BU'JJETIN-No. lI�

Recent British Columbia spring and coho salmon tagging experiments, and a comparison with those conducted from 1925 to 1930

By

D. J. MILNE Fisherw8 Research Board 01 Canada Biological Station, Nanaimo, B.C.

PUBL ISHED BY THE FISHERIES RESEARCH

BOARD OF CANADA UNDER THE CON TROL OF

THE HONOURABLE THE MINISTER OF FISHERIES

fTAWA, 1957 fer50'cents

��, ------� BULLETIN No. 113

Recent British Columbia spring and coho salmon tagging experiments, and a comparison with those conducted from 1925 to 1930

By D. J. MILNE Fisheries Research Board of Canada Biological Station, Nanaimo, B.C.

PUBLISHED BY THE FISHERIES RESEARCH BOARD OF CANADA UNDER THE CONTROL OF THE HONOURABLE THE MINISTER OF FISHERIES

OTTAWA, 1957 w. E. RICKER

N. M. CARTER Editors

(ii) Bulletins of the Fisheries Research Board of Canada are published from time to time to present popular and scientific information concerning fishes and some other aquatic animals; their environment and the biology of their stocks; means of capture; and the handling, processing and utilizing of fish and fishery products. In addition, the Board publishes the following: An A nnual Report of the work carried on under the direction of the Board. The Journal of the Fisheries Research Board of Canada, containing the results of scientific investigations. Atlantic Progress Reports, consisting of brief articles on investigations at the Atlantic stations of the Board. Pacific Progress Reports, consisting of brief articles on investigations at the Pacific stations of the Board. The price of this Bulletin is 50 cents (Canadian funds, postpaid). Orders should be addressed to the Queen's Printer, Ottawa, Canada. Remittance made payable to the Receiver General of Canada should accompany the order. All publications of the Fisheries Research Board of Canada still in print are available for purchase from the Queen's Printer. Bulletin No. 110 is an index and list of publications of the Board and is priced at 75 cents per copy postpaid.

(iii) 91243-l!

CONTENTS

PAGE

ABSTRACT...... (vii) INTRODUCTION...... 1 ACKNOWLEDGMENTS...... 2

METHODS...... 2

REVIEW OF THE 1925-30 TAGGING EXPERIMENTS...... 7 Spring salmon...... , ...... 7 Coho salmon...... 7 SUMMARY OF THE 1949 TAGGING EXPERIMENTS...... 9 Spring salmon...... 9

Coho salmon...... 10 SPRING SALMON EXPERIMENTS FROM 1950 TO 1952...... 11 Queen Charlotte Islands...... 11 West coast of Vancouver Island-Northern part...... 11 -Southern part...... 14 COHO SALMON EXPERIMENTS IN 1950 AND 1951 ...... , ...... 18 Queen Charlotte Islands...... 21

West Coast of Vancouver Island-Northern part...... 22

-Southern part...... 23

COMPARISON OF THE EARLY AND RECENT TAGGING EXPERIMENTS...... 25 Spring salmon...... 25 Coho salmon...... : ...... 30 SUMMARy...... 33

REFERENCES...... 34 ApPENDIX...... 35

(v)

ABSTRACT

From 1949 to 1952 a total of 1,222 spring (chinook) salmon and 2,190 coho salmon were tagged off the coast of British Columbia to demonstrate the ocean migrations and the rivers of origin of fish exploited by the offshore troll fishery. These represent the Canadian part of a coastwide tagging program carried out from Alaska to California through the coordination of the Pacific Marine Fisheries Commission. Most spring salmon were on long spawning migrations, travelling in a southeasterly direction along the continental shelf towards the larger rivers, such as the Columbia and Fraser. Offthe southwestern part of Vancouver Island many small imma­ ture fish were tagged and recovered from various directions during the next two years. All coho salmon were on shorter spawning migrations, travelling in many directions from offshore waters towards the numerous small coastal streams.

Similar experiments carried out from 1925 to 1930 involved 6,457 spring salmon and 7,126 coho salmon. A comparison between the experiments conducted offthe west coast of Vancouver Island in the two periods revealed the following: For spring salmon, the percentages returned were the same (12.9). However small fish appear to provide fewer returns than large fish and more small fish were caught and tagged in recent years. Also more of the tagged fish were recaptured in the recent period off British Columbia (71% compared to 16%) and in the Fraser River (12% compared to 5%) while less were caught in United States waters (29% compared to 82%) and in the Columbia River (16% compared to 60%). This implies that the trollers are now fishingmore intensively near the tagging areas and on stocks which spawn in British Columbia rivers rather than on the Columbia River stocks which have decreased greatly in size and import­ ance in recent years. For coho salmon, the percentages returned were higher recently (9.2 compared to 6.1) owing to an increase in the fishing intensity, particularly in United States waters.

The outside troll fishery of British Columbia has been shown to catch a mixture of fish from many streams along the Pacific coast. For this reason certain conservation measures for spring and coho salmon should be considered on an international basis.

(vii) INTRODUCTION

In 1948 the Pacific Marine Fisheries Commission resolved to undertake an extensive offshore tagging program along the coast of California, Oregon and Washington to determine the movement of chinook or spring salmon (Oncorhynchus tshawytscha) and coho salmon (0. kisutch) at sea and the rivers of origin of the fish exploited in various areas. Preliminary results of the 1948 and 1949 tagging experiments by the three states have been reported by the Commission in its Bulletin No.2 (1951). Both Alaska and Canada cooperated to make the program coastwide. It was hoped that by this means the inter­ national mixing of stocks could be assessed and, where necessary from a conservational viewpoint, common regulations could be formulated.

Since the Biological Board of Canada had tagged extensively off British Columbia from 1925 to 1930 (Pritchard, 1933) a repetition of these experiments would also afford an opportunity to assess any differences in fishing drain which had occurred over the last twenty years. In recent years the British Columbia troll fleet has increased in size and efficiency and both the fishermen and industry have expressed a growing concern over the conservation of the spring and coho salmon stocks.

In 1949 a commercial trolling vessel was chartered by the Fisheries Research Board of Canada to tag salmon off the weat coast of Vancouver Island. Pre­ liminary results have been reported by Neave (1951). A brief summary of the final results will be given, which include a few late returns.

In 1950 tagging operations on a red�ed scale were repeated off the west coast of Vancouver Island by purchasing live fish from commercial trollers for tagging purposes. All fish tagged were small, less than 8 pounds in weight, with an average length of 25 inches. Tagging locations were off Barkley Sound in the south and off Kyuquot Sound in the north.

In 1951 two tagging experiments were conducted offeach side of the northern end of Vancouver Island and the northern end of Queen Charlotte Islands. Later in the fall, coho salmon were tagged off the southern tip of Vancouver Island from Sooke traps.

In the summer of 1952 spring salmon were tagged from Sooke traps. The results of the experiments from 1950 to 1952 will be discussed in detail.

1 91243-2 ACKNOWLEDGMENTS

The success of the tagging program has been due to the excellent cooperation which has been received from the fishermen and those connected with the fishing industry, who conscientiously returned tags and to others who assisted in the actual tagging operations. In particular, sincere thanks are extended to the following trollers: Messrs. L. Ives, S. Olsen, W. Taylor, G. Holden, L. Ochs, A. Oswald, C. Lindrupson, E. Akerley, and ]. Stace-Smith; to the Fishermen's Co-operative Association and to ]. H. Todd & Sons, Ltd. for assistance rendered. To the taggers, Messrs. W. G. Calderwood (Barkley Sound, 1950), E. A. R. Ball (Kyuquot Sound, 1950 and Sooke, 1951), V. R. Taylor (Quatsino Sound, 1951), W. J. Mitchell (Queen Charlotte Islands, 1951) and D. Denbigh (Sooke, 1952) much appreciation is also due. Mr. E. A. R. Ball and Miss M. Philp were responsible for tabulating many of the data obtained.

METHODS

The majority of the tagged spring and coho salmon were caught by trolling gear; a few of each species were obtained from the traps at Sooke. The former resulted in lower returns than the latter, probably because the former fish were caught by hooks and hauled up from considerable depths by the trolling gear. Trolling is the only fishing method employed in the ocean, on the feeding grounds and along the migration routes; most net fisheriestake place on mature fish either near or on the way to the spawning rivers. The present trolling areas adjacent to British Columbia (Fig. 1) show the places of capture of these two species of salmon. All areas are within 50 miles of shore and 100 fathoms or less in depth, and are either along the continental shelf or in inside waters. Most coho salmon are caught in the top 10 fathoms and within 20 miles from shore, while most spring salmon are taken closer inshore and deeper. Coho salmon are often observed to jump clear of the water surface and are usually found in clear, blue water. Spring salmon seldom jump while feeding in brown, plankton-filled water. The intensity and location of the fisheries affected both the area of tagging from trolling boats and the distribution of the recoveries by both the troll and net fishing fleets. Thus the recoveries indicate only partial ocean migration routes, minimum rates of travel, and in some cases the final destination to the larger spawning streams. In the early experiments (1925-30) numbered metal "cattle" tags were attached to the tail of the fish (Williamson, 1927). Recently (1949-52) plastic "button" tags (Peterson discs) were attached to the back of the fish by means of nickel pins (Fig. 2). Some abrasion and mortality occurs (Milne and Ball,

2 " j "<'.

BRITISH COLUMBIA

PACIFIC OCEAN

N

SEA MILES t 'l--__---' 5»L'O __� ''' O

TROLL FISHING AREAS •

ONE HUNDRED FATHOM LINE - - ---

FIGURE L The troll salmon fishing areas in British Colnmbia,

1956), but fish that were seriously damaged by hooking or handling were not tagged, There is also evidence (Calhoun, Fry and Hughes, 1951) that there is a small loss of tags due to corrosion of the nickel pins near the head, A detailed examination of 88 of the returned pins showed that only three had minor signs of corrosion and none were seriously weakened. The three were in the group of 23 pins which had been out from 200 to 800 days. Despite a re\vard of 50 cents for each recovery, a few of the tags that are recovered are never

3 91243-2! FIGURE 2. The method used to tag salmon. Above-spring salmon in tagging box on troller. Below-Petersen disc tag and how it is attached to fish.

4 returned. For some which are returned complete information is lacking, and in a few instances only the unnumbered tag or "baffle" is recovered. Because of these defects-an unknown mortality, shedding of tags, and incomplete returns -the percentage returned is an inadequate index of the degree of exploitation by the fishery. However, from those returned, a qualitative description of the movements of the fish in the ocean can be made and in some cases the final des­ tination can be determined. At the time of tagging, the fork length (from the tip of the nose to the fork of the tail) of the fish was recorded and a few scales were obtained for age determination. A summary of the length frequencies of the fish tagged and returned for the experiments conducted from 1949 to 1952 is given in Tables A and B of the Appendix, while ages of the spring salmon tagged and returned in the 1950-52 experiments are given in Table C (Appendix). These data are needed to determine how representative the tagged fish are of the commercial catch, and to explain certain differences in the results of experiments conducted in different seasons and areas.

5 BRITISH COLUMBIA

PACIFIC OCEAN

H

SEA MILES o 100

FIGURE 3. The tagging locations showing the number of fish tagged (solid line circles) and the general migration routes of spring salmon tagged from 1925 to 1930.

6 REVIEW OF THE 1925-30 TAGGING EXPERIMENTS

SPRING SALMON In 15 separate experiments conducted in 10 areas, a total of 6,457 spring salmon were tagged off British Columbia from 1925 to 1930 inclusive, of which 761 or 11.8% were recovered. The various tagging locations with numbers of fish tagged and the main migration pattern are shown in Figure 3. The width of the probable migration routes are weighted in magnitude. A general summary of the migration of spring salmon based on these experiments is given by Clemens, Foerster and Pritchard (1939). The majority of the spring salmon travelled long distances in a south­ easterly direction along the continental shelf to the larger rivers; many were captured along the way. Columbia River recoveries predominated: for example, from fish tagged off the west coast of Vancouver Island, 60% of the returns were from the Columbia River and 5% of the returns were from the Fraser River. Few fish were recaptured near the tagging areas and 82% were caught in United States waters. Most fish were recovered in the same year as tagged, except for those tagged off Barkley Sound from which numerous returns were recovered in the two years following the tagging year. Many of the fish tagged off the northeast coast of Vancouver Island were recovered from Georgia Strait (21% of the returns) and Puget Sound (45% of the returns). Those tagged in Georgia Strait showed a more local distribution with 50% of the returns from the Fraser River and only 8% from Puget Sound. As the tagging locations moved northward the distribution of recoveries also shifted to the north. From fish tagged off the northwest coast of the Queen Charlotte Islands, 58% of the returns were from United States waters to the south (26% from the Columbia River) and 2% were from southeastern Alaska. The fish tagged in Hecate Strait showed a more localized distribution in British Columbia waters with only 35% of the returns being from United States waters, mostly from the Columbia River (26%).

COHO SALMON In 19 separate experiments conducted in 10 areas, a total of 7,126 coho salmon were tagged off British Columbia from 1925 to 1930 inclusive, of which 632 or 8.7% were recovered. Tagging locations and the main migration pattern are shown in Figure 4. More coho salmon were tagged in inside waters than in the spring salmon operations. No convenient summary is available, but the experiments have been reported by Williamson (1927, 1929), Williamson and Clemens (1932), Clemens (1930) and Pritchard (1934).

7 The coho salmon radiated in more directions from the tagging areas and migrated shorter distances than the spring salmon. In general they appeared to move in all directions towards numerous small streams along the coast. Many small spawning populations contributed to the fish stock at any tagging locality.

BRITISH COLUMBIA

t

PACIFIC OCEAN

SEA MII..E5 100

FIGURE 4. The tagging locations showing the number of fish tagged (solid line circles) and the general migration routes of coho salmon tagged from 1925 to 1930.

8 All tagged fish were recovered in the same year as tagged since nearly all coho salmon are either caught or spawn in their third year of life. From tagging experiments off the west coast of Vancouver Island, the majority of the recoveries were to the south and east from the Fraser River and Puget Sound, with a few from off the \Vashington coast. From fish tagged off the northeast coast of Vancouver Island most were recovered from Georgia Strait (Fraser River 14% of the returns) with a few from Puget Sound. Less than 10% of the recoveries were from areas north of the point of tagging. Approximately one-quarter of the fish tagged at Sooke traps, on the southern end of Vancouver Island, were recovered in Georgia Strait and Puget Sound, with the majority recaptured in the Fraser River. In Georgia Strait the small cohos, popularly called "bluebacks" radiated from the place of tagging in all directions with one-quarter of the returns from Puget Sound. It appears that they probably spend their entire ocean life in inside waters. The same indication of restricted movement was evident in parts of Hecate Strait where the popula­ tions of many small coastal streams intermingle. From the northern coast of the Queen Charlotte Islands, the recoveries were distributed from southeastern Alaska (19% of the returns) to as far south as Vancouver Island.

Sm1MARY OF THE 1949 TAGGING EXPERIMENTS In 1949 both spring and coho salmon were tagged off the west coast of Vancouver Island and the results of the Canadian experiments have been reported by Neave (1951). Since this report there have been 15 returns for spring salmon and 3 for coho. A complete list of the recoveries is given in the Appendix, in Table D for spring salmon and in Table E for cohoes. A summary of the final returns is as follows:

Tags recovered in Tags Percentage Species Area applied recovered 1949 1950 1951

Spring s:tlmon ...... Barkley Sound ..... 772 29 37 5 9.2

Quatsino Sound .... 45 11 1 - 26.6

Coho salmon ...... Barkley Sound... .. 40 5 - - 12.5

Quatsino Sound .. .. 470 41 - - 8.8

SPRING SALMON Off Barkley Sound, the size of the fish caught in 1949 was small compared to recent years with the result that 68% of those tagged were less than 67. 5 cm. in fork length (about 8 lb.) and 86% were three years of age or less. The large number of recoveries (60% of total returns) in the two years following the tagging

9 91243-3 year indicate that many of the fish tagged were immature fish. The returns in both 1949 and 1950 show a wide dispersal in all directions with 30 or 45% of the recaptures in the tagging area. In 1951, four of the fivereturns were from the Columbia River and the remaining one was captured in the tagging area. Thus the area off Barkley Sound is a feeding area for immature spring salmon. In 1949 a total of 315 spring salmon were tagged off the southwestern part of Vancouver Island by the Washington State Fisheries Department. Preliminary results (Bulletin 2, 1951) from 58 returns (18%) also indicate that this is a feeding area for immature spring salmon with the main migration towards the Fraser and Columbia Rivers. Off Quatsino Sound, on the other hand, the fish were larger and a high percentage was recovered in the tagging year. Most were taken at considerable distances to the southeast and one was recovered in 1950 in the Columbia River. Thus almost all of the fish caught off Quatsino Sound are maturing fish on their spawning migration.

COHO SALMON The coho salmon tagged at both locations in 1949 were maturing fish which migrated in all directions around both ends of Vancouver Island. In 1949 a total of 35 coho salmon were tagged by 'Washington State off the southwestern part of Vancouver Island. The return of 11 tags (Bulletin 2, 1951) or 31% was higher than that in the Canadian experiments but a similar migration in all directions indicates that this is a common feeding ground for coho salmon from many river systems.

10 SPRING SALMON EXPERIMENTS FROM 1950 TO 1952

A summary of the numbers of spring salmon tagged in the five experiments from 1950 to 1952 together with the numbers and percentages of the returns is given in Table I. For comparison the results of the two Canadian experiments conducted in 1949 are also shown in the table.

The areas have been arranged in geographical order from north to south. Following this order each area is discussed in greater detail in the text. Complete data for the recoveries are given in Tables F to J in the Appendix. These are arranged in each area in the order of the date of tagging for each recovery year.

The international aspect of the migrations is apparent but, in the 1949-52 experiments, the number of returns from United States waters and the Columbia River are much less than they were in the 1925 to 1930 experiments. For the experiments off Vancouver Island the recoveries in United States waters have dropped from over 80% to under 40% and the Columbia River recoveries have dropped from over 50% to 25% or under for the two periods. This is an indication of both an increased fishery drain by trolling boats in the tagging areas and a reduction in the stocks of the Columbia River.

QLJEEN CHARLOTTE ISLANDS In 1951 a total of 64 spring salmon were tagged from commercial trollers operating off the northwestern end of the Queen Charlotte Islands; 41 off the west coast and 23 off the north coast. In this area most of the fish are large, averaging over 15 Ibs., and are 4 or 5 years of age. The 7 recoveries were all from the 42 fish tagged at a length of over 75 cms. (30 in.). They were all recovered from 11 to 60 days later after migrating from 350 to 600 miles in a southeastern direction (Fig. S). The fish which was recaptured closest to the tagging area, also gave the fastest rate of travel; approximately 350 miles in 11 days or over 30 miles per day.

It appears that the majority of the fish in this area are maturing fish that are migrating around both sides of the Queen Charlotte Islands to spawn in rivers far to the southeast.

WEST COAST OF VANCOUVER ISLAND NORTHERN PART.�In 1951 the operations were similar to those off the Queen Charlotte Islands. A total of S4 fish were tagged off Quatsino Sound. Most of the fish in this area are large and 3 to 5 years of age. The 8 recoveries

11 91243-3� TABLE I.-Summary 01 spring salmon tagged in British Columbia waters in 1949 to 1952.

...

Geographical distribution of returns a

Number Number Percentage U.S. Tagging area Year tagged returned return Canadian waters Fraser Columbia waters exclusive River River of Alaska

---��- �---�-- ��--�� --�� -----

(/c) % % % % QUEEN CHARLOTTE ISLANDS N.W. COAST ...... 1951 64 7 11.0 57 43 14 -

f-'­ N VANCOUVER ISLAND N. W. COAST Quatsino Sd ...... 1949 45 12 26.6 75 25 25 25 Quatsino Sd ...... 1951 54 8 15.0 62 38 12 25 Kyuquot Sd ...... 1950 22 4 18.2 100 - 25 -

S. W. COAST Barkley Sd ...... 1949 772 71 9.2 75 25 8 14 Barkley Sd ...... 1950 140 38 27.2 61 39 16 16

S. COAST Sooke traps ...... 1952 125 28 22.4 72 28 57 4

.��- ��--�------�--�--- --_.-

- .- - - Total ...... 1,222 168 13.7 -

��- ----_.... _-----

a As percentage of the total tags returned. BRITISH COLUMBIA

N +

o SEA MILES 100

NO RTH WEST TAGGIN G AREAS 0 o NUMBERS 23 41 RETURN AREAS • • NUMBERS 3 4

FIGURE 5. Distribution of recaptured spring salmon tagged offthe west and north coasts of the Queen Charlotte Islands in 1951. are shown in Figure 6. The four fish over 80 cm. (32 in.) in length were recap­ tured the same year, three smaller fish were caught the next year, and one fish of 52 cm. (20 in.) was retaken two years after the tagging year. The majority of these fish were not maturing fish when tagged and the final movement was towards the Columbia and Fraser Rivers. The smallest fish grew from about 4 lbs. to 20 lbs. during the two years in the ocean. This is in contrast to the result in 1949 for this area, when all but one of the recoveries were made in the same year as tagged. In 1950 off Kyuquot Sound only 22 small fish, less than 8 lbs. in weight, were tagged. This is a select sample of 2- and 3-year-old fish since most of the

13 fish caught in this area were 3 and 4 years old. The four recoveries are shown in Figure 7. All of the fish had moved in a southeasterly direction along the west coast of Vancouver Island. One fish was recovered the next year and another fish was recovered two years after tagging, both about 100 miles from the tagging area. The latter fish had grown from 52 to 85 cm. (20 to 33 in.) in the two years.

In summary, the majority of the fish off the northwestern part of Vancouver Island are large maturing fishbut a portion of those less than 15 lbs. are immature feeding fish that will spawn in subsequent years. The direction of movement is southeasterly along the west coast of Vancouver Island.

SOUTHERN PART.-The 1950 tagging off Barkley Sound was similar to that off Kyuquot Sound in that only small fish less than 8 lbs. were tagged. Since more small fish in their third year are available in this area, 140 fish were tagged from commercial trolling boats. The majority were tagged in June and ranged from 39 to 67 cm. in length (16-26 in.). The distribution of the 38 recoveries is shown in Figure 7. The percentage return of 27.2 was almost three times as large as that in 1949 (Table I). Fifty-five percent of the returns were made in the next year after tagging and 23% were recovered in the second year following the tagging. Each year a marked radiation from the tagging area was demon­ strated. Most of the returns in the year of tagging were from areas north of the tagging area and from the Fraser River, while in the later years more returns were from the south in the Columbia River.

COLUMBIA N f o SEA MILES 100

R.

1951 1952 1953 TAGGING ARE A 0 NUMBERS 54 RETURN AREAS • II NUMBERS 4 3

FIGURE 6. Distribution of recaptured spring salmon tagged off Quatsino Sound, Vancouver Island, in 1951.

14 A N

SEA MILES 100

'RASE R.

BARKLEY SO. TAGGING AREAS 0 NUMBERS 140 RET URN AREAS • . NUMBERS 8 2

B COLUMBIA N

SE A MILES 100

FRASE R.

TAGGING AREAS o o RETURN AREAS • NUMB ERS 21

c COLUMBIA Nt SEA MILES 100 FRASE R.

TAGGING AREAS 0 0 RETURN AREAS . • NUMBERS 9

FIGURE 7. Distribution of recaptured spring salmon tagged off Kyuquot and Barkley Sounds, Vancouver Island, in 1950. A-Shows the returns in 1950; B-Shows the returns in 1951; C-Shows the returns in 1952. In summary, nine (24% of the returns) were taken by trollers off the west coast of Vancouver Island north of the tagging area; nine (24%) were recaptured in the tagging area; seven (18%) had moved east through the Strait of Juan de Fuca towards the Fraser River and thirteen (34%) had moved south towards the Columbia River. The first two groups were mostly feeding fish caught by trollers while the latter two groups were all maturing fish taken by gill-netters. These facts indicate a feeding or nursery area of immature fish off Barkley Sound which is followed by a spawning migration to the south or east. From the length and age of the tagged fish and from the lengths supplied by some of the fishermenwhen the fish were recaptured, the average ocean growth of the fish is as follows:

Fork length Dressed weight Age

---

em. in. lb.

When tagged or recovered in 1950 ...... 60 24 6 3 When recovered in 1951 ...... 80 31 14 4 \Vhen recovered in 1952 ...... 90 35 23 5 I This is in close agreement with the growth rate calculated from scale measure­ ments, and confirms the contention that the scale annuli represent true annual increments. During July and August, 1952, a total of 125 spring salmon were tagged from the Sooke traps. These were a selected group of small fish, averaging about 9 lbs., much smaller than the average size of fish caught in the traps. The 28 recoveries were maturing fish from 2 to 4 years old and they were all recaptured later in the same season, mostly by the inside net fisheries (Fig. 8). In detail, sixteen (57%) were taken in the Fraser River area, usually about a week later; six were recaptured in parts of Puget Sound, and four were recaptured again in the traps. Only two moved to the west; one was taken off the Washington coast; the other in the Columbia River. The latter was tagged in a group of nine fish from the most westward trap, three of which were later recovered from the Fraser River; yet it was recovered 39 days later up the Columbia River at Bonneville hatchery.

16 BRITISH COLUMBIA

0' SE A M1L ES'100

TAGGING AREA 0 NUMBERS 125 RETURN AREA • NUMBERS 28

FIGURE 8. Distribution of recaptured spring salmon tagged at Sooke traps, Vancouver Island, in 1952.

17 91243-4 COHO SALMON EXPERIMENTS IN 1950 AND 1951

A summary of the numbers tagged in the seven experiments in 1950 and 1951 together with the numbers and percentages of returns is given in Table II. The results of the two Canadian experiments conducted in 1949 are also included in the table. Complete data for the 1950-51 recoveries are given in Tables K to 0 in the Appendix.

COLUMBIA

N t o SEA MILES 0100

TAGGING AREA 0 NUMBERS 234 RETURN AREAS • NUMBERS I UNKNOWN 21 +

FIGURE 9A. Distribution of recaptured coho salrr:on tagged off the west coast of the Queen Charlotte Islands in 1951.

18 TABLE II.�-Summary of coho salmon tagged in British Columbia waters in 1949 to 1951. :0 "" "'" CN I "'" a "'il-' Geographical distribution of returns

Number Number Percentage U.S. Tagging area Year tagged returned return Canadian waters Alaska waters exclusive of Alaska

-�----

% % % % QUEEN CHARLOTTE ISLANDS N. coast ...... 1951 380 27 7.1 70 0 30

'vV. coast ...... 1951 234 22 9.4 72 5 23

...... VANCOUVER ISLAND \0 N.W. coast Cape Scott ...... 1951 178 12 6.8 66 34 - Quatsino Sd ...... 1951 366 28 7.7 61 39 - Quatsino Sd ...... 1949 470 41 8.7 85 15 - Kyuquot Sd ...... 1950 110 5 4.5 80 20 -

S.W. coast Barkley Sci ...... 1949 40 5 12.5 40 60 -

Barkley Sci ...... 1950 262 41 15.6 57 43 -

S. coast

Sooke traps ...... 1951 150 32 21.3 56 44 - "- -,--

Total ...... - 2,190 213 9.7 - - -

-

a As percentage of the total tags returned. COLUMBIA

N t

TAGGING ARE A 0 NUMBERS 380 RE TURN AR EAS .. NUMBERS 26 I UNKNOWN +

FIGURE 9B. Distribution of recaptured coho salmon tagged off the north coast of the Queen Charlotte Islands in 1951.

The international aspect of the migrations is apparent, particularly in the areas adjacent to Washington and Alaska. In the 1949-51 experiments the number of returns from outside British Columbia waters is greater than in the early experiments twenty years ago. For example, from experiments con­ ducted during the two periods off the Queen Charlotte Islands the percentage of the returns from Alaska has increased from 19% to 27%; from experiments off the west coast of Vancouver Island, the returns from \Vashington have in­ creased from 22% to 30%; and from Sooke trap tags, returns have increased from 27% to 44%. This is probably because the fishing effort by the United States fishermen, both commercial and sport, has increased more in recent years than the effort in waters off British Columbia.

20 QUEEN CHARLOTTE ISLANDS In 1951, a total of 614 coho salmon were tagged off the northwestern end of Queen Charlotte Islands; 234 off the west coast and 380 off the north coast. All fish were maturing fish in their third year. The 27 (7.1%) recoveries for the west and the 22 (9.4%) recoveries for the north are shown in Figures 9A and 9B. From both locations the fish moved northeast and south towards many streams along the coast from Alaska to the Fraser River. Only 4 or 8% of the total returns were recaptured near the Queen Charlotte Islands. Thirteen or 27% of the total returns were recovered in southeastern Alaska. The majority probably spawned in the coastal streams of central and northern British Columbia. Only one fish tagged on the west coast was recovered off the coast of Washington. The fishappear to have migrated to the east from these tagging areas, and then radiate in all directions along the coast of British Columbia. The condition of the fishafter tagging was recorded but no marked difference is apparent in the numbers recovered. The majority of the fish were released in good condition and the few released in poor condition gave a relatively high return. Condition of fish : Good Fair Poor Not recorded

No. tagged ...... 536 57 19 2 No. returned ...... 43 4 2 Percent returned ...... 8.1 7.0 10.5

COLUMBIA

N

0, SEA MILES ,100

TA GGING AREA o NUMBERS 178

RETURN AREAS • NUMBERS 12

FIGURE lOA. Distribution of recaptured coho salmon tagged off Cape Scott, Vancouver Island, in 1951.

21 N

o SEA MILES ,,00

TAGGING AREA 0 NUMBERS 366 USA. RET URN AREAS • NUMBERS 28

FIGURE lOB. Distribution of recaptured coho salmon tagged off Quatsino Sound, Vancouver Island, in 1951.

\;VEST COAST OF VANCOUVER ISLAND NORTHERN PART.-In 1951 the operations were similar to those conducted off the Queen Charlotte Islands; that is, fish were tagged in two locations near the northern tip of Vancouver Island. At Cape Scott, 178 fish were tagged and off Quatsino Sound 366 fish were tagged. The 40 recoveries are shown in Figures lOA and lOB. The results of this experiment in relation to the heavy catch made in this area in 1951 have been reported by Milne (1952). For those tagged off the north,vestern tip of the island, 8 or 67% of the returns showed an eastward movement (Fig. lOA). For those tagged off the west coast, only 5 or 18% went around the north end of the island; the remainder migrated down the west coast of Vancouver Island (Fig. lOB). Thus the fish appear to segregate in this region and travel around both ends of the island. Estimates based on the time of tag returns, showed that the fish moved at an average rate of 5 miles per day and took about a month to migrate the length of Vancouver Island. At the entrance to the Strait of Juan de Fuca the fish appear to segregate with some going along the vVashington coast as far south as Oregon while others migrate eastward into Puget Sound. Almost one-half of the returns were recovered in United States waters which is much higher than in 1949 (11 % of the returns). In 1950 off Kyuquot Sound 110 fish of small size, less than 8 Ibs. in weight, 'were tagged and a low return of 5 fish was obtained. The majority probably migrated southeast along the west coast of Vancouver Island, but some fish below Cape Cook apparently migrated around the north end of the island (Fig. 11).

22 SOUTHERN PART.-The 1950 tagging off Barkley Sound was also of small fish under 8 lbs. in weight, varying in length from 42 to 70 cm. (17 to 27 in.). The recovery from the 262 fishtagged was the highest (15.6%) of the experiments using troll-caught fish. It was higher than the 12.5% returned from the smaller Canadian experiment conducted in this area in 1949 but much lower than the 31% returned from the small 1949 experiment conducted by the State of Washington (P.M.F.C., 1951). The 41 recoveries are shown in Figure 12. The fish radiated in all directions and demonstrated that coho salmon from this area migrate around both ends of Vancouver Island. Similarly to spring salmon, this region appears to provide a common feeding ground for many different populations. One-quarter of the recoveries were from Puget Sound and a total of over 40% were from United States waters. Since all fish were maturing, over were recaptured by the net fisheries as they migrated towards their spawn­ i ing streams.

COLUMBIA

N

o SEA MILES! 100

TAGGING AREA 0 NUMBERS 110 RETURN AREA 0» NUMBERS 5

FIGURE 11. Distribution of recaptured coho salmon tagged off Kyuquot Sound, Vancouver Island, in 1950.

In 1951, during September, 150 fish varying in length from 39 to 78 cm. (15 to 31 in.) were tagged at the southern end of Vancouver Island at Sooke traps. The recovery of 32 or 21.8% is high. This is probably because the fish caught in the traps are in better condition than those caught by troll gear. Six fish were recaptured in the series of five traps at Sooke (Fig. 13). Only two had travelled westward, the remainder going to Puget Sound and the Fraser River. Almost half were recovered in United States waters.

23 In summary, the coho salmon move in many directions off the west coast of Vancouver Island and travel around both ends of the island. This is in contrast to the marked southeast migration of spring salmon.

BRITISH COLUMBIA

TAGGING AREA 0 NU MBERS 262

RETURN AREAS • NUMBERS 40+1 UNKNOWN

FIGURE 12. Distribution of recaptured coho salmon tagged off Barkley Sound, Vancouver Island, in 1950.

TAGGING AREA 0 NUMBERS 150 RETURN AREAS • NUMBERS 31 + I UNKNOWN

FIGURE Distribution of recaptured coho salmon tagged at Sooke traps, 13. . Vancouver Island, in 1951.

24 COMPARISON OF THE EARLY AND RECENT TAGGING EXPERIMENTS

SPRING SALMON

The total numbers of spring salmon tagged in outside waters during the two periods, 1925 to 1927 and 1949 to 1951, are given in Table III along with the distribution of the recoveries from the main regions of the coast.

In the Canadian experiments conducted off the west coast of Vancouver Island the percentage return of 12.9 for the early period is the same as that for the recent period despite the fact that different types of tags were used and that changes in the trolling effort have occurred. However, the distribution of the recoveries differs between the two periods. In the recent experiments only 16% of the recaptures were from the Columbia River and 29% were from United States waters, compared to 60% and 82% respectively in the earlier experiments. The recent recoveries from the Fraser River amounted to 12% of the recaptures, compared to 5% twenty years earlier; and the total recovered in British Columbia waters was 71% of all recaptures, compared to only 16% in the early experiments. These differences indicate a decrease in the size and importance of the Columbia River stock, a greater intensity of trolling near the tagging areas, and an increase in the catch of British Columbia stocks such as that of the Fraser River.

In the \Vashington State experiments conducted in this region 111 1948 and 1949 the percentage return of 17.2 is higher than that in the Canadian experi­ ments (12.9%). Moreover the percentages returned from the Columbia River (10%) and in United States waters (19%) are lower than the returns of 16% and 29% in the 1949 to 1951 Canadian experiments. The 81% recovered in Canadian waters, mostly near tagging areas, is higher than that in the 1949-51 Canadian experiments (71%). Although the 10% that were returned from the Fraser River is lower than the 12% recovered in the 1949 and 1951 Canadian experiments, it is the same percentage as was returned from the Columbia River. Consequently the Washington State experiments tend to confirm more strongly the conclusions drawn from the Canadian experiments.

In evaluating the previously mentioned changes in the fishery, it should be stressed that more small immature fish are now caught, and were tagged in the years 1949 to 1951, than in the earlier period. For example, the weights of the fish tagged off Barkley Sound differ between the two periods as follows:

25 TABLE IlL-Comparison of recoveries from tagging experiments on spring salmon for the periods 1925-30 and 1948 .. 51.

S.E. Alaska vVcst coast of Vancouver Island Queen Charlotte Islands b (outside areas)

Recovery areas 1925-27 1949-51 1948-49a 1929-30 1951 1950-51

No. % No. % No. % No. % No. % No. %

California coast .... 2 ( 5) ( 7) Oregon coast. 4 (1) 4 (3) 37 (14) 5 (5) Colnmbia River. 235 (60) 21 (16) 7 (10) 69 (26) 31 (29) Washington coast. 26 (7) 12 (9) 4 (6) 12 (5) 2 (29) 10 (10) Puget Sound ... 55 (14) (7) 2 (3) 35 (13) (14) ------.------· N 1 ..----- 1 ..··_·· .. ·- - 1 ---..· ..---1 --.. ·----- .. --- 1 ----- 1 --­ 0- U.S. (except Alaska). (82) (19) 153 (58) 3 (43) 46 (44) Alaska ... 4 (2) 15 (14) ---,------.--.-- ....------... -..------1 ---- Fraser River. 19 (5) 16 (12) 7 (10) 50 (19) (14) 13 (12) B.C. coast. 46 (11) 78 (59) 49 (71) 52 (20) 3 (43) 31 (30)

-�------I ------· I -----·- I ---- I --- - .. ...- ..-.-- - -.--- -...--- ... - --- -._------..-1------British Columbia total. 65 (16) 94 (71) 56 (81) 102 (39) 4 (57) 1 44 (42) Miscellaneous.. 7 (2) 3 (1) ======c l=�===I ==�== I====I ==== I === I ===7= 1 =====I ==== 1 =====1 === 1 ===1==== Total recoveries. 394 (100) 133 (100) 69 (100) 262 (100) 7 (100) 105 (100)

Total number tagged ...... 1 3049 1033 403 1935 64 710

Percentage recovery ...... 1 12.9 12.9 17.2 13.5 11.0 14.8

a Compiled from Pacific Marine Fisheries Commission Blllletin No. 2, 1951. b Compiled from Alaska Department of Fisheries Annual Reports No.2 and 3. 1950-5 1. vVeight in pounds

1-9 10-19 20-29 I 30-39 40-49 50-59 ------1 ------1925 No. tagged . 330 498 206 72 11 8

1926 No. tagged . 337 433 378 158 41 6

1949 No. tagged . 670 98 4 No. returned . 5:::- 12 % returned .. 8.8 12.3

1950 No. tagged .... 140 No. returned ...... 38 . . . . . : : % returned . . I 27.0

In the 1949 experiment the percentage that was returned from tagging small fish was lower than that from tagging larger fish. Based on the length measurements shown in Table A (Appendix) 21 fish or 6.1% were returned from the 344 fish that were smaller than 62.5 cm. whereas 50 fish or 11. 7% were re­ turned from the 428 fish that were tagged at a larger size. The difference in return probably results from both a higher mortality involved in catching and tagging small fish and the higher natural mortality that would occur during the longer period that the small fish remain in the ocean.

In the 1950 experiment all the fish were small but the relatively high return suggests that an intensive fishery must have exploited these immature fish throughout much of their ocean life. Unfortunately the sizes or ages of the recaptured fish are not available for the 1925 and 1926 experiments as recorded by \Villiamson and Clemens (1932). However a comparison can be made of the ages of the spring salmon tagged in 1926, whose ages are given by Mottley (1929); those tagged in 1949, reported by Neave (1951), and those tagged in 1950, given in Table C (Appendix):

Year of life Year Second I Third Fourth Fifth Sixth ---1----I ------0° 1926 3% 28% 50S-; 18% 1 /0

1949. . ./ 84% 13%

1950.. 17% 78% 5%

27 Obviously there was a great difference in the age of the fish being caught and tagged in the two periods. The difference in size and age of fish tagged probably accounts for the return of only 12.9% (Table III) in recent years when smaller and younger fish were tagged and a higher fishing intensity operated, compared to the same return from tagging larger and older fish when the fishing intensity was much lower during the early years of the outside fishery. The tagging of smaller and younger fish has also resulted in relatively more fish being recaptured in the years following the tagging year and in the vicinity of the tagging area, if it is a feeding area. An increase in fishing effort in the tagging area would also increase the number taken near the point of release and reduce the number that could be taken elsewhere at a later time. For the Barkley Sound experiments, the distribution of the returns in percentages is as follows:

Year after tagging Year

1st 2nd 3rd 4th

1925 Total return ...... 61 35 3 1 Return off Vancouver Island ...... 1 6 - - 1926 Total return ...... 77 18 4 1 Return off Vancouver Island ...... 3 1 - - 1949 Total return ...... 41 52 7 - Return off Vancouver Island ...... 24 34 1 - 1950 Total return...... 21 55 24 - Return off Vancouver Island ...... 11 26 11 -

Because of the difference in recapture near the tagging area, the returns in Table III of 16% from the Columbia River and 12% from the Fraser River are relatively low in comparison with the percentage of 60 and 5 obtained in the early experiments. Furthermore, the tagging of more immature fish implies that a higher ocean mortality is involved. This would result in a lower return from the rivers in comparison with the ocean returns, since some fish would be recaught before all the mortality had occurred. For these reasons, the present contribution from the Columbia River is higher than that indicated by the tagging results, probably about 30%, and the contribution from the Fraser River is probably about 20%. If this is true, then in the period between the two experiments the contribution of fish from the Columbia River to the troll fishery off the west coast of Vancouver Island has decreased by about half while that from the Fraser River has increased by about 4 times. In the experiments conducted off the northwest coast of the Queen Charlotte Islands the return of 11.0% in 1951 (Table III) is lower than the return of

28 13.5% in the earlier period. This may also be related to the difference in size and age of the fish tagged or to changes in the season and intensity of the fisheries. The percentage of the returns that were found in United States waters decreased from 58 to 43 between the two periods and no recoveries were made in the Columbia River itself in the recent experiment in contrast to a return of 26% of the recoveries in the early period. The percentage of the total returns that were found in British Columbia waters increased from 39 to 57 in the two periods although the portion recovered from the Fraser River decreased from 19% to 14%. Because the number of fish tagged in 1951 is so small it is of interest to note the results of some larger experiments conducted off south­ eastern Alaska by the Alaska Department of Fisheries in 1950 and 1951. Larger fish were tagged in these experiments and a larger recovery of 14.8 % was obtained. The percentages of the total returns that were made in United States waters (44) and in the Fraser River (12) are similar to those from the 1951 tagging off the Queen Charlotte Islands. \Vhen the recoveries in Alaskan waters (14%) are added to those recovered in British Columbia waters (42%) the total is similar to the 57% recovered in British Columbia waters in the recent tagging off the Queen Charlotte Islands. Apparently few of these large mature fish were recap­ tured near the tagging area with the majority being recaptured while migrating along the coast to the southeast. Since 29% were recovered in the Columbia River, this stock is probably still an important contributor to the outside troll fishery operating offnorthern British Columbia even though the small experiment in 1951 offthe Queen Charlotte Islands did not disclose it.

Parker and Kirlmess (1951) in summing up all of the southeastern Alaska experiments conducted in outside waters from 1950 to 1952 concluded that the Columbia River was probably the largest single source of salmon and that the Fraser River was the second largest. In the area north of Cape Spencer 37% of the returns were from the Columbia River and 5% were from the Fraser River, while in the area south to the Alaskan boundary 24% were from the Columbia River and 16% were from the Fraser River. Also the majority of the returns from the Columbia River belonged to the fall run, especially the races that spawn above Bonneville dam.

The spring or chinook salmon of the Columbia River are commonly classified into three groups, called spring, summer and fall runs, depending on the times when the fish migrate up the river. Since 1949 the large fall chinook stock has declined sharply to record lows in both 1954 and 1955. Because of this recent decline, it is important to know which segment of the stock is fished by the trollers off the British Columbia coast. The peak of the fishing off the Queen Charlotte Islands occurs in June and that off the west coast of Vancouver Island takes place in July. If another month is allowed for the fish to reach the Columbia River then it would appear that most of the maturing fish caught off British Columbia probably belong to the fall chinook stock. However, in recent years many immature fish have also been caught, particularly off the southwestern part of Vancouver Island. The times, when the tagged fish were

29 recovered in the Columbia River, should indicate the group to which these small fish belong. The numbers of tags recovered in the Columbia River during the three spawning migration periods are as follows :

Spring rull Summer rlln Fall run Tl)tal April-May J une-J uly Aug.-Oct. --- No. % No. % No. % No. --� --� --� ------

1925-30-

Queen Charlotte Islands ...... 3 6 10 22 33 72 46

West coast of Vancouver Island .. 7 3 52 24 157 73 216

1949-5 1- West coast of Vancouver Island ... 2 10 - - 17 90 19

Both in the past and at present, it appears that the British Columbia trollers have exploited the fall chinook run to the Columbia River more heavily than the early runs, although in the past when the troll-fishing season was earlier than at present and the Columbia River gillnet fishery was less restricted during the summer period, many summer chinook salmon were also tagged and recaught. Few spring chinook salmon were caught at any time. Thus since 1925 the fall chinook run has provided the majority of the Columbia River fish that are caught by the trollers off British Columbia and the recent decline in this stock is no doubt affecting the outside catch at the seasonal peak of the fishing. In conclusion, the theory of a northwestward feeding migration of young fish coupled with a southeastward spawning migration of adults has now been firmly established for spring salmon caught in the waters along the continental shelf off British Columbia. Fish from the larger rivers predominate in the catches made in a northwesterly direction from their mouths. Because of these long migrations, the intermixing of spring salmon stocks along the Pacific coast gives rise to regulations and problems which are international in scope and which require mutual co-operation between Canada and United States for their solution.

COHO SALMON The total numbers of coho salmon tagged and the recoveries from the iJ lain regions along the coast are summarized in Table IV for the two periods 1925 to 1930 and 1949 to 1951. The number of recoveries in the recent period is higher than that in the earlier period. In the Canadian experiments off the west coast of Vancouver Island the percentages have increased from 6.1 to 9.2 in the two periods and off the northwest coast of the Queen Charlotte Islands from 6.3 to 8.0. Since most of the coho salmon in the two periods were caught in their third year as maturing fish of the same size, the difference in percentage return

30 TABLE IV.-Comparison of recoveries from tagging experiments on coho salmon for the periods 1925-40 and 1948-51.

S.E. Alaskab Sooke traps West coast of Vancouver Island Queen Charlotte Islands (outside areas) Vancouver Island Recovery areas 1925-2 7 1949-5 1 1948-49 " 1929-30 1951 1950-5 1 1928-29 1951

No. % No. % No. % No. % No. % No. % No. % No. % ------

Oregon coast ...... 1 (1) ......

Washington coast ...... 3 (13) 14 (10) 4 (13) .. . . 1 (2) . . . . 1 (3) .. . .

Puget Sound ...... 2 (9) 25 (19) 2 (27) 1 (1) ...... 8 (24) 14 (44)

--- ._------" w U.S. (except Alaska) ...... 5 (22) 40 (30) 6 (40) 1 (1) 1 (2) .. . . 9 (27) 14 (44) ......

Alaska ...... 22 (19) 13 (27) 61 (80) ......

------

British Columbia total ...... 18 (78) 86 (65) 9 (60) 93 (80) 33 (67) 15 (20) 24 (73) 18 (56)

Miscellaneous ...... 6 (5) ...... 2 (4) ......

------

Total recoveries ...... 23 (100) 132 (100) 15 (100) 116 (100) 49 (100) 76 (100) 33 (100) 32 (100)

Number tagged-total ...... 376 . . 1426 . . 59 .. 1828 .. 614 .. 848 .. 157 .. 150 ..

Percentage recovery ...... 6.1 . . 9.2 . . 25 .4 . . 6.3 .. 8.0 . . 9.0 .. 21 .0 . . 21 .8 . .

a Compiled from Pacific Marine Fisheries Commission, Bulletin No. 2, 1951. b Compiled from Alaska Department of Fisheries Annual Reports No. 2 and 3, 1950-51. is probably an indication of the heavier exploitation in recent years. Judging from the changes in the distribution of the recoveries in the two periods it would appear that much of the increase in exploitation occured in United States waters. From taggings off the west coast of Vancouver Island, recoveries from Puget Sound increased from 9% to 19% between the two periods. Smaller experiments conducted in 1948 and 1949 in this region by the \Vashington State Department of Fisheries further confirm that in recent years the rate of recovery is higher than formerly, and that more tagged fish are recovered in United States waters, particularly in Puget Sound. The returns from fish tagged at Sooke traps suggest the same conclusion, since 44% of the recent returns were from Puget Sound, compared to only 24% in the early period. Similarly for the Queen Charlotte Islands experiments, the returns from southeastern Alaska were 27% of the total returning in 1951 compared to 19% in the 1929 to 1930 period. The tagging in the outside waters off southeastern Alaska in 1950 and 1951 showed a marked radiation from the tagging area with 20% recovered in northern British Columbia waters. As compared with spring salmon, the coho salmon migrated shorter distances in all directions from the point of tagging towards the numerous small streams along the coast. They travel more slowly and appear to wander more during both their feeding and spawning migrations. The result is that the ocean distribution of coho salmon from any one river is more restricted along the coast and probably extends farther offshore than that of spring salmon. Never­ theless, fishrecovered in the Fraser River have been tagged as far south as Oregon and as far north as Alaska. The great overlapping and mixing of many snaIl stocks makes regulation difficult but the fish caught by the trollers of British Columbia are primarily a local management problem, except for the area adjacent to either Alaska or \Vashingtoll state.

32 SUMMARY

By tagging fishcaught by commercial trollers offthe coast of British Columbia the ocean movements of adult spring and coho salmon have been demonstrated and in some cases the final destinations determined. The results from tagging 1,222 spring salmon and 2,190 coho salmon during the period from 1949 to 1952 are presented in detail. Similar experiments involving 6,457 spring salmon and 7,127 coho salmon which were conducted from 1925 to 1930 are reviewed and the results compared with those from the recent tagging operations. For spring salmon caught in the outside waters of British Columbia the majority are on long spawning migrations, travelling in a southeasterly direction along the continental shelf towards the larger rivers, such as the Columbia and Fraser. Off the southwestern part of Vancouver Island many small immature fish are also captured while they are foraging in the ocean a year or two prior to the spawning migration. In the case of coho salmon, all the fish are on shorter spawning migrations, travelling in many directions from offshore waters towards numerous small coastal streams as well as larger rivers.

A comparison between the results of the experiments conducted off the west coast of Vancouver Island about twenty years apart disclosed the following changes. For spring salmon the percentages recovered were the same despite an increase in the fishing intensity in recent years. This may be accounted for by the lower returns that result from tagging the smaller and younger fish which are being caught more heavily in recent years. Also more fish were recovered in recent years in waters off British Columbia and in the Fraser River than formerly while fewer fish were recovered in United States waters and in the Columbia River. This implies that the stocks of the Columbia River have decreased in size and importance while the stocks from British Columbia rivers are being harvested more heavily now than in former years. For coho salmon the percentage recovered was higher in the recent period than in the early one probably because of the increase in fishing intensity, particularly in United States waters.

Since the outside troll fishery operates on a mixture of stocks from many rivers and streams along the Pacific coast, local regulations are often inadequate and conservation measures should be co-ordinated throughout the whole stream and ocean range of each stock. For spring salmon taken along the Pacific coast from California to southeastern Alaska and for coho salmon taken in areas adjacent to Alaska and vVashington state, these problems become inter­ national in scope.

33 REFERENCES

ADF. 1950. Annual Report of Alaska Department of Fisheries. Report No. 2, pp. 25-35. 1951. Ibid., Report No. 3, pp. 20-27.

CALHOUN, A. ]., D. H. FRY AND E. P. HUGHES. 1951. Plastic deterioration and metal corrosion in Petersen disc fish tags. California Fish and Game, 37(3): 301-314.

CLEMENS, W. A. 1930. Pacific salmon migration : The tagging of coho salmon on the west coast of Vancouver Island in 1297 and 1928. Bioi. Bd. Canada, Bull. No. 15, 19 pp.

CLEMENS, W. A., R. E. FOERSTER AND A. L. PRITCHARD. 1939. The migration of Pacificsal mon in British Columbia waters. Amer. Assoc. Adv. Sci., Pub. No. 8, pp. 51-59.

MILNE, D. J. 1952. The coho salmon run off the northern part of the west coast of Vancouver Island in 1951. Fish. Res. Bd. Canada, Pacific Prof!.. Rept. , No. 91, pp. 28-30.

MILNE, D. J.. AND E. A. R. BALL. 1956. The mortality of small salmon when caught by trolling and tagged or released untagged. Ibid. , No. 106, pp. 10-13.

MOTTLEY, C. McC. 1929. Pacific Salmon Migration: Report on the study of the scales of the spring salmon, (0. tshawytscha) tagged in 1926 and 1927 off the west coast of Vancouver Island. Contr. Canadian Bioi. and Fish., N.S., 4(30) : 473-494.

NEAVE, FERRIS. 1951. Observations on troll-caught salmon of the west coast of Vancouver Island, 1949. Bull. Pac. Marine Fish. Comrn., No. 2, pp. 93-100.

PARKER, R. R., AND W. KIRKNESS. 1956. King salmon and the ocean troll fishery of south­ eastern Alaska. Alaska Dept. of Fish. , Research Rept. No. 1, 64 pp.

PMFC. 1951. Bulletin No. 2 of the Pacific Marine Fisheries Commission, 101 pp.

PRITCHARD, A. L. 1933. Relation of tagging programs to the conservation of Pacific salmon off the coast of British Columbia. Trans. Am. Fish. Soc. for 1932, 62 : 88-93. 1934. Pacific salmon migration: The tagging of the coho salmon in British Columbia in 1929 and 1930. Bioi. Bd. Canada, Bull. No. 40, 24 pp. 1940. Studies on the age of the coho salmon (0. kisutch) and the spring salmon (0. tshawytscha) in British Columbia. Trans. Roy. Soc. Canada, Sect. V, 34 : 99-120.

WILLIAMSON, H. C. 1927. Pacific salmon migration: Report on the tagging operations in 1925. Contr. Can. Bioi. and Fish. , N.S., 3(9) : 267-306. 1929. Pacific salmon migration: Report on the tagging operations in 1926, with addi­ tional returns from the operations of 1925. Ibid. , 4(29) : 455-470.

WILLIAMSON, H. C., AND W. A. CLEMENS. 1932. Pacific salmon migration: The tagging opera­ tions at Quatsino and Kyuquot in 1927 with additional returns from the operations of 1925 and 1926. Bioi. Bd. Canada, Bull. No. 26, 16 pp.

34 APPENDIX

Tables showing the salmon tagged and recoveries made in the experiments of 1949-52. TABLE A.-Length frequencies of spring salmon tagged and returned in the 1949 to 1952 experiments. The columns at the left are the mid-class length, measured to the fork of the tail. The United States minimum size limit of 26 inches in total length (from tip of nose to tip of tail when the latter is lying in a natural position) would include most of the fish below 62.5 cm. (24.5 in.) in fork length or about 6 Ibs. in dressed weight (gutted with head on).

Queen Charlotte Is. Quatsino Sonnd Kyuquot Sonnd Barkley Sonnd Sooke Traps

Fork length 1951 1949 1951 1950 1949 1950 1952

Tagged Returned Tagged Returned I Tagged Returned Tagged Returned Tagged Returned I Tagged Returned Tagged Returned I I I I I I I I

em. in.

30 11.8 ...... 2 35 13 .8 ...... 4 CN 0- 40 15.8 .. . . 1 .. 1 . . 4 . . 11 . . 2 45 17.7 .. . . 1 . . 2 .. 1 . . 10 . . 15 50 19.7 ...... 2 1 4 1 32 4 11 4 5 I 3 55 21 .7 2 ...... 7 3 85 4 16 8 60 23 .6 5 . . 3 2 3 1 5 .. 200 13 38 10

65 25 .6 5 . . 3 2 3 " 1 . . 180 20 58 15 15 4

70 27.6 5 .. 12 1 9 1 .. . . 146 18 . . . . 19 2

75 29 .5 9 . . 14 4 4 ...... 55 4 . . . . 24 6

80 31 .5 19 3 7 1 7 3 .. . . 29 5 .. . . 35 9

85 33 .5 13 3 4 2 10 1 . . . . 14 3 .. .. 12 2 90 35 .4 11 1 .. . . 2 ...... 4 95 37 .4 4 .. .. 7

100 39 .4 ...... 3

105 41 .3 ...... 1 ----

,. 64 7 45 12 54 8 22 4 772 71 140 38 125 28 Total ...... I I I I I I I I TABLE B.-Length frequencies of coho salmon in the 1949 to 1951 experiments. The columns at the left are mid-class lengths.

Queen Charlotte Is. Quatsino Sound Kyuquot Sound Barkley Sound Sooke Traps

--�------.-.�------.----�----'"- Fork length 19S1 1949 19.11 19.10 1949 19S0 19S1

-- -.------,------.----,, ---. Tagged Returned Tagged Returned Tagged Returned Tagged Returned Tagged Returned Tagged Returned Tagged Returned

------,------.. -- -.------

em in.

30 11.8 eN 3S 13.8 -:r 40 15.8 45 17.7 4 50 19 .7 4 3 3 11 14 4 55 21.7 11 24 1 6 20 11 53 4

60 23.6 64 8S 8 S2 2 35 101 12 21 2 65 25 .6 2S2 19 244 22 141 8 48 77 21 42 9

70 27.6 231 20 106 9 294 2.1 4 11 58 1 2 75 29 .5 46 7 46 3 17 6 80 31 .5 unrecorded 4

------._------�---- Total . ... 614 49 470 41 544 40 110 40 262 41 1.10 32 TABLE C.-Ages of spring salmon tagged and returned in the 1950 to 1952 experiments.

NOTE : Ages for spring salmon tagged in the 1926-30 experiments are given by Mottley (1929) and Pritchard (1940), and for those tagged in the 1949 experiments are given by Neave (1951).

---�

Queen Charlotte Qua tsino Sound Kyuquot Sound Barkley Sound Sooke Traps Islands 1951 1950 1950 1952 Age in years 1951 --�------Tagged Returned Tagged Returned Tagged Returned Tagged Returned Tagged Returned -----

OCEAl'I TYPE

2 ...... � . . . 1 1 15 1 21 4 13 6 eN 00 3 ...... 9 0 11 3 5 2 93 26 78 14

4 ...... 22 5 19 3 ...... 3 0 2 0

5 ...... 12 1 3 0 ......

STREAM TYPE 3 ...... 1 0 ...... 4 2 10 3 I 4...... , ...... , . . . . 3 1 9 3

NOT READABLE .. .. 21 1 19 1 2 1 16 5 13 2

-----

Total ...... 64 7 54 8 22 4 140 38 125 28 TABLE D.-Returns from the tagging of spring salmon offQuatsino and Barkley Sounds, Vancouver Island, in 1949.

Fork length Tag no. Date tagged Locali ty tagged Date caught Locality caught Period out when tagged

em. days

41665 Jul 6, 1949 Quatsino Sound ...... 84 Jul 11, 1949 Solander Island, V.I...... 5 41658 Jul 5, 1949 " " ...... 74 Jul 25, 1949 Fraser River ...... 20 " " 4f659 Jul 5, 1949 ...... 79 Aug 5, 1949 Nootka Sound, V.I ...... 31 " " 41681 Jul 23, 1949 ...... 85 Aug 11, 1949 Fraser River ...... 19 " " 41697 Jul 30, 1949 ...... 66 Aug 17, 1949 Point Roberts, Wash ...... 18 " " 41660 Jul 5, 1949 ...... 66 Aug 23, 1949 Henry Island, Wash ...... 49 " " 41689 Jul 27, 1949 ...... 74 Aug 23, 1949 Astoria, Oregon ...... 27 " " 41690 Jul 28, 1949 ...... 59 Aug 25, 1949 Cape Scott, V.I ...... 28 41701 Jul 31, 1949 " " ...... 73 Aug 29, 1949 Fraser River ...... 29 " " W 41670 Jul 7, 1949 ...... 71 Sep 7, 1949 Celilo Falls, Columbia R ...... 62 '0 " " 42050 Jul 8, 1949 ...... 61 Oct 11, 1949 Sooke Traps ...... 95 " " 41662 Jul 5, 1949 ...... 76 Aug 14, 1950 Columbia River ...... 405 40929 May 13, 1949 Barkley Sound ...... 59 May 29, 1949 Uc1uelet, V.1...... 16 " " 41095 May 21, 1949 ...... 65 Jun 1, 1949 Fraser River ...... 10 " " 40999 May 17, 1949 ...... 60 Jun 20, 1949 Uc1uelet, V.I ...... 34 " " 41434 Jun 14, 1949 ...... 79 Jun 28, 1949 Long Beach, V. I ...... 21 " " - 41518 Jun 18, 1949 ...... 84 Jul , 1949 Sooke Traps ...... " " 41157 May 28, 1949 ...... 71 Jul 14, 1949 Kyuquot, V.I ...... 47 " " - - 41401 Jun 13, 1949 ...... 74 1949 Swiftsure, V.I ...... " " 41185 May 29, 1949 ...... 81 Jul 16, 1949 Long Beach, V.1...... 48 " " 41132 May 24, 1949 ...... 71 Jul 17, 1949 Long Beach, V.l ...... 54 " " 41195 May 29, 1949 ...... 69 Jul 21, 1949 Estevan Point, V.I ...... 53 " " 41602 Jun 26, 1949 ...... 74 Jul 27, 1949 Long Beach, V.I...... 31 " " 41016 May 18, 1949 ...... 62 Aug 4, 1949 Swiftsure, V.I ...... 78 " " 40996 May 17, 1949 ...... 59 Aug 7, 1949 Esperanza, V.I ...... 81 " " 41485 Jun 18, 1949 ...... 61 Aug 14, 1949 Nootka, V.I ...... 57 " " 41521 Jun 18, 1949 ...... 69 Aug 19, 1949 Barkley Sound, V.I ...... 62 TABLE D.-Returns from the tagging of spring salmon off Quatsino and Barkley Sounds, Vancouver Island, in 1949-Concluded - -

Fork length Tag no. Date tagged Loca Ii ty tagged Date caught Locality caught Period out when tagged

---- -_ .. ------_._-----

em. days

41712 Aug 4, 1949 Barkley Sound ...... 67 Aug 24, 1949 Ucluelet, V.I ...... , . .. 20 " " 41541 Jun 22, 1949 ...... 65 Aug 26, 1949 Swiftsure Bank, V.I ..... 65 " " 41579 Jun 23, 1949 , ...... 79 Sep 1, 1949 Esperal1za, V.I ...... 70 " " 41211 May 29, 1949 60 Sep 9, 1949 Barkley Sound, V.I...... 102 " " 41196 May 29, 1949 85 Sep 9, 1949 Fraser River ...... 102 " " 41590 Jun 25, 1949 ...... 64 Sep 10, 1949 Estevan Point, V. I .... 77 " " 41011 May 18, 1949 57 Sep 11, 1949 West coast of V.I...... 116 " " 41552 JUl1 22, 1949 . . 78 Sep 11, 1949 Celilo Falls, Columbia R ...... 81 " " 41723 Aug 13, 1949 56 Sep 16, 1949 Swiftsure Bank, V.I ...... 34 " " '""'" 41147 May 26, 1949 71 Sep 20, 1949 Tillasqua River, Wash . 117 o " " 41206 May 29, 1949 78 Sep 27, 1949 Fraser River . 121 " " 41589 JUl1 25, 1949 72 Oct 10, 1949 Queens Cove, V.I . 107 " " 41555 JUl1 22, 1949 . . 84 Oct 13, 1949 Fraser River . 113 " " 41262 JUl1 3, 1949 65 Oct 20, 1949 Tatchu Point, V.I . 139 " " 41706 Aug 3, 1949 64 Apr 13, 1950 Barkley Sound, V.I ...... 253 " " 41371 JUll 12, 1949 69 May 1, 1950 Cape Beale, V.I...... 323

" " -- - 41474 JUl1 18, 1949 ..... 64 1950 \V'est coast of V.I .... " " 41655 Jul 2, 1949 74 May 18, 1950 Pt. Ano Nuevo, Cal ... 320 " " 41492 JUll 18, 1949 . .. 69 May 23, 1950 Lippy Pt., V.I . , .. 239 " " 40937 May 14, 1949 ...... 64 May 23, 1950 Ucluelet, V.I.. , ...... 343 " " 41278 Jun 3, 1949 65 May 25, 1950 Barkley Sound, V.I. 356 " " 41313 JUl1 8, 1949 65 May 31, 1950 Barkley Sound, V.I. 357 " " 40966 May 15, 1959 66 Jun 4, 1950 Estevan Light, V.I ...... 387 " " 40903 May 10, 1949 ...... 70 JUl1 6, 1950 Barkley Sound, V.I ...... 392 " " 41565 Jun 23, 1949 70 Jun 7, 1950 Lennard Is., V.I...... 349 " " 40922 May 12, 1949 ...... 70 JUIl 12, 1950 Sooke Traps ...... 393 " " 41709 Aug 4, 1949 70 lun 14, 1950 Tillamook Rock, Wash ...... 314 41440 Jun 14, 1949 " " 64 Jun 15, 1950 West coast of V. I...... 366 40976 May 15, 1949 " " 65 Jun 15, 1950 Cape Flattery ...... 394

41402 Jun 13, 1949 ...... 70 Jun 17, 1950 West coast of V. I...... 369 41726 Aug 18, 1949 " " 50 Jun 24, 1950 Barkley Sound, V.I...... 309 41225 Jun 2, 1949 " " 61 Jun 24, 1950 Barkley Sound, V. I...... 389 40942 May 14, 1949 " 69 Jun 28, 1950 Kyuquot, V. l...... 410

" ...... 40948 May 14, 1949 62 Jul 1, 1950 West coast of Queen C. Is . . 413

...... - V. I...... 41022 May 18, 1949 . .. . 67 Jul ,1950 Barkley Sound, 41580 Jun 23, 1949 " . 76 Jul 20, 1950 Hotsprings Cove, V.I...... 392 40967 May 15, 1949 61 Jul 24, 1950 T�=, �I...... 435 41348 Jun 12, 1949 71 Jul 30, 1950 Neah Bay, Wash ...... 413 41109 May 22, 1949 61 Aug 16, 1950 Hotsprings Cove, V. l...... 457 40955 May 14, 1949 " 65 Aug 7, 1950 Umatilla Reef, Wash ...... 460- 41742 Aug 26, 1949 " 51 Aug 15, 1950 Clayoquot ...... 354 4141)1 Jun 18, 1949 " 64 Aug 17, 1950 Lennard Is. , V.I ...... 425 41438 Jun 14, 1949 65 Aug 22, 1950 Columbia River ...... 434

" ...... 41097 May 21, 1949 67 Aug 26, 1950 Columbia River . . . . 458 ,j:>. " ...... 41126 May 24, 1949 ...... 68 Sep 16, 1950 Bonneville, Columbia R ...... 480 41730 Aug 20, 1949 53 Sep 19, 1950 Ucluelet, V. I...... 395 41532 Jun 19, 1949 " 64 Sep 26, 1950 Fraser River ...... 464 41346 Jun 12, 1949 " " 59 Sep 19, 1950 Lennard Is., V.I...... 450 40959 May 15, 1949 " " 70 1950 Fraser River ......

...... 40908 May 10, 1949 . . . . . 66 1950 Columbia River ......

...... V. I...... 41419 Jun 13, 1949 ...... 59 1950 Cape Beale, 41733 Aug 21, 1949 55 Jul 8, 1951 Portland Point, V.l. . . 686 41740 Aug 23, 1949 " 52 Aug 6, 1951 Columbia River ...... '1 713 " " ...... 41246 Jun 2, 1949 ...... 62 1951 Columbia River ...... " 41563 Jun 23, 1949 69 Sep 10, 1951 Columbia River ...... 809 41376 Jun 12, 1949 51 Oct 10, 1951 Columbia River ...... '1 850

NOTE : Re Neave (1951): for Fig. I on page 98 to be complete, 18 additional spring s:llmon recoveries should be added. TABLE E.-Returns from the tagging of coho salmon off Quatsino and Barkley Sounds, Vancouver Island, in 1949

Fork length Tag no. Date tagged Locality tagged Date caught Locality caught Period out when tagged

em. days

41907 May 15, 1949 Barkley Sound ...... 52 Aug 15, 1949 Johnstone Strait ...... 92

" " 41914 May 20, 1949 ...... '" . 48 Sep 5, 1949 Whidbey Is., Wash ...... 108

" " 41926 Jun 22, 1949 • < ••••••• , •••• 62 Sep 19, 1949 Point Roberts, Wash ...... 89

" " 41921 May 10, 1949 ...... 46 Nov 21, 1949 Green River Hatchery, Wash ...... 167

" " 41923 Jun 12, 1949 ...... 59 Nov 21, 1949 Alberni, V.I...... 167 41996 Jul 7, 1949 Quatsino Sound ...... 66 Jul 15, 1949 Cape Caution ...... 8

" " 42065 Jul 9, 1949 ...... 66 Jul 25, 1949 Rivers Inlet ...... 6

" " 42340 Jul 28, 1949 . " ...... 67 Aug 15, 1949 Nootka, V.I...... 18

" " 41948 Jul 6, 1949 ...... 61 Aug 16, 1949 Wakeman Sound ...... 41

" " *'" . N 42261 Jul 27, 1949 ...... 72 Aug 17, 1949 Cherry Point, Wash ...... 21 " " 41949 Jul 6, 1949 ...... 73 Aug 18, 1949 Knights Inlet ...... 43

" " 41941 Jul 5 , 1949 . . ' . . , .... 62 Aug 19, 1949 Egg Island ...... '" ...... 45

" " 42009 Jul 7, 1949 ...... 67 Aug 19, 1949 Kyuquot Sound, V.I...... 44

" " 42021 Jul 7, 1949 . , . , ...... 64 Aug 25, 1949 Kains Island, V.I...... 49

" " 42031 Jul 7, 1949 ...... 65 Aug 29, 1949 Rivers Inlet ...... 53

" " " 41956 Jul 6, 1949 ...... , ...... 66 Aug 30, 1949 ...... 55

" " 42424 Jul 30, 1949 . . . . , 62 Aug 30, 1949 Swiftsure, V. I...... 31

" " 41974 Jul 6, 1949 ...... 59 Sep 2, 1949 Quatsino Sound, V.I...... 58

" " 42194 Jul 23, 1949 ...... , . 59 Sep 6, 1949 Fraser River ...... 45

" " 42282 J ul 27, 1949 ...... 72 Sep 6, 1949 Lennard Is., V.I...... 41 42312 Jul 28, 1949 " " 70 Sep 12, 1949 Bella Bella ...... 46

" " 42394 Jul 29, 1949 ... . . 69 Sep 13, 1949 Fitzhugh Sound ...... 46

" " 42150 Jul 21, 1949 ...... 61 Sep 13, 1949 Ucluelet, V.I...... , ...... 54

" " 42227 Jul 25, 1949 ... . 61 Sep 14, 1949 Johnstone Strait ...... 51

" " 42342 Jul 28, 1949 " . 65 Sep 15, 1949 Port San Juan, V.I...... 40

" " 42432 Jul 31, 1949 ... .., ... .. 66 Sep 18, 1949 Tacoma, Wash. 49 " " V. I...... 42361 Jul 29, 1949 ...... 66 Sep 20, 1949 Swiftsure, 54 42206 Jul 24, 1949 " " 71 Scp 21, 1949 Nicomcld River ...... 59 41994 Jul 7, 1949 " " 66 Sep 21, 1949 Fraser River...... 76 " 42030 Jul 7, 1949 ...... 69 Sep 23, 1949 ...... 74 " " 41959 Jul 6, 1949 ...... 66 Sep 28, 1949 Skagit River, Wash ...... 78 " " 42328 Jul 28, 1949 ...... 66 Oct 3, 1949 ...... 67 " " 42167 Jul 21, 1949 ...... " . . .. . 66 Oct 4, 1949 English Bay ...... 75 " " 42413 Jul 30, 1949 ...... 66 Oct 4, 1949 YeIIow Rock ...... 66 " " 42209 Jul 24, 1949 ...... 66 Oct 11, 1949 Ramsay Arm, ...... 79

41663 Jul 5, 1949 ...... 64 Oct 12, 1949 Fraser River ...... 99 " " 42131 Jul 21, 1949 ...... 56 Oct 14, 1949 Cape Scott, V.I...... 85 42428 Jul 30, 1949 " " 65 Oct 17, 1949 Fraser River...... 79 42145 Jul 21, 1949 " " 66 Oct 24, 1949 Chehalis River, Wash ...... 96

42028 Jul 7, 1949 62 Oct 26, 1949 ...... 111 42010 Jul 7, 1949 66 Oct 30, 1949 White River, Wash ...... 115 42232 Jul 25, 1949 66 Oct 31, 1949 Fraser River...... 98 42038 Jul 7, 1949 " 69 Nov 15, 1949 Skokomish River, Wash...... 131 " " 42267 Jul 27, 1949 ...... 70 Nov 12, 1949 Soos Creek, Wash ...... 98 ..,. " " " w 42396 Jul 29, 1949 ...... 69 Nov 12, 1949 ...... 98 " 42220 Jul 24, 1949 . . , ...... 65 Nov 12, 1949 Johnstone Strait ...... 98

NOTE :-Neave (1951). For Fig. 2 on page 101 to be complete, 6 additional coho salmon recoveries should be added. TABLE F.-Returns from the tagging of spring salmon on the north and west coast of the Queen Charlotte Islands in 1951.

Fork length Period Type of gear Tag No. Date tagged Locality tagged Age Date caught Locality caught when tagged out caught by

em. days

"'" "'" G 3517 Jun 12, 1951 Shag Rock, (North) ...... 78 4, J ul 16, 1951 Cape Cook, V.I...... 34 Troll G3647 J ul 10, 1951 Shag Rock, (North) ...... 86 4, Sep 8, 1951 Grays Harbour, Wash ...... 60 Troll G 3996 Aug 16, 1951 Rose Spit, (North) ...... 85 5, Sep 22, 1951 Umatilla Reef, Wash ...... 37 Troll

G 3525 J un 16, 1951 LaPerouse Rks., (West) .. 87 4, Aug 20, 1951 Port Alberni, V.I...... 65 Troll G 3530 Jun 21, 1951 LaPerouse Rks., (West) ... 81 4, Aug 20, 1951 Pt. Francis, Wash ...... 60 Gill-net G 3553 J un 26, 1951 LaPerouse Rks., (West) .... 78 - Aug 14, 1951 Fraser River ...... 49 Gill-net G 3752 Jul 15, 1951 LaPerouse Rks., (West) .... 88 4, Jul 26, 1951 Solander Is., V.I...... 11 Troll TABLE G.-Returns from the tagging of spring salmon off Quatsino Sound, Vancouver Island, in 1951

I Fork length Period Type of gear Tag No. 1 Date tagged Locali ty tagged Age Date caught Locality caught when tagged out caught by

em. days

G 2001 May 10, 1951 Lippy Pt. , V.I...... 81 41 Jul 17, 1951 Esperanza, V.I...... 68 Troll G 2009 May 28, 1951 Lippy Pt., V.I...... 97 41 Jun 22, 1951 Fraser River .·...... 25 Gill-net

G 2035 Jul 7, 1951 Lippy Pt., V.I...... 81 3, Jul 17, 1951 Quatsino Sound ...... 10 Troll

G 2053 Jul 26, 1951 Kains Island ...... 83 3, Oct 1, 1951 Willapa Bay, Wash ...... 69 Gill-net

- G 2012 Jun 7, 1951 Cape Cook, V.I...... 71 Jul 12, 1952 Swiftsure, V.I...... 400 Troll G 2028 Jul 2, 1951 Kains Island ...... 62 3, Oct 1, 1957. St. Helens, Co.! R...... 455 Gill-net G 2032 Jul 4, 1951 Kains Island ...... 80 4, May 3, 1952 Cape Cook, V.I...... 304 Troll

G 2018 Jun 25, 1951 Kains Island ...... 52 2, Aug 4, 1953 Columbia River ...... 770 Gill-net

*'" --_.. _------U1

TABLE H.-Returns from the tagging of spring salmon offKyu quot Sound, Vancouver Island, in 1950.

Fork length Period Type of gear Tag No. Date tagged Locality tagged Age Date caught Locality caught when tagged out caught by

em. days

4362 1 Jun 2. 1950 Kyuquot Sound ...... 53 - Sep 8, 1950 Barkley Sound ...... 98 Gill-net

43502 Jun 5, 1950 Refuge Cove ...... 57 31 Sep 25, 1950 Fraser River ...... 112 Gill-net 43509 Jul 7, 1950 Kyuquot Sound . 53 21 May 10, 1951 Barkley Sound ...... 308 Troll 43651 May 25, 1950 Kyuquot Sound . 52 3, May 6, 1952 Swiftsure, V.I...... 711 Troll

------TABLE I.-Returns from the tagging of spring salmon off Barkley Sound, Vancouver Island, in 1950

...

Fork length Period Type of gear Tag No. Date tagged Locality tagged Age Date caught Locality caught when tagged out caught by

------�� ------�------

em. days

42903 Barkley Sound ...... 55 31 Sep 2, 1950 Lennard Is., V.I...... 94 Troll

" " 42904 ...... 55 3. Feb 10, 1951 Fraser River ...... Spawned in Alouette R.

42918 Long Beach ...... 64 31 Aug 25, 1950 Hotsprings Cove, V.I...... 70 Troll

42967 Florencia Bay ...... 60 31 Sep 25, 1950 Fraser River ...... 101 Gill-net

" " 42980 ...... 63 31 Aug 9, 1950 Ucluelet, V.I ...... 51 Troll

" " 42982 ...... 63 3 1 Jul 18, 1950 Kyuquot, V.I...... 29 Troll 42987 Amphitrite Pt ...... 66 42 Jul 11, 1950 Fraser River...... 1 3 Gill-net

" " 43012 ...... 56 31 Sep 14, 1950 Willapa Bay., Wash ...... 33 Gill-net

" " "'" 42901 ...... 63 31 Sep 10, 1951 Columbia River ...... 493 Gill-net

0- " 42924 " 67 - Jun 2, 1951 Ucluelet, V.I...... 359 Troll

42929 Barkley Sound ... . . " .. 64 31 Jun 27, 1951 Off Oregon ...... 382 Troll

42936 Amphitrite Pt...... 59 31 Jun 24, 1951 Kyuquot, V.I ...... 3 77 Troll

" " 42943 . ..., 64 31 May 1, 1951 Astoria, Oregon ...... 320 Gill-net

" " 42944 ...... 59 31 Sep 2, 1951 Swiftsure, V.I...... 445 P. seine

" " - 42954 , ...... 58 31 - 1951 Columbia River ...... Gill-net 42958 Florencia Bay ...... 59 32 May 17, 1951 Willapa Bay, Wash ...... 335 Troll

" " 42959 ... . • ••<. 64 31 Scp 20, 1951 Columbia River ...... 46 1 Dip net

" " 42961 ...... 59 3 1 Sep 1, 1951 Hot Spring Cove, V.I ...... 441 Troll

" " 42971 ... 64 31 Oct. 8, 1951 Fraser River ...... 445 Gill-net

" " 42975 ...... 66 31 Sep 12, 1951 Fraser River ...... 450 Gill-net

" " 42979 ...... 63 31 Jul 8, 1951 Long Beach, V.l...... 384 Troll

42986 Amphitrite Pt ...... 60 31 Aug 25, 1951 Off Esperanza, V.I...... 420 Troll

" " 42988 . . ... 49 31 Apr 14, 1951 Cape Beale, V.l...... 290 Troll " " - . 42992 ...... 56 May 21, 1951 Ucluelet, V.I...... 329 Troll

" " 42997 . . , . . ... SO 21 May 16, 1951 Lennard Is., V.I...... 3 14 Troll I " " 42998 J .. "'. ,., . 59 31 Aug 3, 1951 Suislaw R., Oregon ...... 395 Troll " " 43023 Aug 29, 1950 . .. , ...... 51 21 Jun 21, 1951 Tillamook Head, Ore ...... 296 Troll 43029 Sep 3, 1950 Barkley Sound ...... 50 21 Aug 17, 1951 West coast, V.I...... 348 - " " 43038 Sep 5, 1950 ...... 53 21 Oct 25, 1951 Wind R., Wash ...... 415 Seining " 42930 Jun 10, 1950 ...... 56 31 May 3, 1952 Columbia River ...... 692 Gill-net 42931 JUI1 12, 1950 Amphitrite Pt .. .. 65 31 Aug 13, 1952 Ucluelet, V.I...... 792 Troll 42960 Jun 16, 1950 Florencia Bay .... 65 - May - 1952 Columbia River ...... - Gill-net 42973 Juu 19, 1950 Amphitrite Pt ...... 60 31 Sep 7, 1952 Whidbey Is., Wash ...... 810 P. seine 43003 Jul 29, 1950 Long Beach ...... 60 31 Jul 8, 1952 Lennard Is., V.I...... 709 Troll 43006 Aug 10, 1950 Amphitrite Pt ..... 56 31 Sep 23, 1952 Columbia River ...... 774 Gill-net " " - 43024 Aug 30, 1950 ...... 46 Jul 20, 1952 Cape Scott, V.I...... 689 Troll " " 43031 Sep 4, 1950 ...... 56 - Aug 4, 1952 Makah Bay, Wash...... 700 Troll " " 43032 Sep 5, 1950 ...... 64 31 Jul 14, 1952 Barkley Sd., V.I...... 677 Troll

------

,j:>. _1 TABLE J.-Returns from the tagging of spring salmon at Sooke traps, Vancouver Island, in 1952 - --_... _------_.. ------_... __ ...... _-

Fork length Period Type of gear Tag No. Date tagged Locali ty tagged Age Date caught Locality caught when tagged out caught by

em. days G 4906 Jul 7, 1952 Sooke ...... 78 32 Aug 29, 1952 Westport, Wash ...... 52 Troll

" G4908 Jul 7, 1952 ...... 58 21 Jul 17, 1952 Fraser River ...... 10 Gill-net

" " " - G 4912 Jul 7, 1952 ...... 73 32 Aug - 1952 ...... Gill-net

" " G 49 17 Jul 7, 1952 Beechy Head ...... 67 32 Jul 11, 1952 ...... 5 Gill-net

" " G 4928 Jul 7, 1952 Gordon Head ...... 43 21 Jul 18, 1952 ...... 11 Gill-net

" " G4930 Jul 7, 1952 ...... 68 31 Jul 24, 1952 Pt. Roberts, Wash ...... 17 Gill-net

" " G4939 Jul 14, 1952 ...... 74 42 Jul 18, 1952 Salmon Banks, Wash ...... 4 Reef-net

" " G 4945 Jul 14, 1952 ...... 46 21 Sep 1, 1952 Fraser River ...... 49 Gill-net G 4956 Jul 14, 1952 Sooke ...... 50 31 Jul 24, 1952 Skagit R., Wash ...... 10 (spawned)

" G4964 Aug 5, 1952 ...... 74 31 Aug 12, 1952 Fraser River ...... 7 Gill-net " - ,po. G 4972 Aug 5, 1952 ...... 58 Aug 18, 1952 Lummi Is., Wash ...... 3 Reef-net (Xl G 4973 Aug 5, 1952 Gordon Head ...... 70 31 Aug 11, 1952 Fraser River ...... 6 Gill-net

" " G 4974 Aug 5, 1952 ...... 50 31 Aug 18, 1952 Fraser River ...... 13 Gill-net

G4980 Aug 5, 1952 Sooke ...... 68 31 Aug 7, 1952 Sooke trap, V.I...... 2 Traps

" " " " G4986 Aug 5, 1952 ...... 72 42 Aug 7, 1952 ...... 2

" " " " G 4988 Aug 5, 1952 ...... 66 42 Aug 7, 1952 ...... 2

" G 4993 Aug 11, 1952 ...... 77 31 Aug 12, 1952 Iceberg Pt., Wash ...... 1 Reef-net G 4875 Aug 11, 1952 Coal Creek ...... 61 21 - 1952 Fraser River...... - Gill-net

" " G 4878 Aug 11, 1952 ...... 76 31 Sep 19. 1952 Columbia River ...... 39 (spawned)

" " G4879 Aug 11, 1952 ...... 68 21 Aug 18, 1952 Fraser River ...... 7 Gill-net

" " G 4880 Aug 11, 1952 ...... 78 - Aug 20, 1952 Fraser River ...... 9 Gill-net

" " G 4885 Aug 11, 1952 ...... 73 31 Aug 14, 1952 Sooke trap ...... 3 Trap G4889 Aug 11, 1952 Gordon Head ...... 68 31 Aug 19, 1952 Fraser River ...... 8 Gill-net

" " G 4890 Aug 11, 1952 ...... 74 31 Aug 18, 1952 Fraser River ...... 7 Gill-net

" " G4892 Aug 11, 1952 ...... 73 31 Aug 26, 1952 Point Roberts, Wash ...... 15 P. seine

" " G 4893 Aug 11, 1952 ...... 77 31 Aug 18, 1952 Fraser River ...... 7 Gill-net

" " - . - G4894 Aug 11, 1952 ...... 58 31 1952 Fraser River ...... Gill-net G4898 Aug 11, 1952 Beechy Head ...... 43 21 Aug 21, 1952 Fraser River ...... 10 Gill-net TABLE K-Returns from the tagging of coho salmon on the north and west coast of the Queen Charlotte Islands in 1951

Fork length Period Type of gear Tag No. Date tagged Locality tagged Date caught Locality caught when tagged out caught by

NORTH COAST em. days

G 3515 Jun 12, 1951 Shag Rock ...... " " " " " . " . . . 51 Jul 11, 1951 Cape Naden ...... 29 Troll " G 3598 Jul 5 1951 ...... 64 Aug 22, 1951 Banks Island ...... 47 Troll " G 3615 Jul 5, 1951 ...... 67 Aug 20, 1951 Duke Island, Alaska ...... 46 Trap " G 3616 Jul 5, 1951 ...... " . 64 Aug 20, 1951 Seaforth Channel ...... 46 Seine " G 3618 Jul 5, 1951 ...... 67 Aug 15, 1951 Ketchikan, Alaska ...... 41 - " G 3620 Jul 5, 1951 ...... 67 - 1951 - - - " G 3660 Jul 10, 1951 ...... , 63 Aug 18, 1951 Wales Island ...... 39 - G 3883 Aug 12, 1951 Tow Hill ...... 67 Aug 31, 1951 Cape Fox, Alaska ...... 19 Trap G 3890 Aug 14, 1951 Tow Hill ...... 65 Aug 24, 1951 Foggy Point, Alaska ...... 10 Trap ,.p. G 3965 Aug 16, 1951 Rose Spit ...... \0 71 Sep 17, 1951 Namu ...... " ... 32 Troll " G 3967 Aug 16, 1951 ...... 70 Sep 20, 1951 N. Bentinck Arm ...... 35 Gill-net " G 3970 Aug 16, 1951 ...... 69 - 1951 Digby Island ...... - Troll " G 3978 Aug 16, 1951 ...... 75 Sep 8, 1951 Work Channel ...... 23 Troll " G 3992 Aug 16, 1951 ...... 71 - 1951 N. Bentinck Arm ...... - - " G 4001 Aug 16, 1951 ...... 72 Sep 8, 1951 Dundas Island ...... 23 Gill-net G 4022 Aug 17, 1951 N. of Masset Inlet ...... 66 Aug 30, 1951 Annette Island, Alaska ...... 13 Trap G 4027 Aug 17, 1951 N. of Masset Inlet ...... 66 Aug -,1951 San Cristoval Ch., Alaska ...... - Trap

G 4030 " Aug 19, 1951 Masset Inlet ...... 68 Oct 4, 1951 Masset Inlet ...... 46 Gill-net G 4052 Aug 19, 1951 Cape Edensaw ...... 69 Oct 15, 1951 Namu ...... 57 Gill-net " G 4101 Aug 20, 1951 ...... 59 Sep 16, 1951 Fraser River ...... 27 Gill-net " G 4104 Aug 20, 1951 ...... 71 Sep 18, 1951 Langara Island ...... 29 Troll " G 4113 Aug 20, 1951 ...... 71 Aug 27, 1951 Sitklan Island, Alaska ...... 7 Trap " G 4121 Aug 20, 1951 ...... 64 Sep 5, 1951 Port Simpson ...... 18 Gill-net " G 4145 Aug 20, 1951 . . " ...... 69 Sep -,1951 Duke Island, Alaska ...... - Troll " G 4146 Aug 20, 1951 ...... 68 Sep 6, 1951 Principe Channel...... 17 Gill-net " G 4156 Aug 21, 1951 ...... 74 Sep 14, 1951 Fitzhugh Sound ...... 25 Gill-net " G 4204 Aug 26, 1951 ...... 57 Dec 15, 1951 Porcher Island ...... 111 - TABLE K.-Returns from the tagging of coho salmon on the north and west coast of the Queen Charlotte Islands in 1951-Concluded

Fork length Period Type of gear Tag No. Date tagged Locality tagged Date caught Locality caught when tagged out caught by

WEST COAST em. days

G 3543 Jun 26, 1951 LaPerouse Rocks ...... 62 Sep 5, 1951 Johnstone Strait ...... 71 Troll " G 3555 Jun 26, 1951 · ...... 63 Sep 3, 1951 Toba Inlet ...... 69 Troll " G 3571 Jun 27, 1951 ...... 66 Jul 26, 1951 Whale Channel ...... 29 Troll " G 3573 Jun 27, 1951 · ...... 63 ) Aug 10, 1951 Skeena River ...... f.l,4 Gill-net G 3580 Jul 3, 1951 Langara Rock ...... 70 Jul 14, 1951 Dundas Island ...... 11 Troll G 3628 Jul 6, 1951 LaPerouse Rocks ...... 63 Aug 24, 1951 Smith Inlet ...... 47 Gill-net

" - - G 3629 Jul 6. 1951 · ...... 69 - - G 3670 Jul 13, 1951 Cape Knox ...... 74 Aug 21, 1951 Rivers Inlet ...... 39 Gill-net " G 3691 Jul 13, 1951 ...... 69 Jul 24, 1951 Whales Island ...... 1 Gill-net " V1 G 3692 Jul 13, 1951 ...... 60 Sep 27, 1951 Johnstone Strait ...... 45 - o " - G 3717 Jul 14, 1951 ...... 67 Aug 21, 1951 Hakai Pass ...... 38 " G 3718 Jul 14, 1951 ...... 68 Sep 14, 1951 Namu ...... 61 Gill-net G 3739 Jul 15, 1951 LaPerouse Rocks ...... 69 Aug --, 1951 Prince of Wales Island, Alaska .. - Trap " G 3740 Jul 15, 1951 · ...... 61 Aug 18, 1951 George Inlet, Alaska ...... 33 - " G 3757 Jul 15, 1951 ...... 70 Aug 2, 1951 Union Bay, Nass ...... 18 Troll " G 3767 Jul 16, 1951 ...... 64 Aug 4, 1951 Kanagunut Island, Alaska ...... 19 Trap " G 3776 Jul 18, 1951 ...... 69 Sep 17, 1951 Fraser River ...... 61 Gill G 3783 Jul 18, 1951 Cape Knox ...... 63 Oct 18, 1951 Queets, Wash ...... 92 Gill-net G 3789 Jul 20, 1951 Langara Light ...... 62 Aug 26, 1951 Betton Island, Alaska ...... 37 Troll G 3806 Jul 28, 1931 LaPerouse Rocks ...... 65 Aug 31, 1951 Annette Island, Alaska ...... 34 Trap " G 3824 Jul 31, 1951 ...... 73 Sep 37, 1591 Lockport, Queen Charlotte Is.. . 58 - G 3849 Aug 5, 1951 Andrews Point, North Island .. 69 Aug 12, 1951 Squamish River ...... 7 Angler

- TABLE L.-Returns from the tagging of coho salmon off Quatsino Sound and Cape Scott, Vancouver Island, in 1951

Fork length Period Type of gear Tag No. Date tagged Locali ty tagged Date caught Locality caught when tagged out caught by

QUATSINO SOUND em. days

G 2724 Lippy Point ...... 64 64 -

G 2727 Cape Palmerston ...... 68 30 Gill-net

- G 2730 Kains Island ...... 66 59

" G 2741 ...... 65 72 Troll

" - G 2778 • ,0" • . ... . 72 67

" - G 2783 ...... 71 Angler

" G 2796 ...... 70 79 Angler G 2814 " ...... 67 34 P. seine

" - U1 G 2842 ...... 71 53 ...... " G 2843 ...... 68 108 Indian F.

" - G 2847 ...... 66 53

" - G 2850 ...... , . . . . 68 25

" G 2852 ...... 66 35 Troll

" G 2854 ...... 67 60 Gill-net

" - G 2860 ...... 74 31 G 2912 Sea Otter Cove ...... 71 58 -

G 2931 Kains Island ...... , . 68 48 Trap

" G 2933 ....,. , . . . 69 66 Gill-net

" G 2954 ...... 62 10 Troll

G 2970 Cape Cook ...... 73 47 P. seine

" G 2990 .... 70 177 Hatchery

" - G 2974 ...... , ..... 63 Gill-net

" G 3011 ...... 68 39 Angler

G 3209 Kains I slancl ...... 71 64 Gill-net

" G 3216 ...... , . . . . . 68 28 P. seine

" G 3225 g ...... 69 p q 24 Angler TABLE L.-Returns from the tagging of coho salmon off Quatsino Sound and Cape Scott, Vancouver Island, in 1951-Concluded

----

Fork length Period Type of gear Tag No. Date tagged Locality tagged Date caught Locality caught when tagged out caught by

QUATSINO SOUND em. days

G 3243 Aug 18, 1951 Kains Island ...... 73 Jan 25, 1952 Green River, Wash ...... 160 Hatchery

" - G 3247 Aug 22, 1951 ...... 68 Sep 12, 1951 Port San Juan, V.1...... 22

CAPE SCOTT

G 3051 Aug 6. 1951 Steamer grounds ...... 54 Oct 1, 1951 Nehalem River, Oregon ...... 56 Gill-net

" - G 3064 Aug 6, 1951 ...... 62 Sep 19, 1951 Johnstone Strait...... 44

" - G 3071 Aug 7, 1951 ...... , ... Sep 6, 1951 Amphitrite Pt...... 30 Troll U1 " N G 3098 Aug 7, 1951 ...... 68 Sep 27, 1951 Texada Island ...... 51 Gill-net " - G 3106 Aug 7, 1951 ...... 68 - 1951 Umatilla Reef, Wash ...... - " - G 3116 Aug 7, 1951 •••••••••• 0 68 Aug 20, 1951 Ramsey Arm ...... - " G 3134 Aug 8, 1951 ...... 69 Nov 2, 1951 Port Angeles, Wash ...... 87 - " G 3150 Aug 8, 1951 ...... 70 Oct 16, 1951 Port McNeil ...... 70 Gill-net " G 3153 Aug 8, 1951 0 ••••••••••• 69 Oct 3, 1951 Cracraft Island ...... 56 Gill-net " G 3164 Aug 8, 1951 ••••••••••o. 71 Sep 25, 1951 Queen Charlotte Sd ...... 48 - " G 3171 Aug 9. 1951 ...... 71 Nov 14, 1951 Quinault River, Wash ...... 97 Indian F. " G 3179 Aug 9, 1951 ...... 69 Aug 20, 1951 Fitzhugh Sound ...... 11 - TABLE M.-Returns from the tagging of coho salmon off Kyuquot Sound, Vancouver Island, in 1950

Fork length Period Type of gear Tag No. Date tagged Locality tagged Date caught Locality canght when tagged out caught by

em. days

43528 Jul 5, 1950 Kyuqnot, V.I...... 52 Oct 30, 1950 Johnstone Strait ...... 117 P. seine 43601 Jul 7, 1950 Refuge Cove ...... 48 Oct 3, 1950 Fraser Rivero ...... 118 Gill-net 43560 Aug 4, 1950 Solander Island ...... 62 Sep 25, 1950 San Juan Island, Wash ...... 52 P. seine " 43565 Aug 4, 1950 ...... 58 Sep 29, 1950 Otter Point, V.I...... 56 Trap " 43570 Aug 4, 1950 ...... 61 Sep 1, 1950 Nootka ...... 28 Troll

TABLE N.-Returns from the tagging of coho salmon off Barkley Sound, Vancouver Island, in 1950

Fork length Period Type of gear (n Tag No. Date tagged Locality tagged Date caught Locality caught CN when tagged out caught by

em. days 43202 Jun 1, 1950 Barkley Sound ...... 50 Sep 24, 1950 Tatoosh, Wash ...... 116 Troll " 43204 Jun 1, 1950 •••••• 0'•• ... 51 Oct 6, 1950 Fraser River ...... 127 Gill-net 43208 Jun 3, 1950 Long Beach ...... 46 Nov 2, 1950 Salmon Banks, Wash ...... 122 P. seine

" - 43210 Jun 3, 1950 ...... , .. 49 Oct 2, 1950 Puget Sound, Wash. 121 " 43212 Jun 3, 1950 0 ••• 0 •• ·.' ••••• • 52 Oct 2, 1950 Sooke ...... 121 Traps 43225 Jun 3, 1950 Amphitrite Point ...... 64 Aug 7, 1950 Kyuquot, V.I...... 56 Troll " 43230 Jun 8, 1950 ...... 48 Oct 1, 1950 Russian R., V.I...... 113 - 43232 Jun 10, 1950 Barkley Sound ...... 52 Aug 1, 1950 } West coast, V.l...... 52 Troll 43251 Jun 16, 1950 Florellcia Bay ...... 60 Aug 12, 1950 Ucluelet ...... 57 Troll

" - - 43252 JUll 16, 1950 ••• 0 ••••0.' •••. 64 - 1950 Grays Harbour, Wash ...... " 43261 JUll 19, 1950 ...... 59 Aug 19, 1950 Swiftsure, V.I...... 92 Troll 43270 Jun 30, 1950 Amphitrite Point...... 64 Aug 21, 1950 Sooke ...... 52 Traps 43281 Jul 29, 1950 Long Beach ...... 60 Oct 2, 1950 Puget Sound, Wash ...... 65 Gill-net 43282 Aug 7, 1950 Portland Point ...... 64 Aug 21, 1950 Sooke ...... 14 P. seine TABLE N.-Returns from the tagging of coho salmon off Barkley Sound, Vancouver Island, in 1950- Concluded

------_.. _------_.. _------� --� -

Fork length Period Type of gear Tag No. Date tagged Locali ty tagged Date caught Locality caught when tagged out caught by

em. days

43289 Aug 7, 1950 Amphitrite Pt ..... , . . , .., , . , 67 Aug 31, 1950 Sooke . , . , ...... , .., .., .... 24 Traps

" - 43295 Aug 7, 1950 ...... 60 1950 Wash ...... - -

" 43303 Aug 7, 1950 ...... 63 Oct . 9, 1950 Puget Sound, Wash ...... 63 Troll

" 43309 Aug 7, 1950 ...... 63 Nov 10, 1950 Lummi Island, Wash ...., ...... 78 P. seine

" 43323 Aug 7, 1950 ...... 61 Aug 25, 1950 Sooke Bay, V.I ...... 18 P. seine

" 43328 Aug 8, 1950 ...... 66 Oct 23, 1950 Johnstone Strait ...... 76 P. seine

" 43349 Aug 8, 1950 ...... 64 Oct. 8, 1950 Chehalis River, Wash ...... 61 Gill-net

" 43350 Aug 8, 1950 ...... 58 Sep 27, 1950 Fraser River ...... , .., , .... 50 Gill-net

" 43355 Aug 8, 1950 ...... 63 Sep 20, 1950 Uc1uelet ...... , .., ...... 15 Troll

" 43368 Aug 12, 1950 ...... 64 Aug 27, 1950 Nooksack River, Wash ...... 15 Gill-net

" c.n 43371 Aug 12, 1950 ...... 60 Aug 25, 1950 Sherringham Light, V.l...... 13 P. seine "'" " 43372 Aug 12, 1950 ...... 58 Oct. 2, 1950 Puget Sound, Wash ... , .. , , .. , , 51 -

" 43374 Aug 12, 1950 ...... 63 Aug 25, 1950 Boundary Bay, Wash .. , , . , , .., 13 P. seine

" 43378 Aug 12, 1950 ...... 64 Oct -, 1950 - - -

" 43384 Aug 19, 1950 ...... 60 Aug 24, 1950 West coast, V.I...... , . .. 5 Troll

" 43386 Aug 19, 1950 ...... 64 Oct 15, 1950 Squamish ...... , .. , 57 Gill-net

" 43396 Aug 26, 1950 ...... 59 Nov 1, 1950 Jervis Inlet ..., ...... , . , .. , 67 Gill-net

" 43405 Aug 29, 1950 ...... 65 - 1950 Washington ...... , - Troll

" 43416 Aug 31, 1950 ...... 59 Sep 20, 1950 Salmon Banks, Wash ...... , 20 P. seine

" 43419 Aug 31, 1950 ...... 64 Oct 10, 1950 Egmont ...... , . . , . 40 Gill-net

" 43424 Sep 3, 1950 ...... 66 Oct 21, 1950 Johnstone Strait ...... , .. , .... , 48 P. seine

" 43425 Sep 3, 1950 ...... 56 Sep 21, 1950 Johnstone Strait ...... 18 P. seine

" 43427 Sep 3, 1950 ...... 63 Oct 8, 1950 Grays Harbour, Wash ..... , .... 35 Gill-net

" 43428 Sep 3, 1950 ...... 63 Sep 27, 1950 Rolling Bay, Wash ...... , ...... 24 Gill-net

" - 43446 Sep 4, 1950 ...... 64 - 1950 Puget Sound, Wash ...... , .. Gill-net

" 43454 Sep 5, 1950 ...... 64 Oct 11, 1950 Fraser River ...... 36 Gill-net

" 43462 Sep 5, 1950 ...... 58 Oct 5, 1950 Snohomish R., Wash ...... , .. 30 Gill-net

------�----.------.----.-.-�--- � TABLE O.-Returns from the tagging of coho salmon at Sooke traps, Vancouver Island, in 1951

Fork length Period Type of gear Tag No. Date tagged Locality tagged Date caught Locality caught when tagged out caught by

em. days

G 5011 Beachy Head ..., ... 64 Henry Island, Wash ...... 4 Reef-net

" G 5017 ...... , . .. 76 Coal Creek ...... 7 Trap

G 5021 Coal Creek . ' .. , . . 64 Otter Point ...... ' ...... 11 Trap G 5023 68 San Juan Is., ·Wash...... 1 G 5028 " 68 Port Gardiner, Wash ...... 9 Gill-net G 5030 62 Satellite Channel ...... Gill-net G 5035 Sooke Harbour . 69 Swinonish, Wash ...... 6 Angler

" G 5041 78 Anacortes, Wash ...... , ...... 7

G 5051 Otter Point ... 70 Port San Juan, V.I...... 3 P. seine U1 " G 5052 64 . U1 Salmon Banks, Wash ..... , ... . " G 5058 . . . , ..... 75 Gordon Head . , .., ...... 4 Trap G 5065 Coal Creek . , 65 Lnmmi Is., Wash ...... 6 Reef-net G 5066 " 65 Fraser River ...... 13 Gill-net

G 5068 ...... 70 Fraser River ...... 6 Gill-net

G 5075 Sooke Harbour ...... 72 Anacortes, Wash ...... 7

G 5077 Gordon Head . , , , . . , . , 73 Johns Island, Wash ...... 6 Reef-net G 5086 " Nitinat Lake ...... 29 P. seine G 5097 68 Sooke Harbour ... 3 Trap

G 5101 Coal Creek ...... , ... .., . 64 Skokomish R., Wash .... , .. 77 dead in river

" G 5109 ...... , . ... 78 Henry Island, Wash ...... 2 P. seine

G 5118 Sooke Harbour ..., ..... , 77 Sooke ...... 4 Trap

G 5123 Otter Point ...... , , .. 69 Fraser River ...... , . , ...... , . 4 Gill-net

" G 5124 ...... 71 Everett, Wash, ... , ...... 61 Angler

G 5126 Gordon Head , .., . .. 68 Salmon Bank, Wash ...... 4 P. seine

" G 5127 ...... 65 Sooke .... , . , , ...... 12 P. seine

" G 5128 73 Sooke ...... , ..... 12 Angler

" G 5129 p 73 p Orcas Island, Wash. , ..... , .... 3 Angler TABLE O.-Returns from the tagging of coho salmon at Sooke traps, Vancouver Island, in 1951-Coneluded

Fork length Period Type of gear Tag No. Date tagged Locality tagged Date caught Locality caught when tagged out caught by

em. days

G 5130 Sep 20, 1951 Gordon Head ...... 63 Oct 4, 1951 Saltspring Island ...... 14 Gill-net " G 5136 Sep 20, 1951 ...... 62 Sep 20, 1951 Swiftsure ...... 1 P. seine

" - - G 5140 Sep 20, 1951 ...... 69 - 1951 - G 5141 Sep 20, 1951 Coal Creek ...... 69 Sep 20, 1951 Albert Head ...... 3 Angler " G 5143 Sep 20, 1951 ...... 66 Sep 24, 1951 Fraser River ...... 4 Gill-net