THE BEETLES of DORSET An annotated checklist

Adrian Mylward

A note on using the checklist:

The checklist runs to over 150 pages. It has not been designed to be printed but species can be

found by using Control F to open the Search box.

You can then type in part/all of a species name to

find a summary of Dorset records.

Cover image: Cylindera germanica Cliff Tiger Beetle © Bryan Edwards

The Beetles of Dorset (Vice-county 9) In his History of Glanvilles Wootton of 1878 C.W. Dale included records of 920 species of Coleoptera, the vast majority of which were from the neighbourhood of his home there. Then in 1926, at the age of 23, E.J. Pearce published A List of the Coleoptera of Dorset. Pearce pulled together the efforts of many recorders, and he subsequently added three supplements, in 1927, 1929, and 1931, ‘bringing the grand total to date up to 2149 species and varieties’. Unlike Dale, who spent most of his life at Glanvilles Wootton, Pearce did not remain in the county. His 1957 Royal Entomological Society handbook on the Pselaphidae (now a subfamily of the beetle family Staphylinidae) is still much used today. The county list below, containing 2653 species, is the first since that of Pearce. Its design is deliberately simple, to enable it to be easily updated at a time of frequent new discoveries and continual changes in nomenclature. It is published online and the hope is that it will be updated annually. The nomenclature follows the Checklist of Beetles of the British Isles (Duff, A.G. 2018) but some names from the 2012 checklist have been added to help avoid confusion. Subgenera have only been included where they are needed to justify the order of the listing. In Pearce’s lists there are a few dates, but the majority of species names are followed only by a list of localities. These are fairly general: ‘Wareham’ or ‘Upwey’ for example, and the original list contained many species with just ‘“Dorset”; (A. Ford)’ added. The second supplement did include more detailed information on localities from A. Ford for some, but not all, such species. For a variety of reasons, not all of Pearce’s listed species have found their way into the list below. This checklist is not designed to be a guide to finding beetles in Dorset, but simply an inventory of species found in the county. A ‘record’ here is defined by the presence of a species at a certain location on a certain date, irrespective of the number of specimens. Where the number of records exceeds 9 it is listed as ‘many’. Thanks are due to all those who have contributed records of beetles over the years, but particularly to Alison Stewart of the Dorset Environmental Records Centre for all her help and support. Adrian F. Mylward December 2020

1. Family SPHAERIUSIDAE Erichson, Orectochilus villosus (Müller, O.F., 1776) 1845 Pearce: 2 localities First subsequent record: 1947 Number of subsequent records: many Sphaerius acaroides Waltl, 1838 Latest record: 2020 Pearce: no records First record: 1990 Number of records: 7 3. Family HALIPLIDAE Aubé, 1836 Latest record: 2002 Brychius elevatus (Panzer, 1793) 2. Family GYRINIDAE Latreille, 1810 Pearce: 4 localities First subsequent record: 1934 Number of subsequent records: many Gyrinus (Gyrinulus) minutus Fabricius, Latest record: 2020 1798 Pearce: 5 localities First subsequent record: 1937 Haliplus (Haliplidius) confinis Stephens, Number of subsequent records: many 1828 Latest record: 2019 Pearce: 2 localities First subsequent record: 1947 Number of subsequent records: many Gyrinus (Gyrinus) aeratus Stephens, 1835 Latest record: 2019 Pearce: 3 localities First subsequent record: 1977 Number of subsequent records: 2 Haliplus (Haliplidius) obliquus (Fabricius, Latest record: 1980 1787) Pearce: 4 localities Gyrinus (Gyrinus) caspius Ménétries, First subsequent record: 1991 Number of subsequent records: many 1832 Latest record: 2010 Pearce: 1 locality First subsequent record: 1937 Number of subsequent records: many Haliplus (Haliplus) fluviatilis Aubé, 1836 Latest record: 2020 Pearce: 4 localities First subsequent record: 1977 Number of subsequent records: many Gyrinus (Gyrinus) marinus Gyllenhal, Latest record: 2019 1808 Pearce: 2 localities Haliplus (Haliplus) heydeni Wehncke, First subsequent record: 1937 Number of subsequent records: many 1875 Latest record: 2019 Pearce: 4 localities First subsequent record: 1979 Number of subsequent records: many Gyrinus (Gyrinus) substriatus Stephens, Latest record: 2018 1828 Pearce: generally common and widely distributed Haliplus (Haliplus) immaculatus Gerhardt, First subsequent record: 1939 Number of subsequent records: many 1877 Latest record: 2019 Pearce: 1 locality First subsequent record: 1937 Number of subsequent records: 8 Gyrinus (Gyrinus) urinator Illiger, 1807 Latest record: 2005 Pearce: 1 locality First subsequent record: 1974 Number of subsequent records: many Haliplus (Haliplus) lineolatus Latest record: 2020 Mannerheim, 1844 Pearce: no records First record: 2009 Number of records: 1 Latest record: 2009

Haliplus (Haliplus) ruficollis (De Geer, 5. Family HYGROBIIDAE Régimbart, 1774) 1879 (1837) Pearce: 6 localities First subsequent record: 1947 Hygrobia hermanni (Fabricius, 1775) Number of subsequent records: many Pearce: 3 localities Latest record: 2020 First subsequent record: 1935 Number of subsequent records: many Haliplus (Haliplus) sibiricus Motschulsky, Latest record: 2020 1860 Pearce: 6 localities 6. Family DYTISCIDAE Leach, 1815 First subsequent record: 1943 Number of subsequent records: 7 Latest record: 2009 Agabus (Acatodes) congener (Thunberg, 1794) Pearce: no records Haliplus (Liaphlus) flavicollis Sturm, 1834 First record: 2019 Pearce: 3 localities Number of records: 1 First subsequent record: 1979 Latest record: 2019 Number of subsequent records: 5 Latest record: 2014 Agabus (Acatodes) sturmii (Gyllenhal in Haliplus (Liaphlus) fulvus (Fabricius, Schönherr, 1808) Pearce: 4 localities 1801) First subsequent record: 1939 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1937 Latest record: 2019 Number of subsequent records: many Latest record: 2014 Agabus (Agabus) labiatus (Brahm, 1790) Pearce: 3 localities Haliplus (Liaphlus) laminatus (Schaller, First subsequent record: 1934 1783) Number of subsequent records: 7 Pearce: 4 localities Latest record: 2008 First subsequent record: 2009 Number of subsequent records: 1 Latest record: 2009 Agabus (Agabus) uliginosus (Linnaeus, 1760) Pearce: 6 localities Haliplus (Neohaliplus) lineatocollis First subsequent record: 1934 (Marsham, 1802) Number of subsequent records: many Pearce: 7 localities Latest record: 2011 First subsequent record: 1935 Number of subsequent records: many Latest record: 2020 Agabus (Gaurodytes) affinis (Paykull, 1798) Pearce: no records 4. Family NOTERIDAE Thomson, C. First record: 1947 G., 1860 Number of records: many Latest record: 2019 Noterus clavicornis (De Geer, 1774) Pearce: 6 localities Agabus (Gaurodytes) biguttatus (Olivier, First subsequent record: 1935 1795) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1934 Number of subsequent records: many Latest record: 2015 Agabus (Gaurodytes) bipustulatus Agabus (Gaurodytes) unguicularis (Linnaeus, 1767) (Thomson, C.G., 1867) Pearce: generally common and widely distributed Pearce: 1 locality First subsequent record: 1936 First subsequent record: 2001 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 2001

Agabus (Gaurodytes) brunneus (Fabricius, Ilybius aenescens Thomson, C.G., 1870 1798) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1937 First subsequent record: 1993 Number of subsequent records: many Number of subsequent records: 5 Latest record: 2020 Latest record: 2007 Ilybius ater (De Geer, 1774) Agabus (Gaurodytes) conspersus Pearce: 2 localities (Marsham, 1802) First subsequent record: 1947 Number of subsequent records: many Pearce: 1 locality First subsequent record: 1964 Latest record: 2016 Number of subsequent records: 4 Latest record: 1981 Ilybius chalconatus (Panzer, 1796) Pearce: 4 localities (but probably including Agabus (Gaurodytes) didymus (Olivier, montanus) First subsequent record: 1934 1795) Number of subsequent records: 9 Pearce: 6 localities Latest record: 2013 First subsequent record: 1935 Number of subsequent records: many Latest record: 2020 Ilybius fenestratus (Fabricius, 1781) Pearce: 1 locality First subsequent record: 1993 Agabus (Gaurodytes) guttatus (Paykull, Number of subsequent records: 4 1798) Latest record: 2006 Pearce: 2 localities First subsequent record: 1963 Number of subsequent records: many Ilybius fuliginosus (Fabricius, 1792) Latest record: 2013 Pearce: 5 localities First subsequent record: 1947 Number of subsequent records: many Agabus (Gaurodytes) melanarius Aubé, Latest record: 2020 1837 Pearce: no records Ilybius guttiger (Gyllenhal, 1808) First record: 1987 Pearce: no records Number of records: many First record: 1979 Latest record: 2013 Number of records: many Latest record: 2017 Agabus (Gaurodytes) nebulosus (Forster, 1771) Ilybius montanus (Stephens, 1828) Pearce: 4 localities Pearce: no records (but see chalconatus above) First subsequent record: 1938 First record: 1934 Number of subsequent records: many Number of records: many Latest record: 2020 Latest record: 2019

Agabus (Gaurodytes) paludosus Ilybius quadriguttatus (Lacordaire, 1835) (Fabricius, 1801) Pearce: no records Pearce: 6 localities First record: 1947 First subsequent record: 1943 Number of records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019 Platambus maculatus (Linnaeus, 1758) Laccophilus minutus (Linnaeus, 1758) Pearce: 4 localities Pearce: 4 localities First subsequent record: 1947 First subsequent record: 1938 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Colymbetes fuscus (Linnaeus, 1758) Dytiscus circumcinctus Ahrens, 1811 Pearce: generally common and widely distributed Pearce: no records First subsequent record: 1935 First record: 1964 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1964

Nartus grapii (Gyllenhal, 1808) [2012 Dytiscus circumflexus Fabricius, 1801 Rhantus grapii] Pearce: no records Pearce: no records First record: 1937 First record: 1937 Number of records: 4 Number of records: many Latest record: 2007 Latest record: 2016 Dytiscus marginalis Linnaeus, 1758 Rhantus exsoletus (Forster, 1771) Pearce: 7 localities Pearce: “Dorset” (A. Ford) First subsequent record: 1947 First subsequent record: 1947 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2014 Dytiscus semisulcatus Müller, O.F., 1776 Rhantus frontalis (Marsham, 1802) Pearce: 7 localities Pearce: 1 locality First subsequent record: 1937 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2014 Latest record: 2006 Hydaticus seminiger (De Geer, 1774) Rhantus suturalis (MacLeay, 1825) Pearce: no records Pearce: 2 localities First record: 1947 First subsequent record: 1947 Number of records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019 Acilius sulcatus (Linnaeus, 1758) Rhantus suturellus (Harris, 1828) Pearce: 5 localities Pearce: “Dorset” (A. Ford) First subsequent record: 1936 First subsequent record: 1937 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2014 Graphoderus cinereus (Linnaeus, 1758) Liopterus haemorrhoidalis (Fabricius, Pearce: no records 1787) First record: 1984 Pearce: 3 localities Number of records: many First subsequent record: 1947 Latest record: 2015 Number of subsequent records: many Latest record: 2020 Hydroporus angustatus Sturm, 1835 Pearce: 1 locality Laccophilus hyalinus (De Geer, 1774) First subsequent record: 1937 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1947 Latest record: 2020 Number of subsequent records: 8 Latest record: 1993

Hydroporus discretus Fairmaire & Brisout Hydroporus longulus Mulsant & Rey, de Barneville, C., 1859 1861 Pearce: 2 localities Pearce: no records First subsequent record: 1935 First record: 1966 Number of subsequent records: many Number of records: 3 Latest record: 2019 Latest record: 2005

Hydroporus dorsalis s.l. [2012 Hydroporus marginatus (Duftschmid, Suphrodytes dorsalis] 1805) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1980 First subsequent record: 1933 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2005 Latest record: 2004

Hydroporus erythrocephalus (Linnaeus, Hydroporus melanarius Sturm, 1835 1758) Pearce: 1 locality Pearce: 4 localities First subsequent record: 2005 First subsequent record: 1937 Number of subsequent records: 3 Number of subsequent records: many Latest record: 2006 Latest record: 2020 Hydroporus memnonius Nicolai, 1822 Hydroporus ferrugineus Stephens, 1829 Pearce: 6 localities Pearce: no records First subsequent record: 1933 First record: 1933 Number of subsequent records: many Number of records: many Latest record: 2019 Latest record: 2006 Hydroporus necopinatus Fery, 1999 Hydroporus figuratus (Gyllenhal in Pearce: no records Sahlberg, C.R., 1826) First record: 1937 Pearce: no records Number of records: many First record: 2005 Latest record: 2019 Number of records: 2 Latest record: 2005 Hydroporus neglectus Schaum, 1845 Pearce: no records Hydroporus gyllenhalii Schiødte, 1841 First record: 1964 Pearce: 8 localities Number of records: many First subsequent record: 1933 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Hydroporus nigrita (Fabricius, 1792) Pearce: 1 locality Hydroporus incognitus Sharp, 1869 First subsequent record: 1981 Pearce: no records Number of subsequent records: many First record: 1947 Latest record: 2019 Number of records: many Latest record: 2019 Hydroporus obscurus Sturm, 1835 Pearce: 3 localities Hydroporus longicornis Sharp, 1871 First subsequent record: 1937 Pearce: no records Number of subsequent records: many First record: 2017 Latest record: 2020 Number of records: 1 Latest record: 2017 Hydroporus palustris (Linnaeus, 1760) Pearce: generally common and widely distributed First subsequent record: 1933 Number of subsequent records: many Latest record: 2020

Hydroporus planus (Fabricius, 1782) Oreodytes sanmarkii (Sahlberg, C. R., Pearce: 5 localities 1826) First subsequent record: 1933 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2015 Hydroporus pubescens (Gyllenhal, 1808) Pearce: generally common and widely distributed Stictotarsus duodecimpustulatus First subsequent record: 1933 (Fabricius, 1792) Number of subsequent records: many Pearce: 6 localities Latest record: 2019 First subsequent record: 1962 Number of subsequent records: many Hydroporus scalesianus Stephens, 1828 Latest record: 2007 Pearce: no records First record: 1997 Graptodytes bilineatus (Sturm, 1835) Number of records: 2 Pearce: no records Latest record: 1998 First record: 1981 Number of records: 8 Hydroporus striola (Gyllenhal in Sahlberg, Latest record: 2019 C. R., 1826) Pearce: 1 locality Graptodytes flavipes (Olivier, 1795) First subsequent record: 1937 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1947 Latest record: 2013 Number of subsequent records: many Latest record: 2019 Hydroporus tessellatus (Drapiez, 1819) Pearce: 8 localities Graptodytes granularis (Linnaeus, 1767) First subsequent record: 1933 Pearce: 2 localities (1892) Number of subsequent records: many First subsequent record: 1937 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Hydroporus tristis (Paykull, 1798) Pearce: 2 localities Graptodytes pictus (Fabricius, 1787) First subsequent record: 1937 Pearce: 7 localities Number of subsequent records: many First subsequent record: 1947 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Hydroporus umbrosus (Gyllenhal, 1808) Pearce: 2 localities (1890, 1923) Porhydrus lineatus (Fabricius, 1775) First subsequent record: 1937 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1936 Latest record: 2019 Number of subsequent records: many Latest record: 2014 Deronectes latus (Stephens, 1829) Pearce: 1 locality (1892) Stictonectes lepidus (Olivier, 1795) First subsequent record: 1968 Pearce: 7 localities Number of subsequent records: 4 First subsequent record: 1935 Latest record: 2009 Number of subsequent records: many Latest record: 2018 Nebrioporus elegans (Panzer, 1794) Pearce: 7 localities Hydrovatus clypealis Sharp, 1876 First subsequent record: 1933 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1937 Latest record: 2020 Number of subsequent records: many Latest record: 2014

Hygrotus (Coelambus) confluens Hydroglyphus geminus (Fabricius, 1792) (Fabricius, 1787) Pearce: no records Pearce: 3 localities First record: 1964 First subsequent record: 1938 Number of records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019 7. Family CARABIDAE Latreille, 1802 Hygrotus (Hygrotus) inaequalis (Fabricius, 1777) Leistus (Leistus) ferrugineus (Linnaeus, Pearce: 3 localities 1758) First subsequent record: 1937 Pearce: widely distributed Number of subsequent records: many First subsequent record: 1939 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Hygrotus (Hygrotus) versicolor (Schaller, 1783) Leistus (Leistus) fulvibarbis Dejean, 1826 Pearce: 3 localities Pearce: 3 localities First subsequent record: 1966 First subsequent record: 1935 Number of subsequent records: 3 Number of subsequent records: many Latest record: 2006 Latest record: 2019

Hygrotus (Leptolambus) Leistus (Leistus) terminatus (Panzer, 1793) impressopunctatus (Schaller, 1783) Pearce: 2 localities Pearce: no records First subsequent record: 1976 First record: 1981 Number of subsequent records: 3 Number of records: many Latest record: 2015 Latest record: 2020 Leistus (Pogonophorus) rufomarginatus Hygrotus (Leptolambus) novemlineatus (Duftschmid, 1812) (Stephens, 1829) Pearce: no records Pearce: 1 locality First record: 1965 No subsequent records Number of records: many Latest record: 2018 Hygrotus (Leptolambus) parallelogrammus (Ahrens, 1812) Leistus (Pogonophorus) spinibarbis Pearce: 2 localities (Fabricius, 1775) First subsequent record: 1997 Pearce: common and widely distributed Number of subsequent records: 1 First subsequent record: 1939 Latest record: 1997 Number of subsequent records: many Latest record: 2019 Hyphydrus ovatus (Linnaeus, 1760) Pearce: 6 localities Nebria brevicollis (Fabricius, 1792) First subsequent record: 1937 Pearce: abundant (but probably including salina) Number of subsequent records: many First subsequent record: 1935 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Bidessus unistriatus (Goeze, 1777) Pearce: 2 localities Nebria salina Fairmaire & Laboulbène, First subsequent record: 1942 1854 Number of subsequent records: 1 Pearce: no records (but see brevicollis above) Latest record: 1942 First record: 1935 Number of records: many Latest record: 2019 Notiophilus aquaticus (Linnaeus, 1758) Carabus (Archicarabus) nemoralis Müller, Pearce: 4 localities O.F., 1764 First subsequent record: 1935 Pearce: generally common Number of subsequent records: many First subsequent record: 1939 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Notiophilus biguttatus (Fabricius, 1779) Pearce: generally common and widely distributed Carabus (Carabus) arvensis Herbst, 1784 First subsequent record: 1935 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1936 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Notiophilus germinyi Fauvel, 1863 Pearce: 1 locality Carabus (Carabus) granulatus Linnaeus, First subsequent record: 1935 1758 Number of subsequent records: many Pearce: very common Latest record: 2018 First subsequent record: 1946 Number of subsequent records: many Notiophilus palustris (Duftschmid, 1812) Latest record: 2020 Pearce: widely distributed First subsequent record: 1955 Carabus (Hemicarabus) nitens Linnaeus, Number of subsequent records: many Latest record: 2019 1758 Pearce: common on swampy heaths First subsequent record: 1934 Notiophilus quadripunctatus Dejean, 1826 Number of subsequent records: many Pearce: 1 locality (1890) Latest record: 2005 First subsequent record: 1978 Number of subsequent records: 2 Latest record: 1982 Carabus (Megodontus) violaceus Linnaeus, 1758 Pearce: generally common Notiophilus rufipes Curtis, 1829 First subsequent record: 1939 Pearce: 1 locality (1911) Number of subsequent records: many First subsequent record: 1939 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Carabus (Mesocarabus) problematicus Notiophilus substriatus Waterhouse, G.R., Herbst, 1786 Pearce: 4 localities 1833 First subsequent record: 1934 Pearce: 8 localities Number of subsequent records: many First subsequent record: 1955 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Carabus (Morphocarabus) monilis Cychrus caraboides (Linnaeus, 1758) Fabricius, 1792 Pearce: 4 localities Pearce: 3 localities First subsequent record: 1942 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2019 Latest record: 2019

Calosoma sycophanta (Linnaeus, 1758) Cicindela campestris Linnaeus, 1758 Pearce: 1 locality Pearce: common and widely distributed First subsequent record: 2018 First subsequent record: 1936 Number of subsequent records: 1 Number of subsequent records: many Latest record: 2018 Latest record: 2020

Cicindela maritima Dejean in Latreille & Clivina collaris (Herbst, 1784) Dejean, 1822 Pearce: 2 localities Pearce: 1 locality First subsequent record: 1967 First subsequent record: 1964 Number of subsequent records: 2 Number of subsequent records: 1 Latest record: 1989 Latest record: 1964 Clivina fossor (Linnaeus, 1758) Cicindela sylvatica Linnaeus, 1758 Pearce: not common but widely distributed Pearce: 3 localities First subsequent record: 1955 First subsequent record: 1934 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Dyschirius (Dyschiriodes) aeneus (Dejean, Cylindera germanica (Linnaeus, 1758) 1825) Pearce: widely distributed on coast Pearce: 2 localities First subsequent record: 1936 First subsequent record: 1954 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Loricera pilicornis (Fabricius, 1775) Dyschirius (Dyschiriodes) impunctipennis Pearce: abundant Dawson, 1854 First subsequent record: 1938 Pearce: 2 localities Number of subsequent records: many No subsequent records Latest record: 2020 Dyschirius (Dyschiriodes) nitidus (Dejean, Blethisa multipunctata (Linnaeus, 1758) 1825) Pearce: 2 localities (1896, 1925) Pearce: no records First subsequent record: 1977 First record: 1964 Number of subsequent records: 7 Number of records: 1 Latest record: 2017 Latest record: 1964

Elaphrus (Elaphrus) riparius (Linnaeus, Dyschirius (Dyschiriodes) politus (Dejean, 1758) 1825) Pearce: common Pearce: 1 locality First subsequent record: 1940 First subsequent record: 1968 Number of subsequent records: many Number of subsequent records: 7 Latest record: 2020 Latest record: 2014

Elaphrus (Neoelaphrus) cupreus Dyschirius (Dyschiriodes) salinus Duftschmid, 1812 Schaum, 1843 Pearce: 4 localities Pearce: 2 localities First subsequent record: 1935 First subsequent record: 1989 Number of subsequent records: many Number of subsequent records: 5 Latest record: 2019 Latest record: 2013

Elaphrus (Neoelaphrus) uliginosus Dyschirius (Dyschiriodes) tristis Stephens, Fabricius, 1792 1827 [2012 Dyschirius luedersi] Pearce: 1 locality Pearce: 1 locality First subsequent record: 1955 First subsequent record: 1991 Number of subsequent records: 9 Number of subsequent records: many Latest record: 2018 Latest record: 2014 Dyschirius (Dyschirius) thoracicus (Rossi, Trechus (Trechus) quadristriatus 1790) (Schrank, 1781) Pearce: 2 localities Pearce: generally common and widely distributed First subsequent record: 1939 First subsequent record: 1942 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Dyschirius (Eudyschirius) globosus Trechus (Trechus) rubens (Fabricius, (Herbst, 1784) 1792) Pearce: 4 localities Pearce: 1 locality First subsequent record: 1935 First subsequent record: 1972 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 1972

Broscus cephalotes (Linnaeus, 1758) Asaphidion curtum (Heyden, 1870) Pearce: 7 localities Pearce: no records First subsequent record: 1935 First record: 1938 Number of subsequent records: many Number of records: many Latest record: 2018 Latest record: 2019

Aepopsis robinii (Laboulbène, 1849) [2012 Asaphidion flavipes (Linnaeus, 1760) Aepus robinii] Pearce: 8 localities (sensu lato) Pearce: 3 localities First subsequent record: 1988 No subsequent records Number of subsequent records: many Latest record: 2020 Aepus marinus (Strøm, 1783) Pearce: 2 localities Asaphidion pallipes (Duftschmid, 1812) First subsequent record: 1981 Pearce: 3 localities Number of subsequent records: 7 First subsequent record: 1954 Latest record: 2010 Number of subsequent records: many Latest record: 2010 Trechoblemus micros (Herbst, 1784) Pearce: no records Asaphidion stierlini (Heyden, 1880) First record: 1967 Pearce: no records Number of records: 3 First record: 2002 Latest record: 1989 Number of records: 3 Latest record: 2015 Trechus (Epaphius) secalis (Paykull, 1790) Bembidion (Actedium) pallidipenne Pearce: 2 localities (Illiger, 1802) First subsequent record: 1986 Pearce: 1 locality (1910) Number of subsequent records: 1 No subsequent records Latest record: 1986 Bembidion (Bembidion) quadrimaculatum Trechus (Trechus) fulvus Dejean, 1831 (Linnaeus, 1760) Pearce: 3 localities Pearce: 6 localities First subsequent record: 1954 First subsequent record: 1940 Number of subsequent records: 2 Number of subsequent records: many Latest record: 1954 Latest record: 2019

Trechus (Trechus) obtusus Erichson, 1837 Bembidion (Bembidion) quadripustulatum Pearce: 2 localities Audinet-Serville, 1821 First subsequent record: 1943 Pearce: 7 localities Number of subsequent records: many First subsequent record: 1993 Latest record: 2019 Number of subsequent records: 3 Latest record: 2013 Bembidion (Bembidionetolitzkya) Bembidion (Lymnaeum) nigropiceum atrocaeruleum (Stephens, 1828) (Marsham, 1802) Pearce: no records Pearce: 5 localities First record: 1988 First subsequent record: 1986 Number of records: 4 Number of subsequent records: 3 Latest record: 2009 Latest record: 1999

Bembidion (Bembidionetolitzkya) tibiale Bembidion (Metallina) lampros (Herbst, (Duftschmid, 1812) 1784) Pearce: 2 localities Pearce: generally common and widely distributed First subsequent record: 1963 First subsequent record: 1942 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Bembidion (Diplocampa) assimile Bembidion (Metallina) properans Gyllenhal, 1810 (Stephens, 1828) Pearce: 2 localities Pearce: no records First subsequent record: 1963 First record: 1935 Number of subsequent records: many Number of records: many Latest record: 2020 Latest record: 2020

Bembidion (Diplocampa) clarkii (Dawson, Bembidion (Neja) nigricorne Gyllenhal, 1849) 1827 Pearce: 1 locality (1852, 1896) Pearce: 1 locality (1925) First subsequent record: 1948 First subsequent record: 1935 Number of subsequent records: 5 Number of subsequent records: many Latest record: 2018 Latest record: 1981

Bembidion (Diplocampa) fumigatum Bembidion (Nepha) genei Küster, 1847 (Duftschmid, 1812) [2012 Bembidion illigeri] Pearce: no records Pearce: no records First record: 1963 First record: 1957 Number of records: many Number of records: many Latest record: 2020 Latest record: 2020

Bembidion (Emphanes) minimum Bembidion (Notaphemphanes) ephippium (Fabricius, 1792) (Marsham, 1802) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1943 First subsequent record: 1939 Number of subsequent records: 5 Number of subsequent records: 1 Latest record: 2005 Latest record: 1939

Bembidion (Emphanes) normannum Bembidion (Notaphus) varium (Olivier, Dejean, 1831 1795) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1982 First subsequent record: 1943 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2018

Bembidion (Eupetedromus) dentellum Bembidion (Ocydromus) decorum (Zenker (Thunberg, 1787) in Panzer, 1799) Pearce: “Dorset” (A. Ford) Pearce: 1 locality First subsequent record: 1942 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 1979 Bembidion (Ocydromus) saxatile Bembidion (Philochthus) aeneum Germar, Gyllenhal, 1827 1823 Pearce: 7 localities Pearce: 6 localities First subsequent record: 1953 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2019

Bembidion (Peryphanes) deletum Audinet- Bembidion (Philochthus) biguttatum Serville, 1821 (Fabricius, 1779) Pearce: 6 localities Pearce: abundant and widely distributed First subsequent record: 1951 First subsequent record: 1936 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2019

Bembidion (Peryphanes) stephensii Bembidion (Philochthus) guttula Crotch, 1866 (Fabricius, 1792) Pearce: 4 localities Pearce: common and widely distributed First subsequent record: 1954 First subsequent record: 1942 Number of subsequent records: many Number of subsequent records: many Latest record: 2016 Latest record: 2019

Bembidion (Peryphus) bruxellense Bembidion (Philochthus) iricolor Bedel, Wesmael, 1835 1879 Pearce: 1 locality Pearce: 1 locality First subsequent record: 1980 First subsequent record: 2004 Number of subsequent records: 5 Number of subsequent records: 1 Latest record: 2019 Latest record: 2004

Bembidion (Peryphus) bualei Jacquelin du Bembidion (Philochthus) lunulatum Val, 1852 (Geoffroy in Fourcroy, 1785) Pearce: 1 locality Pearce: 5 localities First subsequent record: 2000 First subsequent record: 1952 Number of subsequent records: 1 Number of subsequent records: many Latest record: 2000 Latest record: 2020

Bembidion (Peryphus) femoratum Sturm, Bembidion (Philochthus) mannerheimii 1825 Sahlberg, C.R., 1827 Pearce: no records Pearce: 4 localities First record: 1966 First subsequent record: 1963 Number of records: 8 Number of subsequent records: many Latest record: 2005 Latest record: 2020

Bembidion (Peryphus) maritimum Bembidion (Phyla) obtusum Audinet- (Stephens, 1835) Serville, 1821 Pearce: “Dorset” (A. Ford) Pearce: 8 localities First subsequent record: 1975 First subsequent record: 1949 Number of subsequent records: 1 Number of subsequent records: many Latest record: 1975 Latest record: 2020

Bembidion (Peryphus) tetracolum Say, Bembidion (Princidium) punctulatum 1823 Drapiez, 1820 Pearce: abundant and widely distributed Pearce: no records First subsequent record: 1942 First record: 1999 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1999 Bembidion (Semicampa) gilvipes Sturm, Sinechostictus stomoides (Dejean, 1831) 1825 [2012 Bembidion (Sinechostictus) Pearce: 2 localities stomoides] First subsequent record: 2006 Pearce: no records Number of subsequent records: 3 First record: 1989 Latest record: 2012 Number of records: 1 Latest record: 1989 Bembidion (Trepanedoris) doris (Panzer, 1796) Tachys (Paratachys) bistriatus Pearce: 2 localities (Duftschmid, 1812) First subsequent record: 1981 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1968 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Bembidion (Trepanes) articulatum (Panzer, 1796) Tachys (Paratachys) micros (Fischer von Pearce: 2 localities Waldheim, 1828) First subsequent record: 1943 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1947 Latest record: 2020 Number of subsequent records: many Latest record: 2017 Bracteon litorale (Olivier, 1790) Pearce: no records Tachys (Tachys) scutellaris Stephens, First record: 1914 1828 Number of records: 2 Pearce: no records Latest record: 1989 First record: 1938 Number of records: 6 Cillenus lateralis Samouelle, 1819 Latest record: 2006 Pearce: 2 localities First subsequent record: 1954 Tachyura parvula (Dejean, 1831) [2012 Number of subsequent records: 5 Latest record: 2019 Elaphropus parvulus] Pearce: no records First record: 2004 Ocys harpaloides (Audinet-Serville, 1821) Number of records: 3 Pearce: 7 localities (but probably including (or all) Latest record: 2017 tachysoides) No subsequent sensu stricto records Tachyura walkeriana (Sharp, 1913) [2012

Elaphropus walkerianus] Ocys quinquestriatus (Gyllenhal, 1810) Pearce: no records Pearce: 1 locality First record: 2019 First subsequent record: 1986 Number of records: 1 Number of subsequent records: 5 Latest record: 2019 Latest record: 2004

Pogonus chalceus (Marsham, 1802) Ocys tachysoides (Antoine, 1933) Pearce: 5 localities Pearce: no records (but see harpaloides above) First subsequent record: 1938 First record: 2014 Number of subsequent records: many Number of records: 4 Latest record: 2019 Latest record: 2020 Pogonus littoralis (Duftschmid, 1812) Pearce: 2 localities First subsequent record: 2013 Number of subsequent records: 3 Latest record: 2018

Pogonus luridipennis (Germar, 1822) Pterostichus (Adelosia) macer (Marsham, Pearce: 2 localities 1802) First subsequent record: 1996 Pearce: 3 localities Number of subsequent records: 1 First subsequent record: 1939 Latest record: 1996 Number of subsequent records: 8 Latest record: 2016 Patrobus atrorufus (Strøm, 1768) Pearce: 1 locality Pterostichus (Argutor) vernalis (Panzer, First subsequent record: 1986 1796) Number of subsequent records: 7 Pearce: 3 localities Latest record: 2013 First subsequent record: 1938 Number of subsequent records: many Brachinus crepitans (Linnaeus, 1758) Latest record: 2019 Pearce: 6 localities First subsequent record: 1939 Pterostichus (Bothriopterus) Number of subsequent records: many Latest record: 2019 oblongopunctatus (Fabricius, 1787) Pearce: 4 localities First subsequent record: 1936 Abax parallelepipedus (Piller & Number of subsequent records: many Mitterpacher, 1783) Latest record: 2016 Pearce: generally common and widely distributed First subsequent record: 1938 Pterostichus (Bothriopterus) Number of subsequent records: many Latest record: 2019 quadrifoveolatus Letzner, 1852 Pearce: no records First record: 2015 Pedius longicollis (Duftschmid, 1812) Number of records: 1 [2012 Pterostichus (Pedius) longicollis] Latest record: 2015 Pearce: 3 localities First subsequent record: 1968 Pterostichus (Morphnosoma) melanarius Number of subsequent records: 7 Latest record: 2019 (Illiger, 1798) Pearce: generally common First subsequent record: 1936 Poecilus cupreus (Linnaeus, 1758) Number of subsequent records: many Pearce: 7 localities Latest record: 2019 First subsequent record: 1936 Number of subsequent records: many Latest record: 2019 Pterostichus (Phonias) diligens (Sturm, 1824) Pearce: 2 localities Poecilus kugelanni (Panzer, 1796) First subsequent record: 1935 Pearce: 4 localities Number of subsequent records: many First subsequent record: 2001 Latest record: 2020 Number of subsequent records: 8 Latest record: 2003 Pterostichus (Phonias) strenuus (Panzer, Poecilus lepidus (Leske, 1785) 1796) Pearce: 3 localities Pearce: 9 localities First subsequent record: 1935 First subsequent record: 1946 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Poecilus versicolor (Sturm, 1824) Pterostichus (Platysma) niger (Schaller, Pearce: 4 localities 1783) First subsequent record: 1935 Pearce: generally common Number of subsequent records: many First subsequent record: 1936 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Pterostichus (Pseudomaseus) anthracinus Amara (Amara) communis (Panzer, 1797) (Illiger, 1798) Pearce: 5 localities Pearce: 3 localities First subsequent record: 1969 First subsequent record: 1999 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2018 Amara (Amara) convexior Stephens, 1828 Pterostichus (Pseudomaseus) gracilis Pearce: 8 localities (Dejean, 1828) First subsequent record: 1969 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1940 Latest record: 2020 Number of subsequent records: 8 Latest record: 2011 Amara (Amara) eurynota (Panzer, 1796) Pearce: 1 locality Pterostichus (Pseudomaseus) minor First subsequent record: 1990 Number of subsequent records: 9 (Gyllenhal, 1827) Latest record: 2019 Pearce: 5 localities First subsequent record: 1935 Number of subsequent records: many Amara (Amara) famelica Zimmermann, Latest record: 2019 C., 1832 Pearce: no records Pterostichus (Pseudomaseus) nigrita First record: 1977 Number of records: 2 (Paykull, 1790) Latest record: 1977 Pearce: 12 localities (but probably including rhaeticus) First subsequent record: 1994 Amara (Amara) familiaris (Duftschmid, Number of subsequent records: many 1812) Latest record: 2019 Pearce: 4 localities First subsequent record: 1942 Pterostichus (Pseudomaseus) rhaeticus Number of subsequent records: many Heer, 1837 Latest record: 2020 Pearce: no records (but see nigrita above) First record: 1994 Amara (Amara) lucida (Duftschmid, 1812) Number of records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1953 Number of subsequent records: 2 Pterostichus (Steropus) madidus Latest record: 1999 (Fabricius, 1775) Pearce: abundant Amara (Amara) lunicollis Schiødte, 1837 First subsequent record: 1935 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1934 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Stomis pumicatus (Panzer, 1796) Pearce: 5 localities Amara (Amara) montivaga Sturm, 1825 First subsequent record: 1952 Pearce: no records Number of subsequent records: many First record: 1994 Latest record: 2019 Number of records: 1 Latest record: 1994 Amara (Amara) aenea (De Geer, 1774) Pearce: generally common Amara (Amara) nitida Sturm, 1825 First subsequent record: 1938 Pearce: 1 locality Number of subsequent records: many No subsequent records Latest record: 2020 Amara (Amara) ovata (Fabricius, 1792) Amara (Percosia) equestris (Duftschmid, Pearce: 5 localities 1812) First subsequent record: 1955 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1954 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Amara (Amara) similata (Gyllenhal, 1810) Pearce: 3 localities Amara (Zezea) plebeja (Gyllenhal, 1810) First subsequent record: 1954 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1957 Latest record: 2015 Number of subsequent records: many Latest record: 2019 Amara (Amara) spreta Dejean, 1831 Pearce: 1 locality Curtonotus aulicus (Panzer, 1796) No subsequent records Pearce: 3 localities First subsequent record: 1980 Amara (Amara) tibialis (Paykull, 1798) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1935 Number of subsequent records: many Curtonotus convexiusculus (Marsham, Latest record: 2019 1802) Pearce: 4 localities Amara (Bradytus) apricaria (Paykull, First subsequent record: 1935 1790) Number of subsequent records: many Pearce: 5 localities Latest record: 2019 First subsequent record: 1942 Number of subsequent records: many Zabrus tenebrioides (Goeze, 1777) Latest record: 2019 Pearce: 5 localities First subsequent record: 1980 Amara (Bradytus) consularis (Duftschmid, Number of subsequent records: 3 1812) Latest record: 2005 Pearce: 1 locality First subsequent record: 1942 Oodes helopioides (Fabricius, 1792) Number of subsequent records: many Pearce: 3 localities Latest record: 2004 First subsequent record: 1955 Number of subsequent records: many Amara (Bradytus) fulva (Müller, O.F., Latest record: 2020

1776) Pearce: 3 localities Panagaeus bipustulatus (Fabricius, 1775) First subsequent record: 2012 Pearce: no records Number of subsequent records: 1 First record: 1941 Latest record: 2012 Number of records: 2 Latest record: 1982

Amara (Celia) bifrons (Gyllenhal, 1810) Pearce: 3 localities Chlaenius nigricornis (Fabricius, 1787) First subsequent record: 2002 Pearce: 4 localities Number of subsequent records: 1 First subsequent record: 1940 Latest record: 2002 Number of subsequent records: many Latest record: 2017

Amara (Celia) infima (Duftschmid, 1812) Pearce: 1 locality Chlaenius nitidulus (Schrank, 1781) No subsequent records Pearce: 2 localities First subsequent record: 1930 Number of subsequent records: 2 Latest record: 1933

Chlaenius vestitus (Paykull, 1790) Anisodactylus (Anisodactylus) binotatus Pearce: 7 localities (Fabricius, 1787) First subsequent record: 1964 Pearce: 6 localities Number of subsequent records: many First subsequent record: 1938 Latest record: 2017 Number of subsequent records: many Latest record: 2019 Badister (Badister) bullatus (Schrank, 1798) Anisodactylus (Anisodactylus) Pearce: abundant and widely distributed nemorivagus (Duftschmid, 1812) First subsequent record: 1936 Pearce: common on heaths Number of subsequent records: many First subsequent record: 2001 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Badister (Badister) unipustulatus Bonelli, 1813 Anisodactylus (Hexatrichus) poeciloides Pearce: 1 locality (Stephens, 1828) First subsequent record: 1934 Pearce: 1 locality Number of subsequent records: 3 No subsequent records Latest record: 1958 Scybalicus oblongiusculus (Dejean, 1829) Badister (Baudia) collaris Motschulsky, Pearce: 3 localities 1844 First subsequent record: 2019 Pearce: no records Number of subsequent records: 2 First record: 1954 Latest record: 2019 Number of records: 3 Latest record: 2014 Harpalus (Cryptophonus) melancholicus

Dejean, 1829 Badister (Baudia) dilatatus Chaudoir, Pearce: 1 locality (1885, 1894) 1837 No subsequent records Pearce: no records First record: 1991 Number of records: many Harpalus (Cryptophonus) tenebrosus Latest record: 2015 Dejean, 1829 Pearce: no records First record: 1979 Badister (Baudia) peltatus (Panzer, 1796) Number of records: 7 Pearce: 1 locality Latest record: 2003 No subsequent records

Harpalus (Harpalus) affinis (Schrank, Badister (Trimorphus) sodalis 1781) (Duftschmid, 1812) Pearce: generally common and widely distributed Pearce: 1 locality First subsequent record: 1935 First subsequent record: 1946 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2018

Harpalus (Harpalus) anxius (Duftschmid, Licinus depressus (Paykull, 1790) Pearce: no records 1812) First record: 1963 Pearce: 3 localities Number of records: many First subsequent record: 1935 Latest record: 2011 Number of subsequent records: many Latest record: 2018 Licinus punctatulus (Fabricius, 1792) Pearce: 6 localities First subsequent record: 1937 Number of subsequent records: many Latest record: 2018 Harpalus (Harpalus) attenuatus Stephens, Harpalus (Harpalus) serripes (Quensel in 1828 Schönherr, 1806) Pearce: 3 localities Pearce: 3 localities First subsequent record: 1954 First subsequent record: 1972 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2005

Harpalus (Harpalus) dimidiatus (Rossi, Harpalus (Harpalus) smaragdinus 1790) (Duftschmid, 1812) Pearce: 7 localities Pearce: 1 locality (1914) First subsequent record: 1939 First subsequent record: 1954 Number of subsequent records: many Number of subsequent records: 3 Latest record: 2019 Latest record: 2018

Harpalus (Harpalus) froelichii Sturm, Harpalus (Harpalus) tardus (Panzer, 1818 1796) Pearce: 3 localities Pearce: 2 localities No subsequent records First subsequent record: 1975 Number of subsequent records: many Harpalus (Harpalus) latus (Linnaeus, Latest record: 2015 1758) Pearce: 8 localities Harpalus (Pseudoophonus) rufipes (De First subsequent record: 1953 Geer, 1774) Number of subsequent records: many Pearce: abundant Latest record: 2019 First subsequent record: 1942 Number of subsequent records: many Harpalus (Harpalus) neglectus Audinet- Latest record: 2020 Serville, 1821 Pearce: 10 localities Ophonus (Hesperophonus) azureus First subsequent record: 1935 (Fabricius, 1775) Number of subsequent records: many Pearce: 5 localities Latest record: 2018 First subsequent record: 1953 Number of subsequent records: many Harpalus (Harpalus) pumilus Sturm, 1818 Latest record: 2013 Pearce: 2 localities First subsequent record: 1990 Ophonus (Metophonus) cordatus Number of subsequent records: 1 (Duftschmid, 1812) Latest record: 1990 Pearce: 1 locality First subsequent record: 1953 Harpalus (Harpalus) rubripes Number of subsequent records: 2 (Duftschmid, 1812) Latest record: 1953 Pearce: widely distributed in south of county First subsequent record: 1950 Ophonus (Metophonus) laticollis Number of subsequent records: many Mannerheim, 1825 Latest record: 2019 Pearce: 4 localities No subsequent records Harpalus (Harpalus) rufipalpis Sturm, 1818 Ophonus (Metophonus) melletii (Heer, Pearce: no records 1837) First record: 1937 Pearce: 3 localities Number of records: many No subsequent records Latest record: 2020 Ophonus (Metophonus) parallelus Ophonus (Ophonus) sabulicola (Panzer, (Dejean, 1829) 1796) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1953 First subsequent record: 1947 Number of subsequent records: 1 Number of subsequent records: 1 Latest record: 1953 Latest record: 1947

Ophonus (Metophonus) puncticeps Acupalpus brunnipes (Sturm, 1825) Stephens, 1828 Pearce: 2 localities Pearce: 2 localities First subsequent record: 1953 First subsequent record: 1936 Number of subsequent records: many Number of subsequent records: many Latest record: 2017 Latest record: 2019 Acupalpus dubius Schilsky, 1888 Ophonus (Metophonus) puncticollis Pearce: 4 localities (Paykull, 1798) First subsequent record: 1953 Pearce: 6 localities Number of subsequent records: many First subsequent record: 1936 Latest record: 2020 Number of subsequent records: 1 Latest record: 1936 Acupalpus exiguus Dejean, 1829 Pearce: 6 localities Ophonus (Metophonus) rufibarbis First subsequent record: 1994 Number of subsequent records: many (Fabricius, 1792) Latest record: 2019 Pearce: 2 localities First subsequent record: 1982 Number of subsequent records: many Acupalpus flavicollis (Sturm, 1825) Latest record: 2014 Pearce: 1 locality (1886) First subsequent record: 1995 Number of subsequent records: 8 Ophonus (Metophonus) rupicola (Sturm, Latest record: 2018 1818) Pearce: 5 localities First subsequent record: 1953 Acupalpus meridianus (Linnaeus, 1760) Number of subsequent records: 7 Pearce: 3 localities Latest record: 2005 First subsequent record: 1954 Number of subsequent records: many Latest record: 2013 Ophonus (Metophonus) schaubergerianus (Puel, 1937) Acupalpus parvulus (Sturm, 1825) Pearce: 1 locality Pearce: 3 localities First subsequent record: 2005 First subsequent record: 1953 Number of subsequent records: 1 Number of subsequent records: many Latest record: 2005 Latest record: 2020

Ophonus (Metophonus) subsinuatus Rey, Anthracus consputus (Duftschmid, 1812) 1886 Pearce: 3 localities Pearce: no records First subsequent record: 1979 First record: 1886 Number of subsequent records: many Number of records: 2 Latest record: 2020 Latest record: 1886 Bradycellus caucasicus (Chaudoir, 1846) Ophonus (Ophonus) ardosiacus (Lutshnik, Pearce: no records 1922) First record: 1977 Pearce: 5 localities Number of records: 4 First subsequent record: 1935 Latest record: 2009 Number of subsequent records: many Latest record: 2020 Bradycellus distinctus (Dejean, 1829) Stenolophus skrimshiranus Stephens, 1828 Pearce: 3 localities Pearce: no records No subsequent records First record: 1990 Number of records: many Bradycellus harpalinus (Audinet-Serville, Latest record: 2011 1821) Pearce: 3 localities Stenolophus teutonus (Schrank, 1781) First subsequent record: 1938 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1939 Latest record: 2019 Number of subsequent records: many Latest record: 2014 Bradycellus ruficollis (Stephens, 1828) Pearce: 4 localities Platyderus depressus (Audinet-Serville, First subsequent record: 1938 1821) Number of subsequent records: many Pearce: 4 localities Latest record: 2020 First subsequent record: 1954 Number of subsequent records: many Bradycellus sharpi Joy, 1912 Latest record: 2019 Pearce: 1 locality First subsequent record: 1938 Calathus (Amphyginus) rotundicollis Number of subsequent records: many Dejean, 1828 Latest record: 2020 Pearce: 3 localities First subsequent record: 1975 Bradycellus verbasci (Duftschmid, 1812) Number of subsequent records: many Pearce: 5 localities Latest record: 2017 First subsequent record: 1962 Number of subsequent records: many Calathus (Calathus) fuscipes (Goeze, Latest record: 2019 1777) Pearce: abundant and widely distributed Dicheirotrichus (Dicheirotrichus) gustavii First subsequent record: 1935 Crotch, 1871 Number of subsequent records: many Pearce: 5 localities Latest record: 2020 First subsequent record: 1935 Number of subsequent records: many Calathus (Neocalathus) ambiguus Latest record: 2015 (Paykull, 1790) Pearce: 4 localities Dicheirotrichus (Dicheirotrichus) First subsequent record: 1953 obsoletus (Dejean, 1829) Number of subsequent records: 1 Pearce: 5 localities Latest record: 1953 First subsequent record: 1968 Number of subsequent records: many Calathus (Neocalathus) cinctus Latest record: 2019 Motschulsky, 1850 Pearce: no records Dicheirotrichus (Trichocellus) placidus First record: 1935 (Gyllenhal, 1827) Number of records: many Pearce: 1 locality Latest record: 2020 First subsequent record: 1963 Number of subsequent records: many Calathus (Neocalathus) erratus (Sahlberg, Latest record: 2020 C.R., 1827) Pearce: 5 localities Stenolophus mixtus (Herbst, 1784) First subsequent record: 1935 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1940 Latest record: 2018 Number of subsequent records: many Latest record: 2020

Calathus (Neocalathus) melanocephalus Agonum (Agonum) nigrum Dejean, 1828 (Linnaeus, 1758) Pearce: 1 locality Pearce: abundant and widely distributed (but may First subsequent record: 1979 include cinctus) Number of subsequent records: 9 First subsequent record: 1982 Latest record: 2018 Number of subsequent records: many Latest record: 2019 Agonum (Europhilus) fuliginosum (Panzer, 1809) Calathus (Neocalathus) micropterus Pearce: 5 localities (Duftschmid, 1812) First subsequent record: 1936 Pearce: no records Number of subsequent records: many First record: 1999 Latest record: 2020 Number of records: 1 Latest record: 1999 Agonum (Europhilus) gracile Sturm, 1824 Pearce: 4 localities Calathus (Neocalathus) mollis (Marsham, First subsequent record: 1955 1802) Number of subsequent records: many Pearce: 4 localities Latest record: 2019 First subsequent record: 1979 Number of subsequent records: many Agonum (Europhilus) micans (Nicolai, Latest record: 2018 1822) Pearce: 4 localities Laemostenus (Laemostenus) complanatus First subsequent record: 1938 (Dejean, 1828) Number of subsequent records: many Pearce: 1 locality Latest record: 2019 No subsequent records Agonum (Europhilus) piceum (Linnaeus, Laemostenus (Pristonychus) terricola 1758) (Herbst, 1784) Pearce: 1 locality (1919) Pearce: 4 localities First subsequent record: 1968 First subsequent record: 1939 Number of subsequent records: many Number of subsequent records: many Latest record: 2010 Latest record: 2019 Agonum (Europhilus) scitulum Dejean, Synuchus vivalis (Illiger, 1798) 1828 Pearce: 4 localities Pearce: no records First subsequent record: 1943 First record: 1955 Number of subsequent records: many Number of records: 1 Latest record: 2011 Latest record: 1955

Agonum (Agonum) marginatum (Linnaeus, Agonum (Europhilus) thoreyi Dejean, 1758) 1828 Pearce: 8 localities Pearce: 1 locality (1924) First subsequent record: 1939 First subsequent record: 1955 Number of subsequent records: many Number of subsequent records: many Latest record: 2017 Latest record: 2019

Agonum (Agonum) muelleri (Herbst, 1784) Agonum (Olisares) emarginatum Pearce: generally common and widely distributed (Gyllenhal, 1827) First subsequent record: 1940 Pearce: generally common Number of subsequent records: many First subsequent record: 1940 Latest record: 2019 Number of subsequent records: many Latest record: 2017 Agonum (Olisares) ericeti (Panzer, 1809) Sericoda quadripunctata (De Geer, 1774) Pearce: no records Pearce: 1 locality First record: 1963 No subsequent records Number of records: 4 Latest record: 2006 Perigona nigriceps (Dejean, 1831) Pearce: no records Agonum (Olisares) sexpunctatum First record: 1990 (Linnaeus, 1758) Number of records: 3 Pearce: 1 locality Latest record: 2003 First subsequent record: 2002 Number of subsequent records: many Odacantha melanura (Linnaeus, 1767) Latest record: 2017 Pearce: no records First record: 2005 Agonum (Olisares) versutum Sturm, 1824 Number of records: 7 Pearce: 1 locality (1919) Latest record: 2019 First subsequent record: 2013 Number of subsequent records: 1 Masoreus wetterhallii (Gyllenhal, 1813) Latest record: 2013 Pearce: 2 localities First subsequent record: 1997 Agonum (Olisares) viduum (Panzer, 1796) Number of subsequent records: 2 Pearce: 3 localities Latest record: 2014 First subsequent record: 1953 Number of subsequent records: many Cymindis axillaris (Fabricius, 1794) Latest record: 2019 Pearce: 5 localities First subsequent record: 1939 Anchomenus dorsalis (Pontoppidan, 1763) Number of subsequent records: 4 Pearce: abundant and widely distributed Latest record: 2018 First subsequent record: 1940 Number of subsequent records: many Demetrias (Aetophorus) imperialis Latest record: 2019 (Germar, 1823) Pearce: no records Limodromus assimilis (Paykull, 1790) First record: 1962 [2012 Platynus assimilis] Number of records: many Pearce: 1 locality (1896) Latest record: 2018 First subsequent record: 1954 Number of subsequent records: many Demetrias (Demetrias) atricapillus Latest record: 2019 (Linnaeus, 1758) Pearce: abundant and widely distributed Olisthopus rotundatus (Paykull, 1790) First subsequent record: 1936 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1936 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Demetrias (Demetrias) monostigma Leach in Samouelle, 1819 Oxypselaphus obscurus (Herbst, 1784) Pearce: 1 locality Pearce: 5 localities No subsequent records First subsequent record: 1935 Number of subsequent records: many Latest record: 2020 Calodromius spilotus (Illiger, 1798) Pearce: 5 localities First subsequent record: 1939 Paranchus albipes (Fabricius, 1796) Number of subsequent records: many Pearce: generally common and widely distributed Latest record: 2019 First subsequent record: 1936 Number of subsequent records: many Latest record: 2020

Dromius agilis (Fabricius, 1787) Philorhizus vectensis (Rye, 1873) Pearce: 3 localities Pearce: 3 localities First subsequent record: 1954 First subsequent record: 1964 Number of subsequent records: 4 Number of subsequent records: many Latest record: 1988 Latest record: 2016

Dromius meridionalis Dejean, 1825 Syntomus foveatus (Geoffroy in Fourcroy, Pearce: 3 localities 1785) First subsequent record: 1934 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1935 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Dromius quadrimaculatus (Linnaeus, 1758) Syntomus obscuroguttatus (Duftschmid, Pearce: 6 localities 1812) First subsequent record: 1943 Pearce: 8 localities Number of subsequent records: many First subsequent record: 1953 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Lionychus quadrillum (Duftschmid, 1812) Pearce: 2 localities Syntomus truncatellus (Linnaeus, 1760) First subsequent record: 1953 Pearce: 3 localities Number of subsequent records: 2 First subsequent record: 1979 Latest record: 2006 Number of subsequent records: 8 Latest record: 2016 Microlestes maurus (Sturm, 1827) Pearce: 10 localities Lebia (Lamprias) chlorocephala First subsequent record: 1953 (Hoffmann, J., 1803) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1950 Number of subsequent records: 4 Microlestes minutulus (Goeze, 1777) Latest record: 2006 Pearce: no records First record: 2013 Lebia (Lamprias) cyanocephala Number of records: many Latest record: 2020 (Linnaeus, 1758) Pearce: 1 locality No subsequent records Paradromius linearis (Olivier, 1795) Pearce: 10 localities First subsequent record: 1936 Lebia (Lebia) cruxminor (Linnaeus, 1758) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2009 Number of records: 2 Latest record: 2015 Philorhizus melanocephalus (Dejean, 1825) Drypta dentata (Rossi, 1790) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1936 First subsequent record: 1933 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2011

Philorhizus notatus (Stephens, 1827) Polistichus connexus (Geoffroy in Pearce: 4 localities First subsequent record: 1983 Fourcroy, 1785) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1995 Number of subsequent records: 3 Latest record: 2020 8. Family HELOPHORIDAE Leach, Helophorus (Rhopalohelophorus) dorsalis 1815 (Marsham, 1802) Pearce: 2 localities Helophorus (Empleurus) nubilus First subsequent record: 2014 Number of subsequent records: 1 Fabricius, 1777 Latest record: 2014 Pearce: 8 localities First subsequent record: 1937 Number of subsequent records: 4 Helophorus (Rhopalohelophorus) flavipes Latest record: 1946 Fabricius, 1792 Pearce: generally common Helophorus (Empleurus) porculus Bedel, First subsequent record: 1947 Number of subsequent records: many 1881 Latest record: 2019 Pearce: 1 locality (1904) No subsequent records Helophorus (Rhopalohelophorus) Helophorus (Empleurus) rufipes (Bosc fulgidicollis Motschulsky, 1860 Pearce: 1 locality d’Antic, 1791) First subsequent record: 1947 Pearce: 3 localities Number of subsequent records: 3 First subsequent record: 1990 Latest record: 2008 Number of subsequent records: 6 Latest record: 2013 Helophorus (Rhopalohelophorus) Helophorus (Helophorus) aequalis granularis (Linnaeus, 1760) Pearce: 3 localities Thomson, C.G., 1868 First subsequent record: 1986 Pearce: 6 localities (but probably including Number of subsequent records: many grandis) Latest record: 2019 First subsequent record: 1939 Number of subsequent records: many Latest record: 2019 Helophorus (Rhopalohelophorus) griseus Herbst, 1793 Helophorus (Helophorus) grandis Illiger, Pearce: 3 localities First subsequent record: 1983 1798 Number of subsequent records: many Pearce: no records (but see aequalis above) Latest record: 2012 First record: 1979 Number of records: many Latest record: 2020 Helophorus (Rhopalohelophorus) laticollis Thomson, C.G., 1853 Helophorus (Rhopalohelophorus) Pearce: 1 locality (1892) No subsequent records arvernicus Mulsant, 1846 Pearce: no records First record: 1995 Helophorus (Rhopalohelophorus) Number of records: 3 longitarsis Wollaston, 1864 Latest record: 2017 Pearce: no records First record: 2002 Helophorus (Rhopalohelophorus) Number of records: 3 brevipalpis Bedel, 1881 Latest record: 2012 Pearce: 7 localities First subsequent record: 1934 Helophorus (Rhopalohelophorus) minutus Number of subsequent records: many Fabricius, 1775 Latest record: 2020 Pearce: no records First record: 1947 Number of records: many Latest record: 2020

Helophorus (Rhopalohelophorus) nanus Berosus (Berosus) luridus (Linnaeus, Sturm, 1836 1760) Pearce: no records Pearce: 3 localities First record: 1933 First subsequent record: 1937 Number of records: 2 Number of subsequent records: many Latest record: 2020 Latest record: 2014

Helophorus (Rhopalohelophorus) Berosus (Berosus) signaticollis obscurus Mulsant, 1844 (Charpentier, 1825) Pearce: no records Pearce: 1 locality (1909, 1925) First record: 1979 First subsequent record: 1937 Number of records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Helophorus (Trichohelophorus) alternans Berosus (Enoplurus) fulvus Kuwert, 1888 Gené, 1836 Pearce: 1 locality (1896) Pearce: 1 locality No subsequent records First subsequent record: 1937 Number of subsequent records: 3 Laccobius (Dimorpholaccobius) atratus Latest record: 2019 Rottenberg, 1874 Pearce: no records 9. Family GEORISSIDAE Laporte, First record: 1947 1840 Number of records: many Latest record: 2019 Georissus crenulatus (Rossi, 1794) Pearce: 4 localities Laccobius (Dimorpholaccobius) First subsequent record: 1988 bipunctatus (Fabricius, 1775) Number of subsequent records: many Pearce: 5 localities Latest record: 2016 First subsequent record: 1934 Number of subsequent records: many 10. Family HYDROCHIDAE Thomson, Latest record: 2020 C.G., 1859 Laccobius (Dimorpholaccobius) sinuatus Hydrochus angustatus Germar, 1824 Motschulsky, 1849 Pearce: 7 localities Pearce: 3 localities First subsequent record: 1937 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2011

Hydrochus elongatus (Schaller, 1783) Laccobius (Dimorpholaccobius) striatulus Pearce: 1 locality (Fabricius, 1801) No subsequent records Pearce: 1 locality First subsequent record: 1933 Number of subsequent records: many 12. Family HYDROPHILIDAE Latest record: 2019 Latreille, 1802 Laccobius (Dimorpholaccobius) ytenensis Berosus (Berosus) affinis Brullé, 1835 Sharp, 1910 Pearce: 1 locality Pearce: 1 locality First subsequent record: 1964 First subsequent record: 1952 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2011 Laccobius (Laccobius) colon (Stephens, Anacaena globulus (Paykull, 1798) 1829) Pearce: 6 localities Pearce: 1 locality First subsequent record: 1933 First subsequent record: 1993 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2019 Latest record: 2019 Anacaena limbata (Fabricius, 1792) Laccobius (Laccobius) minutus (Linnaeus, Pearce: 3 localities 1758) First subsequent record: 1963 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1994 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Anacaena lutescens (Stephens, 1829) Pearce: generally common Paracymus scutellaris (Rosenhauer, 1856) First subsequent record: 1986 Pearce: 1 locality Number of subsequent records: many First subsequent record: 2006 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Cymbiodyta marginella (Fabricius, 1792) Pearce: 2 localities (1899, 1922) Hydrophilus piceus (Linnaeus, 1758) First subsequent record: 1947 Pearce: 1 locality (1821) Number of subsequent records: many No subsequent records Latest record: 2020

Hydrobius fuscipes (Linnaeus, 1758) Enochrus (Enochrus) melanocephalus Pearce: generally common (Olivier, 1793) First subsequent record: 1933 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1989 Latest record: 2020 Number of subsequent records: 3 Latest record: 2020 Limnoxenus niger (Gmelin, 1790) Pearce: 1 locality Enochrus (Lumetus) bicolor (Fabricius, First subsequent record: 1964 1792) Number of subsequent records: 1 Pearce: 2 localities Latest record: 1964 First subsequent record: 1981 Number of subsequent records: many Chaetarthria seminulum (Herbst, 1797) Latest record: 2014 Pearce: 1 locality (sensu lato) First subsequent record: 2005 Enochrus (Lumetus) fuscipennis Number of subsequent records: many (Thomson, C.G., 1884) Latest record: 2017 Pearce: no records First record: 2002 Chaetarthria simillima Vorst & Cuppen, Number of records: 4 2003 Latest record: 2009 Pearce: no records First record: 1997 Enochrus (Lumetus) halophilus (Bedel, Number of records: many 1878) Latest record: 2019 Pearce: no records First record: 2000 Anacaena bipustulata (Marsham, 1802) Number of records: 7 Pearce: 3 localities Latest record: 2006 First subsequent record: 1934 Number of subsequent records: many Latest record: 2019 Enochrus (Lumetus) ochropterus Coelostoma orbiculare (Fabricius, 1775) (Marsham, 1802) Pearce: 4 localities Pearce: 2 localities First subsequent record: 1937 First subsequent record: 1937 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019 Dactylosternum abdominale (Fabricius, Enochrus (Lumetus) quadripunctatus 1792) (Herbst, 1797) Pearce: no records Pearce: no records First record: 2003 First record: 1963 Number of records: 4 Number of records: 9 Latest record: 2006 Latest record: 2007 Sphaeridium bipustulatum Fabricius, 1781 Enochrus (Lumetus) testaceus (Fabricius, Pearce: 7 localities 1801) First subsequent record: 1993 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1947 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Sphaeridium lunatum Fabricius, 1792 Pearce: no records Enochrus (Methydrus) affinis (Thunberg, First record: 1969 Number of records: many 1794) Latest record: 2019 Pearce: 2 localities First subsequent record: 1937 Number of subsequent records: many Sphaeridium marginatum Fabricius, 1787 Latest record: 2019 Pearce: no records First record: 1997 Number of records: 8 Enochrus (Methydrus) coarctatus Latest record: 2018 (Gredler, 1863) Pearce: 2 localities First subsequent record: 1937 Sphaeridium scarabaeoides (Linnaeus, Number of subsequent records: many 1758) Latest record: 2015 Pearce: generally common and widely distributed First subsequent record: 1975 Number of subsequent records: many Enochrus (Methydrus) nigritus (Sharp, Latest record: 2019 1873) Pearce: no records First record: 2009 Cercyon (Cercyon) convexiusculus Number of records: 1 Stephens, 1829 Latest record: 2009 Pearce: 2 localities First subsequent record: 1947 Number of subsequent records: many Helochares lividus (Forster, 1771) Latest record: 2020 Pearce: 3 localities First subsequent record: 1963 Number of subsequent records: many Cercyon (Cercyon) depressus Stephens, Latest record: 2019 1829 Pearce: 7 localities Helochares punctatus Sharp, 1869 First subsequent record: 1968 Pearce: 10 localities Number of subsequent records: many First subsequent record: 1935 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Cercyon (Cercyon) granarius Erichson, Cercyon (Cercyon) obsoletus (Gyllenhal, 1837 1808) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1998 First subsequent record: 1997 Number of subsequent records: 2 Number of subsequent records: 6 Latest record: 2013 Latest record: 2019

Cercyon (Cercyon) haemorrhoidalis Cercyon (Cercyon) pygmaeus (Illiger, (Fabricius, 1775) 1801) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1987 First subsequent record: 1939 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2018

Cercyon (Cercyon) impressus (Sturm, Cercyon (Cercyon) quisquilius (Linnaeus, 1807) 1760) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1975 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2018

Cercyon (Cercyon) lateralis (Marsham, Cercyon (Cercyon) sternalis Sharp, 1918 1802) Pearce: no records Pearce: 3 localities First record: 1998 First subsequent record: 1987 Number of records: many Number of subsequent records: many Latest record: 2014 Latest record: 2018 Cercyon (Cercyon) terminatus (Marsham, Cercyon (Cercyon) littoralis (Gyllenhal, 1802) 1808) Pearce: 1 locality Pearce: 7 localities First subsequent record: 1992 First subsequent record: 1947 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2006 Latest record: 2019 Cercyon (Cercyon) tristis (Illiger, 1801) Cercyon (Cercyon) marinus Thomson, Pearce: 1 locality C.G., 1853 First subsequent record: 1987 Pearce: 3 localities Number of subsequent records: many First subsequent record: 2009 Latest record: 2019 Number of subsequent records: 8 Latest record: 2020 Cercyon (Cercyon) unipunctatus (Linnaeus, 1758) Cercyon (Cercyon) melanocephalus Pearce: 2 localities (Linnaeus, 1758) First subsequent record: 1975 Pearce: 5 localities Number of subsequent records: 6 First subsequent record: 1975 Latest record: 2018 Number of subsequent records: many Latest record: 2019 Cercyon (Dicyrtocercyon) ustulatus (Preyssier, 1790) Cercyon (Cercyon) nigriceps (Marsham, Pearce: 1 locality (1892) 1802) First subsequent record: 1982 Pearce: 1 locality (1910) Number of subsequent records: many First subsequent record: 1993 Latest record: 2019 Number of subsequent records: 4 Latest record: 2006 Cercyon (Paracercyon) analis (Paykull, 14. Family HISTERIDAE Gyllenhal, 1798) 1808 Pearce: 2 localities First subsequent record: 1993 Abraeus (Abraeus) perpusillus (Marsham, Number of subsequent records: many Latest record: 2020 1802) Pearce: 1 locality First subsequent record: 1993 Cercyon (Paracycreon) laminatus Sharp, Number of subsequent records: 9 1873 Latest record: 2019 Pearce: no records First record: 1997 Abraeus (Postabraeus) granulum Number of records: 5 Latest record: 2020 Erichson, 1839 Pearce: no records First record: 2019 Cryptopleurum crenatum (Kugelann, Number of records: 1 1794) Latest record: 2019 Pearce: no records First record: 1994 Plegaderus dissectus Erichson, 1839 Number of records: 5 Pearce: no records Latest record: 2014 First record: 2000 Number of records: many Cryptopleurum minutum (Fabricius, 1775) Latest record: 2020 Pearce: 7 localities First subsequent record: 1975 Plegaderus vulneratus (Panzer, 1797) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1992 Number of records: 3 Cryptopleurum subtile Sharp, 1884 Latest record: 2016 Pearce: no records First record: 2003 Acritus (Acritus) nigricornis (Hoffmann, Number of records: 1 Latest record: 2003 J., 1803) Pearce: no records First record: 1969 Megasternum concinnum (Marsham, Number of records: many 1802) Latest record: 2020 Pearce: 5 localities First subsequent record: 1963 Aeletes atomarius (Aubé, 1842) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2004 Number of records: 2 13. Family SPHAERITIDAE Shuckard, Latest record: 2019 1839 Halacritus punctum (Aubé, 1842) Sphaerites glabratus (Fabricius, 1792) Pearce: 2 localities Pearce: no records First subsequent record: 2016 First record: 2000 Number of subsequent records: 1 Number of records: 1 Latest record: 2016 Latest record: 2000 Saprinus aeneus (Fabricius, 1775) Pearce: 2 localities No subsequent records Saprinus semistriatus (Scriba, 1790) Kissister minimus (Laporte, 1840) Pearce: 4 localities Pearce: 4 localities First subsequent record: 1939 First subsequent record: 1952 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019

Saprinus virescens (Paykull, 1798) Carcinops pumilio (Erichson, 1834) Pearce: 3 localities Pearce: no records First subsequent record: 1943 First record: 2000 Number of subsequent records: 6 Number of records: 4 Latest record: 2019 Latest record: 2008

Hypocaccus (Baeckmanniolus) dimidiatus Paromalus flavicornis (Herbst, 1791) (Illiger, 1807) Pearce: no records Pearce: 3 localities First record: 1953 First subsequent record: 1968 Number of records: many Number of subsequent records: 5 Latest record: 2020 Latest record: 2019 Onthophilus striatus (Forster, 1771) Hypocaccus (Hypocaccus) rugifrons Pearce: 5 localities (Paykull, 1798) First subsequent record: 1971 Pearce: 1 locality Number of subsequent records: 4 No subsequent records Latest record: 2007

Gnathoncus communis (Marseul, 1862) Margarinotus (Paralister) neglectus Pearce: no records (Germar, 1813) First record: 1987 Pearce: no records Number of records: 1 First record: 1993 Latest record: 1987 Number of records: 4 Latest record: 2018 Gnathoncus nannetensis (Marseul, 1862) Pearce: 3 localities Margarinotus (Paralister) purpurascens First subsequent record: 1987 (Herbst, 1791) Number of subsequent records: 4 Pearce: 1 locality Latest record: 2020 First subsequent record: 2001 Number of subsequent records: 7 Gnathoncus rotundatus (Kugelann, 1792) Latest record: 2019 Pearce: no records First record: 1954 Margarinotus (Paralister) ventralis Number of records: 1 (Marseul, 1854) Latest record: 1954 Pearce: 3 localities First subsequent record: 1963 Myrmetes paykulli Kanaar, 1979 Number of subsequent records: many Pearce: 2 localities Latest record: 2019 No subsequent records Margarinotus (Promethister) marginatus Dendrophilus punctatus (Herbst, 1791) (Erichson, 1834) Pearce: no records Pearce: 2 localities First record: 2019 First subsequent record: 2002 Number of records: 1 Number of subsequent records: 1 Latest record: 2019 Latest record: 2002 Margarinotus (Ptomister) brunneus Hydraena gracilis Germar, 1824 (Fabricius, 1775) Pearce: no records Pearce: generally common and widely distributed First record: 1987 First subsequent record: 1943 Number of records: 8 Number of subsequent records: 7 Latest record: 2011 Latest record: 2020 Hydraena nigrita Germar, 1824 Margarinotus (Ptomister) merdarius Pearce: 1 locality (Hoffmann, J., 1803) First subsequent record: 1981 Pearce: no records Number of subsequent records: 5 First record: 1993 Latest record: 2019 Number of records: 5 Latest record: 2020 Hydraena pulchella Germar, 1824 Pearce: no records Margarinotus (Ptomister) striola First record: 1830 Number of records: 2 (Sahlberg, C.R., 1819) Latest record: 1861 Pearce: 2 localities First subsequent record: 1991 Number of subsequent records: 4 Hydraena riparia Kugelann, 1794 Latest record: 2019 Pearce: 5 localities First subsequent record: 1933 Number of subsequent records: many Hister quadrimaculatus Linnaeus, 1758 Latest record: 2020 Pearce: 1 locality (1901) No subsequent records Hydraena rufipes Curtis, 1830 Pearce: 1 locality Hister unicolor Linnaeus, 1758 First subsequent record: 1998 Pearce: 3 localities Number of subsequent records: 1 First subsequent record: 1939 Latest record: 1998 Number of subsequent records: many Latest record: 2018 Hydraena testacea Curtis, 1830 Pearce: 1 locality (1815) Atholus bimaculatus (Linnaeus, 1758) First subsequent record: 1979 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1971 Latest record: 2018 Number of subsequent records: 9 Latest record: 2013 Limnebius nitidus (Marsham, 1802) Pearce: 2 localities Atholus duodecimstriatus (Schrank, 1781) First subsequent record: 1990 Pearce: 4 localities Number of subsequent records: many First subsequent record: 2003 Latest record: 2017 Number of subsequent records: 6 Latest record: 2019 Limnebius papposus Mulsant, 1844 Pearce: 5 localities 15. Family HYDRAENIDAE Mulsant, First subsequent record: 1938 1844 Number of subsequent records: 4 Latest record: 2018 Hydraena britteni Joy, 1907 Pearce: no records Limnebius truncatellus (Thunberg, 1794) First record: 1998 Pearce: 8 localities Number of records: 2 First subsequent record: 1980 Latest record: 2017 Number of subsequent records: many Latest record: 2019 Ochthebius (Asiobates) auriculatus Rey, Ochthebius (Ochthebius) poweri Rye, 1886 1869 Pearce: no records Pearce: 1 locality First record: 2005 First subsequent record: 1971 Number of records: 3 Number of subsequent records: many Latest record: 2011 Latest record: 2019

Ochthebius (Asiobates) bicolon Germar, Ochthebius (Ochthebius) punctatus 1824 Stephens, 1829 Pearce: 2 localities Pearce: no records First subsequent record: 2002 First record: 1935 Number of subsequent records: 5 Number of records: many Latest record: 2011 Latest record: 2016

Ochthebius (Asiobates) dilatatus Stephens, Ochthebius (Ochthebius) pusillus 1829 Stephens, 1835 Pearce: 4 localities Pearce: 1 locality First subsequent record: 1962 First subsequent record: 2005 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2020 Latest record: 2006

Ochthebius (Asiobates) minimus Ochthebius (Ochthebius) viridis Peyron, (Fabricius, 1792) 1858 Pearce: 2 localities Pearce: 2 localities First subsequent record: 1938 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2014

Ochthebius (Enicocerus) exsculptus 16. Family PTILIIDAE Erichson, 1845 Germar, 1824 Pearce: 1 locality Ptenidium (Gillmeisterium) nitidum (Heer, No subsequent records 1841) Pearce: 1 locality Ochthebius (Ochthebius) lejolisii Mulsant First subsequent record: 1979 & Rey, 1861 Number of subsequent records: 4 Pearce: no records Latest record: 2019 First record: 2008 Number of records: 5 Ptenidium (Gressnerium) gressneri Latest record: 2014 Erichson, 1845 Pearce: no records Ochthebius (Ochthebius) marinus First record: 2019 (Paykull, 1798) Number of records: 1 Pearce: 3 localities Latest record: 2019 First subsequent record: 1963 Number of subsequent records: many Ptenidium (Matthewsium) laevigatum Latest record: 2020 Erichson, 1845 Pearce: no records Ochthebius (Ochthebius) nanus Stephens, First record: 1995 1829 Number of records: 4 Pearce: 2 localities Latest record: 2019 First subsequent record: 2005 Number of subsequent records: 3 Latest record: 2011

Ptenidium (Ptenidium) fuscicorne Ptilium exaratum (Allibert, 1844) Erichson, 1845 Pearce: no records Pearce: no records First record: 2000 First record: 1990 Number of records: 1 Number of records: 2 Latest record: 2000 Latest record: 1995 Ptilium horioni Rosskothen, 1934 Ptenidium (Ptenidium) longicorne Fuss, Pearce: no records 1868 First record: 1997 Pearce: no records Number of records: 1 First record: 2004 Latest record: 1997 Number of records: 4 Latest record: 2017 Nephanes titan (Newman, 1834) Pearce: 1 locality Ptenidium (Ptenidium) myrmicophilum First subsequent record: 2003 Number of subsequent records: 3 (Motschulsky, 1845) [2012: Ptenidium Latest record: 2003 formicetorum] Pearce: 2 localities First subsequent record: 2003 Actinopteryx fucicola (Allibert, 1844) Number of subsequent records: 2 Pearce: 1 locality Latest record: 2006 No subsequent records

Ptenidium (Ptenidium) punctatum Acrotrichis (Acrotrichis) atomaria (De (Gyllenhal, 1827) Geer, 1774) Pearce: 5 localities Pearce: 4 localities First subsequent record: 2004 First subsequent record: 1975 Number of subsequent records: 7 Number of subsequent records: many Latest record: 2019 Latest record: 2014

Ptenidium (Ptenidium) pusillum Acrotrichis (Acrotrichis) brevipennis (Gyllenhal, 1808) (Erichson, 1845) Pearce: 4 localities Pearce: 1 locality First subsequent record: 1975 No subsequent records Number of subsequent records: many Latest record: 2020 Acrotrichis (Acrotrichis) cognata (Matthews, A., 1877) Ptenidium (Wankowizium) intermedium Pearce: no records Wankowicz, 1869 First record: 1975 Pearce: no records Number of records: 1 First record: 1995 Latest record: 1975 Number of records: 3 Latest record: 2008 Acrotrichis (Acrotrichis) danica Sundt, 1958 Ptiliolum spencei (Allibert, 1844) Pearce: no records Pearce: 1 locality First record: 2010 No subsequent records Number of records: 5 Latest record: 2011 Oligella foveolata (Allibert, 1844) Pearce: no records Acrotrichis (Acrotrichis) fascicularis First record: 1997 (Herbst, 1793) Number of records: 1 Pearce: 2 localities Latest record: 1997 First subsequent record: 1975 Number of subsequent records: many Latest record: 2013

Acrotrichis (Acrotrichis) henrici Acrotrichis (Ctenopteryx) grandicollis (Matthews, A., 1872) (Mannerheim, 1844) Pearce: no records Pearce: no records First record: 1994 First record: 1975 Number of records: 8 Number of records: 4 Latest record: 2004 Latest record: 2006

Acrotrichis (Acrotrichis) insularis Acrotrichis (Ctenopteryx) sanctaehelenae (Mäklin, 1852) Johnson, 1972 Pearce: no records Pearce: no records First record: 2013 First record: 2003 Number of records: 1 Number of records: 3 Latest record: 2013 Latest record: 2003

Acrotrichis (Acrotrichis) intermedia 17. Family LEIODIDAE Fleming, 1821 (Gillmeister, 1845) Pearce: 2 localities Hydnobius punctatus (Sturm, 1807) First subsequent record: 1975 Pearce: 2 localities Number of subsequent records: 7 No subsequent records Latest record: 2020 Triarthron maerkelii Schmidt, W.L.E., Acrotrichis (Acrotrichis) montandonii 1840 (Allibert, 1844) Pearce: 1 locality (1925) Pearce: no records No subsequent records First record: 1992 Number of records: 1 Latest record: 1992 Leiodes badia (Sturm, 1807) Pearce: 1 locality First subsequent record: 1998 Acrotrichis (Acrotrichis) rosskotheni Number of subsequent records: 2 Sundt, 1971 Latest record: 2008 Pearce: no records First record: 1979 Leiodes calcarata (Erichson, 1845) Number of records: 3 Pearce: 3 localities Latest record: 1994 First subsequent record: 1993 Number of subsequent records: 5 Acrotrichis (Acrotrichis) rugulosa Latest record: 2016 Rosskothen, 1935 Pearce: no records Leiodes ferruginea (Fabricius, 1787) First record: 1975 Pearce: 1 locality (1892) Number of records: 2 No subsequent records Latest record: 2015 Leiodes gyllenhalii Stephens, 1829 Acrotrichis (Acrotrichis) sericans (Heer, Pearce: 1 locality 1841) No subsequent records Pearce: 3 localities No subsequent records Leiodes litura Stephens, 1835 Pearce: 2 localities Acrotrichis (Acrotrichis) thoracica (Waltl, First subsequent record: 2019 1838) Number of subsequent records: 1 Pearce: no records Latest record: 2019 First record: 2003 Number of records: 1 Leiodes lunicollis (Rye, 1872) Latest record: 2003 Pearce: 1 locality No subsequent records

Leiodes nigrita (Schmidt, W.L.E., 1841) Agathidium (Agathidium) seminulum Pearce: 3 localities (Linnaeus, 1758) No subsequent records Pearce: 1 locality First subsequent record: 2013 Leiodes rufipennis (Paykull, 1798) Number of subsequent records: 3 Pearce: 3 localities Latest record: 2019 First subsequent record: 2008 Number of subsequent records: 7 Agathidium (Cyphoceble) nigrinum Sturm, Latest record: 2019 1807 Pearce: no records Liocyrtusa vittata (Curtis, 1840) First record: 2015 Pearce: 2 localities Number of records: 3 No subsequent records Latest record: 2017

Colenis immunda (Sturm, 1807) Agathidium (Neoceble) confusum Brisout Pearce: 1 locality de Barneville, C., 1863 First subsequent record: 1987 Pearce: no records Number of subsequent records: 1 First record: 2013 Latest record: 1987 Number of records: 2 Latest record: 2013 Anisotoma humeralis (Herbst, 1791) Pearce: 4 localities Agathidium (Neoceble) marginatum First subsequent record: 1936 Sturm, 1807 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1979 Number of records: 3 Anisotoma orbicularis (Herbst, 1791) Latest record: 2017 Pearce: 1 locality (1892) First subsequent record: 2000 Agathidium (Neoceble) nigripenne Number of subsequent records: 2 Latest record: 2004 (Fabricius, 1792) Pearce: 1 locality First subsequent record: 2007 Amphicyllis globus (Fabricius, 1792) Number of subsequent records: 1 Pearce: 2 localities Latest record: 2007 First subsequent record: 1977 Number of subsequent records: many Latest record: 2015 Agathidium (Neoceble) rotundatum (Gyllenhal, 1827) Pearce: no records Agathidium (Agathidium) atrum (Paykull, First record: 2008 1798) Number of records: 4 Pearce: 3 localities Latest record: 2008 First subsequent record: 1953 Number of subsequent records: 1 Latest record: 1953 Agathidium (Neoceble) varians Beck, 1817 Pearce: 2 localities Agathidium (Agathidium) laevigatum First subsequent record: 2014 Erichson, 1845 Number of subsequent records: 1 Pearce: 1 locality Latest record: 2014 First subsequent record: 1987 Number of subsequent records: many Latest record: 2019 Colon (Eurycolon) latum Kraatz, 1850 Pearce: 2 localities (1927, 1929) No subsequent records Colon (Myloechus) brunneum (Latreille, Choleva (Choleva) glauca Britten, 1918 1806) Pearce: no records Pearce: 2 localities First record: 2008 First subsequent record: 1993 Number of records: 2 Number of subsequent records: 3 Latest record: 2008 Latest record: 1993 Choleva (Choleva) jeanneli Britten, 1922 Nemadus colonoides (Kraatz, 1851) Pearce: no records Pearce: no records First record: 1979 First record: 2014 Number of records: 8 Number of records: 4 Latest record: 2007 Latest record: 2020 Choleva (Choleva) oblonga Latreille, 1806 Nargus (Demochrus) anisotomoides Pearce: no records (Spence, 1813) First record: 2016 Pearce: 3 localities Number of records: 1 First subsequent record: 2015 Latest record: 2016 Number of subsequent records: many Latest record: 2015 Choleva (Cholevopsis) spadicea (Sturm, 1839) Nargus (Demochrus) wilkini (Spence, Pearce: no records 1813) First record: 1994 Pearce: 4 localities Number of records: 4 First subsequent record: 1969 Latest record: 1999 Number of subsequent records: 3 Latest record: 2016 Sciodrepoides fumatus (Spence, 1813) Pearce: 1 locality Nargus (Nargus) velox (Spence, 1813) First subsequent record: 1980 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1978 Latest record: 2020 Number of subsequent records: many Latest record: 2018 Sciodrepoides watsoni (Spence, 1813) Pearce: 8 localities Choleva (Choleva) agilis (Illiger, 1798) First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: 8 Latest record: 2015 Catops chrysomeloides (Panzer, 1798) Pearce: 4 localities Choleva (Choleva) angustata (Fabricius, No subsequent records 1781) Pearce: 3 localities Catops coracinus Kellner, 1846 First subsequent record: 1987 Pearce: no records Number of subsequent records: 6 First record: 1952 Latest record: 2015 Number of records: many Latest record: 2016 Choleva (Choleva) cisteloides (Frölich, 1799) Catops fuliginosus Erichson, 1837 Pearce: 2 localities Pearce: no records No subsequent records First record: 1975 Number of records: many Latest record: 2008 Choleva (Choleva) fagniezi Jeannel, 1922 Pearce: no records First record: 2017 Number of records: 2 Latest record: 2019 Catops fuscus (Panzer, 1794) Ptomaphagus subvillosus (Goeze, 1777) Pearce: 2 localities Pearce: 10 localities First subsequent record: 1997 First subsequent record: 1975 Number of subsequent records: 7 Number of subsequent records: many Latest record: 2019 Latest record: 2019

Catops grandicollis Erichson, 1837 Leptinus testaceus Müller, P.W.J., 1817 Pearce: 4 localities Pearce: 1 locality (1893) First subsequent record: 1975 First subsequent record: 2000 Number of subsequent records: 4 Number of subsequent records: 1 Latest record: 2016 Latest record: 2000

Catops kirbyi (Spence, 1813) 18. Family SILPHIDAE Latreille, 1806 Pearce: 2 localities First subsequent record: 1999 Necrodes littoralis (Linnaeus, 1758) Number of subsequent records: 5 Pearce: 5 localities Latest record: 2019 First subsequent record: 1940 Number of subsequent records: many Catops morio (Fabricius, 1787) Latest record: 2018 Pearce: 4 localities First subsequent record: 1980 Thanatophilus rugosus (Linnaeus, 1758) Number of subsequent records: many Pearce: generally common and widely distributed Latest record: 2018 First subsequent record: 1997 Number of subsequent records: 4 Catops nigricans (Spence, 1813) Latest record: 2019 Pearce: 4 localities First subsequent record: 1968 Thanatophilus sinuatus (Fabricius, 1775) Number of subsequent records: many Pearce: 5 localities Latest record: 2020 First subsequent record: 1934 Number of subsequent records: many Catops tristis (Panzer, 1794) Latest record: 2003 Pearce: 7 localities First subsequent record: 1986 Oiceoptoma thoracicum (Linnaeus, 1758) Number of subsequent records: 7 Pearce: 9 localities Latest record: 2010 First subsequent record: 1992 Number of subsequent records: 9 Apocatops nigrita (Erichson, 1837) [2012: Latest record: 2011 Catops nigrita] Pearce: 2 localities Aclypea opaca (Linnaeus, 1758) First subsequent record: 1980 Pearce: 9 localities Number of subsequent records: many First subsequent record: 1975 Latest record: 2018 Number of subsequent records: 2 Latest record: 1980 Parabathyscia wollastoni (Janson, 1857) Pearce: no records Dendroxena quadrimaculata (Scopoli, First record: 2019 1771) Number of records: 2 Pearce: 3 localities Latest record: 2020 First subsequent record: 1934 Number of subsequent records: 2 Ptomaphagus sericatus (Chaudoir, 1845) Latest record: 1941 [2012: Ptomaphagus medius] Pearce: 1 locality First subsequent record: 1980 Number of subsequent records: 4 Latest record: 2017

Phosphuga atrata (Linnaeus, 1758) [2012: Nicrophorus vestigator Herschel, 1807 Silpha atrata] Pearce: 8 localities Pearce: generally common and widely distributed No subsequent records First subsequent record: 1934 Number of subsequent records: many 19. Family STAPHYLINIDAE Latreille, Latest record: 2020 1802

Ablattaria laevigata (Fabricius, 1775) Phyllodrepa floralis (Paykull, 1789) [2012: Silpha laevigata] Pearce: 2 localities Pearce: generally common First subsequent record: 1975 First subsequent record: 1939 Number of subsequent records: 8 Number of subsequent records: many Latest record: 2017 Latest record: 2020 Dropephylla ioptera (Stephens, 1832) Silpha obscura Linnaeus, 1758 Pearce: 1 locality Pearce: 2 localities First subsequent record: 2000 First subsequent record: 2012 Number of subsequent records: many Number of subsequent records: 3 Latest record: 2019 Latest record: 2016 Dropephylla koltzei Jászay & Hlavac, Silpha tristis Illiger, 1798 2006 Pearce: 5 localities Pearce: no records First subsequent record: 1952 First record: 2018 Number of subsequent records: many Number of records: 3 Latest record: 2019 Latest record: 2019

Nicrophorus humator (Gleditsch, 1767) Dropephylla vilis (Erichson, 1840) Pearce: generally common Pearce: 2 localities First subsequent record: 1936 First subsequent record: 1970 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2020 Latest record: 2019

Nicrophorus interruptus Stephens, 1830 Hapalaraea pygmaea (Paykull, 1800) Pearce: 1 locality Pearce: 2 localities First subsequent record: 1952 First subsequent record: 2020 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2017 Latest record: 2020

Nicrophorus investigator Zetterstedt, 1824 Acrolocha sulcula (Stephens, 1834) Pearce: 8 localities Pearce: no records First subsequent record: 1936 First record: 2015 Number of subsequent records: 7 Number of records: 1 Latest record: 2014 Latest record: 2015

Nicrophorus vespillo (Linnaeus, 1758) Omalium allardii Fairmaire & Brisout de Pearce: generally common and widely distributed First subsequent record: 1936 Barneville, C., 1859 Number of subsequent records: many Pearce: 2 localities Latest record: 2020 No subsequent records

Nicrophorus vespilloides Herbst, 1783 Omalium caesum Gravenhorst, 1806 Pearce: generally common Pearce: 3 localities First subsequent record: 1935 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2018

Omalium excavatum Stephens, 1834 Phloeostiba plana (Paykull, 1792) Pearce: 2 localities Pearce: 1 locality First subsequent record: 2001 First subsequent record: 2001 Number of subsequent records: 2 Number of subsequent records: 2 Latest record: 2020 Latest record: 2019

Omalium italicum Bernhauer, 1902 Xylodromus concinnus (Marsham, 1802) Pearce: no records Pearce: 3 localities First record: 2003 First subsequent record: 1969 Number of records: 9 Number of subsequent records: 6 Latest record: 2020 Latest record: 2006

Omalium laeviusculum Gyllenhal, 1827 Xylodromus depressus (Gravenhorst, Pearce: 4 localities 1802) First subsequent record: 1975 Pearce: no records Number of subsequent records: many First record: 2006 Latest record: 2019 Number of records: 1 Latest record: 2006 Omalium oxyacanthae Gravenhorst, 1806 Pearce: no records Eusphalerum luteum (Marsham, 1802) First record: 1977 Pearce: 4 localities Number of records: 4 First subsequent record: 1936 Latest record: 2010 Number of subsequent records: many Latest record: 2019 Omalium riparium Thomson, C.G., 1857 Pearce: 8 localities Eusphalerum minutum (Fabricius, 1792) First subsequent record: 2004 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1969 Latest record: 2019 Number of subsequent records: many Latest record: 2011 Omalium rivulare (Paykull, 1789) Pearce: generally common Eusphalerum primulae (Stephens, 1834) First subsequent record: 1969 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1963 Latest record: 2020 Number of subsequent records: 5 Latest record: 2018 Omalium rugatum Mulsant & Rey, 1880 Pearce: no records Eusphalerum sorbi (Gyllenhal, 1810) First record: 1969 Pearce: 3 localities Number of records: 5 First subsequent record: 1996 Latest record: 2019 Number of subsequent records: 3 Latest record: 2019 Phloeonomus punctipennis Thomson, C.G., 1867 Eusphalerum torquatum (Marsham, 1802) Pearce: 1 locality Pearce: 3 localities First subsequent record: 2006 No subsequent records Number of subsequent records: many Latest record: 2019 Anthobium atrocephalum (Gyllenhal, 1827) Phloeonomus pusillus (Gravenhorst, 1806) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1979 First subsequent record: 1991 Number of subsequent records: many Number of subsequent records: 4 Latest record: 2019 Latest record: 2007 Anthobium unicolor (Marsham, 1802) Micralymma marinum (Strøm, 1783) Pearce: 4 localities Pearce: 2 localities (1883, 1894) First subsequent record: 1979 First subsequent record: 1953 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 1953

Olophrum piceum (Gyllenhal, 1810) Philorinum sordidum (Stephens, 1834) Pearce: 5 localities Pearce: 3 localities First subsequent record: 1935 First subsequent record: 1982 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2016

Acidota crenata (Fabricius, 1792) Coryphium angusticolle Stephens, 1834 Pearce: 1 locality Pearce: 2 localities First subsequent record: 1979 First subsequent record: 1934 Number of subsequent records: 6 Number of subsequent records: 4 Latest record: 2016 Latest record: 2011

Acidota cruentata Mannerheim, 1830 Metopsia clypeata (Müller, P.W.J., 1821) Pearce: 1 locality Pearce: 3 localities First subsequent record: 1999 First subsequent record: 1979 Number of subsequent records: 1 Number of subsequent records: many Latest record: 1999 Latest record: 2020

Lesteva (Lesteva) hanseni Lohse, 1953 Megarthrus bellevoyei Saulcy, 1862 Pearce: 1 locality Pearce: 3 localities No subsequent records First subsequent record: 1969 Number of subsequent records: 6 Lesteva (Lesteva) longoelytrata (Goeze, Latest record: 2014 1777) Pearce: 3 localities Megarthrus denticollis (Beck, 1817) First subsequent record: 1963 Pearce: no records Number of subsequent records: many First record: 1989 Latest record: 2020 Number of records: 3 Latest record: 2017 Lesteva (Lesteva) pubescens Mannerheim, 1830 Megarthrus depressus (Paykull, 1789) Pearce: 1 locality Pearce: 1 locality First subsequent record: 1963 First subsequent record: 1975 Number of subsequent records: 5 Number of subsequent records: 7 Latest record: 2017 Latest record: 2008

Lesteva (Lestevidia) punctata Erichson, Megarthrus hemipterus (Illiger, 1794) Pearce: 1 locality 1839 No subsequent records Pearce: 2 localities First subsequent record: 2006 Number of subsequent records: 4 Megarthrus prosseni Schatzmayer, 1904 Latest record: 2019 Pearce: 3 localities First subsequent record: 1970 Number of subsequent records: 9 Lesteva (Lestevina) sicula Erichson, 1840 Latest record: 2018 Pearce: 3 localities First subsequent record: 1968 Number of subsequent records: many Latest record: 2019 Proteinus brachypterus (Fabricius, 1792) Amauronyx maerkelii (Aubé, 1844) Pearce: 3 localities Pearce: 3 localities First subsequent record: 1975 No subsequent records Number of subsequent records: many Latest record: 2020 Euplectus karstenii (Reichenbach, 1816) Pearce: 1 locality (1838) Proteinus laevigatus Hochhuth, 1872 First subsequent record: 2003 Pearce: 2 localities Number of subsequent records: many No subsequent records Latest record: 2019

Proteinus ovalis Stephens, 1834 Euplectus kirbii Denny, 1825 Pearce: 5 localities Pearce: no records First subsequent record: 1993 First record: 2006 Number of subsequent records: many Number of records: 4 Latest record: 2019 Latest record: 2019

Micropeplus fulvus Erichson, 1840 Euplectus mutator Fauvel, 1895 Pearce: 1 locality Pearce: no records First subsequent record: 1969 First record: 2020 Number of subsequent records: 8 Number of records: 1 Latest record: 2008 Latest record: 2020

Micropeplus porcatus (Paykull, 1789) Euplectus nanus (Reichenbach, 1816) Pearce: 1 locality Pearce: 1 locality No subsequent records No subsequent records

Micropeplus staphylinoides (Marsham, Euplectus piceus Motschulsky, 1835 1802) Pearce: 1 locality (1890) Pearce: 2 localities First subsequent record: 2018 First subsequent record: 1986 Number of subsequent records: 8 Number of subsequent records: many Latest record: 2019 Latest record: 2019 Euplectus sanguineus Denny, 1825 Micropeplus tesserula Curtis 1828 Pearce: 1 locality Pearce: “Dorset” (A. Ford) No subsequent records First subsequent record: 1986 Number of subsequent records: 1 Euplectus signatus (Reichenbach, 1816) Latest record: 1986 Pearce: 1 locality No subsequent records Bibloporus bicolor (Denny, 1825) Pearce: no records Rybaxis longicornis (Leach, 1817) First record: 2018 Pearce: 5 localities Number of records: many First subsequent record: 1989 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Bibloplectus ambiguus (Reichenbach, 1816) Brachygluta fossulata (Reichenbach, Pearce: 1 locality (1925) 1816) First subsequent record: 2014 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1979 Latest record: 2014 Number of subsequent records: many Latest record: 2020 Trichonyx sulcicollis (Reichenbach, 1816) Pearce: 1 locality (1921) No subsequent records

Brachygluta haematica (Reichenbach, Bryaxis curtisii (Leach, 1817) 1816) Pearce: 3 localities Pearce: 2 localities First subsequent record: 1986 First subsequent record: 1987 Number of subsequent records: 7 Number of subsequent records: 3 Latest record: 2019 Latest record: 2019 Bryaxis puncticollis (Denny, 1825) Brachygluta helferi (Schmidt-Göbel, Pearce: 2 localities 1836) No subsequent records Pearce: 3 localities First subsequent record: 1995 Tychus niger (Paykull, 1800) Number of subsequent records: many Pearce: 2 localities Latest record: 2018 First subsequent record: 1968 Number of subsequent records: 9 Brachygluta waterhousei (Rye, 1869) Latest record: 2019 Pearce: 3 localities First subsequent record: 1966 Pselaphaulax dresdensis Reitter, 1909 Number of subsequent records: 4 Pearce: 1 locality (1925) Latest record: 2020 No subsequent records

Reichenbachia juncorum (Leach, 1817) Pselaphus heisei Herbst, 1791 Pearce: 6 localities Pearce: 4 localities First subsequent record: 1935 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2020 Latest record: 2017

Fagniezia impressa (Panzer, 1803) Claviger testaceus Preyssler, 1790 Pearce: 3 localities Pearce: 5 localities First subsequent record: 1965 First subsequent record: 1936 Number of subsequent records: 2 Number of subsequent records: 2 Latest record: 2015 Latest record: 1991

Tychobythinus glabratus (Rye, 1870) Phloeocharis subtilissima Mannerheim, Pearce: 1 locality 1830 No subsequent records Pearce: 2 localities First subsequent record: 1968 Bythinus burrellii Denny, 1825 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2013 Number of records: 2 Sepedophilus bipunctatus (Gravenhorst, Latest record: 2015 1802) Pearce: 1 locality Bythinus macropalpus Aubé, 1833 First subsequent record: 2019 Pearce: 1 locality Number of subsequent records: 1 First subsequent record: 1987 Latest record: 2019 Number of subsequent records: 4 Latest record: 2016 Sepedophilus constans (Fowler, 1888) Pearce: no records Bryaxis bulbifer (Reichenbach, 1816) First record: 2008 Pearce: 3 localities Number of records: 5 First subsequent record: 1986 Latest record: 2018 Number of subsequent records: many Latest record: 2019 Sepedophilus immaculatus (Stephens, Tachyporus (Tachyporus) chrysomelinus 1832) (Linnaeus, 1758) Pearce: 1 locality (1893) Pearce: 5 localities (but including dispar) First subsequent record: 1979 First subsequent record: 1994 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2020

Sepedophilus littoreus (Linnaeus, 1758) Tachyporus (Tachyporus) dispar (Paykull, Pearce: 2 localities 1789) First subsequent record: 1993 Pearce: no records (but see chrysomelinus) Number of subsequent records: many First record: 1996 Latest record: 2018 Number of records: many Latest record: 2019 Sepedophilus lusitanicus Hammond, 1973 Pearce: no records Tachyporus (Tachyporus) formosus First record: 2002 Matthews, A.H., 1838 Number of records: 6 Pearce: 2 localities Latest record: 2019 First subsequent record: 1994 Number of subsequent records: many Sepedophilus marshami (Stephens, 1832) Latest record: 2020 Pearce: 6 localities First subsequent record: 1975 Tachyporus (Tachyporus) hypnorum Number of subsequent records: many Latest record: 2020 (Fabricius, 1775) Pearce: generally common and widely distributed First subsequent record: 1963 Sepedophilus nigripennis (Stephens, 1832) Number of subsequent records: many Pearce: 6 localities Latest record: 2020 First subsequent record: 1975 Number of subsequent records: many Latest record: 2020 Tachyporus (Tachyporus) obtusus (Linnaeus, 1767) Pearce: 7 localities Sepedophilus pedicularius (Gravenhorst, First subsequent record: 1979 1802) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1989 Number of subsequent records: 4 Latest record: 2007 Tachyporus (Tachyporus) pallidus Sharp, 1871 Pearce: 2 localities Tachyporus (Palporus) nitidulus First subsequent record: 1975 (Fabricius, 1781) Number of subsequent records: many Pearce: 5 localities Latest record: 2019 First subsequent record: 1968 Number of subsequent records: many Latest record: 2020 Tachyporus (Tachyporus) pusillus Gravenhorst, 1806 Pearce: 4 localities Tachyporus (Tachyporus) atriceps First subsequent record: 1979 Stephens, 1832 Number of subsequent records: many Pearce: 7 localities Latest record: 2019 First subsequent record: 1986 Number of subsequent records: many Latest record: 2015 Tachyporus (Tachyporus) solutus Erichson, 1839 Pearce: generally common First subsequent record: 1975 Number of subsequent records: many Latest record: 2019 Tachyporus (Tachyporus) tersus Erichson, Tachinus pallipes (Gravenhorst, 1806) 1839 Pearce: no records Pearce: 2 localities First record: 2000 First subsequent record: 1979 Number of records: 2 Number of subsequent records: 4 Latest record: 2015 Latest record: 2005 Tachinus proximus Kraatz, 1855 Tachyporus (Tachyporus) transversalis Pearce: no records Gravenhorst, 1806 First record: 1994 Pearce: 1 locality (1925) Number of records: 2 First subsequent record: 2014 Latest record: 1994 Number of subsequent records: 5 Latest record: 2019 Tachinus rufipennis Gyllenhal, 1810 Pearce: no records Lamprinodes saginatus (Gravenhorst, First record: 1909 Number of records: 1 1806) Latest record: 1909 Pearce: no records First record: 2019 Number of records: 1 Tachinus rufipes (Linnaeus, 1758) Latest record: 2019 Pearce: generally common and widely distributed First subsequent record: 1939 Number of subsequent records: many Tachinus bipustulatus (Fabricius, 1792) Latest record: 2020 Pearce: 1 locality No subsequent records Tachinus subterraneus (Linnaeus, 1758) Pearce: 3 localities Tachinus corticinus Gravenhorst, 1802 First subsequent record: 1987 Pearce: no records Number of subsequent records: 7 First record: 1986 Latest record: 2020 Number of records: 4 Latest record: 2011 Cilea silphoides (Linnaeus, 1767) Pearce: 3 localities Tachinus elongatus Gyllenhal, 1810 First subsequent record: 1964 Pearce: 1 locality Number of subsequent records: 4 No subsequent records Latest record: 2000

Tachinus humeralis Gravenhorst, 1802 Mycetoporus angularis Mulsant & Rey, Pearce: 2 localities First subsequent record: 1992 1853 Number of subsequent records: 5 Pearce: 1 locality Latest record: 2019 First subsequent record: 1979 Number of subsequent records: many Latest record: 2019 Tachinus laticollis Gravenhorst, 1802 Pearce: no records First record: 1968 Mycetoporus clavicornis (Stephens, 1832) Number of records: many Pearce: 2 localities Latest record: 2017 First subsequent record: 1979 Number of subsequent records: many Latest record: 2014 Tachinus lignorum (Linnaeus, 1758) Pearce: 1 locality No subsequent records Mycetoporus despectus Strand, A., 1830 Pearce: no records First record: 2005 Tachinus marginellus (Fabricius, 1781) Number of records: 1 Pearce: 3 localities Latest record: 2005 First subsequent record: 1979 Number of subsequent records: many Latest record: 2018 Mycetoporus lepidus (Gravenhorst, 1806) Lordithon thoracicus (Fabricius, 1777) Pearce: 4 localities Pearce: 6 localities First subsequent record: 1964 First subsequent record: 1968 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Mycetoporus longulus Mannerheim, 1830 Lordithon trinotatus (Erichson, 1839) Pearce: 2 localities Pearce: 3 localities First subsequent record: 2000 First subsequent record: 1970 Number of subsequent records: 3 Number of subsequent records: many Latest record: 2018 Latest record: 2020

Mycetoporus nigricollis Stephens, 1835 Bolitobius castaneus (Stephens, 1832) Pearce: 1 locality Pearce: 8 localities First subsequent record: 1979 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2007 Latest record: 2009

Mycetoporus punctus (Gravenhorst, 1806) Bolitobius cingulatus Mannerheim, 1830 Pearce: 2 localities Pearce: 5 localities First subsequent record: 2013 First subsequent record: 1979 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2015 Latest record: 2019

Mycetoporus rufescens (Stephens, 1832) Parabolitobius inclinans (Gravenhorst, Pearce: 1 locality 1806) First subsequent record: 1979 Pearce: 1 locality Number of subsequent records: 9 First subsequent record: 1979 Latest record: 2014 Number of subsequent records: 2 Latest record: 2004 Ischnosoma longicorne (Mäklin, 1847) Pearce: 1 locality (1919) Trichophya pilicornis (Gyllenhal, 1810) First subsequent record: 1986 Pearce: no records Number of subsequent records: 1 First record: 1979 Latest record: 1986 Number of records: 2 Latest record: 2019 Ischnosoma splendidum (Gravenhorst, 1806) Habrocerus capillaricornis (Gravenhorst, Pearce: 3 localities 1806) First subsequent record: 1986 Pearce: no records Number of subsequent records: many First record: 1978 Latest record: 2020 Number of records: many Latest record: 2019 Lordithon exoletus (Erichson, 1839) Pearce: 3 localities Gymnusa brevicollis (Paykull, 1800) First subsequent record: 1968 Pearce: 4 localities Number of subsequent records: 9 First subsequent record: 1979 Latest record: 2015 Number of subsequent records: many Latest record: 2014 Lordithon lunulatus (Linnaeus, 1760) Pearce: 3 localities Gymnusa variegata Kiesenwetter, 1845 First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: many No subsequent records Latest record: 2020 Deinopsis erosa (Stephens, 1832) Aleochara (Polystomota) grisea Kraatz, Pearce: 1 locality 1856 First subsequent record: 1953 Pearce: 5 localities Number of subsequent records: 9 First subsequent record: 1978 Latest record: 2017 Number of subsequent records: many Latest record: 2019 Aleochara (Aleochara) curtula (Goeze, 1777) Aleochara (Polystomota) punctatella Pearce: generally common Motschulsky, 1858 First subsequent record: 1933 Pearce: 3 localities Number of subsequent records: many First subsequent record: 2010 Latest record: 2019 Number of subsequent records: 4 Latest record: 2019 Aleochara (Aleochara) lata Gravenhorst, 1802 Aleochara (Xenochara) brevipennis Pearce: 10 localities Gravenhorst, 1806 First subsequent record: 1933 Pearce: 3 localities Number of subsequent records: 5 First subsequent record: 2004 Latest record: 2010 Number of subsequent records: 9 Latest record: 2017 Aleochara (Baryodma) intricata Mannerheim, 1830 Aleochara (Xenochara) cuniculorum Pearce: 3 localities Kraatz, 1858 First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: many No subsequent records Latest record: 2020 Aleochara (Xenochara) diversa (Sahlberg, Aleochara (Ceranota) ruficornis J., 1876) [2012: Aleochara kamila] Gravenhorst, 1802 Pearce: no records Pearce: 1 locality (1890) First record: 1975 No subsequent records Number of records: 1 Latest record: 1975 Aleochara (Coprochara) bilineata Gyllenhal, 1810 Aleochara (Xenochara) fumata Pearce: 1 locality Gravenhorst, 1802 First subsequent record: 2004 Pearce: 1 locality (1894) Number of subsequent records: 4 No subsequent records Latest record: 2019 Aleochara (Xenochara) funebris Aleochara (Coprochara) bipustulata Wollaston, 1864 (Linnaeus, 1760) Pearce: no records Pearce: 8 localities First record: 1975 First subsequent record: 1975 Number of records: 3 Number of subsequent records: many Latest record: 1975 Latest record: 2020 Aleochara (Xenochara) lanuginosa Aleochara (Emplenota) obscurella Gravenhorst, 1802 Gravenhorst, 1806 Pearce: 8 localities Pearce: 10 localities First subsequent record: 1935 First subsequent record: 1968 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019 Aleochara (Xenochara) moesta Phloeopora testacea (Mannerheim, 1830) Gravenhorst, 1802 Pearce: no records Pearce: 2 localities First record: 1970 No subsequent records Number of records: many Latest record: 2019 Aleochara (Xenochara) sanguinea (Linnaeus, 1758) Ilyobates bennetti Donisthorpe, 1914 Pearce: no records Pearce: no records First record: 2004 First record: 1994 Number of records: 1 Number of records: 1 Latest record: 2004 Latest record: 1994

Aleochara (Xenochara) spadicea Ilyobates nigricollis (Paykull, 1800) Pearce: 2 localities (Erichson, 1837) No subsequent records Pearce: 1 locality (1928) First subsequent record: 1995 Number of subsequent records: 3 Calodera aethiops (Gravenhorst, 1802) Latest record: 2015 Pearce: 3 localities No subsequent records Aleochara (Xenochara) sparsa Heer, 1839 Pearce: no records Calodera riparia Erichson, 1837 First record: 1975 Pearce: 1 locality Number of records: 4 First subsequent record: 1984 Latest record: 2004 Number of subsequent records: 7 Latest record: 2000 Aleochara (Xenochara) stichai Likovský, 1965 Calodera rubens Erichson, 1837 Pearce: no records Pearce: 1 locality First record: 1986 No subsequent records Number of records: 1 Latest record: 1986 Tetralaucopora longitarsis (Erichson, 1839) Aleochara (Xenochara) tristis Pearce: 1 locality Gravenhorst, 1806 First subsequent record: 1988 Pearce: 2 localities Number of subsequent records: 2 First subsequent record: 2006 Latest record: 2017 Number of subsequent records: 7 Latest record: 2019 Tetralaucopora rubicunda (Erichson, 1837) Tinotus morion (Gravenhorst, 1802) Pearce: 1 locality Pearce: 2 localities No subsequent records First subsequent record: 1975 Number of subsequent records: 6 Ocalea badia Erichson, 1837 Latest record: 2010 Pearce: no records First record: 2006 Amarochara umbrosa (Erichson, 1837) Number of records: 1 Pearce: 1 locality Latest record: 2006 No subsequent records Ocalea picata (Stephens, 1832) Phloeopora corticalis (Gravenhorst, 1802) Pearce: 1 locality Pearce: no records First subsequent record: 1986 First record: 2018 Number of subsequent records: 6 Number of records: 1 Latest record: 2019 Latest record: 2018

Ocalea rivularis Miller, L., 1851 Oxypoda (Disochara) elongatula Aubé, Pearce: no records 1850 First record: 1963 Pearce: 3 localities Number of records: 2 First subsequent record: 1969 Latest record: 1974 Number of subsequent records: many Latest record: 2020 Ocyusa maura (Erichson, 1837) Pearce: no records Oxypoda (Mycetodrepa) alternans First record: 2011 (Gravenhorst, 1802) Number of records: 3 Pearce: 1 locality Latest record: 2019 First subsequent record: 1968 Number of subsequent records: 7 Ocyusa picina (Aubé, 1850) Latest record: 2020 Pearce: 2 localities First subsequent record: 1989 Oxypoda (Oxypoda) acuminata (Stephens, Number of subsequent records: 8 Latest record: 2017 1832) Pearce: 1 locality First subsequent record: 1968 Hygropora cunctans (Erichson, 1837) Number of subsequent records: 3 Pearce: 1 locality (1925) Latest record: 2019 No subsequent records Oxypoda (Oxypoda) opaca (Gravenhorst, Oxypoda (Atlantoxypoda) lurida 1802) Wollaston, 1857 Pearce: 2 localities Pearce: 1 locality First subsequent record: 1971 First subsequent record: 1998 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 1998 Oxypoda (Oxypoda) spectabilis Märkel, Oxypoda (Bessopora) annularis 1844 Mannerheim, 1830 Pearce: 1 locality (1894) Pearce: no records First subsequent record: 2002 First record: 1979 Number of subsequent records: 2 Number of records: 4 Latest record: 2002 Latest record: 2020 Oxypoda (Oxypoda) vittata Märkel, 1842 Oxypoda (Bessopora) brachyptera Pearce: 1 locality (Stephens, 1832) First subsequent record: 2010 Pearce: 1 locality Number of subsequent records: 1 First subsequent record: 1975 Latest record: 2010 Number of subsequent records: 4 Latest record: 1979 Oxypoda (Podoxya) brevicornis (Stephens, 1832) Oxypoda (Bessopora) formiceticola Pearce: 1 locality Märkel, 1841 First subsequent record: 1975 Pearce: no records Number of subsequent records: 3 First record: 2017 Latest record: 1979 Number of records: 1 Latest record: 2017 Oxypoda (Podoxya) carbonaria (Heer, 1841) Oxypoda (Bessopora) haemorrhoa Pearce: 2 localities (Mannerheim, 1830) No subsequent records Pearce: 2 localities First subsequent record: 2013 Number of subsequent records: 2 Latest record: 2013 Oxypoda (Podoxya) exoleta Erichson, Dinarda dentata (Gravenhorst, 1806) 1839 Pearce: 1 locality Pearce: 1 locality No subsequent records No subsequent records Dinarda maerkeli Kiesenwetter, 1843 Ischnoglossa turcica Wunderle, 1992 Pearce: no records Pearce: 1 locality for Ischnoglossa prolixa, which First record: 2014 could be this species Number of records: 9 First record: 2014 Latest record: 2015 Number of records: 2 Latest record: 2014 Meotica exilis (Knoch in Gravenhorst, 1806) Dexiogyia corticina (Erichson, 1837) Pearce: 2 localities (1890, 1927) Pearce: 1 locality (1904) No subsequent records First subsequent record: 2001 Number of subsequent records: 2 Meotica exillima Sharp, 1915 Latest record: 2004 Pearce: 1 locality First subsequent record: 2001 Crataraea suturalis (Mannerheim, 1830) Number of subsequent records: 3 Pearce: 5 localities Latest record: 2006 First subsequent record: 1969 Number of subsequent records: 5 Meotica filiformis (Motschulsky, 1860) Latest record: 2007 Pearce: no records First record: 1998 Haploglossa gentilis (Märkel, 1844) Number of records: 1 Pearce: 1 locality (1929) Latest record: 1998 First subsequent record: 2020 Number of subsequent records: 2 Tachyusa concinna Heer, 1839 Latest record: 2020 Pearce: no records First record: 1998 Haploglossa marginalis (Gravenhorst, Number of records: 2 1806) Latest record: 1998 Pearce: no records First record: 1987 Tachyusa constricta Erichson, 1837 Number of records: 2 Pearce: no records Latest record: 1988 First record: 1998 Number of records: 3 Haploglossa nidicola (Fairmaire, 1853) Latest record: 2017 Pearce: 1 locality No subsequent records Ischnopoda leucopus (Marsham, 1802) Pearce: no records Haploglossa villosula (Stephens, 1832) First record: 1988 Pearce: 1 locality Number of records: 1 First subsequent record: 1987 Latest record: 1988 Number of subsequent records: many Latest record: 2020 Ischnopoda umbratica (Erichson, 1837) Pearce: 1 locality Thiasophila angulata (Erichson, 1837) No subsequent records Pearce: no records First record: 2014 Thinonoma atra (Gravenhorst, 1806) Number of records: 7 Pearce: 2 localities Latest record: 2014 First subsequent record: 1970 Number of subsequent records: 9 Thiasophila inquilina (Märkel, 1844) Latest record: 2020 Pearce: 2 localities No subsequent records Gnypeta carbonaria (Mannerheim, 1830) Oligota apicata Erichson, 1837 Pearce: 2 localities Pearce: 2 localities First subsequent record: 2017 First subsequent record: 1979 Number of subsequent records: 1 Number of subsequent records: many Latest record: 2017 Latest record: 2019

Gnypeta rubrior Tottenham, 1939 Oligota granaria Erichson, 1837 Pearce: no records Pearce: no records First record: 2013 First record: 2020 Number of records: 3 Number of records: 1 Latest record: 2020 Latest record: 2020

Dilacra luteipes (Erichson, 1837) Oligota inflata (Mannerheim, 1830) Pearce: 2 localities Pearce: 3 localities First subsequent record: 1997 First subsequent record: 1975 Number of subsequent records: 4 Number of subsequent records: 2 Latest record: 2006 Latest record: 1975

Dilacra vilis (Erichson, 1837) Oligota parva Kraatz, 1862 Pearce: no records Pearce: no records First record: 2005 First record: 1996 Number of records: 3 Number of records: 2 Latest record: 2020 Latest record: 2020

Dacrila fallax (Kraatz, 1856) Oligota picipes (Stephens, 1832) Pearce: no records Pearce: 2 localities First record: 2004 First subsequent record: 1975 Number of records: 5 Number of subsequent records: many Latest record: 2020 Latest record: 2019

Cypha discoidea (Erichson, 1839) Oligota pumilio Kiesenwetter, 1858 Pearce: 1 locality (1925) Pearce: no records First subsequent record: 1999 First record: 2013 Number of subsequent records: 1 Number of records: 1 Latest record: 1999 Latest record: 2013

Cypha laeviuscula (Mannerheim, 1830) Oligota punctulata Heer, 1839 Pearce: 1 locality Pearce: 3 localities No subsequent records First subsequent record: 1975 Number of subsequent records: many Cypha longicornis (Paykull, 1800) Latest record: 2015 Pearce: 4 localities First subsequent record: 1970 Oligota pusillima (Gravenhorst, 1806) Number of subsequent records: many Pearce: 2 localities (1896, 1919) Latest record: 2020 First subsequent record: 2000 Number of subsequent records: 3 Cypha punctum (Motschulsky, 1857) Latest record: 2008 Pearce: no records First record: 1975 Myllaena brevicornis (Matthews, A.H., Number of records: 2 1838) Latest record: 1993 Pearce: 4 localities First subsequent record: 1975 Cypha seminulum (Erichson, 1839) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 No subsequent records

Myllaena dubia (Gravenhorst, 1806) Autalia impressa (Olivier, 1795) Pearce: 2 localities Pearce: 3 localities (but may include longicornis) First subsequent record: 1994 First subsequent record: 1975 Number of subsequent records: 6 Number of subsequent records: 9 Latest record: 2016 Latest record: 2019

Myllaena elongata (Matthews, A.H., Autalia longicornis Scheerpeltz, 1947 1838) Pearce: no records (but see impressa) Pearce: no records First record: 1986 First record: 2020 Number of records: 5 Number of records: 1 Latest record: 2019 Latest record: 2020 Autalia rivularis (Gravenhorst, 1802) Myllaena gracilis (Matthews, A.H., 1838) Pearce: 2 localities Pearce: no records First subsequent record: 1975 First record: 2015 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 2015 Pseudopasilia testacea (Brisout de Myllaena infuscata Kraatz, 1853 Barneville, C., 1863) Pearce: 1 locality Pearce: 2 localities First subsequent record: 2003 No subsequent records Number of subsequent records: 6 Latest record: 2019 Leptusa (Dendroleptusa) fumida (Erichson, 1839) Myllaena intermedia Erichson, 1837 Pearce: 2 localities Pearce: 2 localities First subsequent record: 1978 First subsequent record: 2000 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019 Leptusa (Leptusa) pulchella (Mannerheim, Myllaena kraatzi Sharp, 1871 1830) Pearce: 1 locality Pearce: no records First subsequent record: 2016 First record: 2017 Number of subsequent records: 3 Number of records: 3 Latest record: 2018 Latest record: 2019

Myllaena minuta (Gravenhorst, 1806) Leptusa (Pachygluta) ruficollis (Erichson, Pearce: no records First record: 2001 1839) Number of records: 4 Pearce: 2 localities Latest record: 2019 First subsequent record: 1969 Number of subsequent records: many Latest record: 2020 Diglotta mersa (Haliday, 1837) Pearce: 1 locality First subsequent record: 2006 Heterota plumbea (Waterhouse, G.R., Number of subsequent records: 2 1858) Latest record: 2019 Pearce: 8 localities First subsequent record: 1966 Number of subsequent records: 9 Diglotta sinuaticollis (Mulsant & Rey, Latest record: 2020 1870) Pearce: 5 localities First subsequent record: 2000 Bolitochara bella Märkel, 1844 Number of subsequent records: 1 Pearce: 3 localities Latest record: 2000 First subsequent record: 1975 Number of subsequent records: 6 Latest record: 2019 Bolitochara obliqua Erichson, 1837 Gyrophaena (Gyrophaena) manca Pearce: no records Erichson, 1839 First record: 2018 Pearce: no records Number of records: 2 First record: 2000 Latest record: 2018 Number of records: 2 Latest record: 2019 Bolitochara tecta Assing, 2014 [2012: Bolitochara lucida] Gyrophaena (Gyrophaena) minima Pearce: 1 locality (1874) Erichson, 1837 First subsequent record: 2018 Pearce: 1 locality (1891) Number of subsequent records: 1 First subsequent record: 2000 Latest record: 2018 Number of subsequent records: 2 Latest record: 2015 Encephalus complicans Stephens, 1832 Pearce: 1 locality Gyrophaena (Gyrophaena) nana (Paykull, First subsequent record: 1997 1800) Number of subsequent records: many Pearce: 1 locality Latest record: 2017 First subsequent record: 2019 Number of subsequent records: 1 Gyrophaena (Gyrophaena) bihamata Latest record: 2019 Thomson, C.G., 1867 Pearce: no records Gyrophaena (Leptarthrophaena) affinis First record: 2006 Mannerheim, 1830 Number of records: 4 Pearce: 3 localities Latest record: 2018 First subsequent record: 1994 Number of subsequent records: 6 Gyrophaena (Gyrophaena) fasciata Latest record: 2014 (Marsham, 1802) Pearce: 2 localities Agaricochara latissima (Stephens, 1832) First subsequent record: 1968 Pearce: no records Number of subsequent records: 5 First record: 2000 Latest record: 2020 Number of records: 8 Latest record: 2020 Gyrophaena (Gyrophaena) gentilis Erichson, 1839 Homalota plana (Gyllenhal, 1810) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1971 First subsequent record: 1999 Number of subsequent records: 2 Number of subsequent records: 1 Latest record: 2018 Latest record: 1999

Gyrophaena (Gyrophaena) joyi Wendeler, Anomognathus cuspidatus (Erichson, 1924 1839) Pearce: no records Pearce: 1 locality First record: 1975 First subsequent record: 1991 Number of records: 3 Number of subsequent records: 4 Latest record: 2019 Latest record: 2020

Gyrophaena (Gyrophaena) lucidula Silusa rubiginosa Erichson, 1837 Erichson, 1837 Pearce: no records Pearce: 1 locality (1890) First record: 2004 First subsequent record: 2020 Number of records: 1 Number of subsequent records: 1 Latest record: 2004 Latest record: 2020

Hygronoma dimidiata (Gravenhorst, 1806) Amischa analis (Gravenhorst, 1802) Pearce: 6 localities Pearce: 6 localities First subsequent record: 1989 First subsequent record: 1970 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Placusa pumilio (Gravenhorst, 1802) Amischa bifoveolata (Mannerheim, 1830) Pearce: no records Pearce: no records First record: 2004 First record: 1975 Number of records: 1 Number of records: many Latest record: 2004 Latest record: 2006

Placusa tachyporoides (Waltl, 1838) Amischa decipiens (Sharp, 1869) Pearce: 1 locality (1892) Pearce: 1 locality No subsequent records First subsequent record: 1979 Number of subsequent records: many Actocharis readingii Sharp, 1870 Latest record: 2020 Pearce: no records First record: 1930 Amischa forcipata Mulsant & Rey, 1873 Number of records: 1 Pearce: no records Latest record: 1930 First record: 1979 Number of records: 7 Phytosus (Actosus) balticus Kraatz, 1859 Latest record: 2019 Pearce: 3 localities First subsequent record: 2004 Amischa nigrofusca (Stephens, 1832) Number of subsequent records: 3 Pearce: no records Latest record: 2016 First record: 1998 Number of records: 9 Phytosus (Actosus) nigriventris Latest record: 2019 (Chevrolat, 1843) Pearce: 4 localities Nehemitropia lividipennis (Mannerheim, No subsequent records 1830) Pearce: 3 localities Phytosus (Phytosus) spinifer Curtis, 1838 First subsequent record: 1971 Pearce: 4 localities Number of subsequent records: many First subsequent record: 2004 Latest record: 2013 Number of subsequent records: 4 Latest record: 2019 Notothecta flavipes (Gravenhorst, 1806) Pearce: no records Arena tabida (Kiesenwetter, 1850) First record: 1996 Pearce: 2 localities Number of records: 1 No subsequent records Latest record: 1996

Boreophilia eremita (Rye, 1866) Brundinia marina (Mulsant & Rey, 1853) Pearce: no records Pearce: 1 locality (1886) First record: 1998 No subsequent records Number of records: 1 Latest record: 1998 Brundinia meridionalis (Mulsant & Rey, 1853) Hydrosmecta longula (Heer, 1839) Pearce: 1 locality Pearce: 1 locality No subsequent records No subsequent records Ousipalia caesula (Erichson, 1839) Pearce: 2 localities No subsequent records

Dinaraea aequata (Erichson, 1837) Atheta (Alaobia) scapularis (Sahlberg, Pearce: 2 localities C.R., 1831) First subsequent record: 2000 Pearce: 2 localities (1874, 1925) Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: 3 Latest record: 2017 Dinaraea angustula (Gyllenhal, 1810) Pearce: 1 locality (1925) Atheta (Alaobia) trinotata (Kraatz, 1856) First subsequent record: 1980 Pearce: 3 localities Number of subsequent records: 9 First subsequent record: 1975 Latest record: 2020 Number of subsequent records: 3 Latest record: 2006 Dinaraea linearis (Gravenhorst, 1802) Pearce: 1 locality Atheta (Anopleta) corvina (Thomson, First subsequent record: 1975 C.G., 1856) Number of subsequent records: 1 Pearce: no records Latest record: 1975 First record: 1966 Number of records: 3 Dadobia immersa (Erichson, 1837) Latest record: 2019 Pearce: 1 locality No subsequent records Atheta (Atheta) aeneicollis (Sharp, 1869) Pearce: 1 locality (1892) Plataraea brunnea (Fabricius, 1798) First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: 6 No subsequent records Latest record: 2004

Liogluta longiuscula (Gravenhorst, 1802) Atheta (Atheta) aquatica (Thomson, C.G., Pearce: 2 localities 1852) First subsequent record: 2014 Pearce: 4 localities Number of subsequent records: 3 First subsequent record: 2004 Latest record: 2019 Number of subsequent records: 4 Latest record: 2020 Liogluta microptera Thomson, C.G., 1867 Pearce: 2 localities Atheta (Atheta) autumnalis (Erichson, First subsequent record: 1968 1839) Number of subsequent records: many Pearce: 1 locality Latest record: 2011 No subsequent records

Atheta (Alaobia) gagatina (Baudi di Selve, Atheta (Atheta) basicornis (Mulsant & 1848) Rey, 1852) Pearce: 1 locality (1891) Pearce: no records First subsequent record: 1993 First record: 2013 Number of subsequent records: 2 Number of records: 2 Latest record: 1993 Latest record: 2017

Atheta (Alaobia) pallidicornis (Thomson, Atheta (Atheta) britanniae Bernhauer & C.G., 1856) Scheerpeltz, 1926 Pearce: no records Pearce: no records First record: 2017 First record: 1993 Number of records: 1 Number of records: 3 Latest record: 2017 Latest record: 2017 Atheta (Atheta) brunneipennis (Thomson, Atheta (Atheta) oblita (Erichson, 1839) C.G., 1852) Pearce: 2 localities Pearce: no records First subsequent record: 1975 First record: 1975 Number of subsequent records: 3 Number of records: 1 Latest record: 2020 Latest record: 1975 Atheta (Atheta) ravilla (Erichson, 1839) Atheta (Atheta) castanoptera Pearce: 1 locality (1890) (Mannerheim, 1830) First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: 5 First subsequent record: 1968 Latest record: 2020 Number of subsequent records: 6 Latest record: 2020 Atheta (Atheta) triangulum (Kraatz, 1856) Pearce: no records Atheta (Atheta) crassicornis (Fabricius, First record: 1979 Number of records: 9 1792) Latest record: 2020 Pearce: no records First record: 1975 Number of records: many Atheta (Atheta) vaga (Heer, 1839) Latest record: 2020 Pearce: no records First record: 2013 Number of records: 2 Atheta (Atheta) divisa (Märkel, 1844) Latest record: 2014 Pearce: 1 locality First subsequent record: 1975 Number of subsequent records: 1 Atheta (Atheta) xanthopus (Thomson, Latest record: 1975 C.G., 1856) Pearce: 2 localities Atheta (Atheta) fungicola (Thomson, C.G., First subsequent record: 2006 Number of subsequent records: 3 1852) Latest record: 2020 Pearce: 3 localities First subsequent record: 1975 Number of subsequent records: 3 Atheta (Badura) cauta (Erichson, 1837) Latest record: 1998 [2012: Dimetrota cauta] Pearce: no records Atheta (Atheta) graminicola (Gravenhorst, First record: 1975 Number of records: 2 1806) Latest record: 2015 Pearce: 4 localities First subsequent record: 1969 Number of subsequent records: many Atheta (Bessobia) excellens (Kraatz, 1856) Latest record: 2018 Pearce: no records First record: 1979 Number of records: 1 Atheta (Atheta) harwoodi Williams, 1930 Latest record: 1979 Pearce: no records First record: 1989 Number of records: 1 Atheta (Bessobia) fungivora (Thomson, Latest record: 1989 C.G., 1867) Pearce: 1 locality Atheta (Atheta) hypnorum (Kiesenwetter, First subsequent record: 2000 Number of subsequent records: 4 1850) Latest record: 2012 Pearce: 3 localities (1874, 1892, 1919) First subsequent record: 1982 Number of subsequent records: 1 Atheta (Bessobia) occulta (Erichson, 1837) Latest record: 1982 Pearce: no records First record: 2004 Number of records: 1 Latest record: 2004 Atheta (Ceritaxa) subterranea (Mulsant & Atheta (Dimetrota) atramentaria Rey, 1853) [2012: Neohilara subterranea] (Gyllenhal, 1810) Pearce: no records Pearce: 6 localities First record: 1993 First subsequent record: 1970 Number of records: 1 Number of subsequent records: many Latest record: 1993 Latest record: 2019

Atheta (Chaetida) longicornis Atheta (Dimetrota) marcida (Erichson, (Gravenhorst, 1802) 1837) Pearce: 2 localities (1890, 1910) Pearce: 1 locality First subsequent record: 1970 First subsequent record: 2006 Number of subsequent records: 6 Number of subsequent records: 5 Latest record: 2008 Latest record: 2015

Atheta (Coprothassa) melanaria Atheta (Dimetrota) nigripes (Thomson, (Mannerheim, 1830) C.G., 1856) Pearce: 1 locality Pearce: no records First subsequent record: 2011 First record: 1975 Number of subsequent records: 3 Number of records: 3 Latest record: 2019 Latest record: 2015

Atheta (Datomicra) celata (Erichson, Atheta (Dimetrota) setigera (Sharp, 1869) 1837) Pearce: 1 locality Pearce: 1 locality No subsequent records First subsequent record: 1975 Number of subsequent records: 5 Atheta (Dimetrotina) laticollis (Stephens, Latest record: 2020 1832) [2012: Mycetota laticollis] Pearce: 2 localities Atheta (Datomicra) dadopora Thomson, First subsequent record: 1955 C.G., 1867 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2004 Number of records: 2 Atheta (Microdota) aegra (Heer, 1841) Latest record: 2009 Pearce: no records First record: 2017 Atheta (Datomicra) nigra (Kraatz, 1856) Number of records: 1 Pearce: 2 localities Latest record: 2017 First subsequent record: 1971 Number of subsequent records: many Atheta (Microdota) amicula (Stephens, Latest record: 2017 1832) Pearce: 1 locality Atheta (Datomicra) sordidula (Erichson, First subsequent record: 1975 1837) Number of subsequent records: 5 Pearce: 1 locality Latest record: 2020 First subsequent record: 1975 Number of subsequent records: 3 Atheta (Microdota) atricolor (Sharp, 1869) Latest record: 1999 Pearce: 1 locality No subsequent records Atheta (Datomicra) zosterae (Thomson, C.G., 1856) Atheta (Microdota) benickiella Brundin, Pearce: no records 1948 First record: 2006 Pearce: no records Number of records: 1 First record: 1993 Latest record: 2006 Number of records: 1 Latest record: 1993 Atheta (Microdota) indubia (Sharp, 1869) Atheta (Philhygra) hygrobia (Thomson, Pearce: no records C.G., 1856) First record: 1993 Pearce: no records Number of records: 3 First record: 2004 Latest record: 2004 Number of records: 1 Latest record: 2004 Atheta (Microdota) minuscula (Brisout de Barneville, C., 1860) Atheta (Philhygra) hygrotopora (Kraatz, Pearce: no records 1856) First record: 1975 Pearce: no records Number of records: 2 First record: 2013 Latest record: 1999 Number of records: 1 Latest record: 2013 Atheta (Mocyta) amplicollis (Mulsant & Rey, 1873) Atheta (Philhygra) luridipennis Pearce: no records (Mannerheim, 1830) First record: 1998 Pearce: no records Number of records: many First record: 2020 Latest record: 2020 Number of records: 1 Latest record: 2020 Atheta (Mocyta) clientula (Erichson, 1839) Pearce: 1 locality Atheta (Philhygra) malleus Joy, 1913 First subsequent record: 1980 Pearce: 2 localities Number of subsequent records: 9 First subsequent record: 2006 Latest record: 2015 Number of subsequent records: many Latest record: 2018 Atheta (Mocyta) fungi (Gravenhorst, 1806) Pearce: 8 localities Atheta (Philhygra) melanocera (Thomson, First subsequent record: 1975 C.G., 1856) Number of subsequent records: many Pearce: 1 locality Latest record: 2020 First subsequent record: 1989 Number of subsequent records: 2 Atheta (Mocyta) orbata (Erichson, 1837) Latest record: 2006 Pearce: 2 localities No subsequent records Atheta (Philhygra) obtusangula Joy, 1913 Pearce: 1 locality Atheta (Parameotica) difficilis (Brisout de First subsequent record: 2000 Barneville, C., 1860) Number of subsequent records: 1 Pearce: no records Latest record: 2000 First record: 2006 Number of records: 2 Atheta (Philhygra) palustris Latest record: 2017 (Kiesenwetter, 1844) Pearce: 3 localities Atheta (Philhygra) elongatula First subsequent record: 2019 (Gravenhorst, 1802) Number of subsequent records: 1 Pearce: 1 locality Latest record: 2019 First subsequent record: 1989 Number of subsequent records: many Atheta (Philhygra) parca (Mulsant & Rey, Latest record: 2019 1873) Pearce: 1 locality (1928) Atheta (Philhygra) fallaciosa (Sharp, No subsequent records 1869) Pearce: 1 locality No subsequent records

Atheta (Philhygra) volans (Scriba, 1859) Acrotona obfuscata (Gravenhorst, 1802) Pearce: 1 locality Pearce: 1 locality First subsequent record: 2006 No subsequent records Number of subsequent records: 2 Latest record: 2006 Acrotona parvula (Mannerheim, 1830) Pearce: no records Atheta (Rhagocneme) subsinuata First record: 1975 (Erichson, 1839) Number of records: 3 Pearce: 1 locality Latest record: 2015 No subsequent records Acrotona pygmaea (Gravenhorst, 1802) Atheta (Tetropla) liturata (Stephens, 1832) Pearce: 2 localities Pearce: 1 locality (1925) First subsequent record: 2003 First subsequent record: 1975 Number of subsequent records: 4 Number of subsequent records: 4 Latest record: 2006 Latest record: 2020 Adota maritima (Mannerheim, 1843) Atheta (Tetropla) nigritula (Gravenhorst, Pearce: no records 1802) First record: 1976 Pearce: 1 locality (1890) Number of records: 2 First subsequent record: 1968 Latest record: 2004 Number of subsequent records: 2 Latest record: 1996 Dalotia coriaria (Kraatz, 1856) Pearce: no records Atheta (Thinobaena) vestita (Gravenhorst, First record: 1969 Number of records: 4 1806) Latest record: 2020 Pearce: 5 localities First subsequent record: 1964 Number of subsequent records: many Trichiusa immigrata Lohse, 1984 Latest record: 2019 Pearce: no records First record: 1995 Number of records: 5 Acrotona aterrima (Gravenhorst, 1802) Latest record: 2006 Pearce: 1 locality First subsequent record: 1975 Number of subsequent records: 7 Alianta incana (Erichson, 1837) Latest record: 2015 Pearce: 1 locality First subsequent record: 1964 Number of subsequent records: 6 Acrotona benicki Allen, 1940 Latest record: 2015 Pearce: no records First record: 1937 Number of records: 2 Pachnida nigella (Erichson, 1837) Latest record: 2015 Pearce: 1 locality First subsequent record: 1998 Number of subsequent records: 3 Acrotona exigua (Erichson, 1837) Latest record: 2014 Pearce: 1 locality No subsequent records Thamiaraea cinnamomea (Gravenhorst, Acrotona muscorum (Brisout de 1802) Pearce: no records Barneville, C., 1860) First record: 1998 Pearce: no records Number of records: 3 First record: 2003 Latest record: 2019 Number of records: 4 Latest record: 2020 Thamiaraea hospita (Märkel, 1844) Alevonota elegantula (Brisout de Pearce: 1 locality Barneville, C., 1863) No subsequent records Pearce: 1 locality No subsequent records Halobrecta algae (Hardy, 1851) Pearce: 3 localities Alevonota gracilenta (Erichson, 1839) No subsequent records Pearce: no records First record: 2017 Halobrecta algophila (Fenyes, 1909) Number of records: 1 Pearce: no records Latest record: 2017 First record: 2017 Number of records: 1 Drusilla canaliculata Fabricius 1787 Latest record: 2017 Pearce: generally common First subsequent record: 1939 Halobrecta flavipes Thomson, C.G., 1861 Number of subsequent records: many Pearce: 5 localities Latest record: 2020 First subsequent record: 2019 Number of subsequent records: 1 Zyras collaris (Paykull, 1800) Latest record: 2019 Pearce: 3 localities First subsequent record: 1945 Callicerus obscurus Gravenhorst, 1802 Number of subsequent records: 1 Pearce: 2 localities Latest record: 1945 First subsequent record: 1979 Number of subsequent records: 2 Zyras haworthi (Stephens, 1835) Latest record: 2015 Pearce: 2 localities (1884, 1929) First subsequent record: 1987 Callicerus rigidicornis (Erichson, 1839) Number of subsequent records: 2 Pearce: 1 locality Latest record: 1993 No subsequent records Pella cognata (Märkel, 1842) Aloconota (Aloconota) gregaria (Erichson, Pearce: 3 localities 1839) First subsequent record: 1980 Pearce: 1 locality Number of subsequent records: 3 First subsequent record: 1975 Latest record: 2017 Number of subsequent records: many Latest record: 2018 Pella funesta (Gravenhorst, 1806) Pearce: 1 locality (1925) Aloconota (Aloconota) sulcifrons First subsequent record: 1975 Number of subsequent records: 1 (Stephens, 1832) Latest record: 1975 Pearce: 3 localities No subsequent records Pella humeralis (Gravenhorst, 1802) Pearce: 1 locality Aloconota (Disopora) longicollis (Mulsant First subsequent record: 1981 & Rey, 1852) Number of subsequent records: many Pearce: 1 locality Latest record: 2015 First subsequent record: 2004 Number of subsequent records: 1 Latest record: 2004 Pella laticollis (Märkel, 1844) Pearce: 1 locality (1929) First record: 1975 Geostiba circellaris Gravenhorst 1806 Number of records: 1 Pearce: 2 localities Latest record: 1975 First subsequent record: 1971 Number of subsequent records: many Latest record: 2020

Pella limbata (Paykull, 1789) Myrmecopora sulcata (Kiesenwetter, Pearce: 7 localities 1850) First subsequent record: 1975 Pearce: 2 localities Number of subsequent records: 3 First subsequent record: 1975 Latest record: 2019 Number of subsequent records: 6 Latest record: 2005 Lomechusa emarginata (Paykull, 1789) Pearce: 1 locality Myrmecopora uvida (Erichson, 1840) First subsequent record: 1932 Pearce: 7 localities Number of subsequent records: many First subsequent record: 1975 Latest record: 2016 Number of subsequent records: 2 Latest record: 1991 Lomechusa paradoxa Gravenhorst, 1806 Pearce: no records Scaphidium quadrimaculatum Olivier, First record: 2001 1790 Number of records: 2 Pearce: 4 localities Latest record: 2019 First subsequent record: 1968 Number of subsequent records: many Cordalia obscura (Gravenhorst, 1802) Latest record: 2020 Pearce: 5 localities First subsequent record: 1989 Scaphisoma agaricinum (Linnaeus, 1758) Number of subsequent records: many Pearce: 6 localities Latest record: 2020 First subsequent record: 2011 Number of subsequent records: 7 Falagria caesa Erichson, 1837 Latest record: 2019 Pearce: 1 locality First subsequent record: 1995 Scaphisoma boleti (Panzer, 1793) Number of subsequent records: 7 Pearce: 1 locality Latest record: 2015 First subsequent record: 1971 Number of subsequent records: 2 Falagria sulcatula (Gravenhorst, 1806) Latest record: 2014 Pearce: 2 localities First subsequent record: 2000 Siagonium quadricorne Kirby & Spence, Number of subsequent records: 2 Latest record: 2007 1815 Pearce: 1 locality (1921) First subsequent record: 1993 Falagrioma thoracica (Stephens, 1832) Number of subsequent records: many Pearce: 1 locality Latest record: 2019 First subsequent record: 1975 Number of subsequent records: 8 Latest record: 2016 Deleaster dichrous (Gravenhorst, 1802) Pearce: no records First record: 1988 Myrmecopora brevipes Butler, 1909 Number of records: 6 Pearce: 2 localities Latest record: 2013 No subsequent records Coprophilus striatulus (Fabricius, 1792) Myrmecopora oweni Assing, 1997 Pearce: 1 locality Pearce: no records No subsequent records First record: 1996 Number of records: 4 Latest record: 1998 Manda mandibularis (Gyllenhal, 1827) Pearce: 1 locality No subsequent records Syntomium aeneum (Müller, P.W.J., 1821) Bledius (Dicarenus) fergussoni Joy, 1912 Pearce: 1 locality Pearce: 5 localities No subsequent records No subsequent records

Bledius (Bargus) fuscipes Rye, 1865 Bledius (Dicarenus) subniger Schneider, Pearce: 1 locality (1921) O., 1898 First subsequent record: 2000 Pearce: 2 localities (1910, 1928) Number of subsequent records: 1 No subsequent records Latest record: 2000 Bledius (Hesperophilus) atricapillus Bledius (Bargus) longulus Erichson, 1839 (Germar, 1825) Pearce: 1 locality Pearce: 1 locality First subsequent record: 1989 First subsequent record: 1955 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2006 Latest record: 2019

Bledius (Bargus) opacus (Block, 1799) Bledius (Hesperophilus) femoralis Pearce: 1 locality First subsequent record: 1968 (Gyllenhal, 1827) Number of subsequent records: 2 Pearce: no records Latest record: 2005 First record: 1955 Number of records: 3 Latest record: 2001 Bledius (Bargus) pallipes (Gravenhorst, 1806) Bledius (Hesperophilus) gallicus Pearce: 1 locality (1925) First subsequent record: 1953 (Gravenhorst, 1806) Number of subsequent records: 4 Pearce: no records Latest record: 2005 First record: 2013 Number of records: 1 Latest record: 2013 Bledius (Bledius) frisius Lohse, 1978 Pearce: 3 localities First subsequent record: 1936 Ochthephilus angustior (Bernhauer, 1943) Number of subsequent records: many Pearce: no records Latest record: 2015 First record: 2020 Number of records: 1 Latest record: 2020 Bledius (Bledius) spectabilis Kraatz, 1857 Pearce: no records First record: 1979 Ochthephilus aureus (Fauvel, 1871) Number of records: many Pearce: no records Latest record: 2016 First record: 1997 Number of records: 5 Latest record: 2020 Bledius (Bledius) tricornis (Herbst, 1784) Pearce: 1 locality First subsequent record: 2004 Teropalpus unicolor (Sharp, 1900) Number of subsequent records: 1 Pearce: 1 locality (1912) Latest record: 2004 First subsequent record: 1966 Number of subsequent records: 5 Latest record: 2017 Bledius (Bledius) unicornis (Germar, 1825) Thinodromus arcuatus (Stephens, 1834) Pearce: 1 locality Pearce: “Dorset” (A. Ford) First subsequent record: 1996 First subsequent record: 1954 Number of subsequent records: 2 Number of subsequent records: many Latest record: 1997 Latest record: 2020

Carpelimus (Carpelimus) fuliginosus Carpelimus (Trogophloeus) elongatulus (Gravenhorst, 1802) (Erichson, 1839) Pearce: no records Pearce: 4 localities First record: 2013 First subsequent record: 1975 Number of records: 1 Number of subsequent records: 9 Latest record: 2013 Latest record: 2019

Carpelimus (Carpelimus) obesus Carpelimus (Trogophloeus) gracilis (Kiesenwetter, 1844) (Mannerheim, 1830) Pearce: no records Pearce: no records First record: 2006 First record: 1993 Number of records: 1 Number of records: 4 Latest record: 2006 Latest record: 2014

Carpelimus (Carpelimus) pusillus Carpelimus (Trogophloeus) halophilus (Gravenhorst, 1802) (Kiesenwetter, 1844) Pearce: 1 locality Pearce: 4 localities First subsequent record: 1993 No subsequent records Number of subsequent records: 5 Latest record: 2016 Carpelimus (Trogophloeus) subtilis (Erichson, 1839) Carpelimus (Paratrogophloeus) erichsoni Pearce: no records (Sharp, 1871) First record: 2017 Pearce: 6 localities for bilineatus which could be Number of records: 1 either species Latest record: 2017 First subsequent record: 1949 Number of subsequent records: 9 Thinobius brevipennis Kiesenwetter, 1850 Latest record: 2018 Pearce: no records First record: 1990 Carpelimus (Paratrogophloeus) rivularis Number of records: 2 (Motschulsky, 1860) Latest record: 1997 Pearce: 1 locality First subsequent record: 1949 Aploderus caelatus (Gravenhorst, 1802) Number of subsequent records: many Pearce: 1 locality Latest record: 2019 No subsequent records

Carpelimus (Paratrogophloeus) similis Platystethus (Craetopycrus) alutaceus (Smetana, 1967) Thomson, C.G., 1861 Pearce: no records Pearce: no records First record: 1989 First record: 2014 Number of records: 1 Number of records: 7 Latest record: 1989 Latest record: 2018

Carpelimus (Troginus) zealandicus Platystethus (Craetopycrus) capito Heer, (Sharp, 1900) 1839 Pearce: no records Pearce: 1 locality First record: 1975 First subsequent record: 2002 Number of records: 2 Number of subsequent records: 2 Latest record: 2005 Latest record: 2008

Carpelimus (Trogophloeus) corticinus Platystethus (Craetopycrus) cornutus (Gravenhorst, 1806) (Gravenhorst, 1802) Pearce: 4 localities Pearce: 2 localities First subsequent record: 1963 First subsequent record: 1966 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Platystethus (Craetopycrus) nitens Anotylus sculpturatus (Gravenhorst, 1806) (Sahlberg, C.R., 1832) Pearce: generally common Pearce: no records First subsequent record: 1979 First record: 1986 Number of subsequent records: many Number of records: many Latest record: 2019 Latest record: 2020 Anotylus tetracarinatus (Block, 1799) Platystethus (Craetopycrus) nodifrons Pearce: 6 localities Mannerheim, 1830 First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1963 Latest record: 2020 Number of subsequent records: 2 Latest record: 2008 Oxytelus (Epomotylus) sculptus Gravenhorst, 1806 Platystethus (Platystethus) arenarius Pearce: 2 localities (Geoffroy in Fourcroy, 1785) First subsequent record: 1990 Pearce: 5 localities Number of subsequent records: 9 First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2018 Oxytelus (Oxytelus) migrator Fauvel, 1904 Pearce: no records Anotylus complanatus (Erichson, 1839) First record: 2003 Pearce: 4 localities Number of records: 2 First subsequent record: 1975 Latest record: 2003 Number of subsequent records: 8 Latest record: 2020 Oxytelus (Tanycraerus) laqueatus (Marsham, 1802) Anotylus insecatus (Gravenhorst, 1806) Pearce: 4 localities Pearce: 2 localities First subsequent record: 1975 No subsequent records Number of subsequent records: many Latest record: 2019 Anotylus inustus (Gravenhorst, 1806) Pearce: 2 localities Oxyporus rufus (Linnaeus, 1758) First subsequent record: 1995 Pearce: 7 localities Number of subsequent records: many First subsequent record: 2011 Latest record: 2016 Number of subsequent records: 1 Latest record: 2011 Anotylus maritimus Thomson, C.G., 1861 Pearce: 2 localities Eutheia schaumii Kiesenwetter, 1858 First subsequent record: 1977 Pearce: no records Number of subsequent records: 3 First record: 2003 Latest record: 2019 Number of records: 2 Latest record: 2003 Anotylus nitidulus (Gravenhorst, 1802) Pearce: 3 localities Eutheia scydmaenoides Stephens, 1830 First subsequent record: 2008 Pearce: no records Number of subsequent records: 6 First record: 2003 Latest record: 2020 Number of records: 2 Latest record: 2003 Anotylus rugosus (Fabricius, 1775) Pearce: 5 localities Cephennium gallicum Ganglbauer, 1899 First subsequent record: 1963 Pearce: 7 localities Number of subsequent records: many First subsequent record: 1979 Latest record: 2020 Number of subsequent records: many Latest record: 2020

Neuraphes angulatus (Müller, P.W.J. & Stenichnus scutullaris (Müller, P.W.J. & Kunze, 1822) Kunze, 1822) Pearce: 1 locality (1925) Pearce: 2 localities (1899, 1901) First subsequent record: 1979 First subsequent record: 2002 Number of subsequent records: 9 Number of subsequent records: many Latest record: 2016 Latest record: 2020

Neuraphes elongatulus (Müller, P.W.J. & Euconnus fimetarius (Chaudoir, 1845) Kunze, 1822) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1979 First subsequent record: 1987 Number of subsequent records: 6 Number of subsequent records: 1 Latest record: 2014 Latest record: 1987 Euconnus hirticollis (Illiger, 1798) Neuraphes plicicollis (Reitter, 1880) Pearce: 2 localities Pearce: no records First subsequent record: 2006 First record: 2019 Number of subsequent records: 3 Number of records: 4 Latest record: 2017 Latest record: 2019 Scydmaenus rufus Müller, P.W.J. & Neuraphes praeteritus (Rye, 1872) Kunze, 1822 Pearce: no records Pearce: no records First record: 2018 First record: 2003 Number of records: 1 Number of records: 3 Latest record: 2018 Latest record: 2004

Stenichnus bicolor (Denny, 1825) Scydmaenus tarsatus Müller, P.W.J. & Pearce: no records Kunze, 1822 First record: 2018 Pearce: 4 localities Number of records: 2 First subsequent record: 1954 Latest record: 2019 Number of subsequent records: 8 Latest record: 2020 Stenichnus collaris (Müller, P.W.J. & Kunze, 1822) Dianous coerulescens (Gyllenhal, 1810) Pearce: 3 localities Pearce: “Dorset” (A. Ford) First subsequent record: 1975 First subsequent record: 2017 Number of subsequent records: many Number of subsequent records: 2 Latest record: 2020 Latest record: 2020

Stenichnus poweri (Fowler, 1884) Stenus (Hemistenus) aceris Stephens, 1833 Pearce: no records Pearce: 5 localities First record: 1998 First subsequent record: 1975 Number of records: 4 Number of subsequent records: many Latest record: 2000 Latest record: 2020

Stenichnus pusillus (Müller, P.W.J. & Stenus (Hemistenus) fuscicornis Erichson, Kunze, 1822) 1840 Pearce: 4 localities Pearce: no records First subsequent record: 2002 First record: 1975 Number of subsequent records: many Number of records: 3 Latest record: 2020 Latest record: 1996 Stenus (Hemistenus) geniculatus Stenus (Hypostenus) fulvicornis Stephens, Gravenhorst, 1806 1833 Pearce: 1 locality Pearce: 1 locality (1898) First subsequent record: 1979 First subsequent record: 1967 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Stenus (Hemistenus) impressus Germar, Stenus (Hypostenus) kiesenwetteri 1824 Rosenhauer, 1856 Pearce: generally common and widely distributed Pearce: 1 locality (1892) First subsequent record: 1960 First subsequent record: 2000 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2019 Latest record: 2019

Stenus (Hemistenus) ochropus Stenus (Hypostenus) latifrons Erichson, Kiesenwetter, 1858 1839 Pearce: 1 locality Pearce: 6 localities First subsequent record: 1999 First subsequent record: 1964 Number of subsequent records: 3 Number of subsequent records: many Latest record: 2019 Latest record: 2020

Stenus (Hemistenus) ossium Stephens, Stenus (Hypostenus) similis (Herbst, 1784) 1833 Pearce: 8 localities Pearce: 2 localities First subsequent record: 1964 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Stenus (Hypostenus) solutus Erichson, Stenus (Hemistenus) pallipes Gravenhorst, 1840 1802 Pearce: no records Pearce: no records First record: 1997 First record: 2005 Number of records: many Number of records: 5 Latest record: 2019 Latest record: 2014 Stenus (Hypostenus) tarsalis Ljungh, 1810 Stenus (Hemistenus) subaeneus Erichson, Pearce: 6 localities 1840 First subsequent record: 1964 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1975 Latest record: 2006 Number of subsequent records: many Latest record: 2019 Stenus (Metastenus) bifoveolatus Gyllenhal, 1827 Stenus (Hypostenus) cicindeloides Pearce: 2 localities (Schaller, 1783) First subsequent record: 1997 Pearce: 1 locality (1898) Number of subsequent records: many First subsequent record: 1963 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Stenus (Metastenus) binotatus Ljungh, 1804 Stenus (Hypostenus) fornicatus Stephens, Pearce: 1 locality (1920) 1833 First subsequent record: 1997 Pearce: no records Number of subsequent records: 2 First record: 2005 Latest record: 2016 Number of records: 6 Latest record: 2019 Stenus (Metastenus) brevipennis Thomson, Stenus (Metastenus) picipes Stephens, C.G., 1851 1833 Pearce: no records Pearce: 1 locality First record: 2006 First subsequent record: 1964 Number of records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Stenus (Metastenus) butrintensis Smetana, Stenus (Metastenus) pubescens Stephens, 1959 1833 Pearce: no records Pearce: 1 locality First record: 2009 First subsequent record: 1989 Number of records: 4 Number of subsequent records: 4 Latest record: 2011 Latest record: 2006

Stenus (Metastenus) canescens Stenus (Stenus) argus Gravenhorst, 1806 Rosenhauer, 1856 Pearce: no records Pearce: 1 locality (1899) First record: 2008 First subsequent record: 1992 Number of records: 1 Number of subsequent records: 4 Latest record: 2008 Latest record: 2020 Stenus (Stenus) assequens Rey, 1844 Stenus (Metastenus) flavipes Stephens, Pearce: no records 1833 First record: 1967 Pearce: 5 localities Number of records: 1 First subsequent record: 1975 Latest record: 1967 Number of subsequent records: many Latest record: 2020 Stenus (Stenus) ater Mannerheim, 1830 Pearce: 6 localities Stenus (Metastenus) nitidiusculus No subsequent records Stephens, 1833 Pearce: 3 localities Stenus (Stenus) atratulus Erichson, 1839 First subsequent record: 1964 Pearce: 1 locality (1890) Number of subsequent records: many First subsequent record: 2010 Latest record: 2020 Number of subsequent records: 1 Latest record: 2010 Stenus (Metastenus) niveus Fauvel, 1865 Pearce: no records Stenus (Stenus) biguttatus (Linnaeus, First record: 2018 1758) Number of records: 1 Pearce: no records Latest record: 2018 First record: 1989 Number of records: 8 Stenus (Metastenus) pallitarsis Stephens, Latest record: 2009 1833 Pearce: 5 localities Stenus (Stenus) bimaculatus Gyllenhal, First subsequent record: 1983 1810 Number of subsequent records: many Pearce: 6 localities Latest record: 2020 First subsequent record: 1968 Number of subsequent records: many Stenus (Metastenus) picipennis Erichson, Latest record: 2020 1840 Pearce: 1 locality (1899) Stenus (Stenus) boops Ljungh, 1810 First subsequent record: 1992 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1938 Latest record: 2020 Number of subsequent records: many Latest record: 2020

Stenus (Stenus) canaliculatus Gyllenhal, Stenus (Stenus) melanopus (Marsham, 1827 1802) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1966 First subsequent record: 1969 Number of subsequent records: 5 Number of subsequent records: many Latest record: 2009 Latest record: 2017

Stenus (Stenus) circularis Gravenhorst, Stenus (Stenus) nanus Stephens, 1833 1802 Pearce: 1 locality Pearce: 2 localities (1899, 1909, 1923) First subsequent record: 1998 First subsequent record: 2007 Number of subsequent records: many Number of subsequent records: 3 Latest record: 2019 Latest record: 2017 Stenus (Stenus) nitens Stephens, 1833 Stenus (Stenus) clavicornis (Scopoli, 1763) Pearce: no records Pearce: 3 localities First record: 1998 First subsequent record: 1963 Number of records: 8 Number of subsequent records: many Latest record: 2020 Latest record: 2020 Stenus (Stenus) providus Erichson, 1839 Stenus (Stenus) comma LeConte, 1863 Pearce: 3 localities Pearce: 1 locality First subsequent record: 1968 First subsequent record: 1989 Number of subsequent records: many Number of subsequent records: 2 Latest record: 2020 Latest record: 2005 Stenus (Stenus) pusillus Stephens, 1833 Stenus (Stenus) guttula Müller, P.W.J., Pearce: 8 localities 1821 First subsequent record: 1966 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1988 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Stenus (Stenus) subdepressus Mulsant & Rey, 1861 Stenus (Stenus) incrassatus Erichson, 1839 Pearce: no records Pearce: 1 locality First record: 2016 First subsequent record: 2005 Number of records: 1 Number of subsequent records: 2 Latest record: 2016 Latest record: 2015 Stenus (Tesnus) brunnipes Stephens, 1833 Stenus (Stenus) juno (Paykull, 1789) Pearce: generally common Pearce: 6 localities First subsequent record: 1964 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Stenus (Tesnus) crassus Stephens, 1833 Stenus (Stenus) lustrator Erichson, 1839 Pearce: 1 locality Pearce: no records First subsequent record: 1990 First record: 2019 Number of subsequent records: 2 Number of records: 1 Latest record: 1999 Latest record: 2019 Stenus (Tesnus) formicetorum Stenus (Stenus) melanarius Stephens, 1833 Mannerheim, 1843 Pearce: 1 locality Pearce: no records First subsequent record: 1998 First record: 1999 Number of subsequent records: many Number of records: 3 Latest record: 2014 Latest record: 2019 Stenus (Tesnus) nigritulus Gyllenhal, 1827 Lathrobium impressum Heer, 1841 Pearce: 2 localities Pearce: 2 localities First subsequent record: 1997 First subsequent record: 2006 Number of subsequent records: 1 Number of subsequent records: 5 Latest record: 1997 Latest record: 2014

Euaesthetus bipunctatus (Ljungh, 1804) Lathrobium longulum Gravenhorst, 1802 Pearce: no records Pearce: 1 locality First record: 1980 First subsequent record: 1980 Number of records: 4 Number of subsequent records: 7 Latest record: 2009 Latest record: 2019

Euaesthetus laeviusculus Mannerheim, Lathrobium pallidum Nordmann, 1837 1844 Pearce: no records Pearce: 1 locality (1925) First record: 1965 First subsequent record: 2020 Number of records: 1 Number of subsequent records: 1 Latest record: 1965 Latest record: 2020 Lobrathium multipunctum (Gravenhorst, Euaesthetus ruficapillus (Lacordaire, 1802) 1835) Pearce: 7 localities Pearce: 2 localities First subsequent record: 1964 First subsequent record: 2006 Number of subsequent records: many Number of subsequent records: many Latest record: 2016 Latest record: 2015 Tetartopeus angustatus (Lacordaire, 1835) Lathrobium brunnipes (Fabricius, 1792) Pearce: 4 localities Pearce: 8 localities First subsequent record: 1975 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Tetartopeus quadratus (Paykull, 1789) Lathrobium elongatum (Linnaeus, 1767) Pearce: 1 locality (1896) Pearce: 4 localities First subsequent record: 1962 First subsequent record: 1975 Number of subsequent records: 7 Number of subsequent records: many Latest record: 2017 Latest record: 2015 Tetartopeus rufonitidus (Reitter, 1909) Lathrobium fovulum Stephens, 1833 Pearce: no records Pearce: 1 locality First record: 1994 First subsequent record: 2006 Number of records: 2 Number of subsequent records: many Latest record: 2006 Latest record: 2020 Tetartopeus terminatus (Gravenhorst, Lathrobium fulvipenne (Gravenhorst, 1802) 1806) Pearce: 6 localities Pearce: 6 localities First subsequent record: 1954 First subsequent record: 1987 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Achenium depressum (Gravenhorst, 1802) Lathrobium geminum Kraatz, 1857 Pearce: 5 localities Pearce: 2 localities First subsequent record: 1993 First subsequent record: 1975 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2002 Latest record: 2017 Achenium humile (Nicolai, 1822) Pseudomedon obscurellus (Erichson, Pearce: 2 localities 1840) First subsequent record: 1988 Pearce: no records Number of subsequent records: 1 First record: 1997 Latest record: 1988 Number of records: 3 Latest record: 2003 Medon apicalis (Kraatz, 1857) Pearce: 1 locality Pseudomedon obsoletus (Nordmann, First subsequent record: 2008 1837) Number of subsequent records: 1 Pearce: 1 locality Latest record: 2008 No subsequent records

Medon brunneus (Erichson, 1839) Scopaeus laevigatus (Gyllenhal, 1827) Pearce: 3 localities Pearce: no records First subsequent record: 1968 First record: 1989 Number of subsequent records: many Number of records: 7 Latest record: 2019 Latest record: 2006

Medon fusculus (Mannerheim, 1830) Scopaeus minutus Erichson, 1840 Pearce: 1 locality (1886) Pearce: 2 localities First subsequent record: 1993 No subsequent records Number of subsequent records: 2 Latest record: 1994 Scopaeus sulcicollis (Stephens, 1833) Pearce: 2 localities Medon pocofer (Peyron, 1858) First subsequent record: 1991 Pearce: 1 locality Number of subsequent records: 4 First subsequent record: 1998 Latest record: 2015 Number of subsequent records: 3 Latest record: 2017 Rugilus angustatus (Geoffroy in Fourcroy, Sunius melanocephalus (Fabricius, 1792) 1785) Pearce: 2 localities Pearce: 2 localities (1873, 1891) First subsequent record: 1979 First subsequent record: 1999 Number of subsequent records: 8 Number of subsequent records: 1 Latest record: 2019 Latest record: 1999

Sunius propinquus (Brisout de Barneville, Rugilus erichsonii (Fauvel, 1867) Pearce: 3 localities C., 1867) First subsequent record: 1994 Pearce: 2 localities (1898, 1901) Number of subsequent records: many First subsequent record: 1979 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Rugilus geniculatus (Erichson, 1839) Pearce: 3 localities Lithocharis nigriceps Kraatz, 1859 First subsequent record: 1979 Pearce: no records Number of subsequent records: 3 First record: 1971 Latest record: 2020 Number of records: many Latest record: 2020 Rugilus orbiculatus (Paykull, 1789) Pearce: 8 localities Lithocharis ochracea (Gravenhorst, 1802) First subsequent record: 1954 Pearce: 1 locality (1885) Number of subsequent records: many First subsequent record: 2003 Latest record: 2019 Number of subsequent records: 6 Latest record: 2006

Rugilus rufipes Germar, 1836 Paederus (Eopaederus) caligatus Pearce: 4 localities Erichson, 1840 First subsequent record: 1970 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1938 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Rugilus similis (Erichson, 1839) Pearce: 1 locality Paederus (Heteropaederus) fuscipes First subsequent record: 2000 Curtis, 1826 Number of subsequent records: 1 Pearce: 7 localities Latest record: 2000 First subsequent record: 1987 Number of subsequent records: many Rugilus subtilis (Erichson, 1840) Latest record: 2020 Pearce: 1 locality No subsequent records Paederus (Paederus) riparius (Linnaeus, 1758) Astenus immaculatus Stephens, 1833 Pearce: fairly common and widely distributed Pearce: 1 locality (1901) First subsequent record: 1935 First subsequent record: 2013 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2020 Latest record: 2020 Paederus (Poederomorphus) littoralis Astenus lyonessius (Joy, 1908) Gravenhorst, 1802 Pearce: 5 localities Pearce: generally common and widely distributed First subsequent record: 1962 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Astenus procerus (Gravenhorst, 1806) Othius angustus Stephens, 1833 Pearce: 3 localities Pearce: 2 localities No subsequent records First subsequent record: 1975 Number of subsequent records: 4 Astenus pulchellus (Heer, 1839) Latest record: 2012 Pearce: 6 localities First subsequent record: 1999 Othius laeviusculus Stephens, 1833 Number of subsequent records: 2 Pearce: 3 localities Latest record: 2006 First subsequent record: 1990 Number of subsequent records: many Ochthephilum fracticorne (Paykull, 1800) Latest record: 2020 Pearce: 3 localities First subsequent record: 1968 Othius punctulatus (Goeze, 1777) Number of subsequent records: many Pearce: 4 localities Latest record: 2020 First subsequent record: 1968 Number of subsequent records: many Ochthephilum jacquelinii (Boieldieu, Latest record: 2019 1859) Pearce: no records Othius subuliformis Stephens, 1833 First record: 1981 Pearce: 6 localities Number of records: 3 First subsequent record: 1975 Latest record: 2005 Number of subsequent records: many Latest record: 2018 Atrecus affinis (Paykull, 1789) Gyrohypnus fracticornis (Müller, O.F., Pearce: 3 localities 1776) First subsequent record: 1975 Pearce: 12 localities Number of subsequent records: many First subsequent record: 1964 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Leptacinus batychrus (Gyllenhal, 1827) Pearce: 3 localities Gyrohypnus punctulatus (Paykull, 1789) First subsequent record: 2003 Pearce: no records Number of subsequent records: 1 First record: 1994 Latest record: 2003 Number of records: 7 Latest record: 2009 Leptacinus formicetorum Märkel, 1841 Pearce: no records Gyrohypnus wagneri (Scheerpeltz, 1926) First record: 2014 Pearce: no records Number of records: 2 First record: 2003 Latest record: 2014 Number of records: 1 Latest record: 2003 Leptacinus intermedius Donisthorpe, 1936 Pearce: no records Xantholinus (Polydontophallus) elegans First record: 2003 (Olivier, 1795) Number of records: 2 Pearce: 4 localities Latest record: 2011 First subsequent record: 2009 Number of subsequent records: 1 Leptacinus pusillus (Stephens, 1833) Latest record: 2009 Pearce: 1 locality First subsequent record: 1990 Xantholinus (Purrolinus) tricolor Number of subsequent records: many Latest record: 2020 (Fabricius, 1787) Pearce: no records First record: 1939 Phacophallus parumpunctatus (Gyllenhal, Number of records: 1 1827) Latest record: 1939 Pearce: 2 localities First subsequent record: 1995 Xantholinus (Xantholinus) gallicus Number of subsequent records: 3 Latest record: 2003 Coiffait, 1956 Pearce: no records First record: 1986 Nudobius lentus (Gravenhorst, 1806) Number of records: 8 Pearce: no records Latest record: 2015 First record: 2000 Number of records: 2 Latest record: 2004 Xantholinus (Xantholinus) linearis (Olivier, 1795) Pearce: 10 localities Gauropterus fulgidus (Fabricius, 1787) First subsequent record: 1936 Pearce: no records Number of subsequent records: many First record: 1990 Latest record: 2019 Number of records: 4 Latest record: 2013 Xantholinus (Xantholinus) longiventris Gyrohypnus angustatus Stephens, 1833 Heer, 1839 Pearce: 6 localities Pearce: 7 localities First subsequent record: 1963 First subsequent record: 1939 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Megalinus glabratus (Gravenhorst, 1802) Quedius (Microsaurus) cruentus (Olivier, Pearce: 6 localities 1795) First subsequent record: 1936 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1941 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Heterothops binotatus (Gravenhorst, 1802) Pearce: 4 localities Quedius (Microsaurus) fulgidus First subsequent record: 2005 (Fabricius, 1792) Number of subsequent records: 3 Pearce: 1 locality Latest record: 2006 First subsequent record: 1979 Number of subsequent records: 1 Heterothops niger Kraatz, 1868 Latest record: 1979 Pearce: no records First record: 1994 Quedius (Microsaurus) invreae Gridelli, Number of records: 4 Latest record: 1997 1924 Pearce: no records First record: 2007 Heterothops praevius Erichson, 1839 Number of records: 2 Pearce: no records Latest record: 2015 First record: 1969 Number of records: 1 Latest record: 1969 Quedius (Microsaurus) longicornis Kraatz, 1857 Pearce: 1 locality Euryporus picipes (Paykull, 1800) First subsequent record: 1995 Pearce: 1 locality Number of subsequent records: 4 No subsequent records Latest record: 2000

Quedius (Distichalius) cinctus (Paykull, Quedius (Microsaurus) maurus (Sahlberg, 1790) C.R., 1830) Pearce: 4 localities Pearce: no records First subsequent record: 1971 First record: 1970 Number of subsequent records: 5 Number of records: 3 Latest record: 2019 Latest record: 2013

Quedius (Microsaurus) aetolicus Kraatz, Quedius (Microsaurus) mesomelinus 1858 (Marsham, 1802) Pearce: no records Pearce: 5 localities First record: 2019 First subsequent record: 1987 Number of records: 1 Number of subsequent records: 8 Latest record: 2019 Latest record: 2020

Quedius (Microsaurus) brevicornis Quedius (Microsaurus) microps (Thomson, C.G., 1860) Gravenhorst, 1847 Pearce: no records Pearce: no records First record: 1998 First record: 1997 Number of records: 3 Number of records: 2 Latest record: 2020 Latest record: 2014

Quedius (Microsaurus) brevis Erichson, Quedius (Microsaurus) nigrocaeruleus 1840 Fauvel, 1876 Pearce: 1 locality Pearce: 1 locality (1928) First subsequent record: 1936 First subsequent record: 1997 Number of subsequent records: 1 Number of subsequent records: 1 Latest record: 1936 Latest record: 1997

Quedius (Microsaurus) scitus Quedius (Raphirus) boops (Gravenhorst, (Gravenhorst, 1806) 1802) Pearce: no records Pearce: 6 localities First record: 1987 First subsequent record: 1975 Number of records: many Number of subsequent records: many Latest record: 2015 Latest record: 2018

Quedius (Microsaurus) truncicola Quedius (Raphirus) fumatus (Stephens, Fairmaire & Laboulbène, 1856 1833) Pearce: no records Pearce: 3 localities First record: 2018 First subsequent record: 1968 Number of records: 1 Number of subsequent records: many Latest record: 2018 Latest record: 2019

Quedius (Microsaurus) xanthopus Quedius (Raphirus) humeralis Stephens, Erichson, 1839 1832 Pearce: 2 localities Pearce: no records First subsequent record: 1994 First record: 1975 Number of subsequent records: 4 Number of records: 7 Latest record: 2018 Latest record: 2011

Quedius (Quedius) curtipennis Bernhauer, Quedius (Raphirus) lateralis (Gravenhorst, 1908 1802) Pearce: no records (but see fuliginosus) Pearce: 1 locality (1919) First record: 1935 First subsequent record: 1966 Number of records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019

Quedius (Quedius) fuliginosus Quedius (Raphirus) maurorufus (Gravenhorst, 1802) (Gravenhorst, 1806) Pearce: 4 localities (but probably includes Pearce: 5 localities curtipennis) First subsequent record: 1989 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2017 Latest record: 2019 Quedius (Raphirus) nemoralis Baudi di Quedius (Quedius) levicollis Brullé, 1832 Selve, 1848 Pearce: generally common Pearce: no records First subsequent record: 1968 First record: 1993 Number of subsequent records: many Number of records: 5 Latest record: 2020 Latest record: 2009

Quedius (Quedius) molochinus Quedius (Raphirus) nigriceps Kraatz, 1857 (Gravenhorst, 1806) Pearce: 5 localities Pearce: fairly common and widely distributed First subsequent record: 1965 First subsequent record: 2000 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Quedius (Raphirus) nitipennis (Stephens, Quedius (Quedius) simplicifrons 1833) Fairmaire, 1862 Pearce: 3 localities Pearce: 2 localities First subsequent record: 1989 First subsequent record: 1935 Number of subsequent records: 6 Number of subsequent records: 4 Latest record: 2014 Latest record: 2019 Quedius (Raphirus) persimilis Mulsant & Quedius (Velleius) dilatatus (Fabricius, Rey, 1876 1787) Pearce: no records Pearce: no records First record: 1979 First record: 2019 Number of records: many Number of records: 1 Latest record: 2015 Latest record: 2019

Quedius (Raphirus) picipes (Mannerheim, Acylophorus glaberrimus (Herbst, 1784) 1830) Pearce: no records Pearce: generally common and widely distributed First record: 2016 First subsequent record: 1964 Number of records: 1 Number of subsequent records: many Latest record: 2016 Latest record: 2019 Platydracus fulvipes (Scopoli, 1763) Quedius (Raphirus) plancus Erichson, Pearce: no records 1840 First record: 1987 Pearce: no records Number of records: 3 First record: 2019 Latest record: 2014 Number of records: 1 Latest record: 2019 Platydracus latebricola (Gravenhorst, 1806) Quedius (Raphirus) schatzmayri Gridelli, Pearce: 3 localities 1922 First subsequent record: 1944 Pearce: no records Number of subsequent records: 5 First record: 1968 Latest record: 2020 Number of records: many Latest record: 2020 Platydracus stercorarius (Olivier, 1795) Pearce: 14 localities Quedius (Raphirus) scintillans First subsequent record: 1997 Number of subsequent records: many (Gravenhorst, 1806) Latest record: 2019 Pearce: 1 locality (1899) First subsequent record: 1975 Number of subsequent records: many Dinothenarus pubescens (De Geer, 1774) Latest record: 2018 Pearce: 1 locality No subsequent records Quedius (Raphirus) semiaeneus (Stephens, 1833) Tasgius (Rayacheila) globulifer (Geoffroy Pearce: 3 localities in Fourcroy, 1785) First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1939 Latest record: 2016 Number of subsequent records: many Latest record: 2020 Quedius (Raphirus) semiobscurus (Marsham, 1802) Tasgius (Rayacheila) melanarius (Heer, Pearce: 6 localities 1839) First subsequent record: 1938 Pearce: no records Number of subsequent records: many First record: 1979 Latest record: 2020 Number of records: many Latest record: 2019 Quedius (Raphirus) umbrinus Erichson, 1839 Tasgius (Rayacheila) morsitans (Rossi, Pearce: no records 1790) First record: 1986 Pearce: 8 localities Number of records: many First subsequent record: 1935 Latest record: 2017 Number of subsequent records: many Latest record: 2020 Tasgius (Rayacheila) winkleri (Bernhauer, Ocypus (Pseudocypus) fuscatus 1906) (Gravenhorst, 1802) Pearce: generally common and widely distributed Pearce: 3 localities First subsequent record: 1986 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: 2 Latest record: 2020 Latest record: 2005

Tasgius (Tasgius) ater (Gravenhorst, Staphylinus dimidiaticornis Gemminger, 1802) 1851 Pearce: widely distributed in south of county Pearce: 10 localities (listed as caesareus) First subsequent record: 1935 First subsequent record: 1986 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019

Tasgius (Tasgius) pedator (Gravenhorst, Staphylinus erythropterus Linnaeus, 1758 1802) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1982 First subsequent record: 1979 Number of subsequent records: 6 Number of subsequent records: 5 Latest record: 2019 Latest record: 1979 Creophilus maxillosus (Linnaeus, 1758) Ocypus (Angulephallus) brunnipes Pearce: generally common and widely distributed (Fabricius, 1781) First subsequent record: 1939 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1995 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Ontholestes murinus (Linnaeus, 1758) Pearce: 8 localities Ocypus (Matidus) nitens (Schrank, 1781) First subsequent record: 1934 Pearce: 1 locality Number of subsequent records: many No subsequent records Latest record: 2019

Ocypus (Ocypus) olens (Müller, O.F., Ontholestes tessellatus (Geoffroy in 1764) Fourcroy, 1785) Pearce: generally common and widely distributed Pearce: 7 localities First subsequent record: 1936 First subsequent record: 1986 Number of subsequent records: many Number of subsequent records: 4 Latest record: 2020 Latest record: 2019

Ocypus (Pseudocypus) aeneocephalus (De Emus hirtus (Linnaeus, 1758) Geer, 1774) Pearce: 3 localities Pearce: 10 localities No subsequent records First subsequent record: 1935 Number of subsequent records: many Neobisnius lathroboides (Baudi di Selve, Latest record: 2020 1848) Pearce: no records Ocypus (Pseudocypus) fortunatarum First record: 2017 Wollaston, 1871 Number of records: 1 Pearce: 1 locality Latest record: 2017 First subsequent record: 1998 Number of subsequent records: 3 Neobisnius procerulus (Gravenhorst, Latest record: 2019 1806) Pearce: 2 localities No subsequent records

Neobisnius villosulus (Stephens, 1833) Philonthus (Onychophilonthus) Pearce: 1 locality marginatus (Müller, O.F., 1764) First subsequent record: 1994 Pearce: 5 localities Number of subsequent records: 7 First subsequent record: 1963 Latest record: 2017 Number of subsequent records: many Latest record: 2017 Erichsonius cinerascens (Gravenhorst, 1802) Philonthus (Philonthus) addendus Sharp, Pearce: 2 localities 1867 First subsequent record: 1964 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1979 Latest record: 2019 Number of subsequent records: 5 Latest record: 2008 Erichsonius signaticornis (Mulsant & Rey, 1853) Philonthus (Philonthus) albipes Pearce: 2 localities (Gravenhorst, 1802) First subsequent record: 1989 Pearce: 1 locality (1909) Number of subsequent records: 5 First subsequent record: 1971 Latest record: 2006 Number of subsequent records: 7 Latest record: 2018 Bisnius cephalotes (Gravenhorst, 1802) Pearce: 6 localities Philonthus (Philonthus) atratus First subsequent record: 1997 (Gravenhorst, 1802) Number of subsequent records: 4 Pearce: no records Latest record: 2020 First record: 2020 Number of records: 1 Bisnius fimetarius (Gravenhorst, 1802) Latest record: 2020 Pearce: 5 localities First subsequent record: 1964 Philonthus (Philonthus) carbonarius Number of subsequent records: many Latest record: 2020 (Gravenhorst, 1802) Pearce: 3 localities First subsequent record: 1975 Bisnius nigriventris (Thomson, C.G., Number of subsequent records: many 1867) Latest record: 2020 Pearce: 1 locality First subsequent record: 1993 Philonthus (Philonthus) cognatus Number of subsequent records: 1 Latest record: 1993 Stephens, 1832 Pearce: no records First record: 1935 Bisnius puella (Nordmann, 1837) Number of records: many Pearce: no records Latest record: 2020 First record: 1968 Number of records: 1 Latest record: 1968 Philonthus (Philonthus) concinnus (Gravenhorst, 1802) Pearce: no records Bisnius sordidus (Gravenhorst, 1802) First record: 2019 Pearce: 5 localities Number of records: 3 First subsequent record: 1996 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Philonthus (Philonthus) cruentatus Bisnius subuliformis (Gravenhorst, 1802) (Gmelin, 1790) Pearce: 1 locality (1871) Pearce: 5 localities First subsequent record: 1987 First subsequent record: 1963 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2020 Latest record: 2020 Philonthus (Philonthus) debilis Philonthus (Philonthus) longicornis (Gravenhorst, 1802) Stephens, 1832 Pearce: 2 localities Pearce: 6 localities First subsequent record: 1995 First subsequent record: 1971 Number of subsequent records: 6 Number of subsequent records: 3 Latest record: 1999 Latest record: 2013

Philonthus (Philonthus) decorus Philonthus (Philonthus) mannerheimi (Gravenhorst, 1802) Fauvel, 1869 Pearce: 3 localities Pearce: no records First subsequent record: 1975 First record: 1968 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1968

Philonthus (Philonthus) discoideus Philonthus (Philonthus) micans (Gravenhorst, 1802) (Gravenhorst, 1802) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1990 First subsequent record: 1997 Number of subsequent records: 6 Number of subsequent records: 4 Latest record: 2006 Latest record: 2020

Philonthus (Philonthus) ebeninus Philonthus (Philonthus) micantoides (Gravenhorst, 1802) Benick & Lohse, 1956 Pearce: no records Pearce: no records First record: 2019 First record: 2006 Number of records: 1 Number of records: 2 Latest record: 2019 Latest record: 2007

Philonthus (Philonthus) fumarius Philonthus (Philonthus) nigrita (Gravenhorst, 1806) (Gravenhorst, 1806) Pearce: 2 localities Pearce: 3 localities First subsequent record: 2011 First subsequent record: 1987 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2014 Latest record: 2018

Philonthus (Philonthus) intermedius Philonthus (Philonthus) parvicornis (Lacordaire, 1835) (Gravenhorst, 1802) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1963 First subsequent record: 2015 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2019 Latest record: 2015

Philonthus (Philonthus) jurgans Philonthus (Philonthus) politus (Linnaeus, Tottenham, 1937 1758) Pearce: no records Pearce: generally common First record: 1993 First subsequent record: 1935 Number of records: 8 Number of subsequent records: many Latest record: 2020 Latest record: 2015

Philonthus (Philonthus) laminatus Philonthus (Philonthus) quisquiliarius (Creutzer, 1799) (Gyllenhal, 1810) Pearce: 6 localities Pearce: 4 localities First subsequent record: 1969 First subsequent record: 1939 Number of subsequent records: many Number of subsequent records: many Latest record: 2017 Latest record: 2020 Philonthus (Philonthus) rectangulus Philonthus (Philonthus) umbratilis Sharp, 1874 (Gravenhorst, 1802) Pearce: no records Pearce: no records First record: 1995 First record: 1989 Number of records: 2 Number of records: 5 Latest record: 2006 Latest record: 2019

Philonthus (Philonthus) rubripennis Philonthus (Philonthus) varians (Paykull, Stephens, 1832 1789) Pearce: no records Pearce: 8 localities First record: 2017 First subsequent record: 1963 Number of records: 4 Number of subsequent records: many Latest record: 2020 Latest record: 2020

Philonthus (Philonthus) rufipes (Stephens, Philonthus (Philonthus) ventralis 1832) (Gravenhorst, 1802) Pearce: 1 locality (1899) Pearce: 1 locality No subsequent records First subsequent record: 2003 Number of subsequent records: 1 Philonthus (Philonthus) sanguinolentus Latest record: 2003 (Gravenhorst, 1802) Pearce: 5 localities Gabronthus thermarum (Aubé, 1850) First subsequent record: 1955 Pearce: no records Number of subsequent records: many First record: 2003 Latest record: 2016 Number of records: 6 Latest record: 2013 Philonthus (Philonthus) spinipes Sharp, 1874 Gabrius appendiculatus Sharp, 1910 Pearce: no records Pearce: no records First record: 1997 First record: 1999 Number of records: 5 Number of records: 7 Latest record: 2006 Latest record: 2008

Philonthus (Philonthus) splendens Gabrius breviventer (Sperk, 1835) Pearce: 3 localities (Fabricius, 1792) First subsequent record: 1986 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2015 Gabrius keysianus Sharp, 1910 Pearce: 1 locality Philonthus (Philonthus) succicola First subsequent record: 2001 (Thomson, C.G., 1860) Number of subsequent records: 6 Pearce: no records Latest record: 2014 First record: 1975 Number of records: many Latest record: 2013 Gabrius nigritulus (Gravenhorst, 1802) Pearce: 8 localities First subsequent record: 1955 Philonthus (Philonthus) tenuicornis Number of subsequent records: many (Mulsant & Rey, 1853) Latest record: 2011 Pearce: 2 localities First subsequent record: 1986 Gabrius piliger Mulsant & Rey, 1876 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1975 Number of records: 4 Latest record: 2015 Gabrius splendidulus (Gravenhorst, 1802) Anoplotrupes stercorosus (Scriba, 1791) Pearce: 1 locality Pearce: generally common and widely distributed First subsequent record: 1975 First subsequent record: 1936 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Gabrius trossulus (Nordmann, 1837) Geotrupes mutator (Marsham, 1802) Pearce: 4 localities Pearce: widely distributed in coastal part of county First subsequent record: 1999 First subsequent record: 1951 Number of subsequent records: 3 Number of subsequent records: 8 Latest record: 2017 Latest record: 2019

Gabrius velox Sharp, 1910 Geotrupes spiniger (Marsham, 1802) Pearce: 1 locality Pearce: generally common First subsequent record: 1996 First subsequent record: 1994 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2015 Latest record: 2020

Cafius (Cafius) fucicola Curtis, 1830 Geotrupes stercorarius (Linnaeus, 1758) Pearce: 4 localities Pearce: generally common and widely distributed First subsequent record: 1975 First subsequent record: 1935 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2006 Latest record: 2020

Cafius (Cafius) xantholoma (Gravenhorst, Trypocopris pyrenaeus (Charpentier, 1806) 1825) Pearce: very common along coast Pearce: 4 localities First subsequent record: 1975 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019

Cafius (Platygonium) cicatricosus 21. Family TROGIDAE Macleay, 1819 (Erichson, 1840) Pearce: 1 locality Trox perlatus (Goeze, 1777) No subsequent records Pearce: 1 locality No subsequent records Remus sericeus Holme, 1837 Pearce: 8 localities Trox sabulosus (Linnaeus, 1758) First subsequent record: 1966 Pearce: 6 localities Number of subsequent records: 5 First subsequent record: 1940 Latest record: 2020 Number of subsequent records: 1 Latest record: 1940 20. Family GEOTRUPIDAE Latreille, 1802 Trox scaber (Linnaeus, 1767) Pearce: 3 localities Odonteus armiger (Scopoli, 1772) First subsequent record: 1950 Pearce: 2 localities Number of subsequent records: 3 No subsequent records Latest record: 2013

Typhaeus typhoeus (Linnaeus, 1758) 22. Family LUCANIDAE Latreille, 1804 Pearce: very common on heaths First subsequent record: 1935 Sinodendron cylindricum (Linnaeus, 1758) Number of subsequent records: many Pearce: 4 localities Latest record: 2020 First subsequent record: 1975 Number of subsequent records: many Latest record: 2019 Dorcus parallelipipedus (Linnaeus, 1758) Aphodius fimetarius (Linnaeus, 1758) Pearce: 7 localities Pearce: generally common and widely distributed First subsequent record: 1939 (but many of these may be pedellus) Number of subsequent records: many First subsequent record: 1936 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Lucanus cervus (Linnaeus, 1758) Pearce: 6 localities Aphodius foetidus (Herbst, 1783) First subsequent record: 1945 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1935 Latest record: 2020 Number of subsequent records: many Latest record: 2018 23. Family SCARABAEIDAE Latreille, 1802 Aphodius pedellus (De Geer, 1774) Pearce: no records (but see fimetarius) First record: 1935 Aegialia arenaria (Fabricius, 1787) Number of records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1934 Number of subsequent records: many Latest record: 2020 Bodiloides ictericus (Laicharting, 1781) [2012: Aphodius ictericus] Acrossus depressus (Kugelann, 1792) Pearce: 2 localities First subsequent record: 1963 [2012: Aphodius depressus] Number of subsequent records: many Pearce: 4 localities (but at least one is misidentified Latest record: 2019 luridus – DJ Mann) First subsequent record: 2008 Number of subsequent records: 1 Bodilopsis rufa (Moll, 1782) [2012: Latest record: 2008 Aphodius rufus] Pearce: 2 localities Acrossus luridus (Fabricius, 1775) [2012: First subsequent record: 1975 Number of subsequent records: many Aphodius luridus] Latest record: 2019 Pearce: 9 localities First subsequent record: 1978 Number of subsequent records: many Bodilopsis sordida (Fabricius, 1775) Latest record: 2015 [2012: Aphodius sordidus] Pearce: 1 locality Acrossus rufipes (Linnaeus, 1758) [2012: No subsequent records Aphodius rufipes] Pearce: generally common Calamosternus granarius (Linnaeus, First subsequent record: 1936 1767) [2012: Aphodius granarius] Number of subsequent records: many Pearce: generally common Latest record: 2020 First subsequent record: 1950 Number of subsequent records: many Agrilinus ater (De Geer, 1774) [2012: Latest record: 2019 Aphodius ater] Pearce: generally common Chilothorax conspurcatus (Linnaeus, First subsequent record: 1939 1758) [2012: Aphodius conspurcatus] Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 2007 Number of records: 6 Agrilinus constans (Duftschmid, 1805) Latest record: 2017 [2012: Aphodius constans] Pearce: 2 localities First subsequent record: 1999 Number of subsequent records: many Latest record: 2020 Chilothorax distinctus (Müller, O.F., Heptaulacus testudinarius (Fabricius, 1776) [2012: Aphodius distinctus] 1775) Pearce: 3 localities Pearce: 2 localities First subsequent record: 1968 First subsequent record: 1930 Number of subsequent records: 3 Number of subsequent records: 2 Latest record: 2015 Latest record: 1931

Chilothorax paykulli (Bedel, 1908) [2012: Limarus zenkeri (Germar, 1813) [2012: Aphodius paykulli] Aphodius zenkeri] Pearce: 1 locality Pearce: no records No subsequent records First record: 2007 Number of records: 9 Colobopterus erraticus (Linnaeus, 1758) Latest record: 2017 [2012: Aphodius erraticus] Pearce: 5 localities Liothorax niger (Illiger, 1798) [2012: First subsequent record: 1983 Aphodius ater] Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1913 Number of records: 2 Esymus merdarius (Fabricius, 1775) Latest record: 1915 [2012: Aphodius merdarius] Pearce: 5 localities Liothorax plagiatus (Linnaeus, 1767) First subsequent record: 1964 [2012: Aphodius plagiatus] Number of subsequent records: many Pearce: 2 localities Latest record: 2015 First subsequent record: 1954 Number of subsequent records: many Esymus pusillus (Herbst, 1789) [2012: Latest record: 2015 Aphodius pusillus] Pearce: 3 localities Melinopterus prodromus (Brahm, 1790) First subsequent record: 1954 [2012: Aphodius prodromus] Number of subsequent records: many Pearce: 4 localities Latest record: 2019 First subsequent record: 1975 Number of subsequent records: many Euheptaulacus sus (Herbst, 1783) Latest record: 2020 Pearce: 1 locality No subsequent records Melinopterus sphacelatus (Panzer, 1798) [2012: Aphodius sphacelatus] Euheptaulacus villosus (Gyllenhal in Pearce: generally common Schönherr, 1806) First subsequent record: 1936 Pearce: no records Number of subsequent records: many First record: 1939 Latest record: 2020 Number of records: 2 Latest record: 2008 Nimbus contaminatus (Herbst, 1783) [2012: Aphodius contaminatus] Euorodalus coenosus (Panzer, 1798) Pearce: 4 localities [2012: Aphodius coenosus] First subsequent record: 1938 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1979 Latest record: 2017 Number of subsequent records: many Latest record: 2019 Nimbus obliteratus (Panzer, 1823) [2012: Aphodius obliteratus] Eupleurus subterraneus (Linnaeus, 1758) Pearce: 2 localities (but misidentifications – DJ [2012: Aphodius subterraneus] Mann) Pearce: 1 locality First subsequent record: 2014 No subsequent records Number of subsequent records: 8 Latest record: 2019 Otophorus haemorrhoidalis (Linnaeus, Copris lunaris (Linnaeus, 1758) 1758) [2012: Aphodius haemorrhoidalis] Pearce: 1 locality Pearce: 6 localities First subsequent record: 1954 First subsequent record: 1938 Number of subsequent records: 1 Number of subsequent records: many Latest record: 1954 Latest record: 2018 Onthophagus coenobita (Herbst, 1783) Oxyomus sylvestris (Scopoli, 1763) Pearce: 5 localities Pearce: “Dorset” (A. Ford) First subsequent record: 1939 First subsequent record: 1945 Number of subsequent records: many Number of subsequent records: 3 Latest record: 2020 Latest record: 2007 Onthophagus joannae Goljan, 1953 Plagiogonus arenarius (Olivier, 1789) Pearce: 7 localities [2012: Aphodius arenarius] First subsequent record: 1935 Pearce: 1 locality (1921) Number of subsequent records: many No subsequent records Latest record: 2019

Planolinoides borealis (Gyllenhal, 1827) Onthophagus medius (Kugelann, 1792) Pearce: 4 localities [2012: Aphodius borealis] First subsequent record: 1950 Pearce: no records Number of subsequent records: many First record: 1926 Latest record: 2019 Number of records: many Latest record: 2017 Onthophagus nuchicornis (Linnaeus, Rhodaphodius foetens (Fabricius, 1787) 1758) Pearce: 7 localities (but all vouchers checked are [2012: Aphodius foetens] similis – DJ Mann) Pearce: 8 localities (but 4 are from First record: 1864 misidentifications – DJ Mann) Number of records: 5 First subsequent record: 1939 Latest record: 1954 Number of subsequent records: many Latest record: 2017 Onthophagus similis (Scriba, 1790) Pearce: no records (but see nuchicornis) Sigorus porcus (Fabricius, 1792) [2012: First record: 1936 Aphodius porcus] Number of records: many Pearce: 3 localities Latest record: 2020 First subsequent record: 1975 Number of subsequent records: 1 Latest record: 1975 Hoplia philanthus (Fuessly, 1775) Pearce: 8 localities First subsequent record: 1954 Teuchestes fossor (Linnaeus, 1758) [2012: Number of subsequent records: many Aphodius fossor] Latest record: 2019 Pearce: generally common and widely distributed First subsequent record: 1939 Melolontha melolontha (Linnaeus, 1758) Number of subsequent records: many Pearce: generally common and widely distributed Latest record: 2019 First subsequent record: 1968 Number of subsequent records: many Volinus sticticus (Panzer, 1798) [2012: Latest record: 2020 Aphodius sticticus] Pearce: 3 localities Amphimallon fallenii (Gyllenhal, 1817) First subsequent record: 1938 Pearce: no records Number of subsequent records: many First record: 1986 Latest record: 2019 Number of records: 7 Latest record: 1996

Amphimallon solstitiale (Linnaeus, 1758) Clambus armadillo (De Geer, 1774) Pearce: fairly common and widely distributed Pearce: no records First subsequent record: 1942 First record: 2000 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 2000

Omaloplia ruricola (Fabricius, 1775) Clambus pubescens Redtenbacher, 1847 Pearce: 5 localities Pearce: 1 locality First subsequent record: 1994 First subsequent record: 1971 Number of subsequent records: many Number of subsequent records: 5 Latest record: 2015 Latest record: 2018

Serica brunnea (Linnaeus, 1758) Clambus punctulum (Beck, 1817) Pearce: 11 localities Pearce: no records First subsequent record: 1939 First record: 2005 Number of subsequent records: many Number of records: 1 Latest record: 2019 Latest record: 2005

Anomala dubia (Scopoli, 1763) 26. Family SCIRTIDAE Fleming, 1821 Pearce: 4 localities First subsequent record: 1935 Elodes elongatus Tournier, 1868 Number of subsequent records: many Pearce: no records Latest record: 2018 First subsequent record: 2002 Number of subsequent records: 4 Phyllopertha horticola (Linnaeus, 1758) Latest record: 2011 Pearce: generally common First subsequent record: 1935 Elodes minutus (Linnaeus, 1767) Number of subsequent records: many Pearce: 5 localities Latest record: 2019 First subsequent record: 1964 Number of subsequent records: 6 Cetonia aurata (Linnaeus, 1758) Latest record: 2017 Pearce: 7 localities First subsequent record: 1943 Elodes pseudominutus Klausnitzer, 1971 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2005 Number of records: many 24. Family EUCINETIDAE Lacordaire, Latest record: 2016 1857 Odeles marginata (Fabricius, 1798) Nycteus meridionalis Laporte, 1838 [2012: Pearce: 3 localities Eucinetus meridionalis] First subsequent record: 1987 Pearce: no records Number of subsequent records: many First record: 1991 Latest record: 2017 Number of records: 4 Latest record: 2004 Microcara testacea (Linnaeus, 1767) Pearce: generally common 25. Family CLAMBIDAE Fischer von First subsequent record: 1967 Number of subsequent records: many Waldheim, 1821 Latest record: 2020

Calyptomerus dubius (Marsham, 1802) Contacyphon coarctatus (Paykull, 1799) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1975 First subsequent record: 1959 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2003 Latest record: 2020

Contacyphon hilaris (Nyholm, 1944) Scirtes hemisphaericus (Linnaeus, 1758) Pearce: no records Pearce: 4 localities First record: 1935 First subsequent record: 1975 Number of records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Contacyphon laevipennis (Tournier, 1868) 27. Family DASCILLIDAE Guérin- Pearce: no records Méneville, 1843 (1834) First record: 1992 Number of records: many Latest record: 2019 Dascillus cervinus (Linnaeus, 1758) Pearce: 3 localities First subsequent record: 1943 Contacyphon ochraceus (Stephens, 1830) Number of subsequent records: many Pearce: 2 localities Latest record: 2019 First subsequent record: 1939 Number of subsequent records: many Latest record: 2019 28. Family BUPRESTIDAE Leach, 1815

Contacyphon padi (Linnaeus, 1758) Melanophila acuminata (De Geer, 1774) Pearce: 9 localities Pearce: 1 locality First subsequent record: 1966 No subsequent records Number of subsequent records: many Latest record: 2019 Agrilus angustulus (Illiger, 1803) Pearce: 2 localities Contacyphon palustris (Thomson, C.G., First subsequent record: 1939 Number of subsequent records: many 1855) Latest record: 2019 Pearce: 5 localities First subsequent record: 1999 Number of subsequent records: 8 Agrilus biguttatus (Fabricius, 1777) Latest record: 2011 Pearce: no records First record: 1998 Number of records: 1 Contacyphon pubescens (Fabricius, 1792) Latest record: 1998 Pearce: no records First record: 2005 Number of records: 1 Agrilus laticornis (Illiger, 1803) Latest record: 2005 Pearce: 2 localities First subsequent record: 1963 Number of subsequent records: many Contacyphon variabilis (Thunberg, 1787) Latest record: 2019 Pearce: generally common First subsequent record: 1990 Number of subsequent records: many Agrilus sinuatus (Olivier, 1790) Latest record: 2018 Pearce: no records First record: 1951 Number of records: many Prionocyphon serricornis (Müller, P.W.J., Latest record: 2010 1821) Pearce: 1 locality (1841) First subsequent record: 1979 Agrilus viridis (Linnaeus, 1758) Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 1988 Number of records: 2 Latest record: 2019 Hydrocyphon deflexicollis (Müller, P.W.J., 1821) Aphanisticus emarginatus (Olivier, 1790) Pearce: 1 locality Pearce: no records No subsequent records First record: 1953 Number of records: 3 Latest record: 2013 Aphanisticus pusillus (Olivier, 1790) Porcinolus murinus (Fabricius, 1794) Pearce: 3 localities Pearce: 2 localities First subsequent record: 1940 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Trachys compressus Abeille de Perrin, Chaetophora spinosa (Rossi, 1794) 1891 [2012: Trachys subglaber] Pearce: 3 localities Pearce: no records (but the record of troglodytes, First subsequent record: 1991 and subsequent records of troglodytes could refer Number of subsequent records: 7 to this species) Latest record: 2017 First record: 1938 Number of records: many Curimopsis maritima (Marsham, 1802) Latest record: 2020 Pearce: 1 locality First subsequent record: 1989 Trachys minutus (Linnaeus, 1758) Number of subsequent records: 2 Pearce: 1 locality (1834) Latest record: 1998 No subsequent records Curimopsis setigera (Illiger, 1798) Trachys scrobiculatus Kiesenwetter, 1857 Pearce: 1 locality Pearce: 2 localities First subsequent record: 1982 First subsequent record: 1952 Number of subsequent records: 5 Number of subsequent records: many Latest record: 2016 Latest record: 2017 30. Family ELMIDAE Curtis, 1830 29. Family BYRRHIDAE Latreille, 1804 Elmis aenea (Müller, P.W.J., 1806) Simplocaria semistriata (Fabricius, 1794) Pearce: 6 localities Pearce: 6 localities First subsequent record: 1940 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019 Esolus parallelepipedus (Müller, P.W.J., Byrrhus fasciatus (Forster, 1771) 1806) Pearce: 2 localities Pearce: no records First subsequent record: 1937 First record: 1987 Number of subsequent records: many Number of records: 7 Latest record: 2019 Latest record: 2011

Byrrhus pilula (Linnaeus, 1758) Limnius volckmari (Panzer, 1793) Pearce: generally common and widely distributed Pearce: 3 localities First subsequent record: 1935 First subsequent record: 1965 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Byrrhus pustulatus (Forster, 1771) Oulimnius major (Rey, 1889) Pearce: 3 localities Pearce: no records First subsequent record: 1940 First record: 1982 Number of subsequent records: many Number of records: 3 Latest record: 2018 Latest record: 2006

Cytilus sericus (Forster, 1771) Oulimnius troglodytes (Gyllenhal, 1827) Pearce: common Pearce: no records First subsequent record: 1939 First record: 1963 Number of subsequent records: many Number of records: 9 Latest record: 2019 Latest record: 2006 Oulimnius tuberculatus (Müller, P.W.J., Dryops similaris Bollow, 1936 1806) Pearce: no records Pearce: 2 localities First record: 2013 First subsequent record: 1974 Number of records: 3 Number of subsequent records: many Latest record: 2020 Latest record: 2020 Dryops striatellus (Fairmaire & Brisout de Riolus nitens (Müller, P.W.J., 1817) Barnville, C., 1859) [2012: Normandia nitens] Pearce: 5 localities Pearce: 1 locality First subsequent record: 2000 No subsequent records Number of subsequent records: many Latest record: 2020 Riolus subviolaceus (Müller, P.W.J., 1817) Pearce: 1 locality 32. Family LIMNICHIDAE Erichson, First subsequent record: 1975 1846 Number of subsequent records: many Latest record: 2020 Limnichus pygmaeus (Sturm, 1807) Pearce: 2 localities (1886, 1904) Stenelmis canaliculata (Gyllenhal, 1808) First subsequent record: 1966 Pearce: no records Number of subsequent records: many First record: 1991 Latest record: 2017 Number of records: 4 Latest record: 2013 33. Family HETEROCERIDAE MacLeay, 1825 Macronychus quadrituberculatus Müller, P.W.J., 1806 Augyles maritimus (Guérin-Méneville, Pearce: 1 locality (1923) First subsequent record: 1975 1844) Number of subsequent records: 3 Pearce: 1 locality Latest record: 2010 No subsequent records

31. Family DRYOPIDAE Bilberg, 1820 Heterocerus fenestratus (Thunberg, 1784) Pearce: 2 localities (1817) First subsequent record: 1993 Number of subsequent records: many Dryops auriculatus (Geoffroy in Fourcroy, Latest record: 2020 1785) Pearce: 4 localities Heterocerus flexuosus Stephens, 1828 First subsequent record: 2007 Pearce: 2 localities Number of subsequent records: 1 First subsequent record: 1939 Latest record: 2007 Number of subsequent records: 6 Latest record: 2018 Dryops ernesti Gozis, 1886 Pearce: 2 localities Heterocerus fossor Kiesenwetter, 1843 First subsequent record: 1936 Pearce: no records Number of subsequent records: many First record: 1989 Latest record: 2019 Number of records: 3 Latest record: 2011 Dryops luridus (Erichson, 1847) Pearce: 1 locality Heterocerus fusculus Kiesenwetter, 1843 First subsequent record: 1981 Pearce: no records Number of subsequent records: many First record: 1980 Latest record: 2019 Number of records: 2 Latest record: 1995 Heterocerus marginatus (Fabricius, 1787) Eucnemis capucina Ahrens, 1812 Pearce: 3 localities Pearce: no records First subsequent record: 1989 First record: 2018 Number of subsequent records: many Number of records: 2 Latest record: 2017 Latest record: 2019

Heterocerus obsoletus Curtis, 1828 37. Family THROSCIDAE Laporte, Pearce: 1 locality (1896) 1840 No subsequent records Trixagus carinifrons (de Bonvouloir, 34. Family PSEPHENIDAE Lacordaire, 1859) 1854 Pearce: 1 locality First subsequent record: 1990 Eubria palustris (Germar, 1818) Number of subsequent records: many Pearce: 2 localities Latest record: 2016 First subsequent record: 1989 Number of subsequent records: many Trixagus dermestoides (Linnaeus, 1767) Latest record: 2017 Pearce: 3 localities First subsequent record: 1953 36. Family EUCNEMIDAE Eschscholtz, Number of subsequent records: many 1829 Latest record: 2020

Microrhagus pygmaeus (Fabricius, 1792) Trixagus gracilis Wollaston, 1854 Pearce: 2 localities (1839, 1843) Pearce: no records First subsequent record: 1986 First record: 2019 Number of subsequent records: many Number of records: 1 Latest record: 2019 Latest record: 2019

Epiphanis cornutus Eschscholtz, 1829 Trixagus obtusus (Curtis, 1827) Pearce: no records Pearce: 1 locality (1886) First record: 2007 First subsequent record: 1993 Number of records: 8 Number of subsequent records: many Latest record: 2019 Latest record: 2019

Hylis cariniceps (Reitter, 1902) 38. Family ELATERIDAE Leach, 1815 Pearce: no records First record: 1976 Agrypnus murinus (Linnaeus, 1758) Number of records: 9 Pearce: generally common and widely distributed Latest record: 2019 First subsequent record: 1933 Number of subsequent records: many Hylis olexai (Palm, 1955) Latest record: 2019 Pearce: no records First record: 2016 Agriotes acuminatus (Stephens, 1830) Number of records: 7 Pearce: 3 localities Latest record: 2019 First subsequent record: 1935 Number of subsequent records: many Melasis buprestoides (Linnaeus, 1760) Latest record: 2019 Pearce: no records First record: 1975 Agriotes lineatus (Linnaeus, 1767) Number of records: 4 Pearce: 3 localities Latest record: 2008 First subsequent record: 1934 Number of subsequent records: many Latest record: 2019

Agriotes obscurus (Linnaeus, 1758) Sericus brunneus (Linnaeus, 1758) Pearce: generally common Pearce: 3 localities First subsequent record: 1939 First subsequent record: 1998 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Agriotes pallidulus (Illiger, 1807) Procraerus tibialis (Lacordaire, 1835) Pearce: generally common Pearce: no records First subsequent record: 1934 First record: 1993 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1993

Agriotes sordidus (Illiger, 1807) Adrastus pallens (Fabricius, 1792) Pearce: 3 localities Pearce: 3 localities First subsequent record: 1991 First subsequent record: 1968 Number of subsequent records: 8 Number of subsequent records: many Latest record: 2011 Latest record: 2019

Agriotes sputator (Linnaeus, 1758) Melanotus castanipes (Paykull, 1800) Pearce: 8 localities Pearce: no records (but see villosus) First subsequent record: 1934 First record: 1934 Number of subsequent records: many Number of records: many Latest record: 2020 Latest record: 2019

Dalopius marginatus (Linnaeus, 1758) Melanotus villosus (Geoffroy in Fourcroy, Pearce: 9 localities 1785) First subsequent record: 1934 Pearce: common in woods (but could include Number of subsequent records: many castanipes) Latest record: 2020 First subsequent record: 1945 Number of subsequent records: many Ampedus balteatus (Linnaeus, 1758) Latest record: 2019 Pearce: 2 localities First subsequent record: 1968 Hypnoidus riparius (Fabricius, 1792) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1988 Number of subsequent records: many Ampedus cinnabarinus (Eschscholtz, Latest record: 2017 1829) Pearce: no records Athous (Athous) haemorrhoidalis First record: 2019 (Fabricius, 1801) Number of records: 1 Pearce: generally common Latest record: 2019 First subsequent record: 1933 Number of subsequent records: many Ampedus elongatulus (Fabricius, 1787) Latest record: 2020 Pearce: 1 locality First subsequent record: 1952 Athous (Athous) vittatus (Fabricius, 1792) Number of subsequent records: many Pearce: generally common Latest record: 2019 First subsequent record: 1936 Number of subsequent records: many Ampedus sanguinolentus (Schrank, 1776) Latest record: 2018 Pearce: 3 localities First subsequent record: 1933 Number of subsequent records: many Latest record: 2015 Athous (Orthathous) bicolor (Goeze, Aplotarsus incanus (Gyllenhal, 1827) 1777) Pearce: 4 localities Pearce: 11 localities First subsequent record: 1992 First subsequent record: 1939 Number of subsequent records: many Number of subsequent records: many Latest record: 2013 Latest record: 2020 Calambus bipustulatus (Linnaeus, 1767) Athous (Orthathous) campyloides Pearce: 3 localities Newman, 1833 First subsequent record: 1939 Pearce: no records Number of subsequent records: 7 First record: 1992 Latest record: 2018 Number of records: 4 Latest record: 2019 Ctenicera cuprea (Fabricius, 1775) Pearce: 1 locality Cidnopus aeruginosus (Olivier, 1790) No subsequent records Pearce: 1 locality First subsequent record: 2008 Paraphotistus nigricornis (Panzer, 1799) Number of subsequent records: 1 Pearce: 2 localities Latest record: 2008 First subsequent record: 2006 Number of subsequent records: 4 Limonius poneli Leseigneur & Mertlik, Latest record: 2018 2007 [2012: Kibunea minuta] Pearce: 4 localities Prosternon tessellatum (Linnaeus, 1758) First subsequent record: 1939 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1934 Latest record: 2020 Number of subsequent records: many Latest record: 2009 Denticollis linearis (Linnaeus, 1758) Pearce: 5 localities Selatosomus aeneus (Linnaeus, 1758) First subsequent record: 1939 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1982 Latest record: 2020 Number of subsequent records: 2 Latest record: 2000 Hemicrepidius hirtus (Herbst, 1784) Pearce: 5 localities Oedostethus quadripustulatus (Fabricius, First subsequent record: 1936 1792) Number of subsequent records: many Pearce: 1 locality Latest record: 2019 No subsequent records

Stenagostus rhombeus (Olivier, 1790) Cardiophorus asellus Erichson, 1840 Pearce: no records Pearce: 5 localities First record: 1991 First subsequent record: 1933 Number of records: 8 Number of subsequent records: many Latest record: 2013 Latest record: 2018

Actenicerus sjaelandicus (Müller, O.F., 39. Family DRILIDAE Blanchard, 1845 1764) Pearce: 5 localities Drilus flavescens (Geoffroy in Fourcroy, First subsequent record: 1933 1785) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1940 Number of subsequent records: 4 Latest record: 2006 40. Family LYCIDAE Laporte, 1836 Cantharis (Cantharis) fusca Linnaeus, 1758 Platycis minutus (Fabricius, 1787) Pearce: 3 localities Pearce: no records First subsequent record: 1977 First record: 1979 Number of subsequent records: many Number of records: many Latest record: 2018 Latest record: 2018 Cantharis (Cantharis) livida Linnaeus, 41. Family LAMPYRIDAE Rafinesque, 1758 1815 Pearce: 5 localities First subsequent record: 1936 Number of subsequent records: many Lampyris noctiluca (Linnaeus, 1758) Latest record: 2020 Pearce: generally common First subsequent record: 1952 Number of subsequent records: many Cantharis (Cantharis) nigra (De Geer, Latest record: 2020 1774) [2012: Cantharis thoracica] Pearce: 5 localities 42. Family CANTHARIDAE Imhoff, First subsequent record: 1951 Number of subsequent records: many 1856 (1815) Latest record: 2019

Podabrus alpinus (Paykull, 1798) Cantharis (Cantharis) nigricans (Müller, Pearce: 3 localities First subsequent record: 1990 O.F., 1776) Number of subsequent records: 6 Pearce: generally common Latest record: 2011 First subsequent record: 1939 Number of subsequent records: many Latest record: 2014 Cantharis (Cantharis) cryptica Ashe, 1947 Pearce: bicolor, described as common, could refer to this species Cantharis (Cantharis) pallida Goeze, 1777 First record: 1935 Pearce: bicolor, described as common, could refer Number of records: many to this species Latest record: 2020 First record: 1939 Number of records: many Latest record: 2019 Cantharis (Cantharis) decipiens Baudi di Selve, 1872 Cantharis (Cantharis) pellucida Fabricius, Pearce: 3 localities First subsequent record: 1955 1792 Number of subsequent records: many Pearce: generally common Latest record: 2020 First subsequent record: 1939 Number of subsequent records: many Latest record: 2019 Cantharis (Cantharis) figurata Mannerheim, 1843 Cantharis (Cantharis) rufa Linnaeus, 1758 Pearce: no records Pearce: 5 localities First record: 1987 First subsequent record: 1936 Number of records: 4 Number of subsequent records: many Latest record: 1987 Latest record: 2019

Cantharis (Cantharis) flavilabris Fallén, Cantharis (Cantharis) rustica Fallén, 1807 1807 [2012: Cantharis nigra] Pearce: generally common Pearce: generally common and widely distributed First subsequent record: 1968 First subsequent record: 1982 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Cantharis (Cyrtomoptila) lateralis Malthinus flaveolus (Herbst, 1786) Linnaeus, 1758 Pearce: 3 localities Pearce: 2 localities First subsequent record: 1968 First subsequent record: 1967 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Malthinus frontalis (Marsham, 1802) Rhagonycha fulva (Scopoli, 1763) Pearce: 1 locality Pearce: abundant First subsequent record: 1954 First subsequent record: 1934 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2010 Latest record: 2020 Malthinus seriepunctatus Kiesenwetter, Rhagonycha lignosa (Müller, O.F., 1764) 1852 Pearce: generally common Pearce: generally common First subsequent record: 1939 First subsequent record: 1955 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Rhagonycha lutea (Müller, O.F., 1764) Malthodes dispar (Germar, 1824) Pearce: 1 locality Pearce: 1 locality No subsequent records First subsequent record: 1984 Number of subsequent records: many Rhagonycha nigriventris (Thomson, C.G., Latest record: 2015 1864) [2012: Rhagonycha limbata] Pearce: generally common Malthodes fibulatus Kiesenwetter, 1852 First subsequent record: 1954 Pearce: no records Number of subsequent records: many First record: 1982 Latest record: 2019 Number of records: 2 Latest record: 1995 Rhagonycha testacea (Linnaeus, 1758) Pearce: 2 localities Malthodes flavoguttatus Kiesenwetter, First subsequent record: 1968 1852 Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 2008 Number of subsequent records: 2 Rhagonycha translucida (Krynicki, 1832) Latest record: 2008 Pearce: 1 locality No subsequent records Malthodes fuscus (Waltl, 1838) Pearce: 2 localities Silis ruficollis (Fabricius, 1775) First subsequent record: 1989 Pearce: “Dorset” (A. Ford) Number of subsequent records: 5 First subsequent record: 1968 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Malthodes guttifer Kiesenwetter, 1852 Pearce: 3 localities Malthinus balteatus Suffrian, 1851 First subsequent record: 1987 Pearce: 2 localities Number of subsequent records: 8 First subsequent record: 1954 Latest record: 2008 Number of subsequent records: many Latest record: 2020 Malthodes marginatus (Latreille, 1806) Pearce: 6 localities First subsequent record: 1955 Number of subsequent records: many Latest record: 2020

Malthodes minimus (Linnaeus, 1758) Anthrenus (Florilinus) museorum Pearce: 5 localities (Linnaeus, 1760) First subsequent record: 1982 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1941 Latest record: 2020 Number of subsequent records: 1 Latest record: 1941 Malthodes mysticus Kiesenwetter, 1852 Pearce: 1 locality Anthrenus (Helocerus) fuscus Olivier, First subsequent record: 2010 1790 Number of subsequent records: 1 Pearce: 6 localities Latest record: 2010 First subsequent record: 1993 Number of subsequent records: 9 Malthodes pumilus (Brébisson, 1835) Latest record: 2019 Pearce: 2 localities First subsequent record: 1992 Anthrenus (Nathrenus) verbasci Number of subsequent records: many Latest record: 2011 (Linnaeus, 1767) Pearce: 1 locality First subsequent record: 1994 44. Family DERMESTIDAE Latreille, Number of subsequent records: many 1804 Latest record: 2019

Dermestes (Dermestes) lardarius Ctesias serra (Fabricius, 1792) Linnaeus, 1758 Pearce: no records Pearce: 4 localities First record: 1988 No subsequent records Number of records: 5 Latest record: 2004 Dermestes (Dermestinus) maculatus De Geer, 1774 45. Family BOSTRICHIDAE Latreille, Pearce: no records 1802 First record: 1997 Number of records: 1 Bostrichus capucinus (Linnaeus, 1758) Latest record: 1997 Pearce: 1 locality (1854) No subsequent records Dermestes (Dermestinus) murinus Linnaeus, 1758 Rhyzopertha dominica (Fabricius, 1792) Pearce: generally common and widely distributed Pearce: 1 locality First subsequent record: 2011 No subsequent records Number of subsequent records: 2 Latest record: 2014 Lyctus linearis (Goeze, 1777) Pearce: 1 locality Dermestes (Dermestinus) undulatus No subsequent records Brahm, 1790 Pearce: 4 localities 46. Family PTINIDAE Latreille, 1802 First subsequent record: 1935 Number of subsequent records: 6 Latest record: 1995 Ptinomorphus imperialis (Linnaeus, 1767) [2012: Hedobia imperialis] Pearce: 2 localities Attagenus pellio (Linnaeus, 1758) First subsequent record: 1934 Pearce: 7 localities Number of subsequent records: many First subsequent record: 1982 Latest record: 2018 Number of subsequent records: 2 Latest record: 1995 Niptus holoeucus (Faldermann, 1835) Pearce: 4 localities No subsequent records

Epauloecus unicolor (Piller & Ochina ptinoides (Marsham, 1802) Mitterpacher, 1783) [2012: Tipnus Pearce: 1 locality unicolor] First subsequent record: 2000 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1937 Latest record: 2020 Number of subsequent records: 1 Latest record: 1937 Xestobium rufovillosum (De Geer, 1774) Pearce: 3 localities Ptinus (Bruchoptinus) palliatus Perris, First subsequent record: 1953 Number of subsequent records: 2 1847 Latest record: 2007 Pearce: 3 localities No subsequent records Ernobius mollis (Linnaeus, 1758) Pearce: 4 localities Ptinus (Gynopterus) sexpunctatus Panzer, First subsequent record: 2001 1789 Number of subsequent records: many Pearce: 2 localities Latest record: 2019 First subsequent record: 1940 Number of subsequent records: many Latest record: 2007 Hemicoelus fulvicornis (Sturm, 1837) [2012: Anobium fulvicorne] Pearce: 2 localities Ptinus (Pseudoptinus) lichenum First subsequent record: 2000 (Marsham, 1802) Number of subsequent records: many Pearce: 1 locality Latest record: 2020 First subsequent record: 1979 Number of subsequent records: 1 Latest record: 1979 Anobium inexspectatum Lohse, 1954 Pearce: no records First record: 1993 Ptinus (Ptinus) fur (Linnaeus, 1758) Number of records: 8 Pearce: 4 localities Latest record: 2019 First subsequent record: 1940 Number of subsequent records: 3 Latest record: 2004 Anobium punctatum (De Geer, 1774) Pearce: generally common and widely distributed First subsequent record: 1979 Ptinus (Ptinus) subpilosus Sturm, 1837 Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 1996 Number of records: 5 Latest record: 2009 Ptilinus pectinicornis (Linnaeus, 1758) Pearce: 2 localities First subsequent record: 1953 Ptinus (Tectoptinus) tectus Boieldieu, Number of subsequent records: many 1856 Latest record: 2020 Pearce: 1 locality No subsequent records Dorcatoma (Dorcatoma) dresdensis Herbst, 1791 Grynobius planus (Fabricius, 1787) Pearce: no records Pearce: 3 localities First record: 2010 First subsequent record: 1954 Number of records: 4 Number of subsequent records: many Latest record: 2019 Latest record: 2020 Dorcatoma (Pilosodorcatoma) Dryophilus pusillus (Gyllenhal, 1808) chrysomelina Sturm, 1837 Pearce: 1 locality (1895) Pearce: no records First subsequent record: 1999 First record: 2019 Number of subsequent records: 4 Number of records: many Latest record: 2011 Latest record: 2020 Dorcatoma (Pilosodorcatoma) substriata 51. Family CLERIDAE Latreille, 1802 Hummel, 1829 Pearce: no records Tillus elongatus (Linnaeus, 1758) First record: 2007 Pearce: no records Number of records: 3 First record: 1995 Latest record: 2019 Number of records: 5 Latest record: 2018 Dorcatoma (Sternitodorcatoma) flavicornis (Fabricius, 1792) Thanasimus formicarius (Linnaeus, 1758) Pearce: no records Pearce: 8 localities First record: 2019 First subsequent record: 1936 Number of records: 7 Number of subsequent records: many Latest record: 2020 Latest record: 2020

47. Family LYMEXYLIDAE Fleming, Korynetes caeruleus (De Geer, 1775) 1821 Pearce: 2 localities First subsequent record: 1953 Number of subsequent records: 1 Lymexylon navale (Linnaeus, 1758) Latest record: 1953 Pearce: no records First record: 2004 Number of records: 2 Necrobia ruficollis (Fabricius, 1775) Latest record: 2019 Pearce: 2 localities No subsequent records 48. Family PHLOIOPHILIDAE Kiesenwetter, 1863 Necrobia rufipes (De Geer, 1775) Pearce: 2 localities No subsequent records Phloiophilus edwardsii Stephens, 1830 Pearce: 1 locality First subsequent record: 2009 Necrobia violacea (Linnaeus, 1758) Number of subsequent records: 5 Pearce: 6 localities Latest record: 2019 First subsequent record: 1936 Number of subsequent records: 3 Latest record: 2014 49. Family TROGOSSITIDAE Latreille, 1802 52. Family MELYRIDAE Leach, 1815

Tenebroides mauritanicus (Linnaeus, Aplocnemus impressus (Marsham, 1802) 1758) Pearce: 1 locality Pearce: 1 locality (1888) First subsequent record: 2010 No subsequent records Number of subsequent records: 1 Latest record: 2010 Nemozoma elongatum (Linnaeus, 1760) Pearce: 1 locality Dasytes (Dasytes) niger (Linnaeus, 1760) No subsequent records Pearce: no records First record: 2017 Thymalus limbatus (Fabricius, 1787) Number of records: 1 Pearce: 1 locality (1898) Latest record: 2017 No subsequent records Dasytes (Mesodasytes) aeratus Stephens, 1830 Pearce: 7 localities First subsequent record: 1939 Number of subsequent records: many Latest record: 2020

Dasytes (Mesodasytes) plumbeus (Müller, Anthocomus rufus (Herbst, 1784) O.F., 1776) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1968 First subsequent record: 2009 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 2009 53. Family BYTURIDAE Gistel, 1848 Psilothrix viridicoerulea (Geoffroy in Fourcroy, 1785) Byturus ochraceus (Scriba, 1790) Pearce: common along coast Pearce: 2 localities First subsequent record: 1936 First subsequent record: 1968 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Axinotarsus marginalis (Laporte, 1840) Byturus tomentosus (De Geer, 1774) Pearce: no records Pearce: 7 localities First record: 2000 First subsequent record: 1939 Number of records: 2 Number of subsequent records: many Latest record: 2006 Latest record: 2019

Cordylepherus viridis (Fabricius, 1787) 54. Family SPHINDIDAE Jacquelin du Pearce: 3 localities Val, 1860 First subsequent record: 1939 Number of subsequent records: many Latest record: 2020 Sphindus dubius (Gyllenhal, 1808) Pearce: no records First record: 2019 Malachius aeneus (Linnaeus, 1758) Number of records: 4 Pearce: 3 localities Latest record: 2019 First subsequent record: 1940 Number of subsequent records: 2 Latest record: 1993 Aspidiphorus orbiculatus (Gyllenhal, 1808) Pearce: 1 locality (1840) Malachius bipustulatus (Linnaeus, 1758) First subsequent record: 1993 Pearce: abundant and widely distributed Number of subsequent records: many First subsequent record: 1939 Latest record: 2019 Number of subsequent records: many Latest record: 2020 55. Family BIPHYLLIDAE LeConte, Clanoptilus marginellus (Olivier, 1790) 1861 Pearce: 1 locality First subsequent record: 1954 Biphyllus lunatus (Fabricius, 1787) Number of subsequent records: 4 Pearce: 2 localities Latest record: 2020 First subsequent record: 1997 Number of subsequent records: many Cerapheles terminatus (Ménétries, 1832) Latest record: 2019 Pearce: 1 locality First subsequent record: 1983 Diplocoelus fagi (Chevrolat in Guérin- Number of subsequent records: many Méneville, 1837) Latest record: 2019 Pearce: no records First record: 2018 Anthocomus fasciatus (Linnaeus, 1758) Number of records: 1 Pearce: 1 locality Latest record: 2018 First subsequent record: 1994 Number of subsequent records: many Latest record: 2016

56. Family EROTYLIDAE Latreille, Rhizophagus (Eurhizophagus) depressus 1802 (Fabricius, 1792) Pearce: 4 localities Cryptophilus integer (Heer, 1841) First subsequent record: 2001 Pearce: no records Number of subsequent records: 6 First record: 2020 Latest record: 2018 Number of records: 3 Latest record: 2020 Rhizophagus (Rhizophagus) bipustulatus (Fabricius, 1792) Dacne bipustulata (Thunberg, 1781) Pearce: 4 localities Pearce: 1 locality (1921) First subsequent record: 1963 First subsequent record: 1940 Number of subsequent records: many Number of subsequent records: 7 Latest record: 2019 Latest record: 2019 Rhizophagus (Rhizophagus) dispar Dacne rufifrons (Fabricius, 1775) (Paykull, 1800) Pearce: 3 localities Pearce: no records First subsequent record: 1988 First record: 1978 Number of subsequent records: 2 Number of records: many Latest record: 2020 Latest record: 2020

Triplax aenea (Schaller, 1783) Rhizophagus (Rhizophagus) ferrugineus Pearce: no records (Paykull, 1800) First record: 1994 Pearce: 2 localities Number of records: 6 First subsequent record: 1988 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Triplax lacordairii Crotch, 1870 Pearce: 1 locality (1926) Rhizophagus (Rhizophagus) nitidulus First subsequent record: 1945 (Fabricius, 1798) Number of subsequent records: 6 Pearce: no records Latest record: 2019 First record: 1997 Number of records: 3 Triplax russica (Linnaeus, 1758) Latest record: 2019 Pearce: 1 locality (1842) First subsequent record: 1997 Rhizophagus (Rhizophagus) Number of subsequent records: many Latest record: 2016 parallelocollis Gyllenhal, 1827 Pearce: 3 localities First subsequent record: 1963 Tritoma bipustulata Fabricius, 1775 Number of subsequent records: 1 Pearce: 2 localities Latest record: 1963 First subsequent record: 2011 Number of subsequent records: 1 Latest record: 2011 Rhizophagus (Rhizophagus) perforatus Erichson, 1845 Pearce: 4 localities 57. Family MONOTOMIDAE Laporte, First subsequent record: 1993 1840 Number of subsequent records: 5 Latest record: 2019 Rhizophagus (Anomophagus) cribratus Gyllenhal, 1827 Monotoma angusticollis (Gyllenhal, 1827) Pearce: no records Pearce: 1 locality First record: 1980 No subsequent records Number of records: many Latest record: 2019

Monotoma bicolor Villa & Villa, 1835 Telmatophilus typhae (Fallén, 1802) Pearce: no records Pearce: 1 locality First record: 2004 First subsequent record: 2001 Number of records: 3 Number of subsequent records: many Latest record: 2020 Latest record: 2019

Monotoma brevicollis Aubé, 1837 Paramecosoma melanocephalum (Herbst, Pearce: 2 localities 1793) First subsequent record: 2006 Pearce: 2 localities Number of subsequent records: 2 First subsequent record: 2017 Latest record: 2020 Number of subsequent records: 1 Latest record: 2017 Monotoma conicicollis Chevrolat in Guérin-Méneville, 1837 Henoticus serratus (Gyllenhal, 1808) Pearce: 1 locality Pearce: no records First subsequent record: 2014 First record: 1993 Number of subsequent records: 9 Number of records: 2 Latest record: 2017 Latest record: 2001

Monotoma longicollis (Gyllenhal, 1827) Cryptophagus acutangulus Gyllenhal, Pearce: 1 locality 1827 First subsequent record: 1970 Pearce: 1 locality Number of subsequent records: 9 First subsequent record: 2003 Latest record: 2011 Number of subsequent records: 5 Latest record: 2012 Monotoma picipes Herbst, 1793 Pearce: 4 localities Cryptophagus cellaris (Scopoli, 1763) First subsequent record: 1978 Pearce: 1 locality (1898) Number of subsequent records: many No subsequent records Latest record: 2010 Cryptophagus dentatus (Herbst, 1793) Monotoma spinicollis Aubé, 1837 Pearce: 3 localities Pearce: 1 locality First subsequent record: 2000 First subsequent record: 2006 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 2006 Cryptophagus denticulatus Heer, 1841 Monotoma testacea Motschulsky, 1845 Pearce: no records Pearce: no records First record: 1994 First record: 2004 Number of records: 5 Number of records: 2 Latest record: 2018 Latest record: 2009 Cryptophagus distinguendus Sturm, 1845 58. Family CRYPTOPHAGIDAE Pearce: no records Kirby, 1826 First record: 1994 Number of records: 9 Telmatophilus caricis (Olivier, 1790) Latest record: 2018 Pearce: 3 localities First subsequent record: 1983 Cryptophagus jakowlewi Reitter, 1888 Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 2003 Number of records: 3 Latest record: 2020 Cryptophagus laticollis Lucas, 1846 Cryptophagus scutellatus Newman, 1834 Pearce: 3 localities Pearce: no records First subsequent record: 2000 First record: 1975 Number of subsequent records: 4 Number of records: 4 Latest record: 2019 Latest record: 2013

Cryptophagus lycoperdi (Scopoli, 1763) Cryptophagus setulosus Sturm, 1845 Pearce: 3 localities Pearce: 3 localities First subsequent record: 1998 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Cryptophagus micaceus Rey, 1889 Micrambe ulicis (Stephens, 1830) Pearce: no records Pearce: abundant on furze-blossoms First record: 2018 First subsequent record: 1975 Number of records: 2 Number of subsequent records: many Latest record: 2020 Latest record: 2020

Cryptophagus populi Paykull, 1800 Micrambe woodroffei Johnson, 2007 Pearce: 1 locality Pearce: 2 localities No subsequent records First subsequent record: 1993 Number of subsequent records: 2 Cryptophagus pubescens Sturm, 1845 Latest record: 2018 Pearce: 1 locality First subsequent record: 2019 Antherophagus pallens (Linnaeus, 1758) Number of subsequent records: 4 Pearce: 4 localities Latest record: 2019 First subsequent record: 1981 Number of subsequent records: many Cryptophagus punctipennis Brisout de Latest record: 2020 Barneville, C., 1863 Pearce: 2 localities Antherophagus silaceus (Herbst, 1792) First subsequent record: 1999 Pearce: 1 locality Number of subsequent records: 7 No subsequent records Latest record: 2020 Antherophagus similis Curtis, 1835 Cryptophagus ruficornis Stephens, 1830 Pearce: 4 localities Pearce: no records First subsequent record: 1971 First record: 1970 Number of subsequent records: 8 Number of records: 3 Latest record: 2009 Latest record: 2019 Caenoscelis ferruginea (Sahlberg, C.R., Cryptophagus saginatus Sturm, 1845 1792) Pearce: 2 localities Pearce: 1 locality (1891) First subsequent record: 2003 First subsequent record: 2006 Number of subsequent records: 2 Number of subsequent records: 1 Latest record: 2003 Latest record: 2006

Cryptophagus scanicus (Linnaeus, 1758) Atomaria (Anchicera) apicalis Erichson, Pearce: 2 localities 1846 First subsequent record: 1971 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Atomaria (Anchicera) atricapilla Atomaria (Anchicera) nitidula (Marsham, Stephens, 1830 1802) Pearce: 6 localities Pearce: no records First subsequent record: 1975 First record: 1999 Number of subsequent records: many Number of records: 6 Latest record: 2020 Latest record: 2019

Atomaria (Anchicera) basalis Erichson, Atomaria (Anchicera) pusilla (Paykull, 1846 1798) Pearce: 1 locality Pearce: 2 localities No subsequent records First subsequent record: 2020 Number of subsequent records: 1 Atomaria (Anchicera) fuscata (Schönherr, Latest record: 2020 1808) Pearce: 4 localities Atomaria (Anchicera) rubella Heer, 1841 First subsequent record: 1979 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1977 Latest record: 2017 Number of subsequent records: 6 Latest record: 2017 Atomaria (Anchicera) fuscipes (Gyllenhal, 1808) Atomaria (Anchicera) rubida Reitter, 1875 Pearce: 1 locality Pearce: no records No subsequent records First record: 2000 Number of records: 2 Latest record: 2000 Atomaria (Anchicera) gutta Newman, 1834 Atomaria (Anchicera) rubricollis Brisout Pearce: 3 localities First subsequent record: 1994 de Barneville, C., 1863 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1998 Number of records: 1 Latest record: 1998 Atomaria (Anchicera) lewisi Reitter, 1877 Pearce: no records First record: 1975 Atomaria (Anchicera) testacea Stephens, Number of records: 9 1830 Latest record: 2020 Pearce: 3 localities First subsequent record: 1975 Number of subsequent records: many Atomaria (Anchicera) mesomela (Herbst, Latest record: 2020 1792) Pearce: 1 locality First subsequent record: 1964 Atomaria (Anchicera) turgida Erichson, Number of subsequent records: 9 1846 Latest record: 2019 Pearce: no records First record: 2020 Number of records: 1 Atomaria (Anchicera) munda Erichson, Latest record: 2020 1846 Pearce: no records First record: 2019 Atomaria (Atomaria) linearis Stephens, Number of records: 2 1830 Latest record: 2019 Pearce: 3 localities First subsequent record: 1953 Number of subsequent records: many Latest record: 2019

Atomaria (Atomaria) nigrirostris Ahasverus advena (Waltl, 1834) Stephens, 1830 Pearce: no records Pearce: no records First record: 1994 First record: 1966 Number of records: 9 Number of records: 2 Latest record: 2007 Latest record: 2016 Oryzaephilus surinamensis (Linnaeus, Atomaria (Atomaria) nigriventris 1758) Stephens, 1830 Pearce: 3 localities Pearce: 1 locality (1892) First subsequent record: 2001 No subsequent records Number of subsequent records: 4 Latest record: 2010 Atomaria (Atomaria) punctithorax Reitter, 1888 Silvanus bidentatus (Fabricius, 1792) Pearce: no records Pearce: no records First record: 1999 First record: 2004 Number of records: 1 Number of records: 1 Latest record: 1999 Latest record: 2004

Atomaria (Atomaria) umbrina (Gyllenhal, Silvanus unidentatus (Olivier, 1790) Pearce: no records 1827) First record: 1992 Pearce: 1 locality Number of records: 7 First subsequent record: 2019 Latest record: 2019 Number of subsequent records: 1 Latest record: 2019 60. Family CUCUJIDAE Latreille, 1802 Ootypus globosus (Waltl, 1838) Pearce: 2 localities Pediacus dermestoides (Fabricius, 1792) First subsequent record: 1996 Pearce: no records Number of subsequent records: 2 First record: 1971 Latest record: 2019 Number of records: many Latest record: 2019 Ephistemus globulus (Paykull, 1798) Pearce: 2 localities 61. Family PHALACRIDAE Leach, First subsequent record: 1975 1815 Number of subsequent records: many Latest record: 2020 Phalacrus caricis Sturm, 1807 Pearce: 3 localities 59. Family SILVANIDAE Kirby, 1837 First subsequent record: 2008 Number of subsequent records: 6 Uleiota planatus (Linnaeus, 1760) Latest record: 2020 Pearce: no records First record: 2019 Phalacrus championi Guillebeau, 1892 Number of records: 1 Pearce: no records Latest record: 2019 First record: 1992 Number of records: 2 Psammoecus bipunctatus (Fabricius, 1792) Latest record: 2019 Pearce: 3 localities First subsequent record: 1968 Phalacrus corruscus (Panzer, 1797) Number of subsequent records: many Pearce: 4 localities Latest record: 2020 First subsequent record: 1935 Number of subsequent records: 6 Nausibius clavicornis (Kugelann, 1794) Latest record: 2019 Pearce: 2 localities No subsequent records Phalacrus fimetarius (Fabricius, 1775) 62. Family LAEMOPHLOEIDAE Pearce: no records Ganglbauer, 1899 First record: 1975 Number of records: 7 Latest record: 2019 Cryptolestes duplicatus (Waltl, 1839) Pearce: 1 locality No subsequent records Olibrus aeneus (Fabricius, 1792) Pearce: 6 localities First subsequent record: 1938 Cryptolestes ferrugineus (Stephens, 1831) Number of subsequent records: many Pearce: 3 localities Latest record: 2020 First subsequent record: 1999 Number of subsequent records: 8 Latest record: 2020 Olibrus affinis (Sturm, 1807) Pearce: 4 localities First subsequent record: 1975 63. Family KATERETIDAE Kirby, Number of subsequent records: many 1837 Latest record: 2020 Kateretes pedicularius (Linnaeus, 1758) Olibrus corticalis (Panzer, 1797) Pearce: 4 localities Pearce: 3 localities First subsequent record: 1969 First subsequent record: 1941 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Kateretes pusillus (Thunberg, 1794) Olibrus liquidus Erichson, 1845 Pearce: 6 localities Pearce: 4 localities First subsequent record: 1987 First subsequent record: 1943 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2019 Latest record: 2019 Kateretes rufilabris (Latreille, 1807) Olibrus millefolii (Paykull, 1800) Pearce: 6 localities Pearce: no records First subsequent record: 1993 First record: 1997 Number of subsequent records: many Number of records: 3 Latest record: 2020 Latest record: 2019 Brachypterus glaber (Newman, 1834) Olibrus pygmaeus (Sturm, 1807) Pearce: generally common and widely distributed Pearce: 1 locality (1909) First subsequent record: 1939 First subsequent record: 2001 Number of subsequent records: many Number of subsequent records: 5 Latest record: 2020 Latest record: 2015 Brachypterus urticae (Fabricius, 1792) Stilbus oblongus (Erichson, 1845) Pearce: 3 localities Pearce: 1 locality (1911) First subsequent record: 1936 First subsequent record: 1964 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2020 Latest record: 2020 Brachypterolus linariae (Stephens, 1830) Stilbus testaceus (Panzer, 1796) Pearce: no records Pearce: generally common and widely distributed First record: 1940 First subsequent record: 1970 Number of records: 1 Number of subsequent records: many Latest record: 1940 Latest record: 2019 Brachypterolus pulicarius (Linnaeus, Epuraea (Epuraea) marseuli Reitter, 1873 1758) Pearce: 2 localities Pearce: 1 locality First subsequent record: 2000 First subsequent record: 2005 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 2005 Epuraea (Epuraea) melina Erichson, 1843 Brachypterolus vestitus (Kiesenwetter, Pearce: 6 localities 1850) First subsequent record: 1987 Pearce: no records Number of subsequent records: 6 First record: 2017 Latest record: 2016 Number of records: 1 Latest record: 2017 Epuraea (Epuraea) pallescens (Stephens, 1835) 64. Family NITIDULIDAE Latreille, Pearce: 1 locality 1802 First subsequent record: 2006 Number of subsequent records: 5 Latest record: 2017 Epuraea (Epuraea) aestiva (Linnaeus, 1758) Epuraea (Epuraea) silacea (Herbst, 1783) Pearce: 8 localities Pearce: no records First subsequent record: 1979 First record: 1996 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1996

Epuraea (Epuraea) biguttata (Thunberg, Epuraea (Epuraeanella) limbata 1784) (Fabricius, 1787) Pearce: 4 localities Pearce: 1 locality First subsequent record: 1979 No subsequent records Number of subsequent records: many Latest record: 2020 Epuraea (Haptoncus) ocularis Fairmaire, Epuraea (Epuraea) distincta (Grimmer, 1849 Pearce: no records 1841) First record: 2020 Pearce: no records Number of records: 1 First record: 2001 Latest record: 2020 Number of records: many Latest record: 2020 Epuraea (Micruria) melanocephala Epuraea (Epuraea) fuscicollis (Stephens, (Marsham, 1802) Pearce: 2 localities 1835) First subsequent record: 1970 Pearce: no records Number of subsequent records: many First record: 2019 Latest record: 2020 Number of records: 1 Latest record: 2019 Carpophilus marginellus Motschulsky, Epuraea (Epuraea) guttata (Olivier, 1811) 1858 Pearce: no records Pearce: no records First record: 2006 First record: 1990 Number of records: 4 Number of records: many Latest record: 2019 Latest record: 2020

Pria dulcamarae (Scopoli, 1763) Pearce: 3 localities First subsequent record: 1953 Number of subsequent records: many Latest record: 2013 Meligethes aeneus (Fabricius, 1775) Meligethes exilis Sturm, 1845 Pearce: generally common and widely distributed Pearce: no records First subsequent record: 1975 First record: 1975 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1975

Meligethes atramentarius Förster, 1849 Meligethes flavimanus Stephens, 1830 Pearce: no records Pearce: 6 localities First record: 1987 First subsequent record: 1965 Number of records: 1 Number of subsequent records: 8 Latest record: 1987 Latest record: 2020

Meligethes atratus (Olivier, 1790) Meligethes fulvipes Brisout de Barneville, Pearce: 2 localities C., 1863 First subsequent record: 1975 Pearce: 1 locality (1904) Number of subsequent records: many First subsequent record: 1978 Latest record: 2020 Number of subsequent records: 1 Latest record: 1978 Meligethes bidens Brisout de Barneville, C., 1863 Meligethes kunzei Erichson, 1845 Pearce: no records Pearce: no records First record: 1998 First record: 2019 Number of records: 1 Number of records: 1 Latest record: 1998 Latest record: 2019

Meligethes bidentatus Brisout de Meligethes lugubris Sturm, 1845 Barneville, C., 1863 Pearce: 1 locality (1892) Pearce: no records First subsequent record: 2002 First record: 1950 Number of subsequent records: 2 Number of records: 1 Latest record: 2015 Latest record: 1950 Meligethes matronalis Audisio & Meligethes brunnicornis Sturm, 1845 Spornraft, 1990 Pearce: no records Pearce: no records First record: 2012 First record: 2014 Number of records: 5 Number of records: 4 Latest record: 2020 Latest record: 2020

Meligethes carinulatus Förster, 1849 Meligethes morosus Erichson, 1845 Pearce: 4 localities Pearce: 1 locality First subsequent record: 1968 First subsequent record: 1993 Number of subsequent records: many Number of subsequent records: 4 Latest record: 2020 Latest record: 2020

Meligethes difficilis (Heer, 1841) Meligethes nigrescens Stephens, 1830 Pearce: 1 locality (1892) Pearce: 5 localities First subsequent record: 2008 First subsequent record: 1955 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2019 Latest record: 2015

Meligethes erichsonii Brisout de Meligethes obscurus Erichson, 1845 Barneville, C., 1863 Pearce: 4 localities Pearce: no records First subsequent record: 1975 First record: 1998 Number of subsequent records: many Number of records: 7 Latest record: 2020 Latest record: 2020 Meligethes ovatus Sturm, 1845 Nitidula bipunctata (Linnaeus, 1758) Pearce: 1 locality Pearce: 7 localities First subsequent record: 2006 First subsequent record: 1993 Number of subsequent records: 3 Number of subsequent records: 3 Latest record: 2020 Latest record: 2014

Meligethes pedicularius (Gyllenhal, 1808) Nitidula carnaria (Schaller, 1783) Pearce: 1 locality Pearce: 1 locality (1896) First subsequent record: 2004 No subsequent records Number of subsequent records: 1 Latest record: 2004 Nitidula rufipes (Linnaeus, 1767) Pearce: 2 localities Meligethes persicus Faldermann, 1835 No subsequent records Pearce: 2 localities First subsequent record: 1998 Omosita colon (Linnaeus, 1758) Number of subsequent records: 1 Pearce: 4 localities Latest record: 1998 First subsequent record: 1939 Number of subsequent records: 4 Meligethes planiusculus (Heer, 1841) Latest record: 2015 Pearce: 5 localities First subsequent record: 1969 Omosita depressa (Linnaeus, 1758) Number of subsequent records: many Pearce: 1 locality Latest record: 2020 No subsequent records

Meligethes rotundicollis Brisout de Omosita discoidea (Fabricius, 1775) Barneville, C., 1863 Pearce: 3 localities Pearce: 1 locality First subsequent record: 1979 No subsequent records Number of subsequent records: 4 Latest record: 2019 Meligethes ruficornis (Marsham, 1802) Pearce: 2 localities Soronia grisea (Linnaeus, 1758) First subsequent record: 1978 Pearce: 1 locality Number of subsequent records: 3 First subsequent record: 1998 Latest record: 2020 Number of subsequent records: 6 Latest record: 2019 Meligethes serripes (Gyllenhal, 1827) Pearce: 1 locality Soronia punctatissima (Illiger, 1794) No subsequent records Pearce: 2 localities First subsequent record: 1993 Meligethes solidus (Kugelann, 1794) Number of subsequent records: 1 Pearce: no records Latest record: 1993 First record: 1993 Number of records: 5 Pocadius ferrugineus (Fabricius, 1775) Latest record: 2018 Pearce: 4 localities First subsequent record: 1963 Meligethes umbrosus Sturm, 1845 Number of subsequent records: many Pearce: 1 locality Latest record: 2009 No subsequent records Thalycra fervida (Olivier, 1790) Meligethes viridescens (Fabricius, 1787) Pearce: no records Pearce: generally common First record: 2001 First subsequent record: 1939 Number of records: 3 Number of subsequent records: many Latest record: 2004 Latest record: 2019 Cychramus luteus (Fabricius, 1787) 66. Family CERYLONIDAE Billberg, Pearce: 4 localities 1820 First subsequent record: 1987 Number of subsequent records: 8 Latest record: 2006 Cerylon fagi Brisout de Barneville, C., 1867 Pearce: 1 locality Cryptarcha strigata (Fabricius, 1787) No subsequent records Pearce: no records First record: 1975 Number of records: 6 Cerylon ferrugineum Stephens, 1830 Latest record: 2019 Pearce: no records First record: 1963 Number of records: many Cryptarcha undata (Olivier, 1790) Latest record: 2020 Pearce: 1 locality First subsequent record: 1975 Number of subsequent records: 4 Cerylon histeroides (Fabricius, 1792) Latest record: 2019 Pearce: 3 localities First subsequent record: 1988 Number of subsequent records: many Pityophagus ferrugineus (Linnaeus, 1760) Latest record: 2019 Pearce: 3 localities First subsequent record: 1964 Number of subsequent records: many 67. Family ALEXIIDAE Imhoff, 1856 Latest record: 2014 Sphaerosoma pilosum (Panzer, 1793) Glischrochilus (Glischrochilus) Pearce: 1 locality quadripunctatus (Linnaeus, 1758) First subsequent record: 1979 Pearce: 1 locality Number of subsequent records: 2 No subsequent records Latest record: 2015

Glischrochilus (Librodor) hortensis 68. Family ENDOMYCHIDAE Leach, (Geoffroy in Fourcroy, 1785) 1815 Pearce: no records First record: 1999 Symbiotes latus Redtenbacher, 1849 Number of records: 8 Pearce: no records Latest record: 2018 First record: 2018 Number of records: 3 Glischrochilus (Librodor) quadriguttatus Latest record: 2019 (Fabricius, 1777) Pearce: no records Mycetaea subterranea (Fabricius, 1801) First record: 1987 Pearce: 3 localities Number of records: 2 First subsequent record: 1975 Latest record: 2004 Number of subsequent records: many Latest record: 2013 65. Family BOTHRIDERIDAE Erichson, 1845 Endomychus coccineus (Linnaeus, 1758) Pearce: 3 localities First subsequent record: 1954 Anommatus duodecimstriatus (Müller, Number of subsequent records: many P.W.J., 1821) Latest record: 2020 Pearce: 1 locality First subsequent record: 2010 Lycoperdina bovistae (Fabricius, 1792) Number of subsequent records: 2 Pearce: 1 locality Latest record: 2011 First subsequent record: 2000 Number of subsequent records: 5 Latest record: 2003

69. Family COCCINELLIDAE Scymnus (Neopullus) haemorrhoidalis Latreille, 1807 Herbst, 1797 Pearce: 1 locality Coccidula rufa (Herbst, 1783) First subsequent record: 1943 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Scymnus (Neopullus) limbatus Stephens, 1832 Coccidula scutellata (Herbst, 1783) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1994 First subsequent record: 2009 Number of subsequent records: 3 Number of subsequent records: 4 Latest record: 2012 Latest record: 2015 Scymnus (Pullus) auritus Thunberg, 1795 Rhyzobius chrysomeloides (Herbst, 1792) Pearce: no records Pearce: no records First record: 1930 First record: 2019 Number of records: many Number of records: 2 Latest record: 2019 Latest record: 2020 Scymnus (Pullus) suturalis Thunberg, Rhyzobius litura (Fabricius, 1787) 1795 Pearce: generally common and widely distributed Pearce: 1 locality First subsequent record: 1939 First subsequent record: 1978 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Rhyzobius lophanthae (Blaisdell, 1892) Scymnus (Scymnus) femoralis (Gyllenhal, Pearce: no records 1827) First record: 2013 Pearce: no records Number of records: 3 First record: 1940 Latest record: 2019 Number of records: 2 Latest record: 2012 Hyperaspis pseudopustulata Mulsant, 1853 Scymnus (Scymnus) frontalis (Fabricius, Pearce: 6 localities 1787) First subsequent record: 1979 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1936 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Nephus quadrimaculatus (Herbst, 1783) Pearce: no records Scymnus (Scymnus) nigrinus Kugelann, First record: 1990 1794 Number of records: 2 Pearce: 1 locality Latest record: 1994 First subsequent record: 1973 Number of subsequent records: 7 Nephus redtenbacheri (Mulsant, 1846) Latest record: 2020 Pearce: 1 locality First subsequent record: 1975 Scymnus (Scymnus) schmidti Fürsch, 1958 Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1975 Number of subsequent records: many Latest record: 2020 Stethorus pusillus (Herbst, 1797) Tytthaspis sedecimpunctata (Linnaeus, Pearce: no records 1760) First record: 2003 Pearce: 8 localities Number of records: many First subsequent record: 1973 Latest record: 2015 Number of subsequent records: many Latest record: 2020 Chilocorus bipustulatus (Linnaeus, 1758) Pearce: 5 localities Hippodamia tredecimpunctata (Linnaeus, First subsequent record: 1932 1758) Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 2008 Number of records: 1 Chilocorus renipustulatus (Scriba, 1791) Latest record: 2008 Pearce: 4 localities First subsequent record: 1934 Hippodamia variegata (Goeze, 1777) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1980 Number of subsequent records: 7 Exochomus quadripustulatus (Linnaeus, Latest record: 2019 1758) Pearce: 8 localities Aphidecta obliterata (Linnaeus, 1758) First subsequent record: 1936 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1934 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Platynaspis luteorubra (Goeze, 1777) Pearce: 3 localities Adalia bipunctata (Linnaeus, 1758) First subsequent record: 1997 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1939 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Halyzia sedecimguttata (Linnaeus, 1758) Pearce: 7 localities Adalia decempunctata (Linnaeus, 1758) First subsequent record: 1934 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1934 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Psyllobora vigintiduopunctata (Linnaeus, 1758) Coccinella hieroglyphica Linnaeus, 1758 Pearce: 7 localities Pearce: 5 localities First subsequent record: 1942 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2013

Vibidia duodecimguttata (Poda von Coccinella magnifica Redtenbacher, 1843 Neuhaus, 1761) Pearce: no records Pearce: 1 locality First record: 1988 No subsequent records Number of records: many Latest record: 2019 Anisosticta novemdecimpunctata (Linnaeus, 1758) Coccinella septempunctata Linnaeus, 1758 Pearce: 5 localities Pearce: very common and widely distributed First subsequent record: 1939 First subsequent record: 1934 Number of subsequent records: many Number of subsequent records: many Latest record: 2015 Latest record: 2020

Coccinella undecimpunctata Linnaeus, Subcoccinella vigintiquattuorpunctata 1758 (Linnaeus, 1758) Pearce: generally common and widely distributed Pearce: 7 localities First subsequent record: 1934 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Harmonia axyridis (Pallas, 1773) 70. Family CORYLOPHIDAE LeConte, Pearce: no records 1852 First record: 2005 Number of records: many Latest record: 2020 Orthoperus aequalis Sharp, 1885 Pearce: no records First record: 2003 Harmonia quadripunctata (Pontoppidan, Number of records: 1 1763) Latest record: 2003 Pearce: no records First record: 1975 Orthoperus atomus (Gyllenhal, 1808) Number of records: many Pearce: no records Latest record: 2019 First record: 2011 Number of records: 2 Propylea quattuordecimpunctata Latest record: 2018 (Linnaeus, 1758) Pearce: generally common and widely distributed Orthoperus nigrescens Stephens, 1829 First subsequent record: 1932 Pearce: no records Number of subsequent records: many First record: 1993 Latest record: 2020 Number of records: many Latest record: 2020 Anatis ocellata (Linnaeus, 1758) Pearce: 8 localities Sericoderus brevicornis Matthews, A., First subsequent record: 1943 1890 Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 2013 Number of records: many Myrrha octodecimguttata (Linnaeus, Latest record: 2019 1758) Pearce: 9 localities Sericoderus lateralis (Gyllenhal, 1827) First subsequent record: 1942 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1990 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Calvia quattuordecimguttata (Linnaeus, 1758) Corylophus cassidoides (Marsham, 1802) Pearce: 9 localities Pearce: 3 localities First subsequent record: 1933 First subsequent record: 1991 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Myzia oblongoguttata (Linnaeus, 1758) Corylophus sublaevipennis Jacquelin du Pearce: 7 localities Val, 1859 First subsequent record: 1945 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1990 Latest record: 2020 Number of subsequent records: many Latest record: 2018 Rypobius praetermissus Bowestead, 1999 Latridius minutus (Linnaeus, 1767) Pearce: no records Pearce: 3 localities (but these could refer to First record: 1931 assimilis or porcatus) Number of records: 4 First subsequent record: 1979 Latest record: 2006 Number of subsequent records: many Latest record: 2019 71. Family LATRIDIIDAE Erichson, 1842 Latridius porcatus Herbst, 1793 Pearce: no records First record: 2000 Stephostethus alternans (Mannerheim, Number of records: many 1844) Latest record: 2019 Pearce: no records First record: 1993 Number of records: 2 Enicmus brevicornis (Mannerheim, 1844) Latest record: 2014 Pearce: no records First record: 2000 Number of records: 7 Stephostethus angusticollis (Gyllenhal, Latest record: 2019 1827) Pearce: 1 locality Enicmus fungicola Thomson, C.G., 1868 No subsequent records Pearce: no records First record: 2017 Stephostethus lardarius (De Geer, 1775) Number of records: 6 Pearce: 2 localities Latest record: 2019 First subsequent record: 1979 Number of subsequent records: many Enicmus histrio Joy & Tomlin, 1910 Latest record: 2019 Pearce: 1 locality First subsequent record: 1990 Cartodere (Aridia) bifasciata (Reitter, Number of subsequent records: 9 1877) Latest record: 2019 Pearce: no records First record: 1975 Enicmus rugosus (Herbst, 1793) Number of records: many Pearce: no records Latest record: 2020 First record: 2018 Number of records: 2 Cartodere (Aridia) nodifer (Westwood, Latest record: 2019 1839) Pearce: 6 localities Enicmus testaceus (Stephens, 1830) First subsequent record: 1975 Pearce: no records Number of subsequent records: many First record: 1993 Latest record: 2020 Number of records: many Latest record: 2020 Cartodere (Cartodere) constricta (Gyllenhal, 1827) Enicmus transversus (Olivier, 1790) Pearce: 1 locality Pearce: 4 localities First subsequent record: 2018 First subsequent record: 1975 Number of subsequent records: 1 Number of subsequent records: many Latest record: 2018 Latest record: 2020

Latridius assimilis Mannerheim, 1844 Dienerella ruficollis (Marsham, 1802) Pearce: no records Pearce: 1 locality First record: 1975 First subsequent record: 1969 Number of records: 1 Number of subsequent records: 9 Latest record: 1975 Latest record: 2010 Corticaria crenulata (Gyllenhal, 1827) Corticarina similata (Gyllenhal, 1827) Pearce: 4 localities Pearce: no records First subsequent record: 1975 First record: 1993 Number of subsequent records: many Number of records: 2 Latest record: 2020 Latest record: 1993

Corticaria elongata (Gyllenhal, 1827) Cortinicara gibbosa (Herbst, 1793) Pearce: 3 localities Pearce: 8 localities First subsequent record: 1975 First subsequent record: 1994 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2007 Latest record: 2020

Corticaria ferruginea Marsham, 1802 Melanophthalma transversalis (Gyllenhal, Pearce: 3 localities 1827) No subsequent records Pearce: 1 locality First subsequent record: 1994 Corticaria fulva (Comolli, 1837) Number of subsequent records: 1 Pearce: 1 locality (1895) Latest record: 1994 First subsequent record: 1975 Number of subsequent records: 2 72. Family MYCETOPHAGIDAE Latest record: 1975 Latreille, 1815

Corticaria impressa (Olivier, 1790) Triphyllus bicolor (Fabricius, 1777) Pearce: 5 localities Pearce: 1 locality First subsequent record: 1975 First subsequent record: 1996 Number of subsequent records: many Number of subsequent records: 5 Latest record: 2020 Latest record: 2004

Corticaria punctulata Marsham, 1802 Litargus connexus (Geoffroy in Fourcroy, Pearce: 1 locality First subsequent record: 1975 1785) Number of subsequent records: 9 Pearce: 3 localities Latest record: 2013 First subsequent record: 1984 Number of subsequent records: many Latest record: 2020 Corticaria umbilicata (Beck, 1817) Pearce: no records First record: 2019 Mycetophagus (Ilendus) multipunctatus Number of records: 1 Fabricius, 1792 Latest record: 2019 Pearce: no records First record: 1997 Number of records: 7 Corticarina curta (Wollaston, 1854) Latest record: 2020 Pearce: 5 localities First subsequent record: 1989 Number of subsequent records: many Mycetophagus (Mycetophagus) Latest record: 2020 quadripustulatus (Linnaeus, 1760) Pearce: 2 localities Corticarina minuta (Fabricius, 1792) First subsequent record: 1975 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1991 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Mycetophagus (Parilendus) quadriguttatus Müller, P.W.J., 1821 Pearce: no records First record: 1990 Number of records: 2 Latest record: 2000 Mycetophagus (Philomyces) populi Cis bidentatus (Olivier, 1790) Fabricius, 1798 Pearce: 2 localities Pearce: no records First subsequent record: 1966 First record: 2004 Number of subsequent records: 7 Number of records: 2 Latest record: 2018 Latest record: 2004 Cis bilamellatus Wood, 1884 Mycetophagus (Ulolendus) atomarius Pearce: no records (Fabricius, 1787) First record: 1957 Pearce: no records Number of records: many First record: 1969 Latest record: 2020 Number of records: 7 Latest record: 2015 Cis boleti (Scopoli, 1763) Pearce: 5 localities Mycetophagus (Ulolendus) piceus First subsequent record: 1975 Number of subsequent records: many (Fabricius, 1777) Latest record: 2020 Pearce: no records First record: 2004 Number of records: 9 Cis castaneus (Herbst, 1793) Latest record: 2019 Pearce: 2 localities First subsequent record: 1975 Number of subsequent records: many Typhaea stercorea (Linnaeus, 1758) Latest record: 2019 Pearce: 5 localities First subsequent record: 1954 Number of subsequent records: many Cis fagi Waltl, 1839 Latest record: 2020 Pearce: no records First record: 2019 Number of records: 1 73. Family CIIDAE Leach in Samouelle, Latest record: 2019 1819 Cis festivus (Panzer, 1793) Octotemnus glabriculus (Gyllenhal, 1827) Pearce: no records Pearce: 5 localities First record: 2004 First subsequent record: 1997 Number of records: 7 Number of subsequent records: many Latest record: 2019 Latest record: 2019 Cis micans (Fabricius, 1792) Sulcacis nitidus (Fabricius, 1792) Pearce: no records Pearce: no records First record: 2008 First record: 2013 Number of records: 2 Number of records: 1 Latest record: 2011 Latest record: 2013 Cis punctulatus Gyllenhal, 1827 Strigocis bicornis (Mellié, 1849) Pearce: no records Pearce: no records First record: 2018 First record: 2004 Number of records: 1 Number of records: 4 Latest record: 2018 Latest record: 2019 Cis pygmaeus (Marsham, 1802) Orthocis alni (Gyllenhal, 1813) Pearce: no records Pearce: no records First record: 1968 First record: 2008 Number of records: 7 Number of records: 1 Latest record: 2019 Latest record: 2008 Cis submicans Abielle de Perrin, 1874 Abdera flexuosa (Paykull, 1799) Pearce: no records Pearce: no records First record: 1992 First record: 1995 Number of records: many Number of records: 3 Latest record: 2019 Latest record: 2019

Cis vestitus Mellié, 1849 Anisoxya fuscula (Illiger, 1798) Pearce: no records Pearce: no records First record: 2000 First record: 2015 Number of records: many Number of records: 1 Latest record: 2020 Latest record: 2015

Cis villosulus (Marsham, 1802) Melandrya caraboides (Linnaeus 1760) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1992 First subsequent record: 2005 Number of subsequent records: 3 Number of subsequent records: 1 Latest record: 2015 Latest record: 2005

Ennearthron cornutum (Gyllenhal, 1827) Orchesia (Clinocara) minor Walker, 1837 Pearce: 2 localities Pearce: 2 localities First subsequent record: 1992 First subsequent record: 1965 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019

74. Family TETRATOMIDAE Billberg, Orchesia (Clinocara) undulata Kraatz, 1820 1853 Pearce: no records Tetratoma desmarestii Latreille, 1807 First record: 1996 Pearce: no records Number of records: many First record: 2004 Latest record: 2019 Number of records: 1 Latest record: 2004 Orchesia (Orchesia) micans (Panzer, 1793) Tetratoma fungorum Fabricius, 1790 Pearce: 1 locality (1891) Pearce: no records First subsequent record: 2019 First record: 2011 Number of subsequent records: 1 Number of records: 1 Latest record: 2019 Latest record: 2011 Conopalpus testaceus (Olivier, 1790) Hallomenus binotatus (Quensel, 1790) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1954 First subsequent record: 2000 Number of subsequent records: 3 Number of subsequent records: 4 Latest record: 2018 Latest record: 2018 76. Family MORDELLIDAE Latreille, 75. Family MELANDRYIDAE Leach, 1802 1815 Mordella leucaspis Küster, 1849 Abdera biflexuosa (Curtis, 1829) Pearce: 2 localities for M. aculeata could be this Pearce: 1 locality species or M. holomelaena First subsequent record: 2000 First record: 1987 Number of subsequent records: 7 Number of records: 1 Latest record: 2019 Latest record: 1987 Tomoxia bucephala Costa, A., 1854 77. Family RIPIPHORIDAE Pearce: 1 locality (1870) Gemminger, 1870 First subsequent record: 1992 Number of subsequent records: 6 Latest record: 2011 Metoecus paradoxus (Linnaeus, 1760) Pearce: 1 locality (1880) First subsequent record: 1997 Variimorda villosa (Schrank, 1781) Number of subsequent records: 2 Pearce: 1 locality Latest record: 2019 First subsequent record: 1974 Number of subsequent records: many Latest record: 2020 78. Family ZOPHERIDAE Solier, 1834

Mordellistena brevicauda (Boheman, Pycnomerus fuliginosus Erichson, 1842 Pearce: no records 1849) First record: 2008 Pearce: 1 locality Number of records: many No subsequent records Latest record: 2020

Mordellistena humeralis (Linnaeus, 1758) Colydium elongatum (Fabricius, 1787) Pearce: 1 locality Pearce: no records First subsequent record: 2007 First record: 1992 Number of subsequent records: 3 Number of records: 3 Latest record: 2020 Latest record: 2020

Mordellistena neuwaldeggiana (Panzer, Orthocerus clavicornis (Linnaeus, 1758) 1796) Pearce: 1 locality Pearce: 1 locality No subsequent records First subsequent record: 1993 Number of subsequent records: many Latest record: 2012 Synchita humeralis (Fabricius, 1792) Pearce: no records First record: 2009 Mordellistena parvula (Gyllenhal, 1827) Number of records: 1 Pearce: 1 locality Latest record: 2009 First subsequent record: 2002 Number of subsequent records: many Latest record: 2020 Cicones undatus Guérin-Méneville, 1829 Pearce: no records First record: 2008 Mordellistena pumila (Gyllenhal, 1810) Number of records: 1 Pearce: 4 localities Latest record: 2008 First subsequent record: 1953 Number of subsequent records: many Latest record: 2019 Bitoma crenata (Fabricius, 1775) Pearce: 2 localities First subsequent record: 1991 Mordellistena variegata (Fabricius, 1798) Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 2008 Number of records: 3 Latest record: 2012 79. Family TENEBRIONIDAE Latreille, 1802 Mordellochroa abdominalis (Fabricius, 1775) Lagria hirta (Linnaeus, 1758) Pearce: 2 localities Pearce: 12 localities First subsequent record: 1993 First subsequent record: 1938 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2020

Alphitobius diaperinus (Panzer, 1796) Tenebrio obscurus Fabricius, 1792 Pearce: no records Pearce: 3 localities First record: 2004 No subsequent records Number of records: 2 Latest record: 2006 Tribolium castaneum (Herbst, 1797) Pearce: 4 localities Blaps mucronata Latreille, 1804 First subsequent record: 2018 Pearce: 6 localities Number of subsequent records: 1 No subsequent records Latest record: 2018

Eledona agricola (Herbst, 1783) Crypticus quisquilius (Linnaeus, 1760) Pearce: no records Pearce: 2 localities First record: 1988 First subsequent record: 1935 Number of records: 8 Number of subsequent records: many Latest record: 2018 Latest record: 2019

Nalassus laevioctostriatus (Goeze, 1777) Alphitophagus bifasciatus (Say, 1824) Pearce: generally common Pearce: no records First subsequent record: 1933 First record: 2011 Number of subsequent records: many Number of records: 1 Latest record: 2019 Latest record: 2011

Helops caeruleus (Linnaeus, 1758) Diaperis boleti (Linnaeus, 1758) Pearce: 1 locality Pearce: no records First subsequent record: 2011 First record: 1964 Number of subsequent records: 4 Number of records: many Latest record: 2016 Latest record: 2019

Melanimon tibialis (Fabricius, 1781) Corticeus fraxini (Kugelann, 1794) Pearce: 2 localities Pearce: no records First subsequent record: 1934 First record: 1998 Number of subsequent records: many Number of records: 3 Latest record: 2019 Latest record: 2016

Opatrum sabulosum (Linnaeus, 1760) Corticeus unicolor Piller & Mitterpacher, Pearce: 3 localities 1783 First subsequent record: 1936 Pearce: no records Number of subsequent records: many First record: 2016 Latest record: 2020 Number of records: 4 Latest record: 2017 Palorus ratzeburgii (Wissmann, 1848) Pearce: 2 localities Myrmechixenus vaporariorum Guérin- No subsequent records Méneville, 1843 Pearce: no records Phylan gibbus (Fabricius, 1775) First record: 2008 Pearce: 4 localities Number of records: 4 First subsequent record: 1934 Latest record: 2011 Number of subsequent records: many Latest record: 2018 Phaleria cadaverina (Fabricius, 1792) Pearce: 2 localities Tenebrio molitor Linnaeus, 1758 First subsequent record: 1953 Pearce: 1 locality Number of subsequent records: 6 First subsequent record: 1999 Latest record: 2007 Number of subsequent records: 3 Latest record: 2019

Scaphidema metallica (Fabricius, 1792) Ischnomera sanguinicollis (Fabricius, Pearce: 2 localities 1787) First subsequent record: 1997 Pearce: no records Number of subsequent records: 6 First record: 2009 Latest record: 2011 Number of records: 1 Latest record: 2009 Prionychus ater (Fabricius, 1775) Pearce: no records Nacerdes melanura (Linnaeus, 1758) First record: 1998 Pearce: 1 locality (1886) Number of records: 3 First subsequent record: 1964 Latest record: 2019 Number of subsequent records: 6 Latest record: 2018 Gonodera luperus (Herbst, 1783) Pearce: 1 locality Oedemera (Oedemera) lurida (Marsham, First subsequent record: 2018 1802) Number of subsequent records: 1 Pearce: 4 localities Latest record: 2018 First subsequent record: 1974 Number of subsequent records: many Isomira murina (Linnaeus, 1758) Latest record: 2020 Pearce: 7 localities First subsequent record: 1954 Oedemera (Oedemera) nobilis (Scopoli, Number of subsequent records: many Latest record: 2020 1763) Pearce: 5 localities First subsequent record: 1935 Pseudocistela ceramboides (Linnaeus, Number of subsequent records: many 1758) Latest record: 2020 Pearce: 2 localities (1823, 1867) No subsequent records Oedemera (Oncomera) femoralis Olivier, 1803 Mycetochara humeralis (Fabricius, 1787) Pearce: 1 locality Pearce: 1 locality First subsequent record: 1974 No subsequent records Number of subsequent records: many Latest record: 2020 Cteniopus sulphureus (Linnaeus, 1758) Pearce: 1 locality 81. Family MELOIDAE Gyllenhal, 1810 First subsequent record: 1969 Number of subsequent records: 5 Latest record: 2015 Lytta vesicatoria (Linnaeus, 1758) Pearce: 9 localities First subsequent record: 2009 Omophlus pubescens (Linnaeus, 1767) Number of subsequent records: 4 Pearce: 2 localities Latest record: 2020 First subsequent record: 1989 Number of subsequent records: many Latest record: 2019 Meloe (Eurymeloe) rugosus Marsham, 1802 Pearce: no records 80. Family OEDEMERIDAE Latreille, First record: 2003 1810 Number of records: many Latest record: 2020 Ischnomera cyanea (Fabricius, 1792) Pearce: 4 localities for caerulea/cyanea Meloe (Meloe) proscarabaeus Linnaeus, First subsequent record: 1994 1758 Number of subsequent records: 5 Pearce: generally common and widely distributed Latest record: 2019 First subsequent record: 1936 Number of subsequent records: many Latest record: 2020 Meloe (Meloe) violaceus Marsham, 1802 Sphaeriestes stockmanni (Biström, 1977) Pearce: 5 localities [2012: Sphaeriestes ater] First subsequent record: 1970 Pearce: 1 locality (1925) Number of subsequent records: many No subsequent records Latest record: 2020 Vincenzellus ruficollis (Panzer, 1794) Meloe (Treiodous) autumnalis Olivier, Pearce: 1 locality (1881) 1792 First subsequent record: 1992 Pearce: 1 locality Number of subsequent records: 6 No subsequent records Latest record: 2019

Sitaris muralis (Forster, 1771) Salpingus planirostris (Fabricius, 1787) Pearce: no records Pearce: 5 localities First record: 2015 First subsequent record: 1963 Number of records: 3 Number of subsequent records: many Latest record: 2019 Latest record: 2020

84. Family PYROCHROIDAE Latreille, Salpingus ruficollis (Linnaeus, 1760) 1806 Pearce: 1 locality First subsequent record: 2004 Number of subsequent records: many Pyrochroa coccinea (Linnaeus, 1760) Latest record: 2019 Pearce: no records First record: 1993 Number of records: 4 86. Family ANTHICIDAE Latreille, Latest record: 2017 1819

Pyrochroa serraticornis (Scopoli, 1763) Anthicus angustatus Curtis, 1838 Pearce: 6 localities Pearce: 2 localities First subsequent record: 1978 First subsequent record: 1998 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 1998

85. Family SALPINGIDAE Leach, 1815 Anthicus antherinus (Linnaeus, 1760) Pearce: 6 localities Aglenus brunneus (Gyllenhal, 1813) First subsequent record: 1962 Pearce: no records Number of subsequent records: many First record: 2008 Latest record: 2020 Number of records: 1 Latest record: 2008 Anthicus tristis Schmidt, W.L.E., 1842 Pearce: 5 localities Lissodema denticollis (Gyllenhal, 1813) First subsequent record: 1978 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1982 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Cordicollis instabilis (Schmidt, W.L.E., 1842) Sphaeriestes castaneus (Panzer, 1796) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1936 No subsequent records Number of subsequent records: many Latest record: 2020 Sphaeriestes reyi (Abeille de Perrin, 1874) Pearce: 1 locality Cyclodinus constrictus (Curtis, 1838) No subsequent records Pearce: 4 localities First subsequent record: 1975 Number of subsequent records: 8 Latest record: 2011 Cyclodinus salinus (Crotch, 1867) Anaspis (Anaspis) fasciata (Forster, 1771) Pearce: 2 localities Pearce: 2 localities First subsequent record: 2004 First subsequent record: 1954 Number of subsequent records: 3 Number of subsequent records: many Latest record: 2007 Latest record: 2020

Omonadus bifasciatus (Rossi, 1792) Anaspis (Anaspis) frontalis (Linnaeus, Pearce: no records 1758) First record: 1995 Pearce: generally common Number of records: 6 First subsequent record: 1954 Latest record: 2006 Number of subsequent records: many Latest record: 2019 Omonadus floralis (Linnaeus, 1758) Pearce: 8 localities Anaspis (Anaspis) garneysi Fowler, 1889 First subsequent record: 1978 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1987 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Omonadus formicarius (Goeze, 1777) Pearce: no records Anaspis (Anaspis) lurida Stephens, 1832 First record: 1995 Pearce: 1 locality Number of records: many First subsequent record: 1986 Latest record: 2016 Number of subsequent records: many Latest record: 2020 Stricticollis tobias (Marseul, 1879) Pearce: no records Anaspis (Anaspis) maculata (Geoffroy in First record: 1996 Number of records: 3 Fourcroy, 1785) Latest record: 2003 Pearce: generally common First subsequent record: 1975 Number of subsequent records: many Notoxus monoceros (Linnaeus, 1760) Latest record: 2020 Pearce: 2 localities First subsequent record: 1998 Number of subsequent records: many Anaspis (Anaspis) pulicaria Costa, A., Latest record: 2019 1854 Pearce: 9 localities First subsequent record: 1969 87. Family ADERIDAE Csiki, 1909 Number of subsequent records: many Latest record: 2020 Aderus populneus (Creutzer in Panzer, 1796) Anaspis (Anaspis) regimbarti Schilsky, Pearce: 1 locality (1916) 1895 First subsequent record: 1997 Pearce: 9 localities Number of subsequent records: 2 First subsequent record: 1969 Latest record: 2002 Number of subsequent records: many Latest record: 2020 Euglenes oculatus (Paykull, 1798) Pearce: no records Anaspis (Anaspis) thoracica (Linnaeus, First record: 1996 Number of records: many 1758) Latest record: 2020 Pearce: no records First record: 1989 Number of records: many 88. Family SCRAPTIIDAE Gistel, 1848 Latest record: 2018

Scraptia dubia (Olivier, 1790) Pearce: 1 locality (1842) No subsequent records Anaspis (Nassipa) costai Emery, 1876 Leptura quadrifasciata Linnaeus, 1758 Pearce: no records Pearce: 3 localities First record: 1996 First subsequent record: 1955 Number of records: 9 Number of subsequent records: many Latest record: 2013 Latest record: 2020

Anaspis (Nassipa) rufilabris (Gyllenhal, Stictoleptura rubra (Linnaeus, 1758) 1827) Pearce: no records Pearce: 2 localities First record: 2020 First subsequent record: 1963 Number of records: 3 Number of subsequent records: many Latest record: 2020 Latest record: 2020 Stictoleptura scutellata (Fabricius, 1781) 89. Family CERAMBYCIDAE Latreille, Pearce: no records 1802 First record: 2016 Number of records: 1 Latest record: 2016 Prionus coriarius (Linnaeus, 1758) Pearce: 2 localities First subsequent record: 1981 Paracorymbia fulva (De Geer, 1775) Number of subsequent records: 4 Pearce: no records Latest record: 2019 First record: 1987 Number of records: 6 Latest record: 2018 Rhagium (Hagrium) bifasciatum Fabricius, 1775 Pachytodes cerambyciformis (Schrank, Pearce: generally common and widely distributed First subsequent record: 1959 1781) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1945 Number of records: many Latest record: 2008 Rhagium (Megarhagium) mordax (De Geer, 1775) Alosterna tabacicolor (De Geer, 1775) Pearce: 4 localities Pearce: 3 localities First subsequent record: 1958 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Rhagium (Rhagium) inquisitor (Linnaeus, Pseudovadonia livida (Fabricius, 1777) 1758) Pearce: 9 localities Pearce: 1 locality First subsequent record: 1951 No subsequent records Number of subsequent records: many Latest record: 2020 Stenocorus meridianus (Linnaeus, 1758) Pearce: 2 localities Rutpela maculata (Poda von Neuhaus, First subsequent record: 1939 1761) Number of subsequent records: many Pearce: generally common Latest record: 2020 First subsequent record: 1935 Number of subsequent records: many Grammoptera ruficornis (Fabricius, 1781) Latest record: 2020 Pearce: generally common and widely distributed First subsequent record: 1934 Stenurella (Nigrostenurella) nigra Number of subsequent records: many Latest record: 2020 (Linnaeus, 1758) Pearce: 2 localities No subsequent records

Stenurella (Stenurella) melanura Aromia moschata (Linnaeus, 1758) (Linnaeus, 1758) Pearce: 3 localities Pearce: 4 localities First subsequent record: 1934 First subsequent record: 1940 Number of subsequent records: 4 Number of subsequent records: many Latest record: 1998 Latest record: 2020 Callidium violaceum (Fabricius, 1775) Asemum striatum (Linnaeus, 1758) Pearce: 1 locality Pearce: 5 localities No subsequent records First subsequent record: 1935 Number of subsequent records: many Phymatodes testaceus (Linnaeus, 1758) Latest record: 2020 Pearce: 1 locality First subsequent record: 1988 Tetropium gabrieli Weise, 1905 Number of subsequent records: 5 Pearce: no records Latest record: 2006 First record: 1958 Number of records: 1 Poecilium alni (Linnaeus, 1767) Latest record: 1958 Pearce: 4 localities No subsequent records Arhopalus ferus (Mulsant, 1839) Pearce: 1 locality Clytus arietis (Linnaeus, 1758) First subsequent record: 1936 Pearce: 6 localities Number of subsequent records: 8 First subsequent record: 1939 Latest record: 1975 Number of subsequent records: many Latest record: 2020 Arhopalus rusticus (Linnaeus, 1758) Pearce: no records Anaglyptus mysticus (Linnaeus, 1758) First record: 1958 Pearce: 1 locality Number of records: many First subsequent record: 1996 Latest record: 2019 Number of subsequent records: 4 Latest record: 2011 Gracilia minuta (Fabricius, 1781) Pearce: 2 localities Agapanthia villosoviridescens (De Geer, No subsequent records 1775) Pearce: no records Obrium brunneum (Fabricius, 1792) First record: 2009 Pearce: no records Number of records: many First record: 1936 Latest record: 2020 Number of records: many Latest record: 2019 Pogonocherus hispidulus (Piller & Mitterpacher, 1783) Nathrius brevipennis (Mulsant, 1839) Pearce: 3 localities Pearce: no records First subsequent record: 1940 First record: 1904 Number of subsequent records: many Number of records: 1 Latest record: 2020 Latest record: 1904 Pogonocherus hispidus (Linnaeus, 1758) Molorchus minor (Linnaeus, 1758) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1964 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2019 Latest record: 2008 Acanthocinus aedilis (Linnaeus, 1758) Glaphyra umbellatarum (Schreber, 1759) Pearce: 1 locality (1912) Pearce: 1 locality No subsequent records No subsequent records Leiopus linnei Wallin, Nylander & 92. Family CHRYSOMELIDAE Kvamme, 2009 Latreille, 1802 Pearce: no records (but see Leiopus nebulosus) First record: 2019 Donacia aquatica (Linnaeus, 1758) Number of records: 1 Pearce: 2 localities Latest record: 2019 No subsequent records

Leiopus nebulosus (Linnaeus, 1758) Donacia bicolora Zschach, 1788 Pearce: 8 localities (but may be this species or Pearce: 7 localities Leiopus linnei) First subsequent record: 1980 First subsequent record: 2010 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2020 Latest record: 2019

Donacia cinerea Herbst, 1784 Saperda populnea (Linnaeus, 1758) Pearce: 1 locality Pearce: 1 locality No subsequent records First subsequent record: 1980 Number of subsequent records: 2 Latest record: 1997 Donacia clavipes Fabricius, 1792 Pearce: 3 localities First subsequent record: 1935 Phytoecia cylindrica (Linnaeus, 1758) Number of subsequent records: 8 Pearce: 1 locality Latest record: 2014 First subsequent record: 1947 Number of subsequent records: many Latest record: 2020 Donacia crassipes Fabricius, 1775 Pearce: 1 locality Tetrops praeustus (Linnaeus, 1758) No subsequent records Pearce: 4 localities First subsequent record: 1942 Donacia dentata Hoppe, 1795 Number of subsequent records: many Pearce: 1 locality Latest record: 2020 No subsequent records

90. Family MEGALOPODIDAE Donacia impressa Paykull, 1799 Latreille, 1802 Pearce: 2 localities First subsequent record: 1996 Number of subsequent records: 6 Zeugophora subspinosa (Fabricius, 1781) Latest record: 2008 Pearce: 1 locality (1866) No subsequent records Donacia marginata Hoppe, 1795 Pearce: 2 localities 91. Family ORSCODACNIDAE First subsequent record: 1992 Thomson, C. G., 1859 Number of subsequent records: many Latest record: 2020 Orsodacne cerasi (Linnaeus, 1758) Pearce: no records Donacia obscura Gyllenhal, 1813 First record: 1987 Pearce: 1 locality Number of records: 4 No subsequent records Latest record: 2018 Donacia semicuprea Panzer, 1796 Orsodacne humeralis Latreille, 1804 Pearce: 2 localities Pearce: no records First subsequent record: 1994 First record: 2019 Number of subsequent records: many Number of records: 1 Latest record: 2013 Latest record: 2019

Donacia simplex Fabricius, 1775 Bruchidius varius (Olivier, 1795) Pearce: 5 localities Pearce: no records First subsequent record: 1983 First record: 2008 Number of subsequent records: many Number of records: 7 Latest record: 2020 Latest record: 2020

Donacia sparganii Ahrens, 1810 Bruchidius villosus (Fabricius, 1792) Pearce: 1 locality Pearce: 1 locality No subsequent records First subsequent record: 1963 Number of subsequent records: many Donacia thalassina Germar, 1811 Latest record: 2017 Pearce: 2 localities (1895) First subsequent record: 1934 Bruchus atomarius (Linnaeus, 1760) Number of subsequent records: 3 Pearce: 3 localities Latest record: 2013 First subsequent record: 1982 Number of subsequent records: many Donacia versicolorea (Brahm, 1790) Latest record: 2019 Pearce: 1 locality First subsequent record: 1997 Bruchus loti Paykull, 1800 Number of subsequent records: many Pearce: 2 localities Latest record: 2019 First subsequent record: 1954 Number of subsequent records: many Donacia vulgaris Zschach, 1788 Latest record: 2020 Pearce: 1 locality First subsequent record: 1962 Bruchus pisorum (Linnaeus, 1758) Number of subsequent records: many Pearce: 2 localities Latest record: 2014 No subsequent records

Plateumaris (Euplateumaris) discolor Bruchus rufimanus Boheman, 1833 (Panzer, 1795) Pearce: 5 localities Pearce: 6 localities First subsequent record: 1954 First subsequent record: 1932 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019 Bruchus rufipes Herbst, 1783 Plateumaris (Euplateumaris) sericea Pearce: no records (Linnaeus, 1758) First record: 1954 Pearce: generally common Number of records: many First subsequent record: 1969 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Crioceris asparagi (Linnaeus, 1758) Pearce: 6 localities Plateumaris (Plateumaris) rustica (Kunze, First subsequent record: 2001 Number of subsequent records: 2 1818) Latest record: 2016 Pearce: 8 localities First subsequent record: 1992 Number of subsequent records: 1 Lema cyanella (Linnaeus, 1758) Latest record: 1992 Pearce: 6 localities First subsequent record: 1953 Number of subsequent records: many Bruchidius cisti (Fabricius, 1775) Latest record: 2017 Pearce: 4 localities First subsequent record: 1963 Number of subsequent records: many Latest record: 2020 Lilioceris lilii (Scopoli, 1763) Cryptocephalus (Burlinius) pusillus Pearce: no records Fabricius, 1777 First record: 1977 Pearce: 5 localities Number of records: many First subsequent record: 1938 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Oulema duftschmidi (Redtenbacher, 1874) [2012: Oulema rufocyanea] Cryptocephalus (Cryptocephalus) Pearce: no records (but see melanopus) aureolus Suffrian, 1847 First record: 1978 Pearce: 3 localities Number of records: many First subsequent record: 1977 Latest record: 2016 Number of subsequent records: many Latest record: 2020 Oulema melanopus (Linnaeus, 1758) Pearce: generally common and widely distributed Cryptocephalus (Cryptocephalus) (but probably includes duftschmidi) biguttatus (Scopoli, 1763) First subsequent record: 1968 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1938 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Oulema obscura (Stephens, 1831) Pearce: 2 localities Cryptocephalus (Cryptocephalus) First subsequent record: 1955 bipunctatus (Linnaeus, 1758) Number of subsequent records: many Pearce: 5 localities Latest record: 2019 First subsequent record: 1935 Number of subsequent records: many Cryptocephalus (Burlinius) bilineatus Latest record: 2020 (Linnaeus, 1767) Pearce: 1 locality Cryptocephalus (Cryptocephalus) First subsequent record: 2005 hypochaeridis (Linnaeus, 1758) Number of subsequent records: 2 Pearce: 1 locality (1924) Latest record: 2020 First subsequent record: 1943 Number of subsequent records: many Cryptocephalus (Burlinius) fulvus (Goeze, Latest record: 2015 1777) Pearce: 7 localities Cryptocephalus (Cryptocephalus) moraei First subsequent record: 1953 (Linnaeus, 1758) Number of subsequent records: many Pearce: 5 localities Latest record: 2020 First subsequent record: 1997 Number of subsequent records: many Cryptocephalus (Burlinius) labiatus Latest record: 2020 (Linnaeus, 1760) Pearce: 3 localities Cryptocephalus (Cryptocephalus) First subsequent record: 1963 parvulus Müller, O.F., 1776 Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 2001 Number of subsequent records: 3 Cryptocephalus (Burlinius) punctiger Latest record: 2011 Paykull, 1799 Pearce: 1 locality Cryptocephalus (Cryptocephalus) No subsequent records primarius Harold, 1872 Pearce: no records First record: 1949 Number of records: many Latest record: 2019 Cryptocephalus (Cryptocephalus) Cassida prasina Illiger, 1798 sexpunctatus (Linnaeus, 1758) Pearce: 1 locality (1841) Pearce: 1 locality First subsequent record: 1998 No subsequent records Number of subsequent records: many Latest record: 2020 Clytra quadripunctata (Linnaeus, 1758) Pearce: 2 localities Cassida rubiginosa Müller, O.F., 1776 First subsequent record: 1980 Pearce: 9 localities Number of subsequent records: 8 First subsequent record: 1963 Latest record: 2018 Number of subsequent records: many Latest record: 2019 Oomorphus concolor (Sturm, 1807) Pearce: 4 localities Cassida sanguinosa Suffrian, 1844 First subsequent record: 1982 Pearce: no records Number of subsequent records: many First record: 2004 Latest record: 2019 Number of records: 6 Latest record: 2014 Cassida denticollis Suffrian, 1844 Pearce: no records Cassida vibex Linnaeus, 1767 First record: 1982 Pearce: 1 locality Number of records: 1 First subsequent record: 1963 Latest record: 1982 Number of subsequent records: many Latest record: 2019 Cassida flaveola Thunberg, 1794 Pearce: 6 localities Cassida viridis Linnaeus, 1758 First subsequent record: 1968 Pearce: 3 localities Number of subsequent records: many First subsequent record: 1981 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Cassida hemisphaerica Herbst, 1799 Pearce: 2 localities Cassida vittata de Villers, 1789 First subsequent record: 2016 Pearce: 4 localities Number of subsequent records: 2 First subsequent record: 1935 Latest record: 2016 Number of subsequent records: many Latest record: 2018 Cassida murraea Linnaeus, 1767 Pearce: 5 localities Pilemostoma fastuosum (Schaller, 1783) First subsequent record: 1981 Pearce: 1 locality (1877) Number of subsequent records: many First subsequent record: 1991 Latest record: 2020 Number of subsequent records: 1 Latest record: 1991 Cassida nebulosa Linnaeus, 1758 Pearce: no records Timarcha goettingensis (Linnaeus, 1758) First record: 1979 Pearce: 4 localities Number of records: many First subsequent record: 1953 Latest record: 2013 Number of subsequent records: many Latest record: 2020 Cassida nobilis Linnaeus, 1758 Pearce: 6 localities Timarcha tenebricosa (Fabricius, 1775) First subsequent record: 1935 Pearce: generally common and widely distributed Number of subsequent records: 5 First subsequent record: 1932 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Gastrophysa polygoni (Linnaeus, 1758) Prasocuris (Prasocuris) phellandrii Pearce: generally common and widely distributed (Linnaeus, 1758) First subsequent record: 1935 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1955 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Gastrophysa viridula De Geer 1775 Pearce: 2 localities Chrysomela populi Linnaeus, 1758 First subsequent record: 1977 Pearce: 7 localities Number of subsequent records: many First subsequent record: 1935 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Phaedon (Paraphaedon) tumidulus (Germar, 1824) Plagiodera versicolora (Laicharting, 1781) Pearce: generally common and widely distributed Pearce: no records First subsequent record: 1966 First record: 1975 Number of subsequent records: many Number of records: 7 Latest record: 2020 Latest record: 2019

Phaedon (Phaedon) armoraciae Plagiosterna aenea (Linnaeus, 1758) (Linnaeus, 1758) [2012: Chrysomela aenea] Pearce: 6 localities Pearce: no records First subsequent record: 1980 First record: 1994 Number of subsequent records: many Number of records: 2 Latest record: 2020 Latest record: 2007

Phaedon (Phaedon) cochleariae Phratora (Phratora) vulgatissima (Fabricius, 1792) (Linnaeus, 1758) Pearce: 4 localities Pearce: 5 localities First subsequent record: 1980 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Prasocuris (Hydrothassa) glabra (Herbst, Phratora (Phyllodecta) laticollis (Suffrian, 1783) 1851) Pearce: 4 localities Pearce: 5 localities First subsequent record: 1955 First subsequent record: 1981 Number of subsequent records: many Number of subsequent records: many Latest record: 2013 Latest record: 2019

Prasocuris (Hydrothassa) marginella Phratora (Phyllodecta) vitellinae (Linnaeus, 1758) (Linnaeus, 1758) Pearce: 8 localities Pearce: generally common First subsequent record: 1967 First subsequent record: 1982 Number of subsequent records: many Number of subsequent records: 7 Latest record: 2016 Latest record: 2018

Prasocuris (Prasocuris) junci (Brahm, Chrysolina (Chrysolina) banksii 1790) (Fabricius, 1775) Pearce: generally common and widely distributed Pearce: 8 localities First subsequent record: 1927 First subsequent record: 1934 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Chrysolina (Chrysolina) staphylaea Chrysolina (Stichoptera) sanguinolenta (Linnaeus, 1758) (Linnaeus, 1758) Pearce: 4 localities Pearce: 5 localities First subsequent record: 1955 No subsequent records Number of subsequent records: many Latest record: 2019 Chrysolina (Sulcicollis) oricalcia (Müller, O.F., 1776) Chrysolina (Chrysolinopsis) americana Pearce: 2 localities (Linnaeus, 1758) First subsequent record: 1981 Pearce: no records Number of subsequent records: many First record: 2007 Latest record: 2020 Number of records: many Latest record: 2019 Chrysolina (Synerga) herbacea (Duftschmid, 1825) Chrysolina (Colaphodes) haemoptera Pearce: 2 localities (Linnaeus, 1758) First subsequent record: 1979 Pearce: 11 localities Number of subsequent records: many First subsequent record: 1934 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Gonioctena (Gonioctena) viminalis (Linnaeus, 1758) Chrysolina (Colaphosoma) sturmi Pearce: 3 localities (Westhoff, 1882) First subsequent record: 1978 Pearce: 8 localities Number of subsequent records: 9 First subsequent record: 2008 Latest record: 1996 Number of subsequent records: 3 Latest record: 2016 Gonioctena (Goniomena) pallida (Linnaeus, 1758) Chrysolina (Erythrochrysa) polita Pearce: 1 locality (1894) (Linnaeus, 1758) First subsequent record: 1978 Pearce: generally common Number of subsequent records: 7 First subsequent record: 1937 Latest record: 2017 Number of subsequent records: many Latest record: 2020 Gonioctena (Spartophila) olivacea (Forster, 1771) Chrysolina (Hypericia) brunsvicensis Pearce: 2 localities (Gravenhorst, 1807) First subsequent record: 1981 Pearce: 3 localities Number of subsequent records: many First subsequent record: 2006 Latest record: 2020 Number of subsequent records: 2 Latest record: 2008 Luperomorpha xanthodera (Fairmaire, 1888) Chrysolina (Hypericia) hyperici (Forster, Pearce: no records 1771) First record: 2015 Pearce: 8 localities Number of records: 1 First subsequent record: 1955 Latest record: 2015 Number of subsequent records: many Latest record: 2019 Agelastica alni (Linnaeus, 1758) Pearce: no records Chrysolina (Sphaeromela) varians First record: 2019 (Schaller, 1783) Number of records: many Pearce: 1 locality (1893) Latest record: 2020 First subsequent record: 1963 Number of subsequent records: 4 Latest record: 2008 Sermylassa halensis (Linnaeus, 1767) Galerucella (Neogalerucella) calmariensis Pearce: 8 localities (Linnaeus, 1767) First subsequent record: 1963 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1983 Latest record: 2020 Number of subsequent records: many Latest record: 2016 Calomicrus circumfusus (Marsham, 1802) Pearce: 4 localities Galerucella (Neogalerucella) lineola First subsequent record: 1939 (Fabricius, 1781) Number of subsequent records: many Pearce: 6 localities Latest record: 2020 First subsequent record: 1962 Number of subsequent records: many Luperus flavipes (Linnaeus, 1767) Latest record: 2019 Pearce: 4 localities First subsequent record: 1980 Galerucella (Neogalerucella) pusilla Number of subsequent records: 6 Latest record: 2012 (Duftschmid, 1825) Pearce: no records First record: 2002 Luperus longicornis (Fabricius, 1781) Number of records: 6 Pearce: generally common Latest record: 2016 First subsequent record: 1933 Number of subsequent records: many Latest record: 2020 Galerucella (Neogalerucella) tenella (Linnaeus, 1760) Pearce: 5 localities Phyllobrotica quadrimaculata (Linnaeus, First subsequent record: 1982 1758) Number of subsequent records: many Pearce: 4 localities Latest record: 2020 First subsequent record: 1998 Number of subsequent records: 8 Latest record: 2020 Lochmaea caprea (Linnaeus, 1758) Pearce: 7 localities First subsequent record: 1963 Galeruca laticollis Sahlberg, C.R., 1838 Number of subsequent records: many Pearce: 2 localities (1893, 1919) Latest record: 2019 First subsequent record: 1965 Number of subsequent records: 1 Latest record: 1965 Lochmaea crataegi (Forster, 1771) Pearce: generally common First subsequent record: 1964 Galeruca tanaceti (Linnaeus, 1758) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1955 Number of subsequent records: many Latest record: 2020 Lochmaea suturalis (Thomson, C.G., 1866) Pearce: generally common Galerucella (Galerucella) nymphaeae First subsequent record: 1933 (Linnaeus, 1758) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1994 Number of subsequent records: 3 Latest record: 2017 Pyrrhalta viburni (Paykull, 1799) Pearce: 3 localities First subsequent record: 1963 Galerucella (Galerucella) sagittariae Number of subsequent records: 8 (Gyllenhal, 1813) Latest record: 2019 Pearce: 5 localities First subsequent record: 1992 Number of subsequent records: many Latest record: 2018 Mniophila muscorum (Koch, J.D.W., Mantura rustica (Linnaeus, 1767) 1803) Pearce: 3 localities Pearce: 1 locality First subsequent record: 1982 First subsequent record: 1995 Number of subsequent records: 2 Number of subsequent records: 1 Latest record: 1983 Latest record: 1995 Crepidodera aurata (Marsham, 1802) Sphaeroderma rubidum (Graëlls, 1858) Pearce: 9 localities Pearce: 2 localities First subsequent record: 1933 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019 Crepidodera aurea (Geoffroy in Fourcroy, Sphaeroderma testaceum (Fabricius, 1775) 1785) Pearce: 7 localities Pearce: 5 localities First subsequent record: 1954 First subsequent record: 1938 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Apteropeda globosa (Illiger, 1794) Crepidodera fulvicornis (Fabricius, 1792) Pearce: no records Pearce: 4 localities First record: 1982 First subsequent record: 1963 Number of records: 7 Number of subsequent records: many Latest record: 2016 Latest record: 2020

Apteropeda orbiculata (Marsham, 1802) Crepidodera nitidula (Linnaeus, 1758) Pearce: 4 localities Pearce: no records First subsequent record: 1963 First record: 1981 Number of subsequent records: many Number of records: 2 Latest record: 2019 Latest record: 1983

Podagrica fuscicornis (Linnaeus, 1767) Crepidodera plutus (Latreille, 1804) Pearce: no records Pearce: 1 locality First record: 2005 First subsequent record: 1994 Number of records: 5 Number of subsequent records: 8 Latest record: 2007 Latest record: 2019

Podagrica fuscipes (Fabricius, 1775) Chaetocnema (Chaetocnema) arida Pearce: 2 localities Foudras, 1860 First subsequent record: 2006 Pearce: no records Number of subsequent records: 7 First record: 1963 Latest record: 2018 Number of records: many Latest record: 2020 Mantura matthewsii (Curtis, 1833) Pearce: 1 locality Chaetocnema (Chaetocnema) confusa First subsequent record: 1955 Number of subsequent records: 9 (Boheman, 1851) Latest record: 2020 Pearce: 1 locality First subsequent record: 1975 Number of subsequent records: many Mantura obtusata (Gyllenhal, 1813) Latest record: 2019 Pearce: 1 locality First subsequent record: 2005 Number of subsequent records: 2 Latest record: 2007 Chaetocnema (Chaetocnema) hortensis Psylliodes cuprea (Koch, J.D.W., 1803) (Geoffroy in Fourcroy, 1785) Pearce: 3 localities Pearce: 9 localities First subsequent record: 1995 First subsequent record: 1963 Number of subsequent records: 4 Number of subsequent records: many Latest record: 2014 Latest record: 2020 Psylliodes dulcamarae (Koch, J.D.W., Chaetocnema (Chaetocnema) sahlbergii 1803) (Gyllenhal, 1827) Pearce: 1 locality Pearce: 1 locality (1887) First subsequent record: 1953 First subsequent record: 2015 Number of subsequent records: 8 Number of subsequent records: 3 Latest record: 2008 Latest record: 2015 Psylliodes laticollis Kutschera, 1860 Chaetocnema (Chaetocnema) subcoerulea Pearce: no records (Kutschera, 1864) First record: 1987 Pearce: 3 localities Number of records: 5 First subsequent record: 1981 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Psylliodes luteola (Müller, O.F., 1776) Pearce: 2 localities Chaetocnema (Tlanoma) concinna First subsequent record: 1993 Number of subsequent records: many (Marsham, 1802) Latest record: 2008 Pearce: generally common and widely distributed (but probably includes picipes) First subsequent record: 1975 Psylliodes marcida (Illiger, 1807) Number of subsequent records: many Pearce: 1 locality Latest record: 2020 No subsequent records

Chaetocnema (Tlanoma) picipes Stephens, Psylliodes napi (Fabricius, 1792) 1831 Pearce: 4 localities Pearce: no records (but see concinna above) First subsequent record: 1978 First record: 2010 Number of subsequent records: many Number of records: many Latest record: 2020 Latest record: 2020 Psylliodes picina (Marsham, 1802) Psylliodes affinis (Paykull, 1799) Pearce: 2 localities Pearce: 3 localities First subsequent record: 1979 First subsequent record: 1975 Number of subsequent records: many Number of subsequent records: many Latest record: 2010 Latest record: 2020 Altica brevicollis Foudras, 1860 Psylliodes chalcomera (Illiger, 1807) Pearce: generally common Pearce: 1 locality (1904) First subsequent record: 1982 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: 4 Latest record: 2020 Latest record: 2018 Altica helianthemi (Allard, 1859) Psylliodes chrysocephala (Linnaeus, 1758) Pearce: 3 localities Pearce: 1 locality (1897) First subsequent record: 1981 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2016 Latest record: 2020 Altica longicollis (Allard, 1860) Aphthona melancholica Weise, 1888 Pearce: 3 localities Pearce: 3 localities First subsequent record: 1999 First subsequent record: 1954 Number of subsequent records: 9 Number of subsequent records: many Latest record: 2019 Latest record: 2020

Altica lythri Aubé, 1843 Aphthona nigriceps (Redtenbacher, W., Pearce: 4 localities 1842) First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: many First subsequent record: 2020 Latest record: 2019 Number of subsequent records: 1 Latest record: 2020 Altica oleracea (Linnaeus, 1758) Pearce: 5 localities Aphthona nonstriata (Goeze, 1777) First subsequent record: 1975 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1948 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Altica palustris Weise, 1888 Pearce: 1 locality Batophila aerata (Marsham, 1802) First subsequent record: 1933 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1963 Latest record: 2016 Number of subsequent records: many Latest record: 2020 Aphthona atratula Allard, 1859 Pearce: 1 locality Batophila rubi (Paykull, 1799) First subsequent record: 1968 Pearce: 3 localities Number of subsequent records: many First subsequent record: 2011 Latest record: 2016 Number of subsequent records: 4 Latest record: 2015 Aphthona atrocaerulea (Stephens, 1831) Pearce: 3 localities Hermaeophaga mercurialis (Fabricius, First subsequent record: 1994 1792) Number of subsequent records: many Pearce: 4 localities Latest record: 2018 First subsequent record: 1954 Number of subsequent records: many Aphthona euphorbiae (Schrank, 1781) Latest record: 2020 Pearce: 3 localities First subsequent record: 1953 Lythraria salicariae (Paykull, 1800) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1981 Number of records: 3 Aphthona herbigrada (Curtis, 1837) Latest record: 2009 Pearce: 2 localities First subsequent record: 1968 Ochrosis ventralis (Illiger, 1807) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 2017 Number of subsequent records: 2 Aphthona lutescens (Gyllenhal, 1813) Latest record: 2020 Pearce: 6 localities First subsequent record: 1963 Derocrepis rufipes (Linnaeus, 1758) Number of subsequent records: many Pearce: 6 localities Latest record: 2019 First subsequent record: 1954 Number of subsequent records: many Latest record: 2020

Neocrepidodera ferruginea (Scopoli, Longitarsus (Longitarsus) ballotae 1763) (Marsham, 1802) Pearce: generally common and widely distributed Pearce: 2 localities First subsequent record: 1968 First subsequent record: 1951 Number of subsequent records: many Number of subsequent records: 4 Latest record: 2020 Latest record: 2020

Neocrepidodera impressa (Fabricius, Longitarsus (Longitarsus) brunneus 1801) (Duftschmid, 1825) Pearce: 1 locality Pearce: 3 localities First subsequent record: 1937 No subsequent records Number of subsequent records: many Latest record: 2018 Longitarsus (Longitarsus) dorsalis (Fabricius, 1781) Neocrepidodera transversa (Marsham, Pearce: 2 localities 1802) First subsequent record: 1982 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1963 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Longitarsus (Longitarsus) exsoletus (Linnaeus, 1758) Hippuriphila modeeri (Linnaeus, 1760) Pearce: 5 localities Pearce: 5 localities First subsequent record: 1966 First subsequent record: 1953 Number of subsequent records: 9 Number of subsequent records: many Latest record: 2020 Latest record: 2020 Longitarsus (Longitarsus) ferrugineus Longitarsus (Longitarsus) aeneicollis (Foudras, 1860) (Faldermann, 1837) Pearce: 1 locality Pearce: 2 localities No subsequent records First subsequent record: 1995 Number of subsequent records: 2 Latest record: 2008 Longitarsus (Longitarsus) flavicornis (Stephens, 1831) Pearce: 1 locality (but see jacobaeae) Longitarsus (Longitarsus) aeruginosus First subsequent record: 1933 (Foudras, 1860) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1925 Number of records: 1 Latest record: 1925 Longitarsus (Longitarsus) gracilis Kutschera, 1864 Pearce: 6 localities Longitarsus (Longitarsus) agilis (Rye, First subsequent record: 1982 1868) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2009 Number of records: 1 Latest record: 2009 Longitarsus (Longitarsus) holsaticus (Linnaeus, 1758) Pearce: 1 locality (1916) Longitarsus (Longitarsus) atricillus First subsequent record: 1987 (Linnaeus, 1760) Number of subsequent records: 1 Pearce: 7 localities Latest record: 1987 First subsequent record: 1975 Number of subsequent records: many Latest record: 2020

Longitarsus (Longitarsus) jacobaeae Longitarsus (Longitarsus) nigerrimus (Waterhouse, G.R., 1858) (Gyllenhal, 1827) Pearce: 6 localities (but probably includes Pearce: 1 locality flavicornis) No subsequent records First subsequent record: 1975 Number of subsequent records: many Longitarsus (Longitarsus) nigrofasciatus Latest record: 2019 (Goeze, 1777) Pearce: no records Longitarsus (Longitarsus) kutscherae First record: 2006 (Rye, 1872) Number of records: 3 Pearce: 1 locality Latest record: 2019 No subsequent records Longitarsus (Longitarsus) obliteratus Longitarsus (Longitarsus) luridus (Rosenhauer, 1847) (Scopoli, 1763) Pearce: no records Pearce: generally common First record: 1983 First subsequent record: 1975 Number of records: many Number of subsequent records: many Latest record: 2009 Latest record: 2020 Longitarsus (Longitarsus) ochroleucus Longitarsus (Longitarsus) lycopi (Foudras, (Marsham, 1802) 1860) Pearce: 4 localities Pearce: 3 localities First record: 1954 First subsequent record: 1995 Number of records: 1 Number of subsequent records: 6 Latest record: 1954 Latest record: 2020 Longitarsus (Longitarsus) parvulus Longitarsus (Longitarsus) melanocephalus (Paykull, 1799) (De Geer, 1775) Pearce: no records Pearce: 8 localities First record: 1993 First subsequent record: 1975 Number of records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019 Longitarsus (Longitarsus) pellucidus Longitarsus (Longitarsus) membranaceus (Foudras, 1860) (Foudras, 1860) Pearce: no records Pearce: 4 localities First record: 1978 First subsequent record: 2001 Number of records: 3 Number of subsequent records: 8 Latest record: 2018 Latest record: 2020 Longitarsus (Longitarsus) pratensis Longitarsus (Longitarsus) minusculus (Panzer, 1794) (Foudras, 1860) Pearce: 4 localities Pearce: no records First subsequent record: 1963 First record: 2002 Number of subsequent records: many Number of records: 6 Latest record: 2020 Latest record: 2020 Longitarsus (Longitarsus) quadriguttatus Longitarsus (Longitarsus) nasturtii (Pontoppidan, 1763) (Fabricius, 1792) Pearce: no records Pearce: 2 localities First record: 1987 First subsequent record: 2019 Number of records: 1 Number of subsequent records: 1 Latest record: 1987 Latest record: 2019

Longitarsus (Longitarsus) rubiginosus Epitrix pubescens (Koch, J.D.W., 1803) (Foudras, 1860) Pearce: no records Pearce: no records First record: 1993 First record: 1975 Number of records: many Number of records: many Latest record: 2020 Latest record: 2019 Phyllotreta astrachanica Lopatin, 1977 Longitarsus (Longitarsus) rutilus (Illiger, [2012: Phyllotreta diademata] 1807) Pearce: no records Pearce: no records First record: 1926 First record: 1983 Number of records: 3 Number of records: 3 Latest record: 2010 Latest record: 2013 Phyllotreta atra (Fabricius, 1775) Longitarsus (Longitarsus) strigicollis Pearce: 6 localities Wollaston, 1864 [2012: Longitarsus First subsequent record: 1975 Number of subsequent records: many fowleri] Latest record: 2019 Pearce: no records First record: 1998 Number of records: many Phyllotreta consobrina (Curtis, 1837) Latest record: 2017 Pearce: 2 localities First subsequent record: 1975 Number of subsequent records: 6 Longitarsus (Longitarsus) succineus Latest record: 2020 (Foudras, 1860) Pearce: 4 localities First subsequent record: 1980 Phyllotreta cruciferae (Goeze, 1777) Number of subsequent records: many Pearce: 7 localities Latest record: 2019 First subsequent record: 1981 Number of subsequent records: 3 Latest record: 2012 Longitarsus (Longitarsus) suturellus (Duftschmid, 1825) Phyllotreta exclamationis (Thunberg, Pearce: 5 localities First subsequent record: 1955 1784) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1954 Number of subsequent records: many Latest record: 2019 Longitarsus (Longitarsus) tabidus (Fabricius, 1775) Phyllotreta flexuosa (Illiger, 1794) Pearce: 3 localities Pearce: no records First subsequent record: 1982 First record: 2008 Number of subsequent records: 9 Number of records: 1 Latest record: 2011 Latest record: 2008

Longitarsus (Testergus) anchusae Phyllotreta nemorum (Linnaeus, 1758) (Paykull, 1799) Pearce: 5 localities Pearce: 4 localities First subsequent record: 1979 First subsequent record: 1996 Number of subsequent records: many Number of subsequent records: 7 Latest record: 2019 Latest record: 2019 Phyllotreta nigripes (Fabricius, 1775) Epitrix atropae Foudras, 1860 Pearce: 5 localities Pearce: no records First subsequent record: 1955 First record: 2006 Number of subsequent records: many Number of records: 2 Latest record: 2019 Latest record: 2009 Phyllotreta nodicornis (Marsham, 1802) Anthribus nebulosus Forster, 1770 Pearce: 4 localities Pearce: “Dorset” (A. Ford) First subsequent record: 1980 First subsequent record: 1986 Number of subsequent records: 7 Number of subsequent records: 5 Latest record: 1988 Latest record: 2010

Phyllotreta ochripes (Curtis, 1837) Platyrhinus resinosus (Scopoli, 1763) Pearce: no records Pearce: no records First record: 1987 First record: 2005 Number of records: 6 Number of records: 4 Latest record: 2020 Latest record: 2016

Phyllotreta punctulata (Marsham, 1802) Platystomos albinus (Linnaeus, 1758) Pearce: 1 locality Pearce: 2 localities First subsequent record: 1969 First subsequent record: 1980 Number of subsequent records: 1 Number of subsequent records: 9 Latest record: 1969 Latest record: 2018

Phyllotreta tetrastigma (Comolli, 1837) Dissoleucas niveirostris (Fabricius, 1798) Pearce: 1 locality Pearce: 1 locality (1893) First subsequent record: 1987 First subsequent record: 1984 Number of subsequent records: many Number of subsequent records: 2 Latest record: 2016 Latest record: 1984

Phyllotreta undulata (Kutschera, 1860) 95. Family RHYNCHITIDAE Gistel, Pearce: 9 localities 1848 First subsequent record: 1981 Number of subsequent records: many Latest record: 2020 Involvulus icosandriae (Scopoli, 1763) [2012: Involvulus caeruleus] Pearce: 1 locality Phyllotreta vittula (Redtenbacher, 1849) First subsequent record: 1983 Pearce: 3 localities Number of subsequent records: 4 First subsequent record: 1979 Latest record: 2001 Number of subsequent records: many Latest record: 2019 Lasiorhynchites cavifrons (Gyllenhal, 93. Family NEMONYCHIDAE Bedel, 1833) Pearce: 3 localities 1882 First subsequent record: 1995 Number of subsequent records: 3 Cimberis attelaboides (Fabricius, 1787) Latest record: 2014 Pearce: 1 locality First subsequent record: 1980 Lasiorhynchites comatus (Gyllenhal, Number of subsequent records: 3 Latest record: 2009 1833) [2012: Lasiorhynchites olivaceus] Pearce: 1 locality First subsequent record: 1994 94. Family ANTHRIBIDAE Billberg, Number of subsequent records: 1 1820 Latest record: 1994

Anthribus fasciatus Forster, 1770 Neocoenorrhinus (Neocoenorhinidius) Pearce: 1 locality interpunctatus (Stephens, 1831) First subsequent record: 1986 Pearce: 1 locality Number of subsequent records: 4 First subsequent record: 1980 Latest record: 1997 Number of subsequent records: 3 Latest record: 1984

Neocoenorrhinus (Neocoenorrhinus) Deporaus betulae (Linnaeus, 1758) germanicus (Herbst, 1797) Pearce: generally common Pearce: 4 localities First subsequent record: 1934 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: many Latest record: 2018 Latest record: 2019 96. Family ATTELABIDAE Billberg, Neocoenorrhinus (Schoenitemnus) minutus 1820 (Herbst, 1797) Pearce: 4 localities Apoderus coryli (Linnaeus, 1758) First subsequent record: 1978 Pearce: generally common Number of subsequent records: many First subsequent record: 1963 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Tatianaerhynchites aequatus (Linnaeus, 1767) Attelabus nitens (Scopoli, 1763) Pearce: 4 localities Pearce: 6 localities First subsequent record: 1939 First subsequent record: 1981 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2019 Latest record: 1981

Temnocerus coeruleus (Fabricius, 1798) 97. Family APIONIDAE Schönherr, Pearce: 1 locality 1823 First subsequent record: 1980 Number of subsequent records: 8 Latest record: 2020 Apion cruentatum Walton, 1844 Pearce: 2 localities First subsequent record: 1982 Temnocerus longiceps (Thomson, C.G., Number of subsequent records: many 1888) Latest record: 2018 Pearce: no records First record: 1982 Apion frumentarium (Linnaeus, 1758) Number of records: 3 Pearce: 6 localities Latest record: 1999 First subsequent record: 1963 Number of subsequent records: many Temnocerus nanus (Paykull, 1792) Latest record: 2020 Pearce: 2 localities First subsequent record: 1980 Apion haematodes Kirby, 1808 Number of subsequent records: many Pearce: 8 localities Latest record: 2019 First subsequent record: 1935 Number of subsequent records: many Byctiscus betulae (Linnaeus, 1758) Latest record: 2020 Pearce: 5 localities First subsequent record: 1983 Apion rubens Walton, 1837 Number of subsequent records: many Pearce: 4 localities Latest record: 1998 First subsequent record: 1975 Number of subsequent records: many Caenorhinus mannerheimii (Hummel, Latest record: 2020 1823) Pearce: 3 localities Apion rubiginosum Walton, 1837 First subsequent record: 1963 Pearce: no records Number of subsequent records: many First record: 1996 Latest record: 2013 Number of records: 5 Latest record: 2015 Perapion affine (Kirby, 1808) Ceratapion (Ceratapion) carduorum Pearce: no records (Kirby, 1808) First record: 1953 Pearce: 8 localities, but many of these could refer Number of records: 4 to gibbirostre Latest record: 2008 First subsequent record: 1975 Number of subsequent records: many Perapion curtirostre (Germar, 1817) Latest record: 2018 Pearce: 7 localities First subsequent record: 1963 Ceratapion (Ceratapion) gibbirostre Number of subsequent records: many (Gyllenhal, 1813) Latest record: 2020 Pearce: no records, but see carduorum First record: 1963 Perapion hydrolapathi (Marsham, 1802) Number of records: many Pearce: 2 localities Latest record: 2019 First subsequent record: 1970 Number of subsequent records: many Diplapion confluens (Kirby, 1808) Latest record: 2020 Pearce: 3 localities First subsequent record: 1953 Perapion marchicum (Herbst, 1797) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1963 Number of subsequent records: many Diplapion stolidum (Germar, 1817) Latest record: 2020 Pearce: no records First record: 1968 Perapion violaceum (Kirby, 1808) Number of records: 7 Pearce: 6 localities Latest record: 1998 First subsequent record: 1975 Number of subsequent records: many Omphalapion hookerorum (Kirby, 1808) Latest record: 2019 Pearce: 3 localities First subsequent record: 1953 Pseudaplemonus limonii (Kirby, 1808) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1989 Number of records: 3 Exapion difficile (Herbst, 1797) Latest record: 2018 Pearce: no records First record: 1981 Aspidapion (Aspidapion) radiolus Number of records: 5 (Marsham, 1802) Latest record: 2019 Pearce: 4 localities First subsequent record: 1966 Exapion fuscirostre (Fabricius, 1775) Number of subsequent records: many Pearce: 1 locality (1895) Latest record: 2020 First subsequent record: 2008 Number of subsequent records: 2 Aspidapion (Koestlinia) aeneum Latest record: 2008 (Fabricius, 1775) Pearce: 3 localities Exapion genistae (Kirby, 1811) First subsequent record: 1978 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1981 Latest record: 2019 Number of subsequent records: 1 Latest record: 1981 Ceratapion (Acanephodus) onopordi (Kirby, 1808) Exapion ulicis (Forster, 1771) Pearce: 4 localities Pearce: generally common and widely distributed First subsequent record: 1963 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Kalcapion semivittatum (Gyllenhal, 1833) Catapion pubescens (Kirby, 1811) Pearce: no records Pearce: 2 localities First record: 1980 First subsequent record: 1955 Number of records: many Number of subsequent records: many Latest record: 2019 Latest record: 2018

Melanapion minimum (Herbst, 1797) Catapion seniculus (Kirby, 1808) Pearce: 2 localities (1896, 1897) Pearce: 4 localities No subsequent records First subsequent record: 1968 Number of subsequent records: many Squamapion atomarium (Kirby, 1808) Latest record: 2020 Pearce: no records First record: 1941 Cyanapion (Bothryorrhynchapion) afer Number of records: many (Gyllenhal, 1833) Latest record: 2015 Pearce: 1 locality First subsequent record: 1954 Squamapion cineraceum (Wencker, 1864) Number of subsequent records: many Pearce: 1 locality (1904) Latest record: 2018 First subsequent record: 2002 Number of subsequent records: 2 Cyanapion (Bothryorrhynchapion) Latest record: 2020 gyllenhalii (Kirby, 1808) Pearce: 2 localities Squamapion vicinum (Kirby, 1808) No subsequent records Pearce: “Dorset” (A. Ford) First subsequent record: 1984 Cyanapion (Cyanapion) spencii (Kirby, Number of subsequent records: 1 Latest record: 1984 1808) Pearce: 1 locality First subsequent record: 1954 Taeniapion urticarium (Herbst, 1784) Number of subsequent records: 9 Pearce: 4 localities Latest record: 2018 First subsequent record: 1975 Number of subsequent records: many Latest record: 2020 Eutrichapion (Cnemapion) vorax (Herbst, 1797) Pearce: 1 locality Malvapion malvae (Fabricius, 1775) First subsequent record: 2010 Pearce: no records Number of subsequent records: 1 First record: 1952 Latest record: 2010 Number of records: many Latest record: 2020 Eutrichapion (Eutrichapion) ervi (Kirby, Pseudapion rufirostre (Fabricius, 1775) 1808) Pearce: 1 locality Pearce: 3 localities First subsequent record: 1976 First subsequent record: 1953 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Catapion curtisii (Stephens, 1831) Eutrichapion (Eutrichapion) viciae Pearce: 1 locality (Paykull, 1800) First subsequent record: 1975 Pearce: 1 locality Number of subsequent records: 3 First subsequent record: 1975 Latest record: 1983 Number of subsequent records: many Latest record: 2020 Eutrichapion (Psilocalymma) punctiger Ischnopterapion (Ischnopterapion) (Paykull, 1792) modestum (Germar, 1817) Pearce: “Dorset” (A. Ford) Pearce: no records First subsequent record: 1954 First record: 1980 Number of subsequent records: many Number of records: many Latest record: 2016 Latest record: 2020

Hemitrichapion reflexum (Gyllenhal, Oxystoma cerdo (Gerstaecker, 1854) 1833) Pearce: no records Pearce: 1 locality (1893) First record: 2004 First subsequent record: 1954 Number of records: 4 Number of subsequent records: 4 Latest record: 2020 Latest record: 1994 Oxystoma craccae (Linnaeus, 1767) Hemitrichapion waltoni (Stephens, 1839) Pearce: 1 locality Pearce: 2 localities First subsequent record: 1981 First subsequent record: 1939 Number of subsequent records: many Number of subsequent records: many Latest record: 2015 Latest record: 2020 Oxystoma pomonae (Fabricius, 1798) Holotrichapion (Apiops) pisi (Fabricius, Pearce: 2 localities 1801) First subsequent record: 1975 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1954 Latest record: 2020 Number of subsequent records: many Latest record: 2018 Oxystoma subulatum (Kirby, 1808) Pearce: 2 localities Holotrichapion (Holotrichapion) ononis First subsequent record: 1954 Number of subsequent records: many (Kirby, 1808) Latest record: 2018 Pearce: 1 locality First subsequent record: 1966 Number of subsequent records: many Pirapion immune (Kirby, 1808) Latest record: 2020 Pearce: 2 localities First subsequent record: 1963 Number of subsequent records: 8 Holotrichapion (Legaricapion) aethiops Latest record: 2019 (Herbst, 1797) Pearce: 3 localities First subsequent record: 1969 Protopirapion atratulum (Germar, 1817) Number of subsequent records: many Pearce: 3 localities Latest record: 2016 First subsequent record: 1927 Number of subsequent records: many Latest record: 2019 Ischnopterapion (Chlorapion) virens (Herbst, 1797) Stenopterapion (Cobosiotherium) Pearce: 5 localities First subsequent record: 1968 scutellare (Kirby, 1811) Number of subsequent records: many Pearce: 3 localities Latest record: 2020 First subsequent record: 1964 Number of subsequent records: many Latest record: 2018 Ischnopterapion (Ischnopterapion) loti (Kirby, 1808) Stenopterapion (Stenopterapion) meliloti Pearce: 4 localities First subsequent record: 1953 (Kirby, 1808) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1966 Number of records: 7 Latest record: 2018 Stenopterapion (Stenopterapion) tenue Protapion laevicolle (Kirby, 1811) (Kirby, 1808) Pearce: 1 locality Pearce: 2 localities First subsequent record: 1999 First subsequent record: 1955 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Protapion nigritarse (Kirby, 1808) Synapion ebeninum (Kirby, 1808) Pearce: 5 localities Pearce: 1 locality First subsequent record: 1981 First subsequent record: 1927 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Protapion ononidis (Gyllenhal, 1827) Betulapion simile (Kirby, 1811) Pearce: no records Pearce: no records First record: 1954 First record: 1985 Number of records: many Number of records: many Latest record: 2020 Latest record: 2020 Protapion trifolii (Linnaeus, 1768) Protapion apricans (Herbst, 1797) Pearce: 6 localities Pearce: generally common First subsequent record: 1939 First subsequent record: 1953 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Protapion varipes (Germar, 1817) Protapion assimile (Kirby, 1808) Pearce: 3 localities Pearce: 1 locality First subsequent record: 2000 First subsequent record: 1968 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2000 Latest record: 2020 98. Family NANOPHYIDAE Gistel, Protapion difforme (Germar, 1818) 1848 Pearce: 4 localities First subsequent record: 1982 Dieckmanniellus gracilis (Redtenbacher, Number of subsequent records: many Latest record: 2019 1849) Pearce: “Dorset” (A. Ford) First subsequent record: 1964 Protapion dissimile (Germar, 1817) Number of subsequent records: 4 Pearce: 1 locality Latest record: 2020 First subsequent record: 2016 Number of subsequent records: 2 Latest record: 2016 Nanophyes marmoratus (Goeze, 1777) Pearce: 5 localities First subsequent record: 1955 Protapion filirostre (Kirby, 1808) Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 1966 Number of records: many Latest record: 2019 99. Family DRYOPHTHORIDAE Schönherr, 1825 Protapion fulvipes (Geoffroy in Fourcroy, 1785) Sitophilus granarius (Linnaeus, 1758) Pearce: 7 localities Pearce: 4 localities First subsequent record: 1964 No subsequent records Number of subsequent records: many Latest record: 2020

Sitophilus oryzae (Linnaeus, 1763) Tanysphyrus lemnae (Paykull, 1792) Pearce: 4 localities Pearce: 1 locality First subsequent record: 1999 First subsequent record: 1954 Number of subsequent records: 1 Number of subsequent records: 6 Latest record: 1999 Latest record: 2019

100. Family ERIRHINIDAE Schönherr, 102. Family CURCULIONIDAE 1825 Latreille, 1802

Grypus equiseti (Fabricius, 1775) Limnobaris dolorosa (Goeze, 1777) Pearce: 5 localities Pearce: no records (but see t-album) First subsequent record: 1980 First record: 1981 Number of subsequent records: many Number of records: 7 Latest record: 2016 Latest record: 2020

Notaris acridulus (Linnaeus, 1758) Limnobaris t-album (Linnaeus, 1758) Pearce: 4 localities Pearce: 5 localities (but may include dolorosa) First subsequent record: 1978 First subsequent record: 1978 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Notaris scirpi (Fabricius, 1792) Aulacobaris lepidii (Germar, 1824) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1938 First subsequent record: 2017 Number of subsequent records: 9 Number of subsequent records: 2 Latest record: 2017 Latest record: 2019

Thryogenes festucae (Herbst, 1795) Aulacobaris picicornis (Marsham, 1802) Pearce: 1 locality (1895) Pearce: 1 locality First subsequent record: 1994 No subsequent records Number of subsequent records: 2 Latest record: 2019 Baris analis (Olivier, 1791) Pearce: no records Thryogenes fiorii Zumpt, 1928 First record: 1989 Pearce: no records Number of records: many First record: 2005 Latest record: 2018 Number of records: 1 Latest record: 2005 Melanobaris laticollis (Marsham, 1802) Pearce: 1 locality (1886) Thryogenes nereis (Paykull, 1800) First subsequent record: 1981 Pearce: 2 localities Number of subsequent records: 2 First subsequent record: 1992 Latest record: 1982 Number of subsequent records: many Latest record: 2020 Acalyptus carpini (Fabricius, 1792) Pearce: 2 localities Thryogenes scirrhosus (Gyllenhal, 1836) First subsequent record: 1979 Pearce: no records Number of subsequent records: many First record: 2006 Latest record: 2020 Number of records: 1 Latest record: 2006 Anoplus plantaris (Naezen, 1794) Pearce: 2 localities (1911, 1919) Tournotaris bimaculata (Fabricius, 1787) First subsequent record: 1974 Pearce: 1 locality (1923) Number of subsequent records: many First subsequent record: 2015 Latest record: 2015 Number of subsequent records: 1 Latest record: 2015 Anoplus roboris Suffrian, 1840 Cionus alauda (Herbst, 1784) Pearce: no records Pearce: 10 localities First record: 1997 First subsequent record: 1975 Number of records: 3 Number of subsequent records: many Latest record: 2001 Latest record: 2020

Anthonomus bituberculatus Thomson, Cionus hortulanus (Geoffroy in Fourcroy, C.G., 1868 1785) Pearce: no records Pearce: 3 localities First record: 1977 First subsequent record: 1975 Number of records: many Number of subsequent records: many Latest record: 2020 Latest record: 2014

Anthonomus brunnipennis Curtis, 1840 Cionus nigritarsis Reitter, 1904 Pearce: 1 locality (1925) Pearce: 1 locality First subsequent record: 1995 No subsequent records Number of subsequent records: 3 Latest record: 2014 Cionus scrophulariae (Linnaeus, 1758) Pearce: common Anthonomus chevrolati Desbrochers des First subsequent record: 1936 Loges, 1868 Number of subsequent records: many Pearce: 1 locality Latest record: 2020 No subsequent records Cionus tuberculosus (Scopoli, 1763) Anthonomus pedicularius (Linnaeus, Pearce: 3 localities 1758) First subsequent record: 1954 Pearce: 4 localities Number of subsequent records: many First subsequent record: 1978 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Cleopus pulchellus (Herbst, 1795) Pearce: 4 localities Anthonomus pomorum (Linnaeus, 1758) First subsequent record: 1975 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1963 Latest record: 2006 Number of subsequent records: 7 Latest record: 2009 Archarius pyrrhoceras (Marsham, 1802) Pearce: 6 localities Anthonomus rubi (Herbst, 1795) First subsequent record: 1963 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Archarius salicivorus (Paykull, 1792) Pearce: 6 localities Anthonomus rufus Gyllenhal, 1835 First subsequent record: 1967 Pearce: no records Number of subsequent records: many First record: 1986 Latest record: 2019 Number of records: 2 Latest record: 1987 Curculio betulae (Stephens, 1831) Pearce: 2 localities (1881, 1894) Anthonomus ulmi (De Geer, 1775) First subsequent record: 1993 Pearce: 3 localities Number of subsequent records: 4 First subsequent record: 1981 Latest record: 2007 Number of subsequent records: 8 Latest record: 2001 Curculio glandium Marsham, 1802 Dorytomus rufatus (Bedel, 1888) Pearce: 5 localities Pearce: 3 localities First subsequent record: 1978 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020

Curculio nucum Linnaeus, 1758 Dorytomus taeniatus (Fabricius, 1781) Pearce: 5 localities Pearce: 4 localities First subsequent record: 1953 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2016 Latest record: 2020

Curculio rubidus (Gyllenhal, 1835) Dorytomus tortrix (Linnaeus, 1760) Pearce: 1 locality Pearce: 2 localities First subsequent record: 2001 First subsequent record: 1968 Number of subsequent records: 4 Number of subsequent records: 8 Latest record: 2010 Latest record: 2015

Curculio venosus (Gravenhorst, 1807) Dorytomus tremulae (Fabricius, 1787) Pearce: 6 localities Pearce: 1 locality First subsequent record: 1968 First subsequent record: 2010 Number of subsequent records: many Number of subsequent records: 1 Latest record: 2020 Latest record: 2010

Curculio villosus Fabricius, 1781 Ellescus bipunctatus (Linnaeus, 1758) Pearce: 5 localities Pearce: 3 localities First subsequent record: 1981 First subsequent record: 1980 Number of subsequent records: 5 Number of subsequent records: many Latest record: 1986 Latest record: 2017

Dorytomus dejeani Faust, 1883 Cleopomiarus graminis (Gyllenhal, 1813) Pearce: 1 locality Pearce: 1 locality First subsequent record: 1980 First subsequent record: 1955 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Dorytomus filirostris (Gyllenhal, 1835) Cleopomiarus plantarum (Germar, 1824) Pearce: no records Pearce: 1 locality First record: 2020 First subsequent record: 1953 Number of records: 1 Number of subsequent records: 1 Latest record: 2020 Latest record: 1953

Dorytomus longimanus (Forster, 1771) Gymnetron beccabungae (Linnaeus, 1760) Pearce: 1 locality Pearce: 1 locality First subsequent record: 1983 First subsequent record: 1989 Number of subsequent records: 7 Number of subsequent records: 2 Latest record: 2020 Latest record: 2013

Dorytomus melanophthalmus (Paykull, Gymnetron melanarium (Germar, 1821) 1792) Pearce: 1 locality (1893) Pearce: 1 locality First subsequent record: 1982 First subsequent record: 1983 Number of subsequent records: 7 Number of subsequent records: 9 Latest record: 2017 Latest record: 2020 Gymnetron veronicae (Germar, 1821) Rhinusa collina (Gyllenhal, 1813) Pearce: 2 localities Pearce: 2 localities First subsequent record: 1963 No subsequent records Number of subsequent records: many Latest record: 2020 Isochnus sequensi (Stierlin, 1894) Pearce: no records Gymnetron villosulum Gyllenhal, 1838 First record: 2016 Pearce: 3 localities Number of records: 4 First subsequent record: 1981 Latest record: 2017 Number of subsequent records: many Latest record: 2006 Orchestes (Alyctus) rusci (Herbst, 1795) Pearce: 2 localities Mecinus circulatus (Marsham, 1802) First subsequent record: 1967 Pearce: 5 localities Number of subsequent records: many First subsequent record: 1963 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Orchestes (Alyctus) testaceus (Müller, O.F., 1776) Mecinus collaris Germar, 1821 Pearce: 1 locality Pearce: no records No subsequent records First record: 2014 Number of records: 4 Latest record: 2015 Orchestes (Orchestes) alni (Linnaeus, 1758) Pearce: 7 localities Mecinus labilis (Herbst, 1795) First subsequent record: 1968 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1963 Latest record: 2017 Number of subsequent records: many Latest record: 2020 Orchestes (Orchestes) fagi (Linnaeus, Mecinus pascuorum (Gyllenhal, 1813) 1758) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1968 First subsequent record: 1978 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2014

Mecinus pyraster (Herbst, 1795) Orchestes (Orchestes) hortorum Pearce: 8 localities (Fabricius, 1792) [2012: Orchestes First subsequent record: 1975 signifer] Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1954 Number of subsequent records: many Miarus campanulae (Linnaeus, 1767) Latest record: 2018 Pearce: no records First record: 1955 Orchestes (Orchestes) jota (Fabricius, Number of records: many 1787) Latest record: 2019 Pearce: 2 localities First subsequent record: 1953 Rhinusa antirrhini (Paykull, 1800) Number of subsequent records: many Pearce: 4 localities Latest record: 2018 First subsequent record: 1954 Number of subsequent records: many Latest record: 2020 Orchestes (Orchestes) pilosus (Fabricius, Smicronyx reichii (Gyllenhal, 1835) 1781) Pearce: 1 locality Pearce: 4 localities First subsequent record: 2000 First subsequent record: 1954 Number of subsequent records: 6 Number of subsequent records: many Latest record: 2018 Latest record: 2020 Pachytychius haematocephalus Orchestes (Orchestes) quercus (Linnaeus, (Gyllenhal, 1835) 1758) Pearce: “Dorset” (A. Ford) Pearce: 3 localities First subsequent record: 1947 First subsequent record: 1954 Number of subsequent records: 2 Number of subsequent records: many Latest record: 2018 Latest record: 2017 Orthochaetes insignis (Aubé, 1863) Pseudorchestes pratensis (Germar, 1821) Pearce: 2 localities Pearce: 1 locality First subsequent record: 1979 No subsequent records Number of subsequent records: many Latest record: 2018 Rhamphus oxyacanthae (Marsham, 1802) Pearce: no records Orthochaetes setiger (Beck, 1817) First record: 1980 Pearce: 5 localities Number of records: many First subsequent record: 1968 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Rhamphus pulicarius (Herbst, 1795) Pearce: 4 localities Sibinia (Dichotychius) sodalis Germar, First subsequent record: 1979 1824 Number of subsequent records: many Pearce: 1 locality Latest record: 2020 No subsequent records

Tachyerges salicis (Linnaeus, 1758) Sibinia (Sibinia) arenariae Stephens, 1831 Pearce: 5 localities Pearce: 3 localities First subsequent record: 1955 First subsequent record: 1964 Number of subsequent records: many Number of subsequent records: many Latest record: 2017 Latest record: 2016

Tachyerges stigma (Germar, 1821) Sibinia (Sibinia) primita (Herbst, 1795) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1965 First subsequent record: 1937 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2016

Smicronyx coecus (Reich, 1797) Sibinia (Sibinia) pyrrhodactyla (Marsham, Pearce: 1 locality 1802) First subsequent record: 2015 Pearce: 2 localities Number of subsequent records: 2 First subsequent record: 1935 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Smicronyx jungermanniae (Reich, 1797) Pearce: 2 localities Tychius breviusculus Desbrochers des First subsequent record: 1980 Loges, 1873 Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 1993 Number of records: many Latest record: 2019

Tychius crassirostris Kirsch, 1871 Tychius tibialis Boheman, 1843 Pearce: no records Pearce: 1 locality First record: 1926 First subsequent record: 1984 Number of records: 9 Number of subsequent records: many Latest record: 2020 Latest record: 2019

Tychius junceus (Reich, 1797) Bagous frit (Herbst, 1795) Pearce: 4 localities Pearce: 1 locality First subsequent record: 1968 No subsequent records Number of subsequent records: many Latest record: 2018 Bagous limosus (Gyllenhal, 1827) Pearce: no records Tychius lineatulus Stephens, 1831 First record: 2005 Pearce: 1 locality (1886) Number of records: 5 First subsequent record: 1980 Latest record: 2015 Number of subsequent records: 8 Latest record: 2016 Bagous lutulosus (Gyllenhal, 1827) Pearce: no records Tychius meliloti Stephens, 1831 First record: 2009 Pearce: 4 localities Number of records: 1 First subsequent record: 1955 Latest record: 2009 Number of subsequent records: many Latest record: 2020 Bagous tempestivus (Herbst, 1795) Pearce: no records Tychius picirostris (Fabricius, 1787) First record: 1955 Pearce: 4 localities Number of records: 9 First subsequent record: 1966 Latest record: 2018 Number of subsequent records: many Latest record: 2020 Hydronomus alismatis (Marsham, 1802) [2012: Bagous alismatis] Tychius pusillus Germar, 1842 Pearce: 2 localities Pearce: 3 localities First subsequent record: 1981 First subsequent record: 1981 Number of subsequent records: 3 Number of subsequent records: many Latest record: 2006 Latest record: 2018 Amalus scortillum (Herbst, 1795) Tychius schneideri (Herbst, 1795) Pearce: 3 localities Pearce: 2 localities First subsequent record: 1968 First subsequent record: 1970 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2006 Amalorrhynchus melanarius (Stephens, Tychius squamulatus Gyllenhal, 1835 1831) Pearce: 4 localities Pearce: “Dorset” (A. Ford) First subsequent record: 1985 First subsequent record: 1954 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Tychius stephensi Schönherr, 1835 Calosirus terminatus (Herbst, 1795) Pearce: 3 localities Pearce: 3 localities First subsequent record: 1953 First subsequent record: 1966 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2020 Latest record: 2020 Ceutorhynchus alliariae (Brisout de Ceutorhynchus obstrictus (Marsham, Barneville, C., 1860) 1802) Pearce: no records Pearce: 6 localities First record: 1983 First subsequent record: 1978 Number of records: many Number of subsequent records: many Latest record: 2018 Latest record: 2020

Ceutorhynchus assimilis (Paykull, 1792) Ceutorhynchus pallidactylus (Marsham, Pearce: 6 localities 1802) First subsequent record: 1978 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1963 Latest record: 2016 Number of subsequent records: many Latest record: 2020 Ceutorhynchus atomus Boheman, 1845 Pearce: no records Ceutorhynchus pyrrhorhynchus (Marsham, First record: 2001 1802) Number of records: 2 Pearce: 3 localities Latest record: 2012 First subsequent record: 1954 Number of subsequent records: many Ceutorhynchus chalybaeus Germar, 1824 Latest record: 2020 Pearce: 2 localities First subsequent record: 1981 Ceutorhynchus sulcicollis (Paykull, 1800) Number of subsequent records: many Pearce: 1 locality Latest record: 2020 No subsequent records

Ceutorhynchus cochleariae (Gyllenhal, Ceutorhynchus turbatus Schultze, 1903 1813) Pearce: no records Pearce: 2 localities First record: 1981 First subsequent record: 1971 Number of records: 5 Number of subsequent records: many Latest record: 2013 Latest record: 2017 Ceutorhynchus typhae (Herbst, 1795) Ceutorhynchus constrictus (Marsham, Pearce: 2 localities 1802) First subsequent record: 1963 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1983 Latest record: 2020 Number of subsequent records: 5 Latest record: 2011 Coeliodes rana (Fabricius, 1787) Pearce: 5 localities Ceutorhynchus contractus (Marsham, First subsequent record: 1954 1802) Number of subsequent records: many Pearce: 9 localities Latest record: 2019 First subsequent record: 1980 Number of subsequent records: many Coeliodes ruber (Marsham, 1802) Latest record: 2019 Pearce: 5 localities First subsequent record: 1955 Ceutorhynchus erysimi (Fabricius, 1787) Number of subsequent records: many Pearce: 4 localities Latest record: 2019 First subsequent record: 1979 Number of subsequent records: many Coeliodes transversealbofasciatus (Goeze, Latest record: 2020 1777) Pearce: 2 localities First subsequent record: 1978 Number of subsequent records: many Latest record: 2016

Coeliodinus rubicundus (Herbst, 1795) Hadroplontus trimaculatus (Fabricius, Pearce: 3 localities 1775) First subsequent record: 1965 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1980 Latest record: 2019 Number of subsequent records: many Latest record: 2015 Datonychus melanostictus (Marsham, 1802) Micrelus ericae (Gyllenhal, 1813) Pearce: 2 localities Pearce: 4 localities First subsequent record: 1980 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Datonychus urticae (Boheman, 1845) Microplontus campestris (Gyllenhal, Pearce: 1 locality 1837) First subsequent record: 1983 Pearce: 5 localities Number of subsequent records: 1 First subsequent record: 1980 Latest record: 1983 Number of subsequent records: 7 Latest record: 2020 Drupenatus nasturtii (Germar, 1824) Pearce: 1 locality Microplontus melanostigma (Marsham, First subsequent record: 1953 1802) [2012: Microplontus rugulosus] Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1963 Number of subsequent records: many Glocianus distinctus (Brisout de Latest record: 2019 Barneville, C., 1870) Pearce: 1 locality Microplontus triangulum (Boheman, First subsequent record: 1982 1845) Number of subsequent records: many Pearce: no records Latest record: 2018 First record: 1983 Number of records: 1 Glocianus moelleri (Thomson, C.G., 1868) Latest record: 1983 Pearce: no records First record: 2014 Mogulones asperifoliarum (Gyllenhal, Number of records: 1 Latest record: 2014 1813) Pearce: 4 localities First subsequent record: 1978 Glocianus punctiger (Sahlberg, C.R., Number of subsequent records: many 1835) Latest record: 2015 Pearce: no records First record: 1981 Mogulones geographicus (Goeze, 1777) Number of records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1980 Number of subsequent records: 6 Hadroplontus litura (Fabricius, 1775) Latest record: 2011 Pearce: 2 localities First subsequent record: 1954 Nedyus quadrimaculatus (Linnaeus, 1758) Number of subsequent records: many Pearce: generally common Latest record: 2018 First subsequent record: 1966 Number of subsequent records: many Latest record: 2020 Parethelcus pollinarius (Forster, 1771) Trichosirocalus rufulus (Dufour, 1851) Pearce: 9 localities Pearce: 1 locality First subsequent record: 1955 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: 8 Latest record: 2020 Latest record: 2013

Poophagus sisymbrii (Fabricius, 1777) Trichosirocalus troglodytes (Fabricius, Pearce: 2 localities 1787) First subsequent record: 1978 Pearce: 8 localities Number of subsequent records: many First subsequent record: 1963 Latest record: 2011 Number of subsequent records: many Latest record: 2020 Sirocalodes depressicollis (Gyllenhal, 1813) Zacladus exiguus (Olivier, 1807) Pearce: 1 locality (1919) Pearce: 1 locality First subsequent record: 1981 First subsequent record: 1985 Number of subsequent records: many Number of subsequent records: 7 Latest record: 2017 Latest record: 2019

Sirocalodes mixtus (Mulsant & Rey, 1859) Zacladus geranii (Paykull, 1800) Pearce: no records Pearce: 1 locality First record: 1983 First subsequent record: 1963 Number of records: 2 Number of subsequent records: 2 Latest record: 2017 Latest record: 1981

Sirocalodes quercicola (Paykull, 1792) Mononychus punctumalbum (Herbst, Pearce: no records 1784) First record: 1985 Pearce: 2 localities Number of records: 3 First subsequent record: 1975 Latest record: 1985 Number of subsequent records: many Latest record: 2020 Stenocarus ruficornis (Stephens, 1831) Pearce: 2 localities Eubrychius velutus (Beck, 1817) First subsequent record: 1982 Pearce: 1 locality Number of subsequent records: 6 No subsequent records Latest record: 2019 Neophytobius muricatus (Brisout de Thamiocolus viduatus (Gyllenhal, 1813) Barneville, C., 1867) Pearce: 1 locality Pearce: no records No subsequent records First record: 1983 Number of records: many Trichosirocalus dawsoni (Brisout de Latest record: 2020 Barneville, C., 1869) Pearce: 2 localities Neophytobius quadrinodosus (Gyllenhal, First subsequent record: 1963 1813) Number of subsequent records: many Pearce: 1 locality Latest record: 2020 First subsequent record: 1982 Number of subsequent records: many Trichosirocalus horridus (Panzer, 1801) Latest record: 2020 Pearce: 3 localities First subsequent record: 1978 Pelenomus canaliculatus (Fåhraeus, 1843) Number of subsequent records: 3 Pearce: 1 locality (1904) Latest record: 2005 First subsequent record: 2016 Number of subsequent records: 2 Latest record: 2020

Pelenomus commari (Panzer, 1795) [2012: Rhinoncus pericarpius (Linnaeus, 1758) Pelenomus comari] Pearce: 9 localities Pearce: no records First subsequent record: 1963 First record: 2010 Number of subsequent records: many Number of records: 2 Latest record: 2020 Latest record: 2011 Rhinoncus perpendicularis (Reich, 1797) Pelenomus olssoni (Israelson, 1972) Pearce: 3 localities Pearce: no records First subsequent record: 1955 First record: 2009 Number of subsequent records: many Number of records: 5 Latest record: 2020 Latest record: 2020 Tapinotus sellatus (Fabricius, 1794) Pelenomus quadricorniger (Colonnelli, Pearce: no records 1986) First record: 1981 Pearce: no records Number of records: 1 First record: 2016 Latest record: 1981 Number of records: 2 Latest record: 2019 Cossonus parallelepipedus (Herbst, 1795) Pearce: 1 locality Pelenomus quadrituberculatus (Fabricius, No subsequent records 1787) Pearce: 2 localities Rhopalomesites tardii (Curtis, 1825) First subsequent record: 1978 Pearce: no records Number of subsequent records: many First record: 1994 Latest record: 2015 Number of records: 1 Latest record: 1994 Pelenomus waltoni (Boheman, 1843) Pearce: no records Pselactus spadix (Herbst, 1795) First record: 1987 Pearce: no records Number of records: many First record: 1964 Latest record: 2020 Number of records: 3 Latest record: 2005 Phytobius leucogaster (Marsham, 1802) Pearce: 1 locality Pseudophloeophagus truncorum No subsequent records (Stephens, 1831) [2012: Pseudophloeophagus aeneopiceus] Rhinoncus bruchoides (Herbst, 1784) Pearce: no records Pearce: 1 locality First record: 1982 First subsequent record: 1981 Number of records: 8 Number of subsequent records: many Latest record: 2018 Latest record: 2010 Euophryum confine (Broun, 1881) Rhinoncus castor (Fabricius, 1792) Pearce: no records Pearce: 6 localities First record: 1980 First subsequent record: 1935 Number of records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020 Pentarthrum huttoni Wollaston, 1854 Rhinoncus inconspectus (Herbst, 1795) Pearce: 1 locality Pearce: 1 locality First subsequent record: 2000 First subsequent record: 1954 Number of subsequent records: 4 Number of subsequent records: 9 Latest record: 2008 Latest record: 2020 Cryptorhynchus lapathi (Linnaeus, 1758) Strophosoma (Strophosoma) capitatum Pearce: 3 localities (De Geer, 1775) First subsequent record: 1934 Pearce: generally common Number of subsequent records: 7 First subsequent record: 1954 Latest record: 2016 Number of subsequent records: 4 Latest record: 2014 Acalles misellus Boheman, 1844 Pearce: 2 localities Strophosoma (Strophosoma) fulvicorne First subsequent record: 1980 (Walton, 1846) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1937 Number of subsequent records: many Acalles ptinoides (Marsham, 1802) Latest record: 2016 Pearce: 1 locality (1901) First subsequent record: 1979 Strophosoma (Strophosoma) Number of subsequent records: many Latest record: 2020 melanogrammum (Forster, 1771) Pearce: generally common First subsequent record: 1935 Kyklioacalles roboris (Curtis, 1835) Number of subsequent records: many Pearce: no records Latest record: 2019 First record: 1984 Number of records: many Latest record: 2018 Attactagenus plumbeus (Marsham, 1802) Pearce: 8 localities First subsequent record: 1981 Gronops lunatus (Fabricius, 1775) Number of subsequent records: many Pearce: 3 localities Latest record: 2019 First subsequent record: 1952 Number of subsequent records: many Latest record: 2017 Philopedon plagiatum (Schaller, 1783) Pearce: 7 localities First subsequent record: 1932 Graptus triguttatus (Fabricius, 1775) Number of subsequent records: many Pearce: 1 locality Latest record: 2018 First subsequent record: 1967 Number of subsequent records: many Latest record: 2020 Barynotus moerens (Fabricius, 1792) Pearce: no records First record: 1983 Strophosoma (Neliocarus) faber (Herbst, Number of records: 4 1784) Latest record: 1987 Pearce: 2 localities First subsequent record: 1982 Barynotus obscurus (Fabricius, 1775) Number of subsequent records: 8 Pearce: generally common and widely distributed Latest record: 2020 First subsequent record: 1970 Number of subsequent records: many Strophosoma (Neliocarus) nebulosum Latest record: 2019 (Stephens, 1831) Pearce: 3 localities Otiorhynchus (Aranihus) ligneus (Olivier, First subsequent record: 1927 1807) Number of subsequent records: many Pearce: 4 localities Latest record: 2020 First subsequent record: 1979 Number of subsequent records: many Strophosoma (Neliocarus) sus (Stephens, Latest record: 2019 1831) Pearce: very common on heaths Otiorhynchus (Choilisanus) raucus First subsequent record: 1937 (Fabricius, 1777) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 No subsequent records Otiorhynchus (Clypeotiorhynchus) Otiorhynchus (Zadrehus) atroapterus (De crataegi Germar, 1824 Geer, 1775) Pearce: no records Pearce: 5 localities First record: 2014 First subsequent record: 1934 Number of records: 1 Number of subsequent records: many Latest record: 2014 Latest record: 2015

Otiorhynchus (Dorymerus) sulcatus Otiorhynchus (Zustalestus) rugosostriatus (Fabricius, 1775) (Goeze, 1777) Pearce: generally common Pearce: 8 localities First subsequent record: 1934 First subsequent record: 1979 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Otiorhynchus (Metopiorrhynchus) Phyllobius (Dieletus) argentatus singularis (Linnaeus, 1767) (Linnaeus, 1758) Pearce: generally common and widely distributed Pearce: generally common First subsequent record: 1934 First subsequent record: 1934 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2016

Otiorhynchus (Otiorhynchus) tenebricosus Phyllobius (Metaphyllobius) glaucus (Herbst, 1784) [2012: Otiorhynchus (Scopoli, 1763) clavipes] Pearce: 5 localities Pearce: generally common and widely distributed First subsequent record: 1968 First subsequent record: 1953 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2020 Phyllobius (Metaphyllobius) pomaceus Otiorhynchus (Pendragon) desertus Gyllenhal, 1834 Rosenhauer, 1847 Pearce: 7 localities Pearce: no records First subsequent record: 1954 First record: 1983 Number of subsequent records: many Number of records: 3 Latest record: 2020 Latest record: 2004 Phyllobius (Nemoicus) oblongus Otiorhynchus (Pendragon) ovatus (Linnaeus, 1758) (Linnaeus, 1758) Pearce: 6 localities Pearce: 2 localities First subsequent record: 1980 First subsequent record: 1980 Number of subsequent records: many Number of subsequent records: many Latest record: 2012 Latest record: 2020 Phyllobius (Parnemoicus) roboretanus Otiorhynchus (Provadilus) rugifrons Gredler, 1882 (Gyllenhal, 1813) Pearce: generally common Pearce: 6 localities First subsequent record: 1962 First subsequent record: 1983 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2017 Phyllobius (Parnemoicus) viridicollis (Fabricius, 1792) Pearce: 2 localities First subsequent record: 1964 Number of subsequent records: many Latest record: 2012 Phyllobius (Phyllobius) pyri (Linnaeus, Polydrusus (Eurodrusus) confluens 1758) Stephens, 1831 Pearce: 3 localities Pearce: 4 localities First subsequent record: 1934 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Phyllobius (Pterygorrhynchus) Polydrusus (Eurodrusus) pulchellus maculicornis Germar, 1824 Stephens, 1831 Pearce: 4 localities Pearce: 1 locality First subsequent record: 1980 First subsequent record: 1995 Number of subsequent records: many Number of subsequent records: 6 Latest record: 2019 Latest record: 2018

Phyllobius (Subphyllobius) virideaeris Polydrusus (Eustolus) flavipes (De Geer, (Laicharting, 1781) 1775) Pearce: 4 localities Pearce: 1 locality First subsequent record: 1962 First subsequent record: 1981 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2018

Liophloeus tessulatus (Müller, O.F., 1776) Polydrusus (Eustolus) pterygomalis Pearce: 5 localities Boheman, 1840 First subsequent record: 1962 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1963 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Pachyrhinus lethierryi (Desbrochers des Loges, 1875) Polydrusus (Metallites) marginatus Pearce: no records Stephens, 1831 First record: 2011 Pearce: 1 locality Number of records: 1 First subsequent record: 2016 Latest record: 2011 Number of subsequent records: 1 Latest record: 2016 Polydrusus (Eudipnus) formosus (Mayer, 1779) Polydrusus (Polydrusus) tereticollis (De Pearce: 1 locality Geer, 1775) First subsequent record: 2001 Pearce: generally common Number of subsequent records: many First subsequent record: 1978 Latest record: 2020 Number of subsequent records: many Latest record: 2016 Polydrusus (Eudipnus) mollis (Strøm, 1768) Barypeithes sulcifrons (Boheman, 1842) Pearce: 2 localities Pearce: 2 localities First subsequent record: 2014 First subsequent record: 1994 Number of subsequent records: 1 Number of subsequent records: 2 Latest record: 2014 Latest record: 2008

Polydrusus (Eurodrusus) cervinus Brachysomus echinatus (Bonsdorff, 1785) (Linnaeus, 1758) Pearce: 1 locality Pearce: 5 localities First subsequent record: 1982 First subsequent record: 1935 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2020

Brachysomus hirtus (Boheman, 1845) Sitona gemellatus Gyllenhal, 1834 Pearce: 2 localities Pearce: no records No subsequent records First record: 1954 Number of records: many Exomias araneiformis (Schrank, 1781) Latest record: 2020 [2012: Barypeithes araneiformis] Pearce: generally common Sitona hispidulus (Fabricius, 1777) First subsequent record: 1978 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1936 Latest record: 2018 Number of subsequent records: many Latest record: 2020 Exomias pellucidus (Boheman, 1834) [2012: Barypeithes pellucidus] Sitona humeralis Stephens, 1831 Pearce: 1 locality Pearce: 5 localities First subsequent record: 1979 First subsequent record: 1968 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Sciaphilus asperatus (Bonsdorff, 1785) Sitona lateralis Gyllenhal, 1834 [2012: Pearce: 3 localities Sitona ononidis] First subsequent record: 1963 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1982 Latest record: 2020 Number of subsequent records: 8 Latest record: 2020 Andrion regensteinense (Herbst, 1797) Pearce: 4 localities Sitona lineatus (Linnaeus, 1758) First subsequent record: 1927 Pearce: generally common and widely distributed Number of subsequent records: many First subsequent record: 1955 Latest record: 2020 Number of subsequent records: many Latest record: 2020 Charagmus griseus (Fabricius, 1775) Pearce: 1 locality Sitona lineellus (Bonsdorff, 1785) First subsequent record: 1979 Pearce: no records Number of subsequent records: 1 First record: 1982 Latest record: 1979 Number of records: many Latest record: 2020 Coelositona cambricus (Stephens, 1831) Pearce: 6 localities Sitona macularius (Marsham, 1802) First subsequent record: 1981 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1994 Latest record: 2019 Number of subsequent records: 4 Latest record: 2011 Sitona ambiguus Gyllenhal, 1834 Pearce: 2 localities Sitona obsoletus (Gmelin, 1790) [2012: First subsequent record: 1975 Sitona lepidus] Number of subsequent records: many Pearce: 6 localities Latest record: 2020 First subsequent record: 1975 Number of subsequent records: many Sitona cylindricollis Fåhraeus, 1840 Latest record: 2020 Pearce: 2 localities First subsequent record: 1966 Sitona puncticollis Stephens, 1831 Number of subsequent records: many Pearce: 7 localities Latest record: 2018 First subsequent record: 1966 Number of subsequent records: 9 Latest record: 2008

Sitona striatellus Gyllenhal, 1834 Romualdius angustisetulus (Hansen, 1915) Pearce: generally common Pearce: no records (but see scaber) First subsequent record: 1937 First record: 1991 Number of subsequent records: many Number of records: many Latest record: 2020 Latest record: 2020

Sitona sulcifrons (Thunberg, 1798) Romualdius scaber (Linnaeus, 1758) Pearce: generally common [2012: Romualdius bifoveolatus] First subsequent record: 1979 Pearce: 6 localities (but could include Number of subsequent records: many angustisetulus) Latest record: 2020 First subsequent record: 1979 Number of subsequent records: many Sitona suturalis Stephens, 1831 Latest record: 2019 Pearce: 2 localities First subsequent record: 1955 Trachyphloeus alternans Gyllenhal, 1834 Number of subsequent records: many Pearce: 5 localities Latest record: 2020 First subsequent record: 1964 Number of subsequent records: many Sitona waterhousei Walton, 1846 Latest record: 2019 Pearce: 7 localities First subsequent record: 1955 Trachyphloeus digitalis (Gyllenhal, 1827) Number of subsequent records: many Pearce: no records Latest record: 2020 First record: 2002 Number of records: 1 Tanymecus palliatus (Fabricius, 1787) Latest record: 2002 Pearce: 3 localities First subsequent record: 1963 Trachyphloeus scabriculus (Linnaeus, Number of subsequent records: many 1771) Latest record: 2015 Pearce: 1 locality First subsequent record: 1979 Caenopsis fissirostris (Walton, 1847) Number of subsequent records: 9 Pearce: 1 locality Latest record: 2017 First subsequent record: 1979 Number of subsequent records: many Trachyphloeus spinimanus Germar, 1824 Latest record: 2020 Pearce: 1 locality No subsequent records Caenopsis waltoni (Boheman, 1842) Pearce: 2 localities Tropiphorus elevatus (Herbst, 1795) First subsequent record: 1977 Pearce: 2 localities Number of subsequent records: many First subsequent record: 1984 Latest record: 2020 Number of subsequent records: 6 Latest record: 2009 Cathormiocerus aristatus (Gyllenhal, 1827) Brachypera zoilus (Scopoli, 1763) [2012: Pearce: 1 locality (1896) Hypera zoilus] First subsequent record: 1979 Pearce: 8 localities Number of subsequent records: many First subsequent record: 1934 Latest record: 2019 Number of subsequent records: many Latest record: 2019 Cathormiocerus spinosus (Goeze, 1777) Pearce: 1 locality Hypera (Dapalinus) meles (Fabricius, First subsequent record: 1968 Number of subsequent records: many 1792) Latest record: 2020 Pearce: no records First record: 1979 Number of records: many Latest record: 2015 Hypera (Eririnomorphus) conmaculata Hypera (Hypera) venusta (Fabricius, (Herbst, 1795) [2012: Hypera pollux] 1781) Pearce: 3 localities Pearce: 6 localities First subsequent record: 1979 First subsequent record: 1963 Number of subsequent records: many Number of subsequent records: many Latest record: 2020 Latest record: 2019

Hypera (Eririnomorphus) rumicis Hypera (Kippenbergia) arator (Linnaeus, (Linnaeus, 1758) 1758) Pearce: generally common Pearce: 8 localities First subsequent record: 1955 First subsequent record: 1982 Number of subsequent records: many Number of subsequent records: many Latest record: 2019 Latest record: 2019

Hypera (Hypera) melancholica (Fabricius, Limobius mixtus (Boheman, 1834) 1792) [2012: Hypera fuscocinerea] Pearce: 2 localities Pearce: 2 localities No subsequent records No subsequent records Cleonis pigra (Scopoli, 1763) Hypera (Hypera) miles (Paykull, 1792) Pearce: 5 localities [2012: Hypera suspiciosa] First subsequent record: 1934 Pearce: 1 locality Number of subsequent records: 1 First subsequent record: 1979 Latest record: 1934 Number of subsequent records: 5 Latest record: 2020 Coniocleonus nebulosus (Linnaeus, 1758) Pearce: 6 localities Hypera (Hypera) nigrirostris (Fabricius, First subsequent record: 1975 Number of subsequent records: 6 1775) Latest record: 2003 Pearce: generally common First subsequent record: 1975 Number of subsequent records: many Larinus carlinae (Olivier, 1807) [2012: Latest record: 2020 Larinus planus] Pearce: 6 localities Hypera (Hypera) ononidis (Chevrolat, First subsequent record: 1937 Number of subsequent records: many 1863) Latest record: 2020 Pearce: no records First record: 1991 Number of records: 8 Lixus scabricollis Boheman, 1842 Latest record: 2020 Pearce: no records First record: 1998 Number of records: many Hypera (Hypera) plantaginis (De Geer, Latest record: 2020 1775) Pearce: 6 localities First subsequent record: 1955 Rhinocyllus conicus (Frölich, 1792) Number of subsequent records: many Pearce: 8 localities Latest record: 2020 First subsequent record: 1953 Number of subsequent records: many Latest record: 2019 Hypera (Hypera) postica (Gyllenhal, 1813) Magdalis (Edo) ruficornis (Linnaeus, Pearce: 8 localities First subsequent record: 1955 1758) Number of subsequent records: many Pearce: 2 localities Latest record: 2020 First subsequent record: 1981 Number of subsequent records: many Latest record: 2008

Magdalis (Odontomagdalis) armigera Syagrius intrudens Waterhouse, C.O., (Geoffroy in Fourcroy, 1785) 1903 Pearce: 2 localities Pearce: no records First subsequent record: 1979 First record: 2014 Number of subsequent records: many Number of records: 2 Latest record: 2011 Latest record: 2019

Magdalis (Odontomagdalis) carbonaria Pissodes castaneus (De Geer, 1775) (Linnaeus, 1758) Pearce: 2 localities Pearce: no records First subsequent record: 1979 First record: 1988 Number of subsequent records: many Number of records: 4 Latest record: 2019 Latest record: 2011 Pissodes pini (Linnaeus, 1758) Magdalis (Panus) barbicornis (Latreille, Pearce: no records 1804) First record: 1997 Pearce: no records Number of records: 9 First record: 1985 Latest record: 2011 Number of records: 1 Latest record: 1985 Trachodes hispidus (Linnaeus, 1758) Pearce: no records Magdalis (Porrothus) cerasi (Linnaeus, First record: 2001 Number of records: 1 1758) Latest record: 2001 Pearce: 2 localities First subsequent record: 1980 Number of subsequent records: many Orobitis cyanea (Linnaeus, 1758) Latest record: 2019 Pearce: 2 localities First subsequent record: 2002 Number of subsequent records: many Hylobius abietis (Linnaeus, 1758) Latest record: 2017 Pearce: fairly common and widely distributed First subsequent record: 1945 Number of subsequent records: many Platypus cylindrus (Fabricius, 1792) Latest record: 2019 Pearce: no records First record: 1993 Number of records: many Leiosoma deflexum (Panzer, 1795) Latest record: 2019 Pearce: 7 localities First subsequent record: 1938 Number of subsequent records: many Hylastes angustatus (Herbst, 1794) Latest record: 2019 Pearce: no records First record: 1979 Number of records: 7 Leiosoma oblongulum Boheman, 1842 Latest record: 2015 Pearce: 1 locality First subsequent record: 2018 Number of subsequent records: 2 Hylastes ater (Paykull, 1800) Latest record: 2018 Pearce: 3 localities First subsequent record: 1979 Number of subsequent records: many Liparus coronatus (Goeze, 1777) Latest record: 2019 Pearce: 6 localities First subsequent record: 1988 Number of subsequent records: many Hylastes attenuatus Erichson, 1836 Latest record: 2019 Pearce: no records First record: 1979 Number of records: 6 Latest record: 2015 Hylastes opacus Erichson, 1836 Tomicus minor (Hartig, 1834) Pearce: 2 localities Pearce: no records First subsequent record: 1983 First record: 2001 Number of subsequent records: many Number of records: 2 Latest record: 2019 Latest record: 2001

Hylurgops palliatus (Gyllenhal, 1813) Tomicus piniperda (Linnaeus, 1758) Pearce: 3 localities Pearce: 6 localities First subsequent record: 2004 First subsequent record: 1977 Number of subsequent records: many Number of subsequent records: many Latest record: 2016 Latest record: 2016

Hylastinus obscurus (Marsham, 1802) Phloeotribus rhododactylus (Marsham, Pearce: 1 locality 1802) First subsequent record: 1987 Pearce: 1 locality Number of subsequent records: 4 First subsequent record: 1981 Latest record: 2019 Number of subsequent records: many Latest record: 2020 Hylesinus crenatus (Fabricius, 1787) Pearce: 2 localities Pityophthorus pubescens (Marsham, 1802) First subsequent record: 1987 Pearce: 1 locality Number of subsequent records: 6 First subsequent record: 1991 Latest record: 2019 Number of subsequent records: 8 Latest record: 2020 Hylesinus taranio (Danthoine in Bernard, 1788) Ernoporicus fagi (Fabricius, 1798) Pearce: 2 localities Pearce: 1 locality (1825) First subsequent record: 2005 First subsequent record: 2019 Number of subsequent records: 3 Number of subsequent records: 1 Latest record: 2010 Latest record: 2019

Hylesinus varius (Fabricius, 1775) Trypophloeus binodulus (Ratzeburg, 1837) Pearce: 5 localities Pearce: no records First subsequent record: 1975 First record: 1998 Number of subsequent records: many Number of records: 1 Latest record: 2018 Latest record: 1998

Hylesinus wachtli Reitter, 1887 Dryocoetes autographus (Ratzeburg, Pearce: no records 1837) First record: 2008 Pearce: no records Number of records: 1 First record: 1990 Latest record: 2008 Number of records: 1 Latest record: 1990 Kissophagus vicinus (Comolli, 1837) Pearce: 1 locality Dryocoetes villosus (Fabricius, 1792) First subsequent record: 1986 Pearce: 1 locality Number of subsequent records: many First subsequent record: 1975 Latest record: 2020 Number of subsequent records: many Latest record: 2019 Pteleobius vittatus (Fabricius, 1792) Pearce: “Dorset” (A. Ford) Taphrorychus bicolor (Herbst, 1794) First subsequent record: 1975 Pearce: no records Number of subsequent records: 4 First record: 2017 Latest record: 1983 Number of records: 5 Latest record: 2019

Xylocleptes bispinus (Duftschmid, 1825) Xyleborinus saxesenii (Ratzeburg, 1837) Pearce: 1 locality Pearce: 1 locality First subsequent record: 1986 First subsequent record: 2004 Number of subsequent records: many Number of subsequent records: 4 Latest record: 2016 Latest record: 2020

Ips sexdentatus (Boerner, 1766) Xyleborus dryographus (Ratzeburg, 1837) Pearce: no records Pearce: no records First record: 2005 First record: 2004 Number of records: 3 Number of records: 1 Latest record: 2009 Latest record: 2004

Orthotomicus laricis (Fabricius, 1792) Xyleborus monographus (Fabricius, 1792) Pearce: 1 locality Pearce: no records First subsequent record: 1954 First record: 2019 Number of subsequent records: 3 Number of records: 2 Latest record: 2015 Latest record: 2019

Pityogenes bidentatus (Herbst, 1784) Trypodendron domesticum (Linnaeus, Pearce: 1 locality (1895) 1758) First subsequent record: 1989 Pearce: no records Number of subsequent records: 1 First record: 1981 Latest record: 1989 Number of records: many Latest record: 2019 Scolytus intricatus (Ratzburg, 1837) Pearce: 1 locality Trypodendron lineatum (Olivier, 1795) First subsequent record: 1991 Pearce: no records Number of subsequent records: many First record: 2008 Latest record: 2019 Number of records: 1 Latest record: 2008 Scolytus mali (Bechstein in Bechstein & Scharfenberg, 1805) Trypodendron signatum (Fabricius, 1792) Pearce: no records Pearce: no records First record: 1993 First record: 1970 Number of records: 4 Number of records: 6 Latest record: 2016 Latest record: 2020

Scolytus multistriatus (Marsham, 1802) Pearce: 1 locality First subsequent record: 1980 Number of subsequent records: 4 Latest record: 2019

Scolytus rugulosus (Müller, P.W.J., 1818) Pearce: 1 locality First subsequent record: 1970 Number of subsequent records: 8 Latest record: 2015

Scolytus scolytus (Fabricius, 1775) Pearce: 4 localities First subsequent record: 1975 Number of subsequent records: 3 Latest record: 1983