AMERICAN MUSEUM Novitates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 3137, 11 pp. May 26, 1995

Homonymy, Synonymy, and New Combinations in the ()

I. M. KERZHNERI AND R. T. SCHUH2

ABSTRACT Replacement names are proposed for newly dis- signments, and some species are transferred to oth- covered homonyms. Several genera are placed in er genera. The type species ofBrasifiocarnus Car- synonymy, some genera are given new tribal as- valho and Cleotomiris Schuh are fixed. INTRODUCTION J. C. M. Carvalho, in his monumental world names, new synonymy, and revised subfam- catalog ofMiridae (1957-1960), noticed many ily and tribal assignments are also proposed homonymous species-group names and re- for taxa published subsequent to the Carval- placed the junior homonyms by junior syn- ho Catalogue. onyms or new replacement names. However, in some cases such corrections were not made or the earliest names were not used for re- REPLACEMENT GENUS-GROUP placement. Some homonyms not replaced by NAMES Carvalho were placed in synonymy or re- Cearamiris nom. n. is proposed for Cear- placed in later works. Here we attempt an ana Carvalho and Ferreira, 1974 [Rev. Bras- overview of the remaining cases not dealt ileira Biol. 33 (suppl.): 133], a junior hom- with by Carvalho. Two new patronymics are onym of Cearana Jordan and Brauner, 1908 proposed for the late Dr. Carvalho in rec- [Smithson. Misc. Collect. 52(1792): 27], Pi- ognition of his outstanding contributions to sces. Masculine. of the Miridae. Replacement Candidomiris nom. n. is proposed for

' Department of Taxonomy, Zoological Institute, Academy of Sciences, St. Petersburg, Russia 199034. 2Curator, Department of Entomology, American Museum of Natural History; Adjunct Professor, City College, City University ofNew York; Adjunct Professor, Department ofEntomology, Cornell University, Ithaca, New York.

Copyright © American Museum of Natural History 1995 ISSN 0003-0082 / Price $1.40 2 AMERICAN MUSEUM NOVITATES NO. 3137

Kraussella Carvalho, 1987 [Rev. Brasileira = Deraeocoris cordiger (Hahn, 1834), Pale- Biol. 47: 180], a junior homonym of Kraus- arctic. sella Bolivar, 1909 [Bol. Soc. Esp. Hist. Nat. Deraeocoris brunneolus nom. n. is pro- 9: 292], Orthoptera. Masculine. posed for D. brunneus (Poppius, 1915), Cim- Josifovolygus nom. n. is proposed for Tri- icicapsus brunneus Poppius, 1915 [Ann. Mus. cholygus Josifov, 1992 [Reichenbachia 29: Nati. Hung. 13: 84] from Australia, a junior 108], ajunior homonym of Tricholygus Pop- secondary homonym of D. brunneus Pop- pius, 1910 [Swedish Zool. Exped. Kilimand- pius, 1912 [Acta Soc. Sci. Fenn. 41(3): 120, jaro 2:47], Miridae. Masculine. 126], Madagascar. Deraeocoris darjeelingensis nom. n. is pro- REPLACEMENT SPECIES-GROUP posed for D. discoidalis Poppius, 1915 [Ann. NAMES Mus. Natl. Hungarici 13: 80] from India, a junior secondary homonym ofLamprolygus fijiensis nom. n. is proposed signatus var. discoidalis Poppius, 1912 [Acta for A. lateralis Reuter, 1908 [Ann. Naturhist. Soc. Sci. Fenn. 41(3): 113] = Deraeocoris dis- Mus. Wien 22: 184] from Fiji Is., a junior coidalis (Poppius, 1912), tropical Africa. secondary homonym of Phytocoris lateralis Deraeocoris discoidalis (Poppius, 1912), Fall6n, 1829 [Hem. Suec.: 88] = Adelphocoris stat. n., Lamprolygus signatus var. discoidalis seticornis (Fabricius, 1775), Palearctic. Poppius, 1912 [Acta Soc. Sci. Fenn. 41(3): Adelphocoris sichuanus nom. n. is pro- 113] is the valid name forD. signatoides Car- posed for A. apicalis Reuter, 1906 [Annu. valho, 1957 [Arq. Mus. Nac. 44: 80] from Mus. Zool. St. Petersburg 10: 9, 13] from tropical Africa. Note: In renaming the pre- China, a junior secondary homonym ofPhy- occupied D. signatus (Poppius, 1912), Car- tocoris apicalis Hahn, 1833 [Wanz. Ins. 1: valho (1957: 80) gave it the new name D. 220] = Adelphocoris seticornis (Fabrncius, signatoides instead ofusing the varietal name 1775), Palearctic. discoidalis. Calocoris poppiusi nom. n. is proposed for Halticus spegazzinii Berg, 1883 [An. Soc. Calocoris caucasicus Poppius, 1912, Creon- Cienc. Argentina 9: 74] is the valid name for tiades cuacasicus Poppius, 1912 [ Ofv. Finsk. H. pygmaeus (Berg, 1879), Capsuspygmaeus Vet. Soc. Forh. 54A(29): 6], a junior second- Berg, 1879 [Hem. Argentina, p. 290] from ary homonym of Calocoris caucasicus Pop- Argentina, a junior primary homonym of C. pius, 1912 [Ofv. Finsk. Vet. Soc. Forh. pygmaeus Zetterstedt, 1838 [Ins. Lapp., p. 54A(29): 7], both species from the Caucasus. 279] = Tytthuspygmaeus (Zetterstedt, 1838), Campylomma nigrifemur Wagner, 1975 Palearctic. [Not. Entomol. 61: 11] is the valid name for Heterocordylus italicus nom. n. is proposed C. pallida (Wagner, 1957), Sthenaropsis pal- forH.flavipes Wagner, 1953 [Ann. Naturhist. lida Wagner, 1957 [Jahresh. Ver. Vaterl. Na- Mus. Wien 59: 296] from Italy, a junior pri- turkd. Wurttemberg 112: 94] from the south- mary homonym of H. flavipes Nitobe, 1906 ern Palearctic, a junior secondary homonym [Insect World, Gifu, 10: 19], Japan, nomen ofCampylommapallida Usinger, 1946 [Bull. dubium. Note. In Carvalho (1958) flavipes B. P. Bishop Mus. 189: 89], from Guam. Nitobe is given asflaviceps [sic] Matsumura. Campylomma odhiamboi nom. n. is pro- Hyalopeplus lineifer (Walker, 1873), Cap- posed for Campylomma punctipes (Odhiam- sus lineifer Walker, 1873 [Cat. Heteroptera bo, 1959) Sthenarus punctipes Odhiambo, Br. Mus. 6: 122] is the valid name for H. 1959 [Ann. Mag. Nat. Hist. (13)2: 435], vitripennis (StAl, 1855), vitripennis Uganda, a junior secondary homonym of StAl, 1855 [Ofv. Finsk. Vet. Soc. Forh.,,12(4): Capsus punctipes Herrich-Schaeffer, 1835 186] from Oriental and Australian Regions, [Nomencl. Entomol. 1: 1501 = ? Campylom- a junior primary homonym of Capsus vitri- ma verbasci (Meyer-Dur, 1843), Palearctic. pennis Say, 1832 [Descript. Het. Hem. N. Deraeocoris apicatus nom. n. is proposed Am.: 24] = vitripennis (Say, 1832), for D. apicalis Poppius, 1915 [Arch. Naturg. North America. 80A(8): 40], Taiwan, a junior secondary sinicus nom. n. is proposed for homonym of Capsus apicalis Signoret, 1865 Lygocoris (Apolygus) pubescens Zheng and [Ann. Soc. Entomol. France, ser. 4, 5: 125] Wang, 1983, Lygus (Apolygus) pubescens 1995 KERZHNER AND SCHUH: MIRIDAE NOMENCLATURAL CHANGES 3

Zheng and Wang, 1983 [Entomotaxonomia homonym of Orthotylus elongatus Wagner, 5:48,57], from China, ajunior primary hom- 1965 [Reichenbachia 6: 67], from Cyprus. onym of Lygus pratensis pubescens Reuter, Orthotylus subtropicalis nom. n. is pro- 1912 [Ofv. Finsk. Vet. Soc. Forh. 54A(7): posed for Orthotylus missionensis (Carvalho 37], = Lygus rugulipennis Poppius, 1911, and Carpintero, 1986), Melanotrichus mis- from the Palearctic. sionensis Carvalho and Carpintero, 1986 Lygocoris wangi nom. n. is proposed for [Rev. Brasileira Biol. 46: 619], a junior sec- Lygocoris (Apolygus) signatus Zheng and ondary homonym ofOrthotylus missionensis Wang, 1983, Lygus (Apolygus) signatus Zheng Carvalho 1985 [Rev. Brasileira Biol. 45: 280], and Wang, 1983 [Entomotaxonomia 5: 48, both from Argentina. 56], from China, ajunior primary homonym Orthotylus josei nom. n. is proposed for of Lygus [now ] cervinus form sig- Orthotylus cuneatus Carvalho, 1985 [Rev. nata Stichel, 1930 [Illstr. Bestimmungsta- Brasileira Biol. 45: 277], from Argentina, ju- bellen Deut. Wanzen 6,7: 181, from Europe. nior primary homonym ofOrthotylus cunea- Lygocoris zhengi nom. n. is proposed for tus Van Duzee, 1916 [Proc. California Acad. Lygocoris (Apolygus) marginatus Zheng and Sci. (4)6: 117], from California. Wang, 1983, Lygus (Apolygus) margintus Phoenicocoris knighti nom. n. is proposed Zheng and Wang, 1983 [Entomotaxonomia for Phoenicocoris pini Knight, 1968, Lepi- 5: 48, 56], from China, a junior secondary dopsalluspini Knight, 1968 [Brigham Young homonym of Phytocoris marginatus Zetter- Univ. Sci. Bull., Biol. Ser., 9: 53; see Stone- stedt, 1838 [Insecta Lapponica descripta 1: dahl, 1990, Bull. Amer. Mus. Nat. Hist. 198: 272], = (Fallen, 1807), 14], from western USA, a junior secondary from the Palearctic. homonym ofAtractotomus pini Fieber, 1864 Orthotylus costai nom. n. is proposed for [Wien. Entomol. Monatschr. 8: 224], = Orthotylus clarensis (Carvalho, 1990), Me- Phoenicocoris obscurellus (Fallen, 1829), from lanotrichus clarensis Carvalho, 1990 [An. the Palearctic. Acad. Brail. Cienc. 62: 395], a junior sec- Phytocoris femoratus nom. n. is proposed ondary homonym of Orthotylus clarensis for P.femoralis Fieber, 1861 [Eur. Hem.: 260] Carvalho and Costa, 1990 [An. Acad. Brasi. from the Mediterranean, a junior primary Cienc. 62: 310], both from Brazil [see dis- homonym ofP.femoralis Lucas, 1849 [Expl. cussion of Orthotylus under New Combina- Alg.: 82] = Calocoris fulvomaculatus (De tions for this and other species ofOrthotylus]. Geer, 1773), Palearctic. Orthotylus harryi nom. n. is proposed for Phytocorisjorigtooi nom. n. is proposed for Orthotylus pallens (Knight, 1968), Melano- Phytocoris desertorum Nonnaizab and Jorig- trichuspallens Knight, 1968 [Brigham Young too, 1992 [Zool. Res. 13: 318, 322], from Univ. Sci. Biol. China, a primary homonym of Phytocoris Bull., Ser., 11(3): 125], from dersertorum Reuter, 1900 [Ofv. Finsk. Vet. western USA, a junior secondary homonym Soc. Forh. B 42: 251], from North Africa. of Orthotylus pallens (Matsumura, 1911), Phytocoris leucopterus nom. n. is proposed Calocoris pallens Matsumura, 1911 [J. Coll. for P. albipennis Reuter, 1904 [Ofv. Finsk. Agric. Tokyo Imp. Univ., Sapporo 4: 39], Vet. Soc. Forh., 47(4): 3] from Israel, ajunior from Sakhalin Island. primary homonym of P. albipennis Fallen, Orthotylus hazeltoni nom. n. is proposed 1829 [Hem. Suec., p. 107] = Plagiognathus for Orthotylus uniformis (Knight, 1968), Me- albipennis (Fallen, 1829), Holarctic. lanotrichus uniformis Knight, 1968 [Brigham Phytocoris marmoratus Blanchard, 1852, Young Univ. Sci. Bull., Biol. Ser., 11(3): 128], in Gay [Hist. Fis. Polit. Chile, 7: 194] is the from western USA, ajunior secondary hom- valid name for P. chilensis Carvalho, 1955 onym of Orthotylus uniformis Van Duzee, [Rev. Chilena Entomol. 4: 225] from Chile. 1916 [Proc. California Acad. Sci., 4th ser., Note: In renaming the preoccupied P. ad- 6(5): 100], from California. spersus Spinola, 1852, Carvalho (1955) gave Orthotylus leonardi nom. n. is proposed for it a new name P. chilensis, instead of using Orthotylus elongatus (Kelton, 1980) Melan- the synonymic name P. marmoratus. otrichus elongatus, Kelton [Can. Entomol. Phytocoris nonnaizabi nom. n. is proposed 112: 338]., from Canada, a junior secondary for Phytocoris procerus Nonnaizab and Jo- 4 AMERICAN MUSEUM NOVITATES NO. 3137

rigtoo, 1992 [Zool. Res. 13: 314, 321], from Capsus punctipes Mulsant et Rey, 1852 China, a junior primary homonym of Phy- [Ann. Soc. Linn. Lyon 1850-1852:123] from tocorisprocerus Linnavuori, 1975 [Acta Zool. France, a nomen dubium, is a primary hom- Fenn. 12: 21], from Ethiopia. onym ofC. punctipes Herrich-Schaeffer, 1835 Phytocoris raunoi nom. n. is proposed for [Nomencl. Entomol. 1: 150] = ? Campylom- Phytocoris irroratus Linnavuori, 1965 [Acta ma verbasci (Meyer-Dur, 1843). Entomol. Fenn. 21: 51], from Turkey, a ju- Polymeruspallescens (Walker, 1873), Cap- nior primary homonym of Phytocoris irror- sus pallescens Walker, 1873 [Cat. Het. Br. atus Blanchard, 1852, Hemipteros, in Gay Mus., 6: 94] from Canada is ajunior primary [Hist. Fis. Polit. Chile 7: 193], from Chile. homonym of Capsus pallescens Herrich- Phytocoris somersojae nom. n. is proposed Schaeffer, 1835 [Nomencl. Entomol. 1: 49] for Phytocoris artemisiae Linnavuori, 1984 from Europe, a nomen dubium, but as P. [Acta Entomol. Fennica 44: 22], from Iraq, pallescens, known from original description a junior primary homonym of P. artemisiae only, is probably identical with some other Schilling, 1837 [Uebers. Arbeit. Verand. species ofPolymerus from Canada, we regard Schles. Ges. Vaterl. Kultur. 1836: 84] = Ly- renaming to be superfluous. gusgemellatus (Herrich-Schaeffer, 1835), Pa- learctic. Phytocoris stonedahli nom. n. is proposed NEW AND REVISED SYNONYMY AND for Phytocoris hispidus Stonedahl, 1988 [Bull. REVISED STATUS Am. Mus. Nat. Hist. 188: 144], from south- Aoplonema Knight, 1928 western US, a junior primary homonym of Phytocoris hispidus Linnavuori, 1986 [Fauna Aoplonema Knight, 1928: 177. revised status Saudi Arabia 8: 135], from Saudi Arabia. Aoplonema Knight, proposed as a subge- Polymerus vegatus (Van Duzee, 1933) Poe- nus, has subsequently been treated as such. ciloscytus vegatus Van Duzee, 1933 [Proc. We elevated the group to generic status on California Acad. Sci., ser. 4, 21: 28] is the the basis of the obvious differences between valid name for P. nigritulus (Walker, 1873) the species placed in it and Hadronema Uhl- Capsus nigritulus Walker, 1873 [Cat. Het. Br. er, including the structure of the male geni- Mus., 6: 112] from Galapagos Is., a junior talia and other structural features. Included primary homonym of Capsus nigritulus Zet- species are: echinatum, Gruetzmacher and terstedt, 1838 [Ins. Lapp.: 279] = Mecomma Schaffner, 1977; festivum Van Duzee, 1910; ambulans (Fallen, 1807), Palearctic. princeps Uhler, 1894; uhleriVan Duzee, 1928; Psallus michaili nom. n. is proposed for and uniforme Knight, 1928. Psallus niger Josifov, 1992 [Reichenbachia 29: 113], from Korea, ajunior primary hom- Deraeocoris Kirschbaum, 1856 onym ofPsallus graminicola f. niger Stichel, 1956 [Illustr. Bestimmungstabellen Deut. Deraeocoris Kirschbaum, 1856: 191, 208. Wanzen 16: 283], from the northern Pale- Cimicicapsus Poppius, 1915: 41. new synonym arctic. Type specimens ofall three species placed Saturniomiris lugens (Walker, 1873), Cap- by Poppius in Cimicicapsus were obtained sus lugens Walker, 1873 [Cat. Het. Br. Mus., from the Zoological Museum in Helsinki for 6: 199] is the valid name for S. tristis (Walker, examination (the holotype of C. brunneus 1873), Capsus tristis Walker, 1873 [Cat. Het. Poppius is badly damaged and represented Br. Mus., 6: 125] from New Guinea, ajunior by fragments only). These species form a nat- primary homonym of Capsus tristis Scholtz, ural group that also includes Deraeocoris ko- 1847 [Arb. Verand. Schles. Ges. Vaterl. Kult., reanus Linnavuori, as characterized by the 1846: 137] from Poland, a nomen dubium. dense vestiture of the hemelytra and some Note: Capsus lugens, a new replacement name other characters. Because they are so similar proposed by Walker himself, was overlooked to representatives of the large and variable in later works. genus Deraeocoris, recognition of a separate We do not propose new replacement names genus for them is unwarranted. The new for two junior primary homonyms: combinations created by this action are: D. 1995 KERZHNER AND SCHUH: MIRIDAE NOMENCLATURAL CHANGES 5 brunneus (Poppius, 1915), D. elongatus (Pop- dicates that it is actually a species ofSejanus pius, 1915), and D. parviceps (Poppius, 1915). Distant. Thus, Eosthenarus becomes ajunior The first of these names becomes a junior synonym of Sejanus. secondary homonym, and a new replacement Examination of the type specimens of name, Deraeocoris brunneolus, is proposed Sthenarus interruptus Reuter, 1906 [Annu. for it. Mus. Zool. St. Petersburg 10: 79] and S. ni- veoarcuatus Reuter, 1906 [Annu. Mus. Zool. Leptidolon Reuter, 1904 St. Petersburg 10: 80] indicates that they be- long to Sejanus, new combinations. Leptidolon Reuter, 1904: 14. revised synonym Carvalho (1952: 65) synonymized Lepti- dolon Reuter with Plagiognathus Fieber Xenofulvius Bergroth, 1920 without comment. Based on the known dis- Xenofulvius Bergroth, 1920: 79. revised synonym tribution ofPlagiognathus, there is every rea- son to believe that the similarity between Carvalho (1952: 76) synonymized Xeno- Leptidolon vittipenne Reuter and Plagiog- fulvius Bergroth with Ceratocapsus Reuter nathus spp. is superficial, and we are there- without comment. Based on the known dis- fore removing the genus from synonymy. tribution of Ceratocapsus, there is every rea- son to believe that the similarity between Platyscytus Reuter, 1907 Xenofulius firmicornis Bergroth and Cerato- capsus spp. is superficial, and we are therefore Platyscytus Reuter, 1907: 16. Amazonophilus Carvalho and Costa, 1992: 203. removing the genus from synonymy. new synonym Carvalho and Costa (1992) described Atomoscelis modestus (Van Duzee) Amazonophilus with the single included spe- ciesA. bipunctatus Carvalho and Costa. Judg- Tuponia modesta Van Duzee, 1914: 30 (n. sp.). ing from the habitus figure and Atomoscelis modestus: Van Duzee, 1917: 414 (n. illustrations comb.). ofthe male genitalia, bipunctatus is a species Mineocapsus mineatus Knight, 1972:425 (n. sp.). of Platyscytus, and we are so treating it. new synonym Rhinacloa Reuter, 1876 Van Duzee (1914) described Atomoscelis modestus from California. We have exam- Rhinacloa Reuter, 1876: 88. ined many specimens identified as modestus Sinopmiris Carvalho, 1991: 92. new synonym and compared this species with A. onustus Carvalho (199 1) described Sinopmiris with Fieber, 1861 (type species of Atomoscelis) the single included species S. clarus Carval- from the Palearctic; the two are clearly very ho. Judging from the description and the fig- closely related. Knight (1972) described Mi- ures of the male genitalia, this is a species of neocapsus mineatus on the basis ofspecimens Rhinacloa Reuter, and we are so treating it. from Utah. They are clearly conspecific with specimens from Knight's collection identi- Sejanus Distant, 1910 fied as modestus. We are therefore treating mineatus as a junior synonym. Mineocapsus Sejanus Distant, 1910: 20. Knight, 1972, becomes a junior synonym of Eosthenarus Poppius, 1915: 72 (n. gen.). revised Atomoscelis Reuter, 1875, new synonym. synonym Poppius (1915) described the genus Eos- thenarus from Taiwan, designating E. cras- Opuna annulata (Knight) sicornis as Poppius, 1915, the type species. CampylommaannulatusKnight, 1935:197 (n. sp.). Carvalho (1952) placed Eosthenarus in syn- Pararagmus annulicornis Poppius, 1911: 35 (n. onymy with Chlamydatus Curtis. Our ex- sp.). new synonym amination of the type specimen of crassicor- Pararagmus ceylonensis Carvalho, 1955: 225 (n. nis in the Zoological Museum, Helsinki, in- name). new synonym 6 AMERICAN MUSEUM NOVITATES NO. 3137

Pararagmus lindbergi Kiritshenko, 1961: 444 (un- Phyllopidea picta (Uhler) neccesary new name). new synonym Reuteriola annulicornis Hsiao in Hsiao and Meng, Bolteria picta Uhler, 1893: 373 (n. sp., desc.). 1963: 445, 449. new synonym Europiella concinna Reuter, 1909: 84 (n. sp., desc.). Opuna annulatus: Schuh, 1984: 401 (n. comb.). new synonym Hyoidea picta Reuter, 1909: 72 (note). Comparison of specimens of Opuna an- Bolteriapicta hirta Van Duzee, 1916: 244 (n. var.). nulata (Knight) with the original description new synonym of Pararagmus annulicornis Poppius, (= P. Phyllopidea hirta Knight, 1920: 127 (n. comb, ceylonensis Carvalho, 1955) indicates that the note). two are synonyms. Because annulicornis Phyllopidea picta Knight, 1920: 127 (note, n. comb., lectotype designation). Poppius was replaced as a junior secondary Phyllopidea utahensis Knight, 1968: 32 (n. sp., homonym before 1960, this name cannot be desc., host). new synonym used, and, annulata Knight becomes the val- id name for this species. Because annulicornis Lectotype: 9, Am. Fork, Ut., 22.6.91; de- Poppius is the type species ofthe genus, Par- posited in the USNM. aragmus Poppius, 191 1, that genus becomes Lectotype ofSynonym: Bolteria picta hirta a junior synonym of Opuna Kirkaldy, 1902, Van Duzee: 9, G. Alpine Cr., 7-5-15, Tahoe new synonym. Cal., E. P. Van Duzee Collector; deposited Evaluation of the original description of in the CAS. Reuteriola annulicornis Hsiao, 1963, indi- Holotypes of Synonyms: Europiella con- cates that this taxon is also synonymous with cinna Reuter: 9, Claremont, Cal., Baker; de- annulata Knight. Consequently, Reuteriola posited in the Helsinki Museum. Phyllopidea Hsiao becomes a junior synonym of Opuna utahensis Knight: 8, Scipio, UTAH, June 29, new synonym. 1965, H. H. Knight; deposited in the USNM. Discussion: Comparison of the lectotypes Parapsallus vitellinus (Scholtz) of Bolteria picta Uhler, Bolteria picta hirta Van Duzee, and the holotypes of Europiella Capsus vitellinus Scholtz, 1847: 130 (n. sp.). concinna Reuter and Phyllopidea utahensis Parapsallus vitellinus: Kerzhner, 1964: 760 (n. Knight indicates that all four are conspecific; comb.). Parapsallus wagneri Rozhkov and Volkova, 1966: picta has priority. 59 (n. sp.). new synonym Van Duzee described the subspecies picta hirta from "eight female examples beaten Rozhkov and Volkova (1966) under the from sagebrush (Artemesia) [sic] on the shore heading "Parapsallus wagneri Kulik, sp. n. of Fallen Leaf Lake near the Lodge on July (in lit.)" published a short description and 5." We received on loan one specimen la- figures of a mirid common on Larix sibirica beled as the lectotype and nine additional in East Siberia (Cisbaikalia). The authors specimens labeled as paratypes from the Cal- stated that the species was determined as new ifornia Academy of Sciences. Despite the la- by S. A. Kulik and his identification con- bels, there is apparently no published lecto- firmed by E. Wagner, although Kulik in his type designation for this taxon, so we have later publications on Siberian Heteroptera affixed lectotype and paralectotype labels to used the name P. vitellinus, and not P. wag- eight of the specimens. neri. Judging from some of the characters mentioned (two cells ofmembrane, four seg- Pseudoloxops coccineus ments ofantennae) the description ofP. wag- (Meyer-Dur, 1843) neri was made by Rozhkov and Volkova and the name should be credited to them. In our Capsus coccineus Meyer-Dur, 1843: 75. opinion, the Siberian specimens of Parap- Capsus miniatus Parfitt, 1865: 130. new synonym sallus do not differ from European speci- Capsus miniatus Parfitt was described from mens, and the name P. wagneri should be one specimen collected at Exeter (England). placed in synonymy ofthe widely distributed The name is a junior primary homonym of P. vitellinus. C. miniatus Herrich-Schaeffer, 1838 [Wanz. 1995 KERZHNER AND SCHUH: MIRIDAE NOMENCLATURAL CHANGES 7

Ins. 4: 34] which is a junior synonym ofDer- Orthotylus Fieber, 1858 aeocoris schach (Fabricius, 1781). The above Orthotylus Fieber, 1858: 315. synonymy is clear from Parfitt's description Melanotrichus Reuter, 1875: 151. and was apparently known to British authors because Saunders (1892: 294) cited "Exeter, Parfitt" in distribution of "Loxops cocci- Melanotrichus Reuter has been treated as neus." However, we are unable to find any a subgenus of Orthotylus or as a genus, de- mention of Parfitt's name in synonymy and pending on the author. It has also been treat- are forced to establish the synonymy as new. ed as a synonym of Orthotylus. Several au- thors have recently described new species of NEW COMBINATIONS Orthotylinae in Melanotrichus. In our view these generic assignments are poorly founded Badezorus annulicornis (Reuter), because no characters will consistently allow new combination for recognition ofMelanotrichus and its dis- Plagiognathus annulicornis Reuter, 1879: 298 (n. tribution clearly suggests an unnatural assem- sp.). blage. We therefore prefer to treat these spe- Pararagmus subsinuatus Poppius, 1912: 25 (n. sp.). cies as members ofa more broadly construed Pararagmus annulicornis: Kiritshenko, 1926: 226 Orthotylus until such time as more compre- (n. comb., syn.). hensive studies are conducted on the generic Our examination of the male genitalia of classification of Orthotylus and its near rel- Plagiognathus Reuter indicates atives-recognizing that Orthotylus is also annulicornis clearly not monophyletic. The new combi- that the vesica has a very long single terminal nations created by this action are: 0. argen- spine of the type found in Badezorus signa- tinus Carvalho, 1985; 0. minensis Carvalho, ticornis (Reuter, 1904). The species is closely 1985; 0. saltensis Carvalho, 1985; 0. clar- related if not synonymous with Chamaep- ensis Carvalho, 1990; 0. bonaerensis Car- sallus tomentosus (Reuter, 1904), which was transferred to valho and Carpintero, 1986; 0. missionensis Badezorus by Linnavuori Carvalho and Carpintero, 1986; 0. sumal- (1993a). oensis Carvalho and Carpintero, 1986; 0. Michailocoris chinensis (Hsiao), vermelhensis Carvalho and Costa, 1992; 0. new combination joacemensis Carvalho and Costa, 1992; 0. membranosus Carvalho and Costa, 1992; 0. Aretas chinensis Hsiao, 1941: 245 (n. sp.). elongatus Kelton, 1980; 0. Pseudoloxops chinensis: Carvalho, 1958: 127 (n. pallens Knight, comb.). 1968; and 0. uniformis Knight, 1968. Five of these names-clarensis, elongatus, missi- The only material mentioned in the orig- onensis, pallens, and uniformis -become sec- inal description ofA. chinensis was a female ondary homonyms as a result oftheir transfer holotype, but male genitalia were figured. In to Orthotylus. We propose replacement names the U.S. National Museum the species is rep- above. resented by one male labeled by Hsiao him- A final case involves Dichaetocoris brevi- self as "allotype" but possibly being the ho- rostris Knight, 1968 [Brigham Young Univ. lotype. Examination of this specimen shows Sci. Bull., Biol. Ser., 9: 115], which was later that the species was wrongly placed by Hsiao transferred to Melanotrichus by Polhemus, in the genus Aretas Distant (iunior synonym 1985 [Pan-Pac. Entomol. 61: 149]. Because ofPseudoloxops Kirkaldy), Orthotylinae, and of the resulting secondary homonymy with belongs in fact to the genus Michaiocoris Stys, M. brevirostris Knight, 1927 (transferred by Bryocorinae, Eccritotarsini. Michailocoris Kelton in 1978 to Brooksetta Kelton), Pol- chinensis is similar to M. josifovi Stys (Korea, hemus proposed the replacement name M. Far East of Russia), but differs from it in knighti. The replacement name was unnec- having relatively larger eyes and in having essary because the two taxa were not consid- the coloration of the clavus in the male sim- ered synonymous (International Code of ilar to that in the M. josifovi female. Zoological Nomenclature, Article 59a). 8 AMERICAN MUSEUM NOVITATES NO. 3137

FIXATION OF TYPE SPECIES black (rather than pale) with a longitudinal dark stripe on the apical half as in nigriceps. Brasiliocarnus Kerzhner and Schuh Furthermore, these specimens do not have Brasiliocarnus Carvalho, 1984: 377. what might be considered a typical habitus for the Halticini; they are elongate, macrop- Carvalho (1984) proposed the name Bras- terous in both sexes with semitransparent he- iiocarnus with two included species, but failed melytra, and have a well-delimited pronotal to designate a type species. In the Zoological collar and slender hind femora. They do, Record for 1985 [ZR, 1987, pt. 13F: 197], B. however, have typical halticine male geni- bahiensis was indicated as the type, but be- talia, with the right paramere spoon-shaped, cause this fixation is anonymous it is invalid. the left paramere 7-form, and the aedeagus The type of the genus is here designated as without a vesica and spines. Based on these Brasiliocarnus fraudans Stal, 1860 [K. Vet. observations we transfer Dicyphopsis to the Akad. Handl. 2(7): 52]. Halticini. A second species placed in Dicyphopsis, D. Cleotomiris Schuh spectabilis Linnavuori, 1975 (holotype ex- Cleotomiris Schuh, 1984: 81. amined), does not fit the original description Cleoto- of the genus in many significant details. The Schuh (1984) proposed the name eyes are granulose, not touching the anterior miris with four included species. For reasons margin of the pronotum (contrary to Lin- not now obvious, no type species was des- navuori's description), and much wider than ignated. The type of the genus is here des- the vertex (in Dicyphopsis half as wide). The ignated as Cleotomiris schneirlai Schuh, 1984. large cell of the membrane forms a marked This nomenclatural act, which makes the angle whereas in Dicyphopsis it is broadly name available, should be credited to Schuh rounded, and the color pattern is strongly dis- (this paper). similar to that of D. nigriceps. Cassis (1986) examined two specimens of the type series REVISED HIGHER-CATEGORY and stated that they strongly resemble Cam- PLACEMENTS pyloneuropsis Poppius. We agree with his conclusions and place spectabilis Linnavuori Dicyphopsis Poppius, 1914 in Campyloneuropsis. Dicyphopsis Poppius, 1914: 1 1. Dicyphopsis, with the single included spe- Ifephylus Linnavuori, 1993 cies D. nigriceps Poppius, 1914, was de- Ifephylus Linnavuori, 1993b: 207. scribed from one female specimen collected in Tanzania (Kilimandjaro) and placed in the Linnavuori placed his new genus Ifephylus Macrolophinae (now Dicyphina) by its au- in the Pilophorini solely on the basis of pre- thor. The holotype was completely destroyed tarsal structure. Based on the structure ofthe on shipment from the Naturhistoriska Riks- male genitalia, we treat this taxon as belong- museet, Stockholm, to G. Cassis (Cassis, ing to the Phylini. 1986). We have examined specimens in the American Museum of Natural History con- Mendozaphylus sidered by us to be congeneric, but not con- Carvalho and Carpintero, 1991 specific, with D. nigriceps specimens collect- ed in Zaire by N. A. Weber (Stanleyville, Mendozaphylus Carvalho and Carpintero, 199 lb: March 18, 1948) and Ghana by R. T. Schuh 201. and J. A. Slater (Tafo, October, 5, 1967; many Carvalho and Carpintero (199 ib) placed males and females). These specimens fit the Mendozaphylus in the Hallodapini. Based on original description in all details except that the description and habitus figure and illus- the dorsal vestiture is pale (not dark), anten- trations of the male genitalia of the single nal segment 1 lacks a longitudinal ventroap- included species, we are placing this taxon in ical dark stripe, and segment 2 is completely the Phylini. No true Hallodapini have been 1995 KERZHNER AND SCHUH: MIRIDAE NOMENCLATURAL CHANGES 9 previously documented from the Neotropics, nation ofspecimens will be necessary to more and this one lacks the distinctive genitalia of precisely determine its identity. the Old World taxa and Cyrtopeltocoris Reu- ter from North America. Tibiopilus Carvalho and Costa, 1993 Opisthocoris Tibiopilus Carvalho and Costa, 1993: 207. Carvalho and Costa, 1991 Carvalho and Costa (1993) placed Tibio- pilus in the Hallodapini. Based on the de- Opisthocoris Carvalho and Costa, 1991: 206. re- vised family placement scription of the pronotal collar as "deprim- ido" (depressed) as well as the habitus figure Carvalho and Costa (1991) placed Opisth- and the illustrations of the male genitalia of ocoris, with its single included species 0. car- the single included species T. pedunculatus melitanus Carvalho and Costa, 1991, in the Carvalho and Costa, we are transferring this Stenodemini. Based on the habitus figure and taxon to the Phylini. No Hallodapini have illustrations of the pretarsus, we do not be- been documented from the neotropics, and lieve this taxon belongs to the Miridae, but Tibiopilus appears to lack the diagnostic fea- rather to the Lygaeidae, Cyminae. Exami- tures of the group.

REFERENCES Bergroth, E. cricoes de um novo genero & treze es- 1920. List of the Cylapinae (Hem., Miridae) pecies novas da Republica Argentina with descriptions of new Philippine (). Rev. Brasileria Biol. 46: forms. Ann. Soc. Entomol. Belgique 60: 607-625. 67-83. 1991a. Miridos neotropicales CCCXLVI: des- Carvalho, J. C. M. cripcion de un genero y siete especies 1952. On the major classification of the Mir- nuevas de las subfamilia Orthotylinae, idae (Hemiptera). (With keys to sub- con notas y correciones taxonomicas families and tribes and a catalogue of (Hemiptera). An. Acad. Brasileira Cienc. the world genera). An. Acad. Brasileira 63: 33-42. Cienc. 24: 31-110. 1991b. Mirideos neotropicais, CCCXLVII: no- 1955. Analecta Miridologica: miscellaneous vos generos e especies da Republica Ar- observations in some American muse- gentina (Hemiptera). Ibid., 63: 201-204. ums and bibliography. Rev. Chilena 1992. Mirideos neotropicais, CCCXXXVII. Entomol. 4: 221-227. descricao de doce especies novas de Or- 1957-60. Catalogue ofthe Miridae ofthe world, thotylinae (Hemiptera). Rev. Brasileira Parts 1-5. Arq. Mus. Nac. Rio de Ja- Biol. 51: 763-780 (1991). neiro 44: 1-158; 45: 1-216; 47: 1-161; Carvalho, J. C. M., and L. A. A. Costa 48: 1-384; 51: 1-194. 1991. Mirideos neotropicais, CCCL: sobre un 1984. Mirideos neotropicais, CCL: generos e genero e especie curiosa de Carmo do especies do complexo Pseudocarnus- Rio Claro, Minas Gerais (Hemiptera). Antias-Carmelus (Hemiptera). Rev. An. Acad. Brasileira Cienc. 63: 205-207. Brasileira Biol. 44: 377-386. 1992. Mirideos neotropicais, CCCXXXVIII: 1985. Mirideos Neotropicais, CCLIII: Des- novo genero e especies de e no- cricoes de novos generos e especies da vas especies de Orthotylinae (Hemip- tribo Orthotylini Van Duzee (Hemip- tera). Rev. Brasileira Biol. 52: 199-207. tera). Ibid., 45: 249-298. 1993. Mirideos neotropicais, CCCLXXVII: 1991. Mirideos neotropicais, CCCXIII: dois dois generos e especies de Phylinae (He- generos novos da tribo Phylini Douglas miptera). An. Acad. Brasileira Cienc. 65: and Scott (Hemiptera). Bol. Mus. Par- 203-207. aense Emilio Goeldi, nova serie, Zool. Cassis, G. 6: 89-94 (1990). 1986. A systematic study ofthe subfamily Di- Carvalho, J. C. M., and D. L. Carpintero cyphinae (Heteroptera: Miridae). Ann 1986. Mirideos neotropicais, CCLXXIII: des- Arbor, Univ. Microfilms, 400 pp. [Ph.D. 10 AMERICAN MUSEUM NOVITATES NO. 3137

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Aprilis 1906 a D. E. P. Van Duzee col- Miridae) ofthe Indo-Pacific. Bull. Am. lectae. Ibid., 49(5): 27 pp. Mus. Nat. Hist. 177(1): 1-476. Reuter, 0. M. Stichel, W. 1909. Bemerkungen uber nearktische Capsi- 1957. Illustrierte Bestimmungstabellen der den nebst Beschreibung neuer Arten. Wanzen. II. Europa (Hemiptera-Het- Acta Soc. Sci. Fenn. 36(2): 86 pp. eroptera Europae). Fasc. 16-22, pp. 481- Rozhkov, A. S., and L. V. Volkova 704. Berlin: Hermsdorf. 1966. Order Hemiptera, pp. 58-60. In A. S. Uhler, P. R. Roshkov (ed.), Vrediteli listvennitsy si- 1893. Summary of the collections secured by birskoi [Pests ofLarix sibirica]. 328 pp. Mr. E. A. Schwartz in Utah. Proc. En- Moscow, Nauka. [In Russian] tomol. Soc. Washington 2: 366-385. Saunders, E. Van Duzee, E. P. 1892. The Hemiptera Heteroptera ofthe Brit- 1914. A preliminary list of the Hemiptera of ish Islands. A descriptive account ofthe San Diego County, California. Trans. families, genera, and species indigenous San Diego Soc. Nat. Hist. 2: 1-57. to Great Britain and Ireland, with notes 1916. Notes on some Hemiptera taken near on the localities, habitats etc. 350 pp., Lake Tahoe, California. Univ. Califor- 32 pis. London: Reeve. nia Pub. Entomol., Tech. Bull. 1: 229- Scholtz, H. 249. 1847. Prodromus zu einer Rhynchoten-Fauna 1917. Catalogue ofthe Hemiptera ofAmerica von Schlesien. Uebers. Arb. Verand. north of Mexico (excepting the Aphid- Schles. Ges. Vaterl. Kult. 1846: 104- idae, Coccidae, and Aleurodidae). Univ. 164. California Pub. Tech. Bull. Entomol. 2: Schuh, R. T. xiv + 902 pp. 1984. Revision of the Phylinae (Hemiptera, Recent issues of the Novitates may be purchased from the Museum. Lists of back issues of the Novitates, Bulletin, and Anthropological Papers published during the last five years are available free of charge. Address orders to: American Museum ofNatural History Library, Department D, Central Park West at 79th St., New York, N.Y. 10024. TEL: (212) 769-5545. FAX: (212) 769- 5009. E-MAIL: scipubseamnh.org

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