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Volume 11, Issue 1, 2020 ISSN 2191-3099 This Journal Was Published on March 1St, 2020 Schütziana The Gymnocalycium Online Journal Volume 11, Issue 1, 2020 ISSN 2191-3099 This journal was published on March 1st, 2020 Content Wick, Mario Editorial p. 2 Schädlich, Volker The Seeds of the Genus Gymnocalycium Pfeiffer ex Mittler pp. 3-21 Wick, Mario Part 4: Subgenus Macrosemineum Strub, Thomas Gymnocalycium meregallii Bercht 2012 pp. 22-44 Published: March 1st, 2020 Legal notice Publisher: WORKING GROUP SCHÜTZIANA, Mario Wick, Fichtenweg 43, 14547 Fichtenwalde, Germany Editorial team and responsible for the content: www.schuetziana.org/contact.php SCHÜTZIANA is the journal of the WORKING GROUP SCHÜTZIANA. Source of supply: SCHÜTZIANA is available only as a pdf-file via World Wide Web and can be downloaded from: www.schuetziana.org/download.php. The content of the respective article expresses the opinion of the author and not necessarily that of the WORKING GROUP SCHÜTZIANA. The issues of SCHÜTZIANA are free of charge and may be distributed freely. The content and the pictures of the articles of SCHÜTZIANA are property of the author and may not be used for other purposes than reading, printing and storing without permission. © 2020 WORKING GROUP SCHÜTZIANA. Al rights reserved. ISSN 2191–3099 Cover picture: Gymnocalycium hyptiacanthum subsp. uruguayense, rose-coloured flower, locality VoS 2974, Cañada Zana de Los Novos, province Artigas, Uruguay, altitude of finding: 141 m a.s.l. (Picture: V. Schädlich). Schütziana 11(2020)1 p. 1 Editorial Dear Gymnocalycium friends Mario Wick On looking at the Schütziana Internet homepage, you will probably have noticed that a young Chinese student of agricultural sciences from Ningbo, China, has agreed to translate Schütziana into Chinese. The editorial staff is very grateful for that, as we thus add 1.4 billion new readers to our audience – even if there might not be quite as many in the end. This issue is going to complete the series “The seeds of the genus Gymnocalycium Pfeiffer ex Mittler”, presenting the fourth part about the subgenus Macrosemineum. The four parts would certainly have offered enough information for a book; therefore, we have decided to publish all four parts in a special edition of Schütziana. As all the editorial staff are volunteers, this will not be possible right away. New information as to the Gymnos can be reported constantly. This is why we had to revise the distribution maps of the subgenus Macrosemineum shortly before the completion of the first issue 2020. Ludwig Bercht, together with his travel companions, found Gymnocalycium hyptiacanthum subsp. uruguayense in Argentina as well, south of the town Concepción del Uruguay. In the second contribution to this issue, our editorial member Thomas Strub gives us an understanding of Gymnocalycium meregallii, first described by Ludwig Bercht in Schütziana in 2012. This plant was the topic of the International Gymnocalycium Meeting in September 2019. This species has certainly not been distributed widely in collections so far, so that the paper presented here is a valuable contribution to the knowledge of this species as well as Gymnocalycium andreae and Gymnocalycium bruchii, which also occur in the distribution area. As usual, enjoy yourselves reading! We would like to express our warmest thanks to Mrs Iris Blanz (Fernitz, Austria), to Mr Brian Bates (Bolivia) and to Mr Graham Charles (United Kingdom), who support us with the translation into English, to Mrs Larisa Zaitseva for the translation into Russian (Tscheljabinsk, Russia), to Mr Takashi Shimada (Japan) for the translation into Japanese, to Mr Jiahui Lin (China) for the translation into Chinese and to Mr Daniel Schweich (France), who has mirrored our publications under http://www.cactuspro.com/biblio/. Schütziana 11(2020)1 p. 2 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum The Seeds of the Genus Gymnocalycium Pfeiffer ex Mittler Part 4: Subgenus Macrosemineum Volker Schädlich Bergstraße 1, 03130 Spremberg, Germany Email: [email protected] Mario Wick Fichtenweg 43, 14547 Fichtenwalde, Germany Email: [email protected] ABSTRACT The authors present a paper about the seeds of the genus Gymnocalycium. The species are introduced with a picture at their locality, of their habitat and their seeds. A map of the localities of the individual plants completes the publication. Part 4 deals with the subgenus Macrosemineum. KEYWORDS: Cactaceae, Gymnocalycium, Macrosemineum, angelae, buenekeri, denudatum, fleischerianum, horstii, hyptiacanthum, mesopotamicum, netrelianum, paraguayense. INTRODUCTION The fourth part of our paper about the seeds of the genus Gymnocalycium deals with the taxa of the subgenus Macrosemineum. To establish the maps the GIS software QGIS, which is free of charge, was used (https://www.qgis.org/de/site/forusers/download.html). The background of the maps is formed by Google Inc. maps. Gymnocalycium denudatum (Link & Otto) Pfeiffer ex Mittler (1844) is the type species of the subgenus Macrosemineum. Schütziana 11(2020)1 p. 3 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Genus Gymnocalycium Pfeiffer ex Mittler (1844) Fig. 1: General map showing the distribution areas of the subgenera of the genus Gymnocalycium. Schütziana 11(2020)1 p. 4 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Subgenus Macrosemineum Body flat spherical to spherical, also short, column-shaped at older age, often with shoots, fibre roots. Spines ± straight or bent, flowers sprouting from the apex, funnel- or bell-shaped, often determined either male or female (dioecious). Fruits spherical or spindle-shaped, green when ripe, lacerating vertically or becoming soft. Seeds 1.0-2.0 mm in size, helmet-shaped, testa black, large hilum-micropylar area. Occurrence: Uruguay, Brazil (Rio Grande do Sul), southern Paraguay and north-eastern Argentina. Fig. 2: Map of the distribution area of the subgenus Macrosemineum. Schütziana 11(2020)1 p. 5 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium angelae Meregalli (1998) Fig. 3: Gymnocalycium angelae VoS 2919, Cerro de Susini, Province Corrientes, Argentina, 128 m. Fig. 4: Habitat of Gymnocalycium angelae VoS 2919. Schütziana 11(2020)1 p. 6 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Fig. 5: Seeds of Gymnocalycium angelae Gf 1300. Fig. 6: Locality of Gymnocalycium angelae Gf 1300 and VoS 2919. Schütziana 11(2020)1 p. 7 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium buenekeri Swales (1978) Fig. 7: Gymnocalycium buenekeri VoS 2899, Sao Francisco de Assis, Province Rio Grande do Sul, Brazil, 191 m. Fig. 8: Habitat of Gymnocalycium buenekeri VoS 2899. Schütziana 11(2020)1 p. 8 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Fig. 9: Seeds of Gymnocalycium buenekeri LB 586. Fig. 10: Locality of Gymnocalycium buenekeri LB 586 and VoS 2899. Schütziana 11(2020)1 p. 9 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium denudatum (Link & Otto) Pfeiffer ex Mittler (1844) Fig. 11: Gymnocalycium denudatum VoS 3012, northeast from Dom Pedrito, province Rio Grande do Sul, Brazil, 257 m. Fig. 12: Habitat of Gymnocalycium denudatum VoS 3012. Schütziana 11(2020)1 p. 10 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Fig. 13: Seeds of Gymnocalycium denudatum Gf 18, Minas do Camaqua, Kreuzberg, Province Rio Grande do Sul, Brazil. Fig. 14: Localities of Gymnocalycium denudatum Gf 18 and VoS 3012. Schütziana 11(2020)1 p. 11 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium fleischerianum (Meregalli, Metzing & Kiesling) Vala (2003) Fig. 15: Gymnocalycium fleischerianum VoS 2150, west of Itacurubi, Province Cordillera, Paraguay, 205 m. Fig. 16: Habitat of Gymnocalycium fleischerianum VoS 2150. Schütziana 11(2020)1 p. 12 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Fig. 17: Seeds of Gymnocalycium fleischerianum VoS 2150. Fig. 18: Locality of Gymnocalycium fleischerianum VoS 2150. Schütziana 11(2020)1 p. 13 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium horstii Buining (1970) Fig. 19: Gymnocalycium horstii VoS 3026, Santana da Boa Vista, Province Rio Grande do Sul, Brazil, 279 m. Fig. 20: Habitat of Gymnocalycium horstii VoS 3026. Schütziana 11(2020)1 p. 14 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Fig. 21: Seeds of Gymnocalycium horstii Gf 105. Fig. 22: Locality of Gymnocalycium horstii Gf 105 and VoS 3026. Schütziana 11(2020)1 p. 15 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium hyptiacanthum (Lemaire) Britton & Rose (1922) Fig. 23: Gymnocalycium hyptiacanthum subsp. uruguayense VoS 2952, east of Quaraí, Province Rio Grande do Sul, Brazil, 198 m. Fig. 24: Habitat of Gymnocalycium hyptiacanthum subsp. uruguayense VoS 2952. Schütziana 11(2020)1 p. 16 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Fig. 25: Seeds of Gymnocalycium hyptiacanthum subsp. uruguayense VoS 2952. Fig. 26: Locality of Gymnocalycium hyptiacanthum subsp. uruguayense VoS 2952. Schütziana 11(2020)1 p. 17 Schädlich & Wick – The seeds of the Genus Gymnocalycium. Part 4: Subgenus Macrosemineum Gymnocalycium
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