Potential Fossil Yield Classification (PFYC)
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Revised Correlation of Silurian Provincial Series of North America with Global and Regional Chronostratigraphic Units 13 and D Ccarb Chemostratigraphy
Revised correlation of Silurian Provincial Series of North America with global and regional chronostratigraphic units 13 and d Ccarb chemostratigraphy BRADLEY D. CRAMER, CARLTON E. BRETT, MICHAEL J. MELCHIN, PEEP MA¨ NNIK, MARK A. KLEFF- NER, PATRICK I. MCLAUGHLIN, DAVID K. LOYDELL, AXEL MUNNECKE, LENNART JEPPSSON, CARLO CORRADINI, FRANK R. BRUNTON AND MATTHEW R. SALTZMAN Cramer, B.D., Brett, C.E., Melchin, M.J., Ma¨nnik, P., Kleffner, M.A., McLaughlin, P.I., Loydell, D.K., Munnecke, A., Jeppsson, L., Corradini, C., Brunton, F.R. & Saltzman, M.R. 2011: Revised correlation of Silurian Provincial Series of North America with global 13 and regional chronostratigraphic units and d Ccarb chemostratigraphy. Lethaia,Vol.44, pp. 185–202. Recent revisions to the biostratigraphic and chronostratigraphic assignment of strata from the type area of the Niagaran Provincial Series (a regional chronostratigraphic unit) have demonstrated the need to revise the chronostratigraphic correlation of the Silurian System of North America. Recently, the working group to restudy the base of the Wen- lock Series has developed an extremely high-resolution global chronostratigraphy for the Telychian and Sheinwoodian stages by integrating graptolite and conodont biostratigra- 13 phy with carbonate carbon isotope (d Ccarb) chemostratigraphy. This improved global chronostratigraphy has required such significant chronostratigraphic revisions to the North American succession that much of the Silurian System in North America is cur- rently in a state of flux and needs further refinement. This report serves as an update of the progress on recalibrating the global chronostratigraphic correlation of North Ameri- can Provincial Series and Stage boundaries in their type area. -
Strophomenide and Orthotetide Silurian Brachiopods from the Baltic Region, with Particular Reference to Lithuanian Boreholes
Strophomenide and orthotetide Silurian brachiopods from the Baltic region, with particular reference to Lithuanian boreholes PETRAS MUSTEIKIS and L. ROBIN M. COCKS Musteikis, P. and Cocks, L.R.M. 2004. Strophomenide and orthotetide Silurian brachiopods from the Baltic region, with particular reference to Lithuanian boreholes. Acta Palaeontologica Polonica 49 (3): 455–482. Epeiric seas covered the east and west parts of the old craton of Baltica in the Silurian and brachiopods formed a major part of the benthic macrofauna throughout Silurian times (Llandovery to Pridoli). The orders Strophomenida and Orthotetida are conspicuous components of the brachiopod fauna, and thus the genera and species of the superfamilies Plec− tambonitoidea, Strophomenoidea, and Chilidiopsoidea, which occur in the Silurian of Baltica are reviewed and reidentified in turn, and their individual distributions are assessed within the numerous boreholes of the East Baltic, particularly Lithua− nia, and attributed to benthic assemblages. The commonest plectambonitoids are Eoplectodonta(Eoplectodonta)(6spe− cies), Leangella (2 species), and Jonesea (2 species); rarer forms include Aegiria and Eoplectodonta (Ygerodiscus), for which the new species E. (Y.) bella is erected from the Lithuanian Wenlock. Eight strophomenoid families occur; the rare Leptaenoideidae only in Gotland (Leptaenoidea, Liljevallia). Strophomenidae are represented by Katastrophomena (4 spe− cies), and Pentlandina (2 species); Bellimurina (Cyphomenoidea) is only from Oslo and Gotland. Rafinesquinidae include widespread Leptaena (at least 11 species) and Lepidoleptaena (2 species) with Scamnomena and Crassitestella known only from Gotland and Oslo. In the Amphistrophiidae Amphistrophia is widespread, and Eoamphistrophia, Eocymostrophia, and Mesodouvillina are rare. In the Leptostrophiidae Mesoleptostrophia, Brachyprion,andProtomegastrophia are com− mon, but Eomegastrophia, Eostropheodonta, Erinostrophia,andPalaeoleptostrophia are only recorded from the west in the Baltica Silurian. -
Early Silurian Δ13corg Excursions in the Foreland Basin of Baltica, Both
13 1 Early Silurian δ Corg excursions in the foreland basin of Baltica, both familiar 2 and surprising 3 4 Emma U. Hammarlund1,2*, David K. Loydell3, Arne T. Nielsen4, and Niels H. Schovsbo5 5 6 Affiliations: 7 1Translational Cancer Research, Laboratory Medicine, Lund University, Medicon Village 8 404:C3, Scheelevägen 2, 223 63 Lund, Sweden. 9 2Nordic Center for Earth Evolution, University of Southern Denmark, Campusvej 55, 5230 10 Odense M, Denmark. 11 3School of Earth and Environmental Sciences, University of Portsmouth, Burnaby Road, 12 Portsmouth PO1 3QL, United Kingdom. 13 4Department of Geosciences and Natural Resource Management, University of Copenhagen, 14 Øster Voldgade 10, 1350 København K, Denmark. 15 5Geological Survey of Denmark and Greenland, Øster Voldgade 10, DK-1350 Copenhagen K, 16 Denmark. 17 18 *Corresponding author e-mail: [email protected] 19 20 1 21 Abstract 22 The Sommerodde-1 core from Bornholm, Denmark, provides a nearly continuous 23 sedimentary archive from the Upper Ordovician through to the Wenlock Series (lower Silurian), 24 as constrained by graptolite biostratigraphy. The cored mudstones represent a deep marine 25 depositional setting in the foreland basin fringing Baltica and we present high-resolution data 13 26 on the isotopic composition of the section’s organic carbon (δ Corg). This chemostratigraphical 27 record is correlated with previously recognized δ13C excursions in the Upper Ordovician–lower 28 Silurian, including the Hirnantian positive isotope carbon excursion (HICE), the early Aeronian 29 positive carbon isotope excursion (EACIE), and the early Sheinwoodian positive carbon isotope 30 excursion (ESCIE). A new positive excursion of high magnitude (~4‰) is discovered in the 31 Telychian Oktavites spiralis Biozone (lower Silurian) and we name it the Sommerodde Carbon 13 32 Isotope Excursion (SOCIE). -
First Record of the Early Sheinwoodian Carbon Isotope Excursion (ESCIE) from the Barrandian Area of Northwestern Peri-Gondwana
Estonian Journal of Earth Sciences, 2015, 64, 1, 42–46 doi: 10.3176/earth.2015.08 First record of the early Sheinwoodian carbon isotope excursion (ESCIE) from the Barrandian area of northwestern peri-Gondwana Jiri Frýdaa,b, Oliver Lehnertc–e and Michael Joachimskic a Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha 6 – Suchdol, 165 21, Czech Republic b Czech Geological Survey, Klárov 3/131, 118 21 Prague 1, Czech Republic; [email protected] c GeoZentrum Nordbayern, Universität Erlangen, Schlossgarten 5, D-91054 Erlangen, Germany; [email protected], [email protected] d Department of Geology, Lund University, Sölvegatan 12, 223 62 Lund, Sweden; [email protected] e Institute of Geology at Tallinn University of Technology, Ehitajate tee 5, 19086 Tallinn, Estonia Received 15 July 2014, accepted 21 October 2014 Abstract. The δ13C record from an early Sheinwoodian limestone unit in the Prague Basin suggests its deposition during the time of the early Sheinwoodian carbon isotope excursion (ESCIE). The geochemical data set represents the first evidence for the ESCIE in the Prague Basin which was located in high latitudes on the northwestern peri-Gondwana shelf during early Silurian times. Key words: Silurian, early Sheinwoodian, carbon isotope excursion, northern peri-Gondwana, Barrandian area, Prague Basin, Czech Republic. INTRODUCTION represent, beside coeval diamictites in western peri- Gondwana, an evidence of an early Silurian glaciation. The early Sheinwoodian carbon isotope excursion The expression of this glacial in the Prague Basin has (ESCIE) is known from several palaeocontinents in the already been discussed with respect to the deposition Silurian tropics and subtropics (Lehnert et al. -
Miocene Paleontology and Stratigraphy of the Suwannee River Basin of North Florida and South Georgia
MIOCENE PALEONTOLOGY AND STRATIGRAPHY OF THE SUWANNEE RIVER BASIN OF NORTH FLORIDA AND SOUTH GEORGIA SOUTHEASTERN GEOLOGICAL SOCIETY Guidebook Number 30 October 7, 1989 MIOCENE PALEONTOLOGY AND STRATIGRAPHY OF THE SUWANNEE RIVER BASIN OF NORTH FLORIDA AND SOUTH GEORGIA Compiled and edit e d by GARY S . MORGAN GUIDEBOOK NUMBER 30 A Guidebook for the Annual Field Trip of the Southeastern Geological Society October 7, 1989 Published by the Southeastern Geological Society P. 0 . Box 1634 Tallahassee, Florida 32303 TABLE OF CONTENTS Map of field trip area ...... ... ................................... 1 Road log . ....................................... ..... ..... ... .... 2 Preface . .................. ....................................... 4 The lithostratigraphy of the sediments exposed along the Suwannee River in the vicinity of White Springs by Thomas M. scott ........................................... 6 Fossil invertebrates from the banks of the Suwannee River at White Springs, Florida by Roger W. Portell ...... ......................... ......... 14 Miocene vertebrate faunas from the Suwannee River Basin of North Florida and South Georgia by Gary s. Morgan .................................. ........ 2 6 Fossil sirenians from the Suwannee River, Florida and Georgia by Daryl P. Damning . .................................... .... 54 1 HAMIL TON CO. MAP OF FIELD TRIP AREA 2 ROAD LOG Total Mileage from Reference Points Mileage Last Point 0.0 0.0 Begin at Holiday Inn, Lake City, intersection of I-75 and US 90. 7.3 7.3 Pass under I-10. 12 . 6 5.3 Turn right (east) on SR 136. 15.8 3 . 2 SR 136 Bridge over Suwannee River. 16.0 0.2 Turn left (west) on us 41. 19 . 5 3 . 5 Turn right (northeast) on CR 137. 23.1 3.6 On right-main office of Occidental Chemical Corporation. -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2014/02 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age Erathem / Era System / Period GSSP GSSP age (Ma) GSSP GSSA EonothemErathem / Eon System / Era / Period EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon GSSP age (Ma) present ~ 145.0 358.9 ± 0.4 ~ 541.0 ±1.0 Holocene Ediacaran 0.0117 Tithonian Upper 152.1 ±0.9 Famennian ~ 635 0.126 Upper Kimmeridgian Neo- Cryogenian Middle 157.3 ±1.0 Upper proterozoic Pleistocene 0.781 372.2 ±1.6 850 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian 1000 Callovian 166.1 ±1.2 Quaternary Gelasian 2.58 382.7 ±1.6 Stenian Bathonian 168.3 ±1.3 Piacenzian Middle Bajocian Givetian 1200 Pliocene 3.600 170.3 ±1.4 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.62 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Langhian Sinemurian Proterozoic Neogene 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- Hettangian 2050 Burdigalian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 28.1 Gorstian Oligocene Upper Norian 427.4 ±0.5 2500 Rupelian Wenlock Homerian -
Paleogeographic Maps Earth History
History of the Earth Age AGE Eon Era Period Period Epoch Stage Paleogeographic Maps Earth History (Ma) Era (Ma) Holocene Neogene Quaternary* Pleistocene Calabrian/Gelasian Piacenzian 2.6 Cenozoic Pliocene Zanclean Paleogene Messinian 5.3 L Tortonian 100 Cretaceous Serravallian Miocene M Langhian E Burdigalian Jurassic Neogene Aquitanian 200 23 L Chattian Triassic Oligocene E Rupelian Permian 34 Early Neogene 300 L Priabonian Bartonian Carboniferous Cenozoic M Eocene Lutetian 400 Phanerozoic Devonian E Ypresian Silurian Paleogene L Thanetian 56 PaleozoicOrdovician Mesozoic Paleocene M Selandian 500 E Danian Cambrian 66 Maastrichtian Ediacaran 600 Campanian Late Santonian 700 Coniacian Turonian Cenomanian Late Cretaceous 100 800 Cryogenian Albian 900 Neoproterozoic Tonian Cretaceous Aptian Early 1000 Barremian Hauterivian Valanginian 1100 Stenian Berriasian 146 Tithonian Early Cretaceous 1200 Late Kimmeridgian Oxfordian 161 Callovian Mesozoic 1300 Ectasian Bathonian Middle Bajocian Aalenian 176 1400 Toarcian Jurassic Mesoproterozoic Early Pliensbachian 1500 Sinemurian Hettangian Calymmian 200 Rhaetian 1600 Proterozoic Norian Late 1700 Statherian Carnian 228 1800 Ladinian Late Triassic Triassic Middle Anisian 1900 245 Olenekian Orosirian Early Induan Changhsingian 251 2000 Lopingian Wuchiapingian 260 Capitanian Guadalupian Wordian/Roadian 2100 271 Kungurian Paleoproterozoic Rhyacian Artinskian 2200 Permian Cisuralian Sakmarian Middle Permian 2300 Asselian 299 Late Gzhelian Kasimovian 2400 Siderian Middle Moscovian Penn- sylvanian Early Bashkirian -
2009 Geologic Time Scale Cenozoic Mesozoic Paleozoic Precambrian Magnetic Magnetic Bdy
2009 GEOLOGIC TIME SCALE CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST. HIST. ANOM. ANOM. (Ma) CHRON. CHRO HOLOCENE 65.5 1 C1 QUATER- 0.01 30 C30 542 CALABRIAN MAASTRICHTIAN NARY PLEISTOCENE 1.8 31 C31 251 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 70.6 254 2A PIACENZIAN 32 C32 L 630 C2A 3.6 WUCHIAPINGIAN PLIOCENE 260 260 3 ZANCLEAN 33 CAMPANIAN CAPITANIAN 5 C3 5.3 266 750 NEOPRO- CRYOGENIAN 80 C33 M WORDIAN MESSINIAN LATE 268 TEROZOIC 3A C3A 83.5 ROADIAN 7.2 SANTONIAN 271 85.8 KUNGURIAN 850 4 276 C4 CONIACIAN 280 4A 89.3 ARTINSKIAN TONIAN C4A L TORTONIAN 90 284 TURONIAN PERMIAN 10 5 93.5 E 1000 1000 C5 SAKMARIAN 11.6 CENOMANIAN 297 99.6 ASSELIAN STENIAN SERRAVALLIAN 34 C34 299.0 5A 100 300 GZELIAN C5A 13.8 M KASIMOVIAN 304 1200 PENNSYL- 306 1250 15 5B LANGHIAN ALBIAN MOSCOVIAN MESOPRO- C5B VANIAN 312 ECTASIAN 5C 16.0 110 BASHKIRIAN TEROZOIC C5C 112 5D C5D MIOCENE 320 318 1400 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 326 6 C6 APTIAN 20 120 1500 CALYMMIAN E 20.4 6A C6A EARLY MISSIS- M0r 125 VISEAN 1600 6B C6B AQUITANIAN M1 340 SIPPIAN M3 BARREMIAN C6C 23.0 345 6C CRETACEOUS 130 M5 130 STATHERIAN CARBONIFEROUS TOURNAISIAN 7 C7 HAUTERIVIAN 1750 25 7A M10 C7A 136 359 8 C8 L CHATTIAN M12 VALANGINIAN 360 L 1800 140 M14 140 9 C9 M16 FAMENNIAN BERRIASIAN M18 PROTEROZOIC OROSIRIAN 10 C10 28.4 145.5 M20 2000 30 11 C11 TITHONIAN 374 PALEOPRO- 150 M22 2050 12 E RUPELIAN -
Integrated Biostratigraphy of the Lower Silurian of the Aizpute-41 Core, Latvia
Geol. Mag. 140 (2), 2003, pp. 205–229. c 2003 Cambridge University Press 205 DOI: 10.1017/S0016756802007264 Printed in the United Kingdom Integrated biostratigraphy of the lower Silurian of the Aizpute-41 core, Latvia D. K. LOYDELL*, P. MANNIK¨ † &V.NESTOR† *School of Earth and Environmental Sciences, University of Portsmouth, Burnaby Road, Portsmouth PO1 3QL, UK †Institute of Geology, Tallinn Technical University, 7 Estonia Avenue, 10143 Tallinn, Estonia (Received 20 November 2001; accepted 28 November 2002) Abstract –Integrated graptolite, conodont and chitinozoan biostratigraphical data is presented from the Rhuddanian through to lower Sheinwoodian of the Aizpute-41 core, Latvia. Correlation of the biozonation schemes based upon the three groups is achieved from the cyphus through to lowermost riccartonensis graptolite biozones, except for the upper Aeronian and lower Telychian, which lack both chitinozoans and graptolites, and upper lapworthi through to approximately base murchisoni graptolite Biozone, where there is interpreted to be an unconformity. Datum 2 of the Ireviken Event is correlated with a level at the base of or within the murchisoni Biozone. It is possible that the changes in conodont assemblages at Datum 2 on Gotland are the result of an unconformity here. Streptograptus? kaljoi sp. nov., from the lower spiralis graptolite Biozone, is described. Keywords: graptolites, Chitinozoa, Conodonta, Silurian, biostratigraphy. 1. Introduction setting is further off-shore than that of Ohesaare (Fig. 1; see Loydell, Kaljo & Mannik,¨ 1998, for details of the Recognition of the importance of integrated biostrati- Ohesaare core), and the succession within the core graphical studies in the precise cross-facies correlation is significantly more complete, most notably in the required to test models of biotic and facies change in Aeronian. -
Alphabetical List
LIST E - GEOLOGIC AGE (STRATIGRAPHIC) TERMS - ALPHABETICAL LIST Age Unit Broader Term Age Unit Broader Term Aalenian Middle Jurassic Brunhes Chron upper Quaternary Acadian Cambrian Bull Lake Glaciation upper Quaternary Acheulian Paleolithic Bunter Lower Triassic Adelaidean Proterozoic Burdigalian lower Miocene Aeronian Llandovery Calabrian lower Pleistocene Aftonian lower Pleistocene Callovian Middle Jurassic Akchagylian upper Pliocene Calymmian Mesoproterozoic Albian Lower Cretaceous Cambrian Paleozoic Aldanian Lower Cambrian Campanian Upper Cretaceous Alexandrian Lower Silurian Capitanian Guadalupian Algonkian Proterozoic Caradocian Upper Ordovician Allerod upper Weichselian Carboniferous Paleozoic Altonian lower Miocene Carixian Lower Jurassic Ancylus Lake lower Holocene Carnian Upper Triassic Anglian Quaternary Carpentarian Paleoproterozoic Anisian Middle Triassic Castlecliffian Pleistocene Aphebian Paleoproterozoic Cayugan Upper Silurian Aptian Lower Cretaceous Cenomanian Upper Cretaceous Aquitanian lower Miocene *Cenozoic Aragonian Miocene Central Polish Glaciation Pleistocene Archean Precambrian Chadronian upper Eocene Arenigian Lower Ordovician Chalcolithic Cenozoic Argovian Upper Jurassic Champlainian Middle Ordovician Arikareean Tertiary Changhsingian Lopingian Ariyalur Stage Upper Cretaceous Chattian upper Oligocene Artinskian Cisuralian Chazyan Middle Ordovician Asbian Lower Carboniferous Chesterian Upper Mississippian Ashgillian Upper Ordovician Cimmerian Pliocene Asselian Cisuralian Cincinnatian Upper Ordovician Astian upper -
The Palaeontology Newsletter
The Palaeontology Newsletter Contents 90 Editorial 2 Association Business 3 Association Meetings 11 News 14 From our correspondents Legends of Rock: Marie Stopes 22 Behind the scenes at the Museum 25 Kinds of Blue 29 R: Statistical tests Part 3 36 Rock Fossils 45 Adopt-A-Fossil 48 Ethics in Palaeontology 52 FossilBlitz 54 The Iguanodon Restaurant 56 Future meetings of other bodies 59 Meeting Reports 64 Obituary: David M. Raup 79 Grant and Bursary Reports 81 Book Reviews 103 Careering off course! 111 Palaeontology vol 58 parts 5 & 6 113–115 Papers in Palaeontology vol 1 parts 3 & 4 116 Virtual Palaeontology issues 4 & 5 117–118 Annual Meeting supplement >120 Reminder: The deadline for copy for Issue no. 91 is 8th February 2016. On the Web: <http://www.palass.org/> ISSN: 0954-9900 Newsletter 90 2 Editorial I watched the press conference for the publication on the new hominin, Homo naledi, with rising incredulity. The pomp and ceremony! The emotion! I wondered why all of these people were so invested just because it was a new fossil species of something related to us in the very recent past. What about all of the other new fossil species that are discovered every day? I can’t imagine an international media frenzy, led by deans and vice chancellors amidst a backdrop of flags and flashbulbs, over a new species of ammonite. Most other fossil discoveries and publications of taxonomy are not met with such fanfare. The Annual Meeting is a time for sharing these discoveries, many of which will not bring the scientists involved international fame, but will advance our science and push the boundaries of our knowledge and understanding. -
Type of the Paper (Article
life Article Dynamics of Silurian Plants as Response to Climate Changes Josef Pšeniˇcka 1,* , Jiˇrí Bek 2, Jiˇrí Frýda 3,4, Viktor Žárský 2,5,6, Monika Uhlíˇrová 1,7 and Petr Štorch 2 1 Centre of Palaeobiodiversity, West Bohemian Museum in Pilsen, Kopeckého sady 2, 301 00 Plzeˇn,Czech Republic; [email protected] 2 Laboratory of Palaeobiology and Palaeoecology, Geological Institute of the Academy of Sciences of the Czech Republic, Rozvojová 269, 165 00 Prague 6, Czech Republic; [email protected] (J.B.); [email protected] (V.Ž.); [email protected] (P.Š.) 3 Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 165 21 Praha 6, Czech Republic; [email protected] 4 Czech Geological Survey, Klárov 3/131, 118 21 Prague 1, Czech Republic 5 Department of Experimental Plant Biology, Faculty of Science, Charles University, Viniˇcná 5, 128 43 Prague 2, Czech Republic 6 Institute of Experimental Botany of the Czech Academy of Sciences, v. v. i., Rozvojová 263, 165 00 Prague 6, Czech Republic 7 Institute of Geology and Palaeontology, Faculty of Science, Charles University, Albertov 6, 128 43 Prague 2, Czech Republic * Correspondence: [email protected]; Tel.: +420-733-133-042 Abstract: The most ancient macroscopic plants fossils are Early Silurian cooksonioid sporophytes from the volcanic islands of the peri-Gondwanan palaeoregion (the Barrandian area, Prague Basin, Czech Republic). However, available palynological, phylogenetic and geological evidence indicates that the history of plant terrestrialization is much longer and it is recently accepted that land floras, producing different types of spores, already were established in the Ordovician Period.