Opusc. Zool. Budapest, (2007) 2008, 38: 3–82.

An annotated checklist of the fauna of (: Collembola)

1 2 L. DÁNYI and GY. TRASER

Abstract. A checklist of the species of (Hexapoda: Collembola) hitherto recorded from Hungary is presented. Each entry is accompanied by complete references, and remarks where appropriate. The present list contains 414 species.

he Collembola fauna of several countries in critical review of the literature data of Collembola T the world was already overwied in the recent referring to Hungary. past (e.g. Babenko & Fjellberg 2006, Culik & Zeppelini Filho 2003, Skidmore 1995, Waltz & HISTORY Hart 1996, Zhao et al. 1997). The importance of such catalogues was stressed by several authors The first records of Collembola referring to (e.g. Csuzdi et al, 2006: 2) and their topicality is Hungary are some notes on the mass occurrence indicated also by the fact that several cheklists of certain species (Frenzel 1673, Mollerus 1673, referring even to European states were published Steltzner 1881), which however, are without any most recently (e.g. Fiera (2007) on Romania, taxonomical or faunistical value, as it has already Juceviča (2003) on Latvia, Kaprus et al. (2004) on been pointed out by Stach (1922, 1929). The next the Ukrain, Skarzynskiet al. (2002) on Poland). In springtail reference to Hungary is to be found in spite of these facts, the last comprehensive article the zoological book of János Földy (1801), which on the Hungarian springtail fauna was published was the first time the group was mentioned in about 80 years ago (Stach 1929), eventhough such Hungarian language in the scientific literature, critical reviews have a special importance in the eventhough this work doesn’t contain relevant case of this country because of the large changes faunistical records of the taxon. After a long spell of the political borders in the area during the last of time, it were Ottó Herman (1865) and Géza century (Csuzdi & Zicsi 2003, Mahunka & Papp Horváth (1888) who discussed the mass occur- 2005). Because of these changes, there are many rences of aquatica Linnaeus, 1785, and records in literature which were reported as the first serious works on this group in Hungary Hungarian, but in fact lie outside of the present were made by Ödön Tömösváry (1882, 1883, borders of Hungary. This means that between the 1884a, 1884b). Tömösváry described two species early „Hungarian” data we can find several, wich and one varietas new to the science, from which refer to localities at high elevations in the Sminthurus maculatus Tömösváry, 1883 is still Slovakian, Ukrainian or Romanian Carpathians, valid today. Tömösváry reported altogether 40 in the Croatian Dinarids, while some others refer Collembola taxa from historical Hungary, from to the Mediterranean seeshore of the Adria. These which however only 23 records refer to the pre- special habitat-types are lacking from present-day sent-day territory of the country. After Tömösvá- Hungary, so there are a lot of species not present ry’s tragically early death, O.M. Reuter reported in this smaller area. Now we aim to give a clear on some materials of the meticulous collector, base for further faunistical research presenting a Lajos Bíró (Reuter 1895a, 1895b). He described

______1László Dányi, Magyar Természettudományi Múzeum Állattára (Department of Zoology, Hungarian Natural History Mu- seum), Baross u. 13, H-1088 Budapest, Hungary. E-mail: [email protected] 2György Traser, Nyugat-magyarországi Egyetem, Erdőművelési és Erdővédelmi Intézet (University of Western Hungary, Institute of Forest and Wood Protection), Bajcsy-Zsilinszky u. 4, H-9400 , Hungary. E-mail: [email protected]

Dányi & Traser: Checklist of the springtail fauna of Hungary

one species ( pallidipes Reuter, 1895) new to a note written by Bíró, listing the species together the science from Hungary (Reuter 1895a), then with the collecting sites from which the collected reported on 14 species without indicating the pre- materials were identified by Reuter (fig. 1). The cise collecting localities, only writing „Ungarn” note contains some further records which were (Hungary). This is of no use for faunistical analy- missing at Vellay (1900) as well. According to sis as mentioned above, however, more precise Bíró’s draft and Vellay (1900), 13 of the 14 spe- infomation were given later by Imre Vellay in his cies mentioned by Reuter (1895b) from „Ungarn” checklist (Vellay 1900), which was the first com- were in fact collected in the present area of Hun- prehensive work on the Hungarian springtail fau- gary. There are 10 further taxa which were indica- na. It listed 39 species and 6 varietas from the pre- ted only generally as European in Reuter´s work, sent area of the country. Vellay wasn’t a Collem- eventhough Bíró’s note and Vellay name more bola expert, his catalogue is based on Tömösvá- precise localities, lying within present-day Hunga- ry’s data and on the identified material deposited ry. However, the greater part of Reuter’s material in the Hungarian Natural History Museum at that was collected outside of present-day Hungary, time (Vellay 1900). One further source was found thus, many of the data cited by Vellay (1900) in Lajos Bíró’s heritage, which confirms and refer indeed to Poland, the Ukraine, Romania, completes the localities published by Vellay. It is Slovakia, Slovenia or Croatia today .

Figures 1. The two sides of the note writen by Lajos Bíró on the materials collected by him and identified by Reuter.

After Vellay’s checklist was published there work on the Hungarian fauna was a checklist was a braek in Hungarian Collembola research, which contained records of 242 taxa, 141 of till the next period was started by the Pole Jan which refer to the present area of the country. Stach. Stach enriched our knowledge on Col- Even later, during his huge life-work, Stach re- lembola in the world to a great extent and his ported several data on the Hungarian Collembola works include the fauna of Hungary as well. He fauna (Stach 1930-1963). Stach identifies also the wrote a paper on the Apterygota fauna of „North- springtails collected mainly in caves in Hungary west Hungary” (Stach 1922), which, however, by Endre Dudich and Antal Gebhardt. These re- contains only records from north of the River sults were published in the papers of the collectors Danube, referring today not to Hungary, but to (Dudich 1929, 1930, 1932, 1947; Gebhardt 1933, contemporary Slovakia. He produced also some 1934) later. sporadic data of Hungarian springtails and des- cribed three species new to the science from Contemporary with Stach, Hungarian research- Hungary (Stach 1926a, 1926b, 1928). His next ers work on the group as well. József Szent-Ivány

4 Dányi & Traser: Checklist of the springtail fauna of Hungary

published some papers on Collembola faunistics some records from different parts of Hungary (Szent-Ivány 1938, 1940a, 1940b, 1941) discuss- (Paclt 1960), then Wolfgang Dunger described ing the fauna of the Kőszeg Mts. in detail. He two species new to the science from the country reports on 42 species from that region of the (Dunger 1974). One year later Dunger proposed a country (Szent-Ivány 1940a). In the early 40-ies, a work on the springtail fauna of the Börzsöny Mts. biospeleological communication on the Tapolca (Dunger 1975), which contains data of 144 spe- cave (Geyer & Mann 1940) and an anatomical cies, many of them being new to the fauna of paper on the guts of Collembola were published Hungary. (Tóth 1942), in which Szent-Ivány identified at least part of the specimens. After Loksa’s unexpected death, János Farkas worked on regions around Lake Balaton (Farkas The next period of Hungarian Collembola re- 1995, 1996, 1997; Farkas & Szathmáry 1997; Far- search is linked to the name of Imre Loksa. Loksa kas et al. 1998) and Traser’s investigations were investigated the fauna of several areas of the continued as well. He reported on the fauna of the country, such as the Bükk Mts. (Loksa 1956a, Sopron Mountains, the Őrség, the Szigetköz, the 1977), the Buda Mts., the environs of Barcs, the Aggtelek National Park, the Bükk Mts., the Hortobágy, the Kiskunság, the nature reserves of Gerecse Mts. (Traser 1994, 1995, 1997, 1999, Bátorliget and the Mts. (Loksa 1956b, 2002b, 2006, respectively), the Fertő-Hanság 1978a, 1983, 1987, 1991, Loksa & Rubio 1966, (Traser 2002a, 2003; Traser et al. 2006) and the respectively). We have to mention separately his Kiskunság (Sousa et al. 2006, Traser 2002c; huge ecological work on the fauna of the Panno- Traser & Csóka 2000, 2001; Traser & Horváth- nian karstic white-oak low woods (Cerraso-Quer- cetum, Cotino-Quercetum, Quercetum pubescen- Szováti 2004, 2006). Together with co-authors, ti) all around the country´s mountains (Loksa Traser described also seven species new to the 1966). Similar surveys are scarce in the world’s science from Hungary (Traser 2000; Traser & literature. Another focus of Loksa’s interest lied Christian 1992; Traser & Thibaud 1999; Traser et on subterranean species (Loksa 1959a, 1959b, al. 1993; Traser & Weiner 1999). Further species 1960, 1961a, 1961b, 1962, 1967, 1969a), resulting new to the science has been published from Hun- in descriptions of 12 species new to science from gary by Castaño-Meneses et al. (2000), Chen et Hungarian caves. Loksa’s identifications gave the al. (2002) and Christian & Thibaud (1988) during base of Dániel Bajomi’s paper on the fauna of the the last two decades. A further faunistical work Meteor cave (Bajomi 1969) too. was published by Kontschán et al. (2003) on the distribution of Tetrodontophora bielanensis (Wa- Beside Loksa some further researchers investi- ga, 1842) incorporating new data form Hungary gated the Hungarian Collembola fauna at this as well. time. István Kovács published three short commu- nications on Bátorliget and the Aggtelek Karst In addition to the faunistical and taxonomical (Kovács 1953a, 1953b, 1954), then Katalin Csu- works discussed above, we have to mention also ták wrote on feeding biology of some Collembola the great amount of ecological investigations on species (Csuták 1977) and Erzsébet Hornung re- springtails carried out in Hungary. An increasing ported on the fauna of a sandy grassland in the number of ecological papers have been published Kiskunság (Hornung 1986). In the 1980-ies during the last decades, some of which contain György Traser started his studies on springtails, faunistically relevant data as well. The research dealing in his early papers with the Collembola group of Gábor Bakonyi wrote several papers on fauna of the Bakony Mts. and with some ecologi- the role and importance of Collembola in agri- cal questions (Traser 1980, 1981, 1983, 1989). At cultural systems (e.g. Bakonyi 1989, 1998; Bako- this time not only Hungarian but also foreign nyi & Kiss 1986, 1995; Seres et al. 2003, 2006). experts were involved in the exploration of the János Kimmel and István Kiss investigated the springtails fauna. Thus, Jiří Paclt reported on effect of insecticides and pesticides on springtails

5 Dányi & Traser: Checklist of the springtail fauna of Hungary

(Kimmel & Traser 1999; Kiss & Bakonyi 1990, geographical units (as mountains, depressions, 1992; Kiss & Jager 1985, 1987). Miklós Dombos etc.) within Hungary (Plate X.). wrote on the effects of grazing on Collembola communities (Dombos 2001) and János Varga in- Species are listed according to the system of vestigated the bryofauna of the Bükk Mts. (Varga Deharveng (2007), marking the suborders, the 1991, 1992; Varga & Naár 2002; Varga et al. families, the subfamilies and the genera. 2000, 2002a, 2002b; Varga & Oldal 1998; Varga & Vargha 1992). For each species is reported: a) the latin name, the complete name of the author and year of pub- lication; b) the bibliographic quotations concer- MATERIAL AND METHODS ning the studied area listed according to the names

The present catalogue is based on critically re- of the species as originally quoted; c) taxonomical viewed literature records. Generic quotations, like remarks, where possible; d) the genaral geogra- „Hungary” (e.g. Schött 1893; Stach 1930, 1959) phic distribution according to Babenko & Fjell- are ignored in this catalogue, as the changes of the berg (2006), Bretfeld (1999), Christian (1987), political borders in the past make it impossible to Fjellberg (1998), Gisin (1960), Jordana et al. evaluate their real relevance. Citations of already (1997), Pomorski (1998), Potapow (2001), Zim- published data (e.g. Baumgartner-Gamauf 1959, dars & Dunger (1994); e) the list of the published Bretfeld 1999, Potapow 2001, Thibaud et al. occurrences in Hungary arranged according to 2004) or of type localities are not included. There geographical regions; are also some localities published by Stach, which Records are updated to 01.05.2008. were presented erroneously as Hungarian. „Besz- tercebánya” (Stach 1956) lies in Slovakia („Ban- LIST OF THE SPECIES ska Bystrica” today). „Komárom” is a city divided by the River Danube, lying half on Hungarian and Suborder: half on Slovakian territory. Stach (1929, 1956) cited his records from this city as Hungarian, Family: Brachystomellidae however he wrote earlier (Stach 1922: 4) that the material was collected only on the left (northern) Brachystomella Ågren, 1903 side of the river, which is actually Slovakian. Brachystomella curvula Gisin, 1948 There are also some further publications which Brachystomella curvula Gisin: Loksa & Rubio 1966a: could be listed here (e.g. Lakatos et al. 2003; 142; Hornung 1986: 138; Loksa 1987: 79; Christian & Loksa 1969b, Szemerei & Traser 1994), but as Thibaud 1988: 232; Traser 2002c: 48; Traser & they are of rather popular style, they were ignored Horváth–Szováti 2004: 59; Traser & Horváth-Szováti in the present checklist. 2006: 333.

General distribution: Central Europe, North As it was emphasized by Hopkin (1997: 44): America. „national boundaries may cut across areas with Distribution in Hungary: Bakony Mts. (Loksa similar climate or vegetation type. It is more & Rubio 1966a), Kiskunság (Hornung 1986; Lok- relevant from a biological point of view to divide sa 1987; Traser 2002c; Traser & Horváth–Szováti the world into biogeographical regions.” Al- 2004, 2006), Mezőföld (Christian & Thibaud though, according to Christiansen & Bellinger 1988). (1995) the whole area of Hungary belongs to the Ecological note: xerothermophilic. „European” province, in our opinion, patterns in distribution of several species can be recognised on a finer geographic scale as well, which can Brachystomella parvula (Schäffer, 1896) even be of biogegraphical relevance. Accordingly, Brachystomella parvula (Schäffer): Stach 1928: 364, we arranged the reported occurrences into smaller pl. 19.; Stach 1929: 281; Stach 1949b: 310, pl. 32;

6 Dányi & Traser: Checklist of the springtail fauna of Hungary

Loksa 1966: 379; Loksa 1977: 210; Loksa 1978b: 95; Mts. (Loksa 1966; Loksa & Rubio 1966a; Traser Traser 2002c: 48; Traser & Horváth-Szováti 2006: 1980), Balatonfelvidék (Tömösváry 1884a; Tö- 333; Traser et al. 2006: 72; Schöttella (sic!) parvula Schäffer: Loksa 1956a: 413. mösváry 1884b; Vellay 1900; Stach 1949b, 1929; Farkas & Szathmáry 1997), Börzsöny Mts. (Dun- General distribution: cosmopolitan. ger 1975), Buda Mts. (Loksa 1966), Bükk Mts. Distribution in Hungary: Balatonfelvidék (Loksa 1956a), Dráva Plain (Loksa 1978a), (Stach 1928, 1929, 1949b; Loksa 1966), Börzsöny Hortobágy (Loksa 1983), Kiskunság (Traser & Mts. (Loksa 1966), Buda Mts. (Stach 1928, 1929, Horváth-Szováti 2006; Loksa 1987), Kőszeg Mts. 1949b; Loksa 1966), Bükk Mts. (Loksa 1956a, (Traser & Salló 2004; Szent-Ivány 1940a), Me- Loksa 1977), Cserhát (Loksa 1978b), Fertő-Han- csek Mts. (Stach 1949b; Paclt 1960; Loksa 1966), ság (Traser et al. 2006), Keszthely Mts. (Loksa Mts. (Gebhardt 1934), Mezőföld (Stach 1966), Kiskunság (Traser 2002c; Traser & Hor- 1929, 1949b), Nyírség (Loksa 1991), Őrség (Tra- váth-Szováti 2006), Pilis Mts. (Loksa 1966), ser 1995), Sopron Mts. (Traser 2002a), Villány Visegrád Mts. (Loksa 1966). Mts. (Loksa 1966). Ecological note: frequent eurytopic species. Family: Hypogastruridae Ceratophysella bengtssoni (Ågren, 1904) Bonetogastrura Thibaud, 1974 Ceratophysella bengtssoni (Ågren): Stach 1949b: 142, Bonetogastrura cavicola (Börner 1901) pl. 15; Traser 1999: 50; Hypogastrura (Cerato- physella) bengtssoni (Ågren): Loksa 1962: 75, figs 12- Hypogastrura cavicola (Börner): Gebhardt 1933: 39; 16. Bajomi 1969: 242; Hypogastrura (Ceratophysella) ca- vicola (Börner): Loksa 1962: 75, figs 4-11; Neo- General distribution: Europe, North America. gastrura cavicola (Börner): Stach 1949b: 110, pl. 12. Distribution in Hungary: Aggtelek Karst (Tra- General distribution: Europe, Afghanistan. ser 1999), Balatonfelvidék (Stach 1949b; Loksa Distribution in Hungary: Aggtelek Karst (Ba- 1962), Budapest (Stach 1949b; Loksa 1962), jomi 1969), Bükk Mts. (Loksa 1962), Mecsek Bükk Mts. (Loksa 1962), Pilis Mts. (Stach 1949b; Mts. (Gebhardt 1933; Stach 1949b; Loksa 1962). Loksa 1962). Ecological note: occurs mostly in caves. Ecological note: guanophil.

Ceratophysella Börner, 1932 Ceratophysella bispinata Loksa, 1977 Ceratophysella armata (Nicolet, 1841) Hypogastrura (Ceratophysella) bispinata Loksa: Lok- sa 1977: 206; Hypogastrura bispinata Loksa: Loksa Achorutes armatus Nicolet: Tömösváry 1884a: 50; 1980: 110. Tömösváry 1884b: 231; Vellay 1900: 21; Cerato- physella armata (Nicolet): Stach 1949b: 126, pl. 12; Remarks: Species inquirenda according to Thi- Loksa 1978a: 53; Traser 1995: 44; Traser 1999: 50; baud et al. (2004). Maybe junior synonym of Ce- Traser & Horváth-Szováti 2006: 332; Traser & Salló ratophysella silvatica Rusek, 1964. 2004: 230; Ceratophysella armata (Uzel (sic!)): Traser 2002a: 261; Hypogastrura armata (Nicolet): Stach General distribution: Endemic? 1929: 279; Dudich 1932: 48; Gebhardt 1934: 138; Distribution in Hungary: Bükk Mts. (Loksa Szent-Ivány 1940a: 427; Loksa 1956a: 391; Loksa 1977), Mts. (Loksa 1980). 1966: 377; Loksa 1991: 270; Farkas & Szathmáry 1997: 39; Hypogastrura (Ceratophysella) armata (Nicolet): Paclt 1960: 156; Loksa & Rubio 1966a: 140; Ceratophysella cf. laricis Martinova, 1977 Dunger 1975: 15; Traser 1980: 20; Loksa 1983: 71; Loksa 1987: 78; Ceratophysella crassispina (Cass- Ceratophysella cf. laricis Martinova, 1977: Traser agnau): Traser 1999: 50. 1999: 51. General distribution: cosmopolitan. General distribution: East-Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Aggtelek Karst (Tra- ser 1999; Stach 1949b; Dudich 1932), Bakony ser 1999).

7 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ceratophysella denticulata (Bagnall, 1941) Hypogastrura luteospina Stach: Loksa 1966: 377; Loksa 1977: 210; Loksa 1978a: 53; Loksa 1980: 110; Ceratophysella denticulata (Bagnall): Traser 1997: 61; Farkas et al. 1998: 57. Traser 1999: 50; Traser 2002a: 261; Traser & Horváth- Szováti 2006: 332; Hypogastrura (Ceratophysella) General distribution: East and Central Europe. denticulata (Bagnall): Loksa & Rubio 1966a: 140; Distribution in Hungary: Bakony Mts. (Loksa Dunger 1975: 15; Traser 1980: 22; Loksa 1987: 78; & Rubio 1966a; Loksa 1966), Balatonfelvidék Hypogastrura denticulata (Bagnall): Loksa 1966: 377; Loksa 1977: 210; Loksa 1980: 110; Traser 1981: 133; (Loksa 1966), Börzsöny Mts. (Dunger 1975; Lok- Loksa 1991: 270; Varga & Vargha 1992: 315; Farkas sa 1966), Buda Mts. (Loksa 1966), Bükk Mts. 1995: 46; Farkas 1996: 287; Farkas 1997: 56; Farkas et (Loksa 1977), Dráva Plain (Loksa 1978a), Keszt- al. 1998: 57; Dombos 2001: 2041. hely Mts. (Loksa 1966), Mátra Mts. (Loksa 1980), General distribution: Palaearctic. Mecsek Mts. (Loksa 1966), Mt. Somló (Loksa Distribution in Hungary: Aggtelek Karst (Tra- 1966), Pilis Mts. (Loksa 1966), Somogy Hills ser 1999), Bakony Mts. (Loksa & Rubio 1966a; (Farkas et al. 1998), Vértes Mts. (Loksa 1966), Traser 1980), Balatonfelvidék (Farkas 1995, Villány Mts (Loksa 1966), Visegrád Mts. (Loksa 1997), Börzsöny Mts. (Dunger 1975), Bükk Mts. 1966). (Loksa 1966, 1977), Bükk Mts. (Varga & Vargha Ecological note: in mountains’ forest litter. 1992), Kis-Balaton (Farkas 1996, 1997), Kis- kunság (Traser & Horváth-Szováti 2006; Loksa Ceratophysella mosquensis (Becker, 1905) 1987), Mátra Mts. (Loksa 1980), Mezőföld (Dom- bos 2001), Nyírség (Loksa 1991), Somogy Hills Hypogastrura monstruosa Gisin: Loksa 1980: 107, figs (Farkas et al. 1998), Sopron Mts. (Traser 1981, 2a-c; Loksa 1991: 270. 2002a), Szigetköz (Traser 1997, 2002a), Vas-Sop- ron Plain (Traser 1981). General distribution: Central Europe, Siberia. Ecological note: detritophilous. Distribution in Hungary: Mátra Mts. (Loksa 1980), Nyírség (Loksa 1991). Ceratophysella gibbosa (Bagnall, 1940) Ecological note: tyrphophilous.

Hypogastrura (Ceratophysella) gibbosa (Bagnall): Loksa & Rubio 1966a: 140, figs 1-10; Hypogastrura Ceratophysella sigillata (Uzel, 1890) (Ceratophysella) occidentalis Gisin: Loksa & Rubio Hypogastrura sigillata Uzel: Dudich 1929: 42; Stach 1966a: 140. 1929: 279; Dudich 1932: 48; Ceratophysella sigillata (Uzel): Traser 1999: 51. General distribution: Holarctic. General distribution: Holarctic. Distribution in Hungary: Bakony Mts. (Loksa Distribution in Hungary: Aggtelek Karst (Du- & Rubio 1966a). dich 1932; Traser 1999), Balatonfelvidék (Stach Ecological note: in mountains’ forest litter. 1929), Budapest (Dudich 1929; Stach 1929), Pilis Mts. (Dudich 1929; Stach 1929). Ceratophysella granulata Stach, 1949 Ecological note: in mountains’ forest litter. Hypogastrura (Ceratophysella) granulata (Stach): Loksa 1978b: 94; Hypogastrura granulata Stach: Ceratophysella silvatica (Rusek, 1964) Loksa 1966: 377; Loksa 1991: 270; Ceratophysella cf. granulata Stach: Traser 1999: 50. Hypogastrura (Ceratophysella) silvatica Rusek: Dun- ger 1975: 15. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- General distribution: Europe, Carpathian Mts. ser 1999), Cserhát (Loksa 1978b), Nyírség (Loksa Distribution in Hungary: Börzsöny Mts. (Dun- 1991), Pilis Mts. (Loksa 1966). ger 1975). Ecological note: in mountains’ forest litter. Ecological note: in mountains’ forest litter.

Ceratophysella luteospina (Stach, 1920) Ceratophysella succinea (Gisin, 1949) Hypogastrura (Ceratophysella) luteospina (Stach): Ceratophysella succinea (Gisin): Traser 1999: 51; Loksa & Rubio 1966a: 140; Dunger 1975: 15; Hypogastrura cf. succinea Gisin: Traser 2000: 68;

8 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ceratophysella cf. succinea (Gisin): Traser 2002a: Hypogastrura assimilis (Krausbauer, 1898) 261. Hypogastrura manubrialis var. assimilis (Krausbauer): General distribution: Europe, North America. Stach 1929: 280; Hypogastrura (Hypogastrura) ass- Distribution in Hungary: Aggtelek Karst (Tra- imilis (Krausbauer): Dunger 1975: 14; Hypogastrura ser 1999), Fertő-Hanság (Traser 2000, 2002a). assimilis Krausbauer: Traser 1999: 50. Ecological note: troglophilous. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- Ceratophysella toemoesvaryi Loksa, 1964 ser 1999), Börzsöny Mts. (Dunger 1975), Pilis Hypogastrura (Ceratophysella) tömösvaryi (sic!) Lok- Mts. (Stach 1929). sa: Loksa 1964: 83, figs 1-8; Hypogastrura tömösvaryi Ecological note: detritophilous. (sic!) Loksa: Loksa 1966: 378. Remark: Not included in the key of Thibaud et Hypogastrura calceolaris Latzel, 1918 al. (2004: 40). Hypogastrura calceolaris Latzel: Loksa 1991: 270. General distribution: Hungary. Distribution in Hungary: Mecsek Mts. (Loksa Remark: Species inquirenda according to Gisin 1964, 1966), Villány Mts (Loksa 1964, 1966). (1960) and Thibaud et al. (2004). Ecological note: in xerotherm forests. General distribution: West Palaearctic? Distribution in Hungary: Nyírség (Loksa Choreutinula Paclt, 1944 1991).

Choreutinula inermis (Tullberg, 1871) Hypogastrura manubrialis (Tullberg, 1869) Beckerella inermis Tullbreg: Loksa 1956a: 413; Hypogastrura manubrialis (Tullberg): Stach 1929: Hypogastrura inermis Tullberg: Loksa 1966: 377; Tra- 280; Traser 1999: 50; Neogastrura manubrialis (Tull- ser 1999: 49; Hypogastrura (Schoettella) inermis berg): Stach 1949b: 50, pl. 1. (Tullberg): Dunger 1975: 15; Hypogastrura (Schoette- lla) inermis (Tullberg): Loksa 1978a: 53; Hypogast- General distribution: cosmopolitan. rura (Choreutinula) inermis Tullberg: Traser 1994: Distribution in Hungary: Aggtelek Karst (Tra- 102; Ceratophysella inermis (Tullberg): Traser 1999: 51; Schoettella inermis (Tullberg): Traser 2002a: 261; ser 1999), Balatonfelvidék (Stach 1929, 1949b), Choreutinula inermis (Tullberg): Traser & Horváth- Budapest (Stach 1929, 1949b), Mezőföld (Stach Szováti 2006: 332. 1929, 1949b), Pilis Mts. (Stach 1929, 1949b). General distribution: Palaearctic. Ecological note: "compost springtail".

Distribution in Hungary: Aggtelek Karst (Lok- Hypogastrura purpurescens (Lubbock, 1867)1 sa 1966; Traser 1999), Börzsöny Mts. (Dunger Hypogastrura purpurascens (sic!) (Lubbock): Stach 1975), Bükk Mts. (Loksa 1956a, 1966), Dráva 1929: 280; Dudich 1932: 48; Traser 1999: 50; Neo- Plain (Loksa 1978a), Gerecse Mts. (Loksa 1966), gastrura purpurescens (Lubbock): Stach 1949b: 101, Kiskunság (Traser & Horváth-Szováti 2006), Sop- pl. 11; Loksa 1962: 63; Loksa 1991: 270. ron Mts. (Traser 1994, 2002a). General distribution: cosmopolitan. Ecological note: hemiedaphic, in forest litter. Distribution in Hungary: Aggtelek Karst (Stach 1929; Dudich 1932; Traser 1999), Bakony Hypogastrura Bourlet, 1839 Mts. (Stach 1929, 1949b), Bükk Mts. (Loksa 1962), Nyírség (Loksa 1991). Hypogastrura aequepilosa (Stach, 1949) Ecological note: hemiedaphic, troglophilous. Hypogastrura aequipilosa (Stach): Traser 1999: 50.

General distribution: Central Europe. 1 Hypogastrura pseudohispanica mentioned by Loksa (1966 Distribution in Hungary: Aggtelek Karst (Tra- p. 59) from the Villány Mts. and the Mecsek Mts. As Loksa ser 1999). didn’t provide any description or illustration, we have to Ecological note: in mountains’ forest litter. consider it as nomen nudum.

9 Dányi & Traser: Checklist of the springtail fauna of Hungary

Hypogastrura ripperi Gisin, 1952 2006: 72; Hypogastrura (Hypogastrura) vernalis (Carl): Loksa 1983: 71; Loksa 1987: 78; Neogastrura Hypogastrura ripperi Gisin: Traser 1999: 50. vernalis (Carl): Stach 1949b: 66, pl. 3 figs 1-11.

General distribution: Europe, North America. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Balatonfelvidék ser 1999). (Loksa 1960), Budapest (Stach 1929, 1949b), Ecological note: "compost springtail". Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser 2002a; Traser et al. 2006), Hortobágy (Loksa

1983), Kiskunság (Loksa 1987; Traser 2002c; Hypogastrura sahlbergi (Reuter, 1895) Traser & Horváth-Szováti 2006), Mezőföld (Stach Hypogastrura sahlbergi (Reuter): Traser 1999: 50. 1949b), Nyírség (Loksa 1991), Somogy Hills (Farkas et al. 1998). General distribution: Europe, North America. Ecological note: mostly in meadows. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999). Hypogastrura viatica (Tullberg, 1872) Ecological note: mostly hygrophilous. Hypogastrura viatica (Tullberg): Dudich 1929: 42; Stach 1929: 279; Loksa 1956b: 211; Neogastrura viatica (Tullberg): Stach 1949b: 89, pls. 3, 9. Hypogastrura socialis (Uzel, 1890) General distribution: cosmopolitan. Hypogastrura socialis (Uzel): Szent-Ivány 1940a: 427; Distribution in Hungary: Buda Mts. (Dudich Traser 1999: 50; Traser 2002a: 261; Traser et al. 2006: 72. 1929; Stach 1929, 1949b; Loksa 1956b), Me- zőföld (Stach 1949b). General distribution: Holarctic. Ecological note: mostly on coastal regions.

Distribution in Hungary: Aggtelek Karst (Tra- Mesogastrura Bonet, 1930 ser 1999), Fertő-Hanság (Traser et al. 2006), Kőszeg Mts. (Szent-Ivány 1940a), Sopron Mts. Mesogastrura anthrohungarica Loksa, 1959 (Traser 2002a). Mesogastrura anthrohungarica Loksa, 1959b: 78, figs Ecological note: mostly gregarious in early 8-15. spring. General distribution: Hungary. Hypogastrura tullbergi (Schäffer, 1900) Distribution in Hungary: Cserhát (Loksa 1959b). ?Hypogastrura (Hypogastrura) tullbergi (Schäffer): Dunger 1975: 14. Mesogastrura libyca (Caroli, 1914)

Remarks: According to Fjellberg (1998: 26) it Mesachorutes lybicus Caroli: Loksa 1966: 378. is „a true Arctic species” and „records of “tull- bergi” from Continental Europe refer to other General distribution: Europe, Africa, Ecuador. species”. Distribution in Hungary: Mecsek Mts. (Loksa General distribution: Holarctic, circumpolar. 1966). Distribution in Hungary: Börzsöny Mts. (Dun- Ecological note: coprophilous, guanophilous. ger 1975). Mesogastrura tiliophila (Loksa & Rubio, 1966) Hypogastrura vernalis (Carl, 1901) Mesachorutes tiliophilus Loksa & Rubio: Loksa & Hypogastrura vernalis (Carl): Stach 1929: 280; Loksa Rubio 1966a: 140, figs 11-16. 1960: 48; Loksa 1978a: 53; Loksa 1991: 270; Farkas et al. 1998: 57; Traser 2002a: 261; Traser 2002c: 47; Remark: Species dubia according to Thibaud Traser & Horváth-Szováti 2006: 332; Traser et al. et al. (2004: 128).

10 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Hungary. 1978b: 94; Traser 1999: 51; Traser & Csóka 2000: 76; Distribution in Hungary: Bakony Mts. (Loksa Traser & Csóka 2001: 233; Traser 2002c: 47; Traser & Horváth-Szováti 2006: 332. & Rubio 1966a). General distribution: cosmopolitan. Microgastrura Stach, 1922 Distribution in Hungary: Aggtelek Karst (Lok- sa 1966; Traser 1999), Bakony Mts. (Loksa Microgastrura duodecimoculata Stach, 1922 1966), Balatonfelvidék (Loksa 1966), Börzsöny Microgastrura duodecimoculata Stach: Loksa 1966: Mts. (Dunger 1975), Buda Mts. (Loksa 1966), 379; Dunger 1975: 16; Loksa 1978a: 55; Loksa 1978b: 95; Traser 1999: 51. Bükk Mts. (Loksa 1962, 1966, 1977), Cserhát (Loksa 1978b), Gerecse Mts. (Loksa 1966), General distribution: Europe, Caucasus. Keszthely Mts. (Loksa 1966), Kiskunság (Traser Distribution in Hungary: Aggtelek Karst (Lok- & Csóka 2000, 2001; Traser 2002c; Traser & sa 1966; Traser 1999), Börzsöny Mts. (Loksa Horváth-Szováti 2006), Mecsek Mts. (Loksa 1966; Dunger 1975), Buda Mts. (Loksa 1966), 1966), Mt. Somló (Loksa 1966), Pilis Mts. (Loksa Bükk Mts. (Loksa 1966), Cserhát (Loksa 1978b), 1966), Vértes Mts. (Loksa 1966), Villány Mts Dráva Plain (Loksa 1978a), Keszthely Mts. (Lok- (Loksa 1966). sa 1966), Mecsek Mts. (Loksa 1966), Mt. Somló Ecological note: euedaphic. (Loksa 1966), Pilis Mts. (Loksa 1966), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), Visegrád Mts. (Loksa 1966). Willemia denisi Mills, 1932 Willemia denisi Mills: Traser 2002c: 47; Traser & Schoettella Schäffer, 1896 Horváth-Szováti 2006: 332.

Schoettella ununguiculata (Tullberg, 1869) General distribution: Nearctic and Palaearctic Schöttella (sic!) ununguiculata (Tullberg): Stach 1929: to the Far East. 280; Schoettella ununguiculata (Tulberg): Stach Distribution in Hungary: Kiskunság (Traser 1949b: 181, pl. 19; Traser 2002a: 261; Traser 2002c: 47; Hypogastrura unguiculata (sic!) Tullberg: Loksa 2002c; Traser & Horváth-Szováti 2006). 1966: 378; Hypogastrura ununguiculata Tullberg: Lo- Ecological note: in soil and litter. ksa 1991: 270; Hypogastrura (Schoettella) ununguicu- lata (Tullberg): Dunger 1975: 15; Loksa 1978a: 53; Loksa 1978b: 94; Loksa 1983: 71; Loksa 1987: 78. Willemia intermedia Mills, 1934 Willemia intermedia Mills: Traser 1997: 59; Traser General distribution: Holarctic. 2002a: 261; Traser 2002c: 47; Traser & Horváth- Distribution in Hungary: Balatonfelvidék Szováti 2006: 332. (Loksa 1966), Börzsöny Mts. (Dunger 1975), Buda Mts. (Stach 1929, 1949b; Loksa 1966), General distribution: Holarctic. Cserhát (Loksa 1978b), Dráva Plain (Loksa Distribution in Hungary: Kiskunság (Traser 1978a), Hortobágy (Loksa 1983), Keszthely Mts. 2002c; Traser & Horváth-Szováti 2006), Sziget- (Loksa 1966), Kiskunság (Loksa 1987; Traser köz (Traser 1997, 2002a). 2002c), Mezőföld (Stach 1929, 1949b), Nyírség Ecological note: euedaphic, interstitial. (Loksa 1991), Sopron Mts. (Traser 2002a), Vértes Mts. (Loksa 1966). Willemia scandinavica Stach, 1949 Ecological note: hemiedaphic in forests and meadows. Willemia scandinavica Stach: Dunger 1975: 16; Traser 1999: 51; Traser 2002a: 261; Traser 2002c: 47; Traser Willemia Börner, 1901 & Horváth-Szováti 2006: 332.

Willemia anophthalma Börner, 1901 General distribution: Europe, North America. Willemia anophthalma Börner: Loksa 1962: 63; Loksa Distribution in Hungary: Aggtelek Karst (Tra- 1966: 378; Dunger 1975: 16; Loksa 1977: 210; Loksa ser 1999), Börzsöny Mts. (Dunger 1975), Kis-

11 Dányi & Traser: Checklist of the springtail fauna of Hungary

kunság (Traser 2002c; Traser & Horváth-Szováti 1966), Villány Mts (Loksa 1966), Visegrád Mts. 2006), Moson Plain (Traser 2002a). (Loksa 1966). Ecological note: euedaphic, troglophilous. Ecological note: in litter.

Willemia virae Kaprus, 1997 Xenylla brevisimilis Stach, 1949 Willemia virae Kaprus: Traser 2006: 35; Traser & Hor- Xenylla brevisimilis Stach: Dunger 1975: 15; Loksa váth-Szováti 2006: 332; Traser et al. 2006: 72. 1977: 210; Traser 1999: 51; Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; Traser 2002c: 47. General distribution: East and Central Europe. Distribution in Hungary: Fertő-Hanság (Traser General distribution: Europe, North Africa, et al. 2006), Gerecse Mts. (Traser 2006), Kiskun- Canarian Island. ság (Traser & Horváth-Szováti 2006). Distribution in Hungary: Aggtelek Karst (Tra- Ecological note: euedaphic, troglophilous. ser 1999), Börzsöny Mts. (Dunger 1975), Bükk Mts. (Loksa 1977), Kiskunság (Traser & Csóka

2000, 2001; Traser 2002c, 2002c). Xenylla Tullberg, 1869 Ecological note: in litter and moss.

Xenylla boerneri Axelson, 1905 Xenylla corticalis Börner, 1901 Xenylla boerneri Axelson: Loksa & Rubio 1966a: 140; Dunger 1975: 15; Loksa 1978a: 55; Loksa 1983: 71; Xenylla corticalis Börner: Loksa 1978a: 55, figs 1-4; Loksa 1991: 270; Varga & Oldal 1998: 86; Traser Loksa 1983: 72; Traser 1995: 44; Traser 1997: 61; 1999: 51; Traser 2002b: 159; Traser et al. 2006: 72. Traser 1999: 51; Traser 2002a: 261; Traser 2002c: 47; Traser & Horváth-Szováti 2006: 332.

General distribution: Europe. General distribution: Europe Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Bakony Mts. (Loksa & Rubio 1966a; ser 1999), Dráva Plain (Loksa 1978a), Fertő-Han- Traser 2002b), Börzsöny Mts. (Dunger 1975; Tra- ság (Traser 2002a), Hortobágy (Loksa 1983), ser 2002b), Bükk Mts. (Traser 2002b), Bükk Mts. Kiskunság (Traser 2002c; Traser & Horváth-Szo- (Varga & Oldal 1998), Dráva Plain (Loksa 1978a; váti 2006), Őrség (Traser 1995), Sopron Mts. Traser 2002b), Fertő-Hanság (Traser et al. 2006), (Traser 2002a), Szigetköz (Traser 1997). Hortobágy (Loksa 1983; Traser 2002b), Nyírség Ecological note: corticophilous. (Loksa 1991; Traser 2002b). Ecological note: corticophilous, in moss. Xenylla grisea Axelson, 1900

Xenylla brevicauda Tullberg, 1869 Xenylla grisea Axelson: Loksa 1966: 378; Loksa & Rubio 1966a: 140. Xenylla brevicauda Tullberg: Vellay 1900: 21; Stach 1929: 281; Loksa 1966: 378; Traser 1994: 100; Traser 1995: 44; Traser 1999: 51; Traser 2002a: 261; Traser et General distribution: cosmopolitan. al. 2006: 72. Distribution in Hungary: Aggtelek Karst (Lok- sa 1966), Bakony Mts. (Loksa & Rubio 1966a), General distribution: Europe. Bükk Mts. (Loksa 1966). Distribution in Hungary: Aggtelek Karst Ecological note: in litter and on bark. (Loksa 1966; Traser 1999), Bakony Mts. (Loksa 1966), Balatonfelvidék (Loksa 1966), Börzsöny Xenylla maritima Tullberg, 1869 Mts. (Loksa 1966), Buda Mts. (Loksa 1966), Xenylla maritima Tullberg: Stach 1929: 281; Stach Bükk Mts. (Loksa 1966), Fertő-Hanság (Traser et 1949b: 227, pl. 23; Loksa 1966: 378; Dunger 1975: 15; al. 2006), Gerecse Mts. (Loksa 1966), Keszthely Loksa 1977: 210; Loksa 1978a: 55; Loksa 1983: 72; Mts. (Loksa 1966), Kiskunság (Vellay 1900; Hornung 1986: 138; Loksa 1987: 79; Loksa 1991: 270; Stach 1929), Mecsek Mts. (Loksa 1966), Őrség Varga 1991: 85; Varga 1992: 172; Varga & Vargha 1992: 315; Varga & Oldal 1998: 86; Traser 1999: 51; (Traser 1995), Pilis Mts. (Loksa 1966), Sopron Traser 2002a: 261; Traser 2002c: 47; Traser & Mts. (Traser 1994, 2002a), Vértes Mts. (Loksa Horváth-Szováti 2006: 332; Traser et al. 2006: 72;

12 Dányi & Traser: Checklist of the springtail fauna of Hungary

Xenilla (sic!) maritima Tullberg: Farkas et al. 1998: General distribution: Central Europe, Mediter- 57. ranean. General distribution: Palaearctic. Distribution in Hungary: Kiskunság (Traser Distribution in Hungary: Aggtelek Karst (Lok- 2002c; Traser & Horváth-Szováti 2006). sa 1966; Traser 1999), Bakony Mts. (Loksa Ecological note: hemiedaphic, psammophil- 1966), Balatonfelvidék (Loksa 1966), Börzsöny ous. Mts. (Loksa 1966; Dunger 1975), Buda Mts. (Loksa 1966), Bükk Mts. (Loksa 1966, 1977; Family: Varga 1991, 1992; Varga & Vargha 1992; Varga & Oldal 1998), Dráva Plain (Loksa 1978a), Fertő- Subfamily: Frieseinae Hanság (Traser 2002a; Traser et al. 2006), Gere- cse Mts. (Loksa 1966), Hortobágy (Loksa 1983), Friesea Dalla Torre, 1895 Keszthely Mts. (Loksa 1966), Kiskunság (Hor- nung 1986; Loksa 1987; Traser 2002c; Traser & Friesea afurcata (Denis, 1926) Horváth-Szováti 2006), Mecsek Mts. (Loksa 1966), Mezőföld (Stach 1929, 1949b), Mt. Somló Friesea afurcata (Denis): Loksa 1983: 72; Traser & (Loksa 1966), Nyírség (Loksa 1991), Pilis Mts. Horváth-Szováti 2006: 333. (Loksa 1966), Somogy Hills (Farkas et al. 1998), Szigetköz (Traser 2002a), Vértes Mts. (Loksa General distribution: Europe, Lebanon. 1966), Villány Mts (Loksa 1966), Visegrád Mts. Distribution in Hungary: Hortobágy (Loksa (Loksa 1966). 1983), Kiskunság (Traser & Horváth-Szováti Ecological note: xerophilous. 2006). Ecological note: hygrophilous. Xenylla tullbergi Börner, 1903 Friesea albida Stach, 1949 Xenylla biroi Stach: Stach 1926a: 81; Stach 1929: 281; Stach 1949b: 211; Loksa 1966: 378; Xenylla tullbergi Friesea decipiens Steiner: Dunger 1975: 16. Börner: Loksa & Rubio 1966a: 140; Loksa 1978a: 55; Traser 2002a: 261. General distribution: Central Europe, Pyrene- an Mts. General distribution: Europe. Distribution in Hungary: Börzsöny Mts. (Dun- Distribution in Hungary: Bakony Mts. (Loksa ger 1975). 1966; Loksa & Rubio 1966a), Börzsöny Mts. (Loksa 1966), Buda Mts. (Stach 1926a, 1929), Friesea claviseta Axelson, 1900 Budapest (Stach 1949b), Dráva Plain (Loksa 1978a), Szigetköz (Traser 2002a). Friesea claviseta Axelson: Stach 1929: 281; Dunger Ecological note: colline, montane. 1975: 16; Loksa 1978a: 55; Traser 2002a: 261; Traser 2006: 35; Friesea cf. claviseta Axelson: Traser 1997: 61. Xenylla welchi Folsom, 1916 General distribution: Holarctic. Xenylla welchi Folsom: Traser 1999: 51. Distribution in Hungary: Balatonfelvidék (Stach 1929), Börzsöny Mts. (Dunger 1975), Drá- General distribution: cosmopolitan. va Plain (Loksa 1978a), Gerecse Mts. (Traser Distribution in Hungary: Aggtelek Karst (Tra- 2006), Szigetköz (Traser 1997, 2002a). ser 1999). Ecological note: corticophilous. Ecological note: detritophilous, gregarious. Friesea denisi Kseneman, 1936 Xenyllogastrura Denis, 1932 Friesea denisi Kseneman: Loksa 1956a: 391; Traser Xenyllogastrura octoculata (Steiner, 1955) 1999: 51. Xenyllogastrura octoculata (Steiner): Traser 2002c: General distribution: Europe, Alpine-Carpa- 47; Traser & Horváth-Szováti 2006: 333. thian.

13 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Aggtelek Karst (Tra- Ecological note: probably predatory species. ser 1999), Bükk Mts. (Loksa 1956a). Ecological note: in mountainous habitats. Friesea stachi Kseneman, 1936

Friesea geminioculata Loksa, 1964 Friesea stachi Kseneman: Varga 1992: 172; Varga & Oldal 1998: 86. Friesea (Friesea) geminioculata Loksa: Loksa 1964: 85, figs 9-15; Friesea geminioculata Loksa: Loksa General distribution: Eastern Europe, Carpa- 1966: 379. thian Mts.

General distribution: Hungary. Distribution in Hungary: Bükk Mts. (Varga Distribution in Hungary: Bükk Mts. (Loksa 1992; Varga & Oldal 1998). 1964, 1966). Friesea truncata Cassagnau, 1958 Friesea laszloi Palacios-Vargas & Traser, 1996 Friesea truncata Cassagnau: Traser 1999: 52; Traser Friesea laszloi Palacios-Vargas & Traser: Traser 1999: 2002a: 262; Traser 2002c: 48; Traser 2003: 155; Traser 52. 2006: 35; Traser & Horváth-Szováti 2006: 333; Traser et al. 2006: 72. General distribution: Hungary and the Cauca- sus Mts. General distribution: Europe. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Aggtelek Karst (Tra- ser 1999). ser 1999), Fertő-Hanság (Traser 2002a, 2003; Ecological note: xerothermophilic. Traser et al. 2006), Gerecse Mts. (Traser 2006), Kiskunság (Traser 2002c; Traser & Horváth-Szo- Friesea mirabilis (Tullberg, 1871) váti 2006), Szigetköz (Traser 2002a). Friesea mirabilis (Tullberg): Stach 1929: 281; Stach 1949b: 287, pl. 28; Loksa 1960: 49; Paclt 1960: 156; Subfamily: Morulininae Loksa 1966: 378; Loksa & Rubio 1966a: 140; Dunger 1975: 16; Loksa 1978a: 55; Loksa 1978b: 94; Loksa Morulina Börner, 1906 1980: 110; Traser 1980: 20, fig. 2b; Traser 1981: 135; Loksa 1983: 72; Loksa 1987: 79; Loksa 1991: 270; Farkas 1995: 46; Farkas 1996: 287; Farkas 1997: 58; Morulina verrucosa (Börner, 1903) Farkas & Szathmáry 1997: 39; Traser 1997: 61; Farkas Morulina verrucosa (Börner): Stach 1951: 17, pl. 1: et al. 1998: 57; Traser 2002a: 262; Traser & Salló fig. 1, pl. 2, pl.3: figs 1-4. 2004: 230.

General distribution: Holarctic. General distribution: Central Europe. Distribution in Hungary: around the Lake Distribution in Hungary: Sokoróalja (Stach Balaton (Farkas 1995, 1997), Bakony Mts. (Loksa 1951). & Rubio 1966a; Traser 1980), Balatonfelvidék (Stach 1929, 1949b; Loksa 1960; Paclt 1960; Far- kas 1995, 1997; Farkas & Szathmáry 1997), Bör- Subfamily: zsöny Mts. (Dunger 1975), Buda Mts. (Loksa Albanura Deharveng, 1982 1966), Cserhát (Loksa 1978b), Dráva Plain (Lok- sa 1978a), Fertő-Hanság (Traser 2002a), Gerecse Mts. (Loksa 1966), Hortobágy (Loksa 1983), Albanura dudichi (Loksa, 1967) Keszthely Mts. (Loksa 1966), Kis-Balaton (Far- Neanura dudichi Loksa, 1967: 289, figs 1-3. kas 1996, 1997), Kiskunság (Loksa 1987), Kő- cf Albanura dudichi (Loksa): Traser 1999: 52. szeg Mts. (Traser & Salló 2004), Mátra Mts. (Loksa 1980), Mecsek Mts. (Loksa 1966), Nyír- General distribution: Carpathian. ség (Loksa 1991), Somogy Hills (Farkas et al. Distribution in Hungary: Aggtelek Karst (Lok- 1998), Szigetköz (Traser 1997), Vas-Sopron Plain (Traser 1981), Vértes Mts. (Loksa 1966), Villány sa 1967; Traser 1999). Mts (Loksa 1966). Ecological note: silvicolous, troglophilous.

14 Dányi & Traser: Checklist of the springtail fauna of Hungary

Bilobella Caroli, 1912 General distribution: Central Europe. Distribution in Hungary: Kőszeg Mts. (Traser Bilobella aurantiaca (Caroli, 1910) & Salló 2004), Őrség (Traser 1995), Sopron Mts. Achorutes aurantiacus Caroli: Stach 1929: 283; Stach (Traser et al. 1993; Traser 1994, 2002a). 1930: 58, fig. 14/4; Bilobella aurantiaca (Caroli): Ecological note: silvicolous. Stach 1951: 88, pl. 16: fig. 3; Neanura (Lathriopyga) aurantiaca Caroli: Loksa & Rubio 1966a: 142; Ne- anura aurantiaca Caroli: Loksa 1978a: 55; Bilobella Deutonura conjuncta (Stach, 1926) aurantiaca (Caroli): Traser 1995: 44; Traser 2002a: 262; Traser & Salló 2004: 230. Achorutes conjunctus Stach: Stach 1929: 283; La- thriopyga conjuncta (Stach): Stach 1951: 73, pl. 1: fig. General distribution: Central and Southern Eu- 2, pl. 13; Lathiropyga (sic!) conjuncta (Stach): Dunger rope. 1975: 18; Deutonura conjuncta (Stach): Traser 1995: 44; Traser 1997: 61; Traser 1999: 52; Traser 2002a: Distribution in Hungary: Bakony Mts. (Stach 262; Traser 2003: 155; Traser 2006: 35; Traser & 1929, 1930, 1951; Loksa & Rubio 1966a), Dráva Horváth-Szováti 2006: 333; Neanura conjuncta Stach: Plain (Loksa 1978a), Kőszeg Mts. (Traser & Salló Loksa 1966: 379; Loksa 1978b: 95; Traser 1981: 133; 2004), Őrség (Traser 1995), Sopron Mts. (Traser Loksa 1983: 72; Farkas 1995: 46; Farkas 1997: 56; Farkas et al. 1998: 57; Neanura (Lathriopyga) con- 2002a). juncta (Stach): Loksa & Rubio 1966a: 142; Traser 1980: 20. Bilobella braunerae Deharveng, 1981 General distribution: Central Europe. Bilobella braunerae Deharveng: Traser 2002a: 262. Distribution in Hungary: Aggtelek Karst (Tra- General distribution:Central Europe. ser 1999; Loksa 1966), around the Lake Balaton Distribution in Hungary: Fertő-Hanság (Traser (Farkas 1995, 1997), Bakony Mts. (Loksa 1966; 2002a), Sopron Mts. (Traser 2002a). Loksa & Rubio 1966a; Traser 1980), Bala- Ecological note: silvicolous. tonfelvidék (Farkas 1995, 1997), Börzsöny Mts. (Dunger 1975; Loksa 1966), Buda Mts. (Stach Bilobella excolorata (Loksa & Rubio, 1966) 1929, 1951; Loksa 1966), Cserhát (Stach 1929, Neanura excolorata Loksa & Rubio: Loksa & Rubio 1951; Loksa 1978b), Fertő-Hanság (Traser 2002a, 1966a: 142, figs 17-20. 2003), Gerecse Mts. (Traser 2006), Hortobágy (Loksa 1983), Kiskunság (Traser & Horváth- General distribution: Hungary. Szováti 2006), Mezőföld (Stach 1951), Őrség Distribution in Hungary: Bakony Mts. (Loksa (Traser 1995), Pilis Mts. (Loksa 1966), Somogy & Rubio 1966a). Hills (Farkas et al. 1998), Sopron Mts. (Traser 2002a, 1981), Szigetköz (Traser 1997, 2002a), Deutonura Cassagnau, 1979 Vértes Mts. (Loksa 1966), Visegrád Mts. (Loksa 1966). Deutonura albella (Stach, 1920) Deutonura albella (Stach): Traser & Salló 2004: 230; Deutonura phlegraea (Caroli, 1912) Traser 2006: 35. Achorutes phlegraeus Caroli: Stach 1929: 283; Stach 1930: 57, figs 17/2-3.; Deutonura phlegraea (Caroli): General distribution: Central and Southern Eu- Traser 1995: 44; Traser 1999: 52; Traser 2002a: 262; rope. Traser & Horváth-Szováti 2006: 333. Distribution in Hungary: Gerecse Mts. (Traser 2006), Kőszeg Mts. (Traser & Salló 2004). General distribution: Central and Southern Eu- rope. Deutonura benzi Traser, Thibaud & Najt, 1993 Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Kiskunság (Traser & Horváth-Szováti Deutonura benzi Traser, Thibaud & Najt: Traser et al. 1993: 222, figs 3-5; Traser 1994: 100; Traser 1995: 44; 2006), Mátra Mts. (Stach 1929, 1930), Őrség Traser 2002a: 262; Traser & Salló 2004: 230. (Traser 1995), Szigetköz (Traser 2002a).

15 Dányi & Traser: Checklist of the springtail fauna of Hungary

Deutonura stachi (Gisin, 1952) Loksa 1977: 210; Loksa 1978a: 55; Loksa 1978b: 95; Loksa 1980: 110; Loksa 1983: 72; Loksa 1987: 79; Neanura stachi Gisin: Loksa 1966: 379; Varga & Loksa 1991: 270; Varga 1991: 85; Varga 1992: 172; Vargha 1992: 315; Neanura stachi Templeton (sic!): Varga & Vargha 1992: 315; Traser 1995: 44; Varga & Varga 1992: 172; Varga & Oldal 1998: 86; Deutonura Oldal 1998: 86; Traser 1999: 52; Traser & Csóka 2000: cf. stachi (Gisin): Traser 1999: 52. 77; Traser & Csóka 2001: 233; Traser 2002a: 262; General distribution: Eastern Europe, Carpa- Traser 2002c: 48; Traser 2003: 155; Traser & Salló 2004: 230; Traser & Horváth-Szováti 2006: 333; thian Mts. Traser et al. 2006: 72; Neanura (Neanura) muscorum Distribution in Hungary: Aggtelek Karst (Tra- (Templeton): Loksa & Rubio 1966a: 142. ser 1999), Bükk Mts. (Loksa 1966; Varga 1992; Varga & Vargha 1992; Varga & Oldal 1998). General distribution: cosmopolitan. Distribution in Hungary: Aggtelek Karst Endonura Cassagnau, 1979 (Stach 1929; Dudich 1932; Stach 1951; Loksa 1966; Traser 1999), Bakony Mts. (Loksa 1966; Endonura tetrophthalma (Stach, 1929) Loksa & Rubio 1966a), Balatonfelvidék (Loksa Achorutes tetrophthalmus Stach: Stach 1929: 282; 1966), Börzsöny Mts. (Loksa 1966; Dunger Biloba tetrophthalma (Stach) f. principalis: Stach 1975), Buda Mts. (Loksa 1956b, 1966), Bükk 1951: 52, pl. 8; Neanura tetrophthalma (Stach): Loksa 1966: 379; Loksa 1977: 210; Traser 1980: 19, figs 1a- Mts. (Loksa 1956a, 1962, 1966, 1977; Varga b; Traser 1981: 133; Loksa 1991: 270; Neanura tet- 1991, 1992; Varga & Vargha 1992; Varga & Ol- rophthalma tetrophthalma (Stach): Dunger 1975: 17. dal 1998), Cserhát (Stach 1929, 1951; Loksa General distribution: Europe: East Carpathian 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság Mts. (Traser 2002a, 2003; Traser et al. 2006), Hor- Distribution in Hungary: Bakony Mts. (Traser tobágy (Loksa 1983), Keszthely Mts. (Loksa 1980), Balatonfelvidék (Stach 1929, 1951), Bör- 1966), Kiskunság (Vellay 1900; Stach 1929; Lok- zsöny Mts. (Dunger 1975), Bükk Mts. (Loksa sa 1987; Traser & Csóka 2000, 2001; Traser 1977), Keszthely Mts. (Loksa 1966), Nyírség 2002c; Traser & Horváth-Szováti 2006), Kőszeg (Loksa 1991), Sopron Mts. (Traser 1981). Mts. (Szent-Ivány 1940a; Traser & Salló 2004), Mátra Mts. (Loksa 1980), Mecsek Mts. (Loksa 1966), Mt. Somló (Loksa 1966), Nyírség (Kovács Neanura MacGillivray, 1893 1953a; Loksa 1991), Őrség (Traser 1995), Pilis Neanura alba von Törne, 1956 Mts. (Stach 1929, 1951), Sopron Mts. (Traser 2002a), Vác-Pest Danube Valley (Tömösváry Neanura parva (Stach): Dunger 1975: 17 (partim); Neanura alba von Törne: Traser 1995: 44. 1884a; Vellay 1900; Stach 1929), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966). Remark: The data of Neanura parva (Stach) reported by Dunger (1975) included Neanura alba Neanura parva (Stach, 1951) as well. Biloba parva Stach: Stach 1951: 48, pl. 5; Neanura General distribution: Central Europe. parva (Stach): Loksa 1966: 379; Dunger 1975: 17 Distribution in Hungary: Börzsöny Mts. (Dun- (partim); Varga & Oldal 1998: 86; Traser 1999: 52. ger 1975), Őrség (Traser 1995). Ecological note: in mountainous habitats. Remark: The data of Neanura parva (Stach) reported by Dunger (1975) included Neanura alba Neanura muscorum (Templeton, 1835) von Törne as well. Anura muscorum Templeton: Tömösváry 1884a: 50; General distribution: Eastern Europe. Achorutes muscorum Templeton: Stach 1929: 282; Distribution in Hungary: Aggtelek Karst (Lok- Dudich 1932: 48; Szent-Ivány 1940a: 428; Biloba sa 1966; Traser 1999), Balatonfelvidék (Loksa muscorum (Templeton): Stach 1951: 41, pl. 4; Neanura 1966), Börzsöny Mts. (Loksa 1966; Dunger muscorum (Templeton): Vellay 1900: 21; Kovács 1975), Buda Mts. (Loksa 1966), Bükk Mts. (Var- 1953a: 183; Loksa 1956a: 391; Loksa 1956b: 205; ga & Oldal 1998), Cserhát (Stach 1951), Keszt- Loksa 1962: 68; Loksa 1966: 379; Dunger 1975: 17; hely Mts. (Loksa 1966), Mecsek Mts. (Loksa

16 Dányi & Traser: Checklist of the springtail fauna of Hungary

1966), Mt. Somló (Loksa 1966), Pilis Mts. (Loksa 1941), Mátra Mts. (Szent-Ivány 1940a, 1941), 1966), Vértes Mts. (Loksa 1966), Villány Mts Pilis Mts. (Stach 1929), Sopron Mts. (Traser (Loksa 1966), Visegrád Mts. (Loksa 1966). 2002a). Ecological note: silvicolous. Neanura weigneri (Stach, 1922) Neanura weigneri Stach: Kovács 1953a: 183; Loksa Subfamily: Pseudachorutinae 1991: 26. Anurida Laboulbène, 1865 General distribution: Central and Southern Eu- rope (Albania, Hungary). Anurida ellipsoides Stach, 1949 Distribution in Hungary: Nyírség (Kovács Anurida ellipsoides Stach: Loksa 1978b: 95; Traser 1953a; Loksa 1991). 2002a: 261.

Pumilinura Cassagnau, 1979 General distribution: Eastern Europe. Distribution in Hungary: Cserhát (Loksa Pumilinura loksai (Dunger, 1974) 1978b), Sopron Mts. (Traser 2002a). Neanura loksai Dunger: Dunger 1974: 10, figs 5d-g, 6a-b; Dunger 1975: 17; Pumilinura loksai (Dunger): Traser 1999: 52; Traser & Salló 2004: 230. Anurida granulata Agrell, 1943 Anurida granulata Agrell: Dunger 1975: 17; Traser General distribution: Europe. 2002a: 261; Micranurida granulata (Agrell): Traser Distribution in Hungary: Aggtelek Karst (Tra- 2002a: 261. ser 1999), Börzsöny Mts. (Dunger 1974, 1975), Kőszeg Mts. (Traser & Salló 2004). General distribution: Europe. Distribution in Hungary: Börzsöny Mts. (Dun- Thaumanura Börner, 1932 ger 1975), Fertő-Hanság (Traser 2002a), Sopron Mts. (Traser 2002a). Thaumanura carolii (Stach, 1920) Ecological note: arctoalpine. Achorutes carolii Stach: Dudich 1932: 48; Thauma- nura carolii (Stach): Stach 1951: 61, pl. 1: fig. 4, pl. Anurida tullbergi Schött, 1891 12; Traser 1999: 52; Traser 2002a: 262; Neanura caroli (sic!) (Stach): Loksa 1991: 270. Anurida tullbergi Schött: Stach 1929: 282; Stach 1949a: 22, pl. 2: figs 1-10; Farkas 1995: 46; Traser General distribution: Eastern Europe. 1997: 61; Farkas et al. 1998: 57; Traser 2002a: 261.

Distribution in Hungary: Aggtelek Karst (Du- General distribution: Holarctic. dich 1932; Stach 1951; Traser 1999), Börzsöny Distribution in Hungary: around the Lake Ba- Mts. (Stach 1951), Nyírség (Loksa 1991), Pilis laton (Farkas 1995), Balatonfelvidék (Stach 1929, Mts. (Stach 1951), Sokoróalja (Stach 1951), Sop- 1949a), Fertő-Hanság (Traser 2002a), Somogy ron Mts. (Traser 2002a). Hills (Farkas et al. 1998), Szigetköz (Traser 1997, Ecological note: silvicolous. 2002a).

Ecological note: hygrophilous, mostly litoral. Thaumanura oniscoides (Latzel, 1917)

Achorutes oniscoides Latzel: Stach 1929: 283; Szent- Anurida uniformis Gisin, 1953 Ivány 1940a: 428; Szent-Ivány 1941: 21; Thaumanura oniscoides (Latzel): Traser 2002a: 262. Anurida uniformis Gisin: Traser 2002a: 261.

General distribution: Central Europe. General distribution: Palaearctic. Distribution in Hungary: Börzsöny Mts. Distribution in Hungary: Fertő-Hanság (Traser (Stach 1929), Kőszeg Mts. (Szent-Ivány 1940a, 2002a).

17 Dányi & Traser: Checklist of the springtail fauna of Hungary

Micranurida Börner, 1901 General distribution: Palaearctic including Ca- nary Islands. Micranurida cf. anophthalmica Stach, 1949 Distribution in Hungary: Börzsöny Mts. (Dun- Micranurida cf. anophthalmica Stach: Traser 1980: 20. ger 1974, 1975).

General distribution: Central Europe, Carpa- Pratanurida Rusek, 1973 thian Mts. Distribution in Hungary: Bakony Mts. (Traser Pratanurida cassagnaui Rusek, 1973 1980). Pratanurida cassagnaui Rusek: Traser & Horváth-Szo- váti 2006: 333. Micranurida forsslundi Gisin, 1949 Micranurida cf. forsslundi Gisin: Traser 1980: 22; General distribution: Palaearctic. Micranurida forsslundi Gisin: Traser 2002a: 261; Tra- Distribution in Hungary: Kiskunság (Traser & ser 2006: 35. Horváth-Szováti 2006).

General distribution: Europe, Siberia. Pseudachorutella Stach, 1949 Distribution in Hungary: Bakony Mts. (Traser 1980), Gerecse Mts. (Traser 2006), Sopron Mts. Pseudachorutella asigillata (Börner, 1901) (Traser 2002a). Pseudachorutella asigillata (Börner): Dunger 1975: 17; Traser 1999: 52. Micranurida cf. meridionalis Cassagnau, 1952

Micranurida cf. meridionalis (Cassagnau): Traser General distribution: Europe. 2002a: 261. Distribution in Hungary: Aggtelek Karst (Tra-

General distribution: Mediterranean Europe. ser 1999), Börzsöny Mts. (Dunger 1975). Distribution in Hungary: Fertő-Hanság (Traser 2002a). Pseudachorutella clavata (Börner, 1901) Pseudachorutes (Pseudachorutella) clavatus (Börner): Micranurida pygmaea Börner, 1901 Loksa & Rubio 1966a: 142;

Anurida pygmaea (Börner): Loksa & Rubio 1966a: Remark: According to Gama (1964) the speci- 142; Loksa 1977: 210; Loksa 1978a: 55; Loksa 1980: 110; Traser 1997: 61; Traser & Csóka 2000: 77; Traser es is a junior synonym of Pseudachorutella asigil- & Csóka 2001: 237; Traser 2002c: 48; Micranurida lata (Börner), but it is a valid, separate taxon in pygmaea Börner: Dunger 1975: 17; Traser 2002a: 261; Traser & Horváth-Szováti 2006: 333; Traser et al. our opinion. 2006: 72. General distribution: Europe. Distribution in Hungary: Bakony Mts. (Loksa General distribution: Holarctic. & Rubio 1966a). Distribution in Hungary: Bakony Mts. (Loksa & Rubio 1966a), Börzsöny Mts. (Dunger 1975), Pseudachorutes Tullberg, 1871 Bükk Mts. (Loksa 1977), Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser 2002a; Traser et al. Pseudachorutes corticicolus (Schäffer, 1897) 2006), Kiskunság (Traser & Csóka 2000, 2001; Traser 2002c; Traser & Horváth-Szováti 2006), Pseudachorutes corticicola (sic!) (Schäffer): Stach Mátra Mts. (Loksa 1980), Sopron Mts. (Traser 1929: 282; Loksa 1977: 210; Loksa 1978b: 95; Loksa 2002a), Szigetköz (Traser 1997). 1978a: 55; Loksa 1980: 110; Loksa 1983: 72; Loksa Ecological note: acidophilous. 1991: 270; Farkas 1995: 46; Farkas 1997: 56; Farkas & Szathmáry 1997: 39; Pseudachorutes corticicolus (Schäffer): Traser 1999: 52; Pseudachorutes corticola Micranurida sensillata (Gisin, 1953) (sic!) (Schäffer): Farkas 1996: 287; Farkas 1997: 58. Anurida endroedii Dunger: Dunger 1974: 6, figs 3a-e, 5a-c; Dunger 1975: 17. General distribution: Palaearctic.

18 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Aggtelek Karst (Tra- ron Mts. (Traser 2002a), Szigetköz (Traser 1997, ser 1999), around the Lake Balaton (Farkas 1995, 2002a), Villány Mts (Loksa 1966). 1997), Balatonfelvidék (Stach 1929; Farkas 1995, 1997; Farkas & Szathmáry 1997), Bükk Mts. Pseudachorutes pratensis Rusek, 1973 (Loksa 1977), Cserhát (Loksa 1978b), Dráva Pseudachorutes pratensis Rusek: Traser & Horváth- Plain (Loksa 1978a), Hortobágy (Loksa 1983), Szováti 2006: 333. Kis-Balaton (Farkas 1996, 1997), Mátra Mts. (Loksa 1980), Nyírség (Loksa 1991). General distribution: Europe. Ecological note: corticophilous. Distribution in Hungary: Kiskunság (Traser & Horváth-Szováti 2006). Pseudachorutes dubius Krausbauer, 1898 Pseudachorutes dubius Krausbauer: Stach 1929: 281; Pseudachorutes subcrassus Tullberg, 1871 Stach 1949a: 101, pl. 12, pl. 13: figs 1-5; Kovács Pseudachorutes subcrassus Tullberg: Traser 1980: 20; 1953a: 183; Dunger 1975: 17; Loksa 1978a: 55; Loksa Traser 1981: 133; Traser 1997: 61; Traser 1999: 52; 1983: 72; Loksa 1987: 79; Loksa 1991: 270; Traser Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; 1995: 44; Traser 1999: 52; Traser 2002a: 261; Traser Traser 2002a: 261; Traser 2002c: 48; Peudachorutes 2002c: 48; Pseudachorutes (Pseudachorutes) dubius (sic!) subcrassus Tullberg, 1871: Traser 2002b: 159. Krausbauer: Loksa & Rubio 1966a: 142. General distribution: Palaearctic. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Bakony Mts. (Traser 1980), Bükk Mts. ser 1999), Bakony Mts. (Loksa & Rubio 1966a), (Traser 2002b), Fertő-Hanság (Traser 2002a), Balatonfelvidék (Stach 1929), Börzsöny Mts. Kiskunság (Traser & Csóka 2000, 2001; Traser (Dunger 1975), Dráva Plain (Loksa 1978a), Fertő- 2002c), Sopron Mts. (Traser 1981, 2002a), Szi- Hanság (Traser 2002a), Hortobágy (Loksa 1983), getköz (Traser 1997, 2002b), Vas-Sopron Plain Kiskunság (Loksa 1987; Traser 2002c), Mezőföld (Traser 1981). (Stach 1949a), Nyírség (Kovács 1953a; Loksa 1991), Őrség (Traser 1995), Sopron Mts. (Traser Subfamily: Odontellidae 2002a). Axenyllodes Stach, 1949 Pseudachorutes parvulus Börner, 1901 Axenyllodes ukrainus Thibaud & Taraschuk, Pseudachorutes parvulus Börner: Loksa 1966: 379; Dunger 1975: 16; Loksa 1983: 72; Loksa 1987: 79; 1997 Traser 1997: 61; Traser 1999: 52; Traser & Csóka Axenyllodes ukrainus Thibaud & Taraschuk: Traser 2000: 77; Traser & Csóka 2001: 237; Traser 2002a: 2002a: 262. 261; Pseudachorutes (Pseudachorutes) parvulus Bör- ner: Loksa & Rubio 1966a: 142; Pseudachorutes cf. General distribution: Eastern Europe. parvulus Börner: Traser 2002c: 48; Traser 2006: 35. Distribution in Hungary: Fertő-Hanság (Traser 2002a). General distribution: Palaearctic, Japan. Distribution in Hungary: Aggtelek Karst (Lok- Pseudostachia Arlé, 1968 sa 1966; Traser 1999), Bakony Mts. (Loksa 1966; Loksa & Rubio 1966a), Balatonfelvidék (Loksa Pseudostachia populosa (Selga, 1963) 1966), Börzsöny Mts. (Dunger 1975), Buda Mts. (Loksa 1966), Fertő-Hanság (Traser 2002a), Ge- Pseudostachia populosa (Selga): Traser & Horváth- Szováti 2006: 333. recse Mts. (Loksa 1966; Traser 2006), Hortobágy (Loksa 1983), Keszthely Mts. (Loksa 1966), Kis- General distribution: Palaearctic. kunság (Loksa 1987; Traser & Csóka 2000, 2001; Distribution in Hungary: Kiskunság (Traser & Traser 2002c), Mecsek Mts. (Loksa 1966), Sop- Horváth-Szováti 2006).

19 Dányi & Traser: Checklist of the springtail fauna of Hungary

Superodontella Stach, 1949 Xenyllodes Axelson, 1903

Superodontella empodialis (Stach, 1934) Xenyllodes armatus Axelson, 1903 Odontella empodialis Stach: Loksa 1966: 379; Dunger Xenyllodes armatus Axelson: Traser 1999: 51; Traser 1975: 16; Traser 1999: 51. 2002a: 262; Traser 2003: 155.

General distribution: Europe. General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Börzsöny Mts. (Dunger 1975), Mecsek ser 1999), Fertő-Hanság (Traser 2002a), Fertő- Mts. (Loksa 1966). Hanság (Traser 2003).

Superodontella lamellifera (Axelson, 1903) Family: Onychiuridae Odontella lamellifera Axelson: Loksa 1956b: 211; Subfamily: Onychiurinae Loksa 1961b: 224; Loksa & Rubio 1966a: 142; Traser 1997: 61; Superodontella lamellifera (Axelson): Traser 2002a: 262. Deharvengiurus Weiner, 1996

General distribution: cosmopolitan. Deharvengiurus denisi (Stach, 1934) Distribution in Hungary: Aggtelek Karst (Lok- Onychiurus (Onychiurus) denisi Stach: Loksa & Rubio sa 1961b), Bakony Mts. (Loksa & Rubio 1966a), 1966a: 151; Onychiurus (Paronychiurus) denisi Stach: Buda Mts. (Loksa 1956b), Sopron Mts. (Traser Dunger 1975: 18. 2002a), Szigetköz (Traser 1997, 2002a). General distribution: Eastern Europe. Superodontella pseudolamellifera (Stach, 1949) Distribution in Hungary: Bakony Mts. (Loksa Odontella pseudolamellifera Stach: Loksa 1966: 379; & Rubio 1966a), Börzsöny Mts. (Dunger 1975). Loksa 1978b: 95; Odontella pseodolamerrifera (sic!) Ecological note: in caves too. Stach: Traser 1999: 51. Deharvengiurus microchaetosus (Loksa, 1959) General distribution: Europe. Onychiurus microchaetosus Loksa: Loksa 1959b: 75, Distribution in Hungary: Aggtelek Karst (Tra- figs 16-19. ser 1999), Cserhát (Loksa 1978b), Pilis Mts. (Loksa 1966). General distribution: Hungary. Distribution in Hungary: Cserhát (Loksa Superodontella cf. sensillata (Deharveng & 1959b). Izarra, 1979) Superodontella cf. sensillata Deharveng & Izarra: Tra- Deharvengiurus xerophilus (Loksa, 1964) ser 2002a: 262. Onychiurus (Onychiurus) xerophilus Loksa: Loksa 1964: 86, figs 16-23; Onychiurus xerophilus Loksa: General distribution: Europe. Loksa 1966: 380. Distribution in Hungary: Fertő-Hanság (Traser 2002a). General distribution: Hungary. Distribution in Hungary: Mecsek Mts. (Loksa Superodontella cf. stachi (Denis, 1947) 1964, 1966). Odontella cf. stachi Denis: Traser 1999: 51. Deuteraphorura Absolon, 1901 General distribution: Europe Deuteraphorura fimetaria (Linnaeus, 1766) Distribution in Hungary: Aggtelek Karst (Tra- ser 1999). Onychiurus fimetarius (auct.) Stach: Stach 1934: 164,

20 Dányi & Traser: Checklist of the springtail fauna of Hungary

pl. 33; Stach 1954: 173, pl. 24; Loksa 1962: 75, figs General distribution: Central and Eastern Eu- 17-22; Onychiurus fimetarius Linnaeus: Stach 1929: rope. 285; Gebhardt 1934: 139; Kovács 1953b: 90; Kovács 1954: 97; Loksa 1961b: 224; Onychiurus subterraneus Distribution in Hungary: Aggtelek Karst (Tra- Stach: Stach 1929: 285. ser 1999), Hortobágy (Loksa 1983), Mecsek Mts. (Loksa 1966). Remark: Species inquirenda according to Gisin (1960). Heteraphorura Bagnall, 1948 General distribution: West Palaearctic. Distribution in Hungary: Aggtelek Karst (Ko- Heteraphorura variotuberculata (Stach, 1934) vács 1953b, 1954; Loksa 1961b), Buda Mts. Onychiurus (Hymenaphorura) terricola Kos: Dunger (Stach 1929), Budapest (Stach 1954), Bükk Mts. 1975: 18; Onychiurus terricola Kos: Traser 1995: 44; (Stach 1929, Loksa 1962), Mecsek Mts. (Geb- Heteraphorura terricola (Kos): Traser 1999: 53. hardt 1934; Stach 1934, 1954). General distribution: Eastern Europe, Carpa- Deuteraphorura imperfecta (Denis, 1938) thian Mts. Distribution in Hungary: Aggtelek Karst (Tra- Onychiurus imperfectus Denis: Loksa 1977: 210. ser 1999), Börzsöny Mts. (Dunger 1975), Őrség

(Traser 1995). General distribution: Central Europe. Ecological note: in caves too. Distribution in Hungary: Bükk Mts. (Loksa

1977). Hymenaphorura Bagnall, 1948 Ecological note: in caves too.

Hymenaphorura dentifera (Stach, 1934) Deuteraphorura silesiaca (Dunger, 1977) Onychiurus silesiacus Dunger: Traser 1994: 100; Hymenaphorura creatricis Pomorski: Traser & Salló Traser & Thibaud 1999: 106; Deuteraphorura 2004: 230. silesiaca (Dunger): Traser 2002a: 263. Remark: Species inquirenda according to Gisin General distribution: Eastern Europe. (1960). Distribution in Hungary: Sopron Mts. (Traser General distribution: West Palaearctic. 1994, 2002a), Vas-Sopron Plain (Traser & Thi- Distribution in Hungary: Kőszeg Mts. (Traser baud 1999). & Salló 2004). Ecological note: silvicolous, montane. Hymenaphorura pseudosibirica (Stach, 1954) Deuteraphorura silvaria (Gisin, 1952) Onychiurus pseudosibiricus Stach: Stach 1954: 31. Onychiurus (Onychiurus) silvarius Gisin: Dunger 1975: 19; Onychiurus silvarius Gisin: Loksa 1978b: General distribution: Hungary. 95; Deuteraphorura silvaria (Gisin): Traser 2002a: 263; Traser 2006: 34. Distribution in Hungary: Aggtelek Karst (Stach 1954). General distribution: Europe. Ecological note: troglophilous. Distribution in Hungary: Börzsöny Mts. (Dun- ger 1975), Cserhát (Loksa 1978b), Gerecse Mts. Hymenaphorura sibirica (Tullberg, 1876) (Traser 2006), Sopron Mts. (Traser 2002a). Onychiurus sibiricus (Tullberg): Stach 1929: 284; Ecological note: troglophilous. Dudich 1932: 48; Gebhardt 1933: 39; Stach 1934: 138, pls 27-28; Stach 1954: 96, pl. 6; Loksa 1978b: 95; Deuteraphorura variabilis (Stach, 1954) Onychiurus (Protaphorura) sibiricus (Tullberg): Loksa & Rubio 1966a: 146; Onychiurus (Hymenaphorura) Onychiurus variabilis Stach: Loksa 1966: 381; Loksa sibiricus (Tullberg): Dunger 1975: 18; Hymenaphorura 1983: 73; Traser 1999: 53. sibiricus (Tullberg): Traser 1999: 53.

21 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Northern Europe, Sibe- raphorura absoloni (Börner): Traser 1999: 53; Traser ria. 2002a: 262; Traser 2003: 155. Distribution in Hungary: Aggtelek Karst (Du- General distribution: Holarctic. dich 1932; Stach 1934; Traser 1999), Bakony Distribution in Hungary: Aggtelek Karst (Tra- Mts. (Loksa & Rubio 1966a), Börzsöny Mts. ser 1999), Börzsöny Mts. (Dunger 1975), Buda (Stach 1929, 1954; Dunger 1975), Cserhát (Loksa Mts. (Loksa 1966), Dráva Plain (Loksa 1978a), 1978b), Mecsek Mts. (Gebhardt 1933; Stach Fertő-Hanság (Traser 2002a, 2003), Keszthely 1934, 1954). Mts. (Loksa 1966), Mecsek Mts. (Loksa 1966),

Nyírség (Loksa 1991), Pilis Mts. (Stach 1954; Kalaphorura Absolon, 1901 Loksa 1966), Sopron Mts. (Traser 2002a), Vértes

Mts. (Loksa 1966), Villány Mts (Loksa 1966). Kalaphorura burmeisteri (Lubbock, 1873) Ecological note: detritophilous. Onychiurus (Kalaphorura) burmeisteri Lubbock: Ko- vács 1953a: 183 (partim?); Kalaphorura burmeisteri Lubbock: Loksa 1959b: 70 (partim?); Loksa 1991: Oligaphorura Bagnall, 1949 271. Oligaphorura groenlandica (Tullberg, 1876) Remark: According to Dunger (1975) the ear- Onychiurus groenlandicus (Tullberg): Stach 1929: lier Hungarian data of Kalaphorura burmeisteri 284. could included Kalaphorura paradoxa (Schäffer, 1900) as well. General distribution: Holarctic, circumpolar. General distribution: Europe. Distribution in Hungary: Pilis Mts. (Stach Distribution in Hungary: Cserhát (Loksa 1929). 1959b), Nyírség (Kovács 1953a; Loksa 1991). Ecological note: in caves too. Onychiuroides Bagnall, 1948

Kalaphorura paradoxa (Schäffer, 1900) Onychiuroides granulosus (Stach, 1930) Onychiurus (Kalaphorura) paradoxus (Schäffer): Onychiurus granulosus Stach: Stach 1929: 285; Stach Dunger 1975: 18; Kalaphorura paradoxa (Schäffer): 1934: 193, pl. 39; Stach 1954: 146, pl. 17; Loksa 1966: Traser 1999: 53; Traser & Thibaud 1999: 106; Traser 380; Varga 1992: 172; Varga & Oldal 1998: 86; 2002a: 262; Kalaphorura paradoxa var. clavipila Onychiuroides granulosus (Stach): Traser 1999: 53; (Stach): Traser 1999: 53; Onychiurus burmeisteri Lub- Traser 2002a: 263; Traser & Salló 2004: 230. bock: Loksa 1966: 380. General distribution: Europe. Remark: Loksa’s „Onychiurus burmeisteri” Distribution in Hungary: Aggtelek Karst (Lok- data were corrected by (Traser 1999). sa 1966; Traser 1999), Bakony Mts. (Stach 1929, General distribution: Central Europe. 1934, 1954), Bükk Mts. (Loksa 1966; Varga Distribution in Hungary: Aggtelek Karst (Tra- 1992; Varga & Oldal 1998), Kőszeg Mts. (Traser ser 1999; Loksa 1966), Börzsöny Mts. (Dunger & Salló 2004), Sopron Mts. (Traser 2002a). 1975), Vas-Sopron Plain (Traser & Thibaud 1999; Ecological note: troglophilous. Traser 2002a). Ecological note: in caves too. Onychiuroides pseudogranulosus (Gisin, 1951)

Onychiurus pseudogranulosus Gisin: Loksa 1966: 380; Micraphorura Bagnall, 1949 Loksa 1977: 207; Loksa 1991: 272; Traser 1994: 102; Onychiurus (Onychiurus) pseudogranulosus Gisin: Micraphorura absoloni (Börner, 1901) Loksa & Rubio 1966a: 150; Onychiurus cf. pseudo- granulosus Gisin: Traser 1981: 135; Onychiuroides Onychiurus affinis Ågren: Stach 1954: 85, pl. 2; pseudogranulosus Gisin: Traser 1995: 44; Traser 1999: Onychiurus absoloni Börner: Loksa 1966: 380; Loksa 53; Traser 2002a: 263. 1991: 271; Onychiurus (Olygaphorura) absoloni (Börner): Dunger 1975: 18; Loksa 1978a: 56; Mic- General distribution: Europe.

22 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Aggtelek Karst (Lok- General distribution: Eastern Europe. sa 1966; Traser 1999), Bakony Mts. (Loksa & Distribution in Hungary: Mecsek Mts. (Paclt Rubio 1966a), Börzsöny Mts. (Loksa 1966), Buda 1960). Ecological note: silvicolous. Mts. (Loksa 1966), Bükk Mts. (Loksa 1966,

1977), Nyírség (Loksa 1991), Őrség (Traser Onychiurus rectospinatus Stach, 1922 1995), Pilis Mts. (Loksa 1966), Sopron Mts. (Traser 1994, 2002a), Vas-Sopron Plain (Traser Onychiurus rectospinatus Stach: Loksa 1961b: 224; Traser 1999: 54; Traser 2002a: 263. 1981), Visegrád Mts. (Loksa 1966). General distribution: Europe. Onychiuroides subgranulosus (da Gama, 1964) Distribution in Hungary: Aggtelek Karst (Lok- Onychiurus (Paronychiurus) subgranulosus Gama: sa 1961b; Traser 1999), Moson Plain (Traser Dunger 1975: 18. 2002a). Ecological note: troglophilous. General distribution: Europe. Distribution in Hungary: Börzsöny Mts. (Dun- Onychiurus stillicidii (Schiödte 1849) ger 1975). Lipura stillicidii Schiödte: Tömösváry 1884a: 50; Anurophorus stillicidii Schiödte: Vellay 1900: 21. Onychiurus Gervais, 1841 General distribution: Eastern and Southern Onychiurus ambulans (Linnaeus, 1758) Europe. ?Lipura ambulans Linnaeus: Tömösváry 1884a: 50; Distribution in Hungary: Vác-Pest Duna Val- ?Aphorura ambulans Linnaeus: Vellay 1900: 21. ley (Tömösváry 1884a; Vellay 1900). Ecological note: troglophilous. Remark: All data published before description of Onychiurus circulans Gisin, 1952 are dubious. Orthonychiurus Stach, 1954 General distribution: Europe. Distribution in Hungary: Budapest (Tömösvá- Orthonychiurus cf. pseudostachianus (Gisin, ry 1884a; Vellay 1900), Vác-Pest Duna Valley 1956) (Tömösváry 1884a; Vellay 1900). Onychiurus cf. pseudostachianus (Gisin): Traser Ecological note: troglophilous. 2002a: 263.

Remark: The differences from the species Or- Onychiurus circulans Gisin, 1952 thonychiurus stachianus (Bagnall 1939) must be Onychiurus (Onychiurus) circulans Gisin: Loksa 1964: re-evaluated (Pomorski 1998: 182). 94, figs 45-50; Onychiurus circulans Gisin: Loksa General distribution: Western Palaearctic. 1966: 380; Loksa 1977: 210. Distribution in Hungary: Fertő-Hanság (Traser 2002a). Remark: According to Pomorsky (1998) the species is a junior synonym of Onychiurus am- Orthonychiurus rectopapillatus (Stach, 1933) bulans (Linnaeus), but in our opinion it is a valid, Onychiurus rectopapillatus Stach: Traser 1999: 53. separate taxon. General distribution: Europe. General distribution: Central Europe. Distribution in Hungary: Bükk Mts. (Loksa Distribution in Hungary: Aggtelek Karst (Tra- 1977), Gerecse Mts. (Loksa 1964, 1966), Mecsek ser 1999). Mts. (Loksa 1964, 1966). Orthonychiurus schoenviszkyi (Loksa, 1967) Onychiurus pseudinermis Börner, 1903 Onychiurus (Onychiurus) schoenviszkyi Loksa: Loksa Onychiurus pseudinermis Börner: Paclt 1960: 156. 1967: 292, figs 14-17; Onychiurus schoenviszkyi Lok-

23 Dányi & Traser: Checklist of the springtail fauna of Hungary

sa: Bajomi 1969: 243; Onychiurus schoeviszki (sic!) 2002a), Szigetköz (Traser 1997), Vas-Sopron Loksa: Traser 1999: 49. Plain (Traser 1981), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), Visegrád Mts. (Loksa General distribution: Hungary. Distribution in Hungary: Aggtelek Karst (Lok- 1966). sa 1967; Bajomi 1969; Traser 1999). Ecological note: euedaphic.

Protaphorura Absolon, 1901 Protaphorura aurantiaca (Ridley, 1880) Onychiurus latus Gisin: Loksa 1977: 210; Loksa Protaphorura armata (Tullberg, 1869) 1978b: 95; Protaphorura latus (Gisin): Traser 1999: 53; Protaphorura aurantiaca (Ridley): Traser 2002a: Onychiurus armatus (Tullberg): Stach 1929: 284; 262. Dudich 1932: 48; Stach 1934: 122, pl. 25; Szent-Ivány 1940a: 428; Kovács 1953b: 90; Stach 1954: 123, pl. 12; Loksa 1956a: 391; Loksa 1956b: 203; Paclt 1960: General distribution: Europe. 156; Loksa 1962: 63; Loksa 1966: 380; Loksa 1977: Distribution in Hungary: Aggtelek Karst (Tra- 206; Traser 1980: 20, fig. 3; Traser 1981: 133; ser 1999), Bükk Mts. (Loksa 1977), Cserhát (Lok- Hornung 1986: 138; Loksa 1987: 79; Loksa 1991: 272; sa 1978b), Fertő-Hanság (Traser 2002a), Sopron Varga & Vargha 1992: 315; Farkas 1996: 287; Farkas 1997: 58; Farkas & Szathmáry 1997: 39; Traser 1997: Mts. (Traser 2002a). 61; Traser 2002a: 268; Onychiurus armata (sic!) Tullberg: Kovács 1954: 97; Onychorus (sic!) armatus Protaphorura bicampata (Gisin, 1956) Tullberg: Loksa 1961b: 224; Onychiurus (Protapho- rura) armatus (Tullberg): Dunger 1975: 18; Loksa & Onychiurus cf. bicampatus Gisin: Traser 1995: 44; Rubio 1966a: 146; Loksa 1978a: 56; Protaphorura Protaphorura bicampata (Gisin): Traser 2003: 155. armatus (sic!) (Tullberg): Traser 1999: 53; Traser 2002a: 262; Traser 2006: 35; Protaphorura armata General distribution: Central Europe. (Tullberg): Traser 2002c: 48; Traser & Horváth- Distribution in Hungary: Fertő-Hanság (Traser Szováti 2006: 332; Traser et al. 2006: 72. 2003), Őrség (Traser 1995). General distribution: cosmopolitan. Distribution in Hungary: Aggtelek Karst (Du- Protaphorura campata (Gisin, 1952) dich 1932; Stach 1934; Kovács 1953b, 1954; Onychiurus campatus Gisin: Loksa 1991: 271; Varga Stach 1954; Loksa 1961b, 1966; Traser 1999), 1992: 172; Farkas 1995: 46; Farkas et al. 1998: 57; Varga & Oldal 1998: 86; Protaphorura campatus (sic!) Bakony Mts. (Stach 1929; Loksa 1966; Loksa & (Gisin): Traser 1999: 53. Rubio 1966a; Traser 1980), Balatonfelvidék (Stach 1929; Loksa 1966; Farkas & Szathmáry General distribution: Palaearctic. 1997), Börzsöny Mts. (Stach 1929; Loksa 1966; Distribution in Hungary: Aggtelek Karst (Tra- Dunger 1975), Buda Mts. (Stach 1929, 1934, ser 1999), around the Lake Balaton (Farkas 1995), 1954; Loksa 1956b, 1966), Budapest (Stach Bükk Mts. (Varga 1992; Varga & Oldal 1998), 1929), Bükk Mts. (Loksa 1956a, 1962, 1966, Nyírség (Loksa 1991), Somogy Hills (Farkas et 1977; Varga & Vargha 1992), Dráva Plain (Loksa al. 1998). 1978a), Fertő-Hanság (Paclt 1960; Traser 2002a; Traser et al. 2006), Gerecse Mts. (Loksa 1966; Protaphorura cancellata (Gisin, 1956) Traser 2006), Keszthely Mts. (Loksa 1966), Kis- Onychiurus (Protaphorura) cancellatus Gisin: Loksa Balaton (Farkas 1996, 1997), Kiskunság (Hor- 1978a: 56; Onychiurus cancellatus Gisin: Loksa 1983: 73; Traser 1997: 61; Protaphorura cancellatus (sic!) nung 1986; Loksa 1987; Traser 2002c; Traser & (Gisin): Traser 1999: 53; Traser 2002a: 262; Horváth-Szováti 2006), Kőszeg Mts. (Szent-Ivány Protaphorura cancellata (Gisin): Traser 2006: 35. 1940a), Mecsek Mts. (Stach 1934, 1954; Paclt 1960; Loksa 1966), Mezőföld (Stach 1929, 1954), General distribution: Europe. Moson Plain (Traser 2002a), Mt. Somló (Loksa Distribution in Hungary: Aggtelek Karst (Tra- 1966), Nyírség (Loksa 1991), Pilis Mts. (Stach ser 1999), Dráva Plain (Loksa 1978a), Fertő-Han- 1929; Loksa 1966), Sopron Mts. (Traser 1981, ság (Traser 2002a), Gerecse Mts. (Traser 2006),

24 Dányi & Traser: Checklist of the springtail fauna of Hungary

Hortobágy (Loksa 1983), Sopron Mts. (Traser Protaphorura latosimilis (Loksa & Rubio, 1966) 2002a), Szigetköz (Traser 1997). Onychiurus (Protaphorura) latosimilis Loksa & Rubio: Loksa & Rubio 1966a: 149, figs 49-56. Protaphorura dudichi (Loksa & Rubio, 1966) General distribution: Hungary. Onychiurus (Protaphorura) dudichi Loksa & Rubio: Loksa & Rubio 1966a: 148, figs 41-48. Distribution in Hungary: Bakony Mts. (Loksa & Rubio 1966a). General distribution: Hungary. Distribution in Hungary: Bakony Mts. (Loksa Protaphorura metapulvinata (Loksa, 1964) & Rubio 1966a). Onychiurus (Protaphorura) pulvinatus metapulvinatus Loksa: Loksa 1964: 91, figs 34-37; Onychiurus pul- Protaphorura fimata (Gisin, 1952) vinatus metapulvinatus Loksa: Loksa 1966: 380; Ony- chiurus pulvinatus Gisin: Loksa 1966: 90. Onychiurus (Protaphorura) fimatus Gisin: Loksa 1964: 93, figs 42-44; Loksa & Rubio 1966a: 146; Onychiurus fimatus Gisin: Loksa 1966: 380. General distribution: Hungary. Distribution in Hungary: Keszthely Mts. (Lok- General distribution: Europe. sa 1964, 1966), Mecsek Mts. (Loksa 1964, 1966), Distribution in Hungary: Bakony Mts. (Loksa 1964, 1966; Loksa & Rubio 1966a), Balatonfelvi- Villány Mts (Loksa 1964, 1966). dék (Loksa 1966), Buda Mts. (Loksa 1964, 1966), Gerecse Mts. (Loksa 1964, 1966), Keszthely Mts. Protaphorura nemorata (Gisin, 1952) (Loksa 1964, 1966), Mecsek Mts. (Loksa 1964, Protaphorura nemorata (Gisin): Traser 2002a: 262. 1966), Mt. Somló (Loksa 1966), Vértes Mts. (Loksa 1964, 1966), Villány Mts (Loksa 1964, 1966). General distribution: Western Europe. Distribution in Hungary: Sopron Mts. (Traser Protaphorura gisini (Haybach, 1960) 2002a).

Protaphorura gisini (Haybach): Traser 2002c: 48; Protaphorura octopunctata (Tullberg, 1876) Traser & Horváth-Szováti 2006: 332. Onychiurus octopunctatus (Tullberg): Loksa 1983: 73; General distribution: Central Europe. Protaphorura cf. octopunctatus (Tullberg): Traser 1999: 53. Distribution in Hungary: Kiskunság (Traser 2002c; Traser & Horváth-Szováti 2006). General distribution: Europe, Siberia. Distribution in Hungary: Aggtelek Karst (Tra- Protaphorura illaborata (Gisin, 1952) ser 1999), Hortobágy (Loksa 1983). Protaphorura illaborata (Gisin): Traser 2002a: 262. Protaphorura procampata (Gisin, 1956) General distribution: Central Europe, Alps. Distribution in Hungary: Sopron Mts. (Traser Protaphorura procampatus (sic!) (Gisin): Traser 2002a). 2002a: 262; Onychiurus (Protaphorura) procampatus Gisin: Loksa & Rubio 1966a: 146, figs 21-28.

Protaphorura kadici (Loksa, 1967) General distribution: Palaearctic. Onychiurus (Protaphorura) kadici Loksa: Loksa 1967: Distribution in Hungary: Fertő-Hanság (Traser 290, figs 4-13; Onychiurus (Protaphorura) kadici var. 2002a), Bakony Mts. (Loksa & Rubio 1966a). geminiocellatus Loksa: Loksa 1967: 292; Onychiurus kadici Loksa: Traser 1999: 49. General distribution: Hungary. Protaphorura prolata (Gisin, 1956) Distribution in Hungary: Aggtelek Karst (Lok- Protaphorura prolatus (sic!) (Gisin): Traser 2002a: 262; sa 1967; Traser 1999). Traser & Salló 2004: 230.

25 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Europe. Rubio 1966a: 146, 35-40; Onychiurus sublatus Gisin: Distribution in Hungary: Fertő-Hanság (Traser Traser 1994: 100; Traser 1995: 44. 2002a), Kőszeg Mts. (Traser & Salló 2004). General distribution: Central Europe. Ecological note: troglophilous. Distribution in Hungary: Bakony Mts. (Loksa

& Rubio 1966a), Fertő-Hanság (Traser 2002a), Protaphorura pseudostyriaca (Loksa, 1964) Őrség (Traser 1995), Sopron Mts. (Traser 1994, Onychiurus (Protaphorura) pseudostyriacus Loksa: 2002a). Loksa 1964: 88, figs 24-33; Onychiurus pseudosty- riacus Loksa: Loksa 1966: 381. Protaphorura subnemorata (Gisin, 1957) General distribution: Hungary. Onychiurus subnemoratus Gisin: Traser 1994: 100; Distribution in Hungary: Buda Mts. (Loksa Traser 1995: 44; Protaphorura subnemorata (Gisin): 1964, 1966), Pilis Mts. (Loksa 1964, 1966). Traser 2002a: 263; Protaphorura subnemoratus (sic!) (Gisin): Traser & Salló 2004: 230. Protaphorura serbica (Loksa & Bogojevic, 1967) General distribution: Central Europe. Distribution in Hungary: Kőszeg Mts. (Traser Protaphorura serbica (Loksa & Bogojevic): Traser 2002c: 48; Traser & Horváth–Szováti 2004: 59; Traser & Salló 2004), Őrség (Traser 1995), Sopron Mts. & Horváth-Szováti 2006: 332. (Traser 1994, 2002a). Ecological note: in forests and bogs. General distribution: Central Europe.

Distribution in Hungary: Kiskunság (Traser Protaphorura tricampata (Gisin, 1956) 2002c; Traser & Horváth–Szováti 2004, 2006). Ecological note: xerothermophilic. Onychiurus (Protaphorura) tricampatus Gisin: Loksa & Rubio 1966a: 146, figs. 29-34; Onychiurus tri- campatus Gisin: Bajomi 1969: 243; Protaphorura Protaphorura subarmata (Gisin, 1957) tricampata (Gisin): Traser 2002a: 263. Protaphorura subarmatus (sic!) (Gisin): Traser 1999: 53; Traser 2002a: 262; Protaphorura subarmata (Gi- General distribution: Eastern Europe. sin): Traser 2006: 35. Distribution in Hungary: Aggtelek Karst (Ba- General distribution: Europe. jomi 1969), Bakony Mts. (Loksa & Rubio 1966a), Distribution in Hungary: Aggtelek Karst (Tra- Vas-Sopron Plain (Traser 2002a). ser 1999), Fertő-Hanság (Traser 2002a), Gerecse Mts. (Traser 2006), Szigetköz (Traser 2002a). Supraphorura Stach, 1954

Protaphorura subcancellata (Gisin, 1963) Supraphorura furcifera (Börner, 1901) Onychiurus (Protaphorura) subcancellatus Gisin: Lok- Supraphorura furcifera (Börner): Traser 1999: 53; sa 1964: 92, figs 38-41; Loksa & Rubio 1966a: 146; Traser 2002a: 262; Traser et al. 2006: 72. Onychiurus subcancellatus Gisin: Loksa 1966: 380; Csuták 1977: 68; Loksa 1978b: 95; Loksa 1980: 110. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- General distribution: Europe. ser 1999), Fertő-Hanság (Traser et al. 2006), Szi- Distribution in Hungary: Bakony Mts. (Loksa getköz (Traser 2002a). & Rubio 1966a), Cserhát (Csuták 1977; Loksa

1978b), Gerecse Mts. (Loksa 1964, 1966), Mátra Subfamily: Tetrodontophorinae Mts. (Loksa 1980), Vértes Mts. (Loksa 1964, 1966). Tetrodontophora Reuter, 1882

Protaphorura sublata (Gisin, 1957) Tetrodontophora bielanensis (Waga, 1842) Protaphorura sublata (Gisin): Traser 2002a: 263; Tetrodontophora bielanensis (Waga): Stach 1929: 283; Onychiurus (Protaphorura) sublatus Gisin: Loksa & Traser 1999: 52; Kontschán et al. 2003: 108.

26 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Central Europe, Balkan Mesaphorura betschi Rusek, 1979 Peninsula. Mesaphorura betschi Rusek: Traser 1997: 59; Traser Distribution in Hungary: Aggtelek Karst 2002a: 263. (Stach 1929; Traser 1999), Zemplén Mts. (Kon- tschán et al. 2003). General distribution: Western Europe. Ecological note: in montane habitats. Distribution in Hungary: Szigetköz (Traser 1997, 2002a). Family: Tullbergiidae Ecological note: silvicolous.

Subfamily: Tullbergiinae Mesaphorura critica Ellis, 1976 Doutnacia Rusek, 1974 Mesaphorura critica Ellis: Christian & Thibaud 1988: 233; Traser 1999: 54; Traser & Weiner 1999: 112; Doutnacia mols Fjellberg, 1998 Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; Traser 2002a: 263; Traser 2002b: 160; Traser 2002c: Doutnacia mols Fjellberg: Traser 2002b: 160; Traser 48; Traser 2003: 155; Traser & Horváth–Szováti 2004: 2002c: 48; Traser & Horváth-Szováti 2006: 332. 60; Traser 2006: 35; Traser & Horváth-Szováti 2006: 332; Traser et al. 2006: 72. General distribution: Central Europe. Distribution in Hungary: Bükk Mts. (Traser General distribution: Europe. 2002b), Kiskunság (Traser 2002c; Traser & Hor- Distribution in Hungary: Aggtelek Karst (Tra- váth-Szováti 2006). ser 1999), Bükk Mts. (Traser 2002b), Dráva Plain Ecological note: xerophilous. (Christian & Thibaud 1988), Fertő-Hanság (Tra- ser 2002a, 2003; Traser et al. 2006), Gerecse Mts. Doutnacia xerophila Rusek, 1974 (Traser 2006), Kiskunság (Traser & Csóka 2000, Doutnacia xerophila Rusek: Traser 1997: 59; Traser & 2001; Traser 2002b, 2002c; Traser & Horváth– Weiner 1999: 112; Traser & Csóka 2000: 76; Traser & Szováti 2004, 2006), Mezőföld (Christian & Csóka 2001: 233; Traser 2002a: 263; Traser 2002c: 48; Thibaud 1988), Sopron Mts. (Traser 2002b), Szi- Traser 2006: 35; Traser & Horváth-Szováti 2006: 332; getköz (Traser & Weiner 1999; Traser 2002a). Traser et al. 2006: 72. Ecological note: xerothermophilic.

General distribution: Central and Southern Eu- Mesaphorura delamarei Weiner, 1991 rope. Distribution in Hungary: Fertő-Hanság (Traser Mesaphorura delamarei Weiner: Traser 2003: 155. 2002a; Traser et al. 2006), Gerecse Mts. (Traser 2006), Kiskunság (Traser & Csóka 2000, 2001; General distribution: Eastern Europe. Traser 2002c; Traser & Horváth-Szováti 2006), Distribution in Hungary: Fertő-Hanság (Traser Szigetköz (Traser 1997; Traser & Weiner 1999). 2003). Ecological note: xerophilous. Ecological note: detritivorous.

Jevania Rusek, 1978 Mesaphorura hylophila Rusek, 1982 Mesaphorura hylophila Rusek: Traser 2002a: 263; Jevania weinerae Rusek, 1978 Traser 2006: 35; Traser & Horváth-Szováti 2006: 332; Jevania weinerae Rusek: Traser 2006: 35, figs 1-3. Traser et al. 2006: 72.

General distribution: Eastern Europe. General distribution: Palaearctic. Distribution in Hungary: Gerecse Mts. (Traser Distribution in Hungary: Fertő-Hanság (Traser 2006). 2002a; Traser et al. 2006), Gerecse Mts. (Traser Ecological note: silvicolous. 2006), Kiskunság (Traser & Horváth-Szováti 2006), Sopron Mts. (Traser 2002a). Mesaphorura Börner, 1901 Ecological note: eurytopic.

27 Dányi & Traser: Checklist of the springtail fauna of Hungary

Mesaphorura italica (Rusek, 1971) 35; Traser & Horváth-Szováti 2006: 332; Traser et al. 2006: 72. Mesaphorura italica (Rusek): Traser 2002a: 263. General distribution: cosmopolitan. General distribution: Palaearctic, Iraq. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Sopron Mts. (Traser ser 1999), Dráva Plain (Christian & Thibaud 2002a). 1988), Fertő-Hanság (Traser 2002a, 2003; Traser Ecological note: eurytopic. et al. 2006), Gerecse Mts. (Traser 2006), Kiskun- ság (Traser & Csóka 2000, 2001; Traser 2002c; Mesaphorura krausbaueri Börner, 1901 Traser & Horváth–Szováti 2004, 2006), Mezőföld Tullbergia krausbaueri (Börner): Loksa 1956a: 391; (Christian & Thibaud 1988), Sopron Mts. (Traser Loksa 1956b: 203; Loksa 1961b: 224; Loksa 1962: 73; 2002a), Szigetköz (Traser 1997; Traser & Weiner Loksa 1966: 381; Loksa 1977: 207; Loksa 1978a: 56; 1999). Loksa 1978b: 95; Loksa 1980: 110; Traser 1980: 20; Traser 1981: 133; Loksa 1987: 79; Loksa 1991: 273; Ecological note: eurytopic. Tullbergia (Tullbergia) krausbaueri (Börner): Loksa & Rubio 1966a: 151; Tullbergia (Mesaphorura) kraus- Mesaphorura redondoi Jordana & Arbea, 1994 baueri Börner: Dunger 1975: 19; (sic!) krausbaueri Börner: Traser 1994: 102; Mesaphorura Mesaphorura redondoi Arbea & Jordana (sic!): Traser krausbaueri Börner: Traser 1997: 61; Traser 1999: 54; 1997: 59; Traser 2002c: 48. Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; Traser 2002a: 263; Traser 2002a: 263; Traser 2002c: 48; Traser 2002c: 48; Traser 2002c: 48; Traser 2003: Remarks: Jordana et al. (1997) considered this 155; Traser 2006: 35; Traser & Horváth-Szováti 2006: taxon as a subspecies of Mesaphorura isochaeta 332; Traser et al. 2006: 72. Arbea & Jordana, 1990, endemic of the Iberian Peninsula. However, this status is questionable in General distribution: cosmopolitan. the light of the Hungarian occurences. Distribution in Hungary: Aggtelek Karst (Lok- General distribution: ?Europe. sa 1961b; Traser 1999), Bakony Mts. (Loksa Distribution in Hungary: Kiskunság (Traser 1966; Loksa & Rubio 1966a; Traser 1980), Bala- tonfelvidék (Loksa 1966), Börzsöny Mts. (Loksa 2002c), Szigetköz (Traser 1997). 1966; Dunger 1975), Buda Mts. (Loksa 1956b, 1966), Bükk Mts. (Loksa 1956a, 1962, 1966, Mesaphorura rudolfi Rusek, 1987 1977), Cserhát (Loksa 1978b), Dráva Plain (Lok- sa 1978a), Fertő-Hanság (Traser 2002a, 2003; Mesaphorura rudolfi Rusek: Traser 2002a: 263. Traser et al. 2006), Gerecse Mts. (Loksa 1966; Traser 2006), Keszthely Mts. (Loksa 1966), Kis- General distribution: Central Europe. kunság (Loksa 1987; Traser & Csóka 2000, 2001; Distribution in Hungary: Fertő-Hanság (Traser Traser 2002c; Traser & Horváth-Szováti 2006), 2002a). Mátra Mts. (Loksa 1980), Mecsek Mts. (Loksa 1966), Nyírség (Loksa 1991), Pilis Mts. (Loksa Mesaphorura sylvatica (Rusek, 1971) 1966), Sopron Mts. (Traser 1981, 1994), Sziget- köz (Traser 1997, 2002a), Vas-Sopron Plain (Tra- Tullbergia (Mesaphorura) sylvatica Rusek: Dunger ser 1981, 1981), Vértes Mts. (Loksa 1966), Vi- 1975: 19; Mesaphorura sylvatica (Rusek): Traser lány Mts (Loksa 1966), Visegrád Mts. (Loksa 1999: 54; Traser 2002a: 263; Traser 2006: 35. 1966). Ecological note: eurytopic. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- Mesaphorura macrochaeta Rusek, 1976 ser 1999), Börzsöny Mts. (Dunger 1975), Gerecse Mts. (Traser 2006), Szigetköz (Traser 2002a). Mesaphorura macrochaeta Rusek: Traser 1997: 61; Christian & Thibaud 1988: 233; Christian & Thibaud 1988: 233; Traser 1999: 54; Traser & Weiner 1999: Mesaphorura tenuisensillata Rusek, 1974 112; Traser & Csóka 2000: 77; Traser & Csóka 2001: 237; Traser 2002a: 263; Traser 2002c: 48; Traser 2003: Mesaphorura tenuisensillata Rusek: Traser 1997: 59; 155; Traser & Horváth–Szováti 2004: 58; Traser 2006: Traser 2002a: 263.

28 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Palaearctic. Neonaphorura Bagnall, 1935 Distribution in Hungary: Sopron Mts. (Traser 2002a), Szigetköz (Traser 1997). Neonaphorura loksai Traser & Thibaud, 1999 Neonaphorura loksai Traser & Thibaud: Traser & Mesaphorura yosii (Rusek, 1967) Thibaud 1999: 105, figs 1-8; Traser 2002a: 263. Mesaphorura yosii (Rusek): Traser 1997: 59; Traser 2002a: 263. General distribution: Hungary. Distribution in Hungary: Vas-Sopron Plain General distribution: cosmopolitan. (Traser & Thibaud 1999; Traser 2002a). Ecological note: silvicolous, mesophilic. Distribution in Hungary: Szigetköz (Traser

1997, 2002a). Bagnall, 1935

Metaphorura Bagnall, 1936 Neotullbergia ramicuspis (Gisin, 1953)

Metaphorura affinis (Börner, 1902) Neotullbergia ramicuspis (Gisin): Traser & Weiner 1999: 112; Traser 2002a: 263. Tullbergia affinis Börner: Loksa 1966: 381; Loksa 1983: 73; Varga & Oldal 1998: 86; Metaphorura affinis (Börner): Traser 1999: 54; Traser & Weiner General distribution: Western Europe. 1999: 112; Traser 2002a: 263; Traser 2002c: 48; Traser Distribution in Hungary: Fertő-Hanság (Traser 2006: 35; Traser & Horváth-Szováti 2006: 332; Traser 2002a), Szigetköz (Traser & Weiner 1999). et al. 2006: 72. Ecological note: xerothermophilic.

General distribution: Palaearctic. Paratullbergia Womersley, 1930 Distribution in Hungary: Aggtelek Karst (Lok- sa 1966; Traser 1999), Bakony Mts. (Loksa Paratullbergia callipygos (Börner, 1902) 1966), Balatonfelvidék (Loksa 1966), Börzsöny Paratullbergia callipygos (Börner): Traser & Thibaud Mts. (Loksa 1966), Buda Mts. (Loksa 1966), 1999: 106; Traser 2002a: 263. Bükk Mts. (Loksa 1966; Varga & Oldal 1998), Fertő-Hanság (Traser 2002a; Traser et al. 2006), General distribution: Holarctic. Gerecse Mts. (Loksa 1966; Traser 2006), Hor- Distribution in Hungary: Sopron Mts. (Traser tobágy (Loksa 1983), Keszthely Mts. (Loksa 2002a), Vas-Sopron Plain (Traser & Thibaud 1966), Kiskunság (Traser 2002c; Traser & Hor- 1999). váth-Szováti 2006), Mecsek Mts. (Loksa 1966), Ecological note: silvicolous. Mt. Somló (Loksa 1966), Pilis Mts. (Loksa 1966), Szigetköz (Traser & Weiner 1999), Vértes Mts. Paratullbergia macdougalli Bagnall, 1936 (Loksa 1966), Villány Mts (Loksa 1966), Vi- Paratullbergia macdougalli Bagnall: Traser 2002a: segrád Mts. (Loksa 1966). 263; Traser 2003: 155. Ecological note: xerothermophilic. General distribution: Palaearctic. Metaphorura riozoi Castaño-Meneses, Palacios Distribution in Hungary: Fertő-Hanság (Traser Vargas & Traser, 2000 2002a, 2003).

Metaphorura riozoi Castaño-Meneses, Palacios Vargas & Traser: Castaño-Meneses et al. 2000: 1263, figs 1- Scaphaphorura Petersen, 1965 13; Traser 2002a: 263. Scaphaphorura arenaria (Petersen, 1965) General distribution: Central Europe. Scaphaphorura arenaria (Petersen): Traser 2002c: 48. Distribution in Hungary: Fertő-Hanság (Casta- ño-Meneses et al. 2000; Traser 2002a). General distribution: Palaearctic.

29 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Kiskunság (Traser 71; Loksa 1987: 78; Loksa 1991: 270; Farkas 1995: 46; 2002c). Farkas 1996: 287; Farkas & Szathmáry 1997: 39; Farkas 1997: 58; Traser 1997: 62; Farkas et al. 1998: Ecological note: arenicol, in sand dunes. 57; Traser 2002a: 266.

Stenaphorurella Lucianez & Simon, 1992 General distribution: Holarctic. Distribution in Hungary: Balatonfelvidék Stenaphorurella lubbocki (Bagnall, 1935) (Stach 1929; Szent-Ivány 1938, 1940a; Farkas Stenaphorurella lubbocki (Bagnall): Traser 1999: 54; 1995, 1997; Farkas & Szathmáry 1997), Borsodi- Traser 2002a: 263. Mezőség (Szent-Ivány 1938), Borsodi-Mezőség (Szent-Ivány 1940a), Börzsöny Mts. (Dunger General distribution: Western Europe. 1975), Buda Mts. (Dudich 1947), Budapest (Tö- Distribution in Hungary: Aggtelek Karst (Tra- mösváry 1884a; Vellay 1900; Stach 1929; Szent- ser 1999), Sopron Mts. (Traser 2002a). Ivány 1938, 1940a), Bükk Mts. (Herman 1865), Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser Stenaphorurella metaparisi Traser & Weiner, 1999 2002a), Hortobágy (Loksa 1983), Kis-Balaton Stenaphorurella metaparisi Traser & Weiner: Traser & (Farkas 1996, 1997), Kiskunság (Szent-Ivány Weiner 1999: 109, figs 1-5; Traser 2002a: 263. 1938, 1940a; Loksa 1987), Kőszeg Mts. (Szent- Ivány 1940a), Mátra Mts. (Loksa 1980), Nyírség General distribution: Hungary. (Horváth 1888; Kovács 1953a; Loksa 1991), Pilis Distribution in Hungary: Szigetköz (Traser & Mts. (Stach 1929; Szent-Ivány 1938, 1940a), So- Weiner 1999; Traser 2002a). mogy Hills (Farkas et al. 1998), Sopron Mts.

(Traser 2002a), Szigetköz (Traser 1997), Vác-Pest Stenaphorurella quadrispina (Börner, 1901) Duna Valley (Vellay 1900; Stach 1929; Szent- Tullbergia quadrispina Börner: Loksa 1966: 381; Ivány 1938, 1940a). Loksa 1978b: 95; Farkas 1995: 46; Farkas et al. 1998: 57; Tullbergia (Stenaphorura) quadrispina Börner: Ecological note: hygrophilous. Loksa & Rubio 1966a: 151; Dunger 1975: 19; Ste- naphorurella quadrispina (Börner): Traser 1999: 54; Suborder: Traser & Weiner 1999: 112; Traser 2002a: 263. Family: Cyphoderidae General distribution: Mediterranean.

Distribution in Hungary: Aggtelek Karst (Lok- Cyphoderus Nicolet, 1842 sa 1966; Traser 1999), around the Lake Balaton

(Farkas 1995), Bakony Mts. (Loksa & Rubio Cyphoderus albinus Nicolet, 1842 1966a), Börzsöny Mts. (Dunger 1975), Bükk Mts. (Loksa 1966), Cserhát (Loksa 1978b), Fertő-Han- Beckia albinos (Nicolet): Tömösváry 1884a: 51; Beckia argentea (Lubbock): Tömösváry 1884a: 51; ság (Traser 2002a), Gerecse Mts. (Loksa 1966), Cyphoderus argenteus Lubbock: Vellay 1900: 21; Somogy Hills (Farkas et al. 1998), Szigetköz Cyphoderus albinus Nicolet: Stach 1929: 294; Loksa (Traser & Weiner 1999). 1966: 385; Dunger 1975: 24; Loksa 1978a: 61; Loksa 1983: 75; Hornung 1986: 138; Loksa 1987: 79; Traser 1999: 57; Traser 2002a: 266; Traser 2002c: 49; Traser Family: Poduridae & Horváth-Szováti 2006: 334; Traser et al. 2006: 73. Podura Linnaeus, 1758 General distribution: cosmpolitan. Podura aquatica Linnaeus, 1758 Distribution in Hungary: Aggtelek Karst ?Podura aquatica Linnaeus: Herman 1865: 485; (Stach 1929; Traser 1999), Bakony Mts. (Loksa Podura aquatica Linnaeus: Tömösváry 1884a: 51; 1966), Balatonfelvidék (Loksa 1966), Börzsöny Horváth 1888: 164; Vellay 1900: 21; Stach 1929: 279; Mts. (Dunger 1975), Buda Mts. (Loksa 1966), Szent-Ivány 1938: 91; Szent-Ivány 1940a: 427; Dudich Budapest (Tömösváry 1884a; Vellay 1900; Stach 1947: 370, fig. 1; Kovács 1953a: 183; Dunger 1975: 1929), Dráva Plain (Loksa 1978a), Fertő-Hanság 14; Loksa 1978a: 53; Loksa 1980: 110; Loksa 1983: (Traser 2002a; Traser et al. 2006), Gerecse Mts.

30 Dányi & Traser: Checklist of the springtail fauna of Hungary

(Loksa 1966), Hortobágy (Loksa 1983), Keszt- General distribution: cosmopolitan. hely Mts. (Loksa 1966), Kiskunság (Vellay 1900; Distribution in Hungary: Balatonfelvidék Stach 1929; Hornung 1986; Loksa 1987; Traser (Stach 1929; Geyer & Mann 1940; Loksa 1960; 2002c; Traser & Horváth-Szováti 2006), Mecsek Mts. (Loksa 1966), Mt. Somló (Loksa 1966), Pilis Paclt 1960), Fertő-Hanság (Traser 2002a), Vas- Mts. (Loksa 1966), Vác-Pest Duna Valley (Tö- Sopron Plain (Traser 1981). mösváry 1884a; Vellay 1900; Stach 1929), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966). Entomobrya Rondani, 1861 Ecological note: myrmecophilous. Entomobrya albanica Stach, 1922 Cyphoderus bidenticulatus Parona, 1888 Entomobrya albanica Stach: Stach 1963: 77. Cyphoderus bidenticulatus Parona: Paclt 1960: 157; Traser 1997: 62; Traser 2002a: 266; Traser 2002c: 49; Remark: Species inquirenda according to Gisin Traser & Horváth-Szováti 2006: 334. (1960). General distribution: Mediterranean. General distribution: Southeastern Europe. Distribution in Hungary: Balatonfelvidék Distribution in Hungary: Mezőföld (Stach (Paclt 1960), Kiskunság (Traser 2002c; Traser & 1963). Horváth-Szováti 2006), Moson Plain (Traser 2002a), Szigetköz (Traser 1997). Entomobrya albocincta (Templeton, 1835) Ecological note: thermophilic. Degeeria cincta Lubbock: Tömösváry 1884a: 52; Degeeria cincta Lubbock var. nigrocincta Tömösváry: Cyphoderus reuteri Loksa, 1978 Tömösváry 1884a: 52; Entomobrya albocincta Tem- pleton (sic!): Vellay 1900: 20; Vellay 1900: 20; Vellay Cyphoderus reuteri Loksa: Loksa 1978a: 61, figs 9-11. 1900: 20; Stach 1929: 304; Stach 1929: 304; Stach 1929: 304; Entomobrya albocincta var. nigrocincta Tömösváry: Vellay 1900: 20; Stach 1929: 304. General distribution: Central Europe. Distribution in Hungary: Dráva Plain (Loksa General distribution: Western Europe. 1978a). Distribution in Hungary: Budapest (Tömös- váry 1884a; Vellay 1900; Stach 1929), Gödöllő Family: Hills (Tömösváry 1884a; Vellay 1900; Stach Subfamily: Entomobryinae 1929), Vác-Pest Duna Valley (Tömösváry 1884a; Vellay 1900; Stach 1929).

Ecological note: in mulder. Coecobrya Yosii, 1956

Coecobrya magyari (Chen, Wang & Christian- Entomobrya arborea (Tullberg, 1871) sen, 2002) Entomobrya arborea (Tullberg): Stach 1929: 304; Stach 1963: 47. Sinella (Coecobrya) magyari Chen, Wang & Chris- tiansen: Chen et al. 2002: 43, figs 1-18. General distribution: Europe. Distribution in Hungary: Mezőföld (Stach General distribution: Hungary. 1929; Stach 1963). Distribution in Hungary: Gödöllő (Chen et al. Ecological note: thermophilic, on barks and in 2002). mulder. Ecological note: in the soil of indoor plants. Entomobrya atrocincta Schött, 1897 Coecobrya tenebricosa Folsom, 1902 Entomobrya nigrocincta Denis: Stach 1929: 301; Sinella coeca Schött: Stach 1929: 300; Geyer & Mann Entomobrya atrocincta Schött: Stach 1963: 72. 1940: 371; Loksa 1960: 49; Traser 1981: 134; Traser 2002a: 265; Sinella höfti Schäffer: Paclt 1960: 157. General distribution: cosmopolitan.

31 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Mezőföld (Stach 2006), Hortobágy (Loksa 1983), Kiskunság (Lok- 1929, 1963). sa 1987), Mezőföld (Stach 1929, 1963; Dombos Ecological note: thermophilic. 2001), Somogy Hills (Farkas et al. 1998). Ecological note: xerothermophilic. Entomobrya corticalis (Nicolet, 1842) Entomobrya lanuginosa (Nicolet, 1841) Entomobrya cortocalis (Nicolet): Szent-Ivány 1940a: 431; Loksa & Rubio 1966a: 152; Loksa 1978b: 95; Degeeria lanuginosa Nicolet: Tömösváry 1884a: 52;: Traser 1995: 45; Traser 2002a: 265; Traser 2003: 155; 52;: 52; Entomobrya lanuginosa (Nicolet): Vellay Traser & Salló 2004: 230; Traser et al. 2006: 73. 1900: 20; Stach 1929: 303; Traser 1980: 20; Traser 1981: 133; Loksa 1983: 73; Farkas 1995: 46; Traser General distribution: Europe. 1995: 45; Farkas 1996: 287; Farkas 1997: 58; Farkas et al. 1998: 57; Traser 1999: 56; Traser 2002a: 265; Distribution in Hungary: Bakony Mts. (Loksa Traser 2002c: 48; Traser 2003: 155; Traser & Horváth- & Rubio 1966a), Cserhát (Loksa 1978b), Fertő- Szováti 2006: 333; Traser 2006: 36; Entomobrya Hanság (Traser 2003; Traser et al. 2006), Kőszeg lanuginose (sic!) (Nicolet): Dombos 2001: 2040. Mts. (Szent-Ivány 1940a; Traser & Salló 2004), Őrség (Traser 1995), Sopron Mts. (Traser 2002a). General distribution: Palaearctic. Ecological note: corticophilous. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), around the Lake Balaton (Farkas 1995), Bakony Mts. (Traser 1980), Fertő-Hanság (Traser Entomobrya dorsalis Uzel, 1891 2002a, 2003), Gerecse Mts. (Traser 2006), Horto- Entomobrya dorsalis Uzel: Vellay 1900: 20; Stach bágy (Loksa 1983), Kis-Balaton (Farkas 1996, 1929: 304; Szent-Ivány 1940a: 431; Stach 1963: 99; 1997), Kiskunság (Traser 2002c; Traser & Hor- Eutomobrya puncteola Uzel: Stach 1929: 304; Szent- váth-Szováti 2006), Mecsek Mts. (Tömösváry Ivány 1940a: 431; Loksa 1978a: 60; Traser 1994: 100; 1884a; Vellay 1900; Stach 1929), Mezőföld Traser 1999: 56. (Dombos 2001), Őrség (Traser 1995), Somogy

Hills (Farkas et al. 1998), Sopron Mts. (Traser General distribution: Eastern Europe. 1981, 2002a), Vác-Pest Duna Valley (Tömösváry Distribution in Hungary: Aggtelek Karst (Tra- 1884a; Vellay 1900; Stach 1929), Vas-Sopron ser 1999), Cserhát (Stach 1929, 1963), Dráva Plain (Traser 1981). Plain (Loksa 1978a) Kőszeg Mts. (Szent-Ivány Ecological note: on meadows. 1940a), Mezőföld (Stach 1929; Szent-Ivány 1940a), Nyírség (Vellay 1900), Sopron Mts. (Tra- Entomobrya marginata (Tullberg, 1871) ser 1994). Ecological note: atmobiont, corticophilous. ?Entomobrya maritima Reuter: Vellay 1900: 21; Entomobrya marginata Tullberg: Vellay 1900: 21; Szent-Ivány 1940a: 431; Stach 1929: 300; Stach 1963: Entomobrya handschini Stach, 1922 60; Loksa 1978b: 95; Loksa 1980: 111; Loksa 1983: 74; Loksa 1991: 272; Farkas 1995: 46; Farkas 1997: Entomobrya handschini Stach: Stach 1929: 301; Szent- 56; Farkas et al. 1998: 57; Traser 1999: 56; Traser Ivány 1941: 22; Stach 1963: 50; Loksa & Rubio 1966a: 2002a: 265. 152; Dunger 1975: 21; Loksa 1983: 74; Loksa 1987: 79; Farkas et al. 1998: 57; Traser 1999: 56; Dombos Remark: Vellay’s data (Vellay 1900) were cri- 2001: 2043; Traser et al. 2006: 73; Entomobrya hand- schini ab. aethiopella Stach: Stach 1929: 302; tised by Stach (1929). Entomobrya hanschini (sic!) Stach: Farkas 1995: 46. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- General distribution: Europe, Asia Minor. ser 1999), Balatonfelvidék (Stach 1929, 1963; Distribution in Hungary: Aggtelek Karst (Tra- Farkas 1995, 1997), Budapest (Stach 1929, 1963), ser 1999), around the Lake Balaton (Farkas 1995), Cserhát (Loksa 1978b), Fertő-Hanság (Traser Bakony Mts. (Loksa & Rubio 1966a), Börzsöny 2002a), Hortobágy (Loksa 1983), Kiskunság Mts. (Dunger 1975), Buda Mts. (Stach 1929; (Vellay 1900; Stach 1929), Kőszeg Mts. (Szent- Szent-Ivány 1941), Fertő-Hanság (Traser et al. Ivány 1940a), Mátra Mts. (Loksa 1980), Mező-

32 Dányi & Traser: Checklist of the springtail fauna of Hungary

föld (Stach 1929), Nyírség (Loksa 1991), Somogy 85; Varga & Vargha 1992: 315; Traser 1995: 45; Hills (Farkas et al. 1998). Traser 1999: 56; Traser 2002a: 265; Traser 2003: 155; Traser & Salló 2004: 230; Entomobrya muscorum var. Ecological note: xerophilous, corticophilous. elongata (Brook): Stach 1929: 303; Entomobrya mus- corum f. elongata Brook: Szent-Ivány 1940a: 430; Entomobrya multifasciata (Tullberg, 1871) Szent-Ivány 1941: 21.

Entomobrya multifasciata (Tullberg): Vellay 1900: 21; Stach General distribution: Palaearctic. 1963: 34; Loksa 1966: 382; Loksa & Rubio 1966a: 152; Dunger 1975: 21; Loksa 1978a: 60; Loksa 1978b: Distribution in Hungary: Aggtelek Karst (Tra- 95; Loksa 1980: 111; Loksa 1983: 74; Loksa 1987: 79; ser 1999), Bakony Mts. (Loksa & Rubio 1966a), Varga 1992: 172; Varga & Vargha 1992: 315; Farkas Börzsöny Mts. (Dunger 1975), Buda Mts. (Stach et al. 1998: 57; Varga & Oldal 1998: 86; Traser 1999: 1929; Loksa 1956b), Budapest (Szent-Ivány 56; Traser & Csóka 2000: 76; Traser & Csóka 2001: 1940a, 1940b, 1941; Stach 1963), Bükk Mts. 233; Traser 2002a: 265; Traser 2002b: 160; Traser (Varga 1991; Varga & Vargha 1992), Cserehát 2002c: 48; Traser 2003: 155; Traser & Horváth– (Szent-Ivány 1941), Cserhát (Loksa 1978b), Fer- Szováti 2004: 58; Traser & Horváth-Szováti 2006: tő-Hanság (Paclt 1960; Traser 2002a, 2003), 333; Traser et al. 2006: 73; Entomobrya nicoleti var. Gerecse Mts. (Loksa 1966), Kiskunság (Vellay multifasciata (Tullberg): Stach 1929: 301. 1900; Stach 1929), Kőszeg Mts. (Szent-Ivány 1940a, 1940b, 1941; Traser & Salló 2004), Mátra General distribution: cosmopolitan. Mts. (Szent-Ivány 1940b, 1941; Loksa 1980), Me- Distribution in Hungary: Aggtelek Karst (Lok- zőföld (Stach 1963), Őrség (Traser 1995), Vác- sa 1966; Traser 1999), Bakony Mts. (Loksa 1966; Pest Duna Valley (Tömösváry 1884a; Vellay Loksa & Rubio 1966a; Traser 2002b), Balaton- 1900; Stach 1963), Visegrád Mts. (Stach 1929). felvidék (Loksa 1966), Börzsöny Mts. (Loksa Ecological note: in dense vegetation. 1966; Dunger 1975), Buda Mts. (Loksa 1966), Bükk Mts. (Loksa 1966; Varga 1992; Varga & Entomobrya nicoleti (Lubbock, 1868) Vargha 1992; Varga & Oldal 1998; Traser Entomobrya nicoleti var. muscorum (Tullberg): Stach 2002b), Cserhát (Loksa 1978b), Dráva Plain 1929: 301; Entomobrya nicoleti var. fasciata Stach: Stach 1929: 301; Entomobrya nicoleti var. obscura (Loksa 1978a; Traser 2002b), Fertő-Hanság (Tra- (Tullberg): Stach 1929: 301. ser 2003; Traser et al. 2006), Gerecse Mts. (Loksa 1966), Hortobágy (Loksa 1983; Traser 2002b), General distribution: Europe. Keszthely Mts. (Loksa 1966), Kiskunság (Vellay Distribution in Hungary: Buda Mts. (Stach 1900; Loksa 1987; Traser & Csóka 2000, 2001; 1929), Mezőföld (Stach 1929). Traser 2002b, 2002c; Traser & Horváth–Szováti 2004, 2006), Mátra Mts. (Loksa 1980), Mecsek Entomobrya nigriventris Stach, 1930 Mts. (Loksa 1966), Mezőföld (Stach 1929, 1963), Entomobrya nigriventris Stach: Stach 1929: 302; Stach Mt. Somló (Loksa 1966), Nyírség (Traser 2002b), 1963: 41; Hornung 1986: 138; Loksa 1987: 79; Traser Pilis Mts. (Loksa 1966), Somogy Hills (Farkas et 2002c: 48; Traser & Horváth-Szováti 2006: 333; al. 1998), Sopron Mts. (Traser 2002a), Szigetköz Traser et al. 2006: 73. (Traser 2002a), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), Visegrád Mts. (Loksa 1966). General distribution: Europe. Ecological note: xerothermophilic. Distribution in Hungary: Fertő-Hanság (Traser et al. 2006), Kiskunság (Hornung 1986; Loksa Entomobrya muscorum (Nicolet, 1842) 1987; Traser 2002c; Traser & Horváth-Szováti 2006), Mezőföld (Stach 1929, 1963). Degeeria muscorum Nicolet: Tömösváry 1884a: 52; Entomobrya intermedia Brook: Vellay 1900: 20; Ecological note: xerothermophilic. Entomobrya intermedia var. elongata Brook: Vellay 1900: 20; Entomobrya muscorum (Nicolet): Vellay Entomobrya nivalis (Linnaeus, 1758) 1900: 21; Stach 1929: 303; Szent-Ivány 1940a: 430; Szent-Ivány 1940b: 119; Szent-Ivány 1941: 21; Loksa Degeeria nivalis Linnaeus: Tömösváry 1884a: 52;: 52; 1956b: 227; Paclt 1960: 157; Stach 1963: 95; Loksa Entomobrya nivalis (Linnaeus): Vellay 1900: 21; Stach 1966: 383; Loksa & Rubio 1966a: 152; Dunger 1975: 1929: 301; Szent-Ivány 1940a: 431; Stach 1963: 27; 21; Loksa 1978b: 95; Loksa 1980: 111; Varga 1991: Loksa & Rubio 1966a: 152; Dunger 1975: 21; Loksa

33 Dányi & Traser: Checklist of the springtail fauna of Hungary

1991: 272; Varga 1992: 172; Varga & Vargha 1992: ser 1995), Pilis Mts. (Loksa 1966), Szigetköz 315; Traser 1995: 45; Varga & Oldal 1998: 86; Traser (Traser 1997, 2002a), Villány Mts (Loksa 1966). 1999: 56; Traser 2002a: 265; Traser & Salló 2004: 230; Traser & Horváth-Szováti 2006: 333; Traser et al. Ecological note: xerothermophilic. 2006: 73; Entomobrya nivalis var. immaculata Schäf- fer: Stach 1929: 301; Entomobrya nivalis Nicolet Entomobrya schoetti Stach, 1922 (sic!): Loksa 1956b: 207. Entomobrya schötti (sic!) ab. pigmentata Stach: Szent- General distribution: cosmopolitan. Ivány 1941: 22. Distribution in Hungary: Aggtelek Karst General distribution: Eastern Europe. (Traser 1999), Bakony Mts. (Loksa & Rubio Distribution in Hungary: Tétény Plateau 1966a), Börzsöny Mts. (Dunger 1975), Buda Mts. (Szent-Ivány 1941). (Stach 1929; Loksa 1956b), Budapest (Tömösváry Ecological note: xerothermophilic. 1884a; Vellay 1900; Stach 1929, 1963), Bükk Mts. (Varga 1992; Varga & Vargha 1992; Varga Entomobrya spectabilis Reuter, 1890 & Oldal 1998), Fertő-Hanság (Traser et al. 2006), Entomobrya spectabilis Reuter: Loksa 1978a: 60. Gödöllő Hills (Tömösváry 1884a; Vellay 1900; Stach 1929), Kiskunság (Traser & Horváth- General distribution: Northern and Central Eu- Szováti 2006), Kőszeg Mts. (Szent-Ivány 1940a; rope. Traser & Salló 2004), Mátra Mts. (Stach 1929), Distribution in Hungary: Dráva Plain (Loksa Mezőföld (Stach 1929), Nyírség (Loksa 1991), 1978a). Őrség (Traser 1995), Szigetköz (Traser 2002a). Ecological note: synanthropic. Ecological note: atmobiont. Entomobrya superba (Reuter, 1876) Entomobrya pazaristei Denis, 1933 Calistella superba Reuter: Vellay 1900: 21; Entomobrya pazaristei Denis: Loksa & Rubio 1966a: Entomobrya superba (Reuter): Stach 1929: 304; Szent- 152. Ivány 1940a: 430.

General distribution: Balkan Peninsula. General distribution: Europe. Distribution in Hungary: Bakony Mts. (Loksa Distribution in Hungary: Kőszeg Mts. (Szent- & Rubio 1966a). Ivány 1940a), Nyírség (Vellay 1900; Stach 1929; Ecological note: troglophilous. Szent-Ivány 1940a). Ecological note: atmobiont. Entomobrya quinquelineata Börner, 1901 Entomobrya violaceolineata Stach, 1963 Entomobrya quinquelineata Börner: Stach 1929: 302; Loksa 1956a: 413; Stach 1963: 57; Loksa 1966: 383; Entomobrya violaceolineata Stach: Traser & Horváth- Loksa 1978a: 60; Loksa 1983: 74; Hornung 1986: 138; Szováti 2006: 333. Loksa 1987: 79; Traser 1995: 45; Traser 1997: 62; Traser 1999: 56; Traser 2002a: 265; Traser 2003: 155; General distribution: Eastern Europe. Entomobrya quinquelineata f. alineata: Traser 2002a: Distribution in Hungary: Kiskunság (Traser & 265; Entomobrya quinquelineata f. trilineata: Traser Horváth-Szováti 2006). 2002a: 265. Ecological note: in litter.

General distribution: Mediterranean. Entomobryoides Maynard, 1951 Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Bükk Mts. (Loksa 1956a), Cserhát Entomobryoides purpurascens (Packard, 1872) (Stach 1929, 1963), Dráva Plain (Loksa 1978a), Entomobrya purpurascens (Packard): Loksa 1966: Fertő-Hanság (Traser 2002a, 2003), Hortobágy 383; Loksa & Rubio 1966a: 152. (Loksa 1983), Kiskunság (Hornung 1986; Loksa 1987), Mezőföld (Stach 1929, 1963), Őrség (Tra- General distribution: Europe, North America.

34 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Bakony Mts. (Loksa 384; Loksa & Rubio 1966a: 152; Loksa 1977: 211; 1966; Loksa & Rubio 1966a), Balatonfelvidék Loksa 1978a: 61; Loksa 1980: 111; Loksa 1983: 74; (Loksa 1966), Buda Mts. (Loksa 1966), Gerecse Loksa 1991: 273; Farkas 1995: 46; Farkas 1996: 287; Mts. (Loksa 1966), Keszthely Mts. (Loksa 1966), Farkas 1997: 58; Farkas & Szathmáry 1997: 39; Traser 1999: 56; Lepidocyrtus curvicolis (sic!) Bourlet: Mecsek Mts. (Loksa 1966), Vértes Mts. (Loksa Dombos 2001: 2040. 1966), Villány Mts (Loksa 1966).

General distribution: Holarctic. Entomobryoides myrmecophilus (Reuter, 1886) Distribution in Hungary: Aggtelek Karst Sinella myrmecophila Reuter: Szent-Ivány 1940a: 430; (Stach 1929; Dudich 1932; Gebhardt 1934; Loksa Entomobryoides mayrmecophilus (sic!) (Reuter): Tra- 1966; Traser 1999), around the Lake Balaton ser 1999: 55; Entomobryoides myrmecophila (sic!) (Farkas 1995), Bakony Mts. (Loksa & Rubio (Reuter): Traser & Csóka 2000: 77; Traser & Csóka 1966a), Balatonfelvidék (Stach 1929; Gebhardt 2001: 233; Entomobryoides myrmecophilus (Reuter): 1934; Loksa 1960, 1961a; Farkas & Szathmáry Traser 2002c: 48. 1997), Börzsöny Mts. (Loksa 1966), Buda Mts. (Loksa 1966), Budapest (Tömösváry 1884a; Vel- General distribution: Europe. lay 1900; Stach 1929; Gebhardt 1934), Bükk Mts. Distribution in Hungary: Aggtelek Karst (Tra- (Loksa 1962, 1966, 1977), Dráva Plain (Loksa ser 1999), Kiskunság (Traser & Csóka 2000, 1978a), Gerecse Mts. (Loksa 1966), Gödöllő Hills 2001; Traser 2002c), Kőszeg Mts. (Szent-Ivány (Tömösváry 1884a; Vellay 1900; Stach 1929), 1940a). Hortobágy (Loksa 1983), Kis-Balaton (Farkas Ecological note: myrmecophilous. 1996, 1997), Kiskunság (Vellay 1900; Stach 1929; Gebhardt 1934), Mátra Mts. (Loksa 1980), Subfamily: Lepidocyrtinae Mecsek Mts. (Stach 1929; Gebhardt 1934; Paclt 1960), Mezőföld (Stach 1929; Gebhardt 1934; Lepidocyrtus Bourlet, 1839 Dombos 2001), Nyírség (Vellay 1900; Stach 1929; Loksa 1991), Pilis Mts. (Loksa 1966), Vác- Lepidocyrtus arrabonicus Traser, 2000 Pest Duna Valley (Tömösváry 1884a; Vellay Lepidocyrtus arrabonicus Traser: Traser 2000: 65, figs 1900; Stach 1929; Gebhardt 1934), Visegrád Mts. 1-14; Traser 2002a: 265; Traser 2002c: 48; Traser (Loksa 1966), Zemplén Mts. (Tömösváry 1884a; 2003: 156; Traser & Horváth-Szováti 2006: 333. Vellay 1900; Stach 1929). Ecological note: edaphic, troglophilous. General distribution: Central Europe. Distribution in Hungary: Fertő-Hanság (Traser Lepidocyrtus cyaneus Tullberg, 1871 2000, 2002a, 2003), Kiskunság (Traser & Hor- Lepidocyrtus purpureus Lubbock: Vellay 1900: 21; váth-Szováti 2006; Traser 2002c), Moson Plain Lepidocyrtus assimilis Reuter: Vellay 1900: 21; (Traser 2000). Lepidocyrtus cyaneus Tullberg: Stach 1929: 297; Ecological note: mesophilic. Szent-Ivány 1940a: 432; Szent-Ivány 1941: 22; Kovács 1953a: 184; Loksa 1956a: 411; Loksa 1966: 384; Loksa & Rubio 1966a: 152; Loksa 1978a: 61; Loksa Lepidocyrtus cf. pallidus Reuter, 1890 1978b: 95; Loksa 1980: 111; Traser 1981: 135; Loksa Lepidocyrtus cf. pallidus Reuter 1890: Dunger 1975: 1983: 74; Hornung 1986: 138; Loksa 1987: 79; Loksa 22. 1991: 273; Traser 1997: 62; Farkas et al. 1998: 57; Traser 1999: 56; Dombos 2001: 2040; Traser 2002a: 265; Traser 2002c: 48; Traser & Horváth-Szováti General distribution: Europe, North America. 2006: 333; Traser et al. 2006: 73; Traser 2006: 36; Distribution in Hungary: Börzsöny Mts. (Dun- Lepidocyrtus cyaneus var. assimilis Reuter: Stach ger 1975). 1929: 298.

Lepidocyrtus curvicollis (Bourlet, 1839) General distribution: cosmopolitan. Distribution in Hungary: Aggtelek Karst Lepidocyrtus curvicollis Bourlet: Tömösváry 1884a: (Loksa 1966; Traser 1999), Bakony Mts. (Loksa 53; Vellay 1900: 21; Stach 1929: 298; Dudich 1932: 49; Gebhardt 1934: 140; Loksa 1960: 49; Paclt 1960: 1966; Loksa & Rubio 1966a), Börzsöny Mts. 156; Loksa 1961a: 257; Loksa 1962: 63; Loksa 1966: (Loksa 1966), Buda Mts. (Loksa 1966), Budapest

35 Dányi & Traser: Checklist of the springtail fauna of Hungary

(Stach 1929), Bükk Mts. (Loksa 1956a, 1966), (Loksa 1956a, 1966), Dráva Plain (Loksa 1978a), Cserhát (Loksa 1978b), Dráva Plain (Loksa Fertő-Hanság (Traser 2002a; Traser et al. 2006), 1978a), Fertő-Hanság (Traser et al. 2006), Gere- Gerecse Mts. (Loksa 1966; Traser 2006), Horto- cse Mts. (Loksa 1966; Traser 2006), Hortobágy bágy (Loksa 1983), Keszthely Mts. (Loksa 1966), (Loksa 1983), Kiskunság (Vellay 1900; Stach Kis-Balaton (Farkas 1996, 1997), Kiskunság 1929; Hornung 1986; Loksa 1987; Traser 2002c; (Vellay 1900; Stach 1929; Szent-Ivány 1940a; Traser & Horváth-Szováti 2006), Kőszeg Mts. Traser 2002c; Traser & Horváth-Szováti 2006), (Szent-Ivány 1940a), Mátra Mts. (Loksa 1980), Kőszeg Mts. (Szent-Ivány 1940a), Mátra Mts. Mezőföld (Stach 1929; Dombos 2001), Nyírség (Loksa 1980), Mecsek Mts. (Paclt 1960; Loksa (Vellay 1900; Stach 1929; Kovács 1953a; Loksa 1966), Mezőföld (Stach 1929; Dombos 2001), Mt. 1991), Pilis Mts. (Loksa 1966), Somogy Hills Somló (Loksa 1966), Nyírség (Loksa 1991), (Farkas et al. 1998), Sopron Mts. (Traser 2002a), Őrség (Traser 1995), Pilis Mts. (Loksa 1966), Szigetköz (Traser 1997, 2002a), Tétény Plateou Somogy Hills (Farkas et al. 1998), Sopron Mts. (Szent-Ivány 1941), Vas-Sopron Plain (Traser (Traser 1981), Vas-Sopron Plain (Traser 1981), 1981), Visegrád Mts. (Loksa 1966). Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), Visegrád Mts. (Loksa 1966). Lepidocyrtus cf. instratus Handschin, 1924 Ecological note: mesophilic.

Lepidocyrtus cf. instratus Handschin: Traser 1980: 20. Lepidocyrtus lignorum (Fabricius, 1793) General distribution: Central Europe. Distribution in Hungary: Bakony Mts. (Traser Lepidocyrtus lignorum (Fabricius): Vellay 1900: 21; Tömösváry 1882: 124; Dunger 1975: 22; Loksa 1977: 1980). 209; Loksa 1978a: 61; Loksa 1978b: 95; Loksa 1991: Ecological note: alpine. 273; Traser 1995: 45; Traser 1997: 62; Traser 1999: 56; Traser & Csóka 2000: 76; Traser & Csóka 2001: Lepidocyrtus lanuginosus (Gmelin, 1788) 233; Traser 2002a: 265; Traser 2002b: 160; Traser 2002c: 48; Traser & Salló 2004: 230; Traser 2006: 36; Lepidocyrtus pusillus Linnaeus: Vellay 1900: 21; Traser et al. 2006: 73. Lepidocyrtus rivularis Bourlet: Stach 1929: 297; Lepidocyrtus rivularis var. albicans Reuter: Stach 1929: 297; Lepidocyrtus lanuginosus (Gmelin): Stach General distribution: Holarctic. 1929: 297; Szent-Ivány 1940a: 433; Loksa 1956a: 391; Distribution in Hungary: Aggtelek Karst (Tra- Loksa 1956b: 203; Paclt 1960: 157; Loksa 1966: 384; Loksa & Rubio 1966a: 152; Dunger 1975: 22; Loksa ser 1999), Börzsöny Mts. (Dunger 1975), Bükk 1978a: 60; Loksa 1980: 111; Traser 1980: 20; Traser Mts. (Loksa 1977; Traser 2002b), Cserhát (Loksa 1981: 133; Loksa 1983: 74; Loksa 1991: 273; Farkas 1995: 46; Traser 1995: 45; Farkas 1996: 287; Farkas 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság 1997: 56; Farkas & Szathmáry 1997: 39; Farkas et al. (Traser et al. 2006), Gerecse Mts. (Traser 2006), 1998: 57; Traser 1999: 56; Dombos 2001: 2040; Traser Kiskunság (Traser & Csóka 2000, 2001; Traser 2002a: 265; Traser 2002c: 48; Traser 2006: 36; Traser et al. 2006: 73; Traser & Horváth-Szováti 2006: 334; 2002c), Kőszeg Mts. (Traser & Salló 2004), Nyír- Lepidocyrtus lanuginosus f. albicans Reuter: Szent- ség (Loksa 1991), Őrség (Traser 1995), Sopron Ivány 1940a: 433; Lepidocyrtus lanuginosus f. fucata Uzel: Szent-Ivány 1940a: 433; Lepidocyrtus cf. Mts. (Traser 2002a), Szigetköz (Traser 1997, lanuginosus (Gmelin): Loksa 1961b: 224; Traser 1981: 2002a), Zemplén Mts. (Tömösváry 1882; Vellay 133. 1900). General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Lok- Lepidocyrtus nigrescens Szeptycki, 1967 sa 1966; Traser 1999), around the Lake Balaton Lepidocyrtus nigrescens Szeptycki: Dunger 1975: 22; (Farkas 1995, 1997), Bakony Mts. (Loksa 1966; Traser 1999: 57; Traser & Horváth-Szováti 2006: 333. Loksa & Rubio 1966a; Traser 1980), Balaton- felvidék (Stach 1929; Loksa 1966; Farkas 1995, General distribution: Europe. 1997; Farkas & Szathmáry 1997), Börzsöny Mts. Distribution in Hungary: Aggtelek Karst (Tra- (Loksa 1966; Dunger 1975), Buda Mts. (Loksa ser 1999), Börzsöny Mts. (Dunger 1975), Kiskun- 1956b, 1966), Budapest (Stach 1929), Bükk Mts. ság (Traser & Horváth-Szováti 2006).

36 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ecological note: xerothermophilic. Lepidocyrtus serbicus Denis, 1933

Lepidocyrtus serbicus Denis: Dunger 1975: 22. Lepidocyrtus paradoxus Uzel, 1890

Lepidocyrtus paradoxus Uzel: Stach 1929: 298; Szent- General distribution: Europe. Ivány 1940a: 432; Szent-Ivány 1941: 22; Loksa 1978a: Distribution in Hungary: Börzsöny Mts. (Dun- 61; Traser 1981: 133; Farkas 1995: 46; Traser 1995: ger 1975). 45; Farkas 1997: 56; Traser 1997: 62; Farkas et al. 1998: 57; Traser 1999: 57; Traser 2000: 68; Dombos 2001: 2040; Traser 2002a: 265; Traser 2002c: 48; Lepidocyrtus cf. tellecheae Arbea & Jordana, 1989 Traser 2003: 156; Traser & Horváth-Szováti 2006: 334; Traser et al. 2006: 73. Lepidocyrtus cf. tellecheae Arbea & Jordana: Traser & Horváth–Szováti 2004: 59; Traser & Horváth-Szováti 2006: 334. General distribution: Holarctic.

Distribution in Hungary: Aggtelek Karst (Tra- General distribution: Europe. ser 1999), around the Lake Balaton (Farkas 1995, Distribution in Hungary: Kiskunság (Traser & 1997), Balatonfelvidék (Farkas 1995, 1997), Buda Mts. (Stach 1929), Dráva Plain (Loksa 1978a), Horváth–Szováti 2004, 2006). Fertő-Hanság (Szent-Ivány 1941; Traser 2000, 2002a, 2003; Traser et al. 2006), Kiskunság Lepidocyrtus violaceus (Geoffroy, 1762) (Stach 1929; Traser 2002c; Traser & Horváth- Lepidocyrtus violaceus Lubbock: Dunger 1975: 22; Szováti 2006), Kőszeg Mts. (Szent-Ivány 1940a), Traser 2002a: 265; Traser 2006: 36; Lepidocyrtus Mezőföld (Stach 1929; Dombos 2001), Őrség violaceus (Fourcroy (sic!)): Traser 1995: 45; Traser (Traser 1995), Somogy Hills (Farkas et al. 1998), 1999: 57; Traser et al. 2006: 73. Sopron Mts. (Traser 1981, 2002a), Szekszárd General distribution: Holarctic. Hills (Stach 1929), Szigetköz (Traser 1997), Zala Distribution in Hungary: Aggtelek Karst (Tra- Hills (Stach 1929). ser 1999), Börzsöny Mts. (Dunger 1975), Fertő- Ecological note: mostly in open habitats. Hanság (Traser et al. 2006), Gerecse Mts. (Traser 2006), Őrség (Traser 1995), Sopron Mts. (Traser Lepidocyrtus peisonis Traser & Christian, 1992 2002a). Lepidocyrtus peisonis Traser & Christian: Traser & Christian 1992: 119, figs 1-6; Traser 2000: 68; Traser Lepidocyrtus cf. weidneri Hüther, 1971 2002a: 265; Traser 2003: 156; Traser et al. 2006: 73. Lepidocyrtus cf. weidneri Hüther: Traser 2003: 156.

General distribution: Central Europe. General distribution: Central Europe. Distribution in Hungary: Fertő-Hanság (Traser Distribution in Hungary: Fertő-Hanság (Traser & Christian 1992; Traser 2000, 2002a, 2003; 2003). Traser et al. 2006).

Ecological note: hygrophilous. Pseudosinella Schäffer, 1897

Lepidocyrtus ruber Schött, 1902 Pseudosinella aggtelekiensis (Stach, 1929) Lepidocyrtus ruber var. albidus Stach: Stach 1929: 297; Lepidocyrtus ruber Schött: Loksa 1978a: 61; Lepidocyrtus (Pseudosinella) aggtelekiensis Stach: Traser 1980: 20; Farkas 1995: 46; Traser 1995: 45; Stach 1929: 296; Lepidocyrtus aggtelekiensis Stach: Traser 2002a: 265. Dudich 1930: 70; Dudich 1932: 49; Kovács 1953b: 90; Kovács 1954: 97; Traser 1999: 49; Pseudosinella aggtelekiensis Stach: Loksa 1961b: 224; Traser 1999: General distribution: Europe. 57. Distribution in Hungary: around the Lake Ba- laton (Farkas 1995), Bakony Mts. (Traser 1980), General distribution: Central-Europe. Balatonfelvidék (Stach 1929), Dráva Plain (Loksa Distribution in Hungary: Aggtelek Karst 1978a), Fertő-Hanság (Traser 2002a), Őrség (Tra- (Stach 1929; Dudich 1930, 1932; Kovács 1953b, ser 1995), Sopron Mts. (Traser 2002a). 1954; Loksa 1961b; Traser 1999).

37 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ecological note: troglophilous. Pseudosinella duodecimpunctata Denis, 1931

Pseudosinella duodecimpunctata Denis: Loksa 1978a: 61. Pseudosinella alba (Packard, 1873) Pseudosinella alba (Packard): Szent-Ivány 1940a: 433; General distribution: Europe. Loksa 1956a: 391; Loksa 1956b: 203; Loksa 1961a: Distribution in Hungary: Dráva Plain (Loksa 257, figs 25-27; Loksa 1962: 63; Loksa 1966: 384; 1978a). Loksa & Rubio 1966a: 152; Dunger 1975: 23; Loksa 1977: 211; Loksa 1978b: 95; Traser 1981: 133; Traser 1997: 62; Loksa 1991: 273; Traser 1995: 45; Traser Pseudosinella cf. fallax (Börner, 1903) 1999: 57; Traser 2002a: 265; Traser 2003: 156; Traser Pseudosinella cf. fallax Börner, 1903: Traser 1999: 57. & Horváth–Szováti 2004: 59; Traser 2006: 36; Traser & Horváth-Szováti 2006: 334; Traser et al. 2006: 73. General distribution: Southern Europe. General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Tra- Distribution in Hungary: Aggtelek Karst (Lok- ser 1999). sa 1966; Traser 1999), Bakony Mts. (Loksa 1966; Ecological note: in montane habitats. Loksa & Rubio 1966a), Balatonfelvidék (Loksa 1961a, 1966), Börzsöny Mts. (Loksa 1966; Dun- Pseudosinella horaki Rusek, 1985 ger 1975), Buda Mts. (Loksa 1956b, 1966), Bükk Pseudosinella horaki Rusek, 1985: Traser 1999: 57; Mts. (Loksa 1956a, 1962, 1966, 1977), Cserhát Traser 2002a: 266. (Loksa 1978b), Fertő-Hanság (Traser 2002a, 2003; Traser et al. 2006), Gerecse Mts. (Loksa General distribution: Central Europe. 1966; Traser 2006), Keszthely Mts. (Loksa 1966), Distribution in Hungary: Aggtelek Karst (Tra- Kiskunság (Traser & Horváth–Szováti 2004, ser 1999), Moson Plain (Traser 2002a), Sopron 2006), Kőszeg Mts. (Szent-Ivány 1940a), Mt. Mts. (Traser 2002a). Somló (Loksa 1966), Nyírség (Loksa 1991), Őrség (Traser 1995), Pilis Mts. (Loksa 1966), Pseudosinella immaculata (Lie-Pettersen, 1896) Sopron Mts. (Traser 1981, 2002a), Szigetköz (Traser 1997), Vas-Sopron Plain (Traser 1981), Pseudosinella immaculata (Lie-Pettersen): Loksa 1956a: 395; Loksa 1966: 384; Pseudosinella impari- Vértes Mts. (Loksa 1966), Visegrád Mts. (Loksa punctata Gisin (sic!): Loksa 1966: 384; Farkas 1995: 1966). 46; Traser 1999: 57; Traser 2002a: 266. Ecological note: edaphic. General distribution: Europe. Pseudosinella argentea Loksa, 1961 Distribution in Hungary: Aggtelek Karst (Lok- Pseudosinella argentea Loksa: Loksa 1961a: 262, figs sa 1966; Traser 1999), around the Lake Balaton 28-29. (Farkas 1995), Bükk Mts. (Loksa 1956a), Fertő- Hanság (Traser 2002a), Mecsek Mts. (Loksa General distribution: Central Europe. 1966). Distribution in Hungary: Balatonfelvidék (Loksa 1961a). Pseudosinella ksenemani Gisin, 1944

Pseudosinella decipiens Denis, 1924 Pseudosinella ksenemani Gisin: Dunger 1975: 23; Loksa 1978a: 61. Pseudosinella decipiens Denis: Loksa 1966: 384; Loksa & Rubio 1966a: 152; Traser 2002a: 265; Traser General distribution: Europe. 2002c: 48. Distribution in Hungary: Börzsöny Mts. (Dun- General distribution: Europe. ger 1975), Dráva Plain (Loksa 1978a). Distribution in Hungary: Bakony Mts. (Loksa & Rubio 1966a), Buda Mts. (Loksa 1966), Gere- Pseudosinella octopunctata Börner, 1901 cse Mts. (Loksa 1966), Kiskunság (Traser 2002c), Lepidocyrtus (Pseudosinella) octopunctatus Börner: Moson Plain (Traser 2002a). Stach 1929: 296; Pseudosinella octopunctata Börner:

38 Dányi & Traser: Checklist of the springtail fauna of Hungary

Paclt 1960: 157; Dunger 1975: 23; Loksa 1956b: 203; 1995), Pilis Mts. (Loksa 1966), Somogy Hills Loksa 1977: 211; Loksa 1978a: 61; Traser 1980: 20; (Farkas et al. 1998), Sopron Mts. (Traser 1981), Traser et al. 2006: 73. Vas-Sopron Plain (Traser 1981), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), General distribution: Holarctic. Visegrád Mts. (Loksa 1966). Distribution in Hungary: Bakony Mts. (Traser

1980), Balatonfelvidék (Paclt 1960), Börzsöny Pseudosinella zygophora (Schille, 1908) Mts. (Dunger 1975), Buda Mts. (Loksa 1956b), Bükk Mts. (Loksa 1977), Dráva Plain (Loksa Lepidocyrtus zygophorus Schille: Dudich 1932: 49; 1978a), Fertő-Hanság (Traser et al. 2006), Pseudosinella zygophorus (Schille): Traser 1999: 57. Mezőföld (Stach 1929). General distribution: Central Europe. Distribution in Hungary: Aggtelek Karst (Du- Pseudosinella petterseni Börner, 1901 dich 1932; Traser 1999). Pseudosinella petterseni Börner: Loksa 1966: 385; Dunger 1975: 23; Loksa 1977: 211; Loksa 1978b: 95. Seira Lubbock, 1869

General distribution: cosmopolitan. Seira dollfusi (Carl, 1899) Distribution in Hungary: Börzsöny Mts. (Dun- ger 1975), Bükk Mts. (Loksa 1977), Cserhát Pesudosira dollfusi Carl (sic!): Stach 1929: 299; Lepidocyrtus dolfussi Carl (sic!): Szent-Ivány 1941: (Loksa 1978b), Mecsek Mts. (Loksa 1966), 22, fig. 1. Villány Mts (Loksa 1966). General distribution: Western Europe. Pseudosinella sexoculata Schött, 1902 Distribution in Hungary: Buda Mts. (Stach 1929; Szent-Ivány 1941). Pseudosinella sexoculata Schött: Loksa 1966: 385; Traser & Horváth-Szováti 2006: 334; Traser 2002a: 266; Traser 2002c: 48. Seira domestica (Nicolet, 1842)

Seira domestica (Nicolet): Loksa 1966: 384; Traser General distribution: Holarctic. 2002a: 265; Seirra (sic!) domestica (Nicolet): Dombos Distribution in Hungary: Fertő-Hanság (Traser 2001: 2043. 2002a), Kiskunság (Traser & Horváth-Szováti 2006; Traser 2002c), Mecsek Mts. (Loksa 1966), General distribution: Europe. Villány Mts (Loksa 1966). Distribution in Hungary: Bükk Mts. (Loksa 1966), Fertő-Hanság (Traser 2002a), Mezőföld (Dombos 2001). Pseudosinella wahlgreni (Börner, 1907) Ecological note: synanthropic. Pseudosinella wahlgreni (Börner): Loksa 1966: 385; Dunger 1975: 23; Loksa 1977: 209; Traser 1980: 20; Seira pallidipes Reuter, 1895 Loksa 1980: 111; Traser 1981: 133; Loksa 1991: 273; Farkas 1995: 46; Traser 1995: 45; Farkas et al. 1998: Sira pallidipes Reuter (sic!): Reuter 1895a: 114; Stach 57; Loksa & Rubio 1966a: 152; Loksa 1978a: 61; 1929: 299; Seira pallidipes Reuter: Vellay 1900: 21; Loksa 1978b: 95. Loksa 1983: 74; Loksa 1987: 79; Seira pallipes (sic!) Reuter: Hornung 1986: 138.

General distribution: Europe, North Africa. Distribution in Hungary: around the Lake Ba- General distribution: Eastern Europe. laton (Farkas 1995), Bakony Mts. (Traser 1980; Distribution in Hungary: Kiskunság (Reuter Loksa & Rubio 1966a), Börzsöny Mts. (Loksa 1895a; Vellay 1900; Stach 1929; Loksa 1983; 1966; Dunger 1975), Buda Mts. (Loksa 1966), Hornung 1986; Loksa 1987). Bükk Mts. (Loksa 1966, 1977), Cserhát (Loksa Ecological note: in open habitats. 1978b), Dráva Plain (Loksa 1978a), Gerecse Mts. (Loksa 1966), Keszthely Mts. (Loksa 1966), Seira pillichi Stach, 1930 Mátra Mts. (Loksa 1980), Mecsek Mts. (Loksa Pseudosira pillichi Stach (sic!): Stach 1929: 299; 1966), Nyírség (Loksa 1991), Őrség (Traser Loksa 1959c: 389.

39 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Central Europe. General distribution: Europe. Distribution in Hungary: Mezőföld (Stach Distribution in Hungary: Buda Mts. (Stach 1929; Loksa 1959c). 1929), Mezőföld (Stach 1929), Sopron Mts. (Tra- ser 2002a). Seira stachi Loksa, 1990 Ecological note: corticophilous.

Seira stachi Loksa: Loksa 1991: 274, figs 3a-d. Subfamily: Orchesellinae

General distribution: Central Europe. Heteromurus Wankel, 1860 Distribution in Hungary: Nyírség (Loksa Heteromurus major (Moniez, 1889) 1991). Heteromurus major (Moniez): Stach 1929: 294; Loksa 1966: 384; Dunger 1975: 23; Loksa 1983: 74; Farkas Willowsia Shoebotham, 1917 1995: 46; Traser 1999: 56; Traser 2002a: 265; Traser 2006: 36; Traser et al. 2006: 73. Willowsia buski (Lubbock, 1871) General distribution: Central and Southern Seira buskii (sic!) Lubbock: Vellay 1900: 21; Sira Europe. buski Lubbock: Stach 1929: 299; Szent-Ivány 1940a: 432; Willowsia buski (Lubbock): Loksa 1978a: 60; Distribution in Hungary: Aggtelek Karst (Lok- Traser 1999: 56; Traser 2002a: 265; Traser 2003: 156; sa 1966; Traser 1999), around the Lake Balaton Traser & Horváth-Szováti 2006: 334. (Farkas 1995), Bakony Mts. (Loksa 1966), Bala- tonfelvidék (Loksa 1966), Börzsöny Mts. (Loksa General distribution: Holarctic. 1966; Dunger 1975), Buda Mts. (Loksa 1966), Distribution in Hungary: Aggtelek Karst (Tra- Fertő-Hanság (Traser 2002a; Traser et al. 2006), ser 1999), Cserhát (Stach 1929; Szent-Ivány Gerecse Mts. (Loksa 1966; Traser 2006), Horto- 1940a), Dráva Plain (Loksa 1978a), Fertő-Hanság bágy (Loksa 1983), Keszthely Mts. (Loksa 1966), (Traser 2002a, 2003), Kiskunság (Vellay 1900; Mecsek Mts. (Loksa 1966), Mezőföld (Stach Stach 1929; Szent-Ivány 1940a; Traser & Hor- 1929), Mt. Somló (Loksa 1966), Vértes Mts. váth-Szováti 2006), Kőszeg Mts. (Szent-Ivány (Loksa 1966), Villány Mts (Loksa 1966), 1940a), Mezőföld (Stach 1929; Szent-Ivány Visegrád Mts. (Loksa 1966). 1940a), Sopron Mts. (Traser 2002a), "bank of the Ecological note: thermophilic. River Bodrog" (Szent-Ivány 1940a). Ecological note: synanthrop, corticophilous. Heteromurus nitidus (Templeton, 1835) Templetonia nitida Templeton: Vellay 1900: 21; He- Willowsia nigromaculata (Lubbock, 1873) teromurus nitidus (Templeton): Stach 1929: 294; Gebhardt 1934: 139; Szent-Ivány 1941: 23, figs 2-3; Sira platani ab. nigromaculata (Lubbock): Stach 1929: Loksa 1959b: 70; Paclt 1960: 157; Loksa 1961a: 262, 299; Sira platani f. nigromaculata Lubbock: Szent- fig. 30; Loksa 1961b: 224; Loksa 1962: 63; Loksa Ivány 1940a: 432; Kovács 1953a: 184; Loksa 1991: 1966: 384; Dunger 1975: 23; Loksa 1977: 211; Loksa 26; Willowsia nigromaculata (Lubbock): Loksa & 1978a: 61; Loksa 1978b: 95; Traser 1981: 135; Loksa Rubio 1966a: 152. 1983: 74; Loksa 1991: 273; Farkas 1995: 46; Traser 1995: 45; Farkas 1996: 287; Traser 1997: 62; Farkas General distribution: Holarctic. 1997: 58; Farkas et al. 1998: 57; Traser 1999: 56; Traser 2003: 156; Traser 2006: 36; Traser et al. 2006: Distribution in Hungary: Bakony Mts. (Loksa 73; Heteromurus nitidus Absolon (sic!): Gebhardt & Rubio 1966a), Kőszeg Mts. (Szent-Ivány 1933: 38; Heteromurus nitidus var. margaritarius 1940a, 1941), Mezőföld (Stach 1929; Szent-Ivány (Wankel): Stach 1929: 295; Heteromurus nitidus var. 1941), Nyírség (Kovács 1953a; Loksa 1991). margaritaria (sic!) (Wankel): Dudich 1932: 49; He- teromurus nitidus var. margaritarius Absolon (sic!): Ecological note: corticophilous. Gebhardt 1934: 139; Heteromurus nitidus f. mar- garitaria Wankel: Paclt 1960: 157; Heteromurus ni- Willowsia platani (Nicolet, 1842) tidus var. margaritaria Wankel: Loksa & Rubio 1966a: 154; Heteromurus nitidus var. ocellata Stach: Gebhardt Sira platani ab. corticalis (Carl): Stach 1929: 300; 1934: 139; Heteromurus nitidus var. paucidentatus Willowsia platani (Nicolet): Traser 2002a: 265. Stach: Stach 1929: 295; Dudich 1932: 49; Gebhardt

40 Dányi & Traser: Checklist of the springtail fauna of Hungary

1934: 139; Szent-Ivány 1940b: 120; Szent-Ivány 1941: váth-Szováti 2006), Mezőföld (Dombos 2001). 23; Heteromurus nitidus var. principalis Stach: Geb- Ecological note: xerothermophilic. hardt 1934: 139; Heteromurus nitidus f. quadriocellata (sic!) Kseneman: Paclt 1960: 157; Heteromurus nitidus var. quadrioculata Ksenaman: Traser 2002a: 265. Orchesella alticola Uzel, 1891 General distribution: Holarctic. Orchesella alticola Uzel: Loksa 1966: 383; Traser Distribution in Hungary: Aggtelek Karst 1999: 56. (Stach 1929; Dudich 1932; Gebhardt 1934; Loksa 1961b; Traser 1999), around the Lake Balaton General distribution: Western and Central Eu- (Farkas 1995), Bakony Mts. (Loksa 1966; Loksa rope. & Rubio 1966a), Balatonfelvidék (Paclt 1960; Loksa 1961a, 1966), Börzsöny Mts. (Loksa 1966; Distribution in Hungary: Aggtelek Karst (Lok- Dunger 1975), Buda Mts. (Stach 1929; Gebhardt sa 1966; Traser 1999). 1934; Szent-Ivány 1941; Loksa 1966), Bükk Mts. Ecological note: in montane habitats. (Loksa 1962, 1977), Cserhát (Loksa 1959b, 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság Orchesella balcanica Stach, 1960 (Traser 2002a, 2003; Traser et al. 2006), Gerecse Mts. (Loksa 1966; Traser 2006), Hortobágy (Lok- Orchesella balcanica Stach: Loksa 1966: 383. sa 1983), Keszthely Mts. (Loksa 1966), Kis- Balaton (Farkas 1996, 1997), Kiskunság (Vellay General distribution: Southern and Eastern 1900; Stach 1929), Mecsek Mts. (Stach 1929; Europe. Gebhardt 1933; Gebhardt 1934; Szent-Ivány Distribution in Hungary: Mecsek Mts. (Loksa 1940b, 1941; Paclt 1960; Loksa 1966), Nyírség 1966), Villány Mts (Loksa 1966). (Loksa 1991), Őrség (Traser 1995), Pilis Mts. (Stach 1929; Loksa 1966), Somogy Hills (Farkas Ecological note: in montane habitats. et al. 1998), Sopron Mts. (Traser 2002a), Szi- getköz (Traser 1997), Vas-Sopron Plain (Traser Orchesella bifasciata Nicolet, 1842 1981), Vértes Mts. (Loksa 1966), Villány Mts Orchesella bifasciata Nicolet: Vellay 1900: 20; Stach (Loksa 1966), Visegrád Mts. (Loksa 1966). 1929: 306; Szent-Ivány 1938: 92; Szent-Ivány 1940a: Ecological note: edaphic, troglophilous. 433; Loksa 1956b: 205; Paclt 1960: 157; Loksa & Rubio 1966a: 152; Dunger 1975: 22; Loksa 1978a: 60; Loksa 1978b: 95; Hornung 1986: 138; Loksa 1991: Heteromurus sexoculatus Brown, 1926 272; Traser 1999: 56; Traser & Csóka 2000: 78; Traser Heteromurus sexoculatus Brown: Loksa 1983: 74, figs & Csóka 2001: 237; Traser 2002c: 48; Traser et al. 7-13. 2006: 73; Orchesella bifasciata f. intermedia Ågren: Szent-Ivány 1940a: 433; Szent-Ivány 1941: 23, figs 4- 5. General distribution: Asia Minor. Distribution in Hungary: Hortobágy (Loksa General distribution: Europe. 1983). Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Bakony Mts. (Loksa & Rubio 1966a), Orchesella Templeton, 1835 Balatonfelvidék (Paclt 1960), Börzsöny Mts. Orchesella albofasciata Stach, 1960 (Dunger 1975), Buda Mts. (Loksa 1956b), Csehát (Loksa 1978b), Dráva Plain (Loksa 1978a), Fertő- Orchesella albofasciata Stach: Traser & Csóka 2000: Hanság (Traser et al. 2006), Kiskunság (Vellay 78; Traser & Csóka 2001: 233; Dombos 2001: 2040; Traser 2002c: 48; Traser & Horváth-Szováti 2006: 1900; Stach 1929; Szent-Ivány 1938, 1940a; Ho- 333. nung 1986; Traser & Csóka 2000, 2001; Traser 2002c), Kőszeg Mts. (Szent-Ivány 1940a, 1941), General distribution: Europe. Mátra Mts. (Szent-Ivány 1938, 1941), Nyírség Distribution in Hungary: Kiskunság (Traser & (Loksa 1991). Csóka 2000, 2001; Traser 2002c; Traser & Hor- Ecological note: planar to montane.

41 Dányi & Traser: Checklist of the springtail fauna of Hungary

Orchesella bulgarica Stach, 1960 Mecsek Mts. (Tömösváry 1884a; Vellay 1900; Stach 1929), Mezőföld (Stach 1929; Stach 1960; Orchesella bulgarica Stach: Loksa 1966: 383; Dunger 1975: 22. Dombos 2001), Nyírség (Loksa 1991), Somogy Hills (Farkas et al. 1998), Szigetköz (Traser General distribution: Europe, Balkan Penin- 1997), Vác-Pest Duna Valley (Tömösváry 1884a; sula Vellay 1900; Stach 1929), Zemplén Mts. Distribution in Hungary: Balatonfelvidék (Tömösváry 1882; Vellay 1900; Stach 1929). (Loksa 1966), Börzsöny Mts. (Dunger 1975), Keszthely Mts. (Loksa 1966), Mecsek Mts. Orchesella flavescens (Bourlet, 1839) (Loksa 1966), Mt. Somló (Loksa 1966), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966). Orchesella rufescens Linnaeus (sic!): Tömösváry Ecological note: in montane habitats. 1884a: 54; Orchesella rufescens Wulfen (sic!): Vellay 1900: 20; Orchesella rufescens (Bourlet) (sic!): Stach 1929: 305; Orchesella flavescens var. melanocephala Orchesella cincta (Linnaeus, 1758) (Nicolet): Stach 1929: 305; Orchesella flavescens var. pallida Reuter: Stach 1929: 305; Kovács 1953a: 183; Orchesella cincta (Linnaeus): Tömösváry 1882: 123; Loksa 1991: 26; Orchesella flavescens var. quin- Tömösváry 1884a: 54; Vellay 1900: 20; Szent-Ivány quelineata Stach: Stach 1929: 305; Orchesella flaves- 1940a: 434; Loksa 1956a: 413; Stach 1960: 119, pl. 21: cens f. melanocephala Nicolet: Szent-Ivány 1938: 92; figs 1-4; Loksa 1966: 383; Dunger 1975: 22; Loksa Orchesella flavescens f. melanocephala Nicolet: Szent- 1977: 211; Loksa 1978a: 60; Loksa 1980: 111; Loksa Ivány 1940a: 434; Orchesella flavescens f. pallida 1983: 74; Hornung 1986: 138; Loksa 1987: 79; Loksa Reuter: Szent-Ivány 1940a: 434; Orchesella flavescens 1991: 272; Varga 1991: 85; Varga & Vargha 1992: (Bourlet): Szent-Ivány 1940a: 434; Loksa 1956b: 205; 315; Varga 1992: 172; Farkas 1995: 46; Traser 1997: Stach 1960: 36, pl. 5: figs 1-6; Loksa & Rubio 1966a: 62; Varga & Oldal 1998: 86; Traser & Csóka 2000: 78; 152; Loksa 1966: 383; Dunger 1975: 21; Loksa 1978a: Traser 2000: 68; Traser & Csóka 2001: 237; Dombos 60; Loksa 1978b: 95; Loksa 1983: 74; Loksa 1991: 2001: 2040; Varga et al. 2002a: 105; Traser 2002a: 272; Farkas 1995: 46; Traser 1995: 45; Farkas 1996: 265; Traser 2002c: 48; Varga & Naár 2002: 115; Varga 287; Farkas 1997: 56; Farkas & Szathmáry 1997: 39; et al. 2002b: 526; Traser 2003: 155; Traser 2006: 36; Farkas 1997: 58; Farkas et al. 1998: 57; Traser 1999: Traser et al. 2006: 73; Traser & Horváth-Szováti 2006: 56; Traser 2002a: 265; Traser 2003: 156; Traser & 333; Orchesella cincta var. vaga Linnaeus: Vellay Horváth-Szováti 2006: 333; Orchesella cf. flavescens 1900: 20; Stach 1929: 306; Orchesella cincta f. vaga (Bourlet): Traser 1981: 134. Linnaeus: Loksa 1966: 383; Orchesella cincta var. unifasciata (Nicolet): Stach 1929: 306; Orchesella cincta ab. unifasciata Nicolet: Szent-Ivány 1941: 24; General distribution: Palaearctic. Orchesella cincta (Nicolet (sic!)): Farkas et al. 1998: Distribution in Hungary: Aggtelek Karst (Tra- 57. ser 1999), Bakony Mts. (Loksa & Rubio 1966a), General distribution: Holarctic. Balatonfelvidék (Stach 1929; Farkas 1995, 1997; Distribution in Hungary: around the Lake Ba- Farkas & Szathmáry 1997, 1997), Börzsöny Mts. laton (Farkas 1995), Börzsöny Mts. (Dunger (Stach 1929; Dunger 1975), Buda Mts. (Loksa 1975), Buda Mts. (Stach 1929), Budapest (Tö- 1956b), Cserhát (Loksa 1978b), Dráva Plain mösváry 1884a; Vellay 1900; Stach 1929), Bükk (Loksa 1978a), Fertő-Hanság (Traser 2002a, Mts. (Loksa 1956a, 1966, 1977; Varga 1991, 2003), Hortobágy (Loksa 1983), Kis-Balaton 1992; Varga & Vargha 1992; Varga & Oldal (Farkas 1996, 1997), Kiskunság (Traser & 1998; Varga et al. 2002a, 2002b; Varga & Naár Horváth-Szováti 2006), Kőszeg Mts. (Szent-Ivány 2002), Dráva Plain (Loksa 1978a), Fertő-Hanság 1940a), Mátra Mts. (Szent-Ivány 1938), Mezőföld (Szent-Ivány 1941; Traser 2000, 2002a, 2003; (Stach 1929, 1960), Nyírség (Stach 1929; Kovács Traser et al. 2006), Gerecse Mts. (Traser 2006), 1953a; Loksa 1991), Őrség (Traser 1995), Pilis Gödöllő Hills (Tömösváry 1884a; Vellay 1900; Mts. (Loksa 1966), Somogy Hills (Farkas et al. Stach 1929), Hortobágy (Loksa 1983), Kiskunság 1998), Sopron Mts. (Traser 2002a), Szekszárd (Vellay 1900; Stach 1929; Hornung 1986; Loksa Hills (Stach 1929), Szigetköz (Traser 2002a), 1987; Traser & Csóka 2000, 2001; Traser 2002c; Vác-Pest Duna Valley (Tömösváry 1884a; Vellay Traser & Horváth-Szováti 2006), Kőszeg Mts. 1900; Stach 1929), Vas-Sopron Plain (Traser (Szent-Ivány 1940a), Mátra Mts. (Loksa 1980), 1981).

42 Dányi & Traser: Checklist of the springtail fauna of Hungary

Orchesella hungarica Stach, 1929 General distribution: Eastern Europe. Orchesella hungarica Stach: Stach 1929: 306; Loksa Distribution in Hungary: Balatonfelvidék 1959c: 389; Stach 1960: 102, pl. 17: figs 2-3; Loksa (Stach 1960). 1966: 383. Orchesella quinquefasciata (Bourlet, 1841) General distribution: Europe. Distribution in Hungary: Mecsek Mts. (Loksa Orchesella quinquefasciata Bourlet: Loksa 1966: 383. 1966), Mezőföld (Stach 1929; Loksa 1959c; Stach General distribution: Central Europe. 1960), Villány Mts (Loksa 1966). Distribution in Hungary: Bakony Mts. (Loksa 1966), Gerecse Mts. (Loksa 1966). Orchesella multifasciata (Stscherbakow, 1898) Orchesella multifasciata (Stscherbakow): Stach 1929: Orchesella spectabilis Tullberg, 1871 306; Stach 1960: 92, pl. 14: figs 1-4; Loksa & Rubio 1966a: 152; Dunger 1975: 22; Loksa 1978a: 60; Traser Orchesella spectabilis Tullberg: Vellay 1900: 20; 1980: 20; Loksa 1983: 74; Traser 1999: 56; Dombos Stach 1929: 305; Szent-Ivány 1940a: 434; Szent-Ivány 2001: 2040; Traser 2002a: 265; Traser & Horváth- 1941: 24, figs 6-7; Stach 1960: 50, pl. 8: figs 3-4; Szováti 2006: 333; Traser 2006: 36. Loksa 1966: 383; Loksa 1977: 211; Loksa 1983: 74; Loksa 1991: 272; Traser 1997: 62; Traser 1999: 56; Traser 2002a: 265; Orchesella spectabilis var. pallida General distribution: Europe. Tullberg: Stach 1929: 305. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Bakony Mts. (Loksa & Rubio 1966a; General distribution: Europe. Traser 1980), Börzsöny Mts. (Dunger 1975), Distribution in Hungary: Aggtelek Karst (Tra- Dráva Plain (Loksa 1978a), Gerecse Mts. (Traser ser 1999), Buda Mts. (Stach 1929, 1960), Bükk 2006), Hortobágy (Loksa 1983), Kiskunság (Tra- Mts. (Loksa 1977), Cserehát (Szent-Ivány 1941), ser & Horváth-Szováti 2006), Mecsek Mts. (Stach Fertő-Hanság (Traser 2002a), Gerecse Mts. (Lok- 1929, 1960), Mezőföld (Stach 1929, 1960; Dom- sa 1966), Hortobágy (Loksa 1983), Kiskunság bos 2001), Moson Plain (Traser 2002a), Szigetköz (Vellay 1900; Stach 1929), Kőszeg Mts. (Szent- (Traser 2002a). Ivány 1940a), Nyírség (Vellay 1900; Stach 1929; Ecological note: silvicolous, xerothermophil. Loksa 1991), Szigetköz (Traser 1997, 2002a). Ecological note: xerothermophilic. Orchesella orientalis Stach, 1960 Orchesella villosa (Geoffroy, 1762) Orchesella orientalis Stach: Loksa 1991: 274, figs 2a-d. Orchesella villosa (Geoffroy): Tömösváry 1882: 124; General distribution: Eastern Europe. Tömösváry 1884a: 54; Stach 1929: 306; Loksa 1966: 384; Loksa 1978b: 95; Orchesella villosa Linnaeus Distribution in Hungary: Nyírség (Loksa (sic!): Vellay 1900: 20; Orchesella multifasciata (sic!) 1991). Geoffroy: Loksa 1966: 384.

Orchesella pannonica Stach, 1960 General distribution: Europe, North-America. Orchesella pannonica Stach: Stach 1960: 59, pl. 9: figs 4-5. Distribution in Hungary: Aggtelek Karst (Lok sa 1966), Bakony Mts. (Loksa 1966), Balaton- felvidék (Loksa 1966), Börzsöny Mts. (Loksa General distribution: Central Europe. 1966), Buda Mts. (Loksa 1966), Budapest (Tö- Distribution in Hungary: Buda Mts. (Stach mösváry 1884a; Stach 1929; Vellay 1900), Bükk 1960). Mts. (Loksa 1966), Cserhát (Loksa 1978b), Gere- Ecological note: xerothermophilic. cse Mts. (Loksa 1966), Gödöllő Hills (Tömösváry 1884a; Stach 1929; Vellay 1900), Keszthely Mts. Orchesella pulchra Stscherbakow, 1898 (Loksa 1966), Mecsek Mts. (Tömösváry 1884a; Orchesella pulchra Stach (sic!): Stach 1960: 68, pl. 12: Vellay 1900; Stach 1929; Loksa 1966), Mt. Som- figs 1, 4. ló (Loksa 1966), Pilis Mts. (Loksa 1966), Vác-

43 Dányi & Traser: Checklist of the springtail fauna of Hungary

Pest Duna Valley (Tömösváry 1884a; Vellay Anurophorus laricis Nicolet, 1842 1900; Stach 1929), Vértes Mts. (Loksa 1966), Vil- Anurophorus laricis Nicolet: Stach 1929: 287; Loksa lány Mts (Loksa 1966), Visegrád Mts. (Loksa 1966: 381; Loksa 1980: 110; Traser 1995: 45; Traser 1966), Zemplén Mts. (Tömösváry 1882; Vellay 1999: 54; Traser 2002a: 264. 1900; Stach 1929). General distribution: Palaearctic. Orchesella xerothermica Stach, 1960 Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Budapest (Stach 1929), Cserhát (Stach Orchesella xerothermica Stach: Traser 1999: 56; Tra- 1929), Mátra Mts. (Loksa 1980), Mecsek Mts. ser 2002a: 265; Traser et al. 2006: 73. (Loksa 1966), Őrség (Traser 1995), Sopron Mts.

General distribution: Central Europe. (Traser 2002a), Villány Mts (Loksa 1966). Distribution in Hungary: Aggtelek Karst (Tra- Ecological note: mostly corticolous. ser 1999), Fertő-Hanság (Traser et al. 2006), Szi- getköz (Traser 2002a). Anurophorus pseudolaricis Loksa, 1978 Ecological note: xerothermophilic. Anurophorus pseudolaricis Loksa: Loksa 1978a: 56, figs 5-8. Family: Isotomidae General distribution: Hungary. Agrenia Börner, 1906 Distribution in Hungary: Dráva Plain (Loksa 1978a). Agrenia bidenticulata (Tullberg, 1876) Appendisotoma Stach, 1947 Agrenia bidenticulata (Tullberg): Traser 1999: 55. Appendisotoma franzi (Haybach, 1962) General distribution: Arctoalpine. Proisotoma cf. fanzi Haybach: Traser 1980: 20; Distribution in Hungary: Aggtelek Karst (Tra- Proisotoma franzi Haybach: Traser et al. 1993: 224; ser 1999). Traser & Csóka 2000: 77; Traser & Csóka 2001: 237; Traser 2002c: 48; Traser & Horváth-Szováti 2006: 333. Anurophorus Nicolet, 1842 General distribution: Central-Europe. Anurophorus atlanticus Fjellberg, 1974 Distribution in Hungary: Bakony Mts. (Traser 1980), Kiskunság (Traser et al. 1993; Traser & Anurophorus atlanticus Fjellberg: Traser 2002a: 264. Csóka 2000, 2001; Traser 2002c; Traser & Horváth-Szováti 2006). General distribution: Eastern Europe. Ecological note: xerothermophilic. Distribution in Hungary: Sopron Mts. (Traser 2002a). Appendisotoma juliannae Traser, Thibaud & Najt, 1993 Anurophorus cuspidatus Stach, 1947 Appendisotoma juliannae Traser, Thibaud & Najt: Anurophorus cuspidatus Stach: Loksa 1966: 381; Traser et al. 1993: 221, figs 1-2; Traser 2002c: 48; Loksa & Rubio 1966a: 151; Traser 1999: 54. Traser & Horváth-Szováti 2006: 333.

General distribution: Southern and Eastern General distribution: Eastern Europe (Hunga- Europe. ry, Poland). Distribution in Hungary: Aggtelek Karst (Lok- Distribution in Hungary: Kiskunság (Traser et sa 1966; Traser 1999), Bakony Mts. (Loksa & al. 1993; Traser 2002c; Traser & Horváth-Szováti Rubio 1966a). 2006). Ecological note: in montane habitats. Ecological note: silvicolous.

44 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ballistura Börner, 1906 Distribution in Hungary: Bakony Mts. (Loksa & Rubio 1966a), Dráva Plain (Loksa 1978a). Ballistura hankoi (Stach, 1929) Ecological note: silvicolous. Proisotoma hankoi Stach: Stach 1929: 288; Stach 1947: 196, pl. 23: figs 6-7; Traser 1999: 55. Cryptopygus Willem, 1901

General distribution: Central Europe. Cryptopygus bipunctatus (Axelson, 1903) Distribution in Hungary: Aggtelek Karst (Tra- Isotomina bipunctata Axelson: Loksa & Rubio 1966a: ser 1999), Balatonfelvidék (Stach 1929, 1947). 152; Loksa 1966: 382; Loksa 1977: 210; Loksa 1991: Ecological note: hygrophilous. 272; Cryptopygus bipunctatus (Axelson): Dunger 1975: 20; Traser 1999: 55; Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; Traser 2002c: 48; Traser & Ballistura schoetti (Dalla Torre, 1895) Horváth–Szováti 2004: 60; Traser & Horváth-Szováti 2006: 333; Traser et al. 2006: 72; Isotomia bipunctata Proisotoma schoetti (Dalla Torre): Traser 2002a: 264. (Axelson): Loksa 1978a: 57; Traser 1980: 20; Traser 1981: 133; Farkas et al. 1998: 57. General distribution: cosmopolitan. Distribution in Hungary: Fertő-Hanság (Traser General distribution: Palaearctic. 2002a). Distribution in Hungary: Aggtelek Karst (Lok- Ecological note: hygrophilous. sa 1966; Traser 1999), Bakony Mts. (Loksa & Ru- bio 1966a; Loksa 1966; Traser 1980), Balatonfel- Coloburella Latzel, 1918 vidék (Loksa 1966), Börzsöny Mts. (Loksa 1966; Dunger 1975), Buda Mts. (Loksa 1966), Bükk Coloburella reticulata Latzel, 1918 Mts. (Loksa 1966, 1977), Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser et al. 2006), Ge- Coloburella reticulata Latzel, 1918: Loksa 1991: 272. recse Mts. (Loksa 1966), Keszthely Mts. (Loksa 1966), Kiskunság (Traser & Csóka 2000, 2001; General distribution: Central and Southern Eu- Traser 2002c; Traser & Horváth–Szováti 2004, rope. 2006), Mecsek Mts. (Loksa 1966), Mt. Somló Distribution in Hungary: Nyírség (Loksa (Loksa 1966), Nyírség (Loksa 1991), Pilis Mts. 1991). (Loksa 1966), Somogy Hills (Farkas et al. 1998), Ecological note: silvicolous, in mulder. Sopron Mts. (Traser 1981), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), Visegrád Mts. Coloburella cf. vandeli Cassagnau & Delamare, (Loksa 1966). 1951 Ecological note: xerotolerant. Coloburella cf. vandeli Cassagnau & Delamare: Traser 1999: 54. Cryptopygus exilis (Gisin, 1960)

General distribution: Europe. Cryptopygus exilis (Gisin): Traser 2002a: 264. Distribution in Hungary: Aggtelek Karst (Tra- General distribution: Holarctic. ser 1999). Distribution in Hungary: Fertő-Hanság (Traser Ecological note: silvicolous. 2002a). Ecological note: mostly on meadows. Coloburella zangherii (Denis, 1924) Coloburella zangherii (Denis): Loksa & Rubio 1966a: Cryptopygus cf. orientalis (Stach, 1947) 151; Loksa 1978a: 56. Cryptopygus cf. orientalis (Stach): Traser 2002c: 48; Traser & Horváth-Szováti 2006: 333. General distribution: Southern and Central Europe. General distribution: Europe.

45 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Kiskunság (Traser General distribution: Holarctic. 2002c; Traser & Horváth-Szováti 2006). Distribution in Hungary: Kőszeg Mts. (Szent- Ecological note: xerothermophilic. Ivány 1940a). Ecological note: hygrophilous. Cryptopygus ponticus (Stach, 1947) Isotomina pontica Stach: Stach 1947: 267, pl. 32: figs Desoria nivea (Schäffer, 1896) 6-9; Cryptopygus pontica (sic!) (Stach): Traser 2002a: Isotoma albella Pack: Stach 1929: 290; Stach 1947: 264. 407, pl. 44. General distribution: Palaearctic. Distribution in Hungary: Fertő-Hanság (Traser General distribution: Europe. 2002a), Mezőföld (Stach 1947). Distribution in Hungary: Börzsöny Mts. Ecological note: in xerotherm, ruderal habitats. (Stach 1929, 1947). Ecological note: corticophilous. Cryptopygus thermophilus (Axelson, 1900) Desoria olivacea (Tullberg, 1871) Proistoma (Isotomina) thermophila (Axelson): Stach 1929: 287; Isotomia thermophila (Axelson) (sic!): Isotoma olivacea (Tullberg): Stach 1929: 290; Stach Loksa 1978a: 60; Isotomina thermophila (Axelson): 1947: 396, pl. 43; Loksa & Rubio 1966a: 152; Dunger Traser 1981: 135; Loksa 1983: 73; Cryptopygus 1975: 21; Loksa 1978a: 60; Loksa 1980: 110; Loksa thermophilus (Axelson): Traser 1997: 61; Traser 1983: 73; Loksa 1991: 272; Traser 1994: 102; Farkas 2002a: 264; Traser 2002c: 48; Traser & Horváth- 1995: 46; Farkas et al. 1998: 57; Traser & Salló 2004: Szováti 2006: 333. 230.

General distribution: cosmopolitan. Distribution in Hungary: Balatonfelvidék General distribution: Europe. (Stach 1929), Dráva Plain (Loksa 1978a), Fertő- Distribution in Hungary: around the Lake Ba- laton (Farkas 1995), Bakony Mts. (Loksa & Hanság (Traser 2002a), Hortobágy (Loksa 1983), Rubio 1966a), Börzsöny Mts. (Stach 1929, 1947; Kiskunság (Traser 2002c; Traser & Horváth- Dunger 1975), Budapest (Stach 1947), Dráva Szováti 2006), Mezőföld (Stach 1929), Szigetköz Plain (Loksa 1978a), Hortobágy (Loksa 1983), (Traser 1997), Vas-Sopron Plain (Traser 1981). Kőszeg Mts. (Traser & Salló 2004), Mátra Mts. Ecological note: nitrophilous, sometimes inter- (Loksa 1980), Nyírség (Loksa 1991), Pilis Mts. stitial. (Stach 1929, 1947), Somogy Hills (Farkas et al. 1998), Sopron Mts. (Traser 1994). Desoria Nicolet, 1841 Ecological note: hygrophilous, winter active. Desoria fennica (Reuter, 1895) Desoria propinqua (Axelson, 1902) Isotoma fennica (Reuter): Stach 1929: 290; Loksa 1977: 210; Traser 2002a: 264; Traser & Salló 2004: Spinisotoma pectinata Stach: Stach 1926b: 581, pl. 16; 230; Isotoma (Desoria) fennica (Reuter): Traser 1999: Stach 1929: 290; Stach 1947: 347, pl. 38; Isotoma 55. propinqua Axelson: Dunger 1975: 21; Traser 2002a: 264; Isotoma propinqua var. hexaspina Axelson: Tra- General distribution: Europe, North America. ser 1999: 55; Desoria propinqua (Axelson): Traser & Distribution in Hungary: Aggtelek Karst (Tra- Horváth-Szováti 2006: 333. ser 1999), Bükk Mts. (Loksa 1977), Fertő-Hanság (Traser 2002a), Kőszeg Mts. (Traser & Salló General distribution: Holarctic. 2004), Mezőföld (Stach 1929). Distribution in Hungary: Aggtelek Karst (Tra- Ecological note: mostly winter active. ser 1999), Bakony Mts. (Stach 1929, 1947), Bör- zsöny Mts. (Stach 1926b, 1929; Dunger 1975), Desoria neglecta (Schäffer, 1900) Fertő-Hanság (Traser 2002a), Kiskunság (Traser & Horváth-Szováti 2006). Isotoma olivacea var. neglecta Schäffer: Szent-Ivány 1940a: 429. Ecological note: litter dwelling.

46 Dányi & Traser: Checklist of the springtail fauna of Hungary

Desoria tigrina Nicolet, 1842 Folsomia candida Willem, 1902 Isotoma grisea Lubbock: Tömösváry 1884a: 55; Vel- Folsomia candida Willem: Stach 1947: 178, pl. 19; lay 1900: 20; Isotoma olivacea var. grisea (Lubbock): Loksa 1960: 49; Paclt 1960: 156; Loksa 1961a: 262, Stach 1929: 290; Isotoma tigrina Tullberg: Traser figs 18-24; Loksa 1962: 63; Loksa 1991: 272; Farkas 1999: 55. 1995: 46; Farkas 1996: 287; Farkas & Szathmáry 1997: 39; Farkas 1997: 56; Farkas et al. 1998: 57; General distribution: cosmopolitan. Traser 1999: 54; Traser 2002a: 264; ?Folsomia Distribution in Hungary: Aggtelek Karst (Tra- fimetaria (Linnaeus): Stach 1929: 287; Geyer & Mann ser 1999), Balatonfelvidék (Stach 1929), Buda- 1940: 371; Folsomia litsteri Bagnall: Loksa & Rubio 1966a: 151, fig. 57. pest (Stach 1929), Pilis Mts. (Stach 1929), Vác-

Pest Duna Valley (Tömösváry 1884a; Vellay Remarks: Data of Stach (1929) and Geyer & 1900; Stach 1929). Mann (1940) were corrected by Loksa (1960: 49). Ecological note: nitrophilous. According to Potapow (2001) Folsomia litsteri is a juvenile form of Folsomia candida. Desoria violacea (Tullberg, 1877) General distribution: cosmopolitan. Isotoma violacea Tullberg: Loksa & Rubio 1966a: 152; Distribution in Hungary: Aggtelek Karst (Tra- Loksa 1978a: 60; Loksa 1991: 272; Isotoma cf. viola- ser 1999), around the Lake Balaton (Farkas 1995, cea Tullberg: Traser 1999: 55. 1997), Bakony Mts. (Loksa & Rubio 1966a),

General distribution: Europe, Asia. Balatonfelvidék (Stach 1947; Loksa 1960; Paclt Distribution in Hungary: Aggtelek Karst (Tra- 1960; Loksa 1961a; Farkas 1995, 1997; Farkas & ser 1999), Bakony Mts. (Loksa & Rubio 1966a), Szathmáry 1997; Stach 1929), Bükk Mts. (Loksa Dráva Plain (Loksa 1978a), Nyírség (Loksa 1962), Kis-Balaton (Farkas 1996, 1997), Nyírség 1991). (Loksa 1991), Somogy Hills (Farkas et al. 1998), Ecological note: eurytopic. Sopron Mts. (Traser 2002a), Szigetköz (Traser 2002a). Folsomia Willem, 1902 Ecological notes: parthenogenetic, "flower pot springtail".

Folsomia antricola Loksa, 1959 Folsomia fimetaria (Linnaeus, 1758) Folsomia antricola Loksa: Loksa 1959a: 37, figs 1a-f; Loksa 1961b: 224; Bajomi 1969: 242; Folsomia Folsomia fimetaria (Linnaeus): Dudich 1932: 48; atricola (sic!) Loksa: Traser 1999: 54. Kovács 1953b: 90; Kovács 1954: 97; Loksa 1956a: 393; Traser 1999: 54; Traser 2002a: 264; Traser 2002c: 48; Traser & Horváth-Szováti 2006: 333. General distribution: Central Europe. Distribution in Hungary: Aggtelek Karst (Lok- General distribution: Holarctic. sa 1959a, 1961b; Bajomi 1969; Traser 1999). Distribution in Hungary: Aggtelek Karst (Du- Ecological note: troglophilous. dich 1932; Kovács 1953b, 1954; Traser 1999), Bükk Mts. (Loksa 1956a), Kiskunság (Traser Folsomia cf. brevifurca (Bagnall, 1949) 2002c; Traser & Horváth-Szováti 2006), Sopron Mts. (Traser 2002a). Folsomia cf. brevifurca Bagnall: Traser 1997: 62. Ecological note: in ruderal, nitrophilous ha-

bitats. Remark: Species inquirenda according to Pota- pow (2001: 186). General distribution: ?Palaearctic. Folsomia gebhardti Loksa, 1964 Distribution in Hungary: Szigetköz (Traser Folsomia gebhardti Loksa: Loksa 1964: 95, figs 51-57; 1997). Loksa 1966: 381.

47 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Central Europe (Hunga- (Traser 2002c; Traser & Horváth-Szováti 2006), ry, Serbia). Mátra Mts. (Loksa 1980), Őrség (Traser 1995), Distribution in Hungary: Mecsek Mts. (Loksa Somogy Hills (Farkas et al. 1998), Sopron Mts. 1964, 1966), Villány Mts (Loksa 1964, 1966). (Traser 1981, 1994, 2002a), Szigetköz (Traser Ecological note: in xerotherm forests. 1997), Vas-Sopron Plain (Traser 1981, 1981), Vértes Mts. (Loksa 1966). Folsomia kerni Gisin, 1948 Ecological note: eurytopic.

Folsomia kerni Gisin: Loksa & Rubio 1966a: 151, fig. Folsomia penicula Bagnall, 1939 58; Loksa 1978a: 57; Loksa 1978b: 95. Folsomia multiseta Stach: Loksa 1959a: 41, figs 2a-h; General distribution: Central Europe. Loksa 1966: 381; Loksa & Rubio 1966a: 151; Bajomi 1969: 242; Dunger 1975: 19; Loksa 1978a: 57; Loksa Distribution in Hungary: Bakony Mts. (Loksa 1978b: 95; Traser 1980: 20, fig. 4; Varga 1991: 85; & Rubio 1966a), Cserhát (Loksa 1978b), Dráva Loksa 1991: 272; Varga 1992: 172; Varga & Vargha Plain (Loksa 1978a). 1992: 315; Farkas 1995: 46; Farkas 1997: 56; Varga & Ecological note: in ruderal habitats. Oldal 1998: 86; Folsomia penicula Bagnall: Traser 1994: 102; Traser 1995: 45; Traser 1999: 55; Traser 2002a: 264; Traser et al. 2006: 72; Traser 2006: 36. Folsomia lawrencei Rusek, 1984 Folsomia lawrencei Rusek: Traser 2002a: 264. General distribution: Palaearctic. Distribution in Hungary: Aggtelek Karst (Lok- General distribution: Europe. sa 1966; Bajomi 1969; Traser 1999), Bakony Mts. Distribution in Hungary: Sopron Mts. (Traser (Loksa & Rubio 1966a; Traser 1980), Balatonfel- 2002a). vidék (Farkas 1995, 1997), Börzsöny Mts. (Loksa Ecological note: silvicolous. 1966; Dunger 1975), Bükk Mts. (Loksa 1966; Varga 1991, 1992; Varga & Vargha 1992; Varga Folsomia manolachei Baggnall, 1939 & Oldal 1998), Cserhát (Loksa 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser et al. Folsomia nana Gisin: Loksa 1966: 382; Loksa 1978b: 95; Loksa 1978a: 57; Traser 1980: 20; Loksa 1980: 2006), Gerecse Mts. (Traser 2006), Nyírség (Lok- 110; Traser 1981: 133; Loksa 1983: 73; Traser 1994: sa 1991), Őrség (Traser 1995), Pilis Mts. (Loksa 102; Farkas 1995: 46; Traser 1995: 45; Farkas 1996: 1966), Sopron Mts. (Traser 1994, 2002a), Viseg- 287; Traser 1997: 62; Farkas & Szathmáry 1997: 39; rád Mts. (Loksa 1966), Zala Hills (Loksa 1959a). Farkas 1997: 58; Farkas 1997: 56; Farkas et al. 1998: 57; Folsomia quadrioculata (Tullberg): Dunger 1975: Ecological note: silvicolous, mesophilic. 19 (partim); Folsomia manolachei Baggnall: Traser 1999: 54; Traser 2002c: 48; Traser 2002a: 264; Traser Folsomia quadrioculata (Tullberg, 1871) et al. 2006: 72; Traser & Horváth-Szováti 2006: 333; Traser 2006: 36. Folsomia quadrioculata (Tullberg): Szent-Ivány 1938: 92; Szent-Ivány 1940a: 428; Loksa 1956a: 391; Loksa 1956b: 203; Loksa 1962: 73; Loksa & Rubio 1966a: Remark: The data of Folsomia quadrioculata 151; Loksa 1966: 382; Dunger 1975: 19 (partim); (Tullberg) reported by Dunger (1975) included Loksa 1977: 207; Csuták 1977: 68; Loksa 1983: 73; „Folsomia nana Gisin” as well. Loksa 1991: 272; Traser 1994: 102; Traser 1997: 62; General distribution: Palaearctic. Traser 1999: 55; Traser 2002a: 264; Traser 2003: 155; Distribution in Hungary: Aggtelek Karst (Tra- Traser & Horváth-Szováti 2006: 333; Traser 2006: 36; ser 1999), around the Lake Balaton (Farkas 1995, Traser et al. 2006: 72. 1997), Bakony Mts. (Traser 1980), Balatonfel- Remark: The data of Folsomia quadrioculata vidék (Farkas 1995, 1997; Farkas & Szathmáry (Tullberg) reported by Dunger (1975) included 1997), Börzsöny Mts. (Dunger 1975), Cserhát „Folsomia nana Gisin” as well. (Loksa 1978b), Dráva Plain (Loksa 1978a), Fertő- General distribution: Holarctic. Hanság (Traser 2002a; Traser et al. 2006), Gere- Distribution in Hungary: Aggtelek Karst (Lok- cse Mts. (Traser 2006), Hortobágy (Loksa 1983), sa 1966; Traser 1999), Bakony Mts. (Loksa & Kis-Balaton (Farkas 1996, 1997), Kiskunság Rubio 1966a; Loksa 1966), Balatonfelvidék (Lok-

48 Dányi & Traser: Checklist of the springtail fauna of Hungary

sa 1966), Börzsöny Mts. (Loksa 1966; Dunger Folsomides semiparvulus Fjellberg, 1993 1975), Buda Mts. (Loksa 1956b, 1966), Bükk Folsomides semiparvulus Fjellberg: Traser 2002a: 264. Mts. (Loksa 1956a, 1962, 1966, 1977), Cserhát (Csuták 1977), Fertő-Hanság (Traser 2002a, 2003; Traser et al. 2006), Gerecse Mts. (Loksa General distribution: Canary Island, Mexico. 1966; Traser 2006), Hortobágy (Loksa 1983), Distribution in Hungary: Fertő-Hanság (Traser Keszhely Mts. (Loksa 1966), Kiskunság (Traser 2002a). & Horváth-Szováti 2006), Kőszeg Mts. (Szent- Ivány 1940a), Mátra Mts. (Szent-Ivány 1938, Hydroisotoma Stach, 1947 1940a), Mecsek Mts. (Loksa 1966), Mt. Somló (Loksa 1966), Nyírség (Loksa 1991), Pilis Mts. Hydroisotoma schaefferi (Krausbauer, 1898) (Loksa 1966), Sopron Mts. (Traser 1994, 2002a), Szigetköz (Traser 1997), Vértes Mts. (Loksa Hydroisotoma schaefferi (Krausbauer): Traser 2002a: 1966), Villány Mts (Loksa 1966), Visegrád Mts. 264. (Loksa 1966). Ecological note: eurytopic. General distribution: Holarctic. Distribution in Hungary: Sopron Mts. (Traser 2002a). Folsomides Stach, 1922 Ecological note: neustonic, along streams.

Folsomides angularis (Axelson, 1905) Isotoma Bourlet, 1839 Folsomides angularis (Axelson): Loksa 1966: 303; Loksa & Rubio 1966a: 152. Isotoma anglicana Lubbock, 1862 Isotoma anglicana Lubbock: Traser 2002a: 264; Traser General distribution: Holarctic. 2002c: 48; Traser & Horváth-Szováti 2006: 333. Distribution in Hungary: Bakony Mts. (Loksa & Rubio 1966a), Bükk Mts. (Loksa 1966). General distribution: Northern and Central Eu- Ecological note: xerothermophilic. rope. Distribution in Hungary: Fertő-Hanság (Traser Folsomides parvulus Stach, 1922 2002a), Kiskunság (, 2002c; Traser & Horváth- Szováti 2006). Folsomides parvulus Stach: Dunger 1975: 20; Traser 1999: 55; Traser 2002a: 264; Traser 2006: 36; Pro- Ecological note: eurytopic, on open habitats. isotoma parvula Stach: Loksa 1991: 272. Isotoma cf. intermedia Schött, 1902 General distribution: cosmopolitan. Isotoma (Desoria) cf. intermedia Schött: Traser 1999: Distribution in Hungary: Aggtelek Karst (Tra- 55. ser 1999), Börzsöny Mts. (Dunger 1975), Fertő- Hanság (Traser 2002a), Gerecse Mts. (Traser Remark: Species inquirenda according to Pota- 2006), Nyírség (Loksa 1991). pow (2001). Ecological note: xerophilous. General distribution: West Palaearctic. Distribution in Hungary: Aggtelek Karst (Tra- Folsomides portucalensis da Gama, 1961: ser 1999). Folsomides portucalensis da Gama: Traser 2002c: 48; Traser & Horváth–Szováti 2004: 60; Traser & Hor- Isotoma riparia (Nicolet, 1842) váth-Szováti 2006: 333. Isotoma riparia (Nicolet): Traser et al. 2006: 72.

General distribution: Europe, Siberia. General distribution: Northern and Central Eu- Distribution in Hungary: Kiskunság (Traser rope. 2002c; Traser & Horváth–Szováti 2004, 2006). Distribution in Hungary: Fertő-Hanság (Traser Ecological note: xerothermophilic. et al. 2006).

49 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ecological note: hygrophilous, mostly at 1966; Loksa & Rubio 1966a; Traser 1980), Bala- springs. tonfelvidék (Loksa 1966), Börzsöny Mts. (Loksa 1966; Dunger 1975), Buda Mts. (Loksa 1956b, Isotoma viridis Bourlet, 1839 1966), Bükk Mts. (Loksa 1956a, 1966, 1977; Varga & Vargha 1992; Traser 2002b), Cserhát Isotoma viridis ab. annulata (Nicolet): Stach 1929: 291; Isotoma viridis ab. violacea (Lie-Pettersen): Stach (Loksa 1978b), Dráva Plain (Loksa 1978a), Fertő- 1929: 291; Isotoma viridis Bourlet: Stach 1929: 290; Hanság (Traser 2002a, 2003; Traser et al. 2006), Szent-Ivány 1940a: 429; Stach 1947: 422, pl. 46: figs Gerecse Mts. (Loksa 1966; Traser 2006), 6-14; Loksa 1966: 382; Dunger 1975: 20; Loksa Hortobágy (Loksa 1983), Keszthely Mts. (Loksa 1978a: 60; Loksa 1983: 73; Loksa 1991: 272; Farkas 1995: 46; Traser 1997: 62; Farkas et al. 1998: 57; 1966), Kiskunság (Loksa 1987; Traser & Csóka Traser 1999: 55; Traser 2000: 68; Dombos 2001: 2041; 2000, 2001; Traser 2002c; Traser & Horváth- Traser 2002a: 264; Traser 2002c: 48; Traser 2003: 155; Szováti 2006), Kőszeg Mts. (Traser & Salló Traser & Horváth–Szováti 2004: 60; Traser et al. 2006: 2004), Mátra Mts. (Loksa 1980), Mecsek Mts. 72; Traser & Horváth-Szováti 2006: 333; Isotoma viridis f. pallida Nicolet: Szent-Ivány 1940a: 429. (Loksa 1966), Mt. Somló (Loksa 1966), Nyírség (Loksa 1991), Őrség (Traser 1995), Pilis Mts. General distribution: Holarctic. (Loksa 1966), Somogy Hills (Farkas et al. 1998), Distribution in Hungary: Aggtelek Karst (Tra- Sopron Mts. (Traser 1981, 1994, 2002a), Szi- ser 1999), around the Lake Balaton (Farkas 1995), getköz (Traser 1997), Vas-Sopron Plain (Traser Balatonfelvidék (Stach 1929, 1947), Börzsöny 1981), Vértes Mts. (Loksa 1966), Villány Mts (Loksa 1966), Visegrád Mts. (Loksa 1966). Mts. (Loksa 1966; Dunger 1975), Dráva Plain Ecological note: parthenogenetic, eurytopic. (Loksa 1978a), Fertő-Hanság (Traser 2000,

2002a, 2003; Traser et al. 2006), Hortobágy (Lok- Isotomiella paraminor Gisin, 1942 sa 1983), Kiskunság (Traser 2002c; Traser & Horváth–Szováti 2004, 2006), Kőszeg Mts. Isotomiella paraminor Gisin: Loksa 1966: 382; Traser (Szent-Ivány 1940a; Loksa 1966), Mezőföld 1980: 20. (Stach 1929, 1947; Dombos 2001), Nyírség (Lok- General distribution: Europe. sa 1991), Pilis Mts. (Stach 1929, 1947), Somogy Distribution in Hungary: Bakony Mts. (Traser Hills (Farkas et al. 1998), Szigetköz (Traser 1980), Bükk Mts. (Loksa 1966). 1997), Visegrád Mts. (Loksa 1966). Ecological note: montane, silvicolous. Ecological note: eurytopic, on open habitats.

Isotomodes Linnaniemi, 1907 Isotomiella Bagnall, 1939

Isotomiella minor (Schäffer, 1896) Isotomodes productus (Axelson, 1906) Isotomiella minor Schäffer: Loksa 1956a: 391; Loksa 1956b: Isotomodes productus (Axelson): Loksa 1966: 382; 203; Loksa & Rubio 1966a: 151; Loksa 1966: 382; Bajomi Dunger 1975: 19; Traser 1981: 133; Traser 1997: 62; 1969: 242; Dunger 1975: 20; Loksa 1977: 209; Loksa 1978b: Traser 1999: 54; Traser 2002a: 264; Traser 2002c: 48; 95; Loksa 1978a: 57; Traser 1980: 20; Loksa 1980: 110; Traser 1981: 133; Loksa 1983: 73; Loksa 1987: 79; Loksa Traser & Horváth-Szováti 2006: 333. 1991: 272; Varga & Vargha 1992: 315; Traser 1994: 102; Farkas 1995: 46; Traser 1995: 45; Traser 1997: 62; Farkas et General distribution: cosmopolitan. al. 1998: 57; Traser 1999: 55; Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; Traser 2002a: 264; Traser 2002b: Distribution in Hungary: Aggtelek Karst (Lok- 160; Traser 2002c: 48; Traser 2003: 155; Traser & Salló sa 1966; Traser 1999), Börzsöny Mts. (Dunger 2004: 230; Traser 2006: 35; Traser & Horváth-Szováti 2006: 1975), Bükk Mts. (Loksa 1966), Fertő-Hanság 333; Traser et al. 2006: 72. (Traser 2002a), Kiskunság (Traser 2002c; Traser General distribution: cosmopolitan. & Horváth-Szováti 2006), Mecsek Mts. (Loksa Distribution in Hungary: Aggtelek Karst (Lok- 1966), Sopron Mts. (Traser 1981), Szigetköz sa 1966; Bajomi 1969; Traser 1999), around the (Traser 1997), Vas-Sopron Plain (Traser 1981). Lake Balaton (Farkas 1995), Bakony Mts. (Loksa Ecological note: xerothermophilic.

50 Dányi & Traser: Checklist of the springtail fauna of Hungary

Isotomodes sexsetosus da Gama, 1963 Isotomurus maculatus (Schäffer, 1896) Isotomodes sexsetosus da Gama: Dunger 1975: 20; Isotomurus palustris var. maculata Schäffer: Stach Traser 2002b: 160; Traser & Horváth-Szováti 2006: 1929: 292. 333; Traser 2006: 35. General distribution: Europe. General distribution: Central Europe. Distribution in Hungary: Mezőföld (Stach 1929). Distribution in Hungary: Börzsöny Mts. (Dun- Ecological note: in gardens and flower pots. ger 1975; Traser 2002b), Bükk Mts. (Traser 2002b), Gerecse Mts. (Traser 2006), Kiskunság Isotomurus palustris (Müller, 1776) (Traser & Horváth-Szováti 2006). Isotoma palustris Müller: Tömösváry 1884a: 55; Ecological note: xerothermophilic. Vellay 1900: 20; Isotomurus palustris (Müller): Stach 1929: 291; Szent-Ivány 1940a: 429; Stach 1947: 452, Isotomodes trisetosus Denis, 1923 pl. 48; Kovács 1953a: 183; Paclt 1960: 157; Dunger 1975: 21; Loksa 1978b: 95; Loksa 1978a: 60; Loksa Isotomodes trisetosus Denis: Loksa 1966: 382. 1980: 110; Loksa 1983: 73; Hornung 1986: 138; Loksa 1987: 79; Loksa 1991: 272; Farkas 1995: 46; Farkas General distribution: From Indonesia to the 1995: 46; Farkas 1996: 287; Farkas 1997: 56; Farkas & Szathmáry 1997: 39; Farkas 1997: 58; Farkas et al. Palaearctic. 1998: 57; Traser 2000: 68; Traser 2002a: 264; Iso- Distribution in Hungary: Bakony Mts. (Loksa tomurus cf. palustris (Müller): Traser et al. 2006: 72. 1966), Balatonfelvidék (Loksa 1966), Börzsöny Mts. (Loksa 1966), Buda Mts. (Loksa 1966), Ge- General distribution: Holarctic. recse Mts. (Loksa 1966), Keszthely Mts. (Loksa Distribution in Hungary: around the Lake Ba- 1966), Mecsek Mts. (Loksa 1966), Mt. Somló laton (Farkas 1995; Farkas 1997), Balatonfelvidék (Loksa 1966), Pilis Mts. (Loksa 1966), Vértes (Stach 1929; Farkas 1995, 1997; Farkas & Mts. (Loksa 1966), Villány Mts (Loksa 1966), Szathmáry 1997), Börzsöny Mts. (Dunger 1975), Visegrád Mts. (Loksa 1966). Budapest (Stach 1929), Cserhát (Loksa 1978b), Ecological note: xerothermophilic. Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser 2000, 2002a; Traser et al. 2006), Hortobágy Isotomurus Börner, 1903 (Loksa 1983), Kis-Balaton (Farkas 1996, 1997), Kiskunság (Stach 1929, 1947; Paclt 1960; Hor- Isotomurus balteatus (Reuter, 1876) nung 1986; Loksa 1987), Kőszeg Mts. (Szent- Isotomurus palustris var. balteata (Reuter): Stach Ivány 1940a), Mátra Mts. (Loksa 1980), Me- 1929: 292. zőföld (Stach 1929, 1947), Nyírség (Stach 1929; Kovács 1953a; Loksa 1991), Somogy Hills (Far- General distribution: Europe. kas et al. 1998), Sopron Mts. (Traser 2002a), Vác- Distribution in Hungary: Balatonfelvidék Pest Duna Valley (Tömösváry 1884a; Vellay (Stach 1929). 1900; Stach 1929). Ecological note: hygrophilous. Ecological note: hygrophilous.

Isotomurus fucicolus Reuter, 1891 Isotomurus plumosus Bagnall, 1940 Isotomurus plumosous (sic!) Bagnall: Dunger 1975: Isotomurus palustris var. fucicola (Reuter): Stach 21; Loksa 1983: 73; Traser 1997: 62; Isotomurus 1929: 292. plumosus Bagnall: Traser 2002a: 264.

General distribution: Europe. General distribution: Europe. Distribution in Hungary: Balatonfelvidék Distribution in Hungary: Börzsöny Mts. (Dun- (Stach 1929). ger 1975), Fertő-Hanság (Traser 2002a), Hor- Ecological note: litter dwelling. tobágy (Loksa 1983), Szigetköz (Traser 1997).

51 Dányi & Traser: Checklist of the springtail fauna of Hungary

Ecological note: hygrophilous, mostly at early 1947), Dráva Plain (Loksa 1978a), Fertő-Hanság spring. (Traser 2002a; Traser et al. 2006), Kis-Balaton (Farkas 1996, 1997), Kiskunság (Vellay 1900; Isotomurus prasinus (Reuter, 1891) Stach 1929, 1947; Paclt 1960), Somogy Hills Isotomurus palustris var. prasina Reuter: Stach 1929: (Farkas et al. 1998). 292; Isotoma palustris f. prasina Reuter: Szent-Ivány Ecological note: neustonic. 1940a: 429; 1940b: 119; Isotomurus prasinus (Reuter): Traser et al. 2006: 72. Pachyotoma granulata (Stach, 1947)

General distribution: Siberia. Proisotoma granulosa Stach (sic!): Loksa 1956a: 415. Distribution in Hungary: Balatonfelvidék (Stach 1929; Szent-Ivány 1940a, 1940b), Fertő- General distribution: Eastern Europe. Hanság (Traser et al. 2006), Kőszeg Mts. (Szent- Distribution in Hungary: Bükk Mts. (Loksa Ivány 1940a, 1940b). 1956a). Ecological note: hygrophilous. Ecological note: mostly in lichens.

Isotomurus stuxbergi (Tullberg, 1876) Parisotoma Bagnall, 1940

Isotomurus ciliatus Stach: Stach 1947: 476, pl. 53. Parisotoma notabilis (Schäffer, 1896)

Isotoma notabilis Schäffer: Stach 1929: 291; Dudich General distribution: Eastern Europe. 1932: 48; Stach 1947: 374, pl. 40; Loksa 1956a: 391; Distribution in Hungary: Balatonfelvidék Loksa 1956b: 203; Loksa 1962: 63; Loksa 1966: 382; (Stach 1947). Loksa & Rubio 1966a: 152; Bajomi 1969: 242; Dunger Ecological note: neustonic, along streams. 1975: 20; Loksa 1977: 209; Loksa 1978b: 95; Loksa 1978a: 60; Loksa 1980: 110; Traser 1980: 20, fig. 2a; Traser 1981: 133; Loksa 1983: 73; Loksa 1987: 79; Micranurophorus Bernard, 1977 Loksa 1991: 272; Varga 1991: 85; Traser 1994: 102; Traser 1995: 45; Farkas 1995: 46; Traser 1997: 62; Micranurophorus cf. musci Bernard, 1977 Farkas 1997: 56; Farkas et al. 1998: 57; Traser 1999: 55; Traser & Csóka 2000: 76; Traser & Csóka 2001: Micranurophorus cf. schalleri Christian: Traser & 233; Parisotoma notabilis (Schäffer): Traser 2002c: Horváth-Szováti 2006: 333; Traser 2002c: 48. 48; Traser 2002a: 264; Traser 2003: 155; Traser & Salló 2004: 230; Traser 2006: 36; Traser & Horváth- General distribution: Europe, North America. Szováti 2006: 333; Traser et al. 2006: 72. Distribution in Hungary: Kiskunság (Traser General distribution: cosmopolitan. 2002c; Traser & Horváth-Szováti 2006). Distribution in Hungary: Aggtelek Karst (Du- Ecological note: psammophilous species. dich 1932; Loksa 1966; Bajomi 1969; Traser 1999), around the Lake Balaton (Farkas 1995, Pachyotoma Bagnall, 1949 1997), Bakony Mts. (Loksa 1966; Loksa & Rubio 1966a; Traser 1980), Balatonfelvidék (Stach Pachyotoma crassicauda (Tullberg, 1871) 1929, 1947; Loksa 1966; Farkas 1995, 1997), Isotoma crassicauda Tullberg: Vellay 1900: 20; Börzsöny Mts. (Loksa 1966; Dunger 1975), Buda Proisotoma crassicauda (Tullberg): Stach 1929: 289; Mts. (Loksa 1956b, 1966), Bükk Mts. (Loksa Paclt 1960: 156; Farkas 1995: 46; Farkas 1996: 287; 1956a, 1962, 1966, 1977; Varga 1991), Cserhát Farkas 1997: 58; Farkas et al. 1998: 57; Ballistura crassicauda (Tullberg): Stach 1947: 242, pl. 28; (Loksa 1978b), Dráva Plain (Loksa 1978a), Fertő- Pachyotoma crassicauda (Tullberg): Loksa 1978a: 57; Hanság (Traser 2002a, 2003; Traser et al. 2006), Traser 2002a: 264; Traser et al. 2006: 72. Gerecse Mts. (Loksa 1966; Traser 2006), Hor- tobágy (Loksa 1983), Keszthely Mts. (Loksa General distribution: Palaearctic. 1966), Kiskunság (Loksa 1987; Traser & Csóka Distribution in Hungary: around the Lake Ba- 2000, 2001; Traser 2002c; Traser & Horváth- laton (Farkas 1995), Balatonfelvidék (Stach 1929, Szováti 2006), Kőszeg Mts. (Traser & Salló

52 Dányi & Traser: Checklist of the springtail fauna of Hungary

2004), Mátra Mts. (Loksa 1980), Mecsek Mts. 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság (Loksa 1966), Mezőföld (Stach 1929, 1947), Mt. (Traser 2002a), Kis-Balaton (Farkas 1996, 1997), Somló (Loksa 1966), Nyírség (Loksa 1991), Őr- Kiskunság (Loksa 1987; Traser 2002c; Traser & ség (Traser 1995), Pilis Mts. (Loksa 1966), So- Horváth-Szováti 2006), Kőszeg Mts. (Szent-Ivány mogy Hills (Farkas et al. 1998), Sopron Mts. 1940a), Somogy Hills (Farkas et al. 1998), (Traser 1981, 1994, 2002a), Szigetköz (Traser Szigetköz (Traser 1997), Vas-Sopron Plain (Tra- 1997), Vas-Sopron Plain (Traser 1981; Traser ser 1981). 1981), Vértes Mts. (Loksa 1966), Villány Mts Ecological note: thermophilic, frequently in compost. (Loksa 1966), Visegrád Mts. (Loksa 1966).

Ecological note: eurytopic, very common. Proisotoma ripicola Linnaniemi, 1912 Proisotoma Börner, 1901 Proisotoma ripicola Linnaniemi: Stach 1929: 288.

Proisotoma brevidens Stach, 1947 General distribution: Europe. Proisotoma brevidens Stach: Dunger 1975: 20; Traser Distribution in Hungary: Balatonfelvidék 2002a: 264. (Stach 1929). Ecological note: psammophilous species. General distribution: Eastern Europe. Distribution in Hungary: Börzsöny Mts. (Dun- Proisotoma subminuta Denis, 1931 ger 1975), Szigetköz (Traser 2002a). Ecological note: mostly in mulder. Proisotoma subminuta Denis: Loksa 1980: 108, figs 2d-e.

Proisotoma minima (Absolon, 1901) General distribution: Holarctic. Distribution in Hungary: Mátra Mts. (Loksa Proisotoma minima (Absolon): Loksa 1978a: 57; Traser 1999: 55; Traser 2002a: 264; Traser & Horváth- 1980). Szováti 2006: 333. Ecological note: mostly in compost.

General distribution: Holarctic. Pseudanurophorus Stach, 1922 Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Dráva Plain (Loksa 1978a), Fertő- Pseudanurophorus boerneri Stach, 1922 Hanság (Traser 2002a), Kiskunság (Traser & Pseudanurophorus boerneri (Stach): Traser 2002a: Horváth-Szováti 2006). 264. Ecological note: eurytopic. General distribution: Eastern Europe. Proisotoma minuta (Tullberg, 1871) Distribution in Hungary: Fertő-Hanság (Traser 2002a). Proisotoma minuta (Tullberg): Szent-Ivány 1940a: 428; Loksa & Rubio 1966a: 152; Dunger 1975: 20; Ecological note: soil dwelling species. Loksa 1978b: 95; Loksa 1978a: 57; Traser 1980: 20; Traser 1981: 134; Loksa 1987: 79; Farkas 1995: 46; Pseudisotoma Handschin, 1924 Farkas 1996: 287; Traser 1997: 62; Farkas 1997: 56; Farkas et al. 1998: 57; Traser 1999: 55; Traser 2002a: 264; Traser 2002c: 48; Traser & Horváth-Szováti Pseudisotoma sensibilis (Tullberg, 1876) 2006: 333. Isotoma (Pseudisotoma) sensibilis (Tullberg): Dunger 1975: 20; Pseudisotoma sensibilis (Tullberg): Traser General distribution: cosmopolitan. 1999: 55. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), around the Lake Balaton (Farkas 1995, General distribution: Holarctic. 1997), Bakony Mts. (Loksa & Rubio 1966a; Tra- Distribution in Hungary: Aggtelek Karst (Tra- ser 1980), Balatonfelvidék (Farkas 1995, 1997), ser 1999), Börzsöny Mts. (Dunger 1975). Börzsöny Mts. (Dunger 1975), Cserhát (Loksa Ecological note: corticophilous.

53 Dányi & Traser: Checklist of the springtail fauna of Hungary

Subisotoma Stach, 1947 General distribution: Central Europe. Distribution in Hungary: Sopron Mts. (Traser Subisotoma pusilla (Schäffer, 1900) 1994; Traser 2002a). Folsomides pusillus (Schäffer): Loksa & Rubio 1966a: Ecological note: in montane habitats 151; Proisotoma pusilla (Schäffer): Dunger 1975: 20; Loksa 1991: 272; Subisotoma pusilla (Schäffer): Tetracanthella wahlgreni Axelson, 1907 Traser 1999: 55. Tetracanthella wahlgreni Axelson: Stach 1929: 286; General distribution: Palaearctic. Szent-Ivány 1940a: 428; Stach 1947: 42, pl. 2: figs 1-6; Loksa 1956a: 395; Tetracanthella cf. wahlgreni Distribution in Hungary: Aggtelek Karst (Tra- Linnaniemi (sic!): Traser 1999: 54. ser 1999), Bakony Mts. (Loksa & Rubio 1966a), Börzsöny Mts. (Dunger 1975), Nyírség (Loksa General distribution: Palaearctic. 1991). Ecological note: bryophilous, silvicolous. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Buda Mts. (Stach 1929; Szent-Ivány Tetracanthella Schött, 1891 1940a; Stach 1947), Bükk Mts. (Loksa 1956a), Kőszeg Mts. (Szent-Ivány 1940a). Tetracanthella franzi Cassagnau, 1959 Ecological note: in xerophilous moss and lichen. Tetracanthella franzi Cassagnau: Loksa 1980: 110; Traser 1995: 45. Vertagopus Börner, 1906 General distribution: Europe, Alps. Distribution in Hungary: Mátra Mts. (Loksa Vertagopus arboreus (Linnaeus, 1758) 1980), Őrség (Traser 1995). Isotoma (Vertagopus) arborea (Linnaeus): Stach 1929: Ecological note: in litter and moss. 289; Dunger 1975: 20; Vertagopus arborea (sic!) (Linnaeus): Stach 1947: 338, pl. 37: figs 9-11; Traser Tetracanthella hystrix Cassagnau, 1959 2002a: 264.

Tetracanthella hystrix Cassagnau: Dunger 1975: 19. General distribution: Holarctic. Distribution in Hungary: Börzsöny Mts. (Dun- General distribution: Eastern Europe. ger 1975), Cserhát (Stach 1929, 1947), Vas-Sop- Distribution in Hungary: Börzsöny Mts. (Dun- ron Plain (Traser 2002a). ger 1975). Ecological note: corticophilous. Ecological note: rare species.

Vertagopus cinereus (Nicolet, 1841) Tetracanthella pilosa Schött, 1891 Isotoma cinerea Nicolet: Tömösváry 1884a: 55; Vellay Tetracanthella pilosa Schött: Stach 1929: 286; Loksa 1900: 20; Loksa & Rubio 1966a: 152; Isotoma 1966: 381; Loksa & Rubio 1966a: 151. (Vertagopus) cinerea (Nicolet): Stach 1929: 289; Dunger 1975: 20; Vertagopus cinerea (sic!) (Nicolet): General distribution: Eastern Europe. Traser 1999: 54; Vertagopus cinereus (Nicolet): Traser Distribution in Hungary: Bakony Mts. (Loksa 2002a: 264. & Rubio 1966a), Budapest (Stach 1929), Bükk Mts. (Loksa 1966), Mecsek Mts. (Loksa 1966), General distribution: Europe. Villány Mts (Loksa 1966). Distribution in Hungary: Aggtelek Karst (Tra- Ecological note: bryophilous, silvicolous. ser 1999), Bakony Mts. (Loksa & Rubio 1966a), Börzsöny Mts. (Dunger 1975), Budapest (Tömös- Tetracanthella stachi Cassagnau, 1959 váry 1884a; Vellay 1900; Stach 1929), Fertő- Hanság (Traser 2002a), Sopron Mts. (Traser Tetracanthella stachi Cassagnau: Traser 1994: 102; 2002a), Szigetköz (Traser 2002a). Traser 2002a: 264. Ecological note: corticophilous.

54 Dányi & Traser: Checklist of the springtail fauna of Hungary

Vertagopus westerlundi (Reuter, 1898) 1929: 293; Szent-Ivány 1940a: 435; Kovács 1953a: 183; Loksa 1991: 273; (Pogonognathellus) Vertagopus westerlundi (Reuter): Traser 1999: 54. flavescens (Tullb.): Paclt 1960: 157; Loksa & Rubio 1966a: 154; Loksa 1978a: 61; Traser 1999: 57; Tomo- General distribution: Arctoalpine. cerus flavescens Tullberg: Szent-Ivány 1941: 24; Distribution in Hungary: Aggtelek Karst (Tra- Loksa 1966: 385; Loksa 1977: 211; Loksa 1978b: 95; Loksa 1980: 111; Loksa 1991: 273; Traser 1997: 62; ser 1999). Tomocerus flavescens Lubbock (sic!): Tóth 1942: 398; Ecological note: bryophilous species. Pogonognathellus flavescens (Tullberg 1871): Dunger 1975: 23; Traser 2002a: 266; Traser 2003: 156; Tomo- Family: Oncopoduridae cerus cf. flavescens Tullberg: Traser 1981: 135.

Oncopodura Carl & Lebedinsky, 1905 General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Tra- Oncopodura crassicornis Shoebotham, 1911 ser 1999; Loksa 1966), Bakony Mts. (Loksa & Oncopodura crassicornis Shoebotham: Loksa 1956a: Rubio 1966a), Börzsöny Mts. (Tömösváry 1884a; 391; Loksa 1962: 63; Loksa 1966: 385; Bajomi 1969: Vellay 1900; Stach 1929; Dunger 1975), Buda 242; Dunger 1975: 24; Loksa 1977: 211; Loksa 1978b: 95; Traser 1995: 46; Traser 1999: 57; Traser 2002a: Mts. (Stach 1929; Loksa 1966), Budapest (Tö- 266; Traser 2006: 36; Traser et al. 2006: 73. mösváry 1884a; Vellay 1900; Stach 1929), Bükk Mts. (Loksa 1966, 1977), Cserhát (Loksa 1978b), General distribution: Europe. Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser Distribution in Hungary: Aggtelek Karst (Lok- 2002a, 2003), Gerecse Mts. (Loksa 1966), Gödöl- sa 1966; Bajomi 1969; Traser 1999), Bakony Mts. lő Hills (Tömösváry 1884a; Vellay 1900; Stach (Loksa 1966), Balatonfelvidék (Loksa 1966), 1929), Keszthely Mts. (Loksa 1966), Kiskunság Börzsöny Mts. (Loksa 1966; Dunger 1975), Buda (Vellay 1900; Stach 1929), Kőszeg Mts. (Szent- Mts. (Loksa 1966), Bükk Mts. (Loksa 1956a, Ivány 1940a; Tóth 1942), Mátra Mts. (Stach 1962, 1966, 1977), Cserhát (Loksa 1978b), Fertő- 1929; Szent-Ivány 1941; Loksa 1980), Mecsek Hanság (Traser et al. 2006), Gerecse Mts. (Loksa Mts. (Tömösváry 1884a; Vellay 1900; Stach 1966; Traser 2006), Keszthely Mts. (Loksa 1966), Mecsek Mts. (Loksa 1966), Mt. Somló (Loksa 1929; Paclt 1960; Loksa 1966), Mezőföld (Stach 1966), Őrség (Traser 1995), Pilis Mts. (Loksa 1929), Nyírség (Stach 1929; Kovács 1953a; 1966), Sopron Mts. (Traser 2002a), Vértes Mts. Loksa 1991), Pilis Mts. (Loksa 1966), Sopron (Loksa 1966), Villány Mts (Loksa 1966), Vi- Mts. (Traser 2002a), Szigetköz (Traser 1997, segrád Mts. (Loksa 1966). 2002a), Vác-Pest Duna Valley (Tömösváry Ecological note: edaphic, troglophilous. 1884a; Vellay 1900; Stach 1929), Vas-Sopron Plain (Traser 1981), Villány Mts (Loksa 1966), Oncopodura egerszoegensis Loksa, 1961 Visegrád Mts. (Loksa 1966), Zala Hills (Stach 1929), Zemplén Mts. (Tömösváry 1882; Vellay Oncopodura égerszögensis Loksa: Loksa 1961b: 224, 1900; Stach 1929). figs 9-16. Ecological note: hygrophilous, troglophilous.

General distribution: Central Europe. Pogonognathellus longicornis (Müller, 1776) Distribution in Hungary: Aggtelek Karst (Lok- sa 1961b). Tomocerus (Pogonognathus (sic!)) longicornis (Mül- Ecological note: troglophilous. ler): Stach 1929: 293; Szent-Ivány 1938: 92; Szent-Ivány 1940a: 435; Tomocerus longicornis Tull- berg (sic!): Tóth 1942: 398; Tomocerus (Pogono- Family: Tomoceridae gnathellus) longicornis (Müller): Paclt 1960: 157; Tomocerus longicornis (Müller): Traser 1980: 22; Pogonognathellus Paclt, 1944 Traser 1981: 133; Loksa 1983: 75; Loksa 1991: 273; Farkas 1995: 46; Farkas 1996: 287; Farkas & Szath- Pogonognathellus flavescens (Tullberg, 1871) máry 1997: 39; Farkas 1997: 58; Traser 2002a: 268; Varga & Naár 2002: 115; Varga et al. 2002b: 526; Tomocerus plumbeus Linnaeus: Tömösváry 1882: 124; Tomocerus (Pogonognathellus) longicornis (Müller): Tömösváry 1884a: 56; Vellay 1900: 21; Tomocerus Traser 1999: 57; Tomacerus (sic!) longicornis Müller: (Pogonognathus (sic!)) flavescens Tullberg: Stach Varga 1992: 172; Varga & Oldal 1998: 86.

55 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Palaearctic. 57; Traser 2002a: 266; Traser 2003: 156; Traser & Distribution in Hungary: Aggtelek Karst (Tra- Salló 2004: 230; Traser et al. 2006: 73; Tomocerus ser 1999), around the Lake Balaton (Farkas 1995, minor Tullberg (sic!): Loksa 1956a: 409; Tomocerus (Tomocerus) minor (Lubbock): Loksa & Rubio 1966a: 1997), Bakony Mts. (Traser 1980), Balatonfelvi- 154; Loksa 1978a: 61; Traser 1999: 57. dék (Farkas 1995, 1997; Farkas & Szathmáry 1997), Bükk Mts. (Varga 1992; Varga & Oldal 1998; Varga & Naár 2002; Varga et al. 2002b), General distribution: Holarctic. Fertő-Hanság (Paclt 1960; Traser 2002a), Horto- Distribution in Hungary: Aggtelek Karst bágy (Loksa 1983), Kis-Balaton (Farkas 1996, (Stach 1929; Dudich 1932; Traser 1999), Bakony 1997), Kőszeg Mts. (Szent-Ivány 1940a; Tóth Mts. (Loksa & Rubio 1966a; Traser 1980), Bör- 1942), Mátra Mts. (Szent-Ivány 1938), Mecsek zsöny Mts. (Dunger 1975), Bükk Mts. (Loksa Mts. (Stach 1929), Nyírség (Loksa 1991), Pilis 1962, 1956a), Dráva Plain (Loksa 1978a), Fertő- Mts. (Stach 1929), Sopron Mts. (Traser 1981), Vas-Sopron Plain (Traser 1981), Zselic (Paclt Hanság (Traser 2002a, 2003; Traser et al. 2006), 1960). Kőszeg Mts. (Szent-Ivány 1940a; Traser & Salló Ecological note: silvicolous. 2004), Nyírség (Loksa 1991), Őrség (Traser 1995), Somogy Hills (Farkas et al. 1998), Sopron Tomocerina Yosii, 1955 Mts. (Traser 1994, 2002a). Ecological note: hygrophilous, troglophilous. Tomocerina minuta (Tullberg, 1876) Tomocerus minutus Tullberg: Szent-Ivány 1940a: 435; Tomocerus vulgaris (Tullberg, 1871) Dunger 1975: 24; Traser 1995: 46; Tomocerus (Tomo- cerus) cf. minutus (Tullberg): Traser 1999: 57; Tomo- Tomocerus niger Bourlet: Vellay 1900: 21; Tomocerus cerus cf. minutus (Tullberg): Traser 2002a: 266. vulgaris (Tullberg): Stach 1929: 292; Dudich 1932: 49; Szent-Ivány 1940a: 435; Kovács 1953a: 183; Loksa 1959b: 70; Loksa 1962: 73; Dunger 1975: 24; Loksa General distribution: Palaearctic. 1978b: 95; Loksa 1983: 75; Loksa 1991: 273; Varga & Distribution in Hungary: Aggtelek Karst (Tra- Vargha 1992: 315; Farkas 1996: 287; Traser 1997: 62; ser 1999), Börzsöny Mts. (Dunger 1975), Kőszeg Farkas & Szathmáry 1997: 39; Farkas 1997: 58; Traser Mts. (Szent-Ivány 1940a), Őrség (Traser 1995), 2002a: 266; Traser 2002c: 48; Tomoceris (sic!) vul- Sopron Mts. (Traser 2002a). garis (Tullberg): Farkas 1995: 46; Farkas et al. 1998: 57; Tomocerus (Tomocerus) vulgaris (Tullberg): Traser Tomocerus Nicolet, 1842 1999: 57.

Tomocerus baudoti Denis, 1932 General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Du- Tomocerus (Tomocerus) baudoti Denis: Loksa 1978a: 61; Tomocerus baudoti Denis: Loksa 1991: 273; Traser dich 1932; Traser 1999), around the Lake Balaton 1995: 46; Traser 2002a: 266; Traser & Salló 2004: (Farkas 1995), Balatonfelvidék (Farkas & Szath- 230; Tomocerus cf. baudoti (Denis): Traser et al. 2006: máry 1997), Börzsöny Mts. (Dunger 1975), Buda- 73. pest (Stach 1929), Bükk Mts. (Loksa 1962; Varga & Vargha 1992), Cserhát (Loksa 1959b, 1978b), General distribution: Europe. Fertő-Hanság (Traser 2002a), Hortobágy (Loksa Distribution in Hungary: Dráva Plain (Loksa 1983), Kis-Balaton (Farkas 1996, 1997), Kiskun- 1978a), Fertő-Hanság (Traser et al. 2006), Kőszeg ság (Vellay 1900; Stach 1929; Traser 2002c), Kő- Mts. (Traser & Salló 2004), Nyírség (Loksa szeg Mts. (Szent-Ivány 1940a), Mezőföld (Stach 1991), Őrség (Traser 1995), Sopron Mts. (Traser 1929), Nyírség (Vellay 1900; Stach 1929; Kovács 2002a). 1953a; Loksa 1991), Somogy Hills (Farkas et al.

1998), Szerencs Hills (Stach 1929), Szigetköz Tomocerus minor (Lubbock, 1862) (Traser 1997, 2002a). Tomocerus minor (Lubbock): Stach 1929: 293; Dudich 1932: 49; Szent-Ivány 1940a: 434; Loksa 1962: 68; Suborder: NEELIPLEONA Dunger 1975: 24; Traser 1980: 20; Loksa 1991: 273; Traser 1994: 100; Traser 1995: 46; Farkas et al. 1998: Family: Neelidae

56 Dányi & Traser: Checklist of the springtail fauna of Hungary

Megalothorax Willem, 1900 Neelides minutus (Folsom, 1901) Neelides minutus (Folsom): Loksa 1959b: 70; Loksa Megalothorax incertus Börner, 1903 1960: 49; Loksa 1961a: 257; Traser 2002a: 266; Traser 2003: 156; Traser et al. 2006: 73. Neelus (Megalothorax) incertus Börner: Loksa & Ru- bio 1966a: 154; Megalothorax incertus Börner: Dunger 1975: 24; Traser 1999: 58; Traser 2002a: 266. General distribution: Holarctis, South Austra- lia. General distribution: Holarctis, Australia. Distribution in Hungary: Balatonfelvidék Distribution in Hungary: Aggtelek Karst (Tra- (Loksa 1960, 1961a), Cserhát (Loksa 1959b), Fer- ser 1999), Bakony Mts. (Loksa & Rubio 1966a), tő-Hanság (Traser 2002a, 2003; Traser et al. Börzsöny Mts. (Dunger 1975), Fertő-Hanság 2006), Sopron Mts. (Traser 2002a). (Traser 2002a). Ecological note: euedaphic, troglophilous. Neelus Folsom, 1896

Megalothorax minimus Willem, 1900 Neelus murinus Folsom, 1896 Megalothorax minimus Willem: Loksa 1956b: 207; Neelus murinus Folsom: Gebhardt 1934: 140. Loksa 1956a: 391; Loksa 1961b: 228; Loksa 1962: 68; Dunger 1975: 24; Traser 1981: 133; Traser 1994: 102; General distribution: Holarctis, Central Ame- Traser 1995: 46; Traser 1997: 62; Traser 1999: 58; rica, India, Australia. Traser 2002a: 267; Traser 2003: 156; Traser & Salló 2004: 230; Traser & Horváth-Szováti 2006: 334; Distribution in Hungary: Mecsek Mts. (Geb- Traser et al. 2006: 73; Traser 2006: 37; Neelus (Me- hardt 1934). galothorax) minimus Willem: Loksa & Rubio 1966a: Ecological note: edaphic, troglophilous. 154; Loksa 1978b: 95; Traser 1980: 20; Neelus minimus Willem: Loksa 1966: 385; Loksa 1977: 211; Suborder: Farkas et al. 1998: 57. Family: Bourletiellidae General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Lok- Banks, 1899 sa 1961b, 1966; Traser 1999), Bakony Mts. (Lok- sa & Rubio 1966a; Loksa 1966; Traser 1980), Bourletiella (Bourletiella) arvalis (Fitch, 1863) Balatonfelvidék (Loksa 1966), Börzsöny Mts. Smynthurus (sic!) luteus Lubbock: Tömösváry 1883: (Dunger 1975; Loksa 1966), Buda Mts. (Loksa 36; Tömösváry 1884a: 57; Vellay 1900: 20; Bourle- 1956b, 1966), Bükk Mts. (Loksa 1956a, 1962, tiella lutea (Lubbock): Stach 1929: 308; Bourletiella 1966, 1977), Cserhát (Loksa 1978b), Fertő-Han- arvalis (Fitch): Stach 1956: 143; Bourletiella lutea ság (Traser 2002a, 2003; Traser et al. 2006), Lubbock: Varga 1991: 85; Traser 1995: 47. Gerecse Mts. (Loksa 1966; Traser 2006), Keszt- hely Mts. (Loksa 1966), Kiskunság (Traser & General distribution: Holarctic. Distribution in Hungary: Aggtelek Karst (Tö- Horváth-Szováti 2006), Kőszeg Mts. (Traser & mösváry 1883; Vellay 1900; Stach 1929), Borsod Salló 2004), Mecsek Mts. (Loksa 1966), Őrség Depression (Vellay 1900; Stach 1929), Budapest (Traser 1995), Pilis Mts. (Loksa 1966), Somogy (Tömösváry 1883, 1884a; Vellay 1900; Stach Hills (Farkas et al. 1998), Sopron Mts. (Traser 1929), Bükk Mts. (Varga 1991), Cserhát (Stach 1981, 1994, 2002a), Szigetköz (Traser 1997), 1929, 1956), Gödöllő Hills (Tömösváry 1883, Vas-Sopron Plain (Traser 1981), Vértes Mts. 1884a; Vellay 1900; Stach 1929), Kiskunság (Loksa 1966), Villány Mts (Loksa 1966), Vise- (Stach 1929, 1956), Maros-Körös Region (Tö- mösváry 1883; Vellay 1900; Stach 1929), Mátra- grád Mts. (Loksa 1966). alja (Tömösváry 1883; Vellay 1900; Stach 1929), Ecological note: euedaphic, troglophilous. Mecsek Mts. (Tömösváry 1883; Vellay 1900; Stach 1929), Őrség (Traser 1995), Sajó Valley Neelides Caroli, 1912 (Tömösváry 1883), Vác-Pest Duna Valley (Tö

57 Dányi & Traser: Checklist of the springtail fauna of Hungary

mösváry 1883, 1884a; Vellay 1900; Stach 1929), Distribution in Hungary: Bakony Mts. (Loksa Visegrád Mts. (Vellay 1900; Stach 1929), Zem- & Rubio 1966a). plén Mts. (Vellay 1900; Stach 1929). Ecological note: xerotolerant. Ecological note: on open habitats. Deduterosminthurus fenyesi Stach, 1920 Bourletiella (Bourletiella) hortensis (Fitch, 1863) fényesi Stach: Stach 1926a: 82; Stach 1929: 310; Deuterosminthurus pallipes (Bour- (Fitch): Stach 1956: 149; Loksa let): Traser 2002a: 268. 1983: 76; Loksa 1987: 80; Traser 1997: 62; Traser 2002a: 268. General distribution: Holarctic. Distribution in Hungary: Cserhát (Stach 1926 General distribution: Holarctic. a, 1929), Fertő-Hanság (Traser 2002a). Distribution in Hungary: Fertő-Hanság (Traser Ecological note: in dry grass habitats. 2002a), Hortobágy (Loksa 1983), Kiskunság (Stach 1956; Loksa 1987), Szigetköz (Traser Deuterosminthurus pallipes (Bourlet, 1843) 1997). Bourletiella pallipes Bourlet: Loksa 1966: 386; Loksa & Rubio 1966a: 156. Bourletiella (Bourletiella) viridescens Stach, 1920 General distribution: Holarctic. Bourletiella viridescens (Stach): Traser 2002a: 268. Distribution in Hungary: Bakony Mts. (Loksa & Rubio 1966a), Vértes Mts. (Loksa 1966). General distribution: Europe. Distribution in Hungary: Fertő-Hanság (Traser Deuterosminthurus repandus Ågren, 1903 2002a), Szigetköz (Traser 2002a). Deuterosminthurus repandus (Ågren): Szent-Ivány Ecological note: in open dry habitats. 1940a: 436; Dunger 1975: 25; Traser 1995: 47; Dom- bos 2001: 2041; Deuterosminthurus pallipes f. repadus Deuterosminthurus Börner, 1901 (Ågren): Traser 2002a: 268.

General distribution: cosmopolitan. Deuterosminthurus bicinctus Koch, 1840 Distribution in Hungary: Börzsöny Mts. (Dun- Deuterosminthurus bicinctus Koch: Szent-Ivány 1940 ger 1975), Fertő-Hanság (Traser 2002a), Kőszeg a: 436; Traser 1995: 47; Traser 2002a: 268; Traser et Mts. (Szent-Ivány 1940a), Mezőföld (Dombos al. 2006: 73; Bourletiella bicinctus (sic!) (Koch): 2001), Őrség (Traser 1995). Loksa & Rubio 1966a: 156; Deuterosminthurus bi- cinctus var. flava Gisin: Dunger 1975: 25; Bourletiella bicincta (Koch): Loksa 1991: 273; Deuterosminthurus Deuterosminthurus sulphureus (Koch, 1840) flava Gisin: Traser 1995: 47; Traser 1997: 62. Deuterosminthurus sulphureus (Koch): Traser 2002a: 268. General distribution: Palaearctis, Australia. Distribution in Hungary: Bakony Mts. (Loksa General distribution: Europe. & Rubio 1966a), Börzsöny Mts. (Dunger 1975), Distribution in Hungary: Fertő-Hanság (Traser Fertő-Hanság (Traser 2002a; Traser et al. 2006), 2002a), Szigetköz (Traser 2002a). Kőszeg Mts. (Szent-Ivány 1940a), Nyírség (Loksa 1991), Őrség (Traser 1995), Sopron Mts. (Traser Fasciosminthurus Gisin, 1960 2002a), Szigetköz (Traser 1997). Ecological note: mostly in forests. Fasciosminthurus circumfasciatus (Stach, 1956)

Fasciosminthurus cf. circumfasciatus (Stach): Dunger Deuterosminthurus flavus (Gisin, 1946) 1975: 26; Bourletiella (Fasciosminthurus) circumfasci- Bourletiella flava Gisin: Loksa & Rubio 1966a: 156. ata (Stach): Loksa 1978a: 62.

General distribution: Europe. General distribution: Europe, Central Asia.

58 Dányi & Traser: Checklist of the springtail fauna of Hungary

Distribution in Hungary: Börzsöny Mts. (Dun- General distribution: Palaearctic. ger 1975), Dráva Plain (Loksa 1978a). Distribution in Hungary: around the Lake Ba- laton (Farkas 1995), Börzsöny Mts. (Dunger Fasciosminthurus strigatus (Stach, 1922) 1975), Dráva Plain (Loksa 1978a), Mátra Mts. Fasciosminthurus strigatus strigatus (Stach): Traser (Loksa 1980), Nyírség (Loksa 1991), Őrség (Tra- 2002b: 160; Fasciosminthurus strigatus (Stach): Traser ser 1995), Somogy Hills (Farkas et al. 1998). & Horváth-Szováti 2006: 334; Traser et al. 2006: 73. Ecological note: hygrophilous.

General distribution: Central Europe. Heterosminthurus linnaniemii (Stach, 1920) Distribution in Hungary: Börzsöny Mts. (Tra- Heterosminthurus linnaniemii (Stach): Traser 1995: ser 2002b), Bükk Mts. (Traser 2002b), Fertő- 47; Traser 2002a: 268. Hanság (Traser et al. 2006), Kiskunság (Traser & Horváth-Szováti 2006). General distribution: Europe, East Palaearctic. Distribution in Hungary: Fertő-Hanság (Traser Fasciosminthurus virgulatus (Skorikow, 1899) 2002a), Őrség (Traser 1995). Fasciosminthurus virgulatus (Skorikow): Traser 2002a: 268. Heterosminthurus novemlineatus (Tullberg, 1871) General distribution: Southern and Eastern Deuterosminthurus novemlineatus (Tullberg): Stach Europe. 1929: 309; Heterosminthurus novemlineatus (Tulberg): Distribution in Hungary: Fertő-Hanság (Traser Stach 1956: 193, pl. 24; Traser 2002a: 268; Bourle- 2002a). tiella (Heterosminthurus) novemlineata (Tullberg): Loksa 1978a: 62; Bourletiella novemlineata (Tullberg): Loksa 1987: 80; Varga & Vargha 1992: 315. Heterosminthurus Stach, 1955

General distribution: Europe, Russia. Heterosminthurus bilineatus (Bourlet, 1842) Distribution in Hungary: Balatonfelvidék Heterosminthurus bilineatus (Bourlet): Traser 1995: (Stach 1929, 1956), Bükk Mts. (Varga & Vargha 47; Traser 2002a: 268; Traser 2003: 156; Traser et al. 1992), Dráva Plain (Loksa 1978a), Fertő-Hanság 2006: 73. (Traser 2002a), Kiskunság (Loksa 1987).

General distribution: Europe. Ecological note: hygrophilous. Distribution in Hungary: Fertő-Hanság (Traser 2002a, 2003; Traser et al. 2006), Őrség (Traser Family: Dicyrtomidae 1995). Dicyrtoma Bourlet, 1842

Heterosminthurus chaetocephalus Hüther, 1971 Dicyrtoma fusca (Lubbock, 1873) Heterosminthurus chaetocephalus Hüther, 1971: Dicyrtoma fusca (Lucas) (sic!): Szent-Ivány 1940a: Bretfeld 1999: 262; Heterosminthurus sp.: Traser 436; Dunger 1975: 26; Loksa 1983: 77; Loksa 1991: 1995: 47. 274; Farkas 1995: 46; Farkas 1996: 287; Farkas 1997: 58; Farkas & Szathmáry 1997: 39; Farkas et al. 1998: 57; Dicyrtoma fusca (Lubbock): Traser & Csóka 2000: General distribution: Palearctic. 76; Traser & Csóka 2001: 233; Traser 2002a: 267; Distribution in Hungary: Őrség (Traser 1995). Traser 2002c: 49; Traser et al. 2006: 73.

Heterosminthurus insignis (Reuter, 1876) General distribution: Holarctic. Distribution in Hungary: around the Lake Ba- Heterosminthurus insignis (Reuter): Dunger 1975: 25; laton (Farkas 1995), Balatonfelvidék (Farkas & Traser 1995: 47; Bourletiella (Heterosminthurus) insignis (Reuter): Loksa 1978a: 62; Bourletiella Szathmáry 1997), Börzsöny Mts. (Dunger 1975), insignis (Reuter): Loksa 1980: 111; Loksa 1991: 273; Fertő-Hanság (Traser 2002a; Traser et al. 2006), Farkas 1995: 46; Farkas et al. 1998: 57. Hortobágy (Loksa 1983), Kis-Balaton (Farkas

59 Dányi & Traser: Checklist of the springtail fauna of Hungary

1996, 1997), Kiskunság (Traser & Csóka 2000, Ptenothrix ciliata Stach, 1957 2001; Traser 2002c), Kőszeg Mts. (Szent-Ivány 1940a), Nyírség (Loksa 1991), Somogy Hills Dicyrtoma ciliata Stach: Loksa 1983: 77, figs 14-18. (Farkas et al. 1998). General distribution: Central Europe. Dicyrtomina Börner, 1903 Distribution in Hungary: Hortobágy (Loksa 1983). Dicyrtomina ornata (Nicolet, 1842) Ptenothrix leucostrigata Stach, 1957 Dicyrtoma ornata (Nicolet): Loksa & Rubio 1966a: 156; Loksa 1978b: 95; Farkas 1995: 46; Farkas 1996: Ptenothrix cf. leucostrigata Stach: Dunger 1975: 26; 287; Farkas 1997: 58; Farkas et al. 1998: 57. Dicyrtoma leucostrigata (Stach): Loksa 1980: 109, figs 3a-e. General distribution: West Palaearctic. Distribution in Hungary: Bakony Mts. (Loksa General distribution: Eastern Europe. & Rubio 1966a), Balatonfelvidék (Farkas 1995, Distribution in Hungary: Börzsöny Mts. (Dun- 1997), Cserhát (Loksa 1978b), Kis-Balaton (Far- ger 1975), Mátra Mts. (Loksa 1980). kas 1996, 1997), Somogy Hills (Farkas et al. 1998). Ptenothrix setosa (Krausbauer, 1898)

Dicyrtoma setosa (Krausbauer): Loksa 1978a: 63. Dicyrtomina minuta (Fabricius, 1783)

Dicyrtomina minuta (Fabricius): Traser 1997: 62; Tra- General distribution: Central Europe. ser 2002a: 267; Traser & Salló 2004: 230. Distribution in Hungary: Dráva Plain (Loksa

1978a). General distribution: Holarctic.

Distribution in Hungary: Fertő-Hanság (Traser 2002a), Kőszeg Mts. (Traser & Salló 2004), Family: Arrhopalitidae Szigetköz (Traser 1997). Arrhopalites Börner, 1906 Ecological note: planar to montane.

Arrhopalites aggtelekiensis Stach, 1929 Dicyrtomina saundersi (Lubbock, 1862) Arrhopalites pygmaeus aggtelekiensis Stach: Stach Dicyrtomina saundersi (Lubbock): Traser 2002a: 267. 1929: 308; Dudich 1930: 70; Dudich 1932: 49; Geb- hardt 1934: 140; Arrhopalites aggtelekiensis Stach: General distribution: Western Europe. Stach 1945: 19, pl. 4; Stach 1956: 66; Traser 1999: 58. Distribution in Hungary: Szigetköz (Traser 2002a). General distribution: Central Europe. Distribution in Hungary: Aggtelek Karst Ptenothrix Börner, 1906 (Stach 1929; Dudich 1930, 1932; Gebhardt 1934; Ptenothrix atra (Linnaeus, 1758) Stach 1945, 1956; Traser 1999). Ecological note: troglophilous. Ptenothrix atra (Linnaeus): Szent-Ivány 1941: 25; Traser 1999: 58; Traser 2003: 156; Dicyrtoma atra (Linnaeus): Loksa 1991: 274; Traser 2002a: 267. Arrhopalites bifidus Stach, 1945 Arrhopalites bifidus Stach: Loksa 1962: 63; Loksa General distribution: Europe. 1961b: 226, figs 17-22. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), Fertő-Hanság (Traser 2003, 2002a), General distribution: Europe. Mátra Mts. (Szent-Ivány 1941), Nyírség (Loksa Distribution in Hungary: Aggtelek Karst (Lok- 1991). sa 1961b), Bükk Mts. (Loksa 1962). Ecological note: silvicolous. Ecological note: troglophilous.

60 Dányi & Traser: Checklist of the springtail fauna of Hungary

Arrhopalites buekkensis Loksa, 1969 Distribution in Hungary: Bakony Mts. (Loksa Arrhopalites aggtelekiensis buekkensis Loksa: Loksa & Rubio 1966a). 1969a: 359, figs 17-27. Arrhopalites intermedius Loksa, 1969 General distribution: Central Europe. Arrhopalites hungaricus intermedius Loksa: Bajomi Distribution in Hungary: Bükk Mts. (Loksa 1969: 242; Loksa 1969a: 357, figs 1-16. 1969a). General distribution: Europe. Arrhopalites caecus (Tullberg, 1871) Distribution in Hungary: Aggtelek Karst (Ba- Arrhopalites caecus (Tullberg): Stach 1945: 9, pl. 1; jomi 1969; Loksa 1969a). Stach 1956: 73, pl. 9; Loksa 1956a: 395; Loksa 1962: 73; Traser 1997: 62; Traser 2002a: 267; Arrhopalites Arrhopalites loczyi Loksa, 1960 coecus (sic!) (Tullberg): Loksa 1956b: 203. Arrhopalites coecus (sic!) f. loczyi Loksa: Loksa 1961 General distribution: Palaearctic. a: 264, figs 31-34. Distribution in Hungary: Aggtelek Karst (Stach 1945, 1956), Buda Mts. (Loksa 1956b), General distribution: Europe. Bükk Mts. (Loksa 1956a, 1962), Fertő-Hanság Distribution in Hungary: Balatonfelvidék (Traser 2002a), Szigetköz (Traser 1997). (Loksa 1961a). Ecological note: edaphic. Arrhopalites ornatus Stach, 1945 Arrhopalites cf. cochlearifer Gisin, 1947 Arrhopalites ornatus Stach: Dunger 1975: 25. Arrhopalites cf. cochlearifer Gisin: Traser 2002a: 267. General distribution: Southern Europe. General distribution: Western Europe. Distribution in Hungary: Börzsöny Mts. (Dun- Distribution in Hungary: Sopron Mts. (Traser ger 1975). 2002a). Arrhopalites principalis Stach, 1945 Arrhopalites dudichi Loksa & Rubio, 1966 Arrhopalites principalis Stach: Loksa 1991: 276. Arrhopalites cochlearifer dudichi Loksa & Rubio: Loksa & Rubio 1966a: 154, figs 59-65. General distribution: Holarctic. General distribution: Central Europe. Distribution in Hungary: Nyírség (Loksa Distribution in Hungary: Bakony Mts. (Loksa 1991). & Rubio 1966a). Arrhopalites pygmaeus (Wankel, 1860) Arrhopalites hungaricus Loksa, 1967 Arrhopalites pygmaeus (Wankel): Stach 1929: 307; Arrhopalites hungaricus Loksa: Loksa 1967: 294, figs Dudich 1932: 49; Gebhardt 1934: 140; Stach 1945: 30, 18-26; Traser 1999: 58. pl. 8; Stach 1956: 78, pl. 10, 11; Loksa 1959b: 70; Paclt 1960: 156; Loksa 1960: 49; Loksa 1961b: 226, General distribution: Central Europe. figs 23-28; Loksa & Rubio 1966a: 154; Traser 1999: 58; Arrhopalytes (sic!) pygmaeus Absolon (sic!): Distribution in Hungary: Aggtelek Karst (Lok- Gebhardt 1933: 38; Arrhopalites binoculatus Börner: sa 1967; Traser 1999). Loksa 1956a: 391.

Arrhopalites infrasecundarius Loksa & Rubio, General distribution: Holarctic. 1966 Distribution in Hungary: Aggtelek Karst (Du- Arrhopalites infrasecundarius Loksa & Rubio: Loksa dich 1932; Loksa 1961b; Traser 1999), Bakony & Rubio 1966a: 154, figs 65-71. Mts. (Loksa & Rubio 1966a), Balatonfelvidék (Loksa 1960), Bükk Mts. (Loksa 1956a), Cserhát General distribution: Central Europe. (Loksa 1959b), Mecsek Mts. (Stach 1929; Geb-

61 Dányi & Traser: Checklist of the springtail fauna of Hungary

hardt 1934; Stach 1945, 1956; Paclt 1960; Geb- General distribution: Eastern Europe. hardt 1933). Distribution in Hungary: Fertő-Hanság (Traser 2002a). Arrhopalites secundarius Gisin, 1958 Arrhopalites secundarius Gisin: Loksa 1978a: 62. Family: Katiannidae Gisinianus Betsch, 1977 General distribution: Palaearctic. Distribution in Hungary: Dráva Plain (Loksa Gisinianus flammeolus (Gisin, 1957) 1978a). Gisinianus flammeolus (Gisin): Traser 1999: 58.

Arrhopalites sericus Gisin, 1947 General distribution: Central Europe. Arrhopalites sericus Gisin: Loksa 1978b: 95; Traser Distribution in Hungary: Aggtelek Karst (Tra- 1980: 20; Traser 1995: 46. ser 1999).

General distribution: Palaearctic. Sminthurinus Börner, 1901 Distribution in Hungary: Bakony Mts. (Traser 1980), Cserhát (Loksa 1978b), Őrség (Traser Sminthurinus aureus (Lubbock, 1862) 1995). Sminthurinus aureus var. ochropus (Reuter): Stach 1929: 308; Loksa 1978a: 62; Sminthurinus aureus Arrhopalites terricola Gisin, 1958 (Lubbock): Stach 1956: 118, pl. 16: figs 4-11; Loksa 1966: 386; Dunger 1975: 25; Loksa 1978b: 95; Loksa Arrhopalites terricola Gisin: Loksa 1966: 385; Loksa 1978a: 62; Loksa 1983: 76; Varga 1992: 172; Varga & 1978b: 95; Loksa 1978a: 62; Traser 1999: 58; Traser Vargha 1992: 315; Traser 1997: 62; Traser 1999: 58; 2002a: 267. Traser & Csóka 2000: 78; Dombos 2001: 2041; Traser & Csóka 2001: 233; Traser 2002a: 267; Traser 2002c: General distribution: Europe. 49; Traser 2006: 36; Traser & Horváth-Szováti 2006: Distribution in Hungary: Aggtelek Karst (Tra- 334; Traser et al. 2006: 73; Sminthurinus cf. aureus (Lubbock): Traser 1980: 20; Sminthurinus aureus var. ser 1999), Bakony Mts. (Loksa 1966), Balatonfel- atrata Börner: Traser 2002a: 267. vidék (Loksa 1966), Buda Mts. (Loksa 1966), Bükk Mts. (Loksa 1966, 1977), Cserhát (Loksa General distribution: Palaearctic. 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság Distribution in Hungary: Aggtelek Karst (Tra- (Traser 2002a), Keszthely Mts. (Loksa 1966), ser 1999), Bakony Mts. (Traser 1980), Börzsöny Mecsek Mts. (Loksa 1966), Pilis Mts. (Loksa Mts. (Dunger 1975), Buda Mts. (Stach 1956; Lok- 1966), Sopron Mts. (Traser 2002a), Vértes Mts. sa 1966), Budapest (Stach 1929), Bükk Mts. (Loksa 1966), Villány Mts (Loksa 1966). (Loksa 1966; Varga 1992; Varga & Vargha 1992), Cserhát (Loksa 1978b), Dráva Plain (Lok- Arrhopalites thermophilus Loksa, 1964 sa 1978a), Fertő-Hanság (Traser 2002a; Traser et Arrhopalites thermophilus Loksa: Loksa 1964: 96, figs al. 2006), Gerecse Mts. (Traser 2006), Hortobágy 58-65; Loksa 1964: 96, figs 58-65; Dunger 1975: 25; (Loksa 1983), Kiskunság (Traser & Csóka 2000, Arrhopalites thermophila (sic!) Loksa: Loksa 1966: 2001; Traser 2002c; Traser & Horváth-Szováti 386. 2006), Mezőföld (Dombos 2001), Pilis Mts. (Loksa 1966), Sopron Mts. (Traser 2002a), Szi- General distribution: Europe. getköz (Traser 1997). Distribution in Hungary: Börzsöny Mts. (Lok- Ecological note: eurytopic. sa 1964, 1966; Dunger 1975), Pilis Mts. (Loksa 1964). Sminthurinus bimaculatus Axelson, 1902

Arrhopalites ulehlovae Rusek, 1970 Sminthurinus bimaculatus (Axellson): Dunger 1975: 25; Loksa 1977: 211; Loksa 1978b: 95; Loksa 1983: Arrhopalites ulehlovae Rusek: Traser 2002a: 267. 76; Hornung 1986: 138; Loksa 1987: 80; Loksa 1991:

62 Dányi & Traser: Checklist of the springtail fauna of Hungary

273; Traser 2002a: 267; Traser 2003: 156; Traser & General distribution: Palaearctic. Salló 2004: 230. Distribution in Hungary: Bakony Mts. (Traser General distribution: Palaearctic. 1980), Cserhát (Loksa 1978b), Dráva Plain (Lok- Distribution in Hungary: Börzsöny Mts. (Dun- sa 1978a), Kiskunság (Loksa 1987), Nyírség ger 1975), Bükk Mts. (Loksa 1977), Cserhát (Loksa 1991). (Loksa 1978b), Fertő-Hanság (Traser 2002a, Ecological note: eurytopic species. 2003), Hortobágy (Loksa 1983), Kiskunság (Hor- nung 1986; Loksa 1987), Kőszeg Mts. (Traser & Stenognathellus Stach, 1956 Salló 2004), Nyírség (Loksa 1991).

Stenognathellus denisi Cassagnau, 1953 Sminthurinus elegans (Fitch, 1863) Sminthurinus aureus var. quadrilineatus (Tullberg): Sminthurinus denisi (Cassagnau): Dunger 1975: 25. Stach 1929: 308; Sminthurinus elegans (Fitch): Stach 1956: 124, pl. 16: fig. 1; Loksa & Rubio 1966a: 156; General distribution: Southern and Central Eu- Loksa 1966: 386; Loksa 1978a: 62; Loksa 1978b: 95; rope. Traser 1980: 20; Traser 1981: 133; Loksa 1983: 76; Loksa 1987: 80; Traser 1995: 46; Traser 1997: 62; Distribution in Hungary: Börzsöny Mts. (Dun- Traser 1999: 58; Traser 2002a: 267; Traser 2006: 36; ger 1975). Traser et al. 2006: 73. General distribution: Holarctic. Family: Sminthurididae Distribution in Hungary: Aggtelek Karst (Lok- sa 1966; Traser 1999), Bakony Mts. (Loksa 1966; Sminthurides Börner, 1900 Loksa & Rubio 1966a; Traser 1980), Balatonfel- Sminthurides aquaticus (Bourlet, 1842) vidék (Loksa 1966), Börzsöny Mts. (Loksa 1966), Sminthurides aquaticus (Bourlet): Stach 1929: 307; Buda Mts. (Stach 1929, 1956; Loksa 1966), Stach 1956: 22, pl. 1, 2: fig. 1, pl. 7: fig. 1; Paclt 1960: Cserhát (Stach 1929; Loksa 1978b), Dráva Plain 157; Dunger 1975: 25; Loksa 1980: 111; Loksa 1983: (Loksa 1978a), Fertő-Hanság (Traser 2002a; 75; Loksa 1987: 80; Loksa 1991: 273; Farkas 1995: 46; Traser et al. 2006), Gerecse Mts. (Loksa 1966; Farkas 1996: 287; Traser 1997: 62; Farkas 1997: 56; Farkas & Szathmáry 1997: 39; Farkas 1997: 58; Farkas Traser 2006), Hortobágy (Loksa 1983), Keszthely et al. 1998: 57; Traser 2002a: 267; Sminthurides Mts. (Loksa 1966), Kiskunság (Loksa 1987), aquaticus var. levanderi Reuter: Stach 1929: 307; Mecsek Mts. (Loksa 1966), Mezőföld (Stach ?Sminthurides aquaticus f. levanderi Reuter: Kovács 1929, 1956), Mt. Somló (Loksa 1966), Őrség 1953a: 184. (Traser 1995), Pilis Mts. (Loksa 1966), Sopron General distribution: Holarctic. Mts. (Traser 1981, 2002a), Szigetköz (Traser Distribution in Hungary: around the Lake Ba- 1997), Vas-Sopron Plain (Traser 1981), Vértes laton (Farkas 1995, 1997), Balatonfelvidék (Stach Mts. (Loksa 1966), Villány Mts (Loksa 1966), 1929, 1956; Farkas 1995, 1997; Farkas & Szath- Visegrád Mts. (Loksa 1966). máry 1997), Börzsöny Mts. (Dunger 1975), Fertő- Hanság (Traser 2002a), Hortobágy (Loksa 1983), Sminthurinus igniceps (Reuter, 1881) Kis-Balaton (Farkas 1996, 1997), Kiskunság Sminthurinus igniceps (Reuter): Loksa 1991: 273. (Stach 1929, 1956; Paclt 1960; Loksa 1987), Mát- ra Mts. (Loksa 1980), Nyírség (Kovács 1953a; General distribution: Europe, Japan. Loksa 1991), Somogy Hills (Farkas et al. 1998), Distribution in Hungary: Nyírség (Loksa Sopron Mts. (Traser 2002a), Szigetköz (Traser 1991). 1997). Ecological note: hygrophilous, epineustic. Sminthurinus niger (Lubbock, 1867) Sminthurides cf. parvulus (Krausbauer, 1898) Sminthurinus niger (Lubbock): Loksa 1978a: 62; Loksa 1978b: 95; Loksa 1987: 80; Loksa 1991: 273; Sminthurides cf. parvulus (Krausbauer): Traser 2002a: Sminthurinus cf. niger (Lubbock): Traser 1980: 20. 267.

63 Dányi & Traser: Checklist of the springtail fauna of Hungary

General distribution: Europe. General distribution: Holarctic. Distribution in Hungary: Fertő-Hanság (Traser Distribution in Hungary: Aggtelek Karst (Tra- 2002a). ser 1999), around the Lake Balaton (Farkas 1995), Balatonfelvidék (Farkas & Szathmáry 1997), Sminthurides malmgreni (Tullberg, 1877) Börzsöny Mts. (Dunger 1975), Dráva Plain (Lok- Sminthurides malmgreni (Tullberg): Loksa 1991: 273; sa 1978a), Fertő-Hanság (Traser 2002a, 2003; Farkas 1995: 46; Farkas et al. 1998: 57. Traser et al. 2006), Gerecse Mts. (Traser 2006), Hortobágy (Loksa 1983), Kis-Balaton (Farkas General distribution: Holarctic. 1996, 1997), Kiskunság (Traser & Csóka 2000, Distribution in Hungary: around the Lake Ba- 2001; Traser 2002c; Traser & Horváth-Szováti laton (Farkas 1995), Nyírség (Loksa 1991), 2006), Mátra Mts. (Loksa 1980), Mt. Somló (Lok- Somogy Hills (Farkas et al. 1998). sa 1966), Nyírség (Loksa 1991), Somogy Hills Ecological note: hygrophilous, epineustic. (Farkas et al. 1998), Szigetköz (Traser 2002a). Ecological note: mesophilic. Sminthurides pseudassimilis Stach, 1956 Sminthurides speudassimilis (sic!) Stach: Loksa 1980: Family: Sminthuridae 109. Subfamily: Sminthurinae General distribution: Europe, Japan. Distribution in Hungary: Mátra Mts. (Loksa Allacma Börner, 1906 1980). Allacma fusca (Linnaeus, 1758) Sminthurides schoetti Axelson, 1903 Smynthurus (sic!) fuscus De Geer (sic!): Tömösváry Sminthurides schoetti (Axelson): Loksa 1978a: 62. 1883: 34; Sminthurus fuscus Latreille (sic!): Tömös- váry 1884a: 57; Smynthurus (sic!) fuscus (Linnaeus): General distribution: Palaearctic. Vellay 1900: 20; Allacma fusca (Linnaeus): Stach Distribution in Hungary: Dráva Plain (Loksa 1929: 311; Szent-Ivány 1940a: 436; Stach 1956: 217, 1978a). pl. 27; Loksa 1956a: 411; Dunger 1975: 26; Loksa 1983: 77; Traser 1999: 58; Traser 2002a: 268; Smin- thurus (Sminthurus) fuscus (Linnaeus): Loksa & Rubio Sminthurides signatus (Krausbauer, 1898) 1966a: 156; Loksa 1978a: 63; Sminthurus fuscus (Lin- Sminthurides signatus (Krausbauer): Loksa 1991: 273. naeus): Loksa 1978b: 95; Loksa 1980: 111; Loksa 1991: 274.

General distribution: Palaearctic. General distribution: Holarctic. Distribution in Hungary: Nyírség (Loksa Distribution in Hungary: Aggtelek Karst (Tra- 1991). ser 1999), Bakony Mts. (Loksa & Rubio 1966a),

Bódva Valley (Tömösváry 1883; Vellay 1900; Sphaeridia Linnaniemi, 1912 Stach 1929), Börzsöny Mts. (Dunger 1975), Bu- Sphaeridia pumilis (Krausbauer, 1898) dapest (Tömösváry 1883, 1884a; Vellay 1900; Sminthurides pumilis Krausbauer: Loksa 1966: 385; Stach 1929), Bükk Mts. (Loksa 1956a), Cserhát Loksa 1978a: 62; Loksa 1980: 111; Loksa 1991: 273; (Stach 1929, 1956; Loksa 1978b), Dráva Plain Farkas 1995: 46; Farkas 1996: 287; Farkas & (Loksa 1978a), Fertő-Hanság (Traser 2002a), Szathmáry 1997: 39; Farkas 1997: 58; Farkas et al. Hortobágy (Loksa 1983), Kőszeg Mts. (Szent- 1998: 57; Sphaeridia pumilis (Krausbauer): Dunger 1975: 25; Loksa 1983: 76; Traser 1999: 58; Traser & Ivány 1940a), Mátra Mts. (Loksa 1980), Nyírség Csóka 2000: 76; Traser & Csóka 2001: 233; Traser (Loksa 1991), Vác-Pest Duna Valley (Tömösváry 2002a: 267; Traser 2002c: 49; Traser 2003: 156; Traser 1883; Vellay 1900; Stach 1929). 2006: 36; Traser & Horváth-Szováti 2006: 334; Traser et al. 2006: 73. Ecological note: silvicolous, corticophilous.

64 Dányi & Traser: Checklist of the springtail fauna of Hungary

Caprainea Dallai, 1970 1956: 262, pl. 32: fig. 1, pl. 33: figs 1-5; Loksa 1987: 80. Caprainea marginata (Schött, 1893) General distribution: Palaearctic. Sminthurus marginatus Schött: Stach 1929: 310; Stach Distribution in Hungary: Cserhát (Stach 1929; 1956: 252, pl. 31: figs 1-4; Dunger 1975: 26; Loksa 1983: 76; Loksa 1991: 274; Traser 1995: 46; Traser Szent-Ivány 1940a; Stach 1956), Kiskunság 1999: 58; Sminthurus (Sminthurus) marginatus Schött: (Stach 1929; Szent-Ivány 1940a; Stach 1956; Loksa & Rubio 1966a: 156; Sminthurus marginata Loksa 1987), Kőszeg Mts. (Szent-Ivány 1940a). (sic!) Schött: Loksa 1978b: 95; Caprainea marginata Ecological note: xerothermophilic. (Schött): Traser 2002a: 268.

General distribution: Europe. Sminthurus nigromaculatus Tullberg, 1871 Distribution in Hungary: Aggtelek Karst (Tra- Sminthurus (Sminthurus) nigromaculatus Tullberg: ser 1999), Bakony Mts. (Loksa & Rubio 1966a), Loksa & Rubio 1966a: 156; Loksa 1978a: 63; Sminth- urus nigromaculatus Tullberg: Loksa 1983: 77; Loksa Börzsöny Mts. (Dunger 1975), Cserhát (Stach 1991: 274; Traser 2002a: 268. 1929, 1956; Loksa 1978b), Fertő-Hanság (Traser 2002a), Hortobágy (Loksa 1983), Nyírség (Loksa General distribution: Holarctic. 1991), Őrség (Traser 1995). Distribution in Hungary: Bakony Mts. (Loksa Ecological note: xerothermophilic. & Rubio 1966a), Dráva Plain (Loksa 1978a), Fertő-Hanság (Traser 2002a), Hortobágy (Loksa Sminthurus Latreille, 1802 1983), Nyírség (Loksa 1991).

Sminthurus maculatus Tömösváry, 1883 Sminthurus viridis (Linnaeus, 1758) Smynthurus (sic!) maculatus Tömösváry: Tömösváry Smynthurus (sic!) viridis var. cinereoviridis Tullberg: 1883: 36, fig. 1; Tömösváry 1884a: 57; Vellay 1900: Tömösváry 1883: 38; Vellay 1900: 20; Smynthurus 20; Sminthurus viridis var. maculatus (Tömösváry): (sic!) viridis Linnaeus: Tömösváry 1884a: 57; Vellay Stach 1929: 311; Sminthurus maculatus Tömösváry: 1900: 20; Sminthurus viridis Linnaeus: Stach 1929: Stach 1956: 273, pl. 33: figs 6-8; Loksa 1966: 386; 310; Loksa 1980: 111; Loksa 1983: 76; Loksa 1987: Hornung 1986: 138; Varga 1992: 172; Varga & Oldal 80; Loksa 1991: 274; Traser 1995: 46; Farkas & 1998: 87; Dombos 2001: 2041; Traser 2002c: 49; Szathmáry 1997: 39; Traser & Csóka 2000: 78; Traser Traser & Horváth-Szováti 2006: 334; Sminthurus & Csóka 2001: 233; Dombos 2001: 2041; Traser (Sminthurus) maculatus Tömösváry: Loksa 1978a: 62. 2002a: 268; Traser 2003: 156; Sminthurus viridis var. cinereoviridis Tullberg: Stach 1929: 310; Traser 2002 General distribution: Southern and Central a: 268; Sminthurus viridis f. cinereoviridis Tullberg: Europe. Szent-Ivány 1940a: 436. Distribution in Hungary: Budapest (Tömös- General distribution: cosmopolitan. váry 1883, 1884a; Vellay 1900; Stach 1929), Distribution in Hungary: Balatonfelvidék Bükk Mts. (Loksa 1966; Varga 1992; Varga & (Stach 1929; Farkas & Szathmáry 1997), Buda- Oldal 1998), Dráva Plain (Loksa 1978a), Kiskun- pest (Tömösváry 1883; Vellay 1900; Stach 1929; ság (Stach 1929, 1956; Hornung 1986; Traser Szent-Ivány 1940a), Fertő-Hanság (Traser 2002a, 2002c; Traser & Horváth-Szováti 2006), Mező- 2003), Hortobágy (Loksa 1983), Kiskunság (Vel- föld (Tömösváry 1883; Vellay 1900; Stach 1929; lay 1900; Stach 1929; Loksa 1987; Traser & Dombos 2001). Csóka 2000, 2001), Kőszeg Mts. (Szent-Ivány Ecological note: thermophilic. 1940a), Mátra Mts. (Loksa 1980), Mezőföld

(Dombos 2001), Nyírség (Loksa 1991), Őrség Sminthurus multipunctatus Schäffer, 1896 (Traser 1995), Szentendre Island (Tömösváry Sminthurus multipunctatus ab. viridipunctatus Stach: 1883), Vác-Pest Duna Valley (Tömösváry 1884a; Stach 1929: 311; Szent-Ivány 1940a: 436; Sminthurus multipunctatus f. albopunctatus Stach: Szent-Ivány Vellay 1900; Stach 1929; Szent-Ivány 1940a). 1940a: 436; Sminthurus multipunctatus Schäffer: Stach Ecological note: on open habitats.

65 Dányi & Traser: Checklist of the springtail fauna of Hungary

Sminthurus wahlgreni Stach, 1920 46; Traser 1999: 58; Traser & Csóka 2000: 76; Traser & Csóka 2001: 233; Traser 2002a: 268; Traser 2002c: Sminthurus wahlgreni Stach: Loksa 1966: 386. 49.

General distribution: Eastern and Central Eu- General distribution: Europe, North Africa. rope. Distribution in Hungary: Aggtelek Karst (Tra- ser 1999), around the Lake Balaton (Farkas 1995), Distribution in Hungary: Mecsek Mts. (Loksa Bakony Mts. (Loksa & Rubio 1966a), Börzsöny 1966). Mts. (Dunger 1975), Buda Mts. (Stach 1929; Lok- sa 1956b), Bükk Mts. (Loksa 1956a, 1977; Varga Spatulosminthurus Betsch & Betsch-Pinot, 1984 1991; Varga & Oldal 1998), Cserhát (Loksa 1978b), Dráva Plain (Loksa 1978a), Fertő-Hanság Spatulosminthurus flaviceps (Tullberg, 1871) (Traser 2002a), Hortobágy (Loksa 1983), Kis- Sminthurus flaviceps ab. medius Stach: Stach 1929: kunság (Traser & Csóka 2000, 2001; Traser 311; Sminthurus flaviceps Tullberg: Stach 1956: 256, 2002c), Mátra Mts. (Loksa 1980), Nyírség (Ko- pl. 30; Loksa 1966: 386; Dunger 1975: 26; Loksa vács 1953a; Loksa 1991), Őrség (Traser 1995), 1977: 211; Loksa 1980: 111; Sminthurus (Sminthurus) Somogy Hills (Farkas et al. 1998), Sopron Mts. flaviceps var. ornata Uzel: Loksa & Rubio 1966a: 156. (Traser 2002a).

General distribution: Southern and Central DISCUSSION Europe. Distribution in Hungary: Bakony Mts. (Loksa Our review revealed 414 Collembola species 1966; Loksa & Rubio 1966a), Balatonfelvidék recorded so far from Hungary. It’s a rather high (Loksa 1966), Börzsöny Mts. (Dunger 1975), Bu- number referring to this relatively small area, da Mts. (Loksa 1966), Bükk Mts. (Loksa 1977), which can be explained by the geographical si- Cserhát (Stach 1929, 1956), Keszthely Mts. (Lok- tuation of the country. Although Hungary is lo- sa 1966), Mátra Mts. (Loksa 1980), Mecsek Mts. (Loksa 1966), Mt. Somló (Loksa 1966), Villány cated in the middle of the Carpathian Basin, it is a Mts (Loksa 1966). transitional zone and impacts of the Alpin, Carpathian and Illyric regions are significant as Lipothrix Börner, 1906 well. Regarding the number of the species, Hun- gary belongs to the relatively well investigated Lipothrix italica (Cassagnau, 1968) countries in Europe: Austria: more than 450 Sphyrotheca italica Cassagnau: Dunger 1975: 26; (Christian 1987); Poland: 441 (Skarzynski et al. Lipothrix italica (Cassagnau): Traser 2002a: 268; 2002), Romania: 388 (Fiera 2007), the Ukraine: Traser 2003: 156. 527 (Kaprus et al. 2004). However, it has to be mentioned that even in these countries new spe- General distribution: Europe. cies are discovered every year and that numerous Distribution in Hungary: Börzsöny Mts. (Dun- species occur in neighbouring areas which can be ger 1975), Fertő-Hanság (Traser 2003), Sopron expected from Hungary as well. The fauna of the Mts. (Traser 2002a). country isn’t entirely discovered, there are several territories of totally uninvestigated springtails fa- Lipothrix lubbocki (Tullberg, 1872) una in the country yet. Similarily to the case of Sphyrotheca lubbocki (Tullberg): Stach 1929: 310; oribatid mites (Mahunka & Papp 2005), another Dunger 1975: 26; Sphyrotheca lubbocki var. maculata very diverse group of soil , we can highly Axelson: Stach 1929: 310; Sminthurus lubbocki Tull- berg: Kovács 1953a: 184; Loksa 1956b: 203; Loksa expect to find even many further species new to 1956a: 395; Loksa 1977: 211; Loksa 1978b: 95; Loksa the fauna of Hungary in the future. 1980: 111; Loksa 1983: 76; Loksa 1991: 273; Varga 1991: 85; Farkas 1995: 46; Farkas et al. 1998: 57; Acknowledgements. We thank the Hungarian Scientific Varga & Oldal 1998: 87; Sminthurus (Sphyrotheca) Research Grant (OTKA No. 72744) for a grant to the first Lubbocki Tullberg: Loksa & Rubio 1966a: 156; Loksa author. The research was supported by the National R&D 1978a: 62; Lipothrix lubbocki (Tullberg): Traser 1995: Programme (contract no.: 3B023–04) as well.

66 Dányi & Traser: Checklist of the springtail fauna of Hungary

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TRASER, GY. (1980): Adatok a farkasgyepűi bükkösök Folia Entomologica Hungarica (Rovartani Köz- avarszintjének Collembola (ugróvillás) faunájához. lemények), 52: 119–122. Erdészeti és Faipari Tudományos Közlemények, 2: TRASER, GY. & CSÓKA, GY. (2000): Ásotthalmi fenyő– 19–23. és tölgy erdők összehasonlító talajfaunisztikai vizs- TRASER, GY. (1981): Talajinszekticidek hatása a talaj- gálata. In: Horváth, B. (ed.): Tudományos Ered- faunára. Erdészeti és Faipari Tudományos Közle- mények a Gyakorlatban. Alföldi Erdőkért Egyesület, mények, 3(2): 129–138. Kutatói Nap 1998-1999. Baja-Kecskemét-Sopron, pp. TRASER, GY. (1983): Az ugróvillások (Collembola) el- 75–80. oszlásának vizsgálata 50 ccm-es talajmintákban. TRASER, GY. & CSÓKA, GY. (2001): A mezofauna Erdészeti és Faipari Tudományos Közlemények Insecta: Collembola ásotthalmi fenyő- és tölgy 4(2): 169–174. erdők talajában. Erdészeti kutatások, 90: 231–240. TRASER, GY. (1989): Inszekticidek tesztelése labora- TRASER, GY. & SALLÓ, M. (2004): Új ugróvillás (In- tóriumban ugróvillásokkal (Insecta: Collembola). secta: Collembola) adatok a Kőszegi-hegységből. Erdészeti és Faipari Tudományos Közlemények 10: Természetvédelmi Közlemények, 11: 229–232. 89–94. TRASER, GY., SZŰCS, P. & WINKLER, D. (2006): TRASER GY. (1994): A meszezés hatása egy savanyú Collembola diversity of moss habitats in the Sopron bükkös ugróvillás (Insecta: Collembola) faunájára. Region, NW-Hungary. Acta Silvatica et Lignaria Erdészeti Szakmai Konferencia kiadványa, Sopron, Hungarica, 2: 69–80. pp. 99–103. TRASER, GY. & THIBAUD, J–M. (1999): Une nouvelle TRASER, GY. (1995): The Collembola fauna of Őrség espèce du genere Neonaphorura de Hongrie (Col- (Insecta: Collembola). In: Víg, K. (ed.): Natural lembola, Onychiuridae, Tullbergiinae). Revue History of Őrség Landscape conservation Area I. Savaria Múzeum, Szombathely, pp. 43–47. Française d’Entomologie (Nouvelle série), 21(3): TRASER GY. (1997): Egy szigetközi nemesnyáras bio- 105–108. diverzitása: a talajfauna. In: Pájer, J. & Fabich, E. TRASER, GY., THIBAUD, J-M. & NAJT, J. (1993): Deux (eds): V. Erdészeti Szakmai Konferencia kiadványa nouvelles espèces de collemboles (Insecta) de a WoodTech keretében. Sopron, pp. 57–64. Hongrie. Bonner Zoologische Beiträge, 44(3-4): TRASER, GY. (1999): Springtails of the Aggtelek Na- 221–224. tional Park (Hexapoda: Collembola). Mahunka, S. TRASER, GY. & WEINER, W.M. (1999): A new species (ed.): The fauna of the Aggtelek National Park. of Stenaphorurella Luciáňez et Simón (Collem- Akadémiai Kiadó, Budapest, pp. 49–59. bola: Onychiuridae, Tullbergiinae) from Hungary. TRASER, GY. (2000): A new Lepidocyrtus species Folia Entomologica Hungarica, 60: 109–113. (Collembola: Entomobryidae) from Hungary. Folia TRASER, GY. & CSÓKA, GY. (2001): A mezofauna Entomologica Hungarica, 61: 65–69. Insecta: Collembola ásotthalmi fenyő- és tölgy er- TRASER, GY. (2002a): The Collembola of the Fertő– dők talajában. Erdészeti kutatások, 90: 231–240. Hanság National Park. In. Mahunka, S. (ed.): The TRASER, GY. & HORVÁTH-SZOVÁTI, E. (2004): Táj- fauna of the Fertő-Hanság National Park. MTM, hasznosítás hatása a talajfaunára: ugróvillás (Pa- Budapest, pp. 259–270. rainsecta: Collembola) közösségek diverzitása TRASER GY. (2002b): Xerothermic Collembola from Csévharaszton. Alföldi Erdőkért Egyesület, Kecs- the Bükk–Mts NE–Hungary. Folia Historico-Natu- kemét, pp. 51– 62. ralia Musei Matraensis, 26: 159–161. TRASER, GY. & HORVÁTH-SZOVÁTI, E. (2006): Land– TRASER, GY. (2002c): Borókás–nyáras erdők ugróvillás use intensity and diversity parameters: Collembola (Insecta: Collembola) faunája. In: Barna, T. (ed.): (Insecta) communities in Csévharaszt (Hungary). Alföldi Erdőkért Egyesület Kutatói nap 2001-2002., In: Csóka, Gy.; Hirka, A. & Koltay, A. (eds.): Gyula, pp.46–52. Biotic damage in forests. Proceedings of the TRASER GY. (2003): Hansági nemesnyár és éger erdők IUFRO (WP7.03.10) Symposium held in Mátra- ugróvillás (Insecta: Collembola) faunája. Magyar füred, Hungary, September 12-16, 2004, pp. 329–339. Biológiai Társaság, Budapest. III. Kárpát-meden- VARGA, J. (1991): Adatok Felsőtárkány területének cei Biológiai Szimpózium kiadványa, pp. 153–157. bryofaunájára vonatkozóan. Acta Academiae Peda- TRASER, GY. (2006): A gyarmatpusztai vadgesztenyés gogicae Agriensis, Nova Series, 20: 81–88. ugróvillás (Insecta: Collembola) faunájának diver- VARGA, J. (1992): Analysis of the fauna of protected zitása. Folia Musei Historico-Naturalis Bakonyi- moss species. Biological Conservation, 59: 171–173. ensis, Zirc, 23: 33–39. VARGA, J. & NAÁR, Z. (2002): A collembolák mint TRASER, GY. & CHRISTIAN, E. (1992): Lepidocyrtus lehetséges terjesztői a mohákban élő gombáknak. peisonis sp. n., ein neues Springschwanz aus dem Folia historico-naturalia Musei Matraensis, 26: Neusiedlersee-Gebiet (Collembola: Entomobryidae). 115–120.

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VARGA, J., NAÁR, Z. & DOBOLYI, CS. (2000): A moha Acta Academiae Pedagogicae Agriensis, Nova Seri- mint táplálék két Collembola faj számára. Acta es, 21: 307–326. biologica Debrecina. Supplementum Oecologica VELLAY, I. (1900): Ordo Apterygogenea. In: Pasz- Hungarica, 11(1): 323. lavszky, J. (ed.): A Magyar Birodalom Állatvilága VARGA, J., NAÁR, Z. & DOBOLY, C. (2002a): The com- (Fauna Regni Hungariae) III. Arthropoda. Királyi position of intestine content of Orchesella cincta Magyar Természettudományi Társulat, Budapest, (Linné) (Insecta: Collembola). Opuscula Zoologica, pp. 19–22. Budapest, 34: 105–112. WALTZ, R.D. & HART, J.W. (1996): A checklist of Indiana Collembola (Hexapoda). Proceedings of VARGA, J. & NAÁR, Z. & DOBOLYI, CS. (2002b): the Indiana Academy of Sciences, 105: 29–41. Selective feeding of collembolan species Tomoce- ZHAO, L., TAMURA, H. & KE, X. (1997): Tentative rus longicornis (Mull.) and Orchesella cincta (L.) checklist of Collembolan species from China (In- on moss inhabiting fungi. Pedobiologia, 46, 526–538. secta). Publications of the Itako Hydrobiological VARGA, J. & OLDAL, V. (1998): A légkörszennyezés Station, 9: 15–40. hatása a mohák állatközösségére. Acta Academiae ZIMDARS, B. & DUNGER, W. (1994): Synopses on Pa- Paedagogicae Agriensis, Nova Series, 22: 83–92. laearctic Collembola I.: Tullbergiinae Bagnall, VARGA, J. & VARGHA, B. (1992): Adatok az Upponyi– 1935. Abhandlungen und Berichte des Natur- szoros bryofaunájának összetételére vonatkozóan. kundemuseums Görlitz, No. 68(4), 74 pp.

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Plate I. Distribution of Arrhopalites pygmaeus and Microgastrura duodecimoculata in Hungary.

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Plate II. Distribution of Isotomodes trisetosus and Lepidocyrtus ruber in Hungary.

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Plate III. Distribution of Deutonura stachi, Arrhopalites aggtelekiensis, A. bifidus and and A. buekkensis in Hungary.

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Plate IV. Distribution of Tetrodontophora bielanensis and Pseudosinella aggtelekiensis in Hungary.

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Plate V. Distribution of Protaphorura metapulvinata and Folsomia gebhardti in Hungary.

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Plate VI. Distribution of Protaphorura sublata and Heterosminthusus bilineatus in Hungary.

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Plate VII. Distribution of Lepidocyrtus peisonis and Bilobella braunerae in Hungary.

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Plate VIII. Distribution of Protaphorura subnemorata and Deuteraphorura silesiaca in Hungary.

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Plate IX. Distribution of Neotullbergia ramicuspis and Bourletiella viridescens in Hungary.

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Plate X. Map of Hungary with the geographical units used in the text. 1 = Visegrád Mts., 2 = Pilis Mts., 3 = Buda Mts., 4 = Tétény Plateau, 5 = Szentendre Island, 6 = Vác-Pest Danube Valley

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