Vol. 8, No.1· 2, March· June, 1994 125
A revision of the genus Anisostena weise (Coleoptera. Chrysomelidae, Hispinae). Part II. The subgenus Anisostena: Key to the species groups and the ariadne species group.
C. L. Staines 3302 Decker Place Edgewater, Maryland 21037
Abstract The subgenus Anisostena (Anisostena) is divided into three species groups. A key to the species groups is provided and the ariadne species group is revised. Lectotypes are designated for A. ariadne, A. bellula, and A. elegantula.
Key words: Anisostena, lectotype, species group
Introduction Description: Head: last five antennal segments This is the secondin a series ofarticles revising WIder than precedmg; sulcus on mner margm of the genus Anisostena. The first artiCle (Staines, each eye; row ofpunctures on outer margin ofeaeh 1993) summarized the work on the genus and eye. Pronotum: basal margin bisinuate. Elytra: rev ised the subgenera Neostena and Apostena. costate intervals 2 and 6 unite on apical fjfth. T esfor the ariadne s ecies rou were exam· Anisostenas. str. divideswellinto three species e or a species excep . ico r. ec ypes are designatedfOr it ariadne, it bellula, and it elegan- species, A. nunenmacheri Weise, has color morphs tula. In recording the label data from type speci. which fall intotwo species groups. This species will mens, a slash (f) di~ ides data on different labels; a be covered in both species groups. Otherwise the species have only one color morpho question mark (?) indicates that the label data was unreadable. Measurements were taken with an ocular mi· crometer. Pronotal length and WIdth were taken Key to the species groups of Anisostena along the midlines. Elytral width 'Nas measured at (Anisostena) the humeri. Elytrallength was measured from the base to apex along the suture. Total length was I. Dorsum umcolorous mgrlta group measured from the base ofthe frontal sulcus to the Dorsum with more than one color 2 apex of the elytra. 2. Elytra WIthout markmgs arIadne group Elytra vittate or maculate pilatei group
Anisosfena (Anisosfena) ariadne group Type species: A. elegantula (Baly), designated by The speCIes m the ariadrie species group are Mont6s & Viana (1947) distinguished by the reddish pronotum and daIker elytra. Members of this species group are found from New Jersey south to Colombia. 126 Insecta Mundi
Key to the species of the Chatistenu atiudne (Newman). Daly 1864.252, Clotch 1873b:101; Schwarz 1878:460; Horn 1883:299; ariadne species group Donckier 1899:583; Smith 1899:317; Dike 1902:29; 1. Pronotum with transverse basal impression .... 2 Smith 1910:355. Pronotum without transverse basal impression. GhalepU8 ariadne (Newman). Gemminger & Harold ...... 5 1876:3613. Odontota ariadne (Newman). Crotch 1873a:82. 2. Pronotum with median longitudinal carina ...... Ani808tena ariadne (Newman). Weise 1911a:21, ...... leconteii (New man) 1911b:32; Leng 1920:303; Blatchley 1924:43; Brlmely Pronotum without median longitudinal carina 3 1938'230; Loding 1945'136; BI ackweJder 1946'724; FaUlg 1948:41; Papp 1953:57; Uhmann 1957:73; 3. Pronotum with apical or apical and basal margins Kirk 1969'98,1970'94; Ba]sbaugh &Hays 1972'183; dark syangptera (Suffrian Wilcox 1975:140; Riley & Enns 1979:79; Ford & Pronotum entirely red .4 Cavey 1982'189, 1985'49
be~;~~ 4 Head with median sulcus deep, v-shaped Description' Head and elytra bluish-black, eyes elegantulaa y Head with mediam sulcus not v-shaped between pronotumred. Head: vertexconcave, micropunctate; eyes bwoLOr (Smith) median sulcus present, changes to a carina which becomes the keel between the antennae; antenna! 5. Antennal segment 2 longer than 3 6 pit small; sulcus present on inner margin of each Antenna] segment 3 longer than 2 7 eye; row ofpunctures on outer margin ofeacb eye' reddish projection at base of antennae. Antennal 6. 'Vertex of head micropunctate, with row of three segment I 8ubglObose, punctate; II elOngate, ex punctures between median and ocular sulci; pandedapically, lon~er than III; III c~lindrical;IV- prosternum red laterally ...... texana Schaeffer VI transverse, 'VII- transverse, wi er than pre Vertex of head micropunctate, without larger pundules, PIOstelllulll entirely led .. ce~g; ~ ~Ointerl at ap~x; segm~nt~ ni ~J?~IY ...... ariadne (Newman) pu iscen .rond;um: aplcalmargm lac,le er in middle; wider than long; punctures large, deep; 7. Head with median sulcus deep, wide; surface of micropunctate between punctures; sides parallel sulcus punctate, pronotum with scattered punc on basal half, weakly convergent on apical half, tures, disc almost impunctate .. basal margin bisinuate; length 0.7-1.0 mm (avg...... hans_a Sehaeffer 0.8, n 12); width 1.0-1.1 mm (avg. 1.0). Scutellum: Head with median sulcus not deep or wide; sur- (USNM), Miami, V 1920 (ICOM). Du~al Co.- Jackson- ville (USNM); Hilliard, 28-VII-1934 (SEMC). Franklin , , (EGRC). Hardee Co.- Highlands Hamm. St. Pk., Il-XI- , " (FSCA). Hendry Co.- La Belle, 21-IV-1921 (PADA), 16- , VII-1977 , , 29-VIII-1984 (ISAC), 13-IV-1955 (FSCA); Cedar Keys, , , , 28-III-1976 (EGRC). Manatee Co.- Bradenton 22-III- Hwy. 40 at Zay Pra., 30-IV-1980 (EGRC). Martin Co.- , , 23-III-1982 (CNC). Osceola Co.- 7 mi. N Yeehaw Jcn., , (INHS); 6 mi. N Land O'Lakes, 23-VII-1973 (EGRC). (UMMZ); Oldsmar (FSCA); St. Petersburg, 18-26-VIII- n is s an or , , circles; a. leconteii, triangles. (FSCA). 16-III-1927 (USNM), ll-VII-1951 (SEMC). Volusia Co.- Enterprise (MCZC); New Smyrna, 29-VII Specimens examined: UNITED STATES. USA 1948 (SEMC). Wakulla Co.- 20-1'1-1955 (INHS); 2 mi. (USNM). ALABAMA: Baldwin Co.- 1 mi. N Foley, 30 N Live Oak Pt., 7-VIII-1973 (EGRC). Washington CO. '1-1981 (EGRO), Robertsdale, 12-'11-1945 (USNM). EblO, I-1V-1973, 4-1V-1973 (ONO). GEORGIA. FOI t Mt., Barbour Co.- Spring Hill, 4-V-1912 (USNM). 1O-VII-1937 (USNM). Charlton Co.- Okefenoke Swamp, Crenshaw Co.- Oounty Lake, 2B-VIII-19B4 (ADEM). 27 VII 1939, 1 VIII 1939 (SEMO), 29 VII 1939 (INlIS). Lee Co.- Auburn, 20-IV-1975 (CNC); 5 mi. S. Meadow LOUISIANA: S. La., 29-V-1892 (USNM); Pollard, 12 Mill, 13-IX-19BB (ADEM). Mobile Co. Grand Bay, VII 1983 (ONG). Acadia Par. 2.8 mi. S Eunice, 9 V 1933 (USNM); Mobile, 20-IV-1912, VII (USNM); Or- 1981 (EGRC). Beauregard Co.- 16-VIII-1928 (SEMC); ellard, 5 V 1918 (MOZe). Talladega Co. Galt Grove, 5 mi. S DeRidder, 25 III 1982 (EGRC). Natchitoches 17-VI, 18-VI (USNM). Tallapoosa Co.- Horseshoe Bend Par.- 8 mi. E Kisatchie, 7-IX-1985 (EGRC); Kisatchie Natl. Park, 29 VII 1964 (,A.DEM, NDSD). l.RKANSAS: Nat. For., nr. Red Bluff eampgrd., 15 V 1982 (EGRC); Logan Co.- Mt. Magazine, 15-16-VII-1949 (INHS). DIS- Red Dirt, lookouttower, 26-IX-I981 (EGRC); Fire Tower, TRICT OF COLUMBIA: no further data (ICCM). 15 V 1982 (EGRC); Red Dirt W.M.A., 20 VI 1986 FLORIDA: Paradise Key, 24-II-1919 (USNM); St. Johns (EGRC). St. Tammany Par.- Covington, VI (USNM), Bluff (BMNH); W. Pk., 24-XI-34 (FSCA). Alachua Co. 23-VI-1948 (SEMC). Wmn Par.- 8 ml N Wmnheld, 12 Gajnesvjlle (FSCA), 29-III-19BO, I-IV-19BO (CNC); VIII-19GB (IISNM) MARYLAND' Dorchester Co- Newnan's Lake, I-IV-SO (FSCA). Bay Co.- Ebro, I-VI Blackwater Refuge, 10-VII-1943 (FSCA). Somerset Co. 1973 (CNG) Bradford Co.- St.arke, 29-III-1976 (EGRC) Hopewell, 22-VII-1976 (CLS). MISSISSIPPI' Adams Charlotte Co.- 6-IV-1985 (ISAC); 8-IV-84 (FSCA); 5 ml. Co.- Homochito Nat. For., Sandy Ck. W.M.A., Pipes S Punta Gorda, 13-VII-1953 (INHS) Collier Co.- 5 mi Lake, 30-VII-1986 (EGRC). George Co.- Pascagoula S Corkscrew Swamp, 5-Iv-1972 (FSCA); 12 km E Naples, Wildlife M., nr. Lucedale, 12-IV-1987 (EGRC). Jack 8-III-1982 (CNC); Ochopee, 14-VI-1977 (NDSU). Dade son Co.- Hurley, 18-VI-1929 (CUIC); Ocean Springs, Co.- Andytown, 22-III-1973 (CNC); Hialeah, 27-IX-1971 29-VII-1920 (MUIC). Stone Co.- U. M. Forestlands, 18- 128 Insecta Mundi X 1986 (UMIC). MISSOURI. St. Clair 00.- Dtryal Town- III longer than II, IV-V] transverse, decreasing in ship, T37N R26W, secs. 35 & 36, 6-V-1979 (EGRC); length; VII-X transverse, wider than preceding; XI LiGhen Glade, 6 mi. W Osoo01a, 29 V 1971l (EGRC). pointed at apex, Yv'II-XI hirsute, I-III punctate, IV- NEW JERSEY: Atlantic Co.- DaCosta, 3D-VII (USNM), ~ so~e ~et~e Pronotu~' ~n?er tha~ 3 \11 (MC:ZC). Burlington Co. Wharton TraGI., 26 VII with wide' 1983 (CLS). NORTH CAROLINA: Cumberland Co. F'ayetteville, ]4-V-1939 (FaINt:) Pitt Co - Orimesland, V-1919 (EDNC) Wake Co - Raleigh, 14-V-1931, 27-VI- convergent; apical margin darkened; covered with 1936 (EDNC). OKLAHOMA: Kay Co.- Ponca CIty, S-VI lalge, comse panctmes, length O. 7-0.9 nnn (avg. 1947 (CNC) SOUTH CAROLINA' BarnwelJ Co -Edisto 08,0-2); width 06-07 mm (avg 065) Scutellum' Exp. Sta., BlackVIlle, 15-V-1940 (CDCC). Colleton Co. quadrate; micropunctate. Elytra: 8 regular punc Ashepoo, ] 4-VIII-] 935 (CUCC) Dorchester Co.- ture rows plus scutellar row of 1 puncture; inter vals 2, 4, and 6 costate, all evenly raised; intervals 2 and 6 unite on apical fifth, interval 4 does not 1962 (tIDS ). Wil iamsb rg Co.- ~eville, 30-VI- attain union b the diameter ofa uncture' lateral Waynesboro, 4-VIII-1948 (SEMC). TEXAS: Anderson , , Brooks Co.- Falfurrias, 22-III-1987 (ZMA). Kenedy 00. 3 mi. N Norias, II ... y. 77, 28-III-1987 (ZMA), Sarita, 24-III-1987 (ZMA), 15-IX-1975, 14-V-1976 (USNM). Sahine Co. Sabine Nat. For., 3.3 mi. E Pineland, 29 ViI-1975 (TAMU). Tyler Co.- Kirby State Forest, 3 mi. S Warren, 22 V 19l14 (TAMU). VIRGINLA.: Fairfax Co. Larval host plant: unknown. FaUs Church, 17-VI-1923 (USNM). Total: 306. Distribution: Mexico to Colombia. Amsostena bellula (Baly) Specimens examined: COLOMBIA: Bogota Map 2 (BMt-.'H). COSTA RICA. La Caja, San Jose, III-1929 (USNM). Total: 4. Charistena bellula Baly 1864:253. Lectotype (here des ignated). (?)I Type II. T. (white disk with led bOI- der)! Syntype (white disk with blue border)! Baly Anisostena bicolor (Smith) Go1leetionl Gharistena be1lula Baly, Bogota (blue green label)! Lectotype Anisostena beUula (Baly) Maps 2 and 3 designated G. L. Staines 1989 (red label) (BMNH). Two paralectotypes (here designated): E. Coll. Charistena bicolor Smith 1889:94; Donckier 1899:583; LaCarte!Odontota eyanipennisRaieh., NoyaGranata Fall & Goel<01'e1l 1907:200. (green label)! 67-56! Ex. Rev. H. Clark ColLI Syntype Anisostena bicolor (Smith). Weise 1911a:21, 1911b:32; Anisostena beIlllla Balyf Paralectotype Anisostena Leng 1920'303; SGhaeffer 1933:104; Smith & Wilbur beUula (Baly) designated C. L. Staines 1989 (red 1937:103; Papp 1953:58; Uhmann 1957:73; Wilcox label); E (:011 I,afart.el 6'7-56 (reversed)1 F:x Rev H 1975'1411; Thomas & Werner 1981'2 Clark CollI Syntype Anisostena bellnla Balyl Para- lectotype Allisostena bellula (Ealy) deSIgnated C. L. Description: Head with metallic green sheen, ~ain;.s ];~; (~~~ label) (BMNH) Baly ]885'45' elytra dark metallie blue, pronotum red. Head: onc ler 9: . median sulcus present, becomes a keel between Chale usbellula al . Gemmin er&Harold 1876:3613. u u is s ena e u a a y . eise ,a: , 1911b:32; Blackwelder 1946:724; Papp 1953:58' eye; vertex with three punctures; row of three Uhmann 1957:73: Wilcox 1975:140. punctures along SUlCI near eyes; vertex micropunetate; row ofpunetures on outermargin of Description: Head and elytra metallic blue, each eye; no projection at base of antennae; frons pronotum red Head' median sulcus pres€lnt; sulcus andclypeus 1 eddish. AntennalsegmentI subglobose: present on inner margin of each eye; vertex II-IIIcylindrical, III longest; IV-VI transverse; VII micropunetate; areanearbase ofantennae reddish; X transvers€l, wid€lr than pr€lc€lding; XI pointed at row of punctures on outer margin of each eye. apex; VII-XI hirsute. Pronotum: apicalmarginblack; Antennal segment I subglobose; II-III cylindrical, covered with large, deep punctures, more dense Vol. 8, No.1 - 2, March - June, 1994 129 115 110 lOS 100 95 90 85 80 75 70 65 60 \ ) I '\~ y '\ J ~ 2 h/[ o o ~ _5 lUI-- j ) ~I _ 5 / Or-- Au f I" 110 105 100 95 90 85 80 75 70 Map 2. Distribution of Ani80stena species: A. betlula. triangles: A. bicolor. squares: A. cyanoptera, solid circles; A. elegantula, open circles. laterally; lateral margins parallel for basal half, rugose at sides; abdomen reddish, rugose at sides. weakly convergent apically; median tranS""erse Total length: 3.8 4.6 mm (avg. 4.3). impression at base; basal margin bisinuate; area between punctures alutaceous, length O. 7-0.9 IJlIn Larval host plant: Tripsacum dactyloides (avg 09, n 7); width J 0 mm Scute))um' quadrate, (poaceae) (Smith & Wilbur, 1937) alutaceous. Elytra: bluish; eight regular puncture rows plus scutellar row Of 2 punctures; lateral Distribution: Kansas, Arizona, New Mexico, Cali margins smooth, sinuate; apical margin dentate; fornia, and Mexico. intervals 2, 4, and 6 costate, interval 6 most pro nounced; intervals 2 and 6 unite on apical fifth, 4 Specimens examined: MEXICO· BAJA CALIFOR- does not attain union by the diameter oftwo punc NIA SUR- 3 km S Micraflores, 17-X-1978 (FSCA); San tures; length 3.0 3.7 mm (avg. 3.4); width 1.3 1.4 Jose de Cabo (lISNM) NAYARIT- 15 mi S Acaponet.a, mm (avg. 1.4). Legs: reddish; mesofemur serrulate 20-VIII-1964 (TAMU). SINALOA- 38 mi NE Concordia, on inner margin, apex offemora, tibiae, and tarsi nr. Loberas. 3-VII-1982 (AJGC); 4.5 mi N Elota, 17-VII- darker. Venter: rosternum reddish unctate' 1984 (FSCA); Los Mochis, 12-IV-1924 (USNM), 28-V- z meso- an me s rna e, mi N Mazatlan (USNM). UNITED STATES: ARIZONA: no fUlthel data (USNM), CALIFORNIA. Rossmond, 23- 130 Insecta Mundi VII-1940 (SEMC). IDAIIO. OeeIY, 17-VII-1979 (EGRC). Distribution: Cuba, Jamaica, and Hispaniola. KANSAS: Reno Co.- 30-VI-1934 (USNM). Total: 19. Specimens examined: CUBA: no further data (MUIR, MCZC); Baragua, 22-X-1925, 13-X-1928 Anisostena cyanoptera (Suffrian) (MCZC); Camaguey, 30-vII-I923 (OSNM); Cayamas, 5- Map2 ~9~~=~~~a~::~~~~s~~:~;:~~~~,a;o~:sX~: 1923 (2:MA); Hoya Colorada, Hav. Pr., 23-VIII-1911 OdontotacyanopteraSuffrian 1868:227. Holotype: 31576! (USNM); Lomus de Camoa, Habana, 18-IX-1956 (INHS); MLU Halle, "VB Zoologie, S.-Nr. 7/1/10, T.Nr. cyan.! Mins Carloto. Cienfuegos, 1-XII-1927 (FSCA); Omaia, cyanipennis m., cyanoptera Wng. Hrch. 1868, Cuba 24-VII-1927 (USNM); Pinar del Rio, 16-29-V-1933 (ZMA); (gleen label) (MLUII). Santiago d. l. Vegas (USNM), Soledad, Cienfuegos, 4-XI- Chalepus cyanoptera (Suffrian). Gemminger & Harold 1927 (FSCA), 16-X-1926 (USNM), V-VI-1939, 16-X-1926 1876.3613, Donckier 1899.586. (MCZC). JAMAICA. St. Andlew Palish- Kingston, IV- Anisostena cyanoptera (Suffrian). Weise 1911a:21, 1906, 25-29-XII-1961, 11-1970 (USNM). St. Ann Parish- 1911b:32; Leng & Mutchler 1914:457; Blackwelder Clermont, 27 XII 1972 (CNC). St. Catherine Parish Rio 1946:724; Papp 1953:58; Uhmann 1957:74; Cobra Gorge, 1 mi. N Flat Bridge, 27-VI-1958 (INHS). Sanderson 1967.138, Wilcox 1975.140. Total: 32. DesCIiption: Headandely tIa blue, plOnotum led. Head' median sulcus present, faint; sulcuson inner Anisostena elegantuJa (Baly) ~argin of each eye; occiput alutaceous; keel be Map 2 tween antennae; tront concave; sides Of head red dish, smooth; projection at base ofantennae; frons Charistena elegantula Baly 1864:254. Lectotype (here red, with tubercle; clypeuspunetate. Antennalseg designated): E. ColI Chevt. (folded)! Type H. T. ment I globular; II cylindrical, as wide as I; III (white circular label with red margin)! Baly ColLI longer than II, cylindrical; IV·VI transverse, de Charistena elegantula Baly, New Granada (pale creasing in length; VII-XI expanded, with setae; green label)!Leetotype Anisostenaelegantula (Baly) VII-X transverse; XI acutely pointed. Pronotum: designated C. L. Staines 1989 (red label). Baly 1885:45; Donekier 1899:583. Chalepus elegantulus (Baly). Gemminger & Harold eral~arginS para~el f~ ba!;;~l Anisostena elegantula (Baly). Weise 1910:123, 1911a:21; two-thirds, then Blackwelder 1946' 124; Papp 1958'58; IIhmaDD weaky convergen ; su ace etween punctures 1957'74; Wilcox 1975'140 alutaceous; length0.7 0.9mm (avg. 0.8; n 6); width 0.9-1.0 mm (avg. 0.95). Scutellum: metallic blue; Description: Head and pronotum orange, elytra quadrate, alutaeeous. ElytIa. basal margin alcu metallic blue. Head: vertex punctate; median sul- ate overla in baseof ron um' un r r , than head; clypeus with projection below base of antennae, punctate, quadrate Antenna] segment I orangish, subglobular; II-XI blackish; IItransverse; diameter of a puncture; lateral margins sparsely III cylindrIcal, longer than II; IV-VI transverse, dentate; apicalmargins dentate; length 2.6-3.0mm decreasing in size; VII X hirsute, transverse; XI (avg. 2.7), width 1.1-1.4nlJn (avg. 1.35). Venter. hirsute, pointed at apex; I-VI punctate. Pronotum: r:~~~~~r;t~::::a:~ea~t~:~~~:~k:~::s~::~:a:~ basal margin bisinuate; sides malgined, basal im- pression present; covered with large punctures, smooth in middle, alutaceous at sides. Legs: tro more dense laterally; alutaceous between punc chanter blue-black; femur red on basal half, rest tures; width 0.8 mm (a-I); length 0.9 mm. Scutel blue-black, punctate, mesofemul not senate on lum: blue; quadrate; alutaceous. Elytra: blue ex cept fol faint olange malkings at base between intervals 2 & 4; 8 regular puncture rows; scutellar row of 2 punctures; intervals 2, 4, and 6 costate; intervals 2 & 6 unite on apical fIfth, 4 does not Larval host plant. unknown. attain union by the diameter of a puncture; sides Vol. 8, No.1· 2, March - June, 1994 131 margined, sparsely dentate, apiealmalgin dentate, laminate' suture costate' surface between nunc- tures amtaceous; lengtn ~.tI mm; wiutn 1.0 mm. •• . n ..n_ ""nil metasternum orange m center, DIue laterally, - ,• 1 1 •• 1. 1 " . L 1. • center orane:e laterally. others all orane:e. Lee:s: lemur ana (,lUUie ~-uase-, -narK 'l'ot.f111t>nO'th· ::t Fi mm ~~ ,/~ ;' \ $ --\f ~ ::~~i • j .... T 1 L_~.. _1_ .... 1 r ~ '?~t AI ~ I r I "" ~ '\/ _r-.... / :::>peClmens exanllnea: New UranaCla . C;U- ~. D~M~"~ ~; <:lt~ T.()l\ffiT A· • <) 1 1<' l\A" o.ta 1 fi. / V-l~ 16 {C;1'\j1,j). : C;t111\.t'A.::i: 4 ml.l'\jJ!j l'lCnUCalCO, 1<1.VT.1Clf:fif'T'AMTT\ ()AVAf'A. 11",; NM"t.;""- I).VH-Url1 . :--zTml.!\lVl11anermosa, 19-VIlI-1967 f'T'AMlJ). VERA CRUZ: Coatzacoalcos 7- IX-1968 (TAMU). Total: 8. / l ~1 --~-L~_C£;;:.~ __Tl: " Anisostena kansana Schaeffer 1~i ~~~-- j 1'C Anisostena kansana Schaeffer 1933:104. Holotype: ... TTC' TIT Tr ~ • _. M ,, ;A,J 'J Holotype, C. Schaeffer Coil., 1935 BarberI Charles '''''' Anisostena kansana Schaeffer (USNM). Papp 1953:58; Ullmann 1957:74; Wileox 1975:140; Riley Map 3. Distribution of Anisostena speCles: A. bu:olor, & Enns 1982:35. squares: A. kansana, triangles: A. texana, circles. Description: Head and elytra metallie greenish alutaceous; 8 leg-ulal puncture rows plus sutural blue; pronotum reddish-orange with anterior por row of 2 or 3 punctures, each elytron may be tion greenish-blue; legs and antennae black Head: asymmetriCal in the number Of sutural punctures; vertex alutaceous; median sulcus deep, wide; sur at base a depression before the humerus; lateral face Of median sulcus punctate; rugose between margins sinuate, smooth; apical margin sparsely eyes; sulcus present on inner margin of each eye; dentate; intervals 2, 4, and 6 costate; interval 2 row ofpunctures on outer margin ofeach eye; sides mostpronouncedon apicalfifth, 4 mostpronounced ofhead smooth; fIOns reddish; plojection at base of on apical fourth, 6 carinate fOr entire length; inter- antennae; dypeus evenly arcuate. Antennal seg vals 2 and 6 unite on apical fifth, 4 does not attain ment I subglobose, punctate; II transverse, punc- union by the diameter ofa puneture; length 3.1 4.1 tate; III cylindrical, longer than II, punctate with mm (avg. 3.7; n-20); width 1.3-1.7 mm (avg. 1.5). setae; IV VI tranS\'erse; VII X, transYerse, .....ider Legs. mesofemur senulateoninnel malgin, femora than preceding, with setae; XI pointed at apex. punctate at apex; tibiae punctate Venter' Pronotum: lateralmarginssinuate, margined; basal prosternum black in middle, punctate, reddish and margin hisinuate; basal impression absent; surface rugose at sides; meso and metasterna black, with scattered large, deep punctures; disc almost alutaceous in middle, punctate·rugose at sides; impunctate; anteriOr margin greenish·blUe, mark- abdomen alutaceous in middle, lugose at sides. ings wider in middle; surface between punctures Total length: 4.1-5.3 mm (avg. 4.8). alutaceous, length 0.7-1.0 mm (avg. 0.9, n 20), width 0.9-1.3 mm (avg. 1.1). Scutellum: quadrate; greenish-blue; alutaceous. Elytra: greenish-blue; 132 Insecta Mundi Larval host plant: 'PI ipsacum dactypoides 6 unite on apicalftfth, 4 does notattain union by the (poaceae) (Riley & Enns, 1982) diameterof2 punctures; interval 2 raisedfor entire length, 4 more pronounced at base and apex, 6 Distribution: Kansas, Missouri, and Oklahoma. costate from humerus; lateral margins sinuate, serrate; apical margins serrate; surface between Specimens examined: KANSAS. Douglas 00.- puncturesalutaceous, length 3.5-3.7 mm (a\l g. 3.65, Baldwin, 23-V-1971 (SEMC). Reno Co.· Medora, May n-5); width 1.6-1.7 mm (avg. 1.65). Venter: (USNM), 3·,/11-1927 (SEMG). MISSOURI. Bates 00.· prosternum black in middle, alutaceous, red later· Ballard, 5-VI-1978 (EGRC). Dade Co.· Niawathe Pririe, ally, punctate; meso· and metasterna black, 10 mi. N Loekwood (EGRC). Pettis Ceo Friendly Prai alutaceous; abdommal segments black except for rie, 3-VI-1978 (NDSU, CNC, EGRC, TAMU). OKLA- ~ter HOMA: Kay Co· Ponca City, 8-VI 1947 (eNG) last '.vhich is laterally. Legs: black; femur Woodward Co.· Woodward, 19-V-1981 (USNM). Total: 84 4.8 (avg. 4.75) mm. Anisostena leconteii (Baly) Larval host plant: unknown Map 1 Distribution: Virginia to Florida. Chal istena lecontii Baly 1864.252. Holotype. E ColI. Laferte (folded)1 Type H.T. (white disk with red border)1 Baly ColLI Charistena lecontii Baly, North Specimensexamined: NORTH AMERICA(BMNH). America (faded green) (BMNII). meoneet oIigi- FLORIDA- Dade Co.. 19.VII.1954 (FSCA) SOUTH nal spelling. CAROLINA: Charleston Co.· 1 ml. W McClellanVIlle, Charistena leGontei (Baly). Croteh 1873b:101; Horn 2-VI-1973 basal margin bisinuate; lateral edges parallel on 8. l'\skevold (18AO); Auburn University (AUEM), basalhalf, weaklyconvergenton apicalhalf; length W. E. Clark; British Museum (Natural History) O. 7-0.9 mm (avg. 0.8, n 6), width 1.0. Scutellum. (BMNH), S. L. Shute, Canadian National Collec- metallicblue; quadrate; micropunctate Elytra' eight tion (CNG), I. LeSage; Carnegie Museum ofNatu- regular puncture rows plus sutural row of 3 punc ral HIStory (ICCM), R. L. DaVIdson; Clemson Om tures; lateralmargins smooth, parallel; intervals 2, versity (CUCC), J. C. Morse; Cornell University 4, and6 costate, 6 mostpronounced; intervals 2 and (CUIC), E. R. Hoebeke; Florida State Collection of 6 uniteon apicalfnth, 4 does notattain union by the Artlllopods (FSCA), M. C. Thomas, Arthur J. Gil diameteroftwopunctures; apical margins dentate; bert (A,JGC); University of Georgia (UGCA) C I. length 3.4-3.6 mm (avg. 3.5); width 1.4. Venter: Smith;UniversityofIdaho (UICM),F. W. Menckel; prosternumblackinmiddle, redatsides; meso- and IllinoisNaturalHistorySurvey (INHS),K. McGiffin; metasterna black; abdomen black. Legs: black; Martm-Luther Omversltat (MLUH), M. Dorn; femora punctate; mesofemora serrulate on inner University ofMiehigan (UMMZ), M. O'Brien; Mis margin. Totallength: 4.1-4.6 mm (avg. 4.4). sissippi Entomological Museum (MUle), T. L. Schiefer, University of Mississippi (UMIC), P. K. Larval host plant: unknown Lago; Museum ofComparative Zoology (MCZG), C Vogt; North Carolma Department of Agnculture Distribution: Oklahoma, Texas, Arizona, and (EDNC), J. F. Greene; North DakotaState Univer Mexico. sity (NDSU), E. U. Balsbaugh; Pennsylvania De partmentofAgriculture (pABA), A G. V/heeler, E. Specimens examined: MEXICO: CHIAPAS- El G. Riley (EGRC); Snow Entomological Museum 8umidero, 20-Vl-1950 (INHS). OAXACA- 9 mt 8E El (SEMC), R. W. Brooks; Texas A & M Umverslty Camaron, 4600 ft., 2-VI-1974 (EGRC). UNITEDSTATES: (TAMIl), E G Riley; IT S National Museum ARIZONA. Pima Cu.- Baboquivari Mts., 30-VII-1987 (OSNM), R. E. WhIte; Zoologlsch Museum (UICM). St. Cruz Cu.- Nogales, 29-IX-1906, 16-IX- Universiteitvan Amsterdam (ZM1'\), C. van Nidek. 1906, 27 IX 1906 (USNM, UMMZ), 3 X-1906 (INIlS, UMMZ); 8 mi. WPena Blanca Lake, 23-VII-1987 (UICM); J. F. Cavey, USDA, APHIS, Baltimore, MD, and R. Ruby, 13-VlI-1940 (SEMC); Santa Rita Mts., Madera E. V/hite cmnmented on an earlier draft of this Cyn., 14 VIII 1988 (UICM). OKLAHOMA: 'Nichita Nat. manuscript For., 4-8-VI-1926 (CNC). TEXAS: nofurtherdata (ICCM). Aransas Co. 6 VIII 1928 (SEMG). Brooks Co. 4.8 mi. N Encino, 12-IX-1987 (EGRC). Cameron Co.- Boca Ghiea, 30 VI 1938 (SEMG). Cemal Ceo NeVI Braunfels, Literature Cited 24-VIII-1947 (CNC). Dallas Co.- Dallas, 15-IV-1940 Balsbaugh, E. D., & K. L. Hays. 1972. The leaf (INHS) Gillespie Co. 23 VI 1940 (GNG). Harris Co. beetles of Alabama. Auburn AgIicultural Ex~ Deer Park, 13-VIII-1981 (ISAC). Hildago Co.- Delta periment Station Bulletin 441'1-223 I .ake Co Pk, !B-X-1988 (El ml)) .lefiD.vis Co,-llavis Mts., 8-VI-1939 (CNC). Kenedy Co.- 1 mi. S Armstrong on Hwy. 77, 8-X-1984 (EGRC); 35 mi N Norias, l3-X- 1988 (EGRC, TAMU); 13 mi S Sarita, 8-VIII-1972 (USNM). Kleberg Co.- Kingsville, ll-IX-1975 (US1'<~). Leon Co.- Flynn, 7.6 mi. N Normangee, 14-VIII-1982 (3)2:251-258. (EGRC). San Patricio Co.- Welder Wildlife Refuge, 3 V-1979 (UGCA). ValVerde Co.-ll mi. NComstock, 25- Baly, J. S. 1885 Biologia Centrali.Americana, VII-1987 (EGRC). Wharton Co.- 2 mi. E Bermard, 16 Insecta, Coleoptera, Hlspldae. 6(2):1-124. V-1984 (UICM); 4.8 mi. W Jet. 71 & 1300, 20-VI-1984 (EGRC). WiHacy Cu.- 7 mi. Vol POlt Mansfield, 3-X-1986 Blackwelder, R. E. 1946. Checklist of the Co (EGRC). Total: 129. leopterous insects ofMexico, Central America, the West Indies, and South America. United StatesNationalMuseum Bulletin 185:551-763. AcknO\vledgements Thefollowing institutions andindividualshave Blatchley, W. S. 1924. The Chrysomelidae of lent material for this study, the assistance of the Florida. Florida Entomologist 8:39-46. curators responsible is gratefully acknowledged- Archbold Biological Station (ABSC), M. Deyrup; I. 134 Insecta Mundi Brimley, C. S. 1938. TheinsectsofNolth Calolina. tural Expeliment Station Technical Bulletin North Carolina DepartmentofAgriculture. Di- 1038: 1-117. vision of Entomology 560 pp. Leng, C. W., & A. J. Mutchler. 1914 A prelimi- Crotch, G. R. 1873a. Materials for the study of the nary list of Coleoptera of the West IndIes as Phytophaga of the United States. Proceedings recorded to Jan. 1, 1914. Bulletin ofthe .Ameri of the Academy of Natural Science, Philadel- can Museum of Natural History 33:392-493. phis 25.19-83. Lang, CW 1920 Catalogue ofthe Coleoptera of Crotch, G. R. 1873b. Checklistofthe Coleoptera of Amenca, north of MeXICO. John D. Sherman, America, north ofMexico. Naturalists Agency, Mt. Vernon, NY. 470 pp. Salem, MA. 136 pp. Lading, H. P. 1945. Catalogue of the beetles of Donckier, H. 1899. Catalogue Systematique des Alabama. University of Alabama Monograph Hispides. Annales de la Societe Entomologique 11.1-172. de France 68'540-615 Melshelmer, F'. E. 1853. Catalogue of the de Fall, H. C., & T. D. A. Cockerell. 1907. The scribed Coleoptera of the United States. Coleoptera of New Mexico. Transactions ofthe Smithsonian Institution, Washington. 174 pp. American Entomological Society 33.145-272. Monras, F., & M. J. Viana. 1947 Revisi6n Fattig, P. W. 1948. The Chrysomelidae or leaf sistematIca delos Hispidaeargentmos (Insecta, beetles ofGeorgia Emory University Museum Coleop., Chrysomeloid.). Anales del Museo t\r- Bunetm 6:1-47. gentma Cienclas Naturales "Bernardino Rwadavia" 42: 125 324. Ford, E. J., & J. F. Cavey. 1982. Biology and immatule stages of the Hispinae beetle Newman,E. 1841. Entomological notes. TheEnto- Anisostena ariadne (Coleoptera' Chryso- mologist 73- 79 mehdae). Coleoptensts Bulletm 36: 189-192. Papp, C. S. 1953. The Hispinae ofAmerica. Third Ford, E. J., & J. F. Cavey. 1985. Biology and contribution for promoting the scientific re Ian> al descriptions ofsome Malyland IIispinae sults ofthe Intelnational Hylean AnIazon Inst. (Coleoptera: Chrysomelidae). 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Caribbean Journal of Science nical Bulletin 1033: 1 117. 7:135-139. Kirk, V. M. 1970. A list of the beetles of South Schaeffer, C. F. A. ]933 Notes 00 some Hispioi Carolina Part 2- Mountain, pedimont, and and Cassidini and descriptions of new species southerncoastalplam. South CarolmaAgncul- Vol. 8, No.1- 2, March· June, 1994 135 (Coleoptera: Chrysomelidae). Pan Pacific Ento· Thomas, D. B., & F. G. \Verner. 1981. Grass mologist 9: 103-109. feeding insects of the Western Ranges: An an- notated checklist. University of Arizona Agri Schwarz, E. A. 1878 The Coleoptera of Florida cultural Experiment Station Technical Bulle- Proceedings ofthe American PhIlosophICal So tm 243:1-50. ciety 17:353 471. Uhmann, E. 1957. Coleopterorum Catalogus, Smith, J. B. 1889. Some new species of IIispini. Supplementa, Chrysomelidae. IIispinae. palS Entomologia AnIericana 5:94-95. 35(1):1-153. W. Junk, Gravenhage. Smith, J. B. 1899 Insects of New Jersey 27th IIlke, H 1902. Alist of the beetles ofthe District of Annual Report of the State Board of Agricul ColumbIa. Proceedirigs of the United States ture. MacCrellish & Quigley, Trenton. 755 pp. National Museum 25:1 57, Smith,J. B.191O. Annualreportofthe New Jelsey \Veise, J. 1910. Zweitel Beittag ZUl Kenntnis del State Museum. MacCrellish & Quigley, Tren- Hispinen Verhandlunger des Naturfor- ton. 888 pp. schenden Veremes m Brunn 48:115-162. Smith, R. C., & D. A. Wilbur. 1937. Insects Weise, J. 1911a. Coleopterorum CatalogusChryso injurious to alfalfa, grasses, and allied plants. melidae. IIispinae. W. Junk, BeIlin. pars 35.1- Biennial Report of the Kansas Agricultural 94. Experiment Station 8: 100-103. Weise, J. 1911b. Coleoptera: Phytophaga, family Staines, C. L. 1993. A reVISIOn of the genus Chrysomelidae, subfamily Hispinae. in P. Anisostena Weise (Coleoptera: Chrysomelidae, Wytsman, Genera Inscctorum fasc. 125:1-123. Hispinae). PartI. Introduction and the subgen Brussels. ela Neostena and Apostena. Insecta Mundi 7'183-190 Wilcox, J. A 1975 Checklist of the beetles of Canada, UmtedStates, MeXICO, CentralArilerIca Suffrian, E. 1868. Verzeichniss der von Dr. and the West Indies. Vol. 1 part 7, the leaf Gundlach auf der Insel Cuba gesammelten beetles (redversion). Biological Research Insti Chlysomelinen. AIChiv fUI Natmgeschiethe tute of Amelica, Latham, NY. 166 pp. 34: 163-252.