Towards a Consistent Oxfordian/Kimmeridgian Global Boundary: Current State of Knowledge
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Cryptoclidid Plesiosaurs (Sauropterygia, Plesiosauria) from the Upper Jurassic of the Atacama Desert
Journal of Vertebrate Paleontology ISSN: (Print) (Online) Journal homepage: https://www.tandfonline.com/loi/ujvp20 Cryptoclidid plesiosaurs (Sauropterygia, Plesiosauria) from the Upper Jurassic of the Atacama Desert Rodrigo A. Otero , Jhonatan Alarcón-Muñoz , Sergio Soto-Acuña , Jennyfer Rojas , Osvaldo Rojas & Héctor Ortíz To cite this article: Rodrigo A. Otero , Jhonatan Alarcón-Muñoz , Sergio Soto-Acuña , Jennyfer Rojas , Osvaldo Rojas & Héctor Ortíz (2020): Cryptoclidid plesiosaurs (Sauropterygia, Plesiosauria) from the Upper Jurassic of the Atacama Desert, Journal of Vertebrate Paleontology, DOI: 10.1080/02724634.2020.1764573 To link to this article: https://doi.org/10.1080/02724634.2020.1764573 View supplementary material Published online: 17 Jul 2020. Submit your article to this journal Article views: 153 View related articles View Crossmark data Full Terms & Conditions of access and use can be found at https://www.tandfonline.com/action/journalInformation?journalCode=ujvp20 Journal of Vertebrate Paleontology e1764573 (14 pages) © by the Society of Vertebrate Paleontology DOI: 10.1080/02724634.2020.1764573 ARTICLE CRYPTOCLIDID PLESIOSAURS (SAUROPTERYGIA, PLESIOSAURIA) FROM THE UPPER JURASSIC OF THE ATACAMA DESERT RODRIGO A. OTERO,*,1,2,3 JHONATAN ALARCÓN-MUÑOZ,1 SERGIO SOTO-ACUÑA,1 JENNYFER ROJAS,3 OSVALDO ROJAS,3 and HÉCTOR ORTÍZ4 1Red Paleontológica Universidad de Chile, Laboratorio de Ontogenia y Filogenia, Departamento de Biología, Facultad de Ciencias, Universidad de Chile, Las Palmeras 3425, Santiago, Chile, [email protected]; 2Consultora Paleosuchus Ltda., Huelén 165, Oficina C, Providencia, Santiago, Chile; 3Museo de Historia Natural y Cultural del Desierto de Atacama. Interior Parque El Loa s/n, Calama, Región de Antofagasta, Chile; 4Facultad de Ciencias Naturales y Oceanográficas, Universidad de Concepción, Barrio Universitario, Concepción, Región del Bío Bío, Chile ABSTRACT—This study presents the first plesiosaurs recovered from the Jurassic of the Atacama Desert that are informative at the genus level. -
Report of the Meeting of the Kimmeridgian Working Group
Volumina Jurassica, 2015, Xiii (2): 153–158 Report of the Meeting of the Kimmeridgian Working Group Andrzej WIERZBOWSKI1 Convenor of the Kimmeridgian W.G. The Kimmeridgian Working Group Meeting organized under auspices of the International Subcomission on Ju- rassic Stratigraphy took place in Poland between 18 and 21 May 2015 at the Polish Geological Institute – National Research Institute in Warsaw, with a one and a half day excursion to the Jurassic outcrops in the Wieluń Upland (Polish Jura). It was arranged to discuss new advances in recognition of a unique base of the Kimmeridgian Stage, the Oxfordian/Kimmeridgian boundary since 2006 when the first proposal for the GSSP was made (see Wierzbowski et al., 2006). Fifteen researchers were present, mainly members of the Kimmeridgian Working Group: C. D’Arpa (Geological Museum, University of Palermo), M. Barski (Geological Faculty, University of Warsaw), E. Głowniak (Geological Faculty, University of Warsaw), J. Grabowski (Polish Geological Institute – NRI), S. Hesselbo (University of Exeter, chairmen of ISJS), M. Hodbod (Polish Geological Institute – NRI), B. Matyja (Geological Faculty, University of War- saw), A. Mironenko (Russia), N. Morton (former ISJS chairman, France), M. Rogov (Geological Institute, Russian Academy of Science, Moscow), G. Schweigert (Staatliches Museum, Stuttgart), J. Smoleń (Polish Geological Institute – NRI), K. Sobień (Polish Geological Institute – NRI), A. Wierzbowski (Polish Geological Institute – NRI), H. Wierz bowski (Polish Geological Institute – NRI), J. Wright (Department of Earth Sciences, Royal Holloway, University of London). Twelve presentations were given (including two posters). These included several aspects of the Oxfordian/ Kimmeridgian boundary such as ammonite biostratigraphy, dinoflagellate cyst biostratigraphy, magnetostratigraphy, geochemistry (both carbon and oxygen records as well as other geochemical data along with geochemical studies which enable the recognition of the paleoenvironmental changes in the succession). -
First Three-Dimensionally Preserved in Situ Record of an Aptychophoran Ammonite Jaw Apparatus in the Jurassic and Discussion of the Function of Aptychi
Berliner paläobiologische Abhandlungen 10 321-330 Berlin 2009-11-11 First three-dimensionally preserved in situ record of an aptychophoran ammonite jaw apparatus in the Jurassic and discussion of the function of aptychi Günter Schweigert Abstract: A unique specimen of the microconch ammonite Lingulaticeras planulatum Berckhemer in Ziegler, 1958 comes from a tempestite bed within the Upper Jurassic lithographic limestones of Scham- haupten in Franconia (Painten Formation, uppermost Kimmeridgian). The shell is unique because it retains the complete jaw apparatus in the body chamber. The articulation of the Lamellaptychus and the corresponding upper beak are well preserved. The function of the aptychus is discussed in general, and an operculum function is thought to be unlikely. The formation of strongly calcified aptychi in aspidoceratids and some oppeliid ammonoids is interpreted as an added ballast weight to stabilize the conch for swimming in the water column. Keywords: Ammonites, aptychus, preservation, functional morphology, Upper Jurassic, lithographic lime- stones, Franconia, Germany Zusammenfassung: Ein einzigartig erhaltenes Exemplar des mikroconchen Ammoniten Lingulaticeras planulatum Berckhemer in Ziegler, 1958 aus einer Tempestitlage des oberjurassischen Plattenkalks von Schamhaupten in Franken (Painten-Formation, oberstes Kimmeridgium) enthält noch den vollständigen Kieferapparat in seiner Wohnkammer.Es zeigt die perfekte Artikualation des Lamellaptychus mit dem dazu- gehörenden Oberkiefer. Die Funktion des Aptychus wird allgemein diskutiert und eine Deckelfunktion für unwahrscheinlich gehalten. Die Ausbildung stark verkalkter Aptychen wie in Aspidoceraten und manchen Oppeliiden wird als zusätzliches Tariergewicht gedeutet, um das Gehäuse in starker bewegtem Wasser zu stabilisieren. Schlüsselwörter: Ammoniten, Aptychus, Erhaltung, Plattenkalke, Funktionsmorphologie, Oberjura, Franken, Deutschland Address of the author: Dr. Günter Schweigert, Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart. -
New Data on the Jaw Apparatus of Fossil Cephalopods
New dataon the jaw apparatus of fossil cephalopods YURI D. ZAKHAROV AND TAMAZ A. LOMINADZE \ Zakharov, Yuri D. & Lominadze, Tamaz A. 19830115: New data on the jaw apparatus of fossil LETHAIA cephalopods. Lethaia, Vol. 16, pp. 67-78. Oslo. ISSN 0024-1164. A newly discovered fossil cephalopod jaw apparatus that may belong to Permian representatives of the Endocochlia is described. Permorhynchus dentatus n. gen. n. sp. is established on the basis of this ~ apparatus. The asymmetry of jaws in the Ectocochlia may be connected with the double function of the ventral jaw apparatus, and the well-developed, relatively large frontal plate of the ventral jaw should be regarded as a feature common to all representatives of ectocochlian cephalopods evolved from early Palaeozoic stock. Distinct features seen in the jaw apparatus of Upper Pcrmian cndocochlians include the pronounced beak form of both jaws and the presence of oblong wings on the ventral mandible. o Cephalopoda. jaw. operculum. aptychus, anaptychus, Permorhynchus n.gen.• evolution. Permian. Yuri D. Zakharov llOpllll Ilscumpueeu« Gaxapoe], Institute of Biology and Pedology, Far-Eastern Scientific Centre. USSR Academy of Science, Vladivostok 690022, USSR (EUOJl020-n9~BnlHbliiuucmu my m Ilaot.neeocmo-cnoro Ha."~H020 uenmpav Axaoeuuu 'HayK CCCP, Bnaoueocmo« 690022, CCCP); Tamaz A. Lominadze ITa.'W3 Apl.j1L10BUl.j Jlouunaoee), Institute of Palaeobiology of Georgian SSR Academy of Science. Tbilisi 380004. USSR (Hncmumvm naJle06UOJl02UU Atcaoeuuu naytc TpY3UHUjKOii CCP. T6'LJUCU 380004. CCP; 19th August. 1980 (revised 1982 06 28). The jaw apparatus of Recent cephalopods is re Turek 1978, Fig. 7, but not the reconstruction in presented by two jaw elements (Fig. -
A Review of the Classification of Jurassic Aspidoceratid Ammonites – the Superfamily Aspidoceratoidea
VOLUMINA JURASSICA, 2020, XVIII (1): 47–52 DOI: 10.7306/VJ.18.4 A review of the classification of Jurassic aspidoceratid ammonites – the Superfamily Aspidoceratoidea Horacio PARENT1, Günter SCHWEIGERT2, Armin SCHERZINGER3 Key words: Superfamily Aspidoceratoidea, Aspidoceratidae, Epipeltoceratinae emended, Peltoceratidae, Gregoryceratinae nov. subfam. Abstract. The aspidoceratid ammonites have been traditionally included in the superfamily Perisphinctoidea. However, the basis of this is unclear for they bear unique combinations of characters unknown in typical perisphinctoids: (1) the distinct laevaptychus, (2) stout shells with high growth rate of the whorl section area, (3) prominent ornamentation with tubercles, spines and strong growth lines running in parallel over strong ribs, (4) lack of constrictions, (5) short to very short bodychamber, and (6) sexual dimorphism characterized by minia- turized microconchs and small-sized macroconchs besides the larger ones, including changes of sex during ontogeny in many cases. Considering the uniqueness of these characters we propose herein to raise the family Aspidoceratidae to the rank of a superfamily Aspi- doceratoidea, ranging from the earliest Late Callovian to the Early Berriasian Jacobi Zone. The new superfamily includes two families, Aspidoceratidae (Aspidoceratinae, Euaspidoceratinae, Epipeltoceratinae and Hybonoticeratinae), and Peltoceratidae (Peltoceratinae and Gregoryceratinae nov. subfam.). The highly differentiated features of the aspidoceratoids indicate that their life-histories -
301083243.Pdf
AMEGHINIANA ISSN 0002-7014 Revista de la Asociación Paleontológica Argentina Tomo XII Diciembre de 1975 N?4 THE INDO-PACIFIC AMMONITE MAY AITES IN THE OXFORDIAN OF THESOUTHERN ANDES By P. N. STIPANICIC', G. E. G. WESTERMANN 2 and A. C. RJCCARDI3 ABSTRACT: Oxfordian Iitho- and biostratigraphy of the Chilean and Argentine Andes is reviewed (P. N. Stipanicic). Within the Chacay Group, the Lower to basal Upper Oxfordian La Manga Formation, below, mostly detrital and biogenic, and the Upper Oxfordian Au- quilco Formation, above, mainly chemical, are distinguished. The La Manga Formation (with Gryphaea calceola lumachelle) is rich in ammonite faunas, particularly of thc upper Cordatum to lower Canaliculatum Zones. In Neuquén and Mendoza provinces of Argentina, the Pli- catilis Zone or Middle Oxfordian has yielded Perísphinctes spp., Euaspidoceras spp., Aspido- ceras spp., together with Mayaítes (Araucanites ) stípanícfcí, M. (A.) reyesi, and M. (A.) mulai, Westermann et Riccardi subgen. et spp. nov. The first find of Mayaitidae outside the Indo-Pacific province is discussed in light of _plate-tectonic theory. RESUMEN: La revisron Iito- y bioestratigráfica del Oxfordiano de los Andes de Argentina y Chile (P. N. Stipanicic) ha permitido reconocer dentro del Grupo Chacay: 1) abajo, la Formación La Manga, mayormente detrítica y biogénica, del Oxfordiano inferior-superior basa!, y 2) arriba, la Formación Auquilco, mayormente química, del Oxfordiano superior. La For- mación La Manga (con lumachelas de Gryphaea calceola) contiene abundante cantidad de amonitas, particularmente de las Zonas de Cordatum superior a Canaliculatum inferior. En las provincias de Mendoza y Neuquén,Argentina, la Zona de Plicatilis (Oxfordiano medio) contiene Perispbinctes spp., Euaspidoceras spp., Aspidoceras spp., conjuntamente con Mayaites (Araucanites) stipanicici, M. -
Schmitz, M. D. 2000. Appendix 2: Radioisotopic Ages Used In
Appendix 2 Radioisotopic ages used in GTS2020 M.D. SCHMITZ 1285 1286 Appendix 2 GTS GTS Sample Locality Lat-Long Lithostratigraphy Age 6 2s 6 2s Age Type 2020 2012 (Ma) analytical total ID ID Period Epoch Age Quaternary À not compiled Neogene À not compiled Pliocene Miocene Paleogene Oligocene Chattian Pg36 biotite-rich layer; PAC- Pieve d’Accinelli section, 43 35040.41vN, Scaglia Cinerea Fm, 42.3 m above base of 26.57 0.02 0.04 206Pb/238U B2 northeastern Apennines, Italy 12 29034.16vE section Rupelian Pg35 Pg20 biotite-rich layer; MCA- Monte Cagnero section (Chattian 43 38047.81vN, Scaglia Cinerea Fm, 145.8 m above base 31.41 0.03 0.04 206Pb/238U 145.8, equivalent to GSSP), northeastern Apennines, Italy 12 28003.83vE of section MCA/84-3 Pg34 biotite-rich layer; MCA- Monte Cagnero section (Chattian 43 38047.81vN, Scaglia Cinerea Fm, 142.8 m above base 31.72 0.02 0.04 206Pb/238U 142.8 GSSP), northeastern Apennines, Italy 12 28003.83vE of section Eocene Priabonian Pg33 Pg19 biotite-rich layer; MASS- Massignano (Oligocene GSSP), near 43.5328 N, Scaglia Cinerea Fm, 14.7 m above base of 34.50 0.04 0.05 206Pb/238U 14.7, equivalent to Ancona, northeastern Apennines, 13.6011 E section MAS/86-14.7 Italy Pg32 biotite-rich layer; MASS- Massignano (Oligocene GSSP), near 43.5328 N, Scaglia Cinerea Fm, 12.9 m above base of 34.68 0.04 0.06 206Pb/238U 12.9 Ancona, northeastern Apennines, 13.6011 E section Italy Pg31 Pg18 biotite-rich layer; MASS- Massignano (Oligocene GSSP), near 43.5328 N, Scaglia Cinerea Fm, 12.7 m above base of 34.72 0.02 0.04 206Pb/238U -
The Russian Platform As a Key Region for Volgian
Rivista Italiana di Prleontoloeia e Stratigrafia volume 1l0 no. l , pr IlI ;' ,t'.;r; I .a,prirzoo+ THE RUSSIAN PLATFORM AS A KEY REGION FOR VOLGIAN/TITHONIAN CORRELATION: A REVIE\tr OF THE MEDITERRANEAN FAUNAL ELEMENTS AND AMMONITE BIOSTRATIGRAPHY OF THE VOLGIAN STAGE MIKHAIL A. ROGOV' Receit-ted September 19, 2002; accepted October 3, 2003 Key uords. Jurassic, Volgian, Tithonian, Boreal-Teth1'an correla- Introduction tion, amnronites, biostratigraphy, Russian Platform. Over the last few decades several papers devot- Abstract. A review of the ammonite faunas and biostratieraphy ed to the analysis of mixed Boreal-Tethyan associations of the uppern.rost Kimmeridgian - lowermosr Middle Volgian of Central of Mollusca, and Tithonian-Volgian correlations (Zeiss Russia is presented. The major role of the Sub-n-rediterranean ammonites in the Volgian assemblages is described, providins additional detail to 2001Kutek & Zeiss 1997;Hoedemaeker 1991; Sachs et the existine scheme of Tithonian-Volgian correlations. The base of the a\.1979: Mesezhnikov 1989; Sey tr Kalacheva1993,1.997; Tithonian and the base of the Volgian are coincident, based on the asso- Schweigert 1.993, 1.994, 2000; Scherzinger Er Schweigert ciation of Neoc/retocerds steraspis-Lingulaticeras soLenoides.The Klimovi 1999) have been published. Unfortunately, material from Zone corresponds to nrost of the Hybonotum Zone. However, the top the Russian Platform was either not studied during those of the Hybonotum Zone, based on the distribution of Paralingulati- ceras, Íalls within the Sokolovi Zone.The allocation of r Tenuicostata investigations, or only ammonites never described and il- Subzone (with two horizons: neoburgense and puschi) in the Pseudo- lustrated before were mentioned. -
Restudy of a Phylloceratid Ammonite from Peikang, Taiwan
PETROLEUM GEOLOGY OF TAIWAN NO. 16, P. 51-57, 1 PL., DEC. 1979 Restudy of a Phylloceratid Ammonite from Peikang, Taiwan TATSURO MATSUMOTO Kyushu University, Fukuoka, Japan ABSTRACT This is a result of my restudy on a phylloceratid ammonite which was previously obtained as one of the samples from the Well PK-2 drilled in western Taiwan by the Chinese Petroleum Corporation and described by Lin (1961). The ammonite is closely allied to but not quite identical with Holcophylloceras caucasicum (Sayn), from the Aptian of the Caucasus. Taking all the available data into consideration, I propose to call this ammonite Holcophylloceras caucasicum taiwanum Lin et Huang. This concept of geographical sub species is compatible with the conclusions of age correlation by Matsumoto et al. (1965) andT. C. Huang (1978). INTRODUCTION The Cretaceous ammonites and other molluscan fossils were described by Matsumoto et al.. (1965) on some samples from the two wells near Peikang, PK-2 and PK-3, drilled into the coastal plain of western Taiwan by the Chinese Petroleum Corporation. On that occasion a phylloceratid ammonite described by Lin (1961) was not included in the samples on loan. Fortunately I was invited to visit the Paleontological Laboratory of the Chinese Petroleum Corporation at Miaoli in April 1979, when Dr. Weng-Rong Chi kindly showed and let me study that specimen. This paper is to report the result of my restudy. PALEONTOLOGICAL DESCRIPTION Subclass Ammonoidea Order Phylloceratida Family Phylloceratidae Zittel, 1894 Genus Holcophylloceras Spath, 1927 Type-species.-Phylloceras mediterraheum Neumayr, 1871, by original designation. Generic diagnosis. See Arkell et al, 1957, p. -
Palaeoecology and Palaeoenvironments of the Middle Jurassic to Lowermost Cretaceous Agardhfjellet Formation (Bathonian–Ryazanian), Spitsbergen, Svalbard
NORWEGIAN JOURNAL OF GEOLOGY Vol 99 Nr. 1 https://dx.doi.org/10.17850/njg99-1-02 Palaeoecology and palaeoenvironments of the Middle Jurassic to lowermost Cretaceous Agardhfjellet Formation (Bathonian–Ryazanian), Spitsbergen, Svalbard Maayke J. Koevoets1, Øyvind Hammer1 & Crispin T.S. Little2 1Natural History Museum, University of Oslo, P.O. Box 1172 Blindern, 0318 Oslo, Norway. 2School of Earth and Environment, University of Leeds, Leeds LS2 9JT, United Kingdom. E-mail corresponding author (Maayke J. Koevoets): [email protected] We describe the invertebrate assemblages in the Middle Jurassic to lowermost Cretaceous of the Agardhfjellet Formation present in the DH2 rock-core material of Central Spitsbergen (Svalbard). Previous studies of the Agardhfjellet Formation do not accurately reflect the distribution of invertebrates throughout the unit as they were limited to sampling discontinuous intervals at outcrop. The rock-core material shows the benthic bivalve fauna to reflect dysoxic, but not anoxic environments for the Oxfordian–Lower Kimmeridgian interval with sporadic monospecific assemblages of epifaunal bivalves, and more favourable conditions in the Volgian, with major increases in abundance and diversity of Hartwellia sp. assemblages. Overall, the new information from cores shows that abundance, diversity and stratigraphic continuity of the fossil record in the Upper Jurassic of Spitsbergen are considerably higher than indicated in outcrop studies. The inferred life positions and feeding habits of the benthic fauna refine our understanding of the depositional environments of the Agardhfjellet Formation. The pattern of occurrence of the bivalve genera is correlated with published studies of Arctic localities in East Greenland and northern Siberia and shows similarities in palaeoecology with the former but not the latter. -
Contributions in BIOLOGY and GEOLOGY
MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - --- -
Facies, Phosphate, and Fossil Preservation Potential Across a Lower Cambrian Carbonate Shelf, Arrowie Basin, South Australia
Palaeogeography, Palaeoclimatology, Palaeoecology 533 (2019) 109200 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Facies, phosphate, and fossil preservation potential across a Lower Cambrian T carbonate shelf, Arrowie Basin, South Australia ⁎ Sarah M. Jacqueta,b, , Marissa J. Bettsc,d, John Warren Huntleya, Glenn A. Brockb,d a Department of Geological Sciences, University of Missouri, Columbia, MO 65211, USA b Department of Biological Sciences, Macquarie University, Sydney, New South Wales 2109, Australia c Palaeoscience Research Centre, School of Environmental and Rural Science, University of New England, Armidale, New South Wales 2351, Australia d Early Life Institute and Department of Geology, State Key Laboratory for Continental Dynamics, Northwest University, Xi'an 710069, China ARTICLE INFO ABSTRACT Keywords: The efects of sedimentological, depositional and taphonomic processes on preservation potential of Cambrian Microfacies small shelly fossils (SSF) have important implications for their utility in biostratigraphy and high-resolution Calcareous correlation. To investigate the efects of these processes on fossil occurrence, detailed microfacies analysis, Organophosphatic biostratigraphic data, and multivariate analyses are integrated from an exemplar stratigraphic section Taphonomy intersecting a suite of lower Cambrian carbonate palaeoenvironments in the northern Flinders Ranges, South Biominerals Australia. The succession deepens upsection, across a low-gradient shallow-marine shelf. Six depositional Facies Hardgrounds Sequences are identifed ranging from protected (FS1) and open (FS2) shelf/lagoonal systems, high-energy inner ramp shoal complex (FS3), mid-shelf (FS4), mid- to outer-shelf (FS5) and outer-shelf (FS6) environments. Non-metric multi-dimensional scaling ordination and two-way cluster analysis reveal an underlying bathymetric gradient as the main control on the distribution of SSFs.