TULANE STUDIES in GEOLOGY and PALEONTOLOGY Volume 23
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TULANE STUDIES IN GEOLOGY AND PALEONTOLOGY Volume 23, Numbers 1-3 September 26, 1990 CENOZOIC MURICIDAE OF THE WESTERN ATLANTIC REGION PARTVIII-MUREXS .S ., HAUSTELLUM , CHICOREUS , ANDHEXAPLEX; ADDITIONS AND CORRECTIONS EMILY H . VOKES TULANE UNIVERSITY CONTENTS Page I. ABSTRACT . 1 IL INTRODUCTION . 1 III. STRATIGRAPHIC CORRELATION . ................. 2 IV. ACKNOWLEDGMENTS . ................. 5 V. SYSTEMATIC DESCRIPTIONS . .. ..... ... .... .. 5 VI. LOCALITY DATA . 80 VIL LITERATURE CITED ... ....... ... ... .................. .. ... 87 VIII. APPENDIX I: Alphabetical list of species . 92 IX. APPENDIX II: Stratigraphical list of species . 94 Table 1. Genera of the subfamily Muricinae that occur in the western Atlantic . 2 Table 2. Fossil species originally covered in part I . ........ ..... ..... 9 I. ABSTRACT Chicoreus (Siratus) sextoni and Chicoreus A total of 165 names have been applied (Phyllonotus) louisae , from the Chipola to western Atlantic species referred herein Formation, Florida ; Chicoreus (Phyl to the muricid genera Murex s.s., Haustel lonotus) initialis, from the Weches Forma lum, Hexaplex, and Chicoreus , including tion, Texas ; and Hexaplex isthmicus , from its four subgenera: Chicoreus s.s., Siratus , the Gatun Formation, Panama . Phyllonotus, and Naquetia . Of these, 101 are valid and are treated systematically; II. INTRODUCTION however, the two species of Murex s.s. The first part of this study appeared in present are only questionably referred to 1963 as a monograph on the genus Murex the western Atlantic fauna . Of the remain sensu stricto. In the succeeding years six ing 99 valid species there are 16 of Haustel additional parts appeared ; the last, that on lum, 22 of Chicoreus s.s., 28 of Siratus , 19 the genera Calotrophon and Attiliosa , was of Phyllonotus , one of Naquetia, and 13 of published in 1976. It was my plan at the Hexaplex. Of these 99 species, eight occur conclusion of the study of the subfamily in Eocene beds, with the oldest known Muricinae, which was completed with this from the Middle Eocene Weches Forma latter part, to present a part VIII - Sum tion of Texas; seven occur in Oligocene mary and Corrections for the entire sub beds; 31 in Miocene formations; and 46 in family, before embarking upon the sub Plio-Pleistocene strata . There are 42 families Muricopsinae and Ocenebrinae. species in the Recent fauna , but only 18 However, in 1976, my husband Harold are confined to the Recent, the remainder and I made the first of many collecting trips are both living and fossil. Of this number 9 to the Dominican Republic. This first visit are new species and are named herein. resulted in so many new taxa that a deci They are : Haustellum mimiwilsoni, from sion was made to do a separate paper on the Cumana Formation of Venezuela and the Dominican muricids, in which these the Rio Banano Formation of Costa Rica; taxa would be described before being in Chicoreus (Chicoreus) jungi , from the corporated into the proposed Summary . Grand Bay Formation, Carriacou, West In Little did I suspect that this project would dies; Chicoreus (Chicoreus) veronica and require 13 years and culminate finally in a Chicoreus (Siratus ) habros , from the different Part 8 - the Neogene Paleontology Agueguexquite Formation, Mexico ; of the northern Dominican Republic Chicoreus (Siratus) miltos, from the Con (Vokes, 1989b), which forms only a portion cepcion Inferior Formation , Mexico ; of a major study on the fauna of the Mio- 1 1/ 2 Tulane Studies in Geology and Paleontology Vol. 23 Pliocene of that country. It is fortunate that changes will be discussed with the relevant I waited, for in the Dominican report 32 taxonomic groups. A general reference species of Muricinae are treated, of which chart appears below as Table 1, to indicate 13 are described therein. Nineteen of the where supraspecific taxa are currently 32 are species of the genera covered in the placed. present work; the remainder will be treated in subsequent parts. III. STRATIGRAPHIC CORRELAT ION When Part I of Cenozoic Muricidae was In the interval since this serie s began published in 1963, the Tulane Geological great strides have been made in long. Collections consisted of material from only range geologic correlation by means of 555 localities; today the localities number planktic foraminifera and calcareou s nan over 1500. The Cenozoic localities have in nofossils. Rather than repeat the discus creased from 160 to more than 1000. Re sion for every species that occurs in these cent collecting localities represented have formations it seems more useful to list all of grown from 42 to nearly 600. Thus, a great them together with the most recent infor deal of n ew inform ation has been accumu mation on correlations . It should be noted lated. that the formations discussed here are only Much of this has been published al those in which species of Muricinae occur. ready , as in the case of the Dominican Re This is not intended to be a definitive anal public material. I have described new ysis of all Neogene formations in the west species of Murex (1967a), and new species ern Atlantic. For the best synthesis of the of Chicoreus (1974a). Petuch (1987, and stratigraphy (Upper Cretaceous to Holo elsewhere) has named several new forms . cene) of the Atlantic and Gulf Coastal Plain But the tim e has come to assimilate this the reader is referred to Carter (1984). scattered information into one (currently) Unfortunately, there is a recurring prob complete summary of all species of lem in that the formations in which mol Muricinae , fossil and Recent, presently lusks are most abundant are those near known from the western Atlantic region. shore facies where the planktic foraminif In addition to new material , several era and calcareous nannofossils are least taxonomic changes shifting species into dif likely to be encountered. In the deep ferent genera, and genera into differ ent water faci es where the latter are most subfamilies have been made. These abundant, mollusks are almost nonexis- TABLE 1 Genera of the subfamily Muricinae that occur in the western Atlantic. Cover ed in this paper Mure x s.s. [?no ne in the W es tern Atlantic] Haustellum Chico re us Chicoreus s.s., Siratus , Phyllonotus , Naquetia Hexa plex To be covered in next pap er Pterynotus Pterynotus s. s., Pterochelu s, Purpur ellus Poirieria Poi rieri a s.s., Pa zie lla , Pa zi notus , Flexopteron Aspella Dermo murex Dermomurex s.s., Takia , Gracilimur ex, Tri alatella Calotrophon Attiliosa Nos. 1-3 Cenozoic Muricidae - VIII 3 tent. Thus, there are many formations for dard by which all others were measured. which we have no better correlation data Unfortunately, due to lack of knowledge of than we did 30 years ago. the living Caribbean fauna, in particular In general, there has been little change deeper water forms, the Lyellian tech in the stratigraphic assignments of the nique of percent of living species gave an Paleogene formations. An excellent corre age that was older. More recent planktic lation chart for the Paleogene of the Gulf evidence places the Bowden shell bed in Coastal Plain has been given by Dockery the basal Pleistocene (N.22) rather than (1980, pp. 22-23). The Paleocene, Eocene, Middle Miocene, as it was originally con and Oligocene units are essentially un sidered by Woodring (see Lamb and changed . However, the Silverdale beds of Beard, 1972, p. 32). North Carolina, originally placed in the The Shoal River Formation of north Early Miocene are now placed in the latest western Florida is still placed in the Middle Oligocene (Chickasawhayan Stage), as is Miocene (Carter, 1984, zones N.8 -13). The the Tampa Limestone of peninsular Yellow River Formation (formerly the Florida. "Yoldia Faunizone" of the Choctaw The Chipola Formation of northwestern hatchee Formation, now Group) has been Florida was the subject of a detailed study demonstrated to be Middle Miocene (N .14) by Akers (1972), who determined that the on the basis of the rich planktic fauna formation is Burdigalian (planktic zones (Akers, 1972, p. 12). In the Atlantic Coastal N.4-8) rather than Helvetian as I once Plain section the Calvert, Choptank, and suggested (Vokes, 1965b, p. 205). The re St. Marys formations of Maryland and semblances to the Helvetian beds in south northeastern Virginia, and the Kirkwood ern France are evidently due more to ecol Formation of New Jersey, are still consid ogy than time . The Cantaure Formation of ered to be Middle Miocene (Blackwelder Venezuela monographed by Jung (1965) and Ward, 1976; Gibson, 1983). In the also has been dated as Burdigalian Caribbean, the Grand Bay Formation of (Hunter and Bartok, 1974, p . 147) and is the island of Carriacou has been shown by correlated with the Quiroz fauna of the La Robinson and Jung (1972, p. 124) also to be Rosa Formation ( = Agua Clara Forma of Middle Miocene age. tion; see Bolli, 1972). Most formations formerly placed in the The Pirabas Limestone of Brazil has Late Miocene are now Pliocene and in the been referred to ages ranging from Cre U.S. only the Red Bay Formation of the taceous (resulting in citations of the genus Choctawhatchee Group (Akers, 1972, p. Murex as dating back to the Cretaceous) to 13), the St. Marys Formation of southeast Middle Miocene . In a recent discussion of ern Virginia ( = Eastover Formation: Ward the problem (Vokes, 1989b, p. 28), I con and Blackwelder, 1980) and North cluded that these beds probably correlated Carolina (Gibson, 1983) remain in the Late with the Tampa Limestone and thus were Miocene. Early Miocene, possibly correlating with The Tubara Group of Colombia is no the Baitoa Formation of the Dominican Re longer considered to be Early to Late public. However, if the Tampa should Miocene but is probably Late Miocene to prove to be older, then it is possible that Early Pleistocene in age.