Research articles NAT. NAT. HIST. BUL L. SIAM SOC. 44: 11-22 ， 1996

BOTANICAL NOTES ON THE FLORA OF NORTHERN : 51

J.F. J.F. Maxwe 1l 2

ABSTRACT

Twelve taxa of vascular l1 0wering are preported as new records the for l1 0ra of of Thailand. This in cI udes the genera Acti l1 ostemma' Griff. (Cucurbitaceae) ， Epilobium L (Onagraceae) ， and Phytolacca L. (Phytolaccaceae). Fruits of four other species are described along along with the seeds of three of them.

SCHISANDRACEAE

Schisandr α， verrucosa Gagnep. NEW RECORD ，FRUITS AND SEEDS DESCRIBED ， figure figure 1

1 reported this genus ， viz. S. perulata Gagnep. ， as a new record for the flora of Tha i1 and recent1y (MAXWELL ， 1992). Now another species can be added ，i.e. S. verrucosa Gagnep. which GAGNEPAIN (1938) described along with S. perulata. Schisandra verrucosa (Latin (Latin = warty ，i.e. stems) was described from flowering material collected in Tonkin ，N. . 1 collected mature fruiting material of this species in Jae Sawn National Park ， Lampang Province ，below Bah Miang V il1 age at 925 m on 30 November 1995 (Maxwell 95 ・1194). It was growing in a partly open ， disturbed area in primary evergreen ，seasonal ， hardwood forest on granite bedrock. Fruiting Fruiting pedicels glabrous ， pale light green ，7-22 mm 10ng. Bract remnants often present ，ovate ，broadly rounded at the tip ， c. 2x 1 .5 mm. Fruiting receptacles accrescent ， glabrous ，spike-1ike ， pale light green ，2 4- 70 mm long. Fruiting carpels (monocarps) sessile ， spaced ，globose (with 1 seed) or elllipsoid (2-s 田 ded) ，slightly juicy ， pa1e 1ight green and later later 1ight orangish. c. 7-10 x 5-9 mm. Seeds slight1y laterally compressed ，subglobose and somewhat reniform ， sides rounded; testa smooth; 6-7 x 5-6 mm. Hilum lateral ，on the the emarginate side. Key to the two known species of Schisandra in Thailand: 1. 1. Stems smooth ， often angu1ar; leaf blades elliptic to broadly ovate ，up to 10 x6

cm. …………....・ H ・-…・・ H ・H ・...... ・ H ・.....・ H ・....…・…-…....・ H ・....…… S. perulata Gagnep. 1. 1. Stems verrucose ，always terete; leaf blades ovate-lanceo1ate ，up to 19 x 6.5 cm.

….....・ H ・. S. verrucosa Gagnep.

1 Continued from Nat. Bull. Hist. Siam Soc. 42 (1 994) 259-262. 2 Herbarium ， Department of Biology ， University ， Chiang Mai 50200 ，Thailand. Received Received 25 December 1995; accepted 5 March 1996.

11 11 12 12 J.F. MAXWELL

RUTACEAE

Glycosmis Glycosmis pseudoracemosa (Guill.) Swingle NEW RECORD ，figure 2

STONE (1985) notes that this species ，described in 1912 from material collected in Tonkin ， N. Vietnam ， is also known from Kwangsi Province ， southern. . It is distinguished distinguished by having unifoliate leaves with clearly articulated petioles ，compact inflorescences inflorescences which 紅 e as long as or slightly longer than the petioles (i. e. c. 5 mm long) ， and and 5-10cular ovaries.

1 found flowering material (Maxwell 95 ・ 1291) of this species in the northern part of Jae Jae Sawn National Park ，Wahng Nua District ，Lampang Province ， at Pah Ngahm Station area area at 650 m on 6 December 1995 ，佃d again near that locale at 600 m on 28 April 1996 (Maxwell (Maxwell 96-643). It was found in open ， rugged limestone terrain in degraded deciduous forest forest with much bamboo and grows as a tr 巴elet or shrub 1.5-2 m tall in dense undergrowth.

Za nth o. 砂 lum evodia φ lium Guill. NEW RECORD ，白 gure 3

This This species is very appropriately named since the material ， in fruit ， that 1 found in Doi Chiang Dao Wildlife Sanctuary ， has trifoliate ，less frequently unifoliate leaves ， that closely closely resemble those of Euodia (Rutaceae) that are alternate and not opposite as in the latter. latter. Described Described in 1945 from three collections from Annam in Vietnam ，this species differs from from all others in the region by having unifoliate or trifoliate leaves. The material that 1 found found (Maxwell 95 ・1128) was growing near the east end of Doi Luang Valley in primary ， evergreen ，seasonal ，hardwood forest in shaded ， rugged limestone terrain at 1750 m and was collected on 9 November 1995. It It grows as an understorey tree ι8 m tall with a dbh of ι9 cm. The bark is thin ， very very coarsely roughened ，tan ， and with spine-tipped tubercles.

LEGUMINOSAE ，Papilionoideae

Uraria Uraria poilanei Dy Phon NEW RECORD

Dy Phon (VAN THUAN et al.， 1987) notes that this species is expected to be found in Thailand. Thailand. It was described in 1987 based on material collected by E. Poilan in Xieng Kh ouang Province ， at 900 m in “mixed deciduous" forest on limestone bedrock. 1 have made thr 田 collections of this species ，all on granite bedrock ，viz. two at Doi

Kuhn Dahn National Park ，Lampoon Province (Maxwell 93 ・1436 ，flowers ，22 November

1993 1993 and 94 ・ 151 ，fruits ， 31 January 1994) and in the southern part of Doi Chiang Dao Wildlife Wildlife Sanc 加紅y， Chiang Mai Pr ovince (Maxwell 95-1039 ，flowers ，3 November 1995). 官le two specimens from Doi Kuhn Dahn were made in the same place in degraded primary primary evergreen ，seasonal hardwood forest at 1175 m ， while the Chiang Dao one was collected collected at 925 m in disturbed ，mixed evergreen + deciduous ，seasonal hardwood forest with with much bamboo. It grows as a deciduous herb-shrub 1- 1. 5m tall (Chiang Dao) or as BOTANICAL NOTES 13 an an evergreen shrub 1.5-2.5 m tall (Doi Kuhn Dahn). This This species is distunguished by having glandular-sticky indumentum on most parts ， trifoliate trifoliate leaves with relatively large leaflets ， and “semi-lunar" pod segments. lt is very closely closely related to U. Lacei Craib (type Burma) which is known from northem Thailand ， Burma ，southem China ，Laos ， and Vietnam. Uraria Lacei lacks glandular indumentum ， has smaller smaller leaflets ， and more rectangular pod segments.

CUCURBIT ACEAE

A Ct inostemma tenerum Griff. NEW GENERIC AND SPECIES RECORD ， figure 4

Griffith Griffith established this genus and described this species in 1837 based on material from East Benga l. The genus is distinguished by having 5，equal ，un i1 ocular ，free stamens; small ，circumscissile (pyxidium) fruits; and exalate ，compressed ，denticulate seeds. 1 have collected this species twice ， both staminate ，on the eastem lowlands of Doi Chiang Chiang Dao W i1 dlife Sanctuary ， (Maxwell 89 ・1540 ，800 m ， 16 December 1989) in mixed evergreen + deciduous ，seasonal ，hardwood forest at the base of of the limestone esca 中ment and (Maxwell 95 ・845 ，550 m ，7 October 1995) in an open thicket thicket with much secondary growth in similar forest ， but on shale bedrock. This This species is a vine with very polymorphic leaf blades ranging from ovate to suborbicular suborbicular with acute to deeply cordate bases ， and ranging in size from 7.5-20 cm long x 3.5- 1. 7 cm wide ， with petioles 2-8.5 cm long. According to KERAUDREN-AYMONIN (1975) (1975) A. tenerum ranges from NE lndia ，China ，Japan ，Korea ，Taiwan ， and N. Vietnam.

Momordica Laotica Gagnep. NEW RECORD

According According to the most recent regional revision of the Cucurbitaceae (KERAUDREN- AYMONIN ， 1975) this species is endemic to Laos. Gagnepain described this species in 1918 1918 based on staminate material collected in Louang Prabang which ，apparently ， is the only only material known.

1 found staminate material of this species (Maxwell 95 ・736) at Jae Sawn National P 訂 k，Muang Bahn District ，Lampang Pr ovince on 23 September 1995 in an open ，disturbed area area in degraded ，fire-prone ，seasonal ， deciduous forest with much bamboo on granite bedrock. bedrock. lt lt differs slightly from M. subanguLata B l.， which is known from lndia ，lndo-China ， Peninsular Peninsular Malaysia ， and Java; in having slight mo 中hological differences in the bracts and sepals. sepals.

MELASTOMATACEAE

Osbeckia steLLata Ham. ex Ker-Gaw l. v低 rostrata (D. Don) C. Han. NEW RECORD ， figure figure 5 14 14 J. F. MAXWELL

This This variety has a long taxonomic history and numerous synonyms (HANSEN ， 1977). It It was originally described as Osbeckia rostrata D. Don in 1825 on material from Nepa l. It It is known from the Himalayas ，India ，Burma ，Laos ， and S. Vietnam. Osbeckia Osbeckia stellata has five varieties ，three of which are found in northern Thailand ，viz. var. var. crinita (B 血. ex Naud.) C. Han. and var. marginulata (Wall. ex C l.) C. Han. ， both of which which are common ， and now var. rostrata ， which 1 know from only one place. Var. Var. rostrata is very common in Doi Chiang Dao Wildlife Sanctuary ， Chiang Mai Province Province in the vicinity between Huay Mae Gawk Station and the west side of Doi Luang -the -the limestone massif. 1 collected flowering material of var. rostrata at 1475 m on 1 September September 1995 in seasonal ，mixed evergreen hardwood+pine forest on granite bedrock (Maxwe 1l 95 ・658). Simon Gar 命ler also collected this variety (G 訂 dner H 137) in similar forest forest on shale bedrock at 1150 m on 6 December 1995 slightly west of where 1 got it. It is is a deciduous ，perennial herb 1- 1. 5m tall which differs from var. marginulata ， which is found found at lower elevations (c. 65 0- 1250 m) in the area ， in being nearly glabrous throughou t. Var. Var. crinita ， found in evergreen hardwood + pine forest from 1225-1900 m 加 dsympa 凶c with with var. rostrata ，differs in having hispid indumentum on most parts and flowers in October-November ， i. e. when v低 rostrata is fruiting.

ONAGRACEAE

Epilobium Epilobium brev 扮lium D. Don ssp. trichoneurum (Hauss.) Raven NEW GENE 則 CAND SSP. SSP. RECORD ，figure 6

According According to RA VEN (1 977) this taxon is found throughout SE Asia and Ph i1i ppines but but not ，apparently ，百 lailand. 1 found mostly fruiting material of ssp. trichoneurum (Greek: small small hairs on the nerves) in the upper valley just below the summit of Doi Luang in Doi Chiang Chiang Dao Wildlife Santuary ， Chiang M 泊 Province at 1950 m on 7 November 1995 (Maxwell (Maxwell 95 ・1097). It is found in former primary evergreen ，seasonal ，hardwood forest which which has been cleared for opium cultivation and is now overgrown with Eupatorium adenophorum Spreng. (Compositae) in rugged limestone terrain. Ssp. trichoneurum must be be relatively new to Thailand since CRAIB (1 931) and SMITINAND (1966) do not include it it or the genus Epilobium in their enumerations. Although most of the plants found were in in fruit ，a few had their last flowers which enabled me to identify them. It is an annual herb herb and is presently very common in the 紅 'ea in which 1 found it. GAGNEPAIN (1 921) ， as as E. trichoneuron Hauskn. ， gives a complete description as well as a plate.

ASCLEPIADACEAE

Heterostemma gracile Kerr FRUITS AND SEEDS DESCRIBED

Kerr Kerr (1 939) described this species from his own collection (Kerr 6535) made on Doi Chiang Chiang Dao (Wildlife Sanctuary ， Chiang Mai Province) at 1400 m in November 1922. collected 1 collected fruiting material of this species at 1750 m on 9 November 1995 (Maxwell sOTAN ICAL NOTES 15

Figur e 1. Schis{/ l1 dra ver rll cosa Ga g n ep.

(Sc hi 日Indr accae)

T hi s is lh c おcco nd spec i es 0 1' lhi s gen us kn ow n in Tha il and . Gι1 9ne pain (1938) i ni c lucl cs rOl I¥ おpec ics ror Indo -C h ina. lh c liflh. liflh. S. crassifal ia Pi err e cx F in . &

Gagncp. = Kad Sl/ ra helerocli l(/ (R oxb.) C raib (Sc hi日IIl cl raccae) (K en g. 19 72) PhO lO: S iri porn Kopac hon

no tA 0'糊刷。 CMUH ・_.. 、均 uII V 01 $c l・問 f ，C iII 凹 gM al U川W 繍 V C河川。 '''g t.101.lhoUor刈

U.hy …叫ー """'"'- .. 柄、， 4園出【叶 '一一10<;0l"'" -.. ~ " ‘叫 ・炉 叶曲畑 町民

，-.....， ，-.....， -'制" ・6 ，，_ ，'''''.，

価低岡島川、，..山" 防相 .."

Fi呂ur c 2. C I¥叱 ()S I1 U S jJ selfdoracelll 口町 α (G uill .) Sw in glc (Rula ce a e) In cluclin 呂 lhi ss pec icぉ ancl acco rdin g lO Slonc ( 1985) lh ere arc now 13 spcc i cs 0 1 lh is gcn us kn ow n rrO I1l Thailancl . Ph OlO S irip o rn K op achon

，剣山 "岨 1・ 同 w 低刈"'品 、， "μ品五込~五色~(也lll ， ).・ h 旬・

陶剛山~. 出-お・崎町叫‘・瓜叫・・ ""...... 1. ‘"・'. h.'''''''' d~' 瑚 ~.・ "・凶~"" "" “・・ 3 一一.叩 ー雷~. b ，-..山 .， 9')) 山叩 . ・ P 闘~ . ..問".州1.-，..山.. .. ~州… 山 r …圃品川叩・ ・句."・ u 凶N' 叩吋 I... .~ r山・，u...... __ "叫リ」 ・.... .11 :....."".1 ・ .. .t、・・8 ・ぬ M ・ E ・... .. ，~ . 陶 1"'" ・11.'11 “a‘" .・ 2 ・・'-' "'~.-・・・ 匂.~ ，..w ・m ・・M 川伽 一向 1 ・a 問、ぃ 白..・. ~ro ... d噸 vι. ‘・ 一一 ぷ込 四 1綱高 '--.1 ". ~..・・""，榊 …一蜘 ， 16 J. F. MA XWE LL

Figllr e 3. Zanlhoxylu l1 l eνod ia efo liu lII G lI ill (R lI lacea 巴)

Accorcling Accorcling loC raib (1 926) lh ere are fi v巴

olh 巴r specie s 0 1' lhi s gen ll s in T hail ancl of

w hi ch onl y Z. acanlhopodiu 川 O c. is also known from Oo i C hi ang Oao. PhOlO S irip o rn Kopachon

山叫 一州岡一 ゆ

川町 恥川 叩

'~'，' C Of"I"'<'nNQr1阿 抑制u 向 II~W ~ー叫!

o.o，c l 1'''' "，"' ;'_ れ'o...l ~".4' 向日 A ‘'*~ .， r 叫い岨~ V，J l・，

J-1ifl 陶叫".、." 加

…tJ~( .f 刈"一一、 切』川よ遇

F igllre 4. A Cl inos lemma lel1 erU fII G ri fl (C lI cllr bil aceae) r 円 'f';" T hi ss pec ies ， as fa r as 1 am awa re，is onl y known from slamin al巴 m at巴ri al from Oo i C hi ang Oao in Thailancl. Il al so mi g hl

be present in lI nicl enlifi ecl or mi sicl enlili 巴cl herb arillm m al巴rial. Fema lea nd frlliling malerial ，th erefore ，mll Sl be pr ese nl rh ere， alb eil diffi cllll lo fincl. Ph O lO: Siri po rn Kopachon

C¥ BOTAN ICA L NOTES 17

Figul 巴 5. Osbe c kia .lI e ll a{{/ H al1l . ex K er-Gaw l. va l ros !ra!a (0. Oon) C. H an1

(M elζa 叫削l凶叫 S叫lOlll f.l 刈l凶a∞cea 出e) T he thr ee know n va ri eti es 01' O. s! e!!a !a are n ot on ly read il y cli stin gui sh巴d 11l 0rph olog icall y. but also h ave diffe rent phenolog i es as we ll asS O I1l 巴 habitat dilT，巴!巴 nce s. Ph oto: Sir ipo rn Kopac hon

HO~4 0' 仰山崎 CMI.II1・，_旬町、.~凶 y OlScI<射程・，_，刷.."、省、" .. ，.. 加 IIMOII" 舗-

¥、

f Figur c 6. Epi!obiu l/ l b revifoli u lII O . Oon ss p

!ri cho l/ eU IiIlIl (H auss .) R aven (O nagraceae) T hi ss pecies has establi shed it s巴11" onl y rec 巴ntl y in T hai land . 11l 0st probably fro l1l south e rn C hina . Photo: Sirip o rn Kopacho n

私ド

1; ， . u 18 J.F. M AXWE L L

Fig llr e 7. Phyto!a cc α o lll e ric OllO L. (P hylolacca- cea 巴)

Thi ss pec i巴5，al 50 rec 巴nll y eS labli s hecl in

Thail 山 lcl ，is qllil e rar e on Ooi C hi ang Oao.

blll blll mighl al討o be I" ollncl in olh er lI plancl areas areas in nonh c rn Thai la ncl. Ph olo S irip o rn K opac hon

制附町 α 仙附 川 ……い 山岨 h hm

，t0 ，t0

〆 ‘旬、 1. ， ~・ ‘

Figur e 8. F;c Ll S lI er ν()S (I H ey. ex R O lh va r. lI ervo.¥ '{/ ρ¥ (Mo ra ce a e)

~ '¥ '，Z S in ce lh 巴 prop er icl 巴nlili ca lion 0 1" man y (' spec l巴S 0 1" Fi clI S is qllil e clil Ti Cllll ，lh cre 、、ン/ l11 ig hl be unicl enli f iecl or in co rrec ll y ノ in cle nlili 巴cl spcc il11 巴ns 0 1" thi s laxo n in olh erh erbari a. Ph O lO: Sirip o rn 1くopac hon

川 出 1同肌 rO ~"" 1 o !…削ong M o>l1.l nlv"'''h C 仇畑、 0 1.1凶'" 副畑、d BOT ANICAL NOTES 19

95-1124) 95-1124) in ap 紅t1 y open area in seasonal ， primary evergreen hardwood forest in rugged limestone limestone terrain from the same 紅 'ea that Kerr did. Kerr' s materi a1 was flowering ， thus the the fruits can be described here. Folli c1 es spreading at 180 ・ormore to 血e pedicel ，terete ，glabrous ， dark green ，gradually narrowed narrowed to 白e ac 胸 tip ，67 ・103 x 3.5 -4 mm. Seeds linear ，flat ，brown ， 15 x 1. 5 mm. Coma si1 ky ，white ，about 槌 long as the seed. 百lis species is readily distinguished from other species in this genus by its membranous leaf leaf blades which are acute at the tip and acute to rounded at the distinc t1 y 3-plinerved base. base. The largest blades on my material are 12 x8 cm which 釘 'e slightly larger than Kerr described described (11 x 6.3 cm). As far as 1 know ，this species is only known from where Kerr and and 1 collected it.

Periploca Periploca purpurea Kerr FRUITS AND SEEDS DESCRIBED

This This species was described in 1938 from flowering material that Kerr (5576) collected in in early June 1921 from c. 1600 m in open evergreen forest on Doi Chiang Dao (Kerr ， 1938). 百le species was fruiting on 9 November 1995 when 1 collected it (Maxwell 95 ・1138) at 1775 m in prim 紅 y evergreen ，seasonal ，hardwood forest in rugged limestone te 町剖 n in the same 釘 'ea that Kerr must have found it ，i. e. 血ev a1 1ey below Doi Luang. Fortunately ，one specimen had a few flowers ，undoubtedly the last ofthe rainy season ，血 us 1 was able to identi 町it. It is a woody climber with a bas a1 diameter of 3.5 cm. Th eb 杭 is is thin ， peeling and roughened ，blackish; white latex is abundan t. Folli c1 es paired ，spreading at c. 45 ・(i. e. “V へlike) ，s阿 ght ，uniformly cylindric ，slightly narrowed narrowed at the tip ，finely s住iate when dry ，glabrous ，green ，9-12 'cm x4 mm. Seeds linear ，3 ・angled ，住 uncate at the tip ， acute at the base; funicular sc 紅 a distinct ， medial line on one side for a1 most the entire length of the seed ，glabrous ，blackish (6 ー) 12-14 x 1. 5- 1. 75 mm. Coma si1 ky ，white ，3 0- 35 mm long. Vegetatively 出is species is distinguished by its relatively sma l1， linear-lanceolate leaf blades blades with a prom 泊ently acuminate tip and fine in 住amargin a1 nerve. As with Heterostemma gracile gracile Kerr ， P. purpurea is only known ， as far 槌 1 can determine ，企 om the type locality. Subsequently Subsequently 1 collected the first flowering materi a1 of the year of this species on 出e summit area of Doi Nahng ， which is adjacent to Doi Luang ， at 1650 m on 18 March 1996

(Maxwell (Maxwell 96 ・390).

PHYTOLACCACEAE

Phytolacca Phytolacca americana L. NEW GENERIC AND SPECIES RECORD ，fig 町 e7

LARSEN (1992) notes 白紙出e genus Phytolacca L. has yet to be recorded from Th ailand. Two species ， viz. P. americana L.組 d P. acinosa Roxb. 釘 'e ，however ，known from Indo-China Indo-China (LARSEN ， 1989). 1 found flowering material of P. americana in Doi Chiang Dao W i1 dlife Sanctuary ， Chiang Chiang Mai Pr ovince at 白e east end of Doi Luang V a1 1ey at 1750 m on 8 November 1995 (Maxwe l1 95-1122). It was growing in thin so i1 on rugged limestone terrain in a shaded 20 J.F. MAXWELL

place. place. in deg t: aded ， primary evergreen ，seasonal hardwood forest as 叩 erect ，branching ， succulent succulent herb 1. 5m tall. From my observations it seems to be an annual herb 佃 d quite rare. rare. Phytolacca Phytolacca americana ，originally from North America ，is now found throughout much of 白e world. According to LARSEN (1 989) it is cultivated in tempera 句 areas for its fruits which 紅 e used to colour wine and also for medicinal pu 中 oses by people in Laos. and Vietnam. Vietnam. It s presence on Doi Chiang Dao would seem to be due to bird dispersal from Indo-China. Indo-China.

LORANTHACEAE

Helixanthera Helixanthera pierrei D 佃 s. NEW RECORD

Danser Danser (1938) described this species and indicated 出at it is known only from material coll~cted by Pierre in Tpong Province ，on 血e westem coast of at 1500 m. It is is distinguished by having distinct fibers at broken points; 5-merous ，sessile flowers; and styles styles which reach the base of the anthers. The gibbous ，cupular ，suborbicular bracts are also also distinc t.

1 found flowering material of 血is species ne 紅 Mae Mawn Village in Jae Sawn National P 紅 k，Muang Bahn District ，Lampang Province ， at 850 m elevation in mixed evergreen + deciduous ，seasonal ，hardwood forest on 17 Febru 紅 Y 1996 (Maxwell 96-258). It was growing growing on older stems of Rourea minor (Gaertn.) Leenh. ssp. minor (Connaraceae) ，組 evergreen evergreen woody climber ， in a partly. shaded area below the .f orest canopy. The following notes notes supplement Danser's description. Shrub with stemslbranches up to c. 1m long; oldest oldest stems dull blackish ， younger ones light green and tur 凶ng grey. Inflorescence axes ， bracts ， calyx dull greenish and dull reddish ， tuming red. dull Petals dull red ，anthers dull dark dark red;. leaf blades dark glossy green above. bright light green undemeath. Danser Danser appropriately named 血is species in honour of Jean Baptiste Louis Pierr.e (1 833 ・1905) ，a famous and prolific French botanis t.

MORACEAE

Ficus Ficus nervosa Hey. ex Roth var. nervosa NEW RECORD ，figure 8

CORNER (1965) notes 白紙 this variety is found in Sri Lanka ，India ，Burma ，China ， Hong Kong ，Taiwan ， and. Vietnam. GAG 悶 PAIN'S (1 928) description and illustrations confirm confirm the identification of Maxwell 95 ・ 1278 from Jae Sawn National Park ，Muang Bahn (Pan) (Pan) District ，Lampang Province. It was collected in an open area along a stream at Mae Mee Nawk (K 紅 en) Village at 550 m on 4 December 1995 in very degraded ，fire-prone ， deciduous deciduous forest with much bamboo on shale bedrock. The specimens were from a 位'ee 15 m tall ，ddh 50 cm ， with thin ，smooth ， grey bark and and sparse white latex. The species is immediately recognized by its axillary ，smooth ， glabrous ， globose figs 1. 5-2 cm diameter which are on peduncles 5-7 mm long. 百le BOT ANICAL NOTES 21

bracts ，which 訂 e inserted at the base of the peduncle ，紅 e sheathing ，2 ・lobed to the middle ， and and c. 1. 5 mm long. Unlike most other species of Ficus in Thailand ， the absence of bracts at at the base of the figs of Ficus nervosa is quite diagnostic. The closely reticulate (i.e. nervose) nervose) tertiary venation on the leaf blades is also characteristic and recalls the basic pattem pattem found in many species of Artocarpus (Moraceae). The other variety of this species ，var. minor King ，is found in Sri Lanka and peninsular India. India.

DIOSCOREACEAE

Dioscorea Dioscorea velut 伊es Pr. & Burk. FRUITS DESCRIBED

PRAIN & BURKILL (1 927 ， 1934) described this species in 1914 based on male and femal 巴 flowering material collected in the Southem Shan States of Burma. A mal 巴 specimen was collected by Kerr (5581) in June 1921 (at 1600-1800 m) and by Banzinger (637 at 1530 1530 m) on 15 July 1990 ， both in rugged Ii mestone terrain in Doi Chiang Dao Wildlife Sanctuary ， Chiang Mai Province.

collected 1 collected fruiting material of this species (Maxwell 95 ・ 1075) on 5 November 1995 at at 1625 m elevation in the rugged Ii mestone valley below Doi Luang in Doi Chiang Dao Wildlife Wildlife Sanctuary in an open ，fire-damaged ， former evergreen ，seasonal hardwood forest which which was cleared for opium cultivation and is now overgrown with weeds and secondary growth. growth. Infructescences Infructescences racemose ，pendulous ， 37-56 cm long; axes very finely and sp 紅 sely puberulous ，glabrescent ，slighly angular ，green. Pedicels solitary ，2 0- 35 mm apart ，reflexed ， 5-6 mm long. Bracteoles 2 ，inserted at different levels near the middle of the pedicel ， adpressed ， narrowly triangular ， to 1.5 x. 0.5 mm. Capsules elliptic ， broadly emarginate at the the tip ，truncate at the base ，glabrous ，green; overall size (including wings) 22-23 x 30-32 mm. Seeds immature. Vegetatively Vegetatively this species is easily recognized by its simple leaves with leaf blades which which are acuminate at the tip ， deeply cordate at the base ， and conspicuous ，fine ，persistent pubescence pubescence on the undersurface. A male collection (Maxwell 95-1073) was made in the same place as the fruiting materia l. Finally ， the original dimentions of up to 9x 5 cm for the the leaf blades can be increased to 15 x 10 cm with the petioles up to 9 cm long.

ACKNOWLEDGMENTS

My sincere thanks 紅 e given to Dr. J.F. Veldkamp and Dr. W. Hetterscheid of the R 討ksherbarium ，Leiden ，Netherlands for their continued collaboration and support. They were were kind enough to provide me with important literature information and constructive comments for some of the taxa discussed in this pape r. 22 22 J.F. MAXWELL

RE 陪 RENCES

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