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Chapter 5: the Shoot System I: the Stem
Chapter 5 The Shoot System I: The Stem THE FUNCTIONS AND ORGANIZATION OF THE SHOOT SYSTEM PRIMARY GROWTH AND STEM ANATOMY Primary Tissues of Dicot Stems Develop from the Primary Meristems The Distribution of the Primary Vascular Bundles Depends on the Position of Leaves Primary Growth Differs in Monocot and Dicot Stems SECONDARY GROWTH AND THE ANATOMY OF WOOD Secondary Xylem and Phloem Develop from Vascular Cambium Wood Is Composed of Secondary Xylem Gymnosperm Wood Differs from Angiosperm Wood Bark Is Composed of Secondary Phloem and Periderm Buds Are Compressed Branches Waiting to Elongate Some Monocot Stems Have Secondary Growth STEM MODIFICATIONS FOR SPECIAL FUNCTIONS THE ECONOMIC VALUE OF WOODY STEMS SUMMARY ECONOMIC BOTANY: How Do You Make A Barrel? 1 KEY CONCEPTS 1. The shoot system is composed of the stem and its lateral appendages: leaves, buds, and flowers. Leaves are arranged in different patterns (phyllotaxis): alternate, opposite, whorled, and spiral. 2. Stems provide support to the leaves, buds, and flowers. They conduct water and nutrients and produce new cells in meristems (shoot apical meristem, primary and secondary meristems). 3. Dicot stems and monocot stems are usually different. Dicot stems tend to have vascular bundles distributed in a ring, whereas in monocot stems they tend to be scattered. 4. Stems are composed of the following: epidermis, cortex and pith, xylem and phloem, and periderm. 5. Secondary xylem is formed by the division of cells in the vascular cambium and is called wood. The bark is composed of all of the tissues outside the vascular cambium, including the periderm (formed from cork cambium) and the secondary phloem. -
Pollination and Botanic Gardens Contribute to the Next Issue of Roots
Botanic Gardens Conservation International Education Review Volume 17 • Number 1 • May 2020 Pollination and botanic gardens Contribute to the next issue of Roots The next issue of Roots is all about education and technology. As this issue goes to press, most botanic gardens around the world are being impacted by the spread of the coronavirus Covid-19. With many Botanic Gardens Conservation International Education Review Volume 16 • Number 2 • October 2019 Citizen gardens closed to the public, and remote working being required, Science educators are having to find new and innovative ways of connecting with visitors. Technology is playing an ever increasing role in the way that we develop and deliver education within botanic gardens, making this an important time to share new ideas and tools with the community. Have you developed a new and innovative way of engaging your visitors through technology? Are you using technology to engage a Botanic Gardens Conservation International Education Review Volume 17 • Number 1 • April 2020 wider audience with the work of your garden? We are currently looking for a variety of contributions including Pollination articles, education resources and a profile of an inspirational garden and botanic staff member. gardens To contribute, please send a 100 word abstract to [email protected] by 15th June 2020. Due to the global impacts of COVID-19, BGCI’s 7th Global Botanic Gardens Congress is being moved to the Australian spring. Join us in Melbourne, 27 September to 1 October 2021, the perfect time to visit Victoria. Influence and Action: Botanic Gardens as Agents of Change will explore how botanic gardens can play a greater role in shaping our future. -
Listado De Todas Las Plantas Que Tengo Fotografiadas Ordenado Por Familias Según El Sistema APG III (Última Actualización: 2 De Septiembre De 2021)
Listado de todas las plantas que tengo fotografiadas ordenado por familias según el sistema APG III (última actualización: 2 de Septiembre de 2021) GÉNERO Y ESPECIE FAMILIA SUBFAMILIA GÉNERO Y ESPECIE FAMILIA SUBFAMILIA Acanthus hungaricus Acanthaceae Acanthoideae Metarungia longistrobus Acanthaceae Acanthoideae Acanthus mollis Acanthaceae Acanthoideae Odontonema callistachyum Acanthaceae Acanthoideae Acanthus spinosus Acanthaceae Acanthoideae Odontonema cuspidatum Acanthaceae Acanthoideae Aphelandra flava Acanthaceae Acanthoideae Odontonema tubaeforme Acanthaceae Acanthoideae Aphelandra sinclairiana Acanthaceae Acanthoideae Pachystachys lutea Acanthaceae Acanthoideae Aphelandra squarrosa Acanthaceae Acanthoideae Pachystachys spicata Acanthaceae Acanthoideae Asystasia gangetica Acanthaceae Acanthoideae Peristrophe speciosa Acanthaceae Acanthoideae Barleria cristata Acanthaceae Acanthoideae Phaulopsis pulchella Acanthaceae Acanthoideae Barleria obtusa Acanthaceae Acanthoideae Pseuderanthemum carruthersii ‘Rubrum’ Acanthaceae Acanthoideae Barleria repens Acanthaceae Acanthoideae Pseuderanthemum carruthersii var. atropurpureum Acanthaceae Acanthoideae Brillantaisia lamium Acanthaceae Acanthoideae Pseuderanthemum carruthersii var. reticulatum Acanthaceae Acanthoideae Brillantaisia owariensis Acanthaceae Acanthoideae Pseuderanthemum laxiflorum Acanthaceae Acanthoideae Brillantaisia ulugurica Acanthaceae Acanthoideae Pseuderanthemum laxiflorum ‘Purple Dazzler’ Acanthaceae Acanthoideae Crossandra infundibuliformis Acanthaceae Acanthoideae Ruellia -
A Dissertation Submitted to the Graduate Division of the University of Hawai'i in Partial Fulfillment of the Requirements for the Degree Of
FLOWERING IN HELICONIA ROSTRATA RUIZ & PA VON A DISSERTATION SUBMITTED TO THE GRADUATE DIVISION OF THE UNIVERSITY OF HAWAI'I IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY IN HORTICULTURE DECEMBER 2000 BY Norberto Maciel Dissertation Committee: Richard A. Criley, Chairperson Kent D. Kobayashi Robert Pauli Willian S. Sakai David Webb IN MEMORIAM Antonio Oliveira De Sousa (My Father) Because pursuing this goal I did not share his last moments 111 ACKNOWLEDGMENTS I would like to express my sincere gratitude to my chairperson. Dr. Richard A. Criley for inviting me come to the University of Hawaii, his guidance, and understanding. I very much appreciate my other committee members Dr. Kent D. Kobayashi, Dr. Robert Pauli, Dr. William S. Sakai, and Dr. David D. Webb for their assistance and suggestions. Thanks to: Dr. Osamu Kawabata for the suggestions in the statistical analysis; Dr. David D. Webb and Dr. Adelheid Kuehnle for the help with equipment and chemicals; and Mr Bob Hirano and the Lyon Arboretum for providing material of Heliconia rostrata used in one of the experiments. My special thanks to Mr Ronald Matsuda and Craig Okasaki of the Magoon facility for the great help. I want to express my gratitude to faculty, staff and colleagues in the Department of Horticulture for sharing with me their skills, help, and friendship. I will never forget the help and kindness of the friends that I meet in Hawaii, especially for the scholarly help from Derrick Agboka, Renee and Adrian Ares, Douglas Gaskill, Michael Melzer, Javier Mendez, Monica Mejia, Teresa Restom and Mario Serracin. -
2011 Vol. 14, Issue 3
Department of Botany & the U.S. National Herbarium The Plant Press New Series - Vol. 14 - No. 3 July-September 2011 Island Explorations and Evolutionary Investigations By Vinita Gowda or over a century the Caribbean eastward after the Aves Ridge was formed On joining the graduate program region, held between North and to the West. Although the Lesser Antilles at The George Washington University FSouth America, has been an active is commonly referred to as a volcani- in Washington, D.C., in the Fall of area of research for people with interests cally active chain of islands, not all of the 2002, I decided to investigate adapta- in island biogeography, character evolu- Lesser Antilles is volcanic. Based on geo- tion in plant-pollinator interactions tion, speciation, as well as geology. Most logical origin and elevation all the islands using a ‘multi-island’ comparative research have invoked both dispersal and of the Lesser Antilles can be divided into approach using the Caribbean Heliconia- vicariance processes to explain the distri- two groups: a) Limestone Caribbees (outer hummingbird interactions as the study bution of the local flora and fauna, while arc: calcareous islands with a low relief, system. Since I was interested in under- ecological interactions such as niche dating to middle Eocene to Pleistocene), standing factors that could influence partitioning and ecological adaptations and b) Volcanic Caribbees (inner arc: plant-pollinator mutualistic interactions have been used to explain the diversity young volcanic islands with strong relief, between the geographically distinct within the Caribbean region. One of dating back to late Miocene). islands, I chose three strategic islands of the biggest challenges in understanding the Lesser Antilles: St. -
Networks in a Large-Scale Phylogenetic Analysis: Reconstructing Evolutionary History of Asparagales (Lilianae) Based on Four Plastid Genes
Networks in a Large-Scale Phylogenetic Analysis: Reconstructing Evolutionary History of Asparagales (Lilianae) Based on Four Plastid Genes Shichao Chen1., Dong-Kap Kim2., Mark W. Chase3, Joo-Hwan Kim4* 1 College of Life Science and Technology, Tongji University, Shanghai, China, 2 Division of Forest Resource Conservation, Korea National Arboretum, Pocheon, Gyeonggi- do, Korea, 3 Jodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, United Kingdom, 4 Department of Life Science, Gachon University, Seongnam, Gyeonggi-do, Korea Abstract Phylogenetic analysis aims to produce a bifurcating tree, which disregards conflicting signals and displays only those that are present in a large proportion of the data. However, any character (or tree) conflict in a dataset allows the exploration of support for various evolutionary hypotheses. Although data-display network approaches exist, biologists cannot easily and routinely use them to compute rooted phylogenetic networks on real datasets containing hundreds of taxa. Here, we constructed an original neighbour-net for a large dataset of Asparagales to highlight the aspects of the resulting network that will be important for interpreting phylogeny. The analyses were largely conducted with new data collected for the same loci as in previous studies, but from different species accessions and greater sampling in many cases than in published analyses. The network tree summarised the majority data pattern in the characters of plastid sequences before tree building, which largely confirmed the currently recognised phylogenetic relationships. Most conflicting signals are at the base of each group along the Asparagales backbone, which helps us to establish the expectancy and advance our understanding of some difficult taxa relationships and their phylogeny. -
Woody and Herbaceous Plants Native to Haiti for Use in Miami-Dade Landscapes1
Woody and Herbaceous Plants Native to Haiti For use in Miami-Dade Landscapes1 Haiti occupies the western one third of the island of Hispaniola with the Dominican Republic the remainder. Of all the islands within the Caribbean basin Hispaniola possesses the most varied flora after that of Cuba. The plants contained in this review have been recorded as native to Haiti, though some may now have been extirpated due in large part to severe deforestation. Less than 1.5% of the country’s original tree-cover remains. Haiti’s future is critically tied to re- forestation; loss of tree cover has been so profound that exotic fast growing trees, rather than native species, are being used to halt soil erosion and lessen the risk of mudslides. For more information concerning Haiti’s ecological plight consult references at the end of this document. For present purposes all of the trees listed below are native to Haiti, which is why non-natives such as mango (the most widely planted tree) and other important trees such as citrus, kassod tree (Senna siamea) and lead tree (Leucanea leucocephala) are not included. The latter two trees are among the fast growing species used for re-forestation. The Smithsonian National Museum of Natural History’s Flora of the West Indies was an invaluable tool in assessing the range of plants native to Haiti. Not surprisingly many of the listed trees and shrubs 1 John McLaughlin Ph.D. U.F./Miami-Dade County Extension Office, Homestead, FL 33030 Page | 1 are found in other parts of the Caribbean with some also native to South Florida. -
SPECIES IDENTIFICATION GUIDE National Plant Monitoring Scheme SPECIES IDENTIFICATION GUIDE
National Plant Monitoring Scheme SPECIES IDENTIFICATION GUIDE National Plant Monitoring Scheme SPECIES IDENTIFICATION GUIDE Contents White / Cream ................................ 2 Grasses ...................................... 130 Yellow ..........................................33 Rushes ....................................... 138 Red .............................................63 Sedges ....................................... 140 Pink ............................................66 Shrubs / Trees .............................. 148 Blue / Purple .................................83 Wood-rushes ................................ 154 Green / Brown ............................. 106 Indexes Aquatics ..................................... 118 Common name ............................. 155 Clubmosses ................................. 124 Scientific name ............................. 160 Ferns / Horsetails .......................... 125 Appendix .................................... 165 Key Traffic light system WF symbol R A G Species with the symbol G are For those recording at the generally easier to identify; Wildflower Level only. species with the symbol A may be harder to identify and additional information is provided, particularly on illustrations, to support you. Those with the symbol R may be confused with other species. In this instance distinguishing features are provided. Introduction This guide has been produced to help you identify the plants we would like you to record for the National Plant Monitoring Scheme. There is an index at -
Plant–Pollinator Interactions and Floral Convergence in Two Species Of
Oecologia DOI 10.1007/s00442-011-2043-8 PLANT-ANIMAL INTERACTIONS - ORIGINAL PAPER Plant–pollinator interactions and floral convergence in two species of Heliconia from the Caribbean Islands Silvana Marte´n-Rodrı´guez • W. John Kress • Ethan J. Temeles • Elvia Mele´ndez-Ackerman Received: 4 April 2010 / Accepted: 26 May 2011 Ó Springer-Verlag 2011 Abstract Variation in interspecific interactions across nectar chambers and long corollas, whereas on Hispaniola, geographic space is a potential driver of diversification and H. bihai flowers resembled those of H. caribaea with local adaptation. This study quantitatively examined vari- longer nectar chambers and shorter corolla tubes. Mor- ation in floral phenotypes and pollinator service of Heli- phological variation in floral traits corresponded with conia bihai and H. caribaea across three Antillean islands. geographic differences or similarities in the major pollin- The prediction was that floral characters would correspond ators on each island. The Hispaniolan mango, Anthraco- to the major pollinators of these species on each island. thorax dominicus, is the principal pollinator of both Analysis of floral phenotypes revealed convergence among H. bihai and H. caribaea on Hispaniola; thus, the similarity species and populations of Heliconia from the Greater of floral phenotypes between Heliconia species suggests Antilles. All populations of H. caribaea were similar, parallel selective regimes imposed by the principal polli- characterized by long nectar chambers and short corolla nator. Likewise, divergence between H. bihai populations tubes. In contrast, H. bihai populations were strongly from Dominica and Hispaniola corresponded with differ- divergent: on Dominica, H. bihai had flowers with short ences in the pollinators visiting this species on the two islands. -
IN VITRO PROPAGATION of RARE SPECIES Ruscus Aculeatus L. and HISTOLOGICAL PECULIARITIES of the REGENERANTS
Analele Universităţii din Oradea - Fascicula Biologie Tom. XIX, Issue: 1, 2012, pp. 67-73 IN VITRO PROPAGATION OF RARE SPECIES Ruscus aculeatus L. AND HISTOLOGICAL PECULIARITIES OF THE REGENERANTS Cristian BANCIU*, Anca AIFTIMIE-PĂUNESCU* *Institute of Biology, Romanian Academy, Bucharest, Romania Corresponding author: Cristian Banciu, Institute of Biology, Romanian Academy, 296 Splaiul Independentei, 060031 Bucharest, Romania, phone: 0040212239072, fax: 0040212219071, e-mail: [email protected] Abstract. The present study belongs to the international efforts for plant conservation in the areas endangered by human activities. Ruscus aculeatus L. is one of the threatened plants (included in all national red list of vascular plants from Romania) that grow in the Natural Park Comana, Southern Romania. Seedlings and fragments of rhizome, from plants grown in the natural habitat have been used for in vitro plant regeneration and multiplication. After successfully rooting and acclimatization of the regenerated plantlets, histological studies have been performed in order to compare the regenerants from in vitro cultures with plants from natural habitat. The results indicated that this plant species can be multiplicated, rooted and acclimatized on synthetic medium (MS supplemented with NAA, IBA, kinetin and BAP) with a good efficiency and the regenerants develop a only a few structural modification under vitroculture conditions, with no major consequences for a normal physiology and plant acclimatization. Keywords: Ruscus aculeatus L., in vitro culture, morpho-anatomy. INTRODUCTION from botanical gardens or natural habitat. In contrast, studies on tissue cultures to obtain bioactive Genus Ruscus belongs to family Ruscaceae sensu compounds are very few [11, 22]. In Romania stricto [18] and includes approximately 10 species, the pharmacological properties of Ruscus extracts were most known being seven as follows: R. -
A “Short & Easy Trip”- Ortinola River, Gorge & Waterfall
Quarterly Bulletin of the Trinidad and Tobago Field Naturalists’ Club July – September 2019 Issue No: 3/2019 CLUB TRIP, 31ST MARCH 2019 A “SHORT & EASY TRIP”- ORTINOLA RIVER, GORGE & WATERFALL by Linton Arneaud and Sarah Evelyn Naturalists being brief by Dan Jaggernauth on the rich biodiversity along a perennial stream. Photo by Linton Arneaud Forty–five enthusiastic naturalists by it. Not too long after, we marched off along the gathered at the foot of the St. Joseph Valley Ortinola trail entering the Lower Montane at 7:23 am for the start of what was described Rainforest of the Northern Range and immediately as a ‘short and easy club trip’. As always, Dan started observing the rich biodiversity in this unique Jaggernauth did not hesitate in getting the logistics environment: a mixture of flora and fauna species out of the way; he was particularly anxious to point from the South American continent and northern out that the conspicuous pyramidal-shape ridge Caribbean islands. behind us was not Mt. El Tucuche (Trinidad’s It was quite challenging to keep up with the second highest peak) and we should not be fooled (Continued on page 3) Page 2 THE FIELD NATURALIST Issue No. 3/2019 Inside This Issue Club Trip Report 1 ORTINOLA RIVER, GORGE AND WATERFALLS Quarterly Bulletin of the - Linton Arneaud and Sarah Evelyn Trinidad and Tobago Field Naturalists’ Club July – September 2019 General Club Trip Report Editors 8 MORNE LA CROIX - Kris Sookdeo Laura Baboolal, Stephanie Warren-Gittens Associate Editor: Rupert Mends General Club Trip Report Contributing writers ARENA FOREST 11 Linton Arneaud, Sarah Evelyn, Kris Sookdeo, Delezia Singh, - Christopher K. -
The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development
The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development A thesis presented to the faculty of the College of Arts and Sciences of Ohio University In partial fulfillment of the requirements for the degree Master of Science Alastair R. Plant August 2013 © 2013 Alastair R. Plant. All Rights Reserved. 2 This thesis titled The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development by ALASTAIR R PLANT has been approved for the Department of Environmental and Plant Biology and the College of Arts and Sciences by Stefan Gleissberg Assistant Professor of Environmental and Plant Biology Robert Frank Dean, College of Arts and Sciences 3 ABSTRACT PLANT, ALASTAIR R., M.S., August 2013, Plant Biology The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development Director of Thesis: Stefan Gleissberg The leaf is almost ubiquitous throughout land plants but due to its complex and flexible developmental program is highly morphologically variable between taxa. Description of the functions of regulatory genes key to leaf development in different evolutionary lineages allows the study of changes in developmental mechanisms through evolutionary time as a means for anatomical and morphological diversification. The roles of homologs of CINCINNATA-like TCP family genes, ARP genes, and Class I KNOX genes were investigated in two members of the Papaveraceae, a basal eudicot lineage positioned in between major angiosperm groups, by phylogenetic analysis, in situ hybridization, expression profiling by quantitative polymerase chain reaction, and virus- induced gene silencing in Eschscholzia californica and Cysticapnos vesicaria. Expression data were similar to those for homologous genes in core eudicot species, however, some gene functions found in core eudicots were not associated with basal eudicot homologs, and so have either been gained or lost from the ancestral state.