Great Basin Naturalist

Volume 46 Number 2 Article 12

4-30-1986

New taxa and combinations in the Utah flora

Stanley L. Welsh Brigham Young University

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Recommended Citation Welsh, Stanley L. (1986) "New taxa and combinations in the Utah flora," Great Basin Naturalist: Vol. 46 : No. 2 , Article 12. Available at: https://scholarsarchive.byu.edu/gbn/vol46/iss2/12

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. NEW TAXA AND COMBINATIONS IN THE UTAH FLORA

Stanley L. Welsh'

Abstract— New taxa include; Cnjptantha cinerea (Ton.) Cronq. var. arenicola Higgins & Welsh; Physaria chambersii Rollins var. sobolifera Welsh (Cruciferae); Phacelia demissa Gray var. minor N. D. Atwood (Hydrophyl laceae); Iris pariensis Welsh (Iridaceae); Astragalus preitssii var. ctitleri Barneby and Pediomelum aromaticum (Payson) Welsh var. tuhiji Welsh (Leguminosae); Abronia nana Wats. var. harrisii Welsh (Nyctaginaceae); Camissonia atwoodii Cronq. (Onagraceae); Habenaria zothecina Higgins & Welsh (Orchidaceae); Aqiiilegia formosa Fisch. in DC. var. fostcri Welsh (Ranunculaceae). New nomenclatural combinations include: Rhus aromatica Ait. var. simplicifolia (Creene) Cronq. (Anacardiaceae); Lomatium kingii (Wats.) Cronq., L. kingii var. alpinum (Wats.) Cronq. (Apiaceae); Cnjpiantha cinerea (Torr.) Cronq. var. laxa (Macbr.) Higgins; Mertcnsia lanceolata (Pursh) DC. var. nivalis (Wats.) Higgins (Boraginaceae); Opuntia erinacea Engelm. var. aurea (Baxter) Welsh (Cactaceae); Arenaria fendleri Gray var. aculeata (Wats.) Welsh, A. fendleri var. eastivoodiae (Rydb.) Welsh, Lychnis apetala L. var. kingii (Wats.) Welsh, Stellaria longipes Goldie var. monantha (Hulten) Welsh (Caryophyllaceae); Draha densifolia Nutt. ex T. & G. var, apiculata (C. L. Hitchc.) Welsh, D. oligospcrma Hook. vay. juniperina (Dorn) Welsli, Physaria acutifolia Rydb. var, stylosa (Rollins) Welsh, Thelypodiopsis sagittata (Nutt.) Schulz var. ovalifolia (R\db.) Welsh (Cruciferae); Lotus plebeius (T. Brandg.) Barneby, Lupinus polyphyllus Lindl. in Edwards var. ammophilus (Greene) Barneby, L

polyphyllus VSLT . humicola {A. Nels.) Barneby, L. argenteus Pursh var . fulvomaculatus (Payson) Barneby, L. argenteus var. palmeri (Wats.) Barneby, Pediomelum aromaticum (Payson) Welsh, P. epipsilum (Barneby) Welsh, Psoralidium lanceolatum (Pursh) Rydb. var. stenophyllum (Rydb.) Welsh, and P. lanceolatum var. stenostachys (Rydb.) Welsh (Leguminosae); Mirabilis linearis (Pursh) Hiemerl var. decipiens (Standi.) Welsh (Nyctaginaceae); Camissonia boothii var. condensata (Munz) Cronq., C. boothii var. villosa (Wats.) Cronq., C. clavaeformis (Torr. & Frem.) Raven var. purpurascens (Wats.) Cronq., C. scapoidea (T. & G.) var. utahensis (Raven) Welsh, Oenothera caespitosa var.

macroglottis (Rydb.) Cronq., Oe . caespitosa var. navajoensis (Wagner, Stockhouse, & Klein) Cronq., Oe. flava (A.

Nels.) Garrett var. acutissima (W. L. Wagner) Welsh, and Oe . primiveris Gray var. bufonis (Jones) Cronq. (Ona- graceae); Papaver radicatum Rottb. var. pygmaeum (Rydb.) Welsh (Papaveraceae); Dodecatheon pulchellum (Raf.) Merr. var. zionense (Eastw.) Welsh (Primulaceae); Aquilegia flavescens Wats. var. rubicunda (Tidestr.) Welsh,

Delphinium andersonii Gray var. scaposum (Greene) Welsh, D . occidentalis (Wats.) Wats. var. barbeyi (Huth) Welsh, and Ranunculus andersonii Gray var. juniperinus (Jones) Welsh (Ranunculaceae); Purshia mexicana (D. Don) Welsh andP. mexicana var. stansburyi (ToTr.)'We\sh (Ro.saceae); Galium mexicanum H.B.K. var. asperrimum (Gray) Higgins

& Welsh (Rubiaceae); Castilleja parvula Rydb. var. revcalii (N. Holmgren) N. D. Atwood and C . rhexifolio Rydb. var. sulphurea (Rydb.) N. D. Atwood (Scrophulariaceae).

Nomenclatural changes are a bookkeeping cited as either "in" or "ex" Welsh. Each cita chore that precedes the publication of most tion is considered as a separate proposal by floras. The following changes of names and the author of the constituent taxon or combi- newly proposed ta.xa are published here prior nation. to the publication of the Utah Flora, a work that has been in progress for some 25 years. Anacardiaceae The names published here are the products of Rhus aromatica Ait. var. simplicifolia the authors of the proposed ta.xa and nomen- (Greene) Cronq. comb. nov. [based on Rhus clatural combinations. Dr. N. Duane Atwood, canadensis var. simplicifolia Greene Bull. Dr. Larry C. Higgins, Dr. Arthur Cronquist, Torrey Bot Club 17: 13. 1890 (Greene sn.. and Dr. Rupert C. Barneby have kindly con- Peach Springs, Arizona, in 1889, at sented to having names and ta.xa currently in HDG!,may be presumed to he the holotype)]. ongoing manuscripts published here so as to

facilitate their use in the Utah Flora, which is Apiaceae nearing completion. The names are to be cited under the authorship of the person Lomatium kingii (Wats.) Cronq. comb.

providing the combination, and are not to be no\ . [based on: Peucedanum graveolens

'Life Science Miis and Department of Hii Bn«hani VounK I'm-

254 .

April 1986 WELSH: New PlantTaxa AND Combinations 255 J,

Wats, in King, Rep. Geol. Explor. 40th Paral- Caryophyllac.'Eae lel 5: 128. 1871; not Benth. & Hook. 1867. Arenaria fendleri Gray var. aculeata Peiwedoniim kingii Wats. Proc. Amer. Acad. (Wats.) Welsh stat. nov. [based on: Arenaria Arts 22: 474. 1887 (Watson 463, Wahsatch and aculeata Wats, in King, Rep. U. S. Geol. Uinta Mts.; holotype at US!; isotype at NY. !)]. Explor. 40th Parallel 5: 40. 1871]. Lomatium kingii (Wats.) Cronq. var. Arenaria fendleri Gray var. eastwoodiae alpinum (Wats.) Cronq. comb. nov. [based (Rydb.) Welsh comb nov. [based on: Arenaria on: Peucedanum graveolens var. alpinum eastwoodiae Rydb. Bull. Torrey Bot. Club 31: Wats, in King, Rep. Geol. Explor. 40th Paral- 406. 1904, and the autonym var. eastwoodiae lel 5: 129. 1871. (Watson 464, East Humboldt

established by A . eastwoodiae var. adeno- Mts., Nev.; isotype at NY!)]. phora Kearnev & Peebles J. Washington Acad. 29:475. 1939]. BORAGINACEAE Lychnis apetala L. var. kingii (Wats.) Cryptantha cinerea (Greene) Cronq. var. Welsh comb. nov. [based on: Lychnis kingii arenicola Higgins & Welsh var. nov. Ab Wats. Proc. Amer. Acad. 12: 247. 1877]. Cryptantha cinerea var. abortiva et al. in Stellaria longipes Goldie var. monantha caule simplici et inflorescentia capitato vel (Hulten) Welsh comb nov. [based on: Stel- subcapitato difFert. laria monantha Hulten Bot. Notiser 1943: Type: USA: Utah. Kane Co. T41S, R5W, 265. 1943, and the autonym var. monantha S26, Johnson Canyon, Lone Pine Point, at established by Stellaria monantha var. alto- 1830 m, in Navajo blow-sand, in a pinyon-ju- caulis Hulten Bot. Notiser 1943: 267. 1943]. niper, ponderosa pine, manzanita, and yucca community, 6 June 1984, L. C. Higgins, S. L. Cruceferae Welsh, & K. Thorne 14296 (Holotype BRY; 2 Draba densifolia Nutt. ex T. G. var. isotypes previously distributed as C & (C. L. nov. cinerea). apiculata Hitchc.) Welsh comb. Additional representative specimens: [based on: Draba apiculata C. L. Hitchc. Univ. Biol. 11: 72. Utah. Kane County, ca 22 or 23 mi S of Alton, Washington Publ. 1941, and the autonym var. apiculata established by 27 May 1965, A. Cronquist 10191; ibid.. Coral Pink Sand Dunes, 15 May 1984, N. D. At- Draba apiculata xar. daviesiae (C. L. Hitchc. Univ. Washington Publ. Biol. 17 489. wood 9636; ibid., 5 May 1966, S. L. Welsh (2): 5299. Washington Co., Pocket Mesa, 8 June 1964]. oligosperma Hook. var. juniperina 1983, L. C. Higgins and A. H. Barnum 13615 Draba (Dorn) Welsh stat. nov. [based on: Draba (all BRY). juniperina Dorn Madrono 25: 101. 1978]. Much material of this taxon passed previ- Physaria acutifolia Rydb. var. stylosa ously under the epithet multicaulis , a name (Rollins) Welsh stat. nov. [based on: Physaria that is synomymous with var. jamesii. The stijlosa Rollins, Contr. Gray Herb. 214: 14. name provided above allows recognition of 1984]. this distinctive phase of the species. Cryptantha cinerea (Greene) Cronq. var. Physaria chambersii Rollins var. sobolifera var. nov. Physaria hixa (Macbr.) Higgins comb. nov. [based on: Welsh A chambersii Rollins stricto in caudicibus ramifican- Oreocari/a multicaulis var. laxa Macbr. sensu tibus evolutis et foliis angustioribus et acutis Contr. Gray Herb. 48: 35. 1916]. differt. Mertensia lanceolata (Pursh) DC. var. ni- Garfield Co., Escalante valis (Wats.) Higgins comb. nov. based on Type: USA: Utah. of Sweetwater Creek, above Mertensia paniculata var. nivalis Wats. Rep. Mtns., head (east) ofWidtsoe, bristlecone pine community Geol. Explor. 40th Parallel 5: 239. 1871]. on Wasatch Limestone Formation, 2 Aug. 1981, N. Atwood 8164 (Holotype BRY; Cactaceae D. Isotype, one distributed previously as P. Opuntia erinacea Engelm. var. aurea chambersii). (Baxter) Welsh comb. nov. [based on: Opun- Additional specimens: Garfield Co., tia aurea Baxter Calif. Cactus 27, 28, 1933]. T34S, RIW, S22, center of Sl/4, Dixie NF, 256 Great Basin Naturalist Vol. 46, No. 2

Escalante Mtns., head of Escalante Canyon, Iridaceae 27.5 km 288 degrees from Escalante, 2,898 m, Iris pariensis Welsh sp. nov. Planta simiUs 60 degree slope, S exposure, with scattered I ride missoiiriensis sensu latissimo sed in foliis Douglas-fir on raw, white, marl limestone, 20 angustioribus (2.5-5 mm nee 3-12 mm) June 1981, S. Goodrich 15666 (BRY); ibid., bracteis suboppositis et angustioribus (2-3 T35S, R4/2W, S22, Pink Wasatch Formation, mm nee 6-12 mm) et rhizomatibus gracil- at west end of Red Canyon, road to Bryce ioribus (5-10 nee 10-20 mm) differt. Canyon National Park, in ponderosa pine- Rhizomes less than 10 thick, clothed limber pine community, 8 June 1982, Mutz mm with shredded fibrous leaf bases; leaves sev- and Zarnekee 82-70 (BRY). eral to many on both fertile stems and innova- This variety is partially sympatric with both tions, 2-5 mm wide, (4) 7-24 cm long, strami- varieties chamhcrsii and memhronacea , and neous to brownish or purplish basally; flower the soboliferous condition might be an ecolog- stem 4 cm tall, with several sheathing leaves ical response to creeping substrates. The free almost throughout; flower 1; spathaceous recognition here calls attention of other work- bracts linear-attenuate, 2-3 mm wide, 5-6 ers to these plants and allows for further work. cm long, parallel, subopposite, herbaceous; Thehjpodiopsis sagittata (Nutt.) Schulz var. ovary ca 12 mm long; perianth apparently ovalifolia (Rydb.) Welsh comb. nov. [based white, the tube 15 mm long; sepals ca 6 cm on: Thehjpodium ovalifolium Rvdb. Bull. Tor- long and 1 cm wide; petals narrowly oblance- rey Bot. Club 30: 253. 1903], olate, ca 6 cm long and 8 mm wide; style branches ca 2.8 cm long, the crests ca 7 Hydrophyllaceae mm long; anthers ca 13 mm long; capsule un- Material oiPhacelia demissa Gray from the known. Uinta Basin differs subtly in several ways from Type: USA: Utah, Kane Co., T43S, RIE, that elsewhere in Utah. The plants are finely East Clark Bench, south of US Highway 89, but not densely glandular-puberulent through- sandy soil, found in a semidesert grass shrub out, not densely glandular- villous to glandu- communitv at 1,403 m, Mav 1976, Vane O. lar puberulent; the corolla tube is bright yel- Campbell 42 (Holotype BRY). low, not pale yellow; the sepals are fully twice This distinctive narrow-leaved Iris species as long as the capsule, not less than twice as is compared to / . missoiiriensis in the diagno- long; the seeds are 4-12, not 10-16; and the sis, but its relationships apparently lie with plants average smaller. They are here desig- those far to the west in the coastal states. More nated as follows; material will provide both a better description Phacelia demissa Gray var. minor N. D. and a better possibility of determining proba- Atwood var. nov. Similis var. demissae in ble relationships. forma sed in pubescentibus, corollae tubis flavis, sepalis longioribus, seminibus pauci- Leguminosae oribus et minoribus differt. Type: USA: Utah. Uintah Co., T3S, R23E, Astragalus preussii Gray var. cutleri 84, ca 1.6 km E of Red Fleet Dam, at 1,891 m, Barneby, var. nov., cum var. preussii legu- in a desert shrub community, on clay hills, 31 mine stipitato congruens sed ab ea statura May 1979, N. D. Atwood 7247 (Holotype pumila (caulibus 2-11 nee 10-40 cm altis) e BRY; isotypes 2, distributed previously as radice anmra ortis, foliolis paucis majusculis

Phacelia demissa Gray). 5-11 et 12-20 .X 7-12 mm usque, petalisque Additional representative specimens: albidis pallide coerulescentibus (nee praeter Utah. Uintah Co., Uinta Basin, Utah Highwav alas saturate purpureis) diversa. 121, 10.6 km east of Lapoint, T5S, R20E, S4, Type: U.S.A. Utah. San Juan Co.: common 1,675 m, abundant on gravellv hills, 25 Mav gravel outwash. Copper Canvon, 1 mi from 1978, N. H. Holmgren, P. K. Holmgren, and R. mouth, 18 June 1938, Hugh C. Cutler 2283 C. Barneby 8758 (BRY); ibid.. Orchard Creek, (Holotype NY; isotypes, CAS, WIS); ibid., Dinosaur National Monument, Mancos Shale, bench between Copper Canvon and No Man's Clav soil, Atriple.x-Sarcobatus association, ca Mesa, 22 Mav 1985, N. D. Atwood 11073 1,455 m, 8 May 1955, S. L. Welsh 159 (BRY). (Paratypes BRY, NY). April 1986 WELSH: New PlantTax.\ AND Combinations 257

In my monograph (Mem. New York Bot. aromaticum sensu stricto in floribus mi- Card. 13: 576. 1964) the type of var. cutleri noribus (6-7.3 mm nee 10-13 mm), calyce et was noticed as unusual in the species as repre- dentibus minoribus (3.5-5.3 nee 10-11 mm sented in Utah Canyonlands for few and large et 1.6-2.9 mm nee 5-7 mm respecte), et leaflets but was passed over as a set of caulibus decumbentibus differt. seedlings showing only juvenile foliage. The Type: USA: Utah, San Juan Co., Rone Bai- rediscovery by Duane Atwood of the same ley Mesa, ca 17 km W of La Sal, T30S, R23E, plant, found in a uniform population of small S19, at 1,900 m, pinyon-juniper community, monocarpic individuals such as are never seen on mesa summit red siltstone of the Entrada elsewhere in the range of var. preiissi, show Formation, 4 June 1985, S. L. Welsh & E. that I misjudged the significance of the differ- Neese 23500 (Holotype BRY, 4 isotypes to be ences. Moreover, the flowers were described distributed). by Cutler as varying from white to pale blue Additional SPECIMENS: Utah, San Juan Co., and by Atwood as white when fresh, acquiring T31S, R21E, S6, South Sixshooter Peak plat- a bluish keel-tip when dried. In southeastern form, Davis Canyon, at 1,708 m, on Moenkopi Utah the petals of var. preiissii are consis- Formation in a shadscale community, 11 May tently vivid purple except for pallid wingtips. 1982, S. L. Welsh, B. T. Welsh, and M. Chatter- In the same region the leaflets of the longer ley 21092; ibid.. Rone Bailey Mesa, 1 July 1984, leaves are at once more numerous (17-23) and J. S. Tuhy 1580; ibid., 4 June 1985, E. Neese & smaller (6-15 x 3-6 mm) than those of var. S. L. Welsh 16990; ibid., 19 June 1985, L. C. & cutleri. Leaves of allopatric var. laxiflorus E. Higgins 15812 (all BRY). similar to of var. cutleri in Gray may be those This phase of P. aromaticum is remote from size and number, but its pod is sessile, its the known populations near Bedrock, Colo- stems taller, and its range of dispersal differ- rado (the type locality), and in the Fisher Tow- ent. ers area vicinity east of Moab. The plants dif- Lotus plebeius (T. Brandg.) Barneby [based fer in subtle ways besides those cited in the on: Hosackia plebeia T. Brandg., Proc. Calif diagnosis. When viewed from afar they ap- Acad. Sci. II, 2: 144. 1889 (Brandegee s.n. 30 pear grasslike in patches between sparse ju- Apr 1889, "El Rancho Viejo" [near Calmalli, nipers on a rimrock situation atop Rone Bailey Baja California Norte, +/- 28° 15' N]]. Mesa. The plants tend to sprawl, rather than Lupinus polyphyllus Lindl. in Edwards stand erect as in the typical variety. The ex- var. ammophihis (Greene) Barneby stat. nov. tensive rhizome system is similar, however,

[based on: Lupinus ammophihis Greene Pit- to the typical material. The variety is named tonia4: 136. 1900]. in honor of its discoverer atop Rone Bailey Lupinus polyphyllus Lindl. in Edwards Mesa, Mr. Joel Tuhy. var. humicola (A. Nels.) Barneby comb. nov. Pediomelum epipsilum (Barneby) Welsh [based on: Lupinus humicola A. Nels. Bull. comb. nov. [based on: Psoralea epipsila Torrey Bot. Club 25: 204. 1898]. Barneby Leaf!. W. Bot. 3: 193. 1943, type Lupinus argenteus Pursh var. fulvomacu- from 17 mi E of Kanab toward Jepson Spring, /a

Nyctaginaceae Type: USA: Utah. Kane Co., Smoky Creek Dugway, ca 17 mi NE of Clen Canyon City, Most specimens oiAhronia nana Wats, are Straight Cliffs Formation, 28 August 1973, N. strictly acaulescent and have best developed D. Atwood 5957 (holotvpe NY!; isotype leaves mainly less than twice as long as broad. BRY!).

These represent var. nana . However, in the Robust, taprooted annual to 15 dm tall, Swell, County, Utah, there San Rafael Emery with a single erect, excurrently branched with leaves 2-4 occur caulescent plants main stem; leaves scattered along the stem but times longer than broad. These are named as somewhat basally disposed, at least some of follows: the cauline ones evidently petiolate, with Abronia nana Wats. var. harrisii Welsh broadly ovate to ovate-oblong or elliptic, var. nov. Similis Abroniae nanae Wats var. l)asally rounded to truncate or subcordate, nanae in anthocarpis floribus et bracteis sed in irregularly denticulate blade 4-6 x 2.5-5 cm; caulibus et foliis longioribus differt. herbage glandular-puberulent; flowers axil- Type: USA: Utah. EmervCo., T20S, RUE, lary to small bracts on the lateral branches and S22, San Rafael Swell, ca 1 mi S of San Rafael toward end of the main stem, in bud crowded, River campground, 1,586 m elev., Eri- erect, and nearly sessile, but becoming more ogonum- Astragalus-Hilaria community, 9 widely spaced and evidently pedicellate at

June 1979, J. Harris 364 (Holotype BRY;' Iso- anthesis, fairly showy and adapted to cross types, two distributed previously as A . ellip- pollination; petals purple, 7-14 mm long tica A. Nels.). somewhat asymmetrically (adaxially) dis Additional specimens: Utah. Emery Co, posed; stamens 4 + 4, unequal, declined; an T19S, RIOE, S35, San Rafael Swell, on road to thers ca 1.5-2 mm long; fruits 12-25 mm Wedge Overlook, ca 4 mi S of Buckhorn Flat long, spreading-deflexed, on spreading-de- crossroads, pinyon-juniper communitv, 13 flexed pedicels 3-5 mm long; seeds biseriate June 1979, K. Despain 517 (BRY); do, T20S, in each locule, plumply obovate from a sub- RUE, S22, San Rafoel Swell, Bottleneck Peak stipitate base, 1.5-1.8 mm long, very finely area, off the west side of Cottonwood Draw cellular-reticulate.

road, ca 2 mi S of Bridge Campground, 14 This species is known only by several col- June 1979, K. Despain 564 (BRY). The plant is lections by Duane Atwood from the type lo- named after Dr. James Harris, student of the cality on open slopes on the southwest side of San Rafael Swell flora. the Kaiparowits Plateau, ca 27 km by road Mirabilis linearis (Pursh) Heimer var. de- northeast of Glen Canyon City, Kane Co., cipiens (Standi.) Welsh stat nov. [based on: Utah. Flowering is from August to October. AlUonia decipiens Standi. N. Amer. Fl. 21: Camissonia boothii var. condensata 223. 1918]. (Munz) Cronq. comb. nov. [based on: Oenothera decorticans yar. condensata Munz Bot. Gaz. 85:247. 1928]. Onagraceae Camissonia boothii var. villosa (Wats.)

Camissonia atwoodii Cronq. sp. nov. Spe- Cronq. comb. nov. [based on: Oenothera cies sectionis Chylismae; herba annua, glan- ahjssoides var. villosa Wats., Proc. Amer. Acad.

dulosa-puberula, rotusta, usque ad 15 dm Arts 8: 591. 1873; not Oe . villosa Thunb. 1794; alta, caule excurrenter ramosa, foliis leviter typification will be discussed in the forthcoming basin versus dispositis, laminis subrotundis, treatment in the Intermountain Flora.]. irregulariter denticulatis, 4-6 x 2.5-5 cm; in- Camissonia clavaeformis (Torr. & Frem.) florescentia terminalis, laxe ramosa, floribus Raven var. purpurascens (Wats.) Cronq. comb, pedicellatis, petalis purpureis 7-14 mm nov. [based on: Oenothera scapoidea var. piir- longis leviter versus axem dispositis, sta- purascens Wats. Proc. Amer. Acad. 8: 595. minibus 4 + 4, inaequalibus, declinatis; fruc- 1973], tus 12-25 mm longi, divaricato-deflexi, pedi- Camissonia scapoidea (T. & G) Raven var. cellis divaricato-deflexis 3—5 mm longis, utahensis (Raven) Welsh comb. nov. [based on: seminibus 1.5-8 mm longis, in (^uociue loculo Oenothera scapoidea ssp. utahensis Raven biseriatis. Univ. California Publ. Bot. 34: 96. 1962]. .

April 1986 WELSH: New Plant Taxa and Combinations 259

Oenothera caespitosa Nutt. var. mainly along the margin of the detrital slope macrog/offis (Rydb.) Cronq. stat. nov. [based adjacent to the back wall of the alcoves. Addi- on: Pachi/lophus macro^lottis Rydb. Bull. tional work is necessary to determine the total Torrey Bot. Club 30: 259. 1903]. distribution. Oenothera caespitosa Nutt. var. navajoen- sis (Wagner, Stockhouse, & Klein) Cronq. PapaVE RACE AE stat. nov. [based on: Oenothera caespitosa ssp. navajoensis Wagner, Stockhouse, & Papaver radicatum Rottb. var. ptjgmaeum Klein Monogr. Syst. Bot. Missouri Bot Card. (Rydb.) Welsh stat nov. [based on: Papaver In press 1985]. pi/gmacum Rvdb. Bull. Torrev Bot Club 29: Oenothera flava (A. Nels.) Garrett var. 159. 1902]. acutissima (W. L. Wagner) Welsh comb. nov.

[based on: Oe . acutissima W. L. Wagner Primulaceae Syst. Bot. 6: 153. 1981, type from Flaming Gorge vicinity, Daggett County, Utah]. Dodecatheon pidcheUum (Raf.) Merr. var. Oenothera primiveris Gray var. bufonis zionense (Eastw.) Welsh stat. nov. [based on: (Jones) Cronq. comb. nov. [based on: Oe Dodecatheon zionense Eastw. Leafl. W. Bot. bufonis Jones, Contr. W. Bot. 8: 28. 1898]. 2: 37. 1937, type from Zion Canyon].

Orciiidaceae Ranunculaceae

Habenaria zothecina Higgins & Welsh sp. Aquilegia flavescens Wats. var. rubicunda nov. Planta Habenaria sparsiflora affinis et (Tidestr.) Welsh stat. nov. [based on: Aquile- similis sed in calcari labium 1.5 longiore et gia rubicunda Tidestr. Amer. Midi. Nat. 1: floribus paucioribus diftert. 168. 1910]. Type: USA: Utah. Grand Co., ca 1.6 km N A peculiar specimen oi Aquilegia taken by of Moab, T25S, R21E, S25, at ca 1,281 m, in a Dr. Robert Foster from Zion Canyon in 1977 hanging garden communitv, 10 |ulv 1985, S. requires description. It has the habit and

L. Welsh & L. C. Higgins 23629 (Holotype flower color of A . fl)nnosa but is glandular BRY; 4 isotypes to be distributed. throughout and both petal spur and blade are Additional collections: Utah. Grand longer than in that taxon. A search of the Co., Arches National Monument, 28 Sept. herbarium at Zion National Park yielded two 1963, S. L. Welsh & G. Moore 2735; ibid., specimens of the same taxon taken many years Negro Bill Canyon vicinity, 3 August 1984, B. ago by Dr. Angus Woodbury. Initial efforts to Franklin 1108. San Juan Co., Natural Bridges relocate the area of collection of the type have National Monument, 13 August 1963, S. L. been fruitless. There are many areas to be Welsh & G. Moore 2410; ibid., 15 August investigated, however, and the Hidden 1963, S. L. Welsh & G. Moore 2503; ibid.. Canyon locality of the Woodbury specimens The Neck vicinitv, 31 Oct. 1964, S. L. Welsh has not been searched recently. Possibly the &G. Moore 3840 (all BRY). specimens result from introgression between This peculiar, few-flowered bog orchid with A. fl)rmosa and A. chnjsantha Gray, both its very long spurs has been known for many abundant in the canyon. The longer spurs and years. It has always been placed previously petal blades suggest such a possibility, but within an expanded version off/, sparsiflora neither of the potential parents are glandular but differs as outlined in the diagnosis and in below the inflorescence. The plant is named other salient features. The leaves are broadly after Dr. Robert Foster, enthusiastic collector rounded initially, becoming obtuse and finally and plant geographer. acute upward. The orchid grows in the vege- Aquilegia formosa Fisch. in DC. var. /os- tative assemblages known as hanging gardens teri Welsh var. nov. A var. formosa in petali along the canyons of the Colorado River in laminis et calcari longioribus et herba glandu- southeastern Utah. They occur there with losis diversa. maidenhair fern, sheathed death camus, and Type: USA: Utah. Washington Co., T41S, other shade-tolerant mesophytes. They occur RlOW, S21, W of Tunnel, 1,373 m, N slope of 260 Great Basin Naturalist Vol. 46, No. 2

Bridge Mt., mountain brush community, on ROSACEAE i detritis, 25 1977, R. & R. Foster Wingate May Purshia mexicana (D. Don) Welsh comb, BRY). 3939 (Holotype nov. [based on: Cowania mexicana D. Don Additional specimens; Utah. Washington Trans. Linnaean Soc. 14:575. 1825]. Co., Zion National Park, Hidden Canyon, in Purshia mexicana (D. Don) Welsh var. 1924, A. Woodbury s.n. (ZNP herbarium, 2 stansburyi (Torr.) Welsh stat. nov. [based on: specimens). Cowania stansburiana Torr. in Stansbury Delphinium andersonii Gray var. scapo- Expl. Surv. Utah 386. 1852]. sum (Greene) Welsh stat. nov. [based on: Del- phinium scaposwn Greene Bot. Gaz. 6: 156. RUBIACEAE

1881]. Galium mexicanum H.B.K. var. asperri- Delphinium occidentalis (Wats. ) Wats. var. mum (Gray) Higgins & Welsh stat. nov. harbeyi (Huth) Welsh comb. nov. [based on: [based on: Galium asperrimum Gray Mem. Delphinium exaltatum var. harbeyi Huth He- Amer. Acad. Arts II. 4: 60. 1849]. lios 10: 35. 1892]. Ranunculus andersonii Gray var. juniperi- SCROPHULARIACEAE nus (Jones) Welsh stat. nov. [based on: Ra- Castilleja parvula Rydb. var. revealii (N. nunculus juniperinus Jones Proc. Calif. Acad. Holmgren) N. D. Atwood stat. nov. [based II. 5: 616. 1895]. on: Castilleja revealii N. Holmgren Bull. Tor- Ranunculus acris L. var. aestivalis (L. Ben- rey Bot. Club 100: 87. 1973]. son) Welsh comb, nov. [based on: Ranunculus Castilleja rhexifolia Rydb. var. sulphurea acriformis var. aestivalis L. Benson Amer (Rydb.) N. D. Atwood stat. nov. Based on: Midi. Naturalist 40: 43. 1948], type from 8.3 Castilleja sulphurea Rydb. Mem. New York mi N of Panguitch]. Bot. Card. 1:359. 1900].