Michilogic Time Line
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Decoupled Evolution of Soft and Hard Substrate Communities During the Cambrian Explosion and Great Ordovician Biodiversification Event
Decoupled evolution of soft and hard substrate communities during the Cambrian Explosion and Great Ordovician Biodiversification Event Luis A. Buatoisa,1, Maria G. Mánganoa, Ricardo A. Oleab, and Mark A. Wilsonc aDepartment of Geological Sciences, University of Saskatchewan, Saskatoon, SK, Canada S7N 5E2; bEastern Energy Resources Science Center, US Geological Survey, Reston, VA 20192; and cDepartment of Geology, The College of Wooster, Wooster, OH 44691 Edited by Steven M. Holland, University of Georgia, Athens, GA, and accepted by Editorial Board Member David Jablonski May 6, 2016 (received for review November 21, 2015) Contrasts between the Cambrian Explosion (CE) and the Great shed light on the natures of both radiations. Because there is still Ordovician Biodiversification Event (GOBE) have long been recog- controversy regarding Sepkoski’s nonstandardized curves of nized. Whereas the vast majority of body plans were established Phanerozoic taxonomic diversity (13–15), a rarefaction analysis as a result of the CE, taxonomic increases during the GOBE were was performed in an attempt to standardize diversity data. The manifested at lower taxonomic levels. Assessing changes of ichno- aims of this paper are to document the contrasting ichnodiversity diversity and ichnodisparity as a result of these two evolutionary and ichnodisparity trajectories in soft and hard substrate com- events may shed light on the dynamics of both radiations. The early munities during these two evolutionary events and to discuss the Cambrian (series 1 and 2) displayed a dramatic increase in ichnodi- possible underlying causes of this decoupled evolution. versity and ichnodisparity in softground communities. In contrast to this evolutionary explosion in bioturbation structures, only a few Results Cambrian bioerosion structures are known. -
Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
Chapter 2 Paleozoic Stratigraphy of the Grand Canyon
CHAPTER 2 PALEOZOIC STRATIGRAPHY OF THE GRAND CANYON PAIGE KERCHER INTRODUCTION The Paleozoic Era of the Phanerozoic Eon is defined as the time between 542 and 251 million years before the present (ICS 2010). The Paleozoic Era began with the evolution of most major animal phyla present today, sparked by the novel adaptation of skeletal hard parts. Organisms continued to diversify throughout the Paleozoic into increasingly adaptive and complex life forms, including the first vertebrates, terrestrial plants and animals, forests and seed plants, reptiles, and flying insects. Vast coal swamps covered much of mid- to low-latitude continental environments in the late Paleozoic as the supercontinent Pangaea began to amalgamate. The hardiest taxa survived the multiple global glaciations and mass extinctions that have come to define major time boundaries of this era. Paleozoic North America existed primarily at mid to low latitudes and experienced multiple major orogenies and continental collisions. For much of the Paleozoic, North America’s southwestern margin ran through Nevada and Arizona – California did not yet exist (Appendix B). The flat-lying Paleozoic rocks of the Grand Canyon, though incomplete, form a record of a continental margin repeatedly inundated and vacated by shallow seas (Appendix A). IMPORTANT STRATIGRAPHIC PRINCIPLES AND CONCEPTS • Principle of Original Horizontality – In most cases, depositional processes produce flat-lying sedimentary layers. Notable exceptions include blanketing ash sheets, and cross-stratification developed on sloped surfaces. • Principle of Superposition – In an undisturbed sequence, older strata lie below younger strata; a package of sedimentary layers youngs upward. • Principle of Lateral Continuity – A layer of sediment extends laterally in all directions until it naturally pinches out or abuts the walls of its confining basin. -
MARS DURING the PRE-NOACHIAN. J. C. Andrews-Hanna1 and W. B. Bottke2, 1Lunar and Planetary La- Boratory, University of Arizona
Fourth Conference on Early Mars 2017 (LPI Contrib. No. 2014) 3078.pdf MARS DURING THE PRE-NOACHIAN. J. C. Andrews-Hanna1 and W. B. Bottke2, 1Lunar and Planetary La- boratory, University of Arizona, Tucson, AZ 85721, [email protected], 2Southwest Research Institute and NASA’s SSERVI-ISET team, 1050 Walnut St., Suite 300, Boulder, CO 80302. Introduction: The surface geology of Mars appar- ing the pre-Noachian was ~10% of that during the ently dates back to the beginning of the Early Noachi- LHB. Consideration of the sawtooth-shaped exponen- an, at ~4.1 Ga, leaving ~400 Myr of Mars’ earliest tially declining impact fluxes both in the aftermath of evolution effectively unconstrained [1]. However, an planet formation and during the Late Heavy Bom- enduring record of the earlier pre-Noachian conditions bardment [5] suggests that the impact flux during persists in geophysical and mineralogical data. We use much of the pre-Noachian was even lower than indi- geophysical evidence, primarily in the form of the cated above. This bombardment history is consistent preservation of the crustal dichotomy boundary, to- with a late heavy bombardment (LHB) of the inner gether with mineralogical evidence in order to infer the Solar System [6] during which HUIA formed, which prevailing surface conditions during the pre-Noachian. followed the planet formation era impacts during The emerging picture is a pre-Noachian Mars that was which the dichotomy formed. less dynamic than Noachian Mars in terms of impacts, Pre-Noachian Tectonism and Volcanism: The geodynamics, and hydrology. crust within each of the southern highlands and north- Pre-Noachian Impacts: We define the pre- ern lowlands is remarkably uniform in thickness, aside Noachian as the time period bounded by two impacts – from regions in which it has been thickened by volcan- the dichotomy-forming impact and the Hellas-forming ism (e.g., Tharsis, Elysium) or thinned by impacts impact. -
University of Michigan University Library
CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN VOL.XIX, NO.8, pp. 105-114 (1 pl., 1 fig.) NOVEMBER25, 1964 A NEW SPIRACULATE BLASTOID, PYRAMIBLASTUS, FROM THE MISSISSIPPIAN HAMPTON FORMATION OF IOWA BY DONALD B. MACURDA, JR. MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN ANN ARBOR CONTRIBUTIONS FROM THE MUSEUM OF PaEONTOLOGY Director: LEWISB. KELLUM The series of contributions from the Museum of Paleontology is a medium for the publication of papers based chiefly upon the collections in the Museum. When the number of pages issued is sufficient to make a volume, a title page and a table of contents will be sent to libraries on the mailing list, and to individuals upon request. A list of the separate papers may also be obtained. Correspondence should be directed to the Museum of Paleontology, The University of Michigan, Ann Arbor, Michigan. VOLUME XIX 1. Silicified Trilobites from the Devonian Jeffersonville Limestone at the Falls of the Ohio, by Erwin C. Stumm. Pages 1-14, with 3 plates. 2. Two Gastropods from the Lower Cretaceous (Albian) of Coahuila, Mexico, by Lewis B. Kellum and Kenneth E. Appelt. Pages 15-22, with 2 figures. 3. Corals of the Traverse Group of Michigan, Part XII, The Small-celled Species of Favosites and Emmonsia, by Erwin C. Stumm and John H. Tyler. Pages 23-36, with 7 plates. 4. Redescription of Syntypes of the Bryozoan Species Rhombotrypa quadrata (Rominger), by Roger J. Cuffey and T. G. Perry. Pages 37-45, with 2 plates. 5. Rare Crustaceans from the Upper Devonian Chagrin Shale in Northern Ohio, by Myron T. -
The Rock and Fossil Record the Rock and Fossil Record the Rock And
TheThe RockRock andand FossilFossil RecordRecord Earth’s Story and Those Who First Listened . 426 Apply . 427 Internet Connect . 428 When on Earth? . 429 Activity . 430 MathBreak . 434 Internet Connect 432, 435 Looking at Fossils . 436 QuickLab . 438 Internet Connect . 440 Time Marches On . 441 QuickLab . 443 Internet Connect . 445 Chapter Lab . 446 Chapter Review . 449 TEKS/TAKS Practice Tests . 451, 452 Feature Article . 453 Time Stands Still Pre-Reading Questions Sealed in darkness for 49 million years, this beetle still shimmers with the same metallic hues that once helped it hide among ancient plants. This rare fossil 1. How do scientists study was found in Messel, Germany. In the same rock formation, the Earth’s history? scientists have found fossilized crocodiles, bats, birds, and 2. How can you tell the age frogs. A living stag beetle (above) has a similar form and of rocks and fossils? color. Do you think that these two beetles would live in 3. What natural or human similar environments? What do you think Messel, Germany, events have caused mass was like 49 million years ago? In this chapter, you will extinctions in Earth’s learn how scientists answer these kinds of questions. history? 424 Chapter 16 Copyright © by Holt, Rinehart and Winston. All rights reserved. MAKING FOSSILS Procedure 1. You and three or four of your classmates will be given several pieces of modeling clay and a paper sack containing a few small objects. 2. Press each object firmly into a piece of clay. Try to leave an imprint showing as much detail as possible. -
The Case of the Diminutive Trilobite Flexicalymene Retrorsa Minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati Region
EVOLUTION & DEVELOPMENT 9:5, 483–498 (2007) Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region Brenda R. Hundaa,Ã and Nigel C. Hughesb aCincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA bDepartment of Earth Sciences, University of California, Riverside, CA 92521, USA ÃAuthor for correspondence (email: [email protected]) SUMMARY Flexicalymene retrorsa minuens from the upper- rate of progress along a common ontogenetic trajectory with most 3 m of the Waynesville Formation of the Cincinnatian respect to size, coupled with growth cessation at a small size, Series (Upper Ordovician) of North America lived ‘‘sequential’’ progenesis, or non-uniform changes in the rate of approximately 445 Ma and exhibited marked reduction in progress along a shared ontogenetic trajectory with respect to maximum size relative to its stratigraphically subjacent sister size, can also be rejected. Rather, differences between these subspecies, Flexicalymene retrorsa retrorsa. Phylogenetic subspecies are more consistent with localized changes in analysis is consistent with the notion that F. retrorsa retrorsa rates of character development than with a global hetero- was the ancestor of F. retrorsa minuens. F. retrorsa minuens chronic modification of the ancestral ontogeny. The evolution has been claimed to differ from F. retrorsa retrorsa ‘‘in size of F. retrorsa minuens from F. retrorsa retrorsa was largely alone,’’ and thus presents a plausible example of global dominated by modifications of the development of characters paedomorphic evolution in trilobites. Despite strong similarity already evident in the ancestral ontogeny, not by the origin of in the overall form of the two subspecies, F. -
Critical Analysis of Article "21 Reasons to Believe the Earth Is Young" by Jeff Miller
1 Critical analysis of article "21 Reasons to Believe the Earth is Young" by Jeff Miller Lorence G. Collins [email protected] Ken Woglemuth [email protected] January 7, 2019 Introduction The article by Dr. Jeff Miller can be accessed at the following link: http://apologeticspress.org/APContent.aspx?category=9&article=5641 and is an article published by Apologetic Press, v. 39, n.1, 2018. The problems start with the Article In Brief in the boxed paragraph, and with the very first sentence. The Bible does not give an age of the Earth of 6,000 to 10,000 years, or even imply − this is added to Scripture by Dr. Miller and other young-Earth creationists. R. C. Sproul was one of evangelicalism's outstanding theologians, and he stated point blank at the Legionier Conference panel discussion that he does not know how old the Earth is, and the Bible does not inform us. When there has been some apparent conflict, either the theologians or the scientists are wrong, because God is the Author of the Bible and His handiwork is in general revelation. In the days of Copernicus and Galileo, the theologians were wrong. Today we do not know of anyone who believes that the Earth is the center of the universe. 2 The last sentence of this "Article In Brief" is boldly false. There is almost no credible evidence from paleontology, geology, astrophysics, or geophysics that refutes deep time. Dr. Miller states: "The age of the Earth, according to naturalists and old- Earth advocates, is 4.5 billion years. -
The Geologic Time Scale Is the Eon
Exploring Geologic Time Poster Illustrated Teacher's Guide #35-1145 Paper #35-1146 Laminated Background Geologic Time Scale Basics The history of the Earth covers a vast expanse of time, so scientists divide it into smaller sections that are associ- ated with particular events that have occurred in the past.The approximate time range of each time span is shown on the poster.The largest time span of the geologic time scale is the eon. It is an indefinitely long period of time that contains at least two eras. Geologic time is divided into two eons.The more ancient eon is called the Precambrian, and the more recent is the Phanerozoic. Each eon is subdivided into smaller spans called eras.The Precambrian eon is divided from most ancient into the Hadean era, Archean era, and Proterozoic era. See Figure 1. Precambrian Eon Proterozoic Era 2500 - 550 million years ago Archaean Era 3800 - 2500 million years ago Hadean Era 4600 - 3800 million years ago Figure 1. Eras of the Precambrian Eon Single-celled and simple multicelled organisms first developed during the Precambrian eon. There are many fos- sils from this time because the sea-dwelling creatures were trapped in sediments and preserved. The Phanerozoic eon is subdivided into three eras – the Paleozoic era, Mesozoic era, and Cenozoic era. An era is often divided into several smaller time spans called periods. For example, the Paleozoic era is divided into the Cambrian, Ordovician, Silurian, Devonian, Carboniferous,and Permian periods. Paleozoic Era Permian Period 300 - 250 million years ago Carboniferous Period 350 - 300 million years ago Devonian Period 400 - 350 million years ago Silurian Period 450 - 400 million years ago Ordovician Period 500 - 450 million years ago Cambrian Period 550 - 500 million years ago Figure 2. -
“Anthropocene” Epoch: Scientific Decision Or Political Statement?
The “Anthropocene” epoch: Scientific decision or political statement? Stanley C. Finney*, Dept. of Geological Sciences, California Official recognition of the concept would invite State University at Long Beach, Long Beach, California 90277, cross-disciplinary science. And it would encourage a mindset USA; and Lucy E. Edwards**, U.S. Geological Survey, Reston, that will be important not only to fully understand the Virginia 20192, USA transformation now occurring but to take action to control it. … Humans may yet ensure that these early years of the ABSTRACT Anthropocene are a geological glitch and not just a prelude The proposal for the “Anthropocene” epoch as a formal unit of to a far more severe disruption. But the first step is to recognize, the geologic time scale has received extensive attention in scien- as the term Anthropocene invites us to do, that we are tific and public media. However, most articles on the in the driver’s seat. (Nature, 2011, p. 254) Anthropocene misrepresent the nature of the units of the International Chronostratigraphic Chart, which is produced by That editorial, as with most articles on the Anthropocene, did the International Commission on Stratigraphy (ICS) and serves as not consider the mission of the International Commission on the basis for the geologic time scale. The stratigraphic record of Stratigraphy (ICS), nor did it present an understanding of the the Anthropocene is minimal, especially with its recently nature of the units of the International Chronostratigraphic Chart proposed beginning in 1945; it is that of a human lifespan, and on which the units of the geologic time scale are based. -
Pennsylvanian Boundary Unconformity in Marine Carbonate Successions
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Dissertations & Theses in Earth and Atmospheric Earth and Atmospheric Sciences, Department of Sciences Summer 6-2014 ORIGIN AND DISTRIBUTION OF THE MISSISSIPPIAN – PENNSYLVANIAN BOUNDARY UNCONFORMITY IN MARINE CARBONATE SUCCESSIONS WITH A CASE STUDY OF THE KARST DEVELOPMENT ATOP THE MADISON FORMATION IN THE BIGHORN BASIN, WYOMING. Lucien Nana Yobo University of Nebraska-Lincoln, [email protected] Follow this and additional works at: http://digitalcommons.unl.edu/geoscidiss Part of the Geochemistry Commons, Geology Commons, Sedimentology Commons, and the Stratigraphy Commons Nana Yobo, Lucien, "ORIGIN AND DISTRIBUTION OF THE MISSISSIPPIAN – PENNSYLVANIAN BOUNDARY UNCONFORMITY IN MARINE CARBONATE SUCCESSIONS WITH A CASE STUDY OF THE KARST DEVELOPMENT ATOP THE MADISON FORMATION IN THE BIGHORN BASIN, WYOMING." (2014). Dissertations & Theses in Earth and Atmospheric Sciences. 59. http://digitalcommons.unl.edu/geoscidiss/59 This Article is brought to you for free and open access by the Earth and Atmospheric Sciences, Department of at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Dissertations & Theses in Earth and Atmospheric Sciences by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. ORIGIN AND DISTRIBUTION OF THE MISSISSIPPIAN – PENNSYLVANIAN BOUNDARY UNCONFORMITY IN MARINE CARBONATE SUCCESSIONS WITH A CASE STUDY OF THE KARST DEVELOPMENT ATOP THE MADISON FORMATION IN THE BIGHORN BASIN, WYOMING. By Luscalors Lucien Nana Yobo A THESIS Presented to the Faculty of The Graduate College at the University of Nebraska In Partial Fulfillment of Requirements For the Degree of Master of Science Major: Earth and Atmospheric Sciences Under the Supervision of Professor Tracy D. -
Geologic Time and Geologic Maps
NAME GEOLOGIC TIME AND GEOLOGIC MAPS I. Introduction There are two types of geologic time, relative and absolute. In the case of relative time geologic events are arranged in their order of occurrence. No attempt is made to determine the actual time at which they occurred. For example, in a sequence of flat lying rocks, shale is on top of sandstone. The shale, therefore, must by younger (deposited after the sandstone), but how much younger is not known. In the case of absolute time the actual age of the geologic event is determined. This is usually done using a radiometric-dating technique. II. Relative geologic age In this section several techniques are considered for determining the relative age of geologic events. For example, four sedimentary rocks are piled-up as shown on Figure 1. A must have been deposited first and is the oldest. D must have been deposited last and is the youngest. This is an example of a general geologic law known as the Law of Superposition. This law states that in any pile of sedimentary strata that has not been disturbed by folding or overturning since accumulation, the youngest stratum is at the top and the oldest is at the base. While this may seem to be a simple observation, this principle of superposition (or stratigraphic succession) is the basis of the geologic column which lists rock units in their relative order of formation. As a second example, Figure 2 shows a sandstone that has been cut by two dikes (igneous intrusions that are tabular in shape).The sandstone, A, is the oldest rock since it is intruded by both dikes.