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In the National Museum of Natural History, Smithsonian Institution Type Specimens of Hispinae (Coleoptera: Chrysomelidae) in the National Museum of Natural History, Smithsonian Institution CHARLES L. STAINES and SUSAN L. STAINES I W9\ I SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • NUMBER 585 SERIES PUBLICATIONS OF THE SMITHSONIAN INSTITUTION Emphasis upon publication as a means of "diffusing knowledge" was expressed by the first Secretary of the Smithsonian. In his formal plan for the institution, Joseph Henry outlined a program that included the following statement: "It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes made from year to year in all branches of knowledge." This theme of basic research has been adhered to through the years by thousands of titles issued in series publications under the Smithsonian imprint, commencing with Smithsonian Contributions to Knowledge in 1848 and continuing with the following active series: Smithsonian Contributions to Anthropology Smithsonian Contributions to Botany Smithsonian Contributions to the Earth Sciences Smithsonian Contributions to the Marine Sciences Smithsonian Contributions to Paleobiology Smithsonian Contributions to Zoology Smithsonian Folklife Studies Smithsonian Studies in Air and Space Smithsonian Studies in History and Technology In these series, the Institution publishes small papers and full-scale monographs that report the research and collections of its various museums and bureaux or of professional colleagues in the world of science and scholarship. The publications are distributed by mailing lists to libraries, universities, and similar institutions throughout the world. Papers or monographs submitted for series publication are received by the Smithsonian Institution Press, subject to its own review for format and style, only through departments of the various Smithsonian museums or bureaux, where the manuscripts are given substantive review. Press requirements for manuscript and art preparation are outlined on the inside back cover. I. Michael Heyman Secretary Smithsonian Institution SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • NUMBER 585 Type Specimens of Hispinae (Coleoptera: Chrysomelidae) in the National Museum of Natural History, Smithsonian Institution Charles L. Staines and Susan L. Staines SMITHSONIAN INSTITUTION PRESS Washington, D.C. 1997 ABSTRACT Staines, C.L. and S.L. Staines. Type Specimens of Hispinae (Coleoptera: Chrysomelidae) in the National Museum of Natural History, Smithsonian Institution. Smithsonian Contributions to Zoology, number 585, 25 pages, 1997.—The Hispinae (Coleoptera: Chrysomelidae) type material in the National Museum of Natural History (USNM) is cataloged. Verbatim label data is given for all type specimens. Type specimens that should be present in the USNM but are not also are given. Specimens with type labels that do not meet the requirements of the International Code of Zoological Nomenclature are excluded and are clearly labelled in the collection. The collection contains 42 holotypes and 17 lectotypes. There are allotypes of four species, paratypes of 92 species, paralectotypes of 16 species, and syntypes of 57 species. A lectotype and two paralectotypes are designated for Microrhopala floridana Schwarz. OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded in the Institution's annual report, Annals of the Smithsonian Institution. SERIES COVER DESIGN: The coral Montastrea cavernosa (Linnaeus). Library of Congress Cataloging-in-Publication Data National Museum of Natural History (U.S.) Type specimens of Hispinae (Coleoptera: Chrysomelidae) in the National Museum of Natural History. Smithsonian Institution / Charles L. Staines and Susan L. Staines. p. cm.—(Smithsonian Contributions to Zoology ; no. 585) Includes bibliographical references. I. Chrysomelidae—Type specimens—Catalogs and collections—Washington (D.C.) 2. National Museum of Natural History (U.S.)—Catalogs. I. Staines, Charles L. II. Staines, Susan L. III. Title. IV. Series. QL1.S54 no. 585 [QL596.C5] 590 s—ds 21 [595.76'48] 96-44556 CIP ® The paper used in this publication meets the minimum requirements of the American National Standard for Permanence of Paper for Printed Library Materials Z39.48—1984. Contents Page Introduction 1 Acknowledgments 1 Methods 1 Catalog of Type Specimens 2 Literature Cited 24 in Type Specimens of Hispinae (Coleoptera: Chrysomelidae) in the National Museum of Natural History, Smithsonian Institution Charles L. Staines and Susan L Staines Introduction METHODS While curating the Hispinae (Coleoptera: Chrysomelidae) in In order to verify the status of the specimens we consulted the National Museum Of Natural History, Smithsonian the original descriptions. In most instances the data from the Institution (NMNH), we noticed numerous specimens labelled labels matched that cited in the original description. In holotype or allotype. This was surprising because collection numerous cases the text stated that a type was deposited in policy states that all primary types are to be kept in a separate either the Smithsonian Institution or the USNM (collections of type collection. Secondary types are kept in the general the former United States National Museum, now in the collection. Proper curation of these specimens led to a review National Museum of Natural History, Smithsonian Institution). and verification of all Hispinae types deposited in the national If the label data did not match the data presented in the collections. original description, further investigation was conducted. Eight The NMNH collection contains 42 holotypes and 17 of Gressitt's paratypes had label data differing from the original lectotypes. There are allotypes of four species, paratypes of 92 description. According to Article 72b-vii (International Code of species, paralectotypes of 16 species, and syntypes of 57 Zoological Nomenclature (ICZN)) these eight specimens are species. not types. However, Recommendation 72B (ICZN) states that ACKNOWLEDGMENTS.—We thank R.E. White, USDA, Sys- external evidence may be used in determining whether a tematic Entomology Lab, for access to the collection. P.J. specimen is a type. Samuelson (pers. comm.) stated that Spangler, Smithsonian Institution, and F.C. Thompson and Gressitt did not always record the label data for each type D.R. Miller, Systematic Entomology Lab, assisted in determin- specimen in his publications but that all specimens examined ing the status of various specimens. G.A. Samuelson, Bishop were clearly labelled. The collection at the Bishop Museum has Museum, assisted in determining the status of some Gressitt many Gressitt type specimens that are not cited in the original types. description. We think that this external evidence is sufficient to render the specimens types. Also, the discrepancies are only a Charles L Staines, Collaborating Scientist, and Susan L. Staines, day or two from the original description. Volunteer, Department of Entomology, National Museum of Natural Another major problem was Uhmann's (1950a) lectotype History, Smithsonian Institution, Washington, D.C. 20560. designations (calling them holotypes) of 18 species he Review Chairman: John M. Burns, National Museum of Natural described earlier (Uhmann, 1930a). Uhmann (1950a) did not History, Smithsonian Institution, Washington, D.C. 20560. Reviewers: R. Wills Flowers, Florida Agricultural and Mechanical mention the label data of the specimens designated as University, Tallahassee, Florida 32307; David G. Furth, National lectotypes but simply listed the species and stated the Museum of Natural History, Smithsonian Institution, Washington, specimens were deposited in the USNM. Seven of the 18 D.C. 20560; Edward G. Riley, Texas Agricultural and Mechanical species have label data that does not match the original University, College Station, Texas 77843-2475; Richard E. White, USDA, ARS, SEL, National Museum of Natural History, Smithsonian description, usually an unlisted locality or the collection date Institution, Washington. DC. 20560. several years different than the published one. Uhmann's type SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY labels are hand written or typed red cards with only holotype or Gressitt (1950:125), paratype; Sumitrosis arnetti Butte paratype. Although the invalid specimens are labelled in a (1969:18), paratypes. similar manner to the valid ones, without a definite Uhmann The former F. Nevermann collection came to the Smith- label we cannot be confident that these specimens are the sonian prior to 1950. Nevermann amassed an extensive lectotypes. Another consideration is that the Gaedike and collection of Costa Rican Hispinae. The Nevermann material Dobler (1971) catalog of the Uhmann types in the Deustschen was identified by Uhmann and contains numerous type Entomologischen Institutes cites paralectotypes of Uhmann, specimens. Nevermann used green labels for locality data. Each 1930a, species with the label data mentioned in the original specimen contains two green labels, the upper one with description. country, date, and "F. Nevermann." The second, affixed The final problem is that Nevermann material, although so that the writing is opposite that of the first label present in the NMNH collection, also is present in other and flush with it, contains locality and collector. In the text of collections, so that the stated type series may be housed the catalog the second green label is referred to as "reversed elsewhere. We could find no external evidence to support five green label." designations,
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