HISTORY DEPARTMENT OF ZOOLOGY

1H E

OREGON STATE UNIVERSITY Department of Zoology

A History

b

Ernst I. Dornfeld

a i id

Charles E. King

Oregon State University Corvallis, Oregon 1989 Foreword

As the Department of Zoology at Oregon State University approached its 100th birth- day, it occurred to me that it was time to update the excellent History which had been completed by ErnstI.Dornfeld in 1966. For most of the time since Ernst's retirement in 1976 the department had been chaired by Charles E. King, and he appeared to be the most appropriate person to undertake this task. This was begun in the fall of 1988, and the resulting extended version of our history is what fills the following pages.

History buffs should know that this version omits some material included in the 1966 edition, but the omissions are minor. Copies of both publications might be sought through the department, the College of Science, the Department of Archives at OSU, or the main library on campus.

As we go to press, we feel that we are in an era of change in the State System of Higher Education, and we are uncertain as to the extent to which such change will affect us. Only time will tell, and therein lies the value of books such as this one. We hope you will discover within it all you hope to find, and more.

Christopher J. Bayne Chairperson Department of Zoology July 1989 Contents Part 1,1889-1966. 1 Part II, 1967- 1988 ...... 13

Appendices

IFaculty (1889-1988) ...... 25

IIAssociated Faculty ...... 28

III Graduate Degrees in Zoology ...... 30

IV Chairpersons of the Department of Zoology . . ..36 Part I. 1889- 1966 by Ernst J. Dornfeld

Corvallis College offered the first college-level curriculum about 1865 when the Rev. W. A. Finley became its president. Before this time, beginning with the incorporation of the college in 1858, the program of studies was that of a pre- paratory academy. The faculty of the collegiate division consisted of the Rev. W. A. Finley, A.M., President and Professor of Languages, and the Rev. Joseph Emery, Professor of Mathematics. The 1867-68 annual catalog of the college shows that first-term freshmen included in their course of study the subject of physiology, taught by Prof. Emery. In 1868 Corvallis College became the Agricul- tural College of the State, and Prof. Emery intro- duced first-term sophomores to a course entitled General Principles of Zoology, using Agassiz' textbook. The physiology and zoology courses Professor Ernst J. Dornfeld, Chairman, were part of the required "general curriculum"; in Depart,nent of Zoology, 1952-76 (photo, 1972). 1870 a course in entomology was added for the "agricultural curriculum." In this year the college acquired a 35-acre farm, which is now part of the zoology and entomology. Shortly, however, the present campus. Classroom instruction, however, latter were transferred to the tutelage of W. N. took place in a building near 5th and Madison Hull, A.M., Professor of Physiology and Drawing Streets, which served this purpose until 1889. (sic!). A similar oddity was the title of W. W. Under the new presidency of B. L. Arnold, Bristow, A.B., Professor of Book-Keeping and Bee A.M., in 1872, biological instruction was organ- Culture.The character of Prof. Hull's courses ized under the "School of Physics," and the 1872- can be gleaned from the following catalog de- 73 catalog describes the area as follows: scriptions (1888-89): "Here come to be considered the general principles "Physiology: The best method of studying the of life on its physical side. Vegetable and Mineral human body is undoubtedly by clinique, but since Physiology, Botany, Zoology. Text Books - Huxley, this is not possible except in medical schools, the Carpenter, Gray or Wood, Agassiz. The more im- next in excellence is by drawing and coloring. Every portant principles will be demonstrated by micro- student draws the vital parts of the body upon the scope. We hope to have a Spectroscope this year, blackboard, generally natural size, and recites from his own drawings. The value of this display power when spectroscopic analysis will be taught." and this sight knowledge cannot be overestimated. Apparently the spectroscope did not material- ize as expected, for the "we hope to have..." "Zoology:Here, again, when the animal forms statement appears in successive catalogs until cannot be secured, the graphic art is invaluable. 1876! Moreover, the courses in zoology, as well as Every student becomes a taxidermist, and therefore those in mineralogy and geology, remained in the possesses the power of preserving many valuable versatile hands of Prof. Emery until 1882. specimens. Insect and animal life in their relation to vegetableor plant life, particularly where they benefit In 1883 Edgar Grim, A.B., was appointed or destroy, should be generally studied." Professor of Chemistry and Agriculture, and succeeded Emery-jp charge of the courses in The year 1889 saw the removal of instruction

I .-1-- College Hall (now Bent on Hall), the first b,,ildi,i,'oji the present campus, completed 1888. Zoology occupied one, later two rooms on the third floor, 1889-1902. PhotocourstesyofUniversity Archives.

tI_ : il\

Laboratory and M useuni oft/ic Department ofZoology and Entomology in College ofUniversity Archives. Hall, 1893. Photo courtesy

2 from the original college building at 5th and and several courses in applied zoology and Madison Streets to the new and first building on applied entomology. Cordley saw to the enlarge- the present campus, the historic structure that is ment of the department staff, adding a succession now Benton Hall. of assistants and instructors. Included in the latter "The college building is new and commodious, were Fred M. McElfresh (1899-1901), William T. and is unsurpassed for beauty of situation." At Shaw (1901-07), John C. Bridwell (1907-1 0), Fred L. the same time, a Department of Zoology and Griffin (1908-1 0), Alice L. Edwards (191 0-16), and Entomology was organized, and placed in charge George F. Sykes (1910-21). Shaw prepared a large of Prof. F. L. Washburn, A.B. (Harvard), who was collection of mounted birds which is now in the also made first Entomologist of the Agricultural Homer Museum. Prof. Cordley was an entomolo- Experiment Station. The department quarters gist, and it was perhaps not accidental that this consisted of a single room which served as class- interest dominated the work of the department. room, laboratory, and museum, and was situated Thus, Harley Frost Wilson, who later became head on the third floor; another room was added a little of the entomology department at Wisconsin, was later. Prof. Washburn taught three courses: appointed assistant professor in 1911. General Zoology, Physiology, and Economic In 1902 the Department of Zoology and Ento- Entomology; in 1893 he added, as electives, mology was moved from its cramped two-room Comparative Anatomy, and Economic Ornithol- quarters in the Administration Building (Benton ogy. Prof. Hull's approach was materially modi- Hall) to the newly completed Agriculture Hall fled: (now Education Hall), which was built at a cost of $42,000. The department was installed on the "In General Zoology the student learns, by compar- third floor, where it occupied seven rooms (office, ing the structure of animals, the principles of classi- fication, familiarizes himself or herself with the life entomology research laboratory, student labora- histories of many, paying special attention to those tory, general laboratory, lecture room, storeroom, animals of economic importance. By drawing speci- and photographic darkroom); the fourth floor mens in the laboratory and writing original descrip- housed the museum. tions of the same, the student is trained to independ- When the School of Agriculture was established ent observation and thought. Textbook: Orton's in 1908, with Prof. Cordley as its first Dean, the Comparative Zoology. Department of Zoology and Entomology became a part of this unit. "In Physiology each student dissects in the labora- A third move took place in 1910, when the tory a typical mammal in order to get a general idea present Agriculture Hall (center unit) was com- of mammalian anatomy, and to better understand pleted. Again, the third floor seemed an appropri- references in textbooks. Drawings of these dissec- ate location, and nine rooms were made available tions are required. Laboratory work further consists (two offices, research laboratory for entomology, of demonstrations illustrating circulation of the physiology laboratory, general zoology laboratory, blood, composition of the blood, mechanism and chemistry of respiration, optical phenomena, reflex two lecture rooms, vault, and photographic action, etc., and the study of tissues with a micro- darkroom). The museum remained in its previous scope. Textbook: Martin's Human Body." quarters on the fourth floor of what was now renamed Science Hall (present Education Hall). Washburn served as Professor of Zoology and In 1914, a separate Department of Entomology Entomology until 1895, when he became Professor was established, and George F. Sykes, M.A. of Biology at the University of Oregon and subse- (Brown), was made Professor and Head of the quently (1902) State Entomologist of Minnesota. Department of Zoology and Physiology. During He was succeeded in the fall of 1895 by Arthur his administration there were the following Burton Cordley, B.S. (Mich. State), who in 1908 appointments to the faculty: Irving H. Blake (1913- also became the first Dean of the School of Agri- 16), Asa Chandler (1914-19), Elizabeth R. Cole culture. Under Cordley's head ship, which ex- (1915-16), Charlotte N. Hurd (1916-19), Howard tended to 1914, the course offerings of the depart- M. Wight (191 6-27), Aravilla Taylor (1920-21), and ment were expanded to include the following: Nathan Fasten (1920-44). Dr. Chandler in later Histology, Embryojogy, Systematic Zoology, years achieved eminence as a parasitologist and Parasitology, Evolution, Advanced Entomology, joined the Rice Institute. A number of new 3 Agriculture Hall (later Science Hall, ,iow Education Hall),completed in 1902. The DepartmentofZoology and Entomology occupied seven rooms on the third floor, 1902-ll0. Photo courtesyofUniversity Archives.

Agriculture Hall. Center unit completedin 1909; wings added in 1911 and 1913. Home of the Department of Zoology from 1910 to 1967. The Department first occupiednine rooms on the thi rd floorofthe center section; later additions included allof the second and third floors in the south wing. PhotocourtesyofUniversity Archives.

4 -S

Frederick L. Washburn (1860- Arthur 8. Cordley (1864-1 936), George F. Sykes (1878-?). Head Nathan Fasten (1887-1953), 1927), Head of Department Head of Department 1895-1914. of Department 1914-1921. The Headof Department 1921-1944. 1 889-1895. Department of Univ. Arch. Orange, ool 8 (1915). Photo hi1 E. J. Dornfeld (1940). Entomology. courses were added, among them Game Propaga- This situation and the economic depression of the tion (Sykes), Animal Parasites (Chandler), Proto- early 1930's blocked any significant progress, zoology (Chandler), Aquaticulture (Sykes), offered no prospect for the improvement of Taxidermy and Zoological Collecting (Wight), facilities, and prevented the development of a Genetics (Sykes; Fasten), and Animal Ecology stable staff. (Wight). During this crucial period, higher education in Nathan Fasten, Ph.D. (Wis.), who had been Oregon was completely reorganized and placed Instructor and Assistant Professor of Zoology at under a single State Board of Higher Education. A the University of Washington from 1914 to 1920, system of allocations was devised under which replaced Prof. Sykes as Head of the Department of Oregon State College was designated as the center Zoology and Physiology in 1921. The following for basic and applied sciences. The School of year the department became part of the newly Science resulted (1932) and major curricula were organized School of Basic Arts & Sciences which authorized for its several departments. While this endured until the foundation of the School of important step laid the "legal" foundation for Science in 1932. The name of the department was expansion, the slow recovery from the economic shortened in 1927 to Department of Zoology. depression hampered progress. The implementa- While an undergraduate major curriculum was tion of the reorganization of higher education not authorized before 1932, graduate study was included transfer of faculty between the Univer- possible and the first M.S. degree was awarded in sity at Eugene and the State College at Corvallis. 1923 to Walter P. Covell. In this manner the Department of Zoology ac- Faculty appointments between 1921 and 1932 quired Prof. Arthur Russell Moore and Prof. included James E. Lynch (1921-22), Florence S. Rosalind Wulzen. Both Moore (1932-33) and Hague (1921-26), Wilber D. Courtney (1922-30), Wulzen (1933-47) had been students at the Univer- John L. Osborn (1923-47), Bess R. Green (1926-29), sity of California of the distinguished physiologist, Kenneth L. Gordon (1927-69), Kimber C. Kuster Jacques Loeb. (1926-27), Edith L. Benedict (1929-38), and William The first bachelor's degree in zoologywas B. Woen (1930-31). James Lynch later became awarded in 1932 to Wendell Lee Ball. In 1935 Professor of Fisheries at Washington, Florence Alfred Taylor received the first Ph.D. and was Hague became Chairman of Biology at Sweet Briar appointed instructor (1935-40). Other staff addi- College, and William Owen became Professor of tions under the leadership of Nathan Fasten Zoology at Wyoming. included Ernst J. Dornfeld (1938-76), Grant A. This was a difficult decade, and one not condu- Swan (1939-43), Delmar I. Ailman (1939-65), and cive to departmental growth. The School of Basic Clifford Grobstein (1940-43). Arts & Sciences was "service" oriented and not Under the leadership of the new Dean of the authorized to offer-major departmental curricula. School of Science, F. A. Gilfillan (1938), and of

5 President A. L. Strand who took office in 1942, scopes for the laboratories of embryology, parasi- strong encouragement was given to the improve- tology, and cell biology. Space shortages on the ment of scientific instruction and staff engage- campus were met by the erection of "temporary" ment in research. The policies were welcomed by quonset huts, Butler huts, and an assortment of "the young men in a hurry" who responded by surplus army barracks. In this way the depart- developing courses of rigorous quality, recruiting ment acquired, in 1948, an aluminized Butler hut graduate students, and engaging in productive to house the museum collections and laboratories investigations. Facilities and teaching loads left for vertebrate natural history and ecology. Its much to be desired, but the blessings of NSF and temporary status proved rather enduring. NIH were still in the future. Research in the By 1950 critical space needs also existed in Department of Zoology during these years in- physiology which had expanded beyond any cluded the discovery and exploration of a nutri- ability to share rooms with invertebrate zoology tional anti-stiffness factor (Wulzen), the behavior and parasitology. At this moment the President's and learning responses of ground squirrels residence was about to be vacated and demol- (Gordon), the developmental cytology of the ished. It was an old frame house that had served ovary (Dornfeld), endocrine aspects of fin regen- its purpose and stood in the way of planned site eration in fishes (Grobstein). improvements. The residence was generously The years of World War 11(1942-45) affected the loaned to the Department of Zoology, and quite progress that had been achieved. Class enroll- appropriately, too, quipped the President, since ments dwindled as young men were called into the rats were already there. The physiologists military service. In the fall of 1944 the students found the mansion roomy and adaptable. The served by the Zoology Department numbered 292; cellar was good for bulk storage and experimental the previous high, in 1941, was 627. There were nimals; the living room and dining room became but two undergraduate majors. Staff members class laboratories; bathtubs made excellent frog, volunteered for additional instructional duties in fish, and crayfish tanks; bedrooms were fine for the Army Specialized Training Program. Gordon ergometer and basal metabolism studies, as well taught geography, Dornfeld and Grobstein taught as for offices. physics. In 1943 Grobstein left to accept a commis- Further faculty appointments were made in sion with the air force. 1948, 1949, and 1950. Dr. Wuizen having retired, With cessation of hostilities in 1945 a new era Hugo M. Krueger, Ph.D. (Mich.), who previously began. Prof. Kenneth L. Gordon was appointed held positions at Michigan and St. Louis Univer- Chairman of the Department of Zoology, replac- sity, and had just spent two years at Beirut, was ing Nathan Fasten who resigned from the faculty. made Professor of Physiology in 1948. The same The return of veterans swelled the enrollment, year, Robert M. Storm, who completed his Ph.D. which reached 814 in the fall of 1946. Addition of as a student of Prof. Gordon, was appointed staff and space were imperative. In this year Ivan Instructor and taught courses in ornithology, Pratt, Ph.D. (Wis.) and Howard H. Hillemann, mammalogy, and herpetology. In 1949 Carl L. Ph.D. (Wis.) were appointed assistant professors. Anderson, Dr. P. H. (Mich.), Professor of Hygiene A specialist in parasitology, Dr. Pratt was to & Health Education in the Division of Physical develop this area and invertebrate zoology. Dr. Education, was added to the Zoology staff on a Hillemann, an embryologist, was selected to part-time basis to teach a course in human biol- succeed Prof. Osborn whose retirement was ogy. Max W. DeLaubenfels, Ph.D. (Stanford), a imminent. The faculty, now consisting of Drs. widely known authority on the systematics of Gordon, Allman, Dornfeld, Hillemann, Osborn, sponges, was brought from the University of Pratt, and Wulzen, immediately addressed their Hawaii to become Professor of Zoology in 1950. collective efforts to developing sound policies of Prof. deLaubenfels took charge of the freshman departmental operation, curricular standards, and course in general zoology. graduate study. These served as a foundation for The postwar influx of students was at the orderly growth in the years ahead. graduate as well as undergraduate level. In the The great increase of students brought state spring of 1951, 29 graduate majors were enrolled. funds for the purchase of much needed laboratory Over a three-year period, 29 Master's degrees and equipment, particularly full sets of good micro- two Ph.D.'s. were conferred. 6 Zoology Staff, May 1947. Upper row (1. to r.): Samuel F. Toevs (grad. asst.), Robert M. Storm (grad. asst.), Walter S. Vincent (grad. asst.), Virginia Weimer (grad. asst.), Amelia Jaramillo (secretary), Samuel W. Lesher

(grad. asst.). Lower row: ErnstI.Dornfeld, Howard H. Hil!emann, John L. Osborn, Ivan Pratt, Kenneth L. Gordon, Rosalind Wuizen, Delmar I. Ailman. 15

Zoology Faculty, June 1955. Standing (1. to r.): Kt'u,,eth L. Gordon, Howard H. Hzlle,nann, Ernst J. Dorufeld, Austin W. Pritchard, Robert M. Storm. Seated: Delman I. Ailman, Hugo M. Krueger, Rosalind WuI:e'n, Carl L. Anderson, Max W. de Lauhenfels.

7 In 1952, after serving eight years as chairman, brief illness in February of 1958, necessitated his Dr. Gordon relinquished this office and was replacement in the fall. Frederick L. Hisaw, Jr., succeeded by Prof. Ernst J. Dornfeld. The depart- Ph.D. (Harvard), was appointed to take charge of ment was now in a period of rapid growth. the large freshman course and to develop gradu- Because of the part-time affiliation of Dr. Krueger ate work in endocrinology. The following year with the Department of Animal Husbandry and Cyrus Mayshark,H.S.D. (Indiana), inherited the the need for expanding physiology, Austin W. course in human biology from Prof. Carl Ander- Pritchard, Ph.D. (Hawaii) was added to the faculty son, whose increased responsibilities in the in 1953. Dr. Pritchard took charge of the introduc- Division of Physical Education required his tory course in physidlogy and developed ad- withdrawal from the zoology staff. vanced instruction in general and comparative The School of Science had some yearsprevi- physiology. Prof. Krueger's specialty became ously acquired an electron microscope, but its mammalian physiology. effective use suffered for lack of a qualified person Alfred Owczarzak, Ph.D. (Wis.) arrived in 1955 to take full-time charge and to train potential to work with Prof. Dornfeld as research associate. investigators in its operation. In 1960 James D. His appointment to the instructional staff two Newstead, who was pursuing doctoral research years later permitted expansion in cellular biol- under Prof. Dornfeld and had special training in ogy. Dr. Owczarzak took over the courses in the electron microscopy laboratory of Dr. H. histology, microtechnique, and experimental Stanley Bennett in Seattle, was given this responsi- embryology, and initiated formal instruction in bility and appointed instructor. histochemistry and tissue culture. The spring of 1962 brought from the University An important step in 1955 was the addition of a administration the long-sought approval to geneticist. A beginning course in this subject had p1roceed with the planning of a new building for been taught since 1914 and parallel courses had the department, the first addition to Cordley Hall. sprung up in a number of other departments. In Calculations and projections indicated that a four- order to consolidate these, as recommended by the story building of about 60,000 net square feet was inter-departmental genetics committee, and required. Another story was to be added to strengthen the field by adding graduate instruc- provide additional research space for the Depart- tion, the appointment of James D. Mohler, Ph.D. ments of Botany and Entomology. The prelimi- (Calif.), was made. Facilities for Dr. Mohler's nary plans for this building were prepared during work were duly installed in the President's the summer, together with grant proposals for mansion where his flies joined the menagerie of federal aid to supplement the anticipated state the physiologists. His course in theoretical funds. Drs. Dornfeld and Owczarzak carried the genetics became a basic contribution to the gradu- principal responsibility for this work, coordinating ate programs in genetics. the requirements of the staff, and translating these In 1957 the first unit of Cord ley Hall, the new into specifications and drawings. The architec- building for the biological sciences, wascom- tural work was assigned to the Portland firm of pleted. While it was originally hoped that this Bear, McNeil, Schneider, Bloodworth & Hawes unit would house the Departments of Botany and The fall of 1962 saw the faculty addition of Zoology, space did not permit and the building Ronald H. Alvarado, Ph.D. (Wash. State) to whom was assigned to Botany, Horticulture, and Ento- fell the task of organizing a physiology course mology. As the Department of Botany & Plant suitable to the needs of pharmacy students and Pathology had previously occupied the second other advanced undergraduates. Dr. Alvarado floor in the south wing of Agriculture Hall, this also shared the instruction of general zoology with space was now available for Zoology. Since there Dr. Hisaw as enrollment in this course had was also pressure to demolish the President's old climbed from 255 to 547 in two years. On the residence, physiology and genetics moved backto graduate level, Dr. Alvarado directed work on these quarters. The contiguity with other zoologi- membrane permeability and ion transfer mecha- cal laboratories had advantages, but the facilities nisms. offered by Agriculture Hall were painfully obso- The same year the department set up an lete and no net gain of space was achieved instructorship with the responsibility of supervis- The death of Prof. de Laubenfels, followinga ing and coordinating the freshman laboratories,

E1 Zoology Faculty, June 1966. Standi,o' (1. to r.): John K. Ellis, La Von C. Johnson, James D. Mohier, Alfred Owczarzak, Ronal H. Alvarado, Austin W. Pritchard, Frederick L. Hisaw, Thomas D. Dnrrozi', Patricia J. Jarris. Seated (I. to r.): Robert M. Storm, Hugo M. Krueger, Howard H. Hillemanu, Kenneth L. Gordon, Ernst I. Dornfc'ld, Ivan Pratt. which now ran into 29 sections. Given to an He was replaced in the Zoology Department by experienced graduate assistant in the last year of John K. Ellis, Ph.D. (Mich.). Also, Prof. Allman, his doctoral work, this appointment was awarded who for many years taught human anatomy and to David E. Kerley, and the following years to applied physiology, reached retirement age and Glenn A. Anderson, John C. Belton, Thomas D. was succeeded by La Von C. Johnson, M.A. (Utah). Darrow, and Eugene C. Fuller. A similar position, Of foremost concern to the department was, of for coordination of the physiology laboratory course, the new building, the preliminary plans of sections, was instituted in 1966, and first held by which were drawn in the summer of 1962. In the George S. Aispach. spring of 1963 the State Legislature gave approval During his sabbatical year of research at the for its construction, and the National Science University of Syracuse, Dr. Mohler's duties were Foundation provided a matching grant of $425,000 temporarily assumed by Lester J. Newman, Ph.D. which was subsequently increased to $446,000. (Wash. Univ., St. Louis), who subsequently joined Unhappily, however, a tax referendum in October the Biology faculty of Portland State College. cancelled the higher education building program, James Newstead had resigned as electron and the whole matter was put off for a biennium. microscopist in 1963 in order to pursue postdoc- Success finally came in the legislative session of toral study. His successor, appointed in 1964 as 1965, and an additional grant of $190,000 was Assistant Professor of Zoology, was Patricia J. received from the National Institutes of Health; Harris, Ph.D. (Calif.). Dr. Harris' research on the the Higher Education Facilities Program (Title I) ultrastructure of the mitotic apparatus at Berkeley also contributed $364,777. By the time of bid had gained international attention. Within a year opening, December 7th, construction costs had Dr. Harris was successful in obtaining grant funds risen so high that there was much apprehension. for a second electron microscope and support for a Surprisingly and to everyone's relief, the bids research program on the structural analysis of cell came below the architects' estimates. The building division. was completed in 1967 at a cost of $3,137,0J0 (State In 1965 Cyrus Mayshark, who had been in of Oregon $2,136,223; federal grants $1,000,777). It charge of the course in human biology, accepted a has gross square footage of 125,106 and a net position at the University of Tennessee as Head of footage of 73,547. the Department tf Hygiene and Health Education. Also of importance to the department was the

9 .7/7 I./_ )- _L_./

The New Addition" to Cordley Hall, completed in 1968. The DepartmentofZoology occupies the first, second, third, and fifth floors. This view is from the East. completion in 1965 of the Marine Science Center at research on the cytogenetics and developmental Newport, located on the south shore of Yaquina genetics of Diptera. Bay. This facility of the University, operating on a Student enrollment in the department stood at twelve-month basis with Dr. Joel W. Hedgpeth as 814 in 1946. A temporary decline followed the the first resident director, has both laboratory and disappearance of the G.I. bill students, but by 1956 field research facilities as well as 38,000 square feet the loss had been regained. Five years later, in the of floor space and a dock to accommodate the fall of 1961, the children of the war veterans OSU oceanographic vessel. arrived and the enrollment surged sharply up- Further additions to the Zoology faculty were ward. This trend continued unabated with 2,187 made in 1966. Since Dr. Pratt's program in students enrolled in zoology classes in the fall of parasitology had grown to consume all of his time, 1965. Also between 1961 and 1965 the number of Richard L. Miller, Ph.D. (Chicago), was appointed undergraduate zoology majors climbed from 22 to to take over the work in invertebrate zoology. 167, graduate majors from 31 to 61. Likewise, the load of Dr. Storm required relief, The number of higher degrees awarded in and John A. Wiens, Ph.D. (Wis.), was brought in to Zoology mounted rapidly after 1948. By 1966 a assume responsibility for ecology and ornithology. total of 113 master's degrees and 57 Ph.D.'s had Both also joined the lecture team in general been conferred. zoology and made it possible for the department Departmental research activity, closely en- to grant "released time" to one staff member each meshed with the program of graduate instruction, term for research at the marine laboratory. has been decisively assisted and accelerated by Prof. Mohier, who for eleven years had capably numerous grants from federal, state, and private managed the area of genetics, was offered a agencies, particularly the National Science Foun- position at the University of Iowa in the fall of dation and the Public Health Service. Through 1966, which he decided to accept. His place was these grants the acquisition of research equipment filled by Paul A. Roberts, Ph.D. (Chicago), who has been substantial, and funds have been pro- since 1962 had been on the staff of the Oak Ridge vided for the salaries of research assistants and National Laboratory where he was engaged in associates. Further help for graduate students has

10 come from an increasing number of teaching awards for outstanding teaching and research. assistantships and from fellowships and trainee- Important work has been performed by depart- ships (NSF, PHS, Rockefeller Foundation, NDEA, mental committees. It has not been possible to and other agencies). detail the valuable contributions of the long list of While space does not permit a full account of graduate teaching and research assistants, or to past and present research programs in the depart- comment on the careers of the graduates who are ment, it may be noted that during the twenty-year scattered throughout the country and in foreign period from 1946 to 1965 the published results lands. Many of the latter have made distinguished appeared in 271 articles, as opposed to about 40 records. Nor would the department have been during the preceding two decades. built without the loyal services of competent This brief history of the Department of Zoology secretaries. has been essentially a chronology. In compressing The centennial of the University will coincide the span of almost eighty years into a few pages, with the first year of the department in Cordley much has had to be omitted. Various members of Hall. To paraphrase the remark of 1889 concern- the staff have held offices in national and regional ing Zoology's first home, "the building is new and professional organizations and have served on commodious, and is unsurpassed for excellence of scientific advisory and editorial committees. Inter- its facilities." It will make an auspicious starting national meetings and sabbaticals have involved point for the next history of the department. J the faculty in foreign travel. There have been

7

11 Part II. 1967-1988 by Charles E. King

Preface

Departments, like organisms, evolve and contributions made by my colleagues to bothour Zoology at Oregon State University (OSU) in 1988 science and our students. And it is also clear that is not what it was in 1967. The history of a depart- these contributions are a two-way street for the ment is a Lamarckian record of cultural evolution success of the department, in large measure, containing obvious analogies to the forces that reflects the success of its students, bothpast and drive biological evolution. Several of the faculty present. lineages in our department have a long history of As Ernst Dornfeld stated in Part I,a brief descent with change. The departments first cell history is essentially a chronology in whichmuch biology position was held by Arthur Cordley at information is omitted. Because Iam dealing with the turn of the century. This position has subse- 22 rather than 78 years, I have also attemptedto quently been occupied by George Sykes (1910- give some impressions of the academic climate 1921), Nathan Fasten (1921-1944), Ernst Dornfeld during this period. This climate includes changes (1938-1976), Robert Hard (1978-1985), and Robert in the discipline as a whole as wellas modifica- Gimlich (1988-present). Radiating from this tions of the federal, state, and privateresources lineage have been other cell biology positions such available to support the department. I have also as those occupied by Alfred Owczarzak (1955- attempted to give more detail on faculty interests 1983), Patricia Harris (1964-1973), and Frank Conte and activities than was possible in Part I. (1963-present). Other lineages have been greatly On behalf of the entire department, faculty and modified as time has passed, and still others are students alike, a number of acknowledgmentsare now extinct. in order. First and foremost, none of this history The selection that drives this process of cultural would have taken place without the continuing evolution is carried out both at the level of the support of the Oregon taxpayer. However, as will College of Science, the immediate source of all be made clear, major developments have also positions, and at the level of the department where reflected the generosity of many alumni and the chairman and the faculty jointly track develop- friends of the department. Prominent on this list ing areas of zoology. The most obvious quantita- are funds the department has received from the tive aspect of a departmentthe number of Braley Estate and for the Wayne and Gladys facultyis determined by the university admini- Valley Chair in Marine Biology. Financial support stration. The dominant qualitative aspectthe has also been received from the department's distribution of subject-area specialties within the faculty--for instance, the current OSU Foundation departmentis usually determined by the current report of donors lists contributions by approxi- faculty. A large stochastic factor also influences mately one-half of the faculty present during the the structure of a department since it is impossible 1987-88 academic year. Particularly notable is the to predict accurately the course of individual gift from our emeritus colleague and his wife, Pro- careers or the vicissitudes of federal and state fessor and Mrs. Hugo Krueger, of the Wulzen- funding. We will see each of these factors at work Krueger Fellowship for a graduate student in in the following pages. physiology. And finally, the department's devel- Updating the departmental history at this time opment and atmosphere also reflect the dedicated seems particularly appropriate since 1989 will services of its staff. In this category, I would rec- mark the start of the second century of the Depart- ognize the many and varied contributions of its ment of Zoology at Oregon State University. In head secretariesClaudia Hamren (1978-80), truth, although I gained an appreciation for the Penny Miller Hardesty (1980-81)and its manage- value of a departmental history while I was ment assistant, Pam McAllister (1979-1987). chairman of the Department of Zoology, I did not Particular gratitude is due Dianne Rowe (1981- look forward to the task of preparing this update. present) who first as head secretary, and then as To my surprise the experience has been an enjoy- administrative assistant, has continuously able one. Too often we take our surroundings for breathed life, spirit, and joy into our day-to-day granted. In gathepng the material for this update, professional lives. J I have renewed mappreciation of the many

13 Faculty

Update on Faculty Covered in Part I New Faculty Three of the faculty chronicled in the first part To expand the area of cellular and molecular of this history are still active. Fred Hisaw* has biology at OSU, three new faculty positions were continued to play an important role in the depart- established by the College of Science in 1963. One ment's non-major physiology instructional pro- of these--cell physiology--was awarded to Frank gram. He teaches the two-course sequence in P. Conte (Ph.D., University of California, mammalian physiology that serves students in Berkeley). Temporary facilities were found for pharmacy and the pie-professional health sciences Frank's research in Weniger Hall under the aegis and one of the three terms of our human anatomy of the Department of General Science. When the and physiology sequence that is required for stu- Cordley Hall addition was occupied in 1967, his dents majoring in health and physical education. faculty position was transferred to Zoology. His Austin Pritchard teaches the other two terms of research has involved investigations of the mecha- the non-major's human anatomy and physiology nisms of salt secretion by epithelial cells found in sequence and shares instruction of the major's animals living in saline habitats, particularly brine physiology course and the 15-credit Marine shrimp and brine flies. This research has taken Biology course taught at the Hatfield Marine Frank to marine and saline lake stations in Japan, Science Center each spring. In addition to these Africa, Italy, France, Australia, and Spain. He has duties, Austin serves as head adviser to the also served as a program director for regulatory Zoology undergraduates and faculty adviser of biology at the National Science Foundation and the Zoology Club. He is currently Assistant Chair- been an invited member of the National Research man of the department. Council Mono Lake Basin Ecosystem Study Paul Roberts was appointed to the department committee. just as Part I of the history came to a close. His In 1967 Paul Roberts of Zoology joined with instructional duties include genetics, evolution, fellow geneticists in Botany and Microbiology to and genetics of organisms. His research on appeal for more support of genetics by the College Drosophilacytogenetics has recently been ex- of Science. Two positions were obtainedone for panded into the area of molecular genetics. Paul is a molecular geneticist to be housed in the Depart- also interested in basic determinants of senescence ment of Biochemistry and a second for a popula- and has conducted National Institutes of Health- tion geneticist to be housed in the Department of sponsored research in experimental gerontology Zoology. using fruit flies as model organisms. Successful search strategies sometimes subsume Seven of the faculty active when Part I ended serendipity. In this search the serendipity oc- have subsequently retired, or died while still curred when a six-year-old girl pointed her finger active. Dates for Ernst Dornfeld, Kenneth Gordon, at a twenty-foot pfle of excavation dirt and said: Howard Hilleman, and Ivan Pratt are listed in Ap- "Daddy, look at that mountain!" "At that point," pendix I. These valued colleagues have since died. recounts Seattle native Peter S. Dawson (Ph.D., Happily, Bob Storm, Alfred Owczarzak, and Hugo University of California, Berkeley), "I knew it was Krueger, although retired, still live in Corvallis time to leave central Illinois." and make regular appearances in the department. Pete's research, which has received generous The remaining active faculty covered in Part I support from the National Science Foundation, have, with one exception, left OSU. Jeff Gonor focuses on genetic factors influencing the outcome continues as a full-time Professor and Head of competition between two species of flour Adviser in the College of Oceanography. Richard beetles. Although his graduate study at Berkeley Miller is now at Temple University, Ron Alvarado was in the Department of Genetics, his approach is at Arizona State University, Pat Harris is at the to this problem broadened to include ecological University of Oregon, and John Wiens went to the factors while he was at the University of Illinois. University of New Mexico and has subsequently This resulted from his association with Charles moved to Colorado State University. King (see below) and was continued after his arrival at Oregon State University in 1969 through Faculty active at the end of J988 are first indicated in extensive interaction with John Wiens. One of the boldface print. 14 major benefits of this interaction was the establish- versity of Michigan. His studies were focused on ment of a new course in Population Biology which various aspects of the ecology and physiology of continues to occupy a central position in the gastropod molluscs (snails and slugs). However, department's advanced undergraduate curricu- after joining our department Chris undertook joint lum. Pete is currently investigating the effects of research with Alfred Owczarzak on a study of the different environments in modifying the course of response of a freshwater snail to infection with natural selection both within species and between blood flukes and then, more broadly, his research the two competing species. became focused on the comparative immunology Following the departure of Richard Miller for of compatibility and resistance in host-parasite Temple University in 1968, the department sought systems. Such changes are encouraged for they to rebuild its strength in marine biology and ad- permit a department to track the rapid shifts in a vertised broadly for a replacement to fill this scientific area; were this not so, readjustment of position. Among the applicants was John E. faculty expertise to recent developments would Morris (Ph.D., UCLA) who was completing a only occur when new faculty were hired. postdoctoral research program at the University of In another context, the re-focusing of Chris' Chicago following two years as a postdoctoral research to problems associated with parasite-host fellow at Wenner Gren's Institute in Stockholm. interactions followed the tradition of his predeces- Although John had conducted research using sor in this position, Ivan Pratt. In 1965, when the marine invertebrates, his central expertise was in Marine Science Center opened, Ivan shifted most cellular aspects of developmental biology. Fortu- of his instructional and research activities to the nately, the department recognized a rare opportu- coast. At about the same time the field of parasi- nity and added John to its faculty. His subsequent tology underwent a great reorientation. The new "marine" research has been on the cell biology of emphasis was less descriptive and focused on retina development in chickens and on the interac- functional questions such as "How are parasites tions between the cell surface of embryos and the able to escape the defense systems of their hosts?" uterine epithelium in mice. He has concentrated and "How are hosts able to decrease the damage on the role of a class of molecules, the proteogly- done by their parasites?" Both are fundamental cans, involved in the interaction of the cell surface questions in the field of immunoparasitology, and with the extracellular matrix. the department is fortunate to have Chris repre- John has been one of the most active faculty senting this important area. members in departmental governance. He has a Since joining our faculty in 1971, Chris Bayne particular talent for stimulating his colleagues to has brought over $1,100,000 of federal research consider issues more carefully. His departmental monies to the University and State of Oregon. service includes the positions of Assistant Chair- Over the same period, his total salary from the man and Acting Chairman of Zoology. John also state has been less than half this amount. Similar maintains a high profile in university affairs, statistics could be gathered for several other fac- having served as Vice President and President of ulty who are now active in the department. On a the OSU Chapter of the American Association of department-wide basis, the information presented University Professors. In addition to continuing below is a record in which we can take pride. his research, which has been funded by both the National Science Foundation and National Insti- Total Total tutes of Health, John is currently Associate Dean Year Faculty SalariesGrant Expenditures of the College of Science. 78/79 $ 425.590 $ 455,618 In 1970 John Morris was assigned more appro- 79/80 472.777 359.667 80/81 419,365 478,618 priate duties in the area of developmental biology, 81/82 452,401 502,290 and the department again decided to search for a 82/83 452,106 893.939 marine invertebrate biologist. This time the search 83/84 461,374 479,586 84/85 427,735 343.481 led to the hiring in 1971 of Christopher J. Bayne 85/86 462.004 518,165 (Ph.D., University of Wales). Following comple- 86/87 521,024 537,556 tion of his Ph.D., Chris participated in a Royal 87/88 580,022 610,742 Society expedition,..to Aldabra in the Indian Ocean $4,701,398 $5.1 79,662 and undertook postdoctoral research at the Uni- 15 Obviously, from an economic standpoint it exceptional productivity, the Dean of the College would be difficult to imagine a better investment of Science has increased their employment to two for Oregon than more growth of this type. full-time positions. With the shift of Chris Bayne into cellular With the retirement of Howard Hilleman in aspects of parasitology, the department's staffing 1975, instructional needs were created in theareas needs in marine biology again returned to the of comparative vertebrate anatomy and embry- forefront. In response, Marjorie Reaka (Ph.D., ology. John A. Ruben (Ph.D., University of University of California, Berkeley) was employed California, Berkeley) was selected as Howard's in 1975. She remained for one year and then left replacement. John's professional background is for the University of Maryland. In 1976 the unusually diverse. After completing a bachelor's department recruited a married couple, Bruce degree in wildlife management at Humboldt State Menge (Ph.D., University of Washington) and College, he obtained a master's degree in paleon- Jane Lubchenco (Ph.D., Harvard University), to tology at Berkeley. He then moved to Berkeley's fill the position in marine biology. Husband-wife Zoology Department for his doctoral research teams in academic life have historically had where he combined environmental physiology difficulties in securing suitable positions for both and functional morphology in a study of snakes. individuals. The traditional answer to this di- This diversity has continued to be reflected in lemma has been for one member, almost always John's research and teaching. Since joining the the wife, either to drop out of science or to accept a OSU Zoology Department, John has conducted less appropriate position (e.g., research assistant, NSF-sponsored research on morphological evolu- part-time lecturer, etc.). Both Bruce and Jane fully tion in snakes, U.S. Park Service-sponsored met the advertised criteria for the position in research on the John Day Fossil Beds in eastern marine biology, and both were highly desirable Oregon, an NSF-funded project on the effects of appointees. Therefore, a new arrangement for ash from the Mount St. Helens eruption on salmon OSU tenure-track faculty was set up. Both were smolt, and a Sea Grant investigation of the biology appointed independently to half-time positions of exercise-induced hypercalcemia in fish. John with the understanding that they would be teaches Natural History of the Vertebrates, Com- independently evaluated and each would be parative Anatomy, and Physiological Ecology. assigned half the normal duties of a full-time When John Wiens left for New Mexico in 1978, faculty member. This arrangement has been to the a substantial gap was created in the department's great advantage of the Department of Zoology for vertebrate ecology staff. The advertised position we have gained two productive colleagues for the drew over 200 applications, and it was with great "price" of one. The primary benefit to Bruce and difficulty that the search committee reduced this Jane, in Bruces words, is that "both of uscan number to a short list of ten individuals. Five of teach, do research, have equivalent tenure-track these ten were invited to campus. After extensive positions in the same institution, AND have interviews, Andrew R. Blaustein (Ph.D., Univer- sufficient time to be intimately involved in parent- sity of California, Santa Barbara), who had served ing our two sons." the department as an instructor for General Because of their extraordinary talents and zeal Zoology, was appointed to the position. for scholarly research, Jane and Bruce have joined Andy's doctoral research had been a field study the ranks of the world's leaders in experimental of rodents in which he had found that competition marine ecology. Bruce's research focus is the influences their spatial, temporal, and reproduc- determination of ecological forces producing tive patterns. Shortly after joining our depart- community structure in rocky intertidal systems. ment, he initiated joint research with his postdoc- His concentration tends to be on prey-predator toral associate, Richard OHara, who had noticed relationships. Jane pursues similar problemsat that the larvae of certain frog species formaggre- the population level by studying marine plant- gations in mountain lakes. Starting with this ob- herbivore interactions. Her most recent work has servation, he demonstrated that these aggrega- been an investigation of plant chemicals, so-called tions were probably composed primarily of secondary compounds, that possess anti-herbivore siblings, and that individuals were able torecog- effects and provide at least one explanation of nize one another by their degree of relatedness. why the world is green. In recognition of their Andy has found that an animal reared in total 16 . k i yr ' 4lSI I A 4 - I (I. I . i1lIHI: / / /i?i I faculty, Juize 1989. Top row, ). Ro!'erl I /,1,1' Zoology IV. I'ri( /iiniI, ,(I('/I( h I 1 1( I'JI)I1(I0/1/. All/lull. hoot. A1nfII lIjjio', :1)111(0'taicljnc,' R HIlllI('l)I, ( / ( ', Kmc', IiuI I uin/ l'.Coule, . Morr, A. RoI'pls, Jl((./i/i/innII'lll'/I, ))III!1,I AJn//iooii (onI/iia:1 In flu Ill'.'!to I /f//',,, liii i;,A / A (I, I') /0 j)/' A A 11)l(. /'/fjp II. I 1!oi,'//.o,j '.1/'/'OIIuII/'/'r'Ill l'S In n.! ,/i', / hI BIll)!, /11;! A. °. i ' isolation can recognize as kin both sibs and half- volved formulating computer simulation models sibs it has never seen before. This research has of zooplankton population growth at the Center helped to resolve a major scholarly debate among for Quantitative Studies of the University of evolutionary and behavioral biologists. While the Washington. Following completion of this re- ability of some organisms to recognize kin has search, I spent a year as an instructor at Yale been known for some time, it was not known University and then went to the University of whether this recognition was based on learned or Illinois as an assistant professor. While there I met innate characteristics. Andy's research suggests another young faculty member in Zoology, Pete that some species have a strong genetic compo- Dawson. For two years, during shared daily nent involved in their kin recognition system. His lunches of canned soup and saltines, I learned research has been sponsored by three separate population genetics from Pete while he leArned National Science Foundation grants as well as an population ecology from me. This experience award from the National Geographic Society. profoundly influenced our future paths in both Currently Andy is in the process of testing several teaching and research. For instance, when I hypotheses concerning the adaptive value of kin accepted subsequent faculty positions at Yale recognition in larval amphibians. University, the University of South Florida, and When Ernst Dornfeld retired in 1976, he left two OSU, my courses were either in genetics or had a positions vacant that of cell biology faculty strong genetic orientation. member and department chairman. The cell My research combines population genetics and biology position was filled in 1978 with the ecology to quantify the population structure and employment of Robert Hard (Ph.D., State Univer- evolutionary dynamics of rotifer populations. I sity of New York at Albany), a specialist in cilia am particularly interested in how these small structure and function. Bob remained with us zpoplankton adapt to seasonal variation in the until 1985 when he was offered a position at the physical and biological environments of their medical school of the State University of New lakes and ponds. In addition to National Science York at Buffalo. His replacement, Robert Gimlich Foundation support, I have obtained support also (Ph.D., University of California, Berkeley), joined from the National Institutes of Health to pursue the department in the Fall of 1988 following post- my research interests in the genetics of ageing. I doctoral fellowships in cell biology at Baylor am currently using mitochondrial DNA analysis College of Medicine and Scripps Clinic and to characterize gene flow in rotifers. Research Foundation. Bob studies the cell biology My discussions with Dean Robert Krauss of the of early development in vertebrates using em- College of Science during negotiations over the bryos of a southern African frog as his experimen- chairmanship centered around the need for tal organism. He is particularly interested in the Zoology to develop a focus in cell biology that operation of cell to cell signalling mechanisms would parallel the strong group that had been during embryonic induction of the larval muscula- assembled in ecological and evolutionary biology. ture and central nervous system. Using morpho- As a result of these discussions, two new positions logical, pharmacological, and molecular genetic were added to the department. In the fall of 1978 approaches to study intracellular signal transduc- the Department held a weekend retreat at the tion and cell-cell communication via specialized Menucha Conference Center in the Columbia junctions, Bob is attempting to delineate early Gorge. One of the major topics of this meeting events during development that determine was how best to use the new positions. After patterns of cell fates. extensive discussion of both the department's Since Ernst had been chairman of the depart- instructional needs and research specialties of cell ment for 25 years, both the dean of the college and biology that held great promise for major concep- the faculty of the department thought it appropri- tual progress, two areas were selected for the new ate to bring in a replacement from outside the appointments: neurobiology and behavioral university with a new perspective. The position endocrinology. was announced nationally and in 1977, after an The first of these positions was filled by Philip extensive search, I (Charles E. King, Ph.D., H. Brownell (Ph.D., University of California, University of Washington) was named the new Riverside). Following completion of his doctoral chairman. My postdoctoral work, in 1965, in- work on the neurobiology of prey localization by 18 scorpions, Phil moved to the University of Califor- has published an article entitled "Look out for nia, San Francisco, where he held Sloan Founda- Newts" in that favorite journal of all grade school tion and National Institutes of Health postdoctoral students:Ranger Rick.With such foresight in fellowships to study neural control of cardiovis- recruiting, it seems likely that Frank will continue ceral function in the sea hareAplysia. Thiswork to maintain a laboratory that is bulging at its was continued, again with NIH support, after his seams with bright young students and post-docs. arrival at OSU in the fall of 1979. Phil has recently When faculty are added to a department, they returned to his studies relating sensory physiology are expected to participate in the education of the to behavior in scorpions. He has found that the next generation of scholars by serving as major pectines, comblike structures of hitherto unknown professors to graduate students. Collectively, the function, are actually chemosensory organs that five assistant professors employed from 1976 to function in prey and predator detection and also I 979Drs. Blaustein, Brownell, Lubchenco, in mating behavior. This research is being spon- Menge, and Moorehave graduated 17 students sored by the National Science Foundation. with the M.S. degree and 12 students with the An appointee for the second new position was Ph.D. Doctoral students graduating from these found closer to home. Frank L. Moore (Ph.D., five laboratories have gone on to become faculty University of Colorado) was employed by the members and postdoctoral fellows at Stanford, OSU Department of General Science on a tempo- Brown, and Notre Dame, at the Universities of rary faculty line from 1975 to 1979. He quickly New Mexico, Missouri, California, Maryland, gained a reputation as one of the finest teachers in Colorado, South Carolina and the Virgin Islands, the department. He also obtained National and to become staff members at the Cabrillo Science Foundation Support for his research on Marine Museum, the National Institutes of Health, the reproductive endocrinology of the rough- and Hopkins Marine Station. The five Zoology skinned newt and began producing an outstand- faculty currently have two master's and 16 doc- ing series of publications that would make this toral students working under their tutelage. organism a new model for studies of endocrine A signal event occurred in 1984 when Robert M. control of reproductive behavior. Since the Storm retired after being associated with the Department of General Science did not have a department as a graduate student and then a tenure-track position available, the Department of faculty member for 45 years. When I joined the Zoology was able to arrange the addition of Frank department in 1977, I frequently heard faculty, to its faculty. Two years later he was awarded a students, and office staff saying "Doc" did this, or National Institutes of Health Research Career "Doc" did that, or ask "Doc," or "Doc" said, etc. In Development Award (RCDA). This award, an environment in which doctorates are more perhaps the most prestigious available to a young common than short speeches, the appellation scientist, provided full salary to Frank for a period "Doc" caused me a certain amount of confusion. I of five years so that he could concentrate his soon learned, however, that there was only one energies on his research program. Although real "Doc" and his last name was Storm. In Frank maintained an active role in the depart- addition to acting as head advisor to all Zoology ment's graduate program during his RCDA, his undergraduates, as faculty advisor to the Zoology NIH grant reimbursed the department for his Club, and as Assistant Chairman of the depart- salary so we were able to employ other young ment, Doc maintained a full instructional load and scientists (Drs. Paul Shirk and Jack Rose) to teach was active in herpetological research and writing Frank's undergraduate courses. up to the time of his retirement. As befits a Carter Frank's RCDA ended in 1986. He then resumed Awardee for Inspirational Teaching, his courses in full instructional duties and his research also herpetology, ornithology, zoogeography, and continued at a vigorous rate. He is currently vertebrate biology were among the most popular investigating the role of small neuropeptides in in the department. Such faculty members are not regulating male sexual responsiveness. These replaceable; one merely searches for another studies have brought five postdoctoral associates person to begin again the process of building a to Frank's laboratory. One of his papers deserves program in the area. special note. AlQng with his wife, Dr. Kathleen The department chose to advertise broadly in Moore of the OSU Philosophy Department, Frank the area of vertebrate ecology. As with the last 19 els on the energetic costs and benefits of territori- ality. He successfully tested these models with Hawaiian damselfish and Sierran hummingbirds. Mark is currently studying interactions between larval recruitment, predation, and reef structure in determining the distribution and abundance of fishes in the Virgin Islands. He has constructed 52 artificial reefs providing various numbers and sizes of shelter holes, and will manipulate the densities of predatory fishes on these reefs. Closer to home, Mark has joined with colleagues in the College of Oceanography to study a deep rocky reef on the edge of the continental shelf off Oregon using manned submersibles. The reef is a nursery habitat for commercially important fish species. This study is providing unprecedented data on interannual variation in communities of deep-reef fishes. A significant new addition to our faculty occurred in 1979 when Arthur J. Boucot (Ph.D., Harvard University) of the OSU Geology Depart- ment was given a joint appointment in Zoology. this relationship became increasingly closer and in 1986 his line was formally transferred to our Robert "Doc" Storm. Photo taken in the late I 970s department. Art is one of OSU's most distin- near Denio, Nevada. guished professors. His bibliography contains over 380 entries of scientific articles, books, and search for an ecologist, a large number of applica- other publications, and his numerous honors and tions was received. Two candidates emerged as awards include not only academic citations but, highly desirable colleagues. After much discus- stemming from his Air Corps service in the 1940s, sion with the upper administration, the depart- the Distinguished Flying Cross, Air Medal, and six ment received permission to hire both individuals. Oak Leaf Clusters. Following discharge from the F. Lynn Carpenter (Ph.D., University of California, military in 1945, Art obtained his doctorate in the Berkeley), an established faculty member at the field of invertebrate paleontology at Harvard and University of California, Irvine, whose research then worked for the U.S. Geological Survey before focused on the ecology of hummingbirds, was assuming faculty positions at the Massachusetts appointed Associate Professor. Unfortunately, Institute of Technology and later the California personal considerations led to her return to Institute of Technology. He came to OSU in 1969 California after two years on our faculty. as Chairman of the Geology Department. In The other person identified in the search for addition to extensive field work collecting fossil Doc's replacement was Mark A. Hixon (Ph.D., marine brachiopods in the United States, Art has University of California, Santa Barbara). Mark's conducted research in both Western and Eastern doctoral research included the first published Europe, the Soviet Union, Antarctica, New Zeal- experimental demonstration of competition and, Australia, Malaya, Brazil, Bolivia, Peru, South between natural populations of fishes. He was Africa, Inner Mongolia, Manchuria, China, and jointly appointed to the faculties of Zoology and Ghana. He has contributed extensively to our Oceanography in 1984 after postoctoral stints at understanding of rates and patterns of evolution the Universities of Hawaii and California. Mark's and extinction. At an age when most people start research specialty is behavioral and community thinking of retirement, Art continues to work ecology of reef fishes and hummingbirds. He also seven days (and evenings) a week and to produce brings expertise to the department in theoretical more than 10 papers (and usually a book or two) ecology and has produced a series of elegant mod- each year. He has had continuous support from 20 the National Science Foundation and has just been area of behavioral biology. This area had the awarded an Accomplishment-Based grant; that is, advantage of building on our current strengths in because of the quality and quantity of his past behavioral ecology, genetics, endocrinology, and research, he was not required to submit a full neurobiology. The field of behavior is one that competitive proposal to NSF for grant renewal. lends itself to study at all levels of biological Art teaches courses in evolution, biogeography, organization from the molecule to the community. and paleontology and interacts extensively with Moreover, it is clear that new and exciting devel- both faculty and graduate students in our ecology opments will continue to take place in this area for and evolution group. many years to come. After completing two terms (8 years) as chair- Under Chris' stewardship, and due largely to man, and feeling frustrated at trying to maintain the efforts of OSU President John Byrne and the department's strengths while resources were College of Science Dean Frederick Home, the de- being withdrawn by the upper administration (see partment has received its first endowed professor- below), I expressed my desire to put aside admin- ship, the Wayne and Gladys Valley Chair in istrative duties and return to full-time teaching Marine Biology. This position has been made and research at the end of the 1985-86 academic possible by a $1,500,000 gift to the department and year. The department held an election to recom- will be occupied by a distinguished faculty mend a replacement to the dean, and Chris Bayne member with internationally recognized excel- became the new department chairman in Septem- lence in some area of research with marine ani- ber, 1986. mals. The search for this individual is currently One of Chris' first acts was to arrange a week- underway. end faculty retreat at the Oregon House in After a three-year term of accomplishment and Yachats. The meeting had the dual purpose of of frustrations over the continuing budgetary attempting to consolidate gains made by the woes of the 1980s (see below), Chris Bayne de- department in recent years and to reach new cided to relinquish the chairman's position in 1989 decisions that would lead to increased national so that he could devote more time and energy to and international visibility of OSU Zoology. After research and teaching. The department now looks extensive debate, the department unanimously forward with eager anticipation to that period of decided to create a focus by concentrating future its future history during which its next chairper- faculty appointments, whenever possible, in the son, Jane Lubchenco, will be at the helm. J

21 The Zoology Club

In response to frequent comments from our paper on their selected topic. This course has been undergraduates that they wanted to become better a great success, but it has been necessary to limit acquainted with each other and the faculty, in 1979 enrollment to 20 students because of financial the department established the Zoology Club. Doc restrictions. Limited funds from the Zoology Storm volunteered to act as faculty advisor and to account of the OSU Foundation are used to rent help in arranging speakers. At club meetings, vans for the trip. Students pay for all other members of the zoology faculty or biologists from expenses, which are held to a minimum by another OSU department or a nearby institution camping and cooking as a group. This trip has present informal talks on their research. Two field been cited by a number of department majors as trips and one field course have been sponsored by crystallizing their interests in zoology and stimu- the club each year. The field trips are to the Friday lating them to continue their education in gradu- Harbor (Washington) Marine Laboratory in the ate school. fall, and to the Malheur Refuge in eastern Oregon The Zoology Club also provides input to the each spring. Over half the department's faculty department for changes in its curriculum and and many of its graduate students have partici- undergraduate policies. From the time of its pated in these field trips, providing ideal settings formation, undergraduate members have been for informal interactions. asked to evaluate faculty classroom performance Since 1981 the Club has sponsored an annual when promotion and tenure decisions are being spring break field course to the deserts of southern made. California and Arizona. Each student generally Since Doc Storm's retirement, departmental focuses on one group of desert animals and keeps liaison with the Club has been jointly provided by a complete field notebook of observations and Austin Pritchard and our colleague from the collections. In the spring quarter the specimens Biology Program, Dr. Joe Beatty. Joe, who ob- are prepared and added to the department's tained his Ph.D. in zoology under Doc's guidance, collections while the students continue theirpro- has been a regular leader on the annual field trips. jects with library research and preparation ofa

22 Faculty Meetings Minutes

Departmental faculty meetings sometimes seem -from the minutes of 16 February 1982: like a "good news -bad news" joke. In the former "Next year the College of Science will be required to category, the following items of cheer and whim- reduce its budget base by 3 percent. One entry-level sey have appeared in the minutes: faculty position and the department research assis- tant will be removed from Zoology. Thus a total of -from the minutes of 27 February 1980: approximately $30,000 is being lost by our depart- "The Zoology Department Christmas party will be ment. The department's reduction will be met by (1) on Friday, February 29, 7:00 p.m. at Greenberry elimination of the General Zoology instructorship, Grange. Dinner will be a potluck and families are (2) elimination of the RA, and (3) removal of $3,200 from our course budget (which totals $22,140 this invited." year)." -from the minutes of 27 May 1981: -from the minutes of 14 April 1982: "Student credit hours (SCH) in the College of Sci- ence increased by 12 percent from 1973/74 to 1980/ "Dr. King stated that the main business today is to discuss the budget picture and review the determi- 81 (183,000205,000).In that same period, SCH generated by Zoology faculty increased by 29.5 nation of who gets cut and by how much. In the most recent round of budget cuts, the College of Science percent (11,24314,564)." has had an additional reduction of $85,000. The $240,000 in base-budget reductions will not be re- -from the minutes of 13 January 1982: turned to the college budget in the future. "Beaver Open House will be held on January 30, "In addition, Pharmacy will be sending approxi- 1982. Dr. Storm asked for a volunteer to represent mately students zoology human the department. Dr. Brownell asked, "What is Bea- labs starting next year.It will cost an additional ver Open House?" The assembled faculty immedi- $1,500- $2,000 to supply these students with cats for ately suggested that he be presented with an oppor- dissection. We do not have these funds and so far tunity to acquire this knowledge." have been unable to obtain them from either the Dean of the College of Science or the Vice President The "bad news" side is also amply represented, for Administration. In general discussion it was Indeed, no history of the department in the 1980's suggested that students might be asked to supply would be complete without a record of its serious their own cats; perhaps an arrangement could be and continuing budgetary problems. These made for purchase of cats from the bookstore. The problems are not unique to our department; they department's accountant was asked to check on this originate with the Oregon economy and are possibility, and after several minutes of 'catty' ban- passed to us through the State Board of Higher ter, she was informed by the bookstore that this was Education. They have resulted in faculty losses, 'not a desirable line of inventory'." played a role in the resignation of two department -from the minutes of 13 April 1983: chairmen, and frustrated both faculty and stu- dents alike. Perhaps the best way to report these "Dr. King reported on a recent meeting with Dean problems is to reprint portions of faculty meeting Sugihara. It is highly likely that there will he a cut in minutes. next year's budget. The Teaching Assistant and Supplies budgets are in particular jeopardy. The -from the minutes of 22 September 1981: hall telephones used for emergencies and by our graduate students have been removed to save money. "The $300,000 budget cut to the College of Science Xerox charges have also been increased." will affect the Department of Zoology in some way, as yet unknown. Departmental administrative costs -from the minutes of 7 March 1984: have already been cut. Further budgetary decreases will involve reduction in essential departmental "The Chancellor has reduced the OSU budget by activities." $885,000; $110,000 of this will be passed on to the College of Science.Zoology will lose one more faculty position (that of Dr. Owczarzak who re-

23 cently retired) and 6.5 terms of Graduate Teaching standing the fact that its high productivity is no Assistant service." insurance against loss of resources. However, fiscal insecurities are obviously debilitating when -from the minutes of 11 February 1987: they result in loss of badly needed faculty and "Dr. Bayne distributed a memorandum from the graduate student positions. Equally debilitating Biological Sciences Librarian for Kerr Library which are the frustrations of having to do less than our detailed the need for book and journal requests. best as teachers and as scholars because of out- Approximately $10,000 has been allocated for zool- dated equipment and absence of supplies. Valued ogybooksand faculty and graduate students should colleagues have left OSU because of this situation forward their requests." and, unless this trend is reversed, more will leave in the future. - -from the minutes of 24 February and 4 May Part I of the Zoology history ended on a high 1988: note with the anticipation of moving into a new "The Kerr Library anticipates cancellation of journal building having adequate space and modern subscriptions to save funds. A saving of over $5,000 resources. Part II ends on a different note. As the will be realized by cancellation of zoology journal 1988-1989 academic year draws to a close, new subscriptions." budget cuts severely affecting the department have been announced. Moreover the fiscal -from the minutes of 4 May 1988: outlook for the next biennium remains clouded. At the same time the Department of Zoology has "The College of Science has required our depart- matured into a nationally recognized group of ment to cut next year's budget by 1 percent. Depart- faculty, alumni, and students who are bound to- mental response to this permanent cut will be to reduce the Lab Tech II position to 70 percent time gether by a common perception that the privileges and to reduce funds in the graduate teaching assis- of association we have been accorded must be tant budget." preserved for those to come in the days and years ahead. This would therefore seem to be an appro- As the above notes illustrate, the decade of the priate point at which to wait for the future to '80s has been a difficult one for the State of Ore- become the past so that this chronicle can be gon. The Department of Zoology has accepted continued. U with a commendable measure of grace and under-

24 Appendix I Department of Zoology Faculty 1889-1989

Ailman, Delmar I. Conte, Frank P. B.S. (Michigan State Normal) 1928; M.S. A.B. (UC Berkeley) 1951; Ph.D. (UC Berkeley) (Michigan) 1931; Ph.D.(Michigan) 1936 1961 Asst. Prof. 1960-65, Assoc Prof. 1965- Assoc. Prof. 1939-48, Prof. 1949-65, Emeritus 71, Prof. 1971-present 1965-1983 Cord ley, Arthur B. Alvarado, Ronald H. B.S. (Michigan State) 1888; MS. (Cornell) 1900 B.A. (California) 1956; M.S. (Washington State) Prof. and Head of Department 1895-1914 1959; Ph.D. (Washington State) 1962 Asst. Prof. 1962-66, Assoc. Prof. 1966-74 Courtney, Wilbur D. B.S. (Washington State) 1922; M.S. (Washing- Bayne, Christopher J. ton) 1929 B.S. (Wales) 1963; Ph.D. (Wales) 1966 Instr. 1922-29, Asst. Prof. 1929-30 Res. Assoc. (Michigan) 1968-71, Asst. Prof. 1971-76, Assoc. Prof. 1976-82, Prof. 1982- Dawson, Peter S. present, Chairman of Department, 1986-89 B.S. (Washington State, Pullman) 1960; Ph.D. (UC Berkeley) 1964 Assoc. Prof. 1969-75, Prof. Benedict, Edith L. 1975-present; Chairman, Genetics Program B.S. (Northwestern) 1928; M.A. (Northwest- 1977-84 ern) 1929 Instr. 1929-38 deLaubenfels, Max W. B.A. (Oberlin) 1916; M.A. (Stanford) 1926; Blake, Irving H. Ph.D. (Stanford) 1929 B.A. (Bates) 1911; M.A. (Brown) 1912 Prof. 1950-58 Instr. 1913-16 Domfeld, Ernst J. Blaustein, Andrew R. B.S. (Marquette) 1933; M.A. (Wisconsin) B.A. (UC Santa Barbara) 1971; M.S. (Nevada) Ph.D. (Wisconsin) 1937 1973; Ph.D. (UCSB) 1978 Instr. 1938-42, Asst. Prof. 1942-45, Assoc. Prof. Instr. 1978-79, Asst. Prof. 1979-84, Assoc. Prof 1945-50, 1984-1989, Prof. 1989-present Prof. 1950-76, Chairman of Dept. 1952-76. Emeritus Prof. 1976-83 Boucot, Arthur J. A.B. (Harvard) 1948; Ph.D. (Harvard) 1953 Edwards, Alice L. Prof. of Geology 1969-present; Prof. of Zool- B.S. (Oregon State) 1906 ogy 1979-present Instr. 1910-16

Bridwell, John C. Fasten, Nathan B.S. (Baker) 1901 B.S. (C.C.N.Y.) 1910; Ph.D. (Wisconsin) 1914 Instr. 1907-10 Assoc. Prof. 1920-21, Prof. and Head of Dept. 1921-44 BrowneD, Philip H. B.A. (UC Berkeley) 1970; Ph.D. (UC Riverside) Gimlich, Robert L. 1976 B.S. (UC Santa Barbara) 1980; Ph.D. (UC Asst. Prof. 1979-85, Assoc. Prof. 1985-present Berkeley) 1985 Asst. Prof. 1988-present Carpenter, F. Lynn B.A. (UC Riverside) 1966; Ph.D. (UC Berkeley) Gordon, Kenneth L. 1972 B.A. (Colorado College) 1923; M.A. (Missouri) Associate Prof. 1984-87 1925; Ph.D. (Cornell) 1936 lnstr. 1927-29, Asst. Prof. 1929-40, Assoc. Prof. Chandler, Asa C. 1940-45, Prof. 1945-52, Chairman of Dept. B.A. (Cornell) 1911; M.S. (California) 1912; 1945-52, Emeritus Prof. 1969-83 Ph.D. (California) 1914 Instr. 1914-1 7,Asst. Prof. 1917-19

25 Green, Bess R. Krueger, Hugo M. B.A. (Illinois) 1907; M.A. (Illinois) 1910 B.A. (Denver) 1924; M.A. (Denver) 1926; Ph.D. Asst. Prof. 1926-29 (Michigan) 1930 Prof. 1948-62, Emeritus 1962-present Griffin, Fred L. B.S. (Oregon State) 1908 Lubchenco, Jane Instr. 1908-10 B.A. (Colorado Coll.) 1969; M.S. (Washington) 1971; Ph.D. (Harvard) 1975 Grobstein, Clifford Asst. Prof. 1977-82, Assoc. Prof. 1982-88, Prof. B.S. (C.C.N.Y.) 1936; MA. (UCLA) 1938; Ph.D. 1988-present, Chairperson of Department (UCLA) 1940 1989- Instr. 1940-43 McElfresh, Fred M. Hague, Florence S. B.S.; Instr. 1899-1901 B.A. (Kansas) 1911; M.A. (Kansas) 1914; Ph.D. (Illinois) 1921 Menge, Bruce A. Instr. 1921-24, Asst. Prof. 1924-26 B.A. (Minnesota) 1965; Ph.D. (Washington) 1970 Hard, Robert P. Asst. Prof. 1976-79, Assoc. Prof. 1979-85; Prof. B.S. (Washington) 1967; M.S. (Washington) 1985-present 1970; Ph.D. (St. Univ. of NY) 1975 Asst. Prof. 1978-85 Miller, Richard L. B.S. (Chicago) 1962; Ph.D. (Chicago) 1965 Harris, Patricia J. Asst. Prof. 1966-67 B.A. (UC Berkeley) 1954; MS. (Yale) 1958; Ph.D. (UC Berkeley) 1962 Mohler, James D. Assoc. Prof. 1964-73 BA. (Missouri) 1949; M.A. (Missouri) 1950; Ph.D. (California) 1955 Hillemann, Howard H. Asst. Prof. 1955-60, Assoc. Prof. 1960-66 B.S. (Marquette U., Milwaukee) 1933; M.A. (Wisconsin) 1939; Ph.D. Moore, Arthur R. (Wisconsin) 1942 B.A. (Nebraska) 1904; Ph.D. (California) 1911 Asst. Prof. 1946-52, Assoc. Prof. 1952-56, Prof Prof. 1932-33 1956-75 Emeritus 1975-85 Moore, Frank L. Hisaw, Frederick L., Jr. BA. (College of Wooster) 1967; M.A. (Colo- B.S. (Missouri) 1950, M.S. (Missouri) 1952; rado) 1970; Ph.D. (Colorado) 1974 Ph.D. (Harvard) 1955 Asst. Prof. 1979-82, Assoc. Prof. 1982-87, Prof. Asst. Prof. 1958-61, Assoc. Prof. 1961-1989 1987-present

Hixon, Mark A. Morris, John A. B.S. (UC Santa Barbara) 1973; M.A. (UCSB) B.A. (Stanford) 1958; MS. (Hawaii) 1960; 1974; Ph.D. (UCSB) 1979 Ph.D. (UCLA) 1966 Asst. Prof. 1984-1989, Assoc. Prof. 1989- Asst. Prof. 1968-1974, Assoc. Prof. 1974-82, present Prof. 1982-present Hurd, Charlotte N. Osborn, John L. B.S. (UC Berkeley); M.S. (UC Berkeley) Ph.C. (Michigan) 1915; B.A. (Kansas) 1922; Instr. 1916-19 M.A. (Nebraska) 1923; Ph.D. (Washington) 1939 King, Charles E. Instr. 1923-34, Asst. Prof. 1934-46, Assoc. Prof. A.B. (Emory) 1958; M.S. (Florida State) 1960; 1946-47, Emeritus 1947-? Ph.D. (Washington) 1965 Prof. 1977-present, Chairman of Department, Owczarzak, Alfred 1977=86 B.S. (Cornell) 1944; Ph.D. (Wisconsin) 1953 Instr. 1955-57, Asst. Prof. 1957-61, Assoc. Prof. 1961-83, Emeritus 1983-present. 26 Pratt, Ivan Swan, Grant A. B.A. (College of Emporia, Kansas) 1932; M.S. B.S. (Oregon State) 1922 (Kansas State) 1935; Ph.D. (Wisconsin) 1938 Asst. Prof. 1939-43 Asst. Prof. 1946-47, Assoc. Prof. 1947-52, Prof. 1952-73 Sykes, George F. Ph.D. (Brown) 1909; M.A. (Brown) 1910 Pritchard, Austin W. Instr. 1910-1 3, Asst. Prof. 1913-14, Prof. and B.A. (Stanford) 1948; M.A. (Stanford) 1949; Head of Dept. 1914-21 Ph.D. (Hawaii) 1953 Instr. 1953-55, Asst. Prof. 1955-59, Assoc. Prof. 1959-65, Prof. 1965- Taylor, Alfred present B.A. (Oregon) 1932; M.A. (Oregon State) 1934; Ph.D. (Oregon State) 1935 Reaka, Marjorie Instr. 1935-40 B.A. (Kansas) 1965; M.A. (Kansas) 1967; Ph.D. (UC Berkeley) 1974 Taylor, Aravilla M. Asst. Prof. 1975 B.S. (Chicago) 1915; M.S. (Chicago) 1916; Ph.D. (Chicago) 1919 Roberts, Paul A. Asst. Prof. 1920-21 B.S. (Illinois) 1953; M.D. (Illinois) 1957; M.Ed. (Chicago T.C.) 1962; Ph.D. (Chicago) 1962 Washburn, Frederick L. Assoc. Prof. 1966-73, Prof. 1973-present B.A. (Harvard) 1882 Professor and Head of Dept. 1889-95 Ruben, John A. B.S. (Humboldt) 1968; M.A. (UC Berkeley) Wiens, John A. 1970; Ph.D. (UC Berkeley) 1975 B.S. (Oklahoma) 1961; M.S. (Wisconsin) 1963; Asst. Prof. 1975-83, Assoc. Prof. 1984-present Ph.D. (Wisconsin) 1966 Asst. Prof. 1966-69, Assoc. Prof. 1969-75, Prof. Shaw, William T. 1975-78, Chairman of Department 1976-78 B.Ag. (Minnesota) 1898; B.S. (Minnesota) 1899; M.S. (Michigan State) 1900 Wight, Howard M. Instr. 1901-07 B.S. (Bates) 1915; M.S. (Oregon State) 1916 Instr. 1916-20, Asst. Prof. 1920-27 Storm, Robert M. B.Ed. (N. Illinois) 1939; M.S. (Oregon State) Wulzen, Rosalind 1941; Ph.D. (Oregon State) 1948 B.S. (UC Berkeley) 1904; M.S. (UC Berkeley) Instr. 1948-50, Asst. Prof. 1950-55, Assoc. Prof 1910; Ph.D. (UC Berkeley) 1914; Sc.D. (Ore- 1955-62, Prof. 1962-84, Emeritus 1984-present gon) 1943 Asst. Prof. 1933-41, Assoc. Prof. 1941-46, Prof. 1946-47, Emeritus 1947-80

Anecdote

Preparing this history brought me into contact 1 Dec 88--I asked Chairman Chris Bayne to with OSU's little-known Office of Archives, which obtain information on retirement dates of several serves as a repository for sundry arcane materials. Zoology faculty. This office is the major official source of historical 2 Dec 88--Chris addressed a letter to Archives information since University policy requires requesting the information. departments to forward personnel records to 7 Dec 88--Archives asked the College of Archives when faculty and students depart or Science to provide them with the retirement dates. retire from the University. We thank Archives for 9 Dec 88--The College contacted Chris Bayne considerable help in assembling information. to obtain the information for Archives. Sometimes research progressed less well than we 9 Dec 88--Chris, knowing I was working on wished, as illustrated by the following sequence of the departmental history, asked if I could help the events: College of Science gather some information on retired Zoology faculty. 1 27 Appendix II Associated Faculty 1889-1989

Alspach, George S. Fuller, Eugene C. B.A. (Antioch); M.S. (Oregon State) 1967; Ph.D. B.S. (Nevada) 1960; M.S. (Nevada) 1962 (Oregon State) 1972 Instr. 1966-72 Instr. 1966-67 Gonor, Jefferson J. Anderson, Carl L. B.S. (Southwestern Louisianna) 1953; Ph.D. B.S. (Michigan) 1928; M.S. (Michigan) 1932; (Washington) 1964 Dr. P.H. (Michigan) 1934 Asst. Prof. 1965-79 Prof. 1949-59 Janis, Christine Anderson, Glenn A B.A. (Cambridge) 1973; M.A. (Harvard) 1974; B.S. (Washington State) 1953; M.A. (Washing Ph.D. (Harvard) 1978. - ton State) 1958; Ph.D. (Oregon State) 1964 Instr. 1977-78 Instr. 1963-64 Johnson, La Von C. Baer, Dee B.A. (Brigham Young) 1957; M.A. (Utah) 1959; B.S. (Illinois) 1953; Ph.D. (UC Berkeley) 1962 Ph.D. (Utah) 1967. Prof. 1984-present Asst. Prof. 1965-69

Beatty,J.J. Kerley, David E. B.S. (Missouri) 1970; M.S. (Missouri) 1973; B.S. (Oregon State) 1958; M.S. (Oregon State) Ph.D. (Oregon State) 1979 1962 Instructor (Zoology and Biology Program) Instr. 1962-63 1979-present Kuster, Kimber C. Belton, John C. B.S. (Michigan) 1925; M.S. (Michigan) 1926 B.S. (Lewis & Clark) 1957; M.S. (Oregon State) Instr. 1926-27 1962 Instr. 1964-65 Loker, Eric Sam B.S. (Cornell) 1972; M.S. (Michigan) 1974; Buckley, Patricia Ph.D. (Iowa State) 1979 B.S. (Ohio University) 1948; M.S. (Oregon Instr./Res. Assoc. 1979-83 State) 1958; Ph.D. (Oregon State) 1964 Inst/ Asst.Prof. /Res. Assoc. 1978-82 Lynch, James E. BA. (Nebraska) 1917; M.A. (Nebraska) 1921 Chilgren, John D. Instr. 1921-22 B.S., (Gonzaga) 1965; MS (Washington State) 1968; Ph.D. (Washington State) 1974. Mayshark, Cyrus Asst. Professor 1974-78 B.A. (Williams) 1949; M.Ed. (Boston) 1952; MS. (Harvard) 1952; H.S.D. (Indiana) 1954 Cole, Elizabeth R. Asst. Prof. 1959-62, Assoc. Prof. 1962-65 B.A. (Brown) lnstr. 1915-16 Mullen, Terry L. B.A. (Central Washington State College) 1969; Darrow, Thomas D. MS. (C.W.S.C.) 1971; B.A. (Nebraska State) 1960; M.A. (South Ph.D. (Oregon State) 1975 Dakota) 1961 Instr. 1978-1979 Instr. 1965-66 Muske, Linda Ellis, John K. B.A. (Tufts) 1968; Ph.D. (Oregon) 1983 B.Ed. (S. Illinois) 1943; B.S. (S. Illinois) 1944; Asst. Prof. 1989 M.P.H. (Michigan) 1948; Ph.D. (Michigan) 1963 Newman, Lester J. Assoc. Prof. 1965-74, Prof. 1975-85, Emeritus A.B. (Washington University, St. Louis) 1955; 1985-present M.A. (Michigan) 1960; Ph. D. (Washington University, St.

28 Louis) 1963 Romerdahl, Cynthia A. Asst. Prof. 1963-64 B.S. (Washington State) 1978; Ph.D. (UC Berkeley) 1984 Newstead, James D. Asst. Prof. 1988-present B.A. (British Columbia) 1954; M.A. (British Columbia) 1956; Ph.D. (Oregon State) 1962 Rose, Jack Instr. 1960-62, Asst. Prof. 1962-63 B.S. (Ohio State) 1975; M.S. (Ohio State) 1979; Ph.D. (Oregon State) 1985 O'Hara, Richard Asst. Prof. 1985-86 B.S. (Michigan State) 1972; M.S. (Michigan State) 1974; Ph.D. (Oregon State) 1981 Shirk, Paul D. Asst. Prof./Res. Assoc. 1981-present B.A. (Northern Iowa) 1970; M.S. (Texas A&M) 1975; Ph.D. (Texas A&M) 1978 Owen, William B. Asst. Prof. 1981-84 B.A. (Kentucky) 1927; M.A. (Minnesota) 1929 Instr. 1930-31 Taylor, Alfred B.A. (Oregon) 1932; M.A. (Oregon State) 1934; Ph.D. (Oregon State) 1935 Paul, Dorothy Instr. 1935-40 B.A. (Radcliffe College) 1963; D.E.S. (Faculte des Sciences; Marseille, France) 1965; Ph.D. Turner, Teresa (Stanford) 1970 B.S. (Michigan State) 1974; M.A. (California Asst. Prof. 1976-77 State, Hayward) 1978; Ph.D. (Oregon State) 1982 Polls, Gary Instr. /Res. Asst. 1984-86 B.S. (Loyolla, Los Angeles) 1969; M.A. (UC 1976; Ph.D. Riverside) 1977 Yamada, Sylvia Behrens Asst. Prof. 1977-79 B.Sc. (British Columbia) 1968; M.Sc. (British Columbia) 1971; Ph.D. (Oregon) 1974 Potter, Sandy W. Asst. Prof.! Res. Assoc. 1984-present B.A. (Minnesota) 1960; M.S. (Minnesota) 1967; Ph.D. (Arizona) 1970 Asst. Prof. 1983-85; Asst. Prof. Research 1986- present

/

29 Appendix III GraduateDegrees in Zoology

Year M.A. M.S. Ph.D.

1923 Walter P. Covell 1926 Nettie L. Murray 1934 Alfred Taylor 1935 Alfred Taylor 1938 Raymond W. Coopey 1939 William Graf Maurice S. Tarshis 1941 Daniel W. Slater Robert M. Storm 1942 Sr. Gemma Piennett 1943 Gilbert M. Shearer William Graf 1944 Hildegard Lamfrom Robert L. Livezey 1947 Virginia L. Weimar 1948 Bertha D. Cutress Robert M. Storm Walter S. Vincent Jr. 1949 Philip C. Dumas Myrtle D. Beatty Fred G. Evenden Jr. Donald J. Reish Charles E. Cutress Robert M. Yancey Robert W. Merriam Ernst W. Retzlaff Rudolph E. Rieschel Aryani. Roest Robert C. von Borstel Kenneth M. Walker 1950 James H. Berrian Herbert G. Adams Harold V. Gallaher Claude C. Alexander Robert D. Bratz Richard A. Pimentel Richard M. Ritland Robert R. Sowell Roland D. Walters 1951 Donald G. Dunlap Frank W. Adams Elwin E. Bennington Earl Gregoire Delpha D. Dunlap Marvin F. Hill John A. McGowan Donald M. Purdy Donald W. Twohy William C. Van Arsdel III

1952 William C. Burns Denzel E. Ferguson Robert D. Bratz Darrell L. Davis John C. Giesler Albert C. Hawbecker Lloyd R. Yonce Donald V. Hemphill Richard A. Pimentel 1953 Lydia Beyerlein Wesley J. Birge Philip C. Dumas Richard H. Eddy Jr. 1954 Vernon E. Thatcher Lewis E. Aldrich Jr. Richard E. Freiburg David R. McIntyre Donald L. Lehmann Robert S. Tether James E. McCauley Aryan I. Roest

1955 Franklin W. Sturges Robert A. Knight Wesley J. Birge

30 Year M.A. M.S. Ph.D.

Forrest D. Tibbitts Vernon E. Duiker Simone T.-H. Truong Marvin F. Hill Andrew C. Olson Jr. Kenneth M. Walker

1956 Morris E. Weaver Nathan W. Cohen Charles G. Hansen Donald G. Humphrey 1957 Hilda L. Ching Denzel E. Ferguson Galen E. Clothier Franklin W. Sturges Stanley N. Wilkes

1958 Hugh P. Stanley Jack Foster Forrest D. Tibbitts

1959 John W. Goertz William C. Van Arsdel III Oliver W. Johnson Morris E. Weaver Robert E. Moore Kenneth R. Porter 1960 Glenn R. Stewart Robert L. Puyear Lewis E. Aldrich Gertrude L. S. Swedberg Asa C. Thoresen Norman F. Weatherly Elisha S. Tikasingh

1961 Eugene D. Bawdon Albert G. Canaris Alan C. Heath Galen E. Clothier Norman C. Leeling Clifford V. Davis James E. Monroe Hugh P. Stanley Ronald J. White John H. Wirtz

EllisI.Wyatt

1962 William M. Beck James M. Ford John C. Belton James D. M. Newstead Grace A. Hamilton Harold Watling David E. Kerley Lewis M. Turner Jr.

1963 Elver H. Voth George F. Crandell Audrey L. Braswell Jr. Howard C. Morse Charles G. Danforth John C. Neeley

1964 Gordon W. Martin Jean E. Kneeland Glenn A. Anderson Sheridan V. Merritt Joan D. Beltz Clarence A. Porter John D. DeMartini Gilbert A. Rinard Alan G. Heath Ronald L. Ritschard Glenn R. Stewart Steven R. Thompson Rosalind W.-P. Yuen

1965 Larry T. Spencer Francis P. Belcik Frank W. Adams John A. Dorsch Hugh C. Black FrankJ. Longo Oliver W. Johnson 31 Year M.A. M.S. Ph.D.

1965 Ralph R. Moldenhauer Thomas G. Meade Melvin L. Schamberger Robert L. Puyear Loren E. Van Tassel

1966 Elaine C.-L. Chan Glen W. Clothier John C. Belton James W. York Robert R. Hollenbeck Carl A. Forss Tod S. Johnson Cornelis Laban Christopher 0. Maser Gordon W. Martin James M. Shumake Clarence A. Porter - James G. Wernz Ronald L. Ritschard Herman E. Wyandt Stanley N. Wilkes Patrick T.-C. Wong (Genetics)

1967 Hans W. Behrisch George S. Aispach Thomas D. Darrow James R. Barnes Eugene G. Fuller Edmund D. Brodie Jr. Frank J. Longo Richard C. Gethmann Howard C. Morse John D. Haertel John M. Smith Carol M. Inhelder Steven R. Thompson Joseph S. Lidrich Russell A. Whitehead Edwards R. Long Robert H. Parsons Kasparek (Thompson) Beatrice M. Wyse

1968 Stanley H. Anderson Carol D. Baker James C. Hanson Jeffrey L. Brigg Samuel Lin Peter A Nyberg Richard T. Carter Raul J. V. Moreno Elver H. Voth Louise G. Chatlynne David J. Wall Ronald J. White James R. Hall

1969 Flash Gibson Daniel 1.-H. Law Ronald G. Altig Linda S. Gibson Anne Moody Edmund D. Brodie Michael C. Hartman William E. Noonan Thomas H. Dietz Thomas W. Moon William E. Sype John J. Dropp Barbara W. Saigo Thomas G. Wicklund Ralph R. Moldenhauer Randall W. Smith Eric Panitz

1970 William L. Bliss Dale L. Bures Stanley H.Anderson James F. Traynor Michael A. Duvall John D. Haertel Dennis S. Greenley David E. Kerley Alfred W. Hanson Stephen J. Kleinschuster Shirley A. Hull Joseph S. Lidrich Michael A. Lee Robert I-I. Parsons

Michael G. 0'Rand James 0. Roberts Irwin Polls Melvin L. Schamberger Daniel F. Stiffler Patrick 1.-C. Wong Kent L. Thornburg Richard J. Ulbricht 32 Year M.A. M.S. Ph.D.

1971 Robert E. Batie Jeffrey L. Briggs Norman W. Castillo Glen W. Clothier Corynn D. Converse Flash Gibson Judith A. Elkins Sheldon R. Johnson James D. Hansen Stephen G. Martin Dan L. Hawkins John C. Neeley Robert P. Heckey Lester E. Walker Margaret L Macavoy Antoinette M. Poole Robert F. Ramig Patricia A. Roberts Elizabeth A. Roseberry Dennis L. Wilson

1972 Karen I. Miller Gary L. Gill George S. Alspach Jr. Spencer H. Hamada James R. Barnes Pamela H. Linn Randall E. Brown Joan C. Moran Louise G. Chatlynne John W. Pike Bill P. Lovejoy Jeannine H. Riazance Albert Mozejko Ronald A. Nussbaum Paul J. A. Presidente Kent L. Thornburg Alan B. Thum David R. Voth Robert A. Wasmer Herman E. Wyandt Jr. Ellis J. Wyatt

1973 Patricia J. Elvin Eugene M. Burreson Viroj Boonkoom Paul C. Droukas Richard T. Carter Anne L. Edwards Alfred W. Hanson Theresa L. Felten Michael C. Hartman Gregory R. Garman Shirley A. Hull Frederick E. Jarvis Donald E. Kroodsma Alice M. Lindahl William E. Noonan Robert L. Rodgers James M. Scott Daniel F. Stiffler

1974 Seth R. Hootman Robert E. Ba tie Dwight J. Kimberly Norman. D. Morrison John T. Rotenberry Terry L. Muller Ellen K. Shurtleff Michael F. Salamone Cynthia K. Tait James B. States Daniel C.-N. Tsui Francis T. Takahashi Jr. Gregory J. Wastek Richard J. Ulbricht

1975 David L. Alford Eugene M. Burreson / Jane R. Benson Robert C. Clover David W. Canoy Spencer H. Hamada 33 Year M.A. M.S. Ph.D

1975 Lowell V. Diller Terry L. Mullen Christine M. Hall (Gniewosz)Gary L. Peterson David E. Hill Peter C. Rothlisberg Bert Marsh William E. Sype Mary H. Norton Lori S. Rynd James M. Stone

1976 Lavon L. Bartel Weldon S. Bosworth Jr. Richard D. Bayer Spencer H. Hamada Kam N. Chow Daniel T.-H. Law Charles J. Costa Janet Pennington Keith C. Danielson Bretton W. Kent DiAnne T. Motubu Carolyn J. Mullikin

1977 Gail M. Breed David J. Broderick Sai-Chung Chan Richard L. Miller Ronald L. Haines Clara V. Riddle Susan E. Htfie1d Russel A. Riddle Wayne A. Hoffman Donald K. Seavy Roy B. Mason H James T. Taylor Robert D. Pietruszka Beatrice Van Home

1978 David M. Chapin Theresa L. Felton Dale F. DeWan Daniel B. Matlock Robert S. DeWoskin John T. Rotenberry Dennis W. Heinemann Paul S. Rutledge Beth M. Musser Robert L. White (Carl) Eric Orr Robert C. Roelke Darrel E. Ward Randy Webb

1979 Eric K. Birks Bruce R. Bartlett Charles E. Hart Joseph J. Beatty James S. Heppler Bruce L. Boese Jerry N. Moore Charles L. Davis George R. Schmidt Laurie A. MacPhail Douglas C. Ure Paul B. Samollow Daniel C. N. Tsui

1980 Hailu Ebba Charles L. Davis Steven H. Eddy Steven Craig Hand David B. Herbst Hugh C. Hanlin Lori S. Rynd

1981 John F. Carpenter Karen I. Miller 34 Year M.A. M.S. Ph.D.

1981 David F. Zirkie Richard K. O'Hara William R. Rice

1982 Melinda Brown Monica Geber Audrey A. Millemann Diana K. Padilla Saroj Pandey Emile J. Schoffelen Joan M. Wittier Mary A. Yui Kenneth R. Yates

1983 Brett A. Adams Steven D. Gaines Carla M. D'Antonio Robert J. Lowy Kathy A. Lumas Teresa Turner Jill Meisenhelder

1984 Sunny K. Boyd Christopher P. Marsh Edith F. Moore Robert T. Zoeller Kathryn B. Staley Alice E. Weaver

1985 Carolyn S. Fairfax Sandra P. Spielvogel Shelyl M. Furgal Diana K. Hews Annette M. Olson Laura Pierce Yuqi Zhao

1986 Alice F. Brown Carl A. Boswell Patricia S. Estes Arlene T. Doyle David B. Herbst Lani West

1987 Nancy C. Treneman-Groth

1988 Brenda L. Wiens Sunny K. Boyd Terence M. Farrell

1989 Claire A. Fuller Craig M. Flory Deanna H. Olson Catherine R. Propper

MAIS DEGREES

1977 Nancy L. Eldred Zoology, Psychology, General Science 1978 Sandra L. Rambo Anthropology, History, Zoology

35 Appendix IV Zoology Chairpersons

Pre-Department Formation

1868-1882 Rev. Joseph Emery, A.M. Professor of Mathematics

1882-1889 Edgar Grim, B.S. Professor of Agriculture and Chemistry W. N. Hull, A.M. Professor of Physiology and Drawing

Post-Department Formation

1889-1895 F. L. Washburn 1895-1914 Arthur B. Cordley 1914-1921 George F. Sykes 1921-1944 Nathan Fasten 1945-1952 Kenneth L. Gordon 1952-1976 Ernst J. Dornfeld 1977-1986 Charles E. King 1986-1989 Christopher J. Bayne 1989- Jane Lubchenco

36 Department of Zoology Oregon State University Cordley Hall 3029 Corvallis, OR 97331-2914 Phone (503) 737-3705

Desktop publishing (Aldus PageMaker) and layout by Eva M. Millemann College of Science Oregon State University