DOCSLIB.ORG

Phylogenetic Paleobiology: Phenotypic Diversification and Evolutionary Radiation In

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Phylogenetic Paleobiology: Phenotypic Diversification and Evolutionary Radiation In Phylogenetic Paleobiology: Phenotypic Diversification and Evolutionary Radiation in Paleozoic Crinoids DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By David F. Wright Graduate Program in Geological Sciences The Ohio State University 2016 Dissertation Committee: William I. Ausich, Advisor Matthew R. Saltzman Lawrence A. Krissek John V. Freudenstein Copyrighted by David F. Wright 2016 Abstract Phylogenetic paleobiology is an interdisciplinary research program at the nexus of paleontology, systematics, and evolutionary biology. Specifically, phylogenetic paleobiology integrates the deep-time data, techniques, and geologic perspectives of paleontology with phylogeny-based statistical and computational approaches in macroevolutionary biology. Chapters contained within this dissertation revolve around two primary themes: (1) understanding patterns of biodiversity change over geologic time, and (2) assembling a clearer picture of the phylogeny, evolution, and geologic history of marine invertebrates, especially the Crinoidea (Echinodermata). Under the umbrella of phylogenetic paleobiology, the primary objective of this dissertation is to help bridge the disciplinary gap between specimen-based paleontology and statistical approaches in phylogenetic comparative methods. The chapters herein use advanced statistical phylogenetic methods, such as Bayesian “tip-dating” approaches, to infer phylogenetic trees of fossil species, quantify rates of phenotypic evolution, and document patterns of morphospace occupation among fossil members of the Crinoidea (Echinodermata). In addition to these broader studies, a major taxonomic revision of fossil and extant Crinoidea (Echinodermata) is proposed herein, as well as taxonomic description of a new genus of fossil crinoid from the Ordovician (Katian) of Ontario. ii This work is jointly (and lovingly) dedicated to (1) my parents, for being supportive of my decision to pursue academic interests; (2) to my wife Lena, for her encouragement, friendship, and kindness; and (3) to my dog Gluey, for showing me how beautiful the world can be. iii Acknowledgments This dissertation would not have been possible without the assistance and support of many individuals. I would like to thank to following people for their gracious assistance to museum collections: K. Hollis (United States Museum of Natural History), T. Ewin (Natural History Museum), P. Mayer (Field Museum of Natural History), and K. Riddigton (Lapworth Museum of Geology). J. Koniecki is thanked for collecting and generously donating specimens used in this dissertation. This dissertation was supported by numerous student research grants, including those from the Paleontological Society, Sigma Xi, the Palaeontological Association, the American Federation of Mineralogical and Geological Societies, the Association of Applied Paleontological Sciences, Friends of Orton Hall, and a Presidential Fellowship from The Ohio State University. Additional support for this research was provided by the National Science Foundation’s Assembling the Echinoderm Tree of Life (DEB 10364116) to W.I. Ausich. I have been very fortunate in my academic career to have had wonderful and gracious academic mentors who have helped guide my path. Greg Farley is thanked for igniting my earliest interests in science (pretty late by most standards!), letting me borrow his personal copy of S.J. Gould’s The Flamingo’s Smile, and encouraging me to seek a university degree. Bruce Lieberman is thanked for warmly encouraging my interests in macroevolutionary biology and paleontology when I was an undergraduate at the iv University of Kansas. I thank Alycia Stigall for her delightful (and highly contagious) enthusiasm for specimen-based paleontology. I cannot overstate how much my academic trajectory benefited from her guidance and training when I was a M.Sc. student at Ohio University. Alycia has continued to be a mentor and I thank her for her support and friendship. I would especially like to thank my Ph.D. advisor, Bill Ausich, for his invaluable (and seemingly encyclopaedic) knowledge of fossil crinoids and willingness to have a scientific discussion with me at a moment’s notice. Bill has been an excellent Ph.D. advisor, mentor, and friend. Most of all, I thank Bill for instilling in me a passion for studying fossil crinoids and introducing me to the delectable wonder of Bell’s Two- Hearted Ale. Many ideas presented in this dissertation were greatly aided through informal conversations with friends and colleagues over the years, including Dave Bapst, Tom Baumiller, Lena Cole, Tom Kammer, David Polly, Dan Rabosky, Graham Slater, Jeff Thompson, and Peter Wagner. I would also like to thank those who provided specific feedback on earlier drafts of the chapters presented herein and/or peer-reviewed the published portions of this dissertation, including W.I. Ausich, S.R. Cole, A. Gavryushkina, T.W. Kammer, G.D. Sevastopulo, G.J. Slater, and two anonymous reviewers. Stig Bergström and Dale Gnidovic are thanked for contributing an atmosphere of fossil-based comraderie that makes me proud to have been a part of Ohio State paleontology. I am thankful to have participated in several workshops that greatly improved my programming skills in R and led to my better understanding of analytical approaches in v the fields of paleontology and phylogenetic biology. I would like to thank the organizers, instructors, and fellow participants of the 2013 Analytical Methods in Paleobiology course held in Sydney, Australia; the National Evolutionary Synthesis Center sponsored “Paleobiological and Phylogenetic Approaches to Macroevolution” course held in Durham, NC; the Computational Macroevolution workshop at the 2015 meeting of the Society of Systematic Biologists in Ann Arbor, MI; and the “Phylogeny developeR bootcamp” held at the University of Massachusetts Boston’s field station in Nantucket, MA. I thank Lena Cole for putting up with my perambulating (and at times incomprehensible) vocalizations on various topics related to this dissertation. No one has listened to me ramble, rant, and think out loud about inferring phylogenies, statistical methods, comparative biology, and macroevolution more than her. Lena also read and helped edit my manuscript drafts and generously provided assistance with specimen illustration and photography. I have greatly benefited from her company both in and out of the office and will always appreciate her accompanying me on my daily walks to get coffee. I especially thank Lena for being my best friend, office mate, colleague, and loving spouse. I’ve never been able to relate to the well-circulated blog posts and comics on social media that promulgate the stereotype of unhappy, disgruntled graduate students. My experience has been the exact opposite. These four years as a Ph.D. student at Ohio State have been the best of my life. Many thanks to all of those whom I have not mentioned yet were a part of this wonderful experience. Cheers. –Davey vi Vita 2010................................................................B.S. Geology, University of Kansas 2012................................................................M.S. Geological Sciences, Ohio University 2012 to 2016 .................................................Graduate Teaching Associate, School of Earth Sciences, The Ohio State University 2016 to present ..............................................Presidential Fellow, The Ohio State University Publications Wright, D.F., W.I. Ausich, S.R. Cole, M.E. Peter, and E.C. Rhenberg, in press, Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata). Journal of Paleontology. Wright, D.F. in press, Bayesian estimation of fossil phylogenies and the evolution of early to middle Paleozoic crinoids (Echinodermata). Journal of Paleontology. Wright, D.F., 2015, Fossils, homology, and “Phylogenetic Paleo-ontogeny”: a reassessment of primary posterior plate homologies in fossil and living crinoids with insight from developmental biology. Paleobiology, 41:570-591. Ausich, W.I., T.W. Kammer, E.C. Rhenberg, and D.F. Wright, 2015, Early phylogeny of crinoids within the Pelmatozoan clade. Palaeontology, 58, 937-952. Wright, D.F. and W.I. Ausich, 2015, From the stem to the crown: phylogeny and diversification of pan-cladid crinoids. Pp. 199-202 in Progress in Echinoderm vii Palaeobiology. S. Zamora and I. Rábono (eds.), Cuadernos del museo Geominero, 19, Instituto Geológico y Minero de España, 292 pp. Ausich, W.I., T.W. Kammer, D.F. Wright, S.R. Cole, M.E. Peter, and E.C. Rhenberg, 2015, Toward a phylogenetic classification of the Crinoidea (Echinodermata). Pp. 29-32 in Progress in Echinoderm Palaeobiology. S. Zamora and I. Rábono (eds.), Cuadernos del museo Geominero, 19, Instituto Geológico y Minero de España, 292 pp. Wright, D.F. and A.L. Stigall, 2014, Species-level phylogenetic revision of the Late Ordovician orthid brachiopod genus Glyptorthis. Journal of Systematic Palaeontology, DOI 10.1080/14772019.2013.839584 Wright, D.F. and A.L. Stigall, 2013, Phylogenetic revision of the Late Ordovician orthid brachiopod genera Plaesiomys and Hebertella from North America. Journal of Paleontology, 87: 1107-1128. Wright, D.F. and A.L. Stigall, 2013, Geological drivers of Late Ordovician faunal change in Laurentia: investigating links between tectonics, speciation, and biotic invasions. PLoS ONE, 8(7): e68353. doi:10.1371/journal.pone.0068353 Wright, D.F. and P.A. Selden,
Load more

Recommended publications
  • Adhesion in Echinoderms
    Adhesion in echinoderms PATRICK FLAMMANG* Laboratoire de Biologie marine, Universite' de Mons-Hainaut, Mons, Belgium Final manuscript acceptance: August 1995 KEYWORDS: Adhesive properties, podia, larvae, Cuvierian tubules, Echinodermata. CONTENTS 1 Introduction 2 The podia 2.1 Diversity 2.2 Basic structure and function 2.3 Adhesivity 3 Other attachment mechanisms of echinoderms 3.1 Larval and postlarval adhesive structures 3.2 Cuvierian tubules 4 Comparison with other marine invertebrates 5 Conclusions and prospects Acknowledgements References 1 INTRODUCTION Marine organisms have developed a wide range of mechanisms allowing them to attach to or manipulate a substratum (Nachtigall 1974). Among 1 these mechanisms, one can distinguish between mechanical attachments (e.g. hooks or suckers) and chemical attachments (with adhesive sub- stances). The phylum Echinodermata is quite exceptional in that all its species, *Senior research assistant, National Fund for Scientific Research, Belgium. I whatever their life style, use attachment mechanisms. These mechanisms allow some of them to move, others to feed, and others to burrow in par- ticulate substrata. In echinoderms, adhesivity is usually the function of specialized structures, the podia or tube-feet. These podia are the exter- nal appendages of the arnbulacral system and are also probably the most advanced hydraulic structures in the animal kingdom. 2 THE PODIA From their presumed origin as simple respiratory evaginations of the am- bulacral system (Nichols 1962), podia have diversified into the wide range of specialized structures found in extant echinoderms. This mor- phological diversity of form reflects the variety of functions that podia perform (Lawrence 1987). Indeed, they take part in locomotion, burrow- ing, feeding, sensory perception and respiration.
    [Show full text]
  • Crinoidea, Echinodermata) from Poland
    Title: Uncovering the hidden diversity of Mississippian crinoids (Crinoidea, Echinodermata) from Poland Author: Mariusz A. Salamon, William I. Ausich, Tomasz Brachaniec, Bartosz J. Płachno, Przemysław Gorzelak Citation style: Salamon Mariusz A., Ausich William I., Brachaniec Tomasz, Płachno Bartosz J., Gorzelak Przemysław. (2020). Uncovering the hidden diversity of Mississippian crinoids (Crinoidea, Echinodermata) from Poland. "PeerJ" Vol. 8 (2020), art. no e10641, doi 10.7717/peerj.10641 Uncovering the hidden diversity of Mississippian crinoids (Crinoidea, Echinodermata) from Poland Mariusz A. Salamon1, William I. Ausich2, Tomasz Brachaniec1, Bartosz J. Pªachno3 and Przemysªaw Gorzelak4 1 Faculty of Natural Sciences, Institute of Earth Sciences, University of Silesia in Katowice, Sosnowiec, Poland 2 School of Earth Sciences, Ohio State University, Columbus, OH, United States of America 3 Faculty of Biology, Institute of Botany, Jagiellonian University in Kraków, Cracow, Poland 4 Institute of Paleobiology, Polish Academy of Sciences, Poland, Warsaw, Poland ABSTRACT Partial crinoid crowns and aboral cups are reported from the Mississippian of Poland for the first time. Most specimens are partially disarticulated or isolated plates, which prevent identification to genus and species, but regardless these remains indicate a rich diversity of Mississippian crinoids in Poland during the Mississippian, especially during the late Viséan. Lanecrinus? sp. is described from the late Tournaisian of the D¦bnik Anticline region. A high crinoid biodiversity occurred during late Viséan of the Holy Cross Mountains, including the camerate crinoids Gilbertsocrinus? sp., Platycrinitidae Indeterminate; one flexible crinoid; and numerous eucladid crinoids, including Cyathocrinites mammillaris (Phillips), three taxa represented by partial cups left in open nomenclature, and numerous additional taxa known only from isolated radial plates, brachial plates, and columnals.
    [Show full text]
  • Two New Crinoids from Lower Mississippian Rocks in Southeastern Kentucky
    TWO NEW CRINOIDS FROM LOWER MISSISSIPPIAN ROCKS IN SOUTHEASTERN KENTUCKY BY GEORGE M. EHLERS AND ROBERT V. KESLING Reprinted from JOURNAL OF PALEONTOLOGY Val. 37, No. 5, September, 1963 JOURNALOF PALEONTOLOGY,V. 37, NO. 5, P. 1028-1041, PLS. 133,134, 3 TEXT-FIGS., SEPTEMBER,1963 TWO NEW CRINOIDS FROM L20\'C7ERMISSISSIPPIAN ROCKS IN SOUTHEASTERN KENTUCKY GEORGE M. EHLERS AKD ROBERT V. ICESLING Museum of Paleontology, The University of Michigan .~BsTR.~~T-AII~~~~specimens collected many years ago bl- the senior author and his students near Mill Springs, Kentucky, are a new species of Agaricocrinzis and a new speries of Actino- crinites. Although only one specimen of each is known, it is well preserved. The new Agnrico- crinus bears a resemblance to A. ponderoszts Wood, and the new Actinocriniles to four species described by Miller & Gurley: A. spergenensis, A. botuztosz~s,A. gibsoni, and A. shnronensis. A preliminary survey of species assigned to Agaricocrinz~ssuggests that revision of the genus is overdue. Although the occurrence of the specimens leaves some doubt as to their stratigraphic posi- tion, we conclude that they both probably weathered from the Fort Payne formation and rolled down the slope onto the New Providence, where they were found. The sites where the crinoids were picked up are now deeply inundated by water impounded by the Wolf Creek dam on the Cumberland River. INTRODUCTION onto the New Providence, \$here they were OTH of the new crinoids described here are found. rZt present, both the New Providence B from Lower Mississippian rocks in the valley formation and the I~asalbeds of the Fort Payne of the Cumberland River in Wayne and Russell are underwater at the type locality of the new Counties, Kentucky.
    [Show full text]
  • "Lophophorates" Brachiopoda Echinodermata Asterozoa
    Deuterostomes Bryozoa Phoronida "lophophorates" Brachiopoda Echinodermata Asterozoa Stelleroidea Asteroidea Ophiuroidea Echinozoa Holothuroidea Echinoidea Crinozoa Crinoidea Chaetognatha (arrow worms) Hemichordata (acorn worms) Chordata Urochordata (sea squirt) Cephalochordata (amphioxoius) Vertebrata PHYLUM CHAETOGNATHA (70 spp) Arrow worms Fossils from the Cambrium Carnivorous - link between small phytoplankton and larger zooplankton (1-15 cm long) Pharyngeal gill pores No notochord Peculiar origin for mesoderm (not strictly enterocoelous) Uncertain relationship with echinoderms PHYLUM HEMICHORDATA (120 spp) Acorn worms Pharyngeal gill pores No notochord (Stomochord cartilaginous and once thought homologous w/notochord) Tornaria larvae very similar to asteroidea Bipinnaria larvae CLASS ENTEROPNEUSTA (acorn worms) Marine, bottom dwellers CLASS PTEROBRANCHIA Colonial, sessile, filter feeding, tube dwellers Small (1-2 mm), "U" shaped gut, no gill slits PHYLUM CHORDATA Body segmented Axial notochord Dorsal hollow nerve chord Paired gill slits Post anal tail SUBPHYLUM UROCHORDATA Marine, sessile Body covered in a cellulose tunic ("Tunicates") Filter feeder (» 200 L/day) - perforated pharnx adapted for filtering & repiration Pharyngeal basket contractable - squirts water when exposed at low tide Hermaphrodites Tadpole larvae w/chordate characteristics (neoteny) CLASS ASCIDIACEA (sea squirt/tunicate - sessile) No excretory system Open circulatory system (can reverse blood flow) Endostyle - (homologous to thyroid of vertebrates) ciliated groove
    [Show full text]
  • Bedrock Geology of Altenburg Quadrangle, Jackson County
    BEDROCK GEOLOGY OF ALTENBURG QUADRANGLE Institute of Natural Resource Sustainability William W. Shilts, Executive Director JACKSON COUNTY, ILLINOIS AND PERRY COUNTY, MISSOURI STATEMAP Altenburg-BG ILLINOIS STATE GEOLOGICAL SURVEY E. Donald McKay III, Interim Director Mary J. Seid, Joseph A. Devera, Allen L. Weedman, and Dewey H. Amos 2009 360 GEOLOGIC UNITS ) ) ) 14 Qal Alluvial deposits ) 13 18 Quaternary Pleistocene and Holocene 17 360 ) 15 360 16 14 0 36 ) 13 Qf Fan deposits ) Unconformity Qal ) & 350 tl Lower Tradewater Formation Atokan ) ) Pennsylvanian 360 ) &cv Caseyville Formation Morrowan 24 360 ) Unconformity ) 17 Upper Elviran undivided, Meu ) Waltersburg to top of Degonia 19 20 Qal 21 22 23 ) 24 ) Mv Vienna Limestone 360 o ) 3 Mts ) 350 Mts Tar Springs Sandstone ) 20 360 ) Mgd 360 30 ) Mgd Glen Dean Limestone ) 21 350 360 Mts 29 ) Qal Hardinsburg Sandstone and J N Mhg Chesterian ) Golconda Formations h Æ Qal Mav anc 28 27 Br ) N oJ 26 25 JN 85 N ) Cypress Sandstone through J Mcpc Dsl 500 Paint Creek Formation JN N ) J o Mts N 5 J s ) Dgt 600 J N 70 J N Mgd Yankeetown Formation s ) Myr Db 80 28 Æ and Renault Sandstone N J 29 N J N ) Sb J Mgd Mississippian o Dgt Ssc 25 Clines o N 25 Msg 27 ) Qal J 80 s 3 Mav Aux Vases Sandstone N J N Mts o MILL J MISSISSIPPI 34 ) Qal J N ) N J Dsl 35 N 26 J o N 25 J Mgd Mgd ) Msg Ste. Genevieve Limestone 500 o Db DITCH J 20 Mgd N N N ) J J o RIVER o N 600 J 80 N ) 10 o J Mav Æ Msl St.
    [Show full text]
  • Evidence for Cospeciation Events in the Host–Symbiont System Involving Crinoids (Echinodermata) and Their Obligate Associates, the Myzostomids (Myzostomida, Annelida)
    Molecular Phylogenetics and Evolution 54 (2010) 357–371 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Evidence for cospeciation events in the host–symbiont system involving crinoids (Echinodermata) and their obligate associates, the myzostomids (Myzostomida, Annelida) Déborah Lanterbecq a,*, Grey W. Rouse b, Igor Eeckhaut a a Marine Biology Laboratory, University of Mons, 6 Av. du Champ de Mars, Bât. Sciences de la vie, B-7000 Mons, Belgium b Scripps Institution of Oceanography, University of California, San Diego, La Jolla, CA 92093-0202, USA article info abstract Article history: Although molecular-based phylogenetic studies of hosts and their associates are increasingly common Received 14 April 2009 in the literature, no study to date has examined the hypothesis of coevolutionary process between Revised 3 August 2009 hosts and commensals in the marine environment. The present work investigates the phylogenetic Accepted 12 August 2009 relationships among 16 species of obligate symbiont marine worms (Myzostomida) and their echino- Available online 15 August 2009 derm hosts (Crinoidea) in order to estimate the phylogenetic congruence existing between the two lin- eages. The combination of a high species diversity in myzostomids, their host specificity, their wide Keywords: variety of lifestyles and body shapes, and millions years of association, raises many questions about Coevolution the underlying mechanisms triggering their diversification. The phylogenetic
    [Show full text]
  • Smithsonian Miscellaneous Collections [Vol
    NEW GENERA OF RECENT FREE CRINOIDS By AUSTIN HOBART CLARK Assistant, Bureau of Fisheries Since the publication of Dr. P. H. Carpenter's great monograph on the recent unstalked crinoids in 1888, the group has received very- little attention from systematists, probably because of the rarity of most of the species and the difficulty in getting together representative material of even the more common ones. Dr. Carpenter included in the family Comatulidse the genera Thaumatocrinus, Eudiocrinus, Promachocrinus (including the subsequently differentiated Decame- trocrinus), Atelecrinus, Antedon, Comatula (=Actinometra) , and Thiolliericrinus. All of these, with the exception of Antedon and Comatula, are comparatively small, strictly homogeneous genera; with them, however, the case is quite different. The genus Antedon was divided by Dr. Carpenter into four "series," and all but the first series into two or more "groups," the characters used in the differ- entiation of the groups and series being (1) the character of the joint between the costals, (2) the number of arms, (3) the number of the distichals, (4) the character of the lower pinnules, (5) the development or absence of covering plates on the ambulacra, and (6) the rounded or "wall-sided" character of the costals and lower brachials. Of all these characters, the first alone is the only one not common to two or more of his series or groups, as diagnosed by him. Taking No. 2, for instance, five of his groups and also several unassigned species are ten-armed ; all the rest have more than ten arms. A number of single species have both ten and more than ten arms, as a result of purely individual variation.
    [Show full text]
  • Progress in Echinoderm Paleobiology
    Journal of Paleontology, 91(4), 2017, p. 579–581 Copyright © 2017, The Paleontological Society 0022-3360/17/0088-0906 doi: 10.1017/jpa.2017.20 Progress in echinoderm paleobiology Samuel Zamora1,2 and Imran A. Rahman3 1Instituto Geológico y Minero de España (IGME), C/Manuel Lasala, 44, 9ºB, 50006, Zaragoza, Spain 〈[email protected]〉 2Unidad Asociada en Ciencias de la Tierra, Universidad de Zaragoza-IGME, Zaragoza, Spain 3Oxford University Museum of Natural History, Parks Road, Oxford, OX1 3PW, United Kingdom 〈[email protected]〉 Echinoderms are a diverse and successful phylum of exclusively Universal elemental homology (UEH) has proven to be one marine invertebrates that have an extensive fossil record dating of the most powerful approaches for understanding homology back to Cambrian Stage 3 (Zamora and Rahman, 2014). There in early pentaradial echinoderms (Sumrall, 2008, 2010; Sumrall are five extant classes of echinoderms (asteroids, crinoids, and Waters, 2012; Kammer et al., 2013). This hypothesis echinoids, holothurians, and ophiuroids), but more than 20 focuses on the elements associated with the oral region, extinct groups, all of which are restricted to the Paleozoic identifying possible homologies at the level of specific plates. (Sumrall and Wray, 2007). As a result, to fully appreciate the Two papers, Paul (2017) and Sumrall (2017), deal with the modern diversity of echinoderms, it is necessary to study their homology of plates associated with the oral area in early rich fossil record. pentaradial echinoderms. The former contribution describes and Throughout their existence, echinoderms have been an identifies homology in various ‘cystoid’ groups and represents a important component of marine ecosystems.
    [Show full text]
  • TREATISE TREATISE Paleo.Ku.Edu/Treatise
    TREATISE TREATISE paleo.ku.edu/treatise For each fossil group, volumes describe and illustrate: morphological features; ontogeny; classification; geologic distribution; evolutionary trends and phylogeny; and All 53 volumes of the Treatise are now available as fully searchable pdf systematic descriptions. Each volume is fully indexed and files on CD, even out-of-print and superseded volumes! Also available contains a complete set of references, and there are many is the complete set of volumes, either on one DVD, or a set of 41 CDs, detailed illustrations and plates throughout the series. each decorated with a different fossil. See below for details. Complete set of 53 Treatise volumes Part B Single DVD or 41 CDs: $1850.00 Protoctista 1: Charophyta, vol. 1 Edited by Roger L. Kaesler, with coordinating author, Monique Feist, Porifera (Revised), vol. 2 Part F leading a team of international specialists, 2005 edited by Roger L. Kaesler; coordinating author, J. Keith Rigby, with First volume of Part B to be published, covering generally plantlike authors R. E. H. Reid, R.M. Finks, and J. Keith Rigby, 2003 Coelenterata, Supplement 1, Rugosa and Tabulata, vol. 1–2 autotrophic protoctists. Future volumes will cover the dinoflagellates, Second volume in the revision of Porifera. General features of the Porifera, edited by C. Teichert, coordinating author Dorothy Hill, 1981 silicoflagellates, ebredians, benthic calcareous algae, coccolithophorids, including glossary, references, and index. Introduction to Paleozoic corals and systematic descriptions for Rugosa; and diatoms. Included in the charophyte volume are introductory chapters 376 p., ISBN 0-8137-3130-5, $75.00, hard copy or CD introduction to and systematic descriptions for Tabulata.
    [Show full text]
  • PROGRAMME ABSTRACTS AGM Papers
    The Palaeontological Association 63rd Annual Meeting 15th–21st December 2019 University of Valencia, Spain PROGRAMME ABSTRACTS AGM papers Palaeontological Association 6 ANNUAL MEETING ANNUAL MEETING Palaeontological Association 1 The Palaeontological Association 63rd Annual Meeting 15th–21st December 2019 University of Valencia The programme and abstracts for the 63rd Annual Meeting of the Palaeontological Association are provided after the following information and summary of the meeting. An easy-to-navigate pocket guide to the Meeting is also available to delegates. Venue The Annual Meeting will take place in the faculties of Philosophy and Philology on the Blasco Ibañez Campus of the University of Valencia. The Symposium will take place in the Salon Actos Manuel Sanchis Guarner in the Faculty of Philology. The main meeting will take place in this and a nearby lecture theatre (Salon Actos, Faculty of Philosophy). There is a Metro stop just a few metres from the campus that connects with the centre of the city in 5-10 minutes (Line 3-Facultats). Alternatively, the campus is a 20-25 minute walk from the ‘old town’. Registration Registration will be possible before and during the Symposium at the entrance to the Salon Actos in the Faculty of Philosophy. During the main meeting the registration desk will continue to be available in the Faculty of Philosophy. Oral Presentations All speakers (apart from the symposium speakers) have been allocated 15 minutes. It is therefore expected that you prepare to speak for no more than 12 minutes to allow time for questions and switching between presenters. We have a number of parallel sessions in nearby lecture theatres so timing will be especially important.
    [Show full text]
  • Reinterpretation of the Enigmatic Ordovician Genus Bolboporites (Echinodermata)
    Reinterpretation of the enigmatic Ordovician genus Bolboporites (Echinodermata). Emeric Gillet, Bertrand Lefebvre, Véronique Gardien, Emilie Steimetz, Christophe Durlet, Frédéric Marin To cite this version: Emeric Gillet, Bertrand Lefebvre, Véronique Gardien, Emilie Steimetz, Christophe Durlet, et al.. Reinterpretation of the enigmatic Ordovician genus Bolboporites (Echinodermata).. Zoosymposia, Magnolia Press, 2019, 15 (1), pp.44-70. 10.11646/zoosymposia.15.1.7. hal-02333918 HAL Id: hal-02333918 https://hal.archives-ouvertes.fr/hal-02333918 Submitted on 13 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. 1 Reinterpretation of the Enigmatic Ordovician Genus Bolboporites 2 (Echinodermata) 3 4 EMERIC GILLET1, BERTRAND LEFEBVRE1,3, VERONIQUE GARDIEN1, EMILIE 5 STEIMETZ2, CHRISTOPHE DURLET2 & FREDERIC MARIN2 6 7 1 Université de Lyon, UCBL, ENSL, CNRS, UMR 5276 LGL-TPE, 2 rue Raphaël Dubois, F- 8 69622 Villeurbanne, France 9 2 Université de Bourgogne - Franche Comté, CNRS, UMR 6282 Biogéosciences, 6 boulevard 10 Gabriel, F-2100 Dijon, France 11 3 Corresponding author, E-mail: [email protected] 12 13 Abstract 14 Bolboporites is an enigmatic Ordovician cone-shaped fossil, the precise nature and systematic affinities of 15 which have been controversial over almost two centuries.
    [Show full text]
  • Two New Brittle Star Species of the Genus Ophiothrix
    Caribbean Journal of Science, Vol. 41, No. 3, 583-599, 2005 Copyright 2005 College of Arts and Sciences University of Puerto Rico, Mayagu¨ez Two New Brittle Star Species of the Genus Ophiothrix (Echinodermata: Ophiuroidea: Ophiotrichidae) from Coral Reefs in the Southern Caribbean Sea, with Notes on Their Biology GORDON HENDLER Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, California 90007, U.S.A. [email protected] ABSTRACT.—Two new species, Ophiothrix stri and Ophiothrix cimar, inhabit shallow reef-platforms and slopes in the Southern Caribbean, and occur together at localities in Costa Rica and Panama, nearly to Colombia. What appears to be an undescribed species resembling O. cimar has been reported from eastern Venezuela. In recent years, reefs where the species were previously observed have deteriorated because of environmental degradation. As a consequence, populations of the new species may have been reduced or eradicated. The new species have previously been mistaken for O. angulata, O. brachyactis, and O. lineata. Ophiothrix lineata, O. stri, and O. cimar have in common a suite of morphological features pointing to their systematic affinity, and a similar pigmentation pattern consisting of a thin, dark, medial arm stripe flanked by two pale stripes. Ophiothrix lineata is similar to Indo-Pacific members of the subgenus Placophiothrix and closely resembles Ophiothrix stri. The latter is extremely similar to O. synoecina, from Colombia, and both can live in association with the rock-boring echinoid Echinometra lucunter. Although O. synoecina is a protandric hermaphrodite that reportedly broods its young externally, the new species are gonochoric and do not brood.
    [Show full text]