Molecular Phylogeny and Divergence Time of Trachypithecus: with Implications for the Taxonomy of T
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Cross-Species-Specific Binding Domain Spezifische Artenübergreifende Bindungsdomäne Domaine De Liaison Hétérospécifique
(19) TZZ ZZ_T (11) EP 2 520 590 B1 (12) EUROPEAN PATENT SPECIFICATION (45) Date of publication and mention (51) Int Cl.: of the grant of the patent: C07K 16/28 (2006.01) C07K 16/30 (2006.01) 15.08.2018 Bulletin 2018/33 C07K 16/32 (2006.01) C07K 16/42 (2006.01) C07K 16/46 (2006.01) C07K 14/725 (2006.01) (2006.01) (2006.01) (21) Application number: 12163656.7 A61K 39/395 A61P 35/00 (22) Date of filing: 03.04.2008 (54) Cross-species-specific binding domain Spezifische artenübergreifende Bindungsdomäne Domaine de liaison hétérospécifique (84) Designated Contracting States: •Kufer,Peter AT BE BG CH CY CZ DE DK EE ES FI FR GB GR 81477 Munich (DE) HR HU IE IS IT LI LT LU LV MC MT NL NO PL PT RO SE SI SK TR (74) Representative: Schiweck, Weinzierl & Koch Designated Extension States: Patentanwälte Partnerschaft mbB AL BA MK RS Landsberger Straße 98 80339 München (DE) (30) Priority: 03.04.2007 EP 07006990 03.04.2007 EP 07006988 (56) References cited: 24.04.2007 US 913668 P WO-A-2005/061547 WO-A-2005/118635 13.03.2008 EP 08004741 WO-A-2007/033230 WO-A-2007/042261 (43) Date of publication of application: • HAYASHI HIROKI ET AL: "A highly effective and 07.11.2012 Bulletin 2012/45 stable bispecific diabody for cancer immunotherapy: cure of xenografted tumors by (62) Document number(s) of the earlier application(s) in bispecific diabody and T-LAK cells", CANCER accordance with Art. 76 EPC: IMMUNOLOGY, IMMUNOTHERAPY : CII JUN 08735001.3 / 2 155 783 2004,, vol. -
Human Evolution: a Paleoanthropological Perspective - F.H
PHYSICAL (BIOLOGICAL) ANTHROPOLOGY - Human Evolution: A Paleoanthropological Perspective - F.H. Smith HUMAN EVOLUTION: A PALEOANTHROPOLOGICAL PERSPECTIVE F.H. Smith Department of Anthropology, Loyola University Chicago, USA Keywords: Human evolution, Miocene apes, Sahelanthropus, australopithecines, Australopithecus afarensis, cladogenesis, robust australopithecines, early Homo, Homo erectus, Homo heidelbergensis, Australopithecus africanus/Australopithecus garhi, mitochondrial DNA, homology, Neandertals, modern human origins, African Transitional Group. Contents 1. Introduction 2. Reconstructing Biological History: The Relationship of Humans and Apes 3. The Human Fossil Record: Basal Hominins 4. The Earliest Definite Hominins: The Australopithecines 5. Early Australopithecines as Primitive Humans 6. The Australopithecine Radiation 7. Origin and Evolution of the Genus Homo 8. Explaining Early Hominin Evolution: Controversy and the Documentation- Explanation Controversy 9. Early Homo erectus in East Africa and the Initial Radiation of Homo 10. After Homo erectus: The Middle Range of the Evolution of the Genus Homo 11. Neandertals and Late Archaics from Africa and Asia: The Hominin World before Modernity 12. The Origin of Modern Humans 13. Closing Perspective Glossary Bibliography Biographical Sketch Summary UNESCO – EOLSS The basic course of human biological history is well represented by the existing fossil record, although there is considerable debate on the details of that history. This review details both what is firmly understood (first echelon issues) and what is contentious concerning humanSAMPLE evolution. Most of the coCHAPTERSntention actually concerns the details (second echelon issues) of human evolution rather than the fundamental issues. For example, both anatomical and molecular evidence on living (extant) hominoids (apes and humans) suggests the close relationship of African great apes and humans (hominins). That relationship is demonstrated by the existing hominoid fossil record, including that of early hominins. -
A Unique Middle Miocene European Hominoid and the Origins of the Great Ape and Human Clade Salvador Moya` -Sola` A,1, David M
A unique Middle Miocene European hominoid and the origins of the great ape and human clade Salvador Moya` -Sola` a,1, David M. Albab,c, Sergio Alme´ cijac, Isaac Casanovas-Vilarc, Meike Ko¨ hlera, Soledad De Esteban-Trivignoc, Josep M. Roblesc,d, Jordi Galindoc, and Josep Fortunyc aInstitucio´Catalana de Recerca i Estudis Avanc¸ats at Institut Catala`de Paleontologia (ICP) and Unitat d’Antropologia Biolo`gica (Dipartimento de Biologia Animal, Biologia Vegetal, i Ecologia), Universitat Auto`noma de Barcelona, Edifici ICP, Campus de Bellaterra s/n, 08193 Cerdanyola del Valle`s, Barcelona, Spain; bDipartimento di Scienze della Terra, Universita`degli Studi di Firenze, Via G. La Pira 4, 50121 Florence, Italy; cInstitut Catala`de Paleontologia, Universitat Auto`noma de Barcelona, Edifici ICP, Campus de Bellaterra s/n, 08193 Cerdanyola del Valle`s, Barcelona, Spain; and dFOSSILIA Serveis Paleontolo`gics i Geolo`gics, S.L. c/ Jaume I nu´m 87, 1er 5a, 08470 Sant Celoni, Barcelona, Spain Edited by David Pilbeam, Harvard University, Cambridge, MA, and approved March 4, 2009 (received for review November 20, 2008) The great ape and human clade (Primates: Hominidae) currently sediments by the diggers and bulldozers. After 6 years of includes orangutans, gorillas, chimpanzees, bonobos, and humans. fieldwork, 150 fossiliferous localities have been sampled from the When, where, and from which taxon hominids evolved are among 300-m-thick local stratigraphic series of ACM, which spans an the most exciting questions yet to be resolved. Within the Afro- interval of 1 million years (Ϸ12.5–11.3 Ma, Late Aragonian, pithecidae, the Kenyapithecinae (Kenyapithecini ؉ Equatorini) Middle Miocene). -
Chapter 18 Body Size and Intelligence in Hominoid Evolution
Chapter 18 ................................ ................................ Body Size and Intelligence in Hominoid ................................ ................................ chapter Evolution ................................ ................................ Carol Ward ................................ Department of Anthropology – Department of Pathology and Anatomical Sciences ................................ 107 Swallow Hall, University of Missouri ................................ Columbia, MO 65211 USA ................................ [email protected] ................................ Mark Flinn ................................ Department of Anthropology, 107 Swallow Hall, University of Missouri ................................ Columbia, MO 65211 USA ................................ [email protected] ................................ David Begun ................................ Department of Anthropology, University of Toronto ................................ Toronto, ON M5S 3G3, Canada ................................ [email protected] ................................ ................................ Word count: text (6526) XML Typescript c Cambridge University Press – Generated by TechBooks. Body Size and Intelligence in Hominoid Evolution Page 1218 of 1365 A-Head 1 Introduction ................................ ................................ 2 Great apes and humans are the largest brained primates. Aside from a few ................................ 3 extinct subfossil lemurs, they are also the largest in body mass. -
Fossil Primates
AccessScience from McGraw-Hill Education Page 1 of 16 www.accessscience.com Fossil primates Contributed by: Eric Delson Publication year: 2014 Extinct members of the order of mammals to which humans belong. All current classifications divide the living primates into two major groups (suborders): the Strepsirhini or “lower” primates (lemurs, lorises, and bushbabies) and the Haplorhini or “higher” primates [tarsiers and anthropoids (New and Old World monkeys, greater and lesser apes, and humans)]. Some fossil groups (omomyiforms and adapiforms) can be placed with or near these two extant groupings; however, there is contention whether the Plesiadapiformes represent the earliest relatives of primates and are best placed within the order (as here) or outside it. See also: FOSSIL; MAMMALIA; PHYLOGENY; PHYSICAL ANTHROPOLOGY; PRIMATES. Vast evidence suggests that the order Primates is a monophyletic group, that is, the primates have a common genetic origin. Although several peculiarities of the primate bauplan (body plan) appear to be inherited from an inferred common ancestor, it seems that the order as a whole is characterized by showing a variety of parallel adaptations in different groups to a predominantly arboreal lifestyle, including anatomical and behavioral complexes related to improved grasping and manipulative capacities, a variety of locomotor styles, and enlargement of the higher centers of the brain. Among the extant primates, the lower primates more closely resemble forms that evolved relatively early in the history of the order, whereas the higher primates represent a group that evolved more recently (Fig. 1). A classification of the primates, as accepted here, appears above. Early primates The earliest primates are placed in their own semiorder, Plesiadapiformes (as contrasted with the semiorder Euprimates for all living forms), because they have no direct evolutionary links with, and bear few adaptive resemblances to, any group of living primates. -
The Placenta of the Colobinae Nghi™N C¯U V“ Nhau Thai Còa Nh„M Khÿ
Vietnamese Journal of Primatology (2008) 2, 33-39 The placenta of the Colobinae Kurt Benirschke University of California San Diego, Department of Pathology, USA 8457 Prestwick Drive La Jolla, CA 92037, USA <[email protected]> Key words: Colobinae, langurs, placenta, bilobed, hemochorial Summary Leaf-eating monkeys have a hemomonochorial placenta that is usually composed of two lobes and these are connected by large fetal vessels. In general, the placenta is similar to that of the rhesus monkey ( Macaca mulatta ) and, like that species, occasional placentas possess only a single lobe. This paper describes the structure, weights and cord lengths of all colobine monkeys examined by the author to date and it provides an overview of the placentation of langurs in general. Nghi™n c¯u v“ nhau thai cÒa nh„m khÿ ®n l∏ T„m tæt ô nh„m khÿ ®n l∏ (leaf-eating monkeys) nhau thai th≠Íng Æ≠Óc tπo bÎi hai thÔy, vµ hai thÔy nµy nËi vÌi nhau bÎi nh˜ng mπch m∏u lÌn tı bµo thai. Nh◊n chung, c†u tπo nhau thai cÒa nh„m nµy giËng Î nh„m khÿ vµng (Macaca mulatta ). Vµ cÚng nh≠ Î khÿ vµng, thÿnh tho∂ng nhau thai chÿ c„ mÈt thÔy. Trong nghi™n c¯u nµy t∏c gi∂ m´ t∂ c†u tπo, c©n n∆ng cÚng nh≠ chi“u dµi nhau thai cÒa c∏c loµi thuÈc nh„m khÿ ®n l∏. Qua Æ„ cung c†p th´ng tin toµn di÷n v“ c†u tπo nhau thai cÒa c∏c loµi khÿ ®n l∏. -
The Socioecology, and the Effects of Human Activity on It, of the Annamese Silvered Langur ( Trachypithecus Margarita ) in Northeastern Cambodia
The Socioecology, and the Effects of Human Activity on It, of the Annamese Silvered Langur ( Trachypithecus margarita ) in Northeastern Cambodia Álvaro González Monge A thesis submitted for the degree of Doctor of Philosophy of the Australian National University School of Archaeology and Anthropology Submitted in March, 2016 Copyright by Álvaro González Monge, 2016 All Rights Reserved Statement of originality The work presented in this thesis is, to the best of my knowledge and belief, original and my own work, except where acknowledged. This material has not been submitted either in whole or in part, for a degree at this or other university Álvaro González Monge In memoriam: GANG HU JOAQUIM JOSEP VEÀ BARÓ Acknowledgements This project wouldn’t have successfully arrived at its conclusion without the help of an astounding amount of people. I wanted to thank many more but I think two and a half pages of this must be testing for many. I’m forever indebted to my academic supervisors, for steering me towards meaningful research and pointing out my endless flaws with endless patience, for the encouragement and heaps of valuable feedback. Whatever useful information in this thesis is largely due to them: Professor Colin Groves, for accepting me as a student which I think is one of the highest honors that can be given to a person in our field of work, and his unquenchable thirst for all mammalian bits of information I brought to his attention. Dr. Alison Behie, for her patience in greatly helping me focus on the particular topics treated in this thesis and her invaluable feedback on my research. -
Fleagle and Lieberman 2015F.Pdf
15 Major Transformations in the Evolution of Primate Locomotion John G. Fleagle* and Daniel E. Lieberman† Introduction Compared to other mammalian orders, Primates use an extraordinary diversity of locomotor behaviors, which are made possible by a complementary diversity of musculoskeletal adaptations. Primate locomotor repertoires include various kinds of suspension, bipedalism, leaping, and quadrupedalism using multiple pronograde and orthograde postures and employing numerous gaits such as walking, trotting, galloping, and brachiation. In addition to using different locomotor modes, pri- mates regularly climb, leap, run, swing, and more in extremely diverse ways. As one might expect, the expansion of the field of primatology in the 1960s stimulated efforts to make sense of this diversity by classifying the locomotor behavior of living primates and identifying major evolutionary trends in primate locomotion. The most notable and enduring of these efforts were by the British physician and comparative anatomist John Napier (e.g., Napier 1963, 1967b; Napier and Napier 1967; Napier and Walker 1967). Napier’s seminal 1967 paper, “Evolutionary Aspects of Primate Locomotion,” drew on the work of earlier comparative anatomists such as LeGros Clark, Wood Jones, Straus, and Washburn. By synthesizing the anatomy and behavior of extant primates with the primate fossil record, Napier argued that * Department of Anatomical Sciences, Health Sciences Center, Stony Brook University † Department of Human Evolutionary Biology, Harvard University 257 You are reading copyrighted material published by University of Chicago Press. Unauthorized posting, copying, or distributing of this work except as permitted under U.S. copyright law is illegal and injures the author and publisher. fig. 15.1 Trends in the evolution of primate locomotion. -
STATUS and SOCIAL ORGANIZATION of HATINH LANGUR (Trachypithecus Hatinhensis Dao, 1970) in DONG HOA and THACH HOA COMMUNES FOREST, QUANG BINH PROVINCE
Management of Forest Resources and Environment STATUS AND SOCIAL ORGANIZATION OF HATINH LANGUR (Trachypithecus hatinhensis Dao, 1970) IN DONG HOA AND THACH HOA COMMUNES FOREST, QUANG BINH PROVINCE Dong Thanh Hai1, Thao A Tung2 1,2Vietnam National University of Forestry SUMMARY Hatinh Langur (Trachypithecus hatinhensis Dao, 1970) is endemic to Central Vietnam and Laos. In Vietnam, this species is restricted to two provinces of Quang Binh and Quang Tri. The species is nationally and globally threatened. Previous studies have shown that the social structure of leaf-eating monkeys is one-male or multimales and multifemales units. Group size ranges from 7 to 20 individuals. Studies on the social structure of T. hatinhensis are mostly carried out in Phong Nha - Ke Bang National Park, while other sites only investigate the status and distribution of species. The objective of this study was to determine the population status and social organization of the Hatinh langur in Dong Hoa and Thach Hoa Forest as a basis for proposing conservation measures. Linetransects and direct count methods was used to collect data. The results showed that there were 9 groups with the total number of 81 individuals. Social organization of groups of T. hatinhensis is similar to those patterns of species in subfamily colobinae that comprise of one-male and multimales and mutifemale units. Average group size is 9 ± 3 individuals. The ratio of adult males to females (AM/AF) is 1:2.5; infants to females (IF/AF) is 1:3; adults to immature individuals is 1:0.84. Keywords: Hatinh langur, population size, social structure. -
Ecology and Conservation of the Laotian Langur Trachypithecus Laotum in a Protected Area of Laos (Southeast Asia)
diversity Article Ecology and Conservation of the Laotian langur Trachypithecus laotum in a Protected Area of Laos (Southeast Asia) Johnny Souwideth 1 , Phaivanh Phiapalath 2, Hai Dong Thanh 1, Peter Brakels 3, Thong Pham Van 4 and Luca Luiselli 5,6,7,* 1 Vietnam National University of Forestry, Street 21, Xuan Mai Town, Chuong My District, Hanoi 100000, Vietnam; [email protected] (J.S.); [email protected] (H.D.T.) 2 Freelance Consultant, Vientiane, Laos; [email protected] 3 IUCN Vientiane Office, 391/24 Bourichane Road, Ban Naxay, Sayshetta District, Vientiane, Laos; [email protected] 4 Save Vietnam’s Wildlife, Cuc Phuong National Park, Cuc Phuong Commune, Nho Quan 430000, Vietnam; [email protected] 5 Institute for Development, Ecology, Conservation and Cooperation, Via G. Tomasi di Lampedusa 33, 00144 Rome, Italy 6 Department of Applied and Environmental Biology, Rivers State University of Science and Technology, P.M.B. 5080 Port Harcourt, Nigeria 7 Département de Zoologie et Biologie Animale, Faculté des Sciences, Université de Lomé, B.P. 1515 Lomé, Togo * Correspondence: [email protected] Abstract: Terrestrial species from the Lao People’s Democratic Republic (PDR) are under high threat due to deforestation and overhunting. Previous studies have even defined these forests as subjected to an “empty forest syndrome”, a condition in which forests that are apparently well preserved are instead almost deprived of vertebrate faunas due to extreme exploitation by local communities. Forest Citation: Souwideth, J.; Phiapalath, specialists, including several primates, are among the most threatened species in the country. The P.; Thanh, H.D.; Brakels, P.; Van, T.P.; Laotian langur (Trachypithecus laotum) is endemic to Lao PDR, is listed as Endangered by the IUCN Luiselli, L. -
Supporting Information
Supporting Information Moya` -Sola` et al. 10.1073/pnas.0811730106 SI Text netic relationships between all these taxa by classifying them all Systematic Framework. In Table S1 we provide a systematic into a single family Afropithecidae with 2 subfamilies (Keny- classification of living and fossil Hominoidea to the tribe level, apithecinae and Afropithecinae). by further including extant taxa and extinct genera discussed in The systematic scheme used here requires several nomencla- this paper. Hominoidea are defined as the group constituted by tural decisions, which deserve further explanation. The nomina Hylobatidae and Hominidae, plus all extinct taxa more closely Kenyapithecini and Kenyapithecinae are adopted instead of related to them than to Cercopithecoidea. Hominidae, in turn, Griphopithecinae and Griphopithecini (see also ref. 25) merely are defined as the group containing Ponginae and Homininae, because the former have priority. It is unclear why neither Begun plus all extinct forms more closely related to them than to (1) nor Kelley (2) specify the authorship of Griphopithecinae (or Hylobatidae. While this broad concept of Hominidae is currently Griphopithecidae), but, to our knowledge, the authorship of the used by many paleoprimatologists (e.g., refs. 1–2), the systematic latter nomina must be attributed to Begun (ref. 4, p. 232: Table position of primitive (or archaic) putative hominoids is far from 10.1), which therefore do not have priority over Kenyapithecinae clear (see below). Begun (3–5) employs the terms ‘‘Eohomi- Andrews, 1992. Griphopithecinae thus remains potentially valid noidea’’ and ‘‘Euhominoidea’’ to informally refer to hominoids only if Kenyapithecus (and Afropithecus, see below) are excluded of primitive and modern aspect, respectively. -
ASP December Newsletter
December 2004 ASP Bulletin Vol. 28, No. 4 Page 1 AMERICAN SOCIETY OF PRIMA TOLOGISTS Toni Ziegler - Executive Secretary F F F Volume 28, Number 4 December 2004 A Message from the President... Season’s in the process of putting together a primate-relevant venue. ASP is going greetings ASP! genomics symposium, a reproductive to play a pivotal role in how attendees This is a busy tract biology pre-meeting symposium, interface with the Congress organizers. time of year for and a teacher’s workshop (under the Specifically, the Congress registration everyone and I auspices of the Education committee). and abstract submission process for the just want to take In conjunction with the reproductive IPS Congress will be adapted from the a few minutes of tract biology symposium and courtesy of ASP technology for these functions. your time to the Oregon National Primate Research This is good for you for at least two update you on Center, we will have some extra money reasons. First, you will benefit from some of the this year to support student travel to the the efficient, user friendly technology things that are happening with ASP, and meeting. If you are a student, please pay that Nancy Capitanio has developed for IPS as well. special attention to the student travel Continued on page 2 The Program and Local Arrangements portions of the announcement. We also committees are hard at work in have some money from John Wiley & Look inside for preparation for ASP 2005 in Portland, Sons, the publisher of AJP, to support OR. Most of the essential information student travel.