412 Scolopacidae: , , stints, snipe and

of a ringed at Lake Nakuru (0°S 36°E), Kenya, on 15 Marsh April 1972, probably on migration northwards when ringed. Moerasruiter Interspecific relationships: It occurs widely alongside the two other ‘inland’ congeneric , Greenshank and stagnatilis T. glareola as well as the Ruff Philomachus pugnax. The breeds mainly in marshes on the Eur- Historical distribution and conservation: There is no asian steppes, extending into the southern parts of the boreal evidence of a change in distribution in southern . How- zone, from the River Volga to Lake Baikal (Hayman et al. ever, the breeding distribution of the Marsh Sandpiper in 1986; Rogacheva 1992). Most migrate to Africa south of the has contracted eastwards owing to the intensification Sahara or to , with relatively few migrating to south- of agriculture and associated drainage. eastern Asia or to . The main concentration in south- ern Africa was in the section of the highveld plateau with L.G. Underhill many pans (26–28°S, 25–31°E). Other areas with high report- ing rates were the Okavango Delta, the lowveld of the east- ern Transvaal, eastern Swaziland and adjacent KwaZulu- Recorded in 1048 grid cells, 23.1% Natal, and the coastal region of the southern and southwestern Total number of records: 7178 Cape Province. Mean reporting rate for range: 7.6% Population sizes, both on the breeding and nonbreeding grounds, are poorly known (Rose & Scott 1994). Summers et al. (1987a) estimated 1000 in coastal southern Africa, but a larger number occurs at in the interior. The Marsh Sandpiper and Greenshank T. nebularia are Reporting rates for vegetation types easily confused, but the former has proportionately longer % 0816 legs, a slimmer and straighter bill, and a more slender neck and head, giving a more delicate ‘jizz’ (Hayman et al. 1986). Okavango 10.2 When flushed it gives either a solitary ‘tewk’ or a ‘tew-tew- Mixed Grasslands 7.7 tew’ call which is twice as fast as, and softer than, the similar Sweet Grasslands 7.0 ‘tjuu-tjuu-tjuu’ call of the Greenshank. East Coast Littoral 6.8 Habitat: Although mainly a species occurring at a wide Mopane 6.6 variety of freshwater wetlands, it also occurs at saltworks, Fynbos 5.9 coastal lagoons and tidal estuaries. It often feeds by wading Miombo 5.5 in shallow water over a muddy substrate, and less often on Arid Woodland 4.6 exposed mud. It rarely occurs on coastal shorelines; the high Sour Grasslands 3.8 reporting rates in some sections of the coast are of birds at Succulent Karoo 3.7 wetlands along the coastal plain. Northern Kalahari 3.6 Movements: The models show that departure starts earlier Grassy Karoo 3.4 (February–March) than for most species, that there Moist Woodland 3.4 were rather few overwintering records April–August, and that Valley Bushveld 3.0 arrival commences in September. This pattern is consistent Central Kalahari 2.8 with breeding in the Russian steppes beginning in April Nama Karoo 1.8 (Hayman et al. 1986). Namib 1.8 There have been no recoveries of the 592 individuals E Zimbabwe Highlands 1.8 ringed in southern Africa (SAFRING). However, there was Alpine Grasslands 0.3 one recovery at Rooiwal Pans (2727BC) on 13 January 1975, Namibian Escarpment 0.3 Scolopacidae: turnstones, sandpipers, stints, snipe and curlews 413

14û

MARSH SANDPIPER 5 1 18û

22û 2 6

26û

3 7 30û Reporting rate (%) > 11.9 5.9 — 11.9 2.0 — 5.8

< 2.0 34û 4 8 18û 22û 26û 14û 30û 10û 34û 40 1 5 20

40 2 6 20

40 3 7 20

40 Occurrence reporting rate (%) 4 8 20

J ASONDJ FMAMJ J ASONDJ FMAMJ Models of seasonality for Zones. Number of records (top to bottom, left to right): Occurrence: 108, 65, 83, 347, 545, 551, 1425, 89.