Download Download
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Fused and Vaulted Nasals of Tyrannosaurid Dinosaurs: Implications for Cranial Strength and Feeding Mechanics
Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics ERIC SNIVELY, DONALD M. HENDERSON, and DOUG S. PHILLIPS Snively, E., Henderson, D.M., and Phillips, D.S. 2006. Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics. Acta Palaeontologica Polonica 51 (3): 435–454. Tyrannosaurid theropods display several unusual adaptations of the skulls and teeth. Their nasals are fused and vaulted, suggesting that these elements braced the cranium against high feeding forces. Exceptionally high strengths of maxillary teeth in Tyrannosaurus rex indicate that it could exert relatively greater feeding forces than other tyrannosaurids. Areas and second moments of area of the nasals, calculated from CT cross−sections, show higher nasal strengths for large tyrannosaurids than for Allosaurus fragilis. Cross−sectional geometry of theropod crania reveals high second moments of area in tyrannosaurids, with resulting high strengths in bending and torsion, when compared with the crania of similarly sized theropods. In tyrannosaurids trends of strength increase are positively allomeric and have similar allometric expo− nents, indicating correlated progression towards unusually high strengths of the feeding apparatus. Fused, arched nasals and broad crania of tyrannosaurids are consistent with deep bites that impacted bone and powerful lateral movements of the head for dismembering prey. Key words: Theropoda, Carnosauria, Tyrannosauridae, biomechanics, feeding mechanics, computer modeling, com− puted tomography. Eric Snively [[email protected]], Department of Biological Sciences, University of Calgary, 2500 University Drive NW, Calgary, Alberta T2N 1N4, Canada; Donald M. Henderson [[email protected]], Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, Alberta T0J 0Y0, Canada; Doug S. -
A Comprehensive Anatomical And
Journal of Paleontology, Volume 94, Memoir 78, 2020, p. 1–103 Copyright © 2020, The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/ licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/20/1937-2337 doi: 10.1017/jpa.2020.14 A comprehensive anatomical and phylogenetic evaluation of Dilophosaurus wetherilli (Dinosauria, Theropoda) with descriptions of new specimens from the Kayenta Formation of northern Arizona Adam D. Marsh1,2 and Timothy B. Rowe1 1Jackson School of Geosciences, the University of Texas at Austin, 2305 Speedway Stop C1160, Austin, Texas 78712, USA <[email protected]><[email protected]> 2Division of Resource Management, Petrified Forest National Park, 1 Park Road #2217, Petrified Forest, Arizona 86028, USA Abstract.—Dilophosaurus wetherilli was the largest animal known to have lived on land in North America during the Early Jurassic. Despite its charismatic presence in pop culture and dinosaurian phylogenetic analyses, major aspects of the skeletal anatomy, taxonomy, ontogeny, and evolutionary relationships of this dinosaur remain unknown. Skeletons of this species were collected from the middle and lower part of the Kayenta Formation in the Navajo Nation in northern Arizona. Redescription of the holotype, referred, and previously undescribed specimens of Dilophosaurus wetherilli supports the existence of a single species of crested, large-bodied theropod in the Kayenta Formation. The parasagittal nasolacrimal crests are uniquely constructed by a small ridge on the nasal process of the premaxilla, dorsoventrally expanded nasal, and tall lacrimal that includes a posterior process behind the eye. -
Tetrapod Biostratigraphy and Biochronology of the Triassic–Jurassic Transition on the Southern Colorado Plateau, USA
Palaeogeography, Palaeoclimatology, Palaeoecology 244 (2007) 242–256 www.elsevier.com/locate/palaeo Tetrapod biostratigraphy and biochronology of the Triassic–Jurassic transition on the southern Colorado Plateau, USA Spencer G. Lucas a,⁎, Lawrence H. Tanner b a New Mexico Museum of Natural History, 1801 Mountain Rd. N.W., Albuquerque, NM 87104-1375, USA b Department of Biology, Le Moyne College, 1419 Salt Springs Road, Syracuse, NY 13214, USA Received 15 March 2006; accepted 20 June 2006 Abstract Nonmarine fluvial, eolian and lacustrine strata of the Chinle and Glen Canyon groups on the southern Colorado Plateau preserve tetrapod body fossils and footprints that are one of the world's most extensive tetrapod fossil records across the Triassic– Jurassic boundary. We organize these tetrapod fossils into five, time-successive biostratigraphic assemblages (in ascending order, Owl Rock, Rock Point, Dinosaur Canyon, Whitmore Point and Kayenta) that we assign to the (ascending order) Revueltian, Apachean, Wassonian and Dawan land-vertebrate faunachrons (LVF). In doing so, we redefine the Wassonian and the Dawan LVFs. The Apachean–Wassonian boundary approximates the Triassic–Jurassic boundary. This tetrapod biostratigraphy and biochronology of the Triassic–Jurassic transition on the southern Colorado Plateau confirms that crurotarsan extinction closely corresponds to the end of the Triassic, and that a dramatic increase in dinosaur diversity, abundance and body size preceded the end of the Triassic. © 2006 Elsevier B.V. All rights reserved. Keywords: Triassic–Jurassic boundary; Colorado Plateau; Chinle Group; Glen Canyon Group; Tetrapod 1. Introduction 190 Ma. On the southern Colorado Plateau, the Triassic– Jurassic transition was a time of significant changes in the The Four Corners (common boundary of Utah, composition of the terrestrial vertebrate (tetrapod) fauna. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Jurassic Algae of the Perachora-Peninsula: Biostratigraphical and Paleoecological Implications
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Beiträge zur Paläontologie Jahr/Year: 1994 Band/Volume: 19 Autor(en)/Author(s): Dragastan Ovidiu, Gielisch Hartwig, Richter Detlef K., Grewer Till, Kaziur Thomas, Kube Bärbel, Radusch Christoph Artikel/Article: Jurassic algae of the Perachora-Peninsula: Biostratigraphical and paleoecological implications 49-81 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien Beitr. Paläont., 19:49-81, Wien 1994 Jurassic algae of the Perachora-Peninsula: Biostratigraphical and paleoecological implications Jurassische Algen der Perachora Halbinsel: Biostratigraphische und paläoökologische Folgerungen by DRAGASTAN, Ovidiu,* GIELISCH, Hartwig**, RICHTER, Detlev K.**, GREWER, TiU**, KAZIUR, Thomas**, KUBE, Bärbel **& RADUSCH, Christoph** DRAGASTAN, O., GIELISCH, H„ RICHTER, D.K., GREWER, T„ KAZIUR, T„ KUBE, B. & RADUSCH, C., 1994. Jurassic algae of the Perachora-Peninsula: Biostratigraphical and paleoecological implications. — Beitr. Palaont., 19:49-81, 9 Figures, 6 Plates, Wien. ceae), Rivularia (Rivulariaceae), Alpinelia graeca n.sp. Contents (Scytonemataceae), Girvanella, Hedstroemia (Porostro- Abstract, Zusammenfassung.................................................. 49 mata) and Microproblematicae. 1. Introduction..............................................................................50 Biostratigraphically, a succession of algae and foramini- 2. Geological setting................................................................. -
The Sauropodomorph Biostratigraphy of the Elliot Formation of Southern Africa: Tracking the Evolution of Sauropodomorpha Across the Triassic–Jurassic Boundary
Editors' choice The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary BLAIR W. MCPHEE, EMESE M. BORDY, LARA SCISCIO, and JONAH N. CHOINIERE McPhee, B.W., Bordy, E.M., Sciscio, L., and Choiniere, J.N. 2017. The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary. Acta Palaeontologica Polonica 62 (3): 441–465. The latest Triassic is notable for coinciding with the dramatic decline of many previously dominant groups, followed by the rapid radiation of Dinosauria in the Early Jurassic. Among the most common terrestrial vertebrates from this time, sauropodomorph dinosaurs provide an important insight into the changing dynamics of the biota across the Triassic–Jurassic boundary. The Elliot Formation of South Africa and Lesotho preserves the richest assemblage of sauropodomorphs known from this age, and is a key index assemblage for biostratigraphic correlations with other simi- larly-aged global terrestrial deposits. Past assessments of Elliot Formation biostratigraphy were hampered by an overly simplistic biozonation scheme which divided it into a lower “Euskelosaurus” Range Zone and an upper Massospondylus Range Zone. Here we revise the zonation of the Elliot Formation by: (i) synthesizing the last three decades’ worth of fossil discoveries, taxonomic revision, and lithostratigraphic investigation; and (ii) systematically reappraising the strati- graphic provenance of important fossil locations. We then use our revised stratigraphic information in conjunction with phylogenetic character data to assess morphological disparity between Late Triassic and Early Jurassic sauropodomorph taxa. Our results demonstrate that the Early Jurassic upper Elliot Formation is considerably more taxonomically and morphologically diverse than previously thought. -
Perinate and Eggs of a Giant Caenagnathid Dinosaur from the Late Cretaceous of Central China
ARTICLE Received 29 Jul 2016 | Accepted 15 Feb 2017 | Published 9 May 2017 DOI: 10.1038/ncomms14952 OPEN Perinate and eggs of a giant caenagnathid dinosaur from the Late Cretaceous of central China Hanyong Pu1, Darla K. Zelenitsky2, Junchang Lu¨3, Philip J. Currie4, Kenneth Carpenter5,LiXu1, Eva B. Koppelhus4, Songhai Jia1, Le Xiao1, Huali Chuang1, Tianran Li1, Martin Kundra´t6 & Caizhi Shen3 The abundance of dinosaur eggs in Upper Cretaceous strata of Henan Province, China led to the collection and export of countless such fossils. One of these specimens, recently repatriated to China, is a partial clutch of large dinosaur eggs (Macroelongatoolithus) with a closely associated small theropod skeleton. Here we identify the specimen as an embryo and eggs of a new, large caenagnathid oviraptorosaur, Beibeilong sinensis. This specimen is the first known association between skeletal remains and eggs of caenagnathids. Caenagnathids and oviraptorids share similarities in their eggs and clutches, although the eggs of Beibeilong are significantly larger than those of oviraptorids and indicate an adult body size comparable to a gigantic caenagnathid. An abundance of Macroelongatoolithus eggs reported from Asia and North America contrasts with the dearth of giant caenagnathid skeletal remains. Regardless, the large caenagnathid-Macroelongatoolithus association revealed here suggests these dinosaurs were relatively common during the early Late Cretaceous. 1 Henan Geological Museum, Zhengzhou 450016, China. 2 Department of Geoscience, University of Calgary, Calgary, Alberta, Canada T2N 1N4. 3 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China. 4 Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada T6G 2E9. 5 Prehistoric Museum, Utah State University, 155 East Main Street, Price, Utah 84501, USA. -
Abstract Book.Pdf
Welcome! Welcome to the VI Symposium on Dinosaur Eggs and Babies, the return of this periodic gathering to the Iberian Peninsula, when it hatched eighteen years ago. From the slopes of the Pyrenees, we have followed the first steps of dinosaurs through France, Argentina, the United States and China. Today, we come back and see the coast where the first theropod embryos were discovered twenty years ago. Since the end of the last century, Paleoology, much like other branches of palaeontology, has evolved thanks to the advance of new methodologies and analytical tools, becoming a progressively more interdisciplinary area of knowledge. Dinosaur babies and embryos, rare findings back when these meetings started, seem to be everywhere now that we learn to look for them under the light of the microscope. New astonishing specimens allow us to understand how Mesozoic dinosaurs mate and reproduce. Oology, our parent discipline in the modern world, has made great advances in understanding the form and function of the egg, and its applications on poultry industry are countless. More than thirty contributions evidence that our field remains small but alive and healthy. We hope that you find in this Symposium an opportunity to share knowledge and open new lines of collaboration. And do not forget to enjoy your stay in Portugal. The host committee CONTENTS How to get to the FCT 6 Acknowledgements 10 PROGRAM 11 ABSTRACTS 14 THE FIRST ORNITHOMIMID EMBRYO IN A SHELL WITH A SINGLE STRUCTURAL LAYER: A CHALLENGE TO ORTHODOXY 15 Araújo R., Lamb J., Atkinson P., Martins R. M. S., Polcyn M.J., Fernandez V. -
Pterosaur Distribution in Time and Space: an Atlas 61
Zitteliana An International Journal of Palaeontology and Geobiology Series B/Reihe B Abhandlungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie B28 DAVID W. E. HONE & ERIC BUFFETAUT (Eds) Flugsaurier: pterosaur papers in honour of Peter Wellnhofer CONTENTS/INHALT Dedication 3 PETER WELLNHOFER A short history of pterosaur research 7 KEVIN PADIAN Were pterosaur ancestors bipedal or quadrupedal?: Morphometric, functional, and phylogenetic considerations 21 DAVID W. E. HONE & MICHAEL J. BENTON Contrasting supertree and total-evidence methods: the origin of the pterosaurs 35 PAUL M. BARRETT, RICHARD J. BUTLER, NICHOLAS P. EDWARDS & ANDREW R. MILNER Pterosaur distribution in time and space: an atlas 61 LORNA STEEL The palaeohistology of pterosaur bone: an overview 109 S. CHRISTOPHER BENNETT Morphological evolution of the wing of pterosaurs: myology and function 127 MARK P. WITTON A new approach to determining pterosaur body mass and its implications for pterosaur fl ight 143 MICHAEL B. HABIB Comparative evidence for quadrupedal launch in pterosaurs 159 ROSS A. ELGIN, CARLOS A. GRAU, COLIN PALMER, DAVID W. E. HONE, DOUGLAS GREENWELL & MICHAEL J. BENTON Aerodynamic characters of the cranial crest in Pteranodon 167 DAVID M. MARTILL & MARK P. WITTON Catastrophic failure in a pterosaur skull from the Cretaceous Santana Formation of Brazil 175 MARTIN LOCKLEY, JERALD D. HARRIS & LAURA MITCHELL A global overview of pterosaur ichnology: tracksite distribution in space and time 185 DAVID M. UNWIN & D. CHARLES DEEMING Pterosaur eggshell structure and its implications for pterosaur reproductive biology 199 DAVID M. MARTILL, MARK P. WITTON & ANDREW GALE Possible azhdarchoid pterosaur remains from the Coniacian (Late Cretaceous) of England 209 TAISSA RODRIGUES & ALEXANDER W. -
Lower Cretaceous Avian-Dominated, Theropod
Lower cretaceous avian-dominated, theropod, thyreophoran, pterosaur and turtle track assemblages from the Tugulu Group, Xinjiang, China: ichnotaxonomy and palaeoecology Lida Xing1,2, Martin G. Lockley3, Chengkai Jia4, Hendrik Klein5, Kecheng Niu6, Lijun Zhang7, Liqi Qi8, Chunyong Chou2, Anthony Romilio9, Donghao Wang2, Yu Zhang2, W Scott Persons10 and Miaoyan Wang2 1 State Key Laboratory of Biogeology and Environmental Geology, China University of Geoscience (Beijing), Beijing, China 2 School of the Earth Sciences and Resources, China University of Geoscience (Beijing), Beijing, China 3 Dinosaur Trackers Research Group, University of Colorado at Denver, Denver, United States 4 Research Institute of Experiment and Detection of Xinjiang Oil Company, PetroChina, Karamay, China 5 Saurierwelt Paläontologisches Museum, Neumarkt, Germany 6 Yingliang Stone Natural History Museum, Nan’an, China 7 Institute of Resources and Environment, Key Laboratory of Biogenic Traces & Sedimentary Minerals of Henan Province, Collaborative Innovation Center of Coalbed Methane and Shale Gas for Central Plains Economic Region, Henan Polytechnic University, Jiaozuo, China 8 Faculty of Petroleum, China University of Petroleum (Beijing) at Karamay, Karamay, China 9 School of Biological Sciences, The University of Queensland, Brisbane, Australia 10 Mace Brown Museum of Natural History, Department of Geology and Environmental Geosciences, College of Charleston, Charleston, United States ABSTRACT Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source Submitted 10 January 2021 of some of the largest Lower Cretaceous track collections from China. They originate Accepted 26 April 2021 from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, 28 May 2021 Published variously divided into four formations and subgroups in the northwestern margin of Corresponding author the Junggar Basin. -
The Fauna from the Tyrannosaurus Rex Excavation, Frenchman Formation (Late Maastrichtian), Saskatchewan
The Fauna from the Tyrannosaurus rex Excavation, Frenchman Formation (Late Maastrichtian), Saskatchewan Tim T. Tokaryk 1 and Harold N. Bryant 2 Tokaryk, T.T. and Bryant, H.N. (2004): The fauna from the Tyrannosaurus rex excavation, Frenchman Formation (Late Maastrichtian), Saskatchewan; in Summary of Investigations 2004, Volume 1, Saskatchewan Geological Survey, Sask. Industry Resources, Misc. Rep. 2004-4.1, CD-ROM, Paper A-18, 12p. Abstract The quarry that contained the partial skeleton of the Tyrannosaurus rex, familiarly known as “Scotty,” has yielded a diverse faunal and floral assemblage. The site is located in the Frenchman River valley in southwestern Saskatchewan and dates from approximately 65 million years, at the end of the Cretaceous Period. The faunal assemblage from the quarry is reviewed and the floral assemblage is summarized. Together, these assemblages provide some insight into the biological community that lived in southwestern Saskatchewan during the latest Cretaceous. Keywords: Frenchman Formation, Maastrichtian, Late Cretaceous, southwestern Saskatchewan, Tyrannosaurus rex. 1. Introduction a) Geological Setting The Frenchman Formation, of latest Maastrichtian age, is extensively exposed in southwestern Saskatchewan (Figure 1; Fraser et al., 1935; Furnival, 1950). The lithostratigraphic units in the formation consist largely of fluvial sandstones and greenish grey to green claystones. Outcrops of the Frenchman Formation are widely distributed in the Frenchman River valley, southeast of Eastend. Chambery Coulee, on the north side of the valley, includes Royal Saskatchewan Museum (RSM) locality 72F07-0022 (precise locality data on file with the RSM), the site that contained the disarticulated skeleton of a Tyrannosaurus rex. McIver (2002) subdivided the stratigraphic sequence at this locality into “lower” and “upper” beds. -
South-East Asia Second Edition CHARLES S
Geological Evolution of South-East Asia Second Edition CHARLES S. HUTCHISON Geological Society of Malaysia 2007 Geological Evolution of South-east Asia Second edition CHARLES S. HUTCHISON Professor emeritus, Department of geology University of Malaya Geological Society of Malaysia 2007 Geological Society of Malaysia Department of Geology University of Malaya 50603 Kuala Lumpur Malaysia All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior permission of the Geological Society of Malaysia ©Charles S. Hutchison 1989 First published by Oxford University Press 1989 This edition published with the permission of Oxford University Press 1996 ISBN 978-983-99102-5-4 Printed in Malaysia by Art Printing Works Sdn. Bhd. This book is dedicated to the former professors at the University of Malaya. It is my privilege to have collabo rated with Professors C. S. Pichamuthu, T. H. F. Klompe, N. S. Haile, K. F. G. Hosking and P. H. Stauffer. Their teaching and publications laid the foundations for our present understanding of the geology of this complex region. I also salute D. ]. Gobbett for having the foresight to establish the Geological Society of Malaysia and Professor Robert Hall for his ongoing fascination with this region. Preface to this edition The original edition of this book was published by known throughout the region of South-east Asia. Oxford University Press in 1989 as number 13 of the Unfortunately the stock has become depleted in 2007. Oxford monographs on geology and geophysics.