port d'Alcúdia, 25-10-1990. Ptiliidae Spedophilus testaceus (Fabricius). During April 1987, specimens of Sepe­ Ptenidium nitidum (Heer). Under dophilus were taken from a huge sea­ logs with ants Iridomyrmex humilis washed trunk of Pinus halepensis Miller. Mayr, Parc de S'Albufera, 23-10-1990. I have compared them in detail with Bri­ tish S. testaceus and refer them to that species. The genus, however, is a diffi­ Stap hylin ida e cult one, and until the Balearic Sepedo­ philus are thoroughly overhauled, it is Dropephylla devillei (Bernhauer) (D. best to regard the identification as pro­ grandiloqua Luze). One on wall under· visional. The site is by the lila d'Alca­ Pinus halepensis Miller, Alcanada, 27- nada; the trunk has since been moved, 10-1990. This is thought to be a new and much of the softer heartwood remo­ species for the archipelago. it is a very ved. by wave action. southern record for D. devillei, although Myrmecopora laesa (Erichson). 70 the species has occured in the French in flood debris on beach, mouth of Alpes-Maritimes. The identification was Torrent de S'Albufereta, 21/25-10-1990. confirmed by Mr.P.M. Hammond, Lon­ Liogluta longiuscula (Gravenhorst). don. In flight, Parc de S'Albufera, 16-4-1987. Carpelimus anthracinus (Rey). Sali­ Atheta pittionii Scheerpeltz. In basi­ nes de S'Albufera, April 1987; on beach diomycete fungus under Pinus halepen­ Badia de Pollen¡;a, October 1990. sis Miller, S'Albufereta, 25-10-1990. The Anotylus nitidulus (Gravenhorst). A identity was confirmed by Herr V. few amongst flood debris on beach by Assing, Hanover. Torrent de S'Albufereta, 25-10-1990; on Atheta triangulum (Kraatz). Probably wall Alcanada, 27-10-1990. fairly widespread, although not cited by Stenus ater Mannerheim. Amongst Tenenbaum (1915). October 1990: in the cement waste, Lago Esperanza, Port Alcúdia area in fungi, under flood debris d'Alcúdia, 19-10-1990. on beaches, and in grass tussocks, Ochthephilum fracticorne (Paykull). Atalaia de la Victoria, 450 m altitude. 50 Bol/. SoCo Hist. Nat. Balears, 36 (1993)

Pselaphidae Buprestidae

Brachygluta globulicollis aubei Tour­ Acmaeodera cylindria Fabricius. nier. 80 in flood debris on beach by April 1987: breeding population in Pinus Torrent de S'Albufereta, 21/25-10-1990; halepensis Mil/er, Alcanada. a few, Pare de S'Albufera, 23-10-1990. Examples of both species have been seen by Dr. C. Besuchet, Geneva. Elateridae

Lacon punctatus (Herbst). 1987, 1990: breeding population in Pinus Scarabaeidae halepensis Mil/er, Cap de Menorca area, west of Alcúdia. Pleurophorus mediterranicus Pittino Agriotes sordibus (IIliger) varo scu­ & Mariani 1986. Although P. caesus tel/atus Schnaufuss. Frequently encoun­ (Creutzer) is widespread on Mallorca, tered in the north coast freshwater wet­ Dr. E. Piattella (Rom e) has, with cus­ lands. tomary expertise, identified oneexample of P. mediterranicus collected on the strandline at Port d'Alcúdia on 27-4- Nitidulidae 1987. This is the first record for Mal/or­ ca, although unrecognised P. mediterra­ Haptoncus luteolus (Erichson). 2 in nicus may exist in collections. flood drift on beach, by Torrent de S'AI­ Rhizotrogus lepidus Schaufuss. On bufereta, 25-10-1990. A specimen has beach near Pont des Anglesos, Pare de been authenticated by Mr. C. Johnson. S'Albufera, 16-4-1987. Endemic to the archipelago. Cryptophagidae

Clambidae Atomaria godarti Guil/ebeau. Mr. C. Johnson has identified one example Clambus minitus complicans Wo­ from flood debris, Badia de Pol/enºa, I/aston. 2 in flood debris on beach at October, 1990. mouth of Torrent de S'Albufereta, 25-10- 1990. Examples have been authenti­ cated by Mr. C. Johnson, Manchenter. Corylophidae Corylophus cassidoides (Marsham). A few in sedge IiUer, Pare de S'Albu­ Limnichidae (sometimes assimilat­ fera, October 1990. ed within Byrrhidae)

Bothriopterus atomus Mulsant. 4 in La tridiida e flood debris on beach, mouth of Torrent de S'Albufereta, 25-10-1990. Corticarina fulvipes (Comolli). Fairly P.F. Whitehead, Observations on Co/eoptera of Mallorca 51

general in sandy places at the coast. de Menorca, near Alcúdia, and Messor Also inland, south of Alcúdia, 1987. sp. at Atalaia de la Victoria.

Nycetophagidae Meloidae

Litargus ba/teatus LeConte. Wide­ Me/de vio/aceus Marsham. In Cy­ spread, normally corticolous species e.g. c/amini-Quercetum i1icis association Cala under bark of olive O/ea europaea Lin­ de Sant Vicent, April 1987. naeus, Port de Pollensa, October 1990; Me/de mediterraneus Müller. Alcú­ in flight, Parc de S'Albufera, 19-4-1987. dia town, 25-10-1990. This species is easily confused with Me/de rugosus Marsham. Tenebrionidae Zonitis fernancastroi Pardo. In flight, Parc de S'Albufera, 16-4-1987. Tentyria schauni Kraatx. Endemic This specimen has been identified by to Arxipelago, widespread on Mallorca. Professor M. Bologna, L'Aquila, to Dredating dipterous larvae (with sapri­ whom it has been presented. Z. fernan­ nus georgius Marseul, Hypocaccus bra­ castroi was described from Morocco and si/iensis (Paykull), Hypocaccus climi­ evidently remains a very scarce west diatus (1IIiger) at dead gavina (barus Palearctic species known from Barcelo­ cachinnans seusu tato), Es Trenc, 22-4- na and recently from France. New to 1987; eating other T. schaumi immo­ the Balearic archipelago. bilised on Road, near Alcúdia, October 1990; eating Apis me/lífera L., Parc de s'Albufera, April 1987. Arthicidae Asida moraguesi Schafuss. Ma­ lIorcan endemic. GenerallY an upland Spiniferes longipilis (Brisout) and species (Español, 1954), but one at Lepta/eus rodriguesi (Latreille). Of the Roman forum, Alcúdia, April 1987. Vall many species of anthicids noted under de Bóquer; Atalaia de la Victoria, Alcú­ flood debris, Badia de Pollen<;a, Octo­ dia. ber 1990, these were the least abun­ Miso/ampus goudoti erichsoni Vau­ dant, with 2 speciments of each. loger. Español (1954) touched on the habitat of this species in the trunks of trees, where they may be sapropha­ Cerambycidae gous. On two occasions I found groups of this species between the annual Arhopalus rusticus (Linnaeus). Re­ rings of dead trees (Pinus ha/epensis latively scarce in the Balearics (Compte Miller) in active colonies of ants. Their 1963). Breeding in sea-washed trunk of potential relationships remain unclear, Pinus ha/epensis Miller, lila d'Alcanada, and the association wiht ants appears April 1987. not to be obligate. The ants involved were Monomorium bicolor Mayr at Cap 52 BolI. Soco Hist. Nat. Balears, 36 (1993)

Chrysomelidae Bruchidae

Crioceris asparagi varo campestris Bruchidius caninus (Kraatz) and Linnaeus. One in orchard, foothills Bruchidius nanus (Germar) were freo behind Alcúdia port, 20-10-1990. The quently encountered in the hinterland, name given, rather than C. campestris, especially on flowers of Papa ver in cul­ is employed on the recommendation of tivated areas. Bruchidius varipictus (Mot­ Dr. M.L. Cox, London. New to the Ba­ schulsky) was frequently observed in learic archipelago. The distribution is the northern coastal zone, especially on centred on Corsica, Sardinia and Si­ flowers of Chrysanthemum coronarium cily, with a citation from the Florence Linnaeus. Bruchidius tritolii (Motschulsky) region of Italy (Dr. M.L. Cox, pers. was taken in flight, Salines de Llevant, comm.). 22-4-1987. Cyrtonus majoricensis Breit. An example of this endemic species was found in the pellet of a raptor, Falco Curculionidae tinnunculus Linnaeus, at 480 m altitude, Atalaia de la Victoria, Alcúdia, 24-10- Sitona discoideus Gyllenhal. One 1990. example on whitewashed wall, Alcana­ Chrysolina banksi (Fabricius). An da, 26-10-1990; it has been authenti­ interesting discussion on the hostplants cated by Mr. R.T. Thompson, London, of this species (Alexander 1984; Foster and presented to the Natural History 1984; Morris 1984; Petitpierre 1985) Museum, London. S. discoideus is a provides clear agreement that Labiatae largely western species extending from and Plantaginaceae both provide hosts. the Atlantic Islands north to Jersey; the Imagines on Plantago, near Alcúdia, 15- easternmost record appears to be from 4-1987. Italy. Evidently new to the Balearic ochroleucus (Marsham) archipelago. In flight, Port d'Alcúdia, 28-10-1990. Sitona intermedius Küster. Dry xe­ Psylliodes laticol/is Kutschera. One ric microsites, Vall de Bóquer, October is flood debris on beach, Badia de Po­ 1990. Host plant Genista (Velazquez et. lIenºa, 25-10-1990. New to the Balearic al., 1990). archipelago Species doubtfouly recorded Brachycerus plicatus (Gyllenhal). from Pollenºa (Heikertinger, como pers.; Fairly widespread in the northern coas­ in Jolivert, 1953) (but see Petitpierre, E. tal zone. On the coast of the Badia de & Palmer, M., this volume). Populations Pollenºa, Pancratium maritimum Lin­ extend from the lonian Island group naeus is a host plant. through Dalmatia and Italy, with a fur­ Brachycerus barbarus barbarus ther population in North Africa (Dr. M.L. (Linnaeus). I found this only at one site Cox, pers. comm.). A pre-Holocene Ba­ in small numbers, Atalaia de la Victoria, learic population satisfies the need for a Alcúdia, (470-490m. altitude) October biogeographical link in the known range; 1990, where Urginea maritima (Lin­ P. laticollis should be sought on other naeus) Baker is a likely host plant. An west Mediterranean islands. example of B. barbarus barbarus has P.F. Whitehead, Observations on Coleoptera of Mallorca 53 been authenticated by Mr. R.T. Thomp­ Conservation and the fauna son. Coniatus tamarisci (Fabricius). Du­ Effective conservation stems from a ring October 1990, as an example of wide range of primary objectives some passive dispersion, we found this spe­ of which overlap, but all of which ulti­ cies in logs of Tamarix swept up by the mately converge for the benefit of all sea on the north coast beaches. life, human culture and spirit. The con­ Pselactus spadix (Herbst). Breeding servation of habitat and niche benefits population in large sea-washed trunk of from effective scale, whereby many spe­ Pinus halepensis Miller, near Cap de cies, including endemics, may flourish. Menorca, April 1987. Without this the conservation of Rhyncolus strangulatus Perris. organisms at species level becomes in­ Breeding population on slopes of Atalaia creasingly difficult, although there are de la Victoria, Alcúdia, October 1990, in numerous successes which combine Pinus halapensis Miller. The oft-quoted rigorous legislation and advanced tech­ view that R. strangulatus is monopha­ nology. What has been achieved on gous on Picea is untenable. Mallorca in recent years gives clear Brachytemnus porcatus (Germar). grounds for optimism, but not compla­ Breeding in sea-washed trunk Pinus cency. The management and conserva­ halepensis Miller, lila d'Alcanada, April tion of the north-coast wetlands sets a 1987. high standard for other Mediterranean Sitophilus zeamais Motschulsky. 32 countries. At the same time, much of on whitewashed wall, 10m. long, 1 m the foredune system of the Badia d'AI­ high, Alcanada, 27-10-1990. Although a cúdia has been lost to coastal develop­ much longer run ot unwhitewashed wall ment. was available, the were concen­ Efects on invertebrate assemblages trated in a small area (several square have been particularly marked; the si­ feet) of whitewashed wall only. There tuation with regard to Hemiptera: Cydni­ was nothing to indicate source. Assem­ dae is of particular concern. Such wide­ blages of on walls, dlfficult to spread Palearctic species as Calathus rationalise, have been mentioned by mollis (Marsham) have suffered frag­ Allen (1989). mentatiorí and isolation of populations. In 1987, a small dune relic was identi­ fied as the only remaining area of open Scolytidae foredune between Alcúdia and Can Pi­ cafort. Dactylotrypes longicollis (Wollaston), Situated near Port d'Alcúdia, its The host plant, Phoenix canariensis beetle fauna included Cicindela flexuosa Chabaud, and beetle are indigenous to Fabricius, Macrothorax morbillosus (Fa­ the Canary Islands. A few assembled on bricius), Amara ingenua Duftschmid, whitewashed wall, Alcanada, 27-10-1990. Harpalus distinguendus Duftschmid, Bra­ D. longicol/is was not cited by Te­ dycellus verbasci (Duftschmid), Philon­ nenbaum (1915), but is presumably not thus dimidiatipennis Erichson, (at damp now rare on Mallorca. spots) Ouedius pal/ipes Lucas (O. his- 54 BolI. Soco Hist. Nat. Balears, 36 (1993)

panicus Bernhauer, Scarabaeus semi­ tessellatus Drapiez, Agabus biguttatus punctatus Fabricius, Blaps lusitanica (Olivier), Laccobius atrocephalus Reitter, Herbst, Stenosis intricata Reitter, Gono­ Platystethus nitens (Sahlberg) (Iarge cephalum rusticum Olivier and Psyllio­ form), Stenus guttula Müller, Nephus des pal/idipennis Rosenhauer. Cydnids redtenbacheri (Mulsant) as well as the included Aethus f1avicornis (Faabricius), orthopteran Tetrix ceperoi (Bolívar). Geotomus sp. and Microscylus brun­ Such spatially restricted pure-water as­ neus (Fabricius). sociations are fragile, vulnerable and By the time of our second visit in important. The frequent habit on Mallor­ 1990 the dunes had been removed to­ ca of impounding the hill streams on tally and replaced by a villa, the ento­ the terraced slopes of the sierras has mofauna having no nearby sites to dis­ effectively conserved and enhanced perso too their aquatic fauna. Valladares & Montes (1991) have reviewed the endemic Hydraena baleari­ ca D'Orchymont on Mallorca. They say: Acknowledgements "Fuentes, manantiales generalmente en­ cauzados o pequeños cursos fluviales I thank all those named in the text, como el de Algendar, constituyen el re­ many friends and eminent specialists for ducido hábitat de la especie. Estos me­ encouragement, in particular Dr. L. Bo­ dios suelen presentar sustrato de are­ rowiec (Wroclaw), Dr. M.L. Cox, M. Pal­ nas y grava, siendo los parámetros físi• mer (Mallorca), J. Vives (Barcelona) and co-químicos del agua bastante varia­ Dr. T. Yelamos (Barcelona). I thank the bles: 10-20°C de temperatura, pH 7.6- appointed referees of the first draft for 7.9 Y 500-1240 de conductividad. La their constructive and meticulous at­ necesidad de una drástica política de tention to detail, which enabled nume­ conservación para estos frágiles y redu­ rous improvements to be made. cidos ecosistemas dulceacuícolas es evidente. De su preservación depende la pervivencia de esta y otras especies Reference de insectos acuáticos endémicos o de distribución muy restringida". Alexander, K. 1984. Foodplants of I found a somewhat isolated popu­ Chrysolina banksi (F.) Chrysome­ lation of Hydraena balearica D'Orchy­ lidae. Col. News. 16:8. mont in the Vall de Bóquer near Port Allen, AA 1989. Beetles and bugs on de Pollenºa in an area of great land­ a Thames-side wall in autumn. En­ scape value. It occurs at a freshwater tomologist's Rec. J. Var., 101 :47- trickle descending the valley side on 49. basic clay; adults of H. balearica were Bias, M. 1977. Los Cerambícidos (Col.) sometimes found amongst thesaturated de las Islas Baleares. Rapp. stems of Adiantum capillus-veneris Comm. Int. Mer. Médit., 24(9):101- Linnaeus. 102 . . The associated fauna includes Bias, M. 1981. Sobra la presencia de Hor­ Chlaenius vestitus (Paykull), Hydroporus mosacus vandalitiae (Heyden 1870) P.F. Whitehead, Observations on Coleoptera of Mallorca 55

en la isla de Mallorca (Coi., Catopi­ García-Avilés, J.A. 1990. Insectos acuá­ dae). Rapp. Comm. Int. Mer. Médit. ticos de Baleares (Odonata, Ephe­ Boettger, C.R. 1921. Carabus morbillo­ meroptera, Heteroptera, Plecoptera sus F. und Otala tigri Gerv., eine y Coleoptera). Doctoral Thesis, Anpassungstudies. Abh. Sencken­ Univ. Complutense Madrid, 690 pp. berg. naturf. Ges., 37(4):321-325. Jeanne, C. & Zaballos, J.P. 1986. Cata­ Compte, A. 1953. Algunos coleópteros logue des coléopteres carabiques de Palma y sus alrededores Bol. de la Peninsule Iberique. Supp. Soco Hist. Nat. Baleares 1 :15-20. Bull. Soco Linn. Bord.:1-200. Compte, A. 1963. Los Cerambycidae de Jolivet, P. 1953. Les las islas Baleares. (Insecta, Co­ des Iles Baléares: Mém. Inst. royo leoptera). Bol. R. Soco Esp. Hist. Se. Nat. Belgique 2(50):1-88. Nat. (B), 61 :175-207. Jorda, J. 1922. Contribució al coneixe­ Compte, A. 1966. Resultados de una ment deis coleópters de les Ba­ expedición zoológica a las islas lears. Butll. Inst. Cat. d'Hist. Nat., Pitiusas. 2. Coleópteros. Bol. R. 2:128-132. Soco Esp. Hist. Nat. (B). 64:239- Jorda, J. 1923 Contribució al coneixe­ 275. ment deis coleópters de les Ba­ Compte, A. 1968. La fauna de Menorca lears. Formes inedites. Butll. Inst. y su origen. Coleoptera (94-142) in: Cat. d'Hist. Nat. 3:103-109. Revista de Menorca special is-sue Jorda, J. 1923b. Contribució al coneixe­ 212pp. ment deis coleópters de les Ba­ Español, F. 1951. Tenebriónidos de las lears. Formes inedites. Butll. Inst. Pitiusas (Baleares occidentales). Cat. d'Hist. Nat., 7:30-34. Eos., 27:7-41. Montes, C. & Soler, A.G. 1986. Lista Español, F. 1954. Los tenebriónidos de faunística y bibliográfica de los co­ Baleares. Trab. Mus. Cienc. Nat. leópteros acuáticos Dryopoidea Barcelona. (N.S. zool.) 5:1-96. (Dryopidae & Elmidae) de la Penín• Español, F. 1955. Los bostríquidos de sula Ibérica e Islas Baleares. Asoc. Cataluña y Baleares (Col.). Publ. Esp. Limnol. Pub., 3:1-38. Inst. Biol. Aplic., 21:107-135. Morris, M.G. 1984. Foodplants of Chryso­ Español, F. 1959. Los cléridos (Col.) de lina banksi (F) Col. News., 17:1-2. Cataluña y Baleares. Publ. Inst. Palmer, M. & Vives, J. Carabidae i Te­ Biol. Apl., 30:105-146. nebrionidae (Coleoptera) de s'Albu­ Español, F. 1976. Sobre els caverníco• fera de Mallorca: dades prelimi­ les terrestres de les Balears. Bol. nars. Boll. Soco Hist. Nat. Balears, Soco Hist. Nat. Balears. 21 :21-24. 36:65-76. Español, F. 1977. Sur quelques Anobii­ Petitpierre, E. 1985. Notas faunisticas y des (Col.) de les IlIes Baleares ecológicas sobre Chrysomelidae (nota 81). Rapp. Comm. int. Mer. (Coleoptera) de Mallorca y Cata­ Medit., 24(9):99-100. lunya. BolI. Soco Hist. Nat. Balears, Foster, A. 1984. A further note on the 29:31-36. foodplants of Chrysolina banksi Petitpierre, E. & Palmer, M. Noves (F.). Col. News. 17:5. aportacions a la fauna coleopte- 56 BolI. Soco Hist. Nat. Ba/ears, 36 (1993)

rológica de les IlIes Balears. Boll. Valladares, L.F. & Montes, C. 1991 b. Soco Hist. Nat. Balears, 36:77-82. Redescripción de Hydraena ba/ea­ Pittino, R. & Mariani, G. 1986. A revi­ rica D'Orchymont 1930 (Coleoptera: sion of the Old World species of ) E/ytron, 5:3-8. the genus Diastictus Muls. and its Velazquez, A.J., Zarazaga, MAA., & allies (Coleoptera, Aphodiidae, Psa­ Uterelo, R. 1990. Curculionoidea mmodiini). G. it. Ent., 3(12):1-165. (Coleoptera) from Navacerrad a, Sie­ Rico, E., Pérez, L.C. & Montes, C. rra de Guadarrama (Spain). Bo/. R. 1990. Lista faunística y bibliográfica Soco Esp. Hist. Nat. (Sec. Bio/.), de los Hydradephaga (Coleoptera: 85(1-4):17-37. Haliplidae, Hygrobiidae, Gyrinidae, Vives, J. & Vives, E. 1978. Carabidos Noteridae, Dystiscidae) de la Pe­ nuevos o interesantes para la pe­ nínsula Ibérica e Islas Baleares. nínsula Ibérica. Misc. Zoo/. 4(2): Asoc. Esp. Limno/. Pub. 7. 165-178. Tenenbaum, S. 1915. Fauna ko/eopte­ Whitehead, P.F. 1990. Carpelimus halo­ r%giczna wysp Ba/earskich. Ge­ philus Kies. (Col., Staphylinidae) bethner i Wolff, Warsaw. and other Coleoptera from north Valladares, L.F. & Montes, C. 1991. Lis­ Somerset. Entomologist's Rec. J. ta faunística y bibliográfica de los Var., 102:292. Hydraenidae (Coleoptera) de la Pe­ Whitehead, P.F. 1992. Recent evidence nínsula Ibérica e Islas Baleares. for the augmentation of British Asoc. Esp. Limno/., 10:1-93. beetle faunas from mainland Europe. Trans. Worcs. Nat. Club. 3:152-158.