Studies and Reports Taxonomical Series 9 (2): 261-366, 2013

A checklist of the subfamily Anthiinae Bonelli, 1813 of the World (Coleoptera: Carabidae)

Martin Häckel1) & Jan Farkač 2)

1) Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, CZ‑165 21 Prague 6, Czech Republic e-mail: [email protected] 2) Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, CZ‑165 21 Prague 6, Czech Republic e-mail: [email protected]

Checklist, Coleoptera, Carabidae, Anthiinae, Catalogue with world distribution

Abstract. In the present checklist of the subfamily Anthiinae (Coleoptera: Carabidae) we summarize taxonomic data including synonyms, type localities and geographic distributions of all listed species described up to July 2013. The subfamily (or according to some authors “group of tribes“) includes in this work the following tribes: Anthiini Bonelli, 1813 (with two subtribes: Anthiina Bonelli, 1813 and Cypholobina Strohmeyer, 1928), Helluonini Hope, 1838 (with two subtribes: Helluonina Hope, 1838 and Omphrina Jedlička, 1941) and Physocrotaphini Chaudoir, 1863, together comprising 41 genera and 350 species. A new synonym is proposed: Termophilum marginatum (Latreille, 1823) syn. nov. of Termophilum sexmaculatum (Fabricius, 1787).

INTRODUCTION

The family Carabidae (Coleoptera: Adephaga) contains more than 35 000 species and is one of the most successful groups of beetles found in all geographic regions and environments with the exception of polar areas. After favorable responses to two catalogues (Broscini Hope, 1838 and Panagaeinae Hope, 1838), we decided to continue publishing such catalogues and chose next to compile another subfamily, the Anthiinae Bonelli, 1813. The species of this subfamily inhabit mainly tropical regions (Afrotropical, Neotropical, Oriental and tropical part of Australia), but distributions of some representatives of the tribe Anthiini Bonelli, 1813 and Helluonini Hope, 1838 (namely the genera Anthia, Colfax, Helluomorphoides and Macrocheilus) often extend into the Holarctic Region, with the highest diversity outside of the tropics recorded in the southern parts of Palearctic and Nearctic subregions. Representatives of the subtribe Helluonina Hope, 1838 inhabit the entire continent of Australia. Some species of Anthiinae are alate (e.g. Physocrotaphini, some genera of Helluonini: Gigadema Thomson, 1859, Macrocheilus Hope, 1838, etc); some are apterous (e.g. most species of Anthiini Bonelli, 1813; some genera of Helluonini: Helluapterus Sloane 1914 or Helluarchus SLoane, 1914); most are nocturnal, and as most carabids are predaceous. Many species are feeding on termites (Anthia Weber, 1801, Netrodera Chaudoir, 1850, Termophilum Basilewsky, 1950). The most speciose genus is Macrocheilus Hope, 1838 represented in tropical regions of Africa and Asia (the catalogue lists 58 species). Currently the subfamily as a whole contains 350 species in 41 genera and three tribes (including 4 subtribes). 261 The system of catalogization is similar to the previous catalogue. Valid taxa are in boldface *We suspect that Termophilum desertorum (Thomson, 1859b) is conspecific with T. babaulti italics and names of authors are not abbreviated. Invalid names (synonyms, junior homonyms, (Bénard, 1921a). If so, Thomson’s name would have priority. In any case, the T. biguttatum etc.) are in standard italics. Names of generic and subgeneric taxa are in capitalized boldface species group (Häckel sn. Obst 1901b) needs to be revised. italics, those of genera extended (e.g. CYCLOLOBA Chaudoir, 1850) and those of subgenera unextended (e.g. CYCLOLOBA s. str.). Names of species are ordered in species groups AF ‑ Africa (if any have been proposed) with names of authors in parentheses or alphabetized (if no groups ANG ‑ Angola have been proposed). Following a species described in a genus different from the currently Ben ‑ Benin accepted combination, the generic name originally used is given in standard italics, and in case BFO ‑ Burkina Faso of a valid taxon the name of the describer is given in parentheses - e.g. villosa (Thunberg, 1806) BOW ‑ Botswana Anthia (parentheses are not used with invalid names - e.g. decemsulcata Bonelli, 1813 Anthia). BUr ‑ Burundi Geographic data are ordered from left to right as follows: continent or its part in boldface, CAM ‑ Cameroun / Cameroon usually further specified in north-south or east-west direction in standard type (e.g. with: s AF COn ‑ Congo (Brazzaville) for countries of southern Africa), and then name of the country. Where the name of the country DOc ‑ Democratic Congo (Kinshasa) accompanying the type locality agrees with the currently recognized English name, it is given eri ‑ Eritrea as a three-letter abbreviation (see the list of abbreviations). In case the name differs from that eti ‑ Ethiopia currently recognized (e.g. the country no longer exists or its name has changed or it does not GAB ‑ Gabon exist in English), it is given in quotes followed by a three-letter abbreviation of the currently GAM ‑ Gambia accepted name in brackets – e.g. “Congo-Belge“ [=DOC]. All geographic data accompanying GBi ‑ Guinea-Bissau the type locality in the species description are underlined, e.g. e AF: KEN, etc. This is followed GHa ‑ Ghana by the name of a province or a region of non-administrative nature, e.g. e AF: TAN: Arusha GUi ‑ Guinea Reg. or c AF: DOC: Orientale Prov., etc. The name of the holotype locality is followed (where ken ‑ Kenya available) by the elevation in meters or feet, GPS coordinates, paratype locality (one or more), lir ‑ Liberia its elevation and GPS coordinates, etc. (paratype localities are ordered alphabetically), and the MAD ‑ Madagascar information is supplemented in plain type by other known occurrences down to regions, to make MAW ‑ Malawi the geographic distributions more complete – e.g. e AF: “Kobbo, dans le pays de Gallas-Raïas“ Mli ‑ Mali [=n ETI: Amhara Reg.: Kobo]; Afar Reg. means that the species is known only from Etiopia, MOS ‑ Mozambique Amhara region, the type locality is Kobo, but finds are known also from another region. Mri ‑ Mauritania Analogously with the previous catalogue, the list of three-letter abbreviations is substantially NAB ‑ Namibia enlarged to include also provinces (regions) of the following countries: Argentina, Australia, NIG ‑ Niger Brazil, Canada, China, India, Mexico, Russia and USA. Maps are provided for Democratic nre ‑ Nigeria Congo, Ethiopia, Madagascar, Mozambique, Namibia, Somalia, South Africa and Tanzania, rca ‑ (Republic of) Central Africa (Centrafrique) whose administrative divisions have recently changed. rci ‑ (Republic of) Ivory Coast (Republique de Côte d´Ivoire) RSa ‑ (Republic of) South Africa Anthia promontorii (Péringuey, 1898) sn. Obst 1901 is an erroneous name recently treated Sen ‑ Senegal as a synonym of Apristus promontorii (Péringuey, 1898) (Carabidae, Lebiini) sn. Mawdsley Sln ‑ Sierra Leone et al. 2011. SOM ‑ Somaliland Anthia tatumana White, 1846 (which is treated by websites Carabidae of the world 2007- STP ‑ São Tomé e Príncipe (St. Thomas and Principe Island) 2002 as “species incertae sedis“ in the genus Anthia Weber, 1801) is treated here according to SUD ‑ Sudan notices of Methuen (1846) and Guérin-Méneville (1852) as a species of the genus Cypholoba SWa ‑ Swaziland Chaudoir, 1850 near the caillaudi/macilenta group. tan ‑ Tanzania Creagris bechuanus Péringuey, 1910 is a synonym (preoccupied name) for Cueta trivirgata TCH ‑ Chad (Gerstaecker, 1885) (Neuroptera, Myrmeleontidae). TOG ‑ Togo Helluo pygmaeus Dejean, 1826 is treated now (sn. Le Conte 1851) as Thalpius Le Conte, UGa ‑ Uganda 1851 (Carabidae, Dryptinae, Zuphiini).

262 263 WSa ‑ Western Sahara CUB ‑ Cuba ZAM ‑ Zambia DOM ‑ Dominican Republic (Hispaniola Island) ZIM ‑ Zimbabwe FRG ‑ Guiane Française (French Guiana) AM ‑ America MEX ‑ United States of Mexico / federal states ARG ‑ Argentina / provinces Bcn ‑ Baja California del Norte Bua ‑ Buenos Aires Bcs ‑ Baja California del Sur Chb ‑ Chubut Cam ‑ Campeche Chc ‑ Chaco Cia ‑ Chiapas Crd ‑ Córdoba Clm ‑ Colima Crr ‑ Corrientes Chh ‑ Chihuahua Ctm ‑ Catamarca Coh ‑ Coahuila Eri ‑ Entre Ríos Dfe ‑ Distrito Federal Juy ‑ Jujuy Dur ‑ Durango Lrj ‑ La Rioja Gue ‑ Guerrero Mdz ‑ Mendoza Guj ‑ Gunajuato Mns ‑ Misiones Hid ‑ Hidalgo Neq ‑ Neuquén Jal ‑ Jalisco Pam ‑ La Pampa Mex ‑ Mexico Rng ‑ Río Negro Mih ‑ Michoacan Sat ‑ Salta Mor ‑ Morelos Scz ‑ Santa Cruz Nay ‑ Nayarit Sfe ‑ Santa Fe Nle ‑ Nuevo Leon Sge ‑ Santiago del Estero Oax ‑ Oaxaca Snl ‑ San Luis Pub ‑ Puebla Tuc ‑ Tucumán Qer ‑ Queretaro Bra ‑ Brazil / federal states Qro ‑ Quintana-Róo Ama ‑ Amazonas (North Region) Sin ‑ Sinaloa Bhi ‑ Bahia (Northeast Region) Slp ‑ San Luis Potosí Eps ‑ Espiritu Santo (Southeast Region) Son ‑ Sonora Goi ‑ Goiás (Central-West Region) Tab ‑ Tabasco Mgr ‑ Mato Grosso (Central-West Region) Tam ‑ Tamaulipas Mgs ‑ Mato Grosso do Sul (Central-West Region) Ver ‑ Veracruz Msg ‑ Minas Gerais (Southeast Region) Yuc ‑ Yucatan Pra ‑ Pará (North Region) Zac ‑ Zacatecas Pnb ‑ Pernambuco (Northeast Region) Par ‑ Paraguay Pui ‑ Piauí (Northeast Region) PUr ‑ Puerto Rico Rdj ‑ Rio de Janeiro (Southeast Region) URU ‑ Uruguay Rgn ‑ Rio Grande do Norte (Northeast Region) TND ‑ Trinidad Rgs ‑ Rio Grande do Sul (South Region) USa ‑ United States of America / federal states Ron ‑ Rondonia (North Region) Aba ‑ Alabama (se) Sac ‑ Santa Catarina (South Region) Ari ‑ Arizona (sw) Spo ‑ São Paulo (Southeast Region) Ark ‑ Arkansas (se) can ‑ Canada / provinces Ont ‑ Ontario

264 265 Col ‑ Colorado (wc) Gan ‑ Gansu Flo ‑ Florida (se) Gax ‑ Guangxi Gor ‑ Georgia (se) Gui ‑ Guizhou Ili ‑ Illinois (ce) Heb ‑ Hebei Ina ‑ Indiana (ce) Hei ‑ Heilongjiang Iow ‑ Iowa (ce) Hen ‑ Henan Kan ‑ Kansas (wc) Hgk ‑ Hong Kong Ken ‑ Kentucky (ce) Hub ‑ Hubei Lou ‑ Louisiana (s) Hun ‑ Hunan Mar ‑ Maryland (e) Jia ‑ Jiangxi Mas ‑ Massachusetts (e) Jil ‑ Jilin Mis ‑ Mississippi (wc) Jis ‑ Jiangsu Mso ‑ Missouri (wc) Lin ‑ Liaoning Mon ‑ Montana (n, nw) Nem ‑ Nei Mongol (Inner Mongolia) Nca ‑ North Carolina (se) Nix ‑ Ningxia Nda ‑ North Dakota (n) Qin ‑ Qinghai Nje ‑ New Jersey (e) Sch ‑ Sichuan Nme ‑ New Mexico (sw) Sgh ‑ Shanghai Nyo ‑ New York (ne) Sha ‑ Shaanxi Ohi ‑ Ohio (ce) Shn ‑ Shandong Okl ‑ Oklahoma (wc) Shx ‑ Shanxi Pen ‑ Pennsylvania (e) Xij ‑ Xinjiang (Uyguristan) Rdi ‑ Rhode Island (e) Xiz ‑ Xizang (Tibet) Sca ‑ South Carolina (se) Yun ‑ Yunnan Sda ‑ South Dakota (wc) Zhe ‑ Zhejiang Ten ‑ Tennessee (se) CYP ‑ Cyprus Tex ‑ Texas (s) IDn ‑ Indonesia Usv ‑ Virgin Islands of the United States (Caribbean Sea) IND ‑ India / federal states Vir ‑ Virginia (e) Adh ‑ Andhra Pradesh Wdc ‑ District of Columbia (Washington City and vicinity) (ne) Aru ‑ Arunachal Pradesh Wis ‑ Wisconsin (ce) Ass ‑ Assam Wvi ‑ West Virginia (e) Ben ‑ West Bengal Wyo ‑ Wyoming (wc) Bih ‑ Bihar Ven ‑ Venezuela Cht ‑ Chhattisgarh Hip ‑ Himachal Pradesh AS ‑ Asia Kas ‑ Jammu & Kashmir AFG ‑ Afghanistan Jhr ‑ Jharkhand Ban ‑ Bangladesh Ker ‑ Kerala CAB ‑ Cambodja Krt ‑ Karnataka CHi ‑ The People‘s Republic of China / provinces Map ‑ Manipur Anh ‑ Anhui Mhy ‑ Meghalaya Bei ‑ Beijing (Pekin) Mhp ‑ Madhya Pradesh Fuj ‑ Fujian Mhr ‑ Maharashtra Gad ‑ Guangdong Nwd ‑ New Delhi

266 267 Ori ‑ Orissa Qld ‑ Queensland Pud ‑ Puducherry Sau ‑ South Australia Pun ‑ Punjab Tas ‑ Tasmania Raj ‑ Rajasthan Vic ‑ Victoria SiD ‑ Sikkim, Darjeeling Wau ‑ Western Australia Tmd ‑ Tamil Nadu PNG ‑ Papua-New Guinea Utk ‑ Uttarkhand Utp ‑ Uttar Pradesh c ‑ central (ec = east-central etc.) irn ‑ Iran e ‑ eastern ISr ‑ Israel n ‑ northern JAP ‑ Japan s ‑ southern JOr ‑ Jordan w ‑ western KAZ ‑ Kazakhstan env. ‑ environs kir ‑ Kirgizstan Is(l).. ‑ island(s) KUW ‑ Kuwait Mt(s) ‑ mountain(s) LAO ‑ Laos Pen. ‑ peninsula Mal ‑ Malaysia Dept. ‑ department MOG ‑ Mongolia Prov(s). ‑ province(s) MYa ‑ Myanmar (Burma) Reg(s). ‑ region(s) NEP ‑ Nepal Riv(s). ‑ river(s) NCO ‑ North Korea (People´s Democratic Republic of Korea) JH ‑ junior homonym Pak ‑ Pakistan nn ‑ nomen novum PHi ‑ Philippines t ‑ type species of the genus RUS ‑ Russia Sg. ‑ subgenus Fea ‑ Far East (Asian) Territory sn. ‑ sensu Esb ‑ East Siberia syn. ‑ synonym Wsb ‑ West Siberia SCO ‑ South Korea (Republic of Korea) Srl ‑ Sri Lanka TAJ ‑ Tajikistan THa ‑ Thailand TUM ‑ Turkmenistan TUr ‑ Turkey TWa ‑ Taiwan (The Republic of China, Taipei) Uae ‑ United Arab Emirates UZB ‑ Uzbekistan Vie ‑ Vietnam

AU ‑ Australia and territories in southwestern Pacific Ocean AUS ‑ Australia / federal states Act ‑ Australian Capital Territory (Canberra) Nsw ‑ New South Wales Nte ‑ Northern Territory

268 269 11 8

10 12 5 1 5 7 13 11 6 2 9 8 3 10 2 6 3 3 4 9 4 3 2

5- 7 11 5 1- 7 - 7- 1 8 4 6 2 4 1 8 9 10 9 6 Fig. 5. Administrative divisions (regions) of Namibia (=NAB) 1. Caprivi, 2. Erongo, 3. Hardap, 4.Karas, Fig. 6. Administrative divisions (provinces) of the Republic of Fig. 2. Administrative divisions (regions) of Ethiopia 5. Okavango, 6. Khomas, 7. Kunene, South Africa (=RSA) (=ETI) 8. Ohangwena, 9. Omaheke, 10. Omusati, 1. Eastern Cape, 2. Free State, 3. Gauteng, 4. KwaZulu-Natal, 1. Addis Ababa, 2. Afar, 3. Amhara, 4. Benishangul- 11. Oshana, 12. Oshikoto, 13. Otjozondjupa 5. Limpopo, 6. Mpumalanga, 7. Nothern Cape, 8. North West Fig. 1. Administra-tive divisions (provinces) of the Gumuz, 5. Dire-Dawa, 6. Gambela, 7. Harari, 8. Oromiya, Province, 9. Western Cape. Democratic Republic of the Congo (=DOC) 9. Somali, 10. Southern Nations, Nationalities, and 1. Bandundu, 2. Bas-Congo, 3. Équateur, 4. Kasai- People´s Region, 11. Tigray Ooccidental, 5. Kasai-Oriental, 6. Katanga, 7. 18 13 Kinshasa, 8. Maniema, 9. Nord-Kiwu, 10. Orientale, 15 5 11. Sud-Kiwu 17 10 16 14 14 12 20 1

1 11 6 7 8 9 -15 2 22 23 10 -16 25 7 5 9 21 3 10 26 18 11 8 24 4 3 4 6 2 5 -7 4 17 11 9 1 4 2 12 1 8 Fig. 3. Administrative Fig. 4. Administrative divisions 19 (provinces) of Mozambique divisions (provinces) of 6 3 13 3 (=MOS) Madagascar (=MAD) 2 Fig. 7. Administrative divisions (regions) of 1.A ntananarivo, 2.A ntsi- 1. Cabo Delgado, 2. Gaza, Somalia (=SOM) ranana, 3. Fianarantsoa, 3. Inhambane, 4. Manica, 5. Maputo 1. Jubbada Hoose (=Lower Juba), 2. Jubbada Fig. 8. Administrative divisions (regions) of Tanzania 4. Mahajanga, 5. Toa- 6 -5 City, 6. Maputo, 7. Nampula, Dhexe (=Middle Juba), 3. Gedo, 4. Bay, (=TAN) masina, 6. Toliara 8. Niassa, 9. Sofala, 10. Tete, 5. Bakool, 6. Shabeellaha Hoose (=Lower 1. Arusha, 2. Dar Es Salaam, 3. Dodoma, 4. Iringa, 5. Kagera, 11. Zambezia Shabele), 7. Banaadir (=Benadir), 8. Shabeellaha 6. Kigoma, 7. Kilimanjao, 8. Lindi, 9. Manyara, 10. Mara, 11. Dhexe (=Middle Shabele), 9. Hiiraan, Mbeya, 12. Morogoro, 13. Mtwara, 14. Mwanza, 15. Pemba 10. Galguduud, 11. Mudug, 12, Nugaal, 13. Bari, north, 16. Pemba south, 17. Pwani, 18. Rukwa, 19. Ruvuma, 14. Sool, 15. Sanaag, 16. Togdheer, 17. Woqooyi 20. Shinyanga, 21. Singida, 22. Tabora, 23. Tanga, 24. Zanzibar Galbeed, 18. Awdal central/south, 25. Zanzibar north, 26. Zanzibar urban/west

270 271 ANTHIINAE Bonelli, 1813 ssp. adulatrix Kolbe, 1894 “Tanganjika-See“ [=w TAN: Kigoma, Rukwa Regions]; ANTHIINI Bonelli, 1813 (subtribes: 2; genera: 10; species: 125) Mbeya, Ruvuma Regions 5 lefebvrei Guérin-Méneville, 1847 e AF: “Abyssinie“ [=ETI]: ANTHIINABonelli, 1813 (genera: 5; species: 55) Tigray Reg.; DJI, ERI, ETI: ANTHIAWeber, 1801; T: Carabus sexguttatus Fabricius, 1775 (species: 14) Oromiya, Tigray Regions; =Pachymorpha Hope, 1838; T: Carabus sexguttatus Fabricius, 1775 KEN: Rift Valley Prov.; SSD =Thoracolobus Gistel, 1857; T: Anthia thoracica Thunberg, 1784 6 lunae Thomson, 1859b e, w AF: BEN, BFO, CAM, =Calanthia Obst, 1901; T: Anthia pulcherrima Bates, 1888 CON, DOC, GHA, KEN, NRE, =Odontanthia Obst, 1901; T: Anthia calida Harold, 1878b RCA, SSD, SUD, TAN, TCH, TOG, UGA ANTHIA s. str. (11 species) ssp. lunae s. str. “Région du Nil Blanc“ [=SSD]; CAM, CON, ne DOC: calida species (Häckel sn. Obst 1901b; = Sg. Odontanthia Obst, 1901b sn. Kleinfeld & Orientale Prov.; NRE, RCA, Puchner2012): SUD, TCH 1 calida Harold, 1878b “innern Guinea: Kabebe“ [=ne =tomentosa Chaudoir, 1861 “Kordofan“ [=SSD] ANG]: Lunda Norte, Malanje, ssp. bordasii Roger, 1899 “Dahomey: cimetière des Uíhe Provinces; s DOC: Bas- Amazones, ďAbomey“ [=s Congo, Kasai-Occidental, BEN: Zou Dept.]; BFO, GHA, Katanga Provinces s NRE, TOG =convexipennis Putzeys, 1880 ANG ssp. martensi Obst, 1901b “Deutsch-Ostafrika: Iringa; Uhehe; Weg von Ubena bis ferox species (Häckel sn. Obst 1901b) Langenburg“ [=sw TAN: Iringa 2 ferox Thomson, 1859a e AF: “Afrique mérid. orient.?“ Reg.]; KEN: Rift Valley Prov.; [=SOM]; ERI, n ETI: Tigray, UGA Oromiya Regions; DJI, SSD =fallax Obst, 1901b “Uhehe, Iringa in Deutsch- [sn. Raffray 1885] Ostafrika“ [=sw TAN: Iringa Reg.] lefebrvei group (Häckel sn. Obst 1901b; = Sg. Anthia s. str. sn. Kleinfeld & Puchner 2012) 3 artemis Gerstaecker, 1884 e AF: “Kilimandjaro“ [=ne sexguttata species (Häckel sn. Obst 1901b; = Sg. Pachymorpha sn. Kleinfeld & Puchner TAN: Kilimanjaro Reg.]; 2012) Arusha, Mwanza Regions; 7 sexguttata (Fabricius, 1775) Carabus s AS: AFG, BAN, IND, IRN, BUR, ETI: Oromiya Reg.; KAZ, NEP, PAK, SRL, TUM, KEN: Rift Valley Prov.; UGA UZB =angulata Obst, 1901b “Südost-Victoria-Nyansa“ ssp. sexguttata s. str. “Suratte Indiae“ [w IND: Gur: [=nw TAN: Mwanza Reg.] Surat]; And, Ass, Ben, Har, 4 bucolica Kolbe, 1894 e AF: BUR, e DOC, KEN, Hip, Jhr, Kas, Ker, Krt, Map, MAW, TAN, UGA, ZAM Mhy, Mhp, Mhr, Nwd, Ori, ssp. bucolica s. str. “bei Bukoba am Westufer des Pun, Raj, SiD, Tmd, Tri, Utk, Victoria-Njansa“ [=nw TAN: Utp; BAN, NEP, PAK, SRL Kagera Reg.]; Iringa Reg.; =thunbergi Fischer von Waldheim, 1806 [IND] BUR, e DOC: Sud-Kiwu Prov.; =orientalis Hope, 1838 Pachymorpha “circa Poonah in India orientali“ KEN: Rift Valley Prov.; MAW, [=ne IND: Mhr: Pune] UGA, ZAM

272 273 =indica Chaudoir, 1861 “Indes orientales“ [=BAN, e Ohangwena, Omusati, Oshana, IND] Oshikoto Regions =elliptica Motschulsky, 1864 “Tranquebar“ [=se IND: Tmd] =ooptera Bates, 1889 “Mossamedes (Sala)“ [=sw ssp. mannerheimi Chaudoir, 1842 “Prov. de Mazendéran, près de anG: Namibe Prov.] Astrabad“ [=IRN: Mazandaran 9 circumscripta Klug, 1853 c, e, s AF: ANG, BOW, DOC, Province: Gorgan]; Amu-Darja KEN, MAW, MOS, NAB, (Perse)“ [=TUM]; KAZ, TUM, RSA, TAN, ZAM, ZIM UZB ssp. circumscripta s. str. “Mossambique: Tette“ [=n ssp. afghana Anichtchenko, 2009 c AFG: Oruzgan [Uruzgan MOS: Tete Prov.]; BOW, Prov.], Sahrestan; 2400m; DOC: Katanga Prov.; KEN, 2500m; Ghazni [Prov.], MAW, NAB: Caprivi, Hardap, Karmay sw Moqur; AFG, PAK Ohangwena, Okavango, Omusati, Oshana, Oshikoto, thoracica group (Häckel sn. Obst 1901b = Sg. Anthia s. str. sn. Kleinfeld & Puchner 2012) Otjozondjupa Regions; 8 cinctipennis [Dupont in litt.] Lequien, 1832 s AF: ANG, BOW, LES, MOS, RSA: Gauteng, Free State, NAB, RSA, SWA, TAN, ZAM, Eastern Cape, KwaZulu- ZIM Natal, Northern Cape, North ssp. cinctipennis s. str. “Cap de Bonne-Espérance“ West Provinces; TAN: Iringa, [=RSA]: Eastern Cape, Free Mbeya, Mtwara, Ruvuma State, Gauteng, KwaZulu- Regions, ZAM, ZIM Natal, Limpopo, Mpumalanga, =striatopunctata Guérin-Méneville, 1847 “Abyssinie“ [=ETI] probably Northern Cape, North West by error Provinces; s ANG: Cuando =vitticollis Bates, 1878 “Lake Nyassa“ [=n MAW] Cubango, Cunene Provinces; =burdoi Fairmaire, 1882a “Zanguebar“ [=e TAN] BOW, LES, MOS: Maputo =ovampoensis Péringuey, 1892a “Ovampoland“ [=n NAB: Prov.; NAB: Erongo, Hardap, Ohangwena, Omusati, Oshana, Karas, Khomas, Kunene, Oshikoto Regions] Omaheke, Otjozondjupa =fochi Bénard, 1919 “Umtali, dans le Manicaland Regions; SWA, ZAM, ZIM (Rhodésia du Sud)“ [=e ZIM: =marginipennis Laporte de Castelnau, 1835 “Cafrerie“ [=MOS, RSA, ZIM] Manicaland Prov.: Mutare] =limbipennis Chaudoir, 1861 MOS ssp. niveicincta Bates, 1889 Humpata [=sw ANG: Namibe =pachyoma Chaudoir, 1883 “Transvaal“ [=ne RSA: Prov.]; Mossamedes (Sala) Gauteng, Limpopo, [=w ANG: Namibe, Zaire Mpumalanga, North-West Provinces] Provinces] =duparqueti Lucas, 1867 “cap Négro“ [=sw ANG: =hottentota Oliff, 1889 “Matabeleland“ [=ZIM] Namibe Prov.: Cabo Negro] =dissimilis Obst, 1901b “Rietfontein (Deutsch- ssp. omostigma Chaudoir, 1866 “Benguéla“ [=w ANG: Südwestafrika)“ [=c NAB: Benguela Prov.]; Bihé, Cunene Omaheke Reg.] Provinces =fairmairei Sternberg, 1907a “Namaland“ [=s NAB: Hardap, 10 maxillosa [Olivier in litt.] (Fabricius, 1781) Carabus s, e AF: “Cap. bon. sp.“ [=s karas Regions] RSA]: Eastern Cape, Free ssp. actaeon Erichson, 1843 ANG: Cunene, Namibe State, Limpopo, Mpumalanga, Provinces; n NAB: Kunene, Northern Cape, North West,

274 275 Western Cape Provinces; s =dohrni Rousseau, 1906 Anthia MOS: Maputo Prov.; s NAB: nn for Anthia portentosa Dohrn, 1882 Hardap, Karas Regions ssp. luculediensis Sternberg, 1906b “Südöstliches D.-Ostafrika: =agilis Thunberg, 1784 Carabus “Capite bonae spei“ [=s RSA] Lukuledi; Mikindani [=TAN: =atra Chaudoir, 1843 “Cap de Bonne-Espérance“ [=s Mtwara Reg.]; Kilwa [=Lindi rSA] Reg.]; Mahenge Distr.; =fabricii Crotch, 1871 [=Morogoro Reg.]; Madibira nn for Carabus maxillosus Fabricius, 1781 [=Mbeya Reg.]; Ugoni“ =algoa Péringuey, 1896 “Mozambique (Rikatla)“ [=s [=Ruvuma Reg.]; s KEN: Coast MOS: Maputo Prov.] Prov. =khoina Péringuey, 1899 “w Karroo: (Fraserburg, Worcester); Namaqualand BAEOGLOSSAChaudoir, 1850; T: Anthia villosa Thunberg, 1806 (species: 2) (Concordia, Garies); nw part of the [Cape] Colony (Calvinia, 1 melanaria (Boheman, 1848) Anthia e AF: “Cafrerie“ [=n RSA, Pella, Bushmanland)“ [=s ZAM, ZIM]: Free State, RSA: Northern Cape, Western Northern Cape, North West, Cape Provinces] Limpopo Provinces; BOW, 11 thoracica (Thunberg, 1784) Carabus s AF: BOW, KEN, LES, MAW, NAB: Caprivi, Erongo, MOS, NAB, RSA, SWA, TAN, Hardap, Khomas, Kunene, ZAM, ZIM Okavango, Omaheke, ssp. thoracica s. str. “Capite bonae spei“ [=s RSA]: Otjozondjupa Regions Eastern Cape, Free State, =anthracina (Guérin-Méneville, 1849) Anthia “Caffraria interior“ [=n RSA, Gauteng, KwaZulu-Natal, ZAM, ZIM] Limpopo, Mpumalanga, 2 villosa (Thunberg, 1806) Anthia s AF: “Caput bonae spei“ [=s Northern Cape, North West, RSA]: Eastern Cape, Free Western Cape Provinces; State, Northern Cape, Western BOW, LES, MAW, s MOS: Cape Provinces Inhambane, Maputo Provinces; =decemsulcata Bonnelli, 1813 Anthia “Cap de Bonne-Espérance“ [=s n NAB: Caprivi, Erongo, RSA] Hardap, Kunene, Ohangwena, Okavango, Omusati, Oshana, CHILANTHIAObst, 1901b; T: Anthia cavernosa Gerstaecker, 1867 (1 species) Oshikoto, Otjozondjupa Regions; SWA, ZAM, ZIM 1 cavernosa Gerstaecker, 1867 e AF: “ad lacum Jipe“ [=ne =fimbriatus (Thunberg, 1784) Carabus “Capite bonae spei“ [=s RSA] TAN: Kilimanjaro Reg.]; =maculicollis Perroud, 1846 Anthia “ ľ intérieur de Natal“ [=se “Mombas“ [=se KEN: Coast RSA: KwaZulu-Natal Prov.] Prov.: Mombasa Distr.]; se =portentosa Dohrn, 1882 Anthia “südafrikanische Ausbeute des SOM: Shabeellaha Dhexe Reg. JH of Fairmaire, 1882 dr. Holub“ [=BOW, ne NAB, RSA, ZAM, ZIM] CYCLOLOBA Chaudoir, 1850; T: Anthia septemguttata Fabricius, 1794 =stigmodera [Chaudoir in litt.] Péringuey, 1896 (species: 2) Anthia “Manica [=e ZIM: Manicaland CYCLOLOBA s. str. (2 species) Prov.]; Ovampoland“ [=n NAB: Ohangwena, Omusati, 1 septemguttata (Fabricius, 1794) Carabus s AF: “Cap. Bon. spei.“ Oshana, Oshikoto Regions] [=RSA]: Eastern Cape, Free

276 277 State, Gauteng, KwaZulu- =nyassica Sternberg, 1906a Anthia “Langenburg (Nyassa-See Natal, Mpumalanga Provinces; bor.)“ [=sw TAN: Mbeya BOW, SWA Reg.:Tukyu] =sexnotatus Thunberg, 1806 Carabus “Promontorium bonae spei“ [=s RSA] TERMOPHILUM Basilewsky, 1950; T: Carabus decimguttatus Linnaeus, =pilosa Boheman, 1848 Anthia “Montes Makkalienses“ [=c 1764 (32 species) RSA: Gauteng Prov.] =Thermophila Hope, 1838; T: Carabus decimguttatus Linnaeus, 1764 2 truncatipennis (Boheman, 1848) Anthia s AF: “Caffraria interior“ [=n =Thermophilum misspelling RSA, ZAM, ZIM]: KwaZulu- Natal, Northern Cape biguttatum group (Häckel sn. Obst 1901b): Provinces; LES biguttatum subgroup (Häckel sn. Kleinfeld & Puchner 2012) =delegorguei Guérin-Méneville, 1847 Anthia “ Natal“ [=se RSA: KwaZulu- 1 biguttatum (Bonelli, 1813) Anthia s AF: type locality not stated; Natal Prov.] “Cap de Bonne-Espérance“ =acuticollis Thomson, 1857a Anthia “ Natal“ [=se RSA: KwaZulu- [=RSA sn. Basilewsky 1980]: Natal Prov.] Eastern Cape, Western Cape Provinces ACANTATNHIA s. str. (1 species) =costata Gory, 1836 Anthia “Cap de Bonne-Espérance“ [=s RSA]: Western Cape Prov. 3 mirabilis (Sternberg, 1906a) Anthia c AF: “San Benito Flufs =sexcostata Chaudoir, 1848 Anthia “Cap de Bonne-Espérance“ [=s (Congo)“ [=c CON: Saint RSA] Benoit (Boundji)]; GAB =reichei [Dejean] Obst, 1901b Anthia “ Cap de Bonne-Espérance“ [=s CALANTHIA s. str. (3 species) RSA] 2 capillatum (Obst, 1901b) Anthia sw AF: “Hereroland (Deutsch- 4 ida (Kolbe, 1894) Anthia e AF: “bei Bukoba am Südwestafrika)“ [=c NAB: Westufer des Victoria-Njansa“ Erongo, Khomas, Kunene, [=nw TAN: Kagera Reg.]; Omaheke, Otjozondjupa Dodoma, Mwanza, Shinyanga, Regions]; s ANG: Cuango Singida, Tabora Regions Cubango, Cunene Provinces 5 oberthuri (Obst, 1906) Anthia e AF: BOW, MAW, n RSA: 3 mimum (Péringuey, 1896) Anthia s AF: “Carnarvon; Fraserburg“ nn for Anthia pulcherrima Bates sn. Obst 1901b limpopo Prov. [nw RSA: Northern Cape =pulcherrima Obst, 1901b Anthia “ Ubemba in Brit.-O.-Afrika Prov.] [=n MAW: Northern Prov.]; 4 namaquum (Péringuey, 1896) Anthia s AF: “Small Namaqualand Ubena-Langenburg in D.- (O‘okiep)“ [nw RSA: Northern O.-Afrika; Konde-Unyika, Cape Prov.]; North West, nordöstlich vom Nyassa-See“ Western Cape Provinces [=sw TAN: Mbeya Reg.]; =propinqua Péringuey, 1896 Anthia “Small Namaqualand (O‘okiep) BOW, n RSA: North West [=nw RSA: Northern Cape Prov.; ZIM Prov.]; Springbokfontein“ 6 pulcherrima (Bates, 1888) Anthia se AF: “River Lujenda, East [=North West Province] Africa“ [=nw MOS: Niassa =affinis Obst, 1901b Anthia “Orlog River, Capland“ [=w Prov.: Lugenda Riv./ sw RSA: Northern Cape Prov.] TAN: Iringa, Mbeya, Rukwa, =parallela Obst, 1901b Anthia “Troe-Troe (West-Capland)“ Ruvuma Regions]; se DOC: [=s RSA: Western Cape Prov.] Katanga Prov.; ZAM

278 279 cephalotes subgroup (Häckel sn. Kleinfeld & Puchner 2012) State, Gauteng, KwaZulu- 5 aequilaterum (Klug, 1853) Anthia se AF: “Mossambique: Natal, Limpopo, Mpumalanga, Tette“ [=n MOS: Tete Prov.]; North West, Western Cape Zambézia Prov.; BOW, s KEN: Provinces; BOW Coast Prov.; MAW, NAB: ssp. kolbei Obst, 1901b “Outyo, Deutsch- Caprivi, Okavango Regions; Südwestafrika“ [=n NAB: TAN: Dodoma, Singida Kunene Reg.: Outjo]; Caprivi, Regions, ZIM Erongo, Khomas, Kunene, =menyharti Brancsik, 1897 Anthia “Zambezi, Boroma“ [=c MOS: Okavango, Ohangwena, Zambézia Prov.: Morrumbala] Omaheke, Omusati, Oshama, 6 babaulti (Bénard, 1921a)* Anthia e AF: “Afrique Orientale: Oshikoto, Otjozondjupa Kedong Valley“ [=s KEN: Rift Regions; s ANG: Cunene Prov.; Valley Prov.]; SSD, n TAN: w BOW; w RSA: Northern Arusha, Mara Regions Cape Prov. =desertorum Thomson, 1859b* Anthia “Région du Nil Blanc“ [=SSD] 10 machadoi (Basilewsky, 1955a) Anthia sw AF: “Moçâmedes: Deserto 7 cephalotes (Guérin-Méneville, 1845) Anthia se AF: “reg. Massilicatzi“ [=s de Moçâmedes“ [=sw ANG: MOS: Gaza, Maputo Provinces]; Namibe Prov.]; nw NAB: BOW; e, n RSA: KwaZulu- Kunene Reg. Natal, Limpopo, Mpumalanga, 11 ovampoa (Péringuey, 1896) Anthia sw AF: ANG, BOW, NAB, RSA Northern Cape, North West ssp. ovampoa s. str. “Ovampoland“ [=n NAB: Provinces; SWA, ZIM Ohangwena, Omusati, Oshana, =bimaculata Perroud, 1846 Anthia “ľ intérieur de Natal“ [=se Oshikoto Regions]; s ANG: RSA: n KwaZulu-Natal Prov.] Cuanza Norte, Cuango- =binotata Perroud, 1846 Anthia “Natal“ [=se RSA: n KwaZulu- Cubango, Cunene Provinces; Natal Prov.] n BOW; se RSA: KwaZulu- =immaculata Wallengreen, 1881 Anthia “Transvaal: Waterberg“ [=ne Natal Prov. RSA: Limpopo Prov.] =alvesi Sternberg, 1906b Anthia “Natal“ [=se RSA: KwaZulu- =torva Péringuey, 1896 Anthia “Transvaal (Middelburg) [=ne Natal Prov.] RSA: Mpumalanga Prov.]; ssp. cunenense [Basilewsky in litt.] Kleinfeld & Puchner, 2012 “N.-W-Namibia, Middle Limpopo [=Limpopo Marienfluss; 17°36‘S, Prov.]; Mozambique 12°31‘E; Junction east branch (Lebombo)“ [=s MOS: Maputo Marienfluss-Cunene Riv., nw Prov.] Kaokoveld“ [=s ANG: Cunene =sternbergi Obst, 1903 Anthia “Cap der guten Hoffnung“ [=s Prov.] RSA] 8 csikii (Obst, 1906) Anthia sw AF: “Kubub, D.-S.-W.- burchelli group (Häckel sn. Obst 1901b): Afrika“ [=s NAB: Karas Reg.]; 12 brevivittatum (Obst, 1901b) Anthia e AF: “Uhehe, Iringa (Deutsch- w RSA: Northern Cape Prov. Ostafrika)“ [=ec TAN: Iringa Reg.]; Morogoro, Pwani, Dar limbatum subgroup (Häckel sn. Kleinfeld & Puchner 2012) Es Saalam Regions 9 limbatum (Dejean, 1831) Anthia sw AF: w BOW, ne NAB, RSA 13 burchelli (Hope, 1832) Anthia c, s, se AF: BOW, DOC, MAW, ssp. limbatum s. str. “Cap de Bonne-Espérance“ MOS, NAB, RSA, TAN, ZAM, [=s RSA]: Eastern Cape, Free ZIM

280 281 ssp. burchelli s. str. unknown [“Cafrerie“ sn. Exploration“ (=n BOW, s Lequien 1832 =BOW, RSA, ZAM) sn. Dohrn 1881]; BOW, ZAM, ZIM]: s MOS: Gaza, n NAB: Ohangwena, Omaheke, Inhambane, Maputo Provinces; Omusati, Oshana, Oshikoto NAB: Caprivi, Erongo, Regions; n RSA: Free State, Kunene Regions; KwaZulu- Northern Cape, North West Natal, Limpopo, North West Provinces Provinces; s TAN: Lindi, =betschuana Obst, 1908a Anthia “Betschuanaland-Protekrorat: Mbeya Regions Mookane (Kalahari); =nuda Sternberg, 1906b Anthia “Sikumba; Maputa im Lookaneng-Severelela südlichen Portugiesisch Ost- (Kalahari); Khakhea-Kgokong Afrika“ [=s MOS: Maputo (Kalahari)“ [=n BOW] Prov.] 16 alternatum (Bates, 1878) Anthia s AF: “Lake Nyassa“ [=sw =maculata Sternberg, 1906b Anthia “Lindi (D.-Ostafrika) [=se TAN/ n MOS]; s DOC: TAN: Lindi Reg.]; Langenburg Katanga Prov.; s KEN: Eastern (Nyassa-See)“ [=sw TAN: Prov.; c, n MOS: Niassa, Mbeya Reg.: Tukuyu] Sofala, Zambézia Provinces; ssp. petersi (Klug, 1853) Anthia “Mossambique: Tette“ [=n c, s, se TAN: Dodoma, Iringa, MOS: Tete Prov.]; Niassa Lindi, Mbeya, Mtwara, Tabora Prov.; s DOC: Katanga Prov.; Regions MAW, ZAM, ZIM 17 andersoni (Chaudoir, 1861) Anthia s, sw AF: ANG, BOW, DOC, ssp. upembanum Basilewsky, 1953 “P. N. U., Mabwe“; 585m [=se NAB, RSA, ZAM, ZIM DOC: Katanga Prov.: Parc ssp. andersoni s. str. “lac N´gami“ [=nw BOW: National de ľUpemba] Ngami Lake]; s ANG: Cuando 14 praesignis (Bates, 1888) Anthia s AF: “Mamboia“ [=nw MOS: Cubango, Cunene Provinces; Niassa Prov.]; s TAN: Ruvuma n NAB: Hardap, Khomas, Reg. Ohangwena, Omaheke, 14 sexmaculatum (Fabricius, 1787) Carabus n AF: “Barbaria“ [=ALG, LIB, Omusati, Oshana, Oshikoto, MOR, TUN]; EGY, MLI, MRI, Otjozondjupa Regions; nw NIG, SSD, SUD, TUN, TCH, RSA: Northern Cape, North WSA; sw AS: SYR West Provinces =marginata Latreille, 1823 Anthia; syn. nov. “El-Ouâh el Bahryeb (Nubie)“ =westermanni Dohrn, 1883 Anthia “ Ad flumen Zambese“ [=BOW, [=SSD] ne NAB] =pharaonum Bedel, 1914 Anthia “désert arabique ďÉgypte“ ssp. bennigseni Sternberg, 1906b Anthia type locality not stated, s DOC: [=EGY] Bandundu, Kasai-Occidental =singularis Bedel, 1914 Anthia “Sud Oranais“ [=ALG] Provinces; n ANG: Malanje =gafsana Bodemeyer, 1931 Anthia TUN: Oasis Gafsa Prov. ssp. crudelis Harold, 1878b Anthia “innern Guinea: Kabebe“ massilicatum group (Häckel sn. Obst 1901): [=probably ne ANG sn. 15 aemilianum (Dohrn, 1881) Anthia s AF: type locality not stated, Chaudoir, 1876]; DOC: Kasai- [“Zambezia,nördlichere Occidental, Katanga Provinces; Grenze der Holub‘schen n ANG: Lunda Norte Prov. ssp. latepunctatum [Basilewsky in litt.] “Bihé“ [=c ANG: Bié Prov.; kleinfeld & Puchner, 2012 ZAM: West [Prov.] Liamana

282 283 env.; 1092m; 15°13‘34“S, =madibirensis Sternberg, 1907d Anthia “Madibira (Uhehe: Deutsch 23°21‘71“E; Huambo, Moxico, Ostafrika)“ [=sw TAN: Mbeya Lunda Sul Provinces; ZIM Reg.] ssp. shabanum [Basilewsky in litt.] Kleinfeld & Puchner, 2012 “Lulua Kapanga“ =schenklingi Sternberg, 1907d Anthia “Deutsch Ostafrika (Manow, [=se DOC: Katanga Prov. Uhehe)“ [=sw TAN: w Iringa 18 calvum (Sternberg, 1907d) Anthia e AF: “Région de Tanganjika“ Reg.] [=se DOC: e Katanga Prov.]; =uhehensis Sternberg, 1907d Anthia “Uhehe (Deutsch Ostafrika)“ sw TAN: Ruvuma Reg.; ne [=sw TAN: Iringa Reg.] ZAM 20 massilicatum (Guérin-Méneville, 1845) Anthia se AF: “reg. Massilicatzi“ =pilosa Obst, 1906 Anthia “Kigonsera, Nordufer vom [=s MOS: Gaza, Maputo Nyassa-See“ [=sw TAN: Provinces]; ANG: Huila Ruvuma Reg.] Prov.; BOW, n NAB: Erongo, 19 fornasinii (Bertoloni, 1849a) Anthia e AF: DOC, KEN, MAW, Omaheke, Otjozondjupa MOS, RSA, TAN, ZAM, ZIM Regions, RSA: Free State, ssp. fornasinii s. str. “Inhambane“ [=se MOS: KwaZulu-Natal, Limpopo, Inhambane Prov.]; Maputo, Mpumalanga, North West Tete Provinces; se DOC: Provinces; ZAM, ZIM Katanga Prov.; MAW, e RSA: =natalensis Perroud, 1846 Anthia “ľ intérieur de Natal“ [=se KwaZulu-Natal, Limpopo, RSA: KwaZulu-Natal Prov.] Mpumalanga Provinces; s =massilicatzi Boheman, 1848 Anthia “Caffraria interior“ [=n RSA, TAN: Iringa, Lindi, Mbeya, ZAM, ZIM] Mtwara, Ruvuma Regions =damarensis Kuntzen, 1919 Anthia “Rietfontein-Nord; Herero“ [=e =unicolor Chaudoir, 1861 Anthia “ Delagoa“ [=s MOS: Maputo NAB: Otjozondjupa, Omaheke Prov.: Baía Maputo] Regions] =aequecostata Sternberg, 1907d Anthia “Blantyre (Brit. Ostafrika, =laevithorax Bénard, 1922c Anthia “Angola (district de Huilla), Östlich vom Shire-Fluß)“ [s passage dela Luassingua“ [=sw MAW: Southern Region]; ANG: Huila Prov.] Uhehe (Deutsch-Ostafrika)“ 21 pilosum (Obst, 1906) Anthia e AF: “Kigonsera“ [=sw TAN: [=sw TAN: s Iringa Reg.] Ruvuma Reg.]; MAW; nw ssp. gorge Kolbe, 1894 Anthia “bei Bukoba am Westufer des MOS: Niassa Prov. Victoria-Njansa“ [=nw TAN: =discedens Sternberg, 1907d Anthia “Kandulu (R. Lubjenda, Nord. Kagera Reg.]; Mara Reg. Portug. Ostafrika)“ [=s TAN: ssp. hildebrandti Harold, 1878a Anthia “Sansibar“ [=TAN: Zanzibar Ruvuma Reg.] Isl. Regions]; Dar es Salaam, e 22 principale Sternberg, 1907d Anthia e AF: “Nyassa-See, Deutsch- Iringa, Pwani, Tabora Regions; Ostafrika (Mahenge [=c TAN: s KEN: Coast Prov. Morogoro Reg.]; Manow) [=w =ampliata Chaudoir, 1878 Anthia “Bagamoyo près de Zanzibar“ Iringa Reg.]; Langenburg am [=e TAN: Zanzibar Isl. Nordende des Nyassa“ [Mbeya Regions] Reg.: Tukyu]; Lindi, Ruvuma ssp. stygne Kolbe, 1894 Anthia “Muansa am Südufer des Regions Victoria-Njansa“ [=nw TAN: Mwanza Reg.]; Iringa, Mbeya, Morogoro Regions; e DOC: Sud Kivu Prov.

284 285 nimrod group (Häckel sn. Obst 1901b): ssp. galla s. str. “Région du Nil Blanc“ [=SSD]; 23 decemguttatum (Linnaeus, 1764) Carabus s AF: “Cap. b. Spei.“ [=s BEN, GHA, CAM, s CON, RSA]: Free State, Northern DOC: Bas-Congo, Orientale Cape, Western Cape Provinces; Provinces; ETI: Southern BOW, s NAB: Karas Reg. Nations, Nationalities and =alboguttatus DeGeer, 1778 Carabus “pris du Cap“ [=s RSA: Cape People‘s Region]; NRE, RCA, Prov.] RCI, SEN, SOM: Banaadir =elongatus DeGeer, 1778 Carabus “pris du Cap“ [=s RSA: Cape Reg.; TCH, TOG Prov.] =georgei Ancey, 1885 Anthia “Wydah, Guinée“ [=BEN] =quadriguttatus [Olivier in litt.] Fabricius, 1792 =bella Obst, 1901b Anthia TOG: “Misahöhe“ Carabus “Cap de Bonne-Espérance“ [=s =obsti Sternberg, 1906a Anthia “Ketu (Dahomé)“ [=se BEN] RSA: Cape Prov.] =senegalensis Sternberg, 1906a Anthia SEN =actaeon Fabricius, 1794 Carabus “Cap de Bonne-Espérance“ [=s =somala Müller, 1938 Anthia “Dimé-Bass Narok (Lago RSA: Cape Prov.] Rodolfo) [=s ETI: Southern =alboguttatus Schoenherr, 1778 Carabus “Cap de Bonne-Espérance“ [=s Nations, Nationalities and RSA: Cape Prov.] People‘s Region]; Somalia =elongatus [Olivier in litt.] Schoenherr, 1778 Carabus “Cap de Bonne-Espérance“ italica (Brava, presso [=s RSA: Cape Prov.] Mogadiscio)“ [=s SOM: =laevicollis [Schoenherr in litt.] Dejean, 1825 Anthia “ Cap de Bonne-Espérance“ [=s Banaadir Reg.] RSA: Cape Prov.] =congoana Basilewsky, 1948 Anthia “Oubangui-Chari“ [=RCA] =guttata Lequien, 1832 Anthia “ Cap de Bonne-Espérance“ [=s ssp. lesnei (Sternberg, 1906a) Anthia “Congo français: Haute Sanga, RSA: Cape Prov.] Nebenflufs des unteren Congo“ =villosa [Westermann in litt.] Lequien, 1832 Anthia “ Cap de Bonne-Espérance“ [=s [=w RCA: Mambéré-Kadéï RSA: Cape Prov.] Prefecture]; CAM, n CON =cruoricollis Mannerheim, 1837 Anthia "Caput bonae Spei" [=s RSA: ssp. luluanum Basilewsky, 1948 “Kafakumbu“ [=se DOC: Western Cape Prov.] Katanga Prov.] =knysna Péringuey, 1896 Anthia “ Cape Colony (Knysna)“ [=s ssp. torleyi (Burgeon, 1929) Anthia UGA: Gula; ne DOC: Orientale RSA: Western Cape Prov.] Prov. =stolida Péringuey, 1896 Anthia “Cape Colony (Knysna?, =ertliana Bénard, 1929a Anthia “Ouganda, Afrique Occidentale Oudtshoorn)“ [=s RSA: anglaise“ [=UGA] Western Cape Prov.] 25 hedenborgi (Boheman, 1848) Anthia ne AF: “Sennaar ad Bahr el =vorax Péringuey, 1896 Anthia “Cape Colony (Ladismith)“ [=s Abiad“ [=se SUD: Sennar]; RSA: Western Cape Prov.] ERI, n ETI: Tigray Reg.; SSD =punctulata Sternberg, 1906b “Kapland und Oranje (D.-SW.- 26 hexastictum (Gerstaecker, 1867) Anthia e AF: ne DOC, s ETI, KEN, Afrika)“ [=c, s RSA] TAN, s SOM =oblongoguttata Heyne & Taschenberg, 1908 Anthia “Kap und Natal“ [=s, se RSA] ssp. hexastictum s. str. “ad lacum Jipe“ [Lake =ne =octoguttata Heyne & Taschenberg, 1908 Anthia “Kap und Natal“ [=s, se RSA] TAN: Kilimanjaro Reg.]; =degeeri de Jong, 1944 Anthia “South Africa“ Zanzibar Isl Regions; ne DOC: =nollothiana Basilewsky, 1948 “Namaqualand“ [=sw RSA: Orientale Prov.; KEN: Coast, Northern Cape Prov.] Eastern, Rift Valley Provinces 24 galla (Thomson, 1859b) Anthia c, e, w AF: BEN, CAM, CON, ssp. fortesculptum (Müller, 1938) Anthia “Bacino de Giuba; [=s DOC, ETI, GHA, NRE, RCA, SOM: Gedo Reg.]; tra Dolo; RCI, SEN, SOM, SSU, SUD, Neghelli“ [=s ETI: Oromiya TCH, TOG, UGA

286 287 Reg.]; n KEN: North Eastern =homoplata in error Prov. =mellyi De Brême, 1844 Anthia “ Port Natal“ [=se RSA: ssp. megaera (Lucas, 1881) “Saulalé, intérieur de Brawa KwaZulu-Natal] sn. Fairmaire 1882b Anthia (Afrique australe)“ [=s SOM: =algoensis Péringuey, 1892a Anthia “ Delagoa Bay“ [=s MOS: Shabeellaha Hoose Reg.: n of Maputo Prov.] Baraawe]; Banaadir, Jubbada =cupiens Péringuey, 1892a Anthia “Ovampoland [=n NAB: Dhexe, Jubbada Hoose, Ohangwena, Omusati, Oshana, Shabeellaha Dhexe Regions Oshikoto Regions]; Upper =somaliana Bénard, 1929b Anthia “Somalia italiana, lungo il Limpopo“ [=ne RSA: Limpopo corso deľUebi Scebeli“ [=s Prov.] SOM: Shabeellaha Hoose =heres Péringuey, 1892a Anthia “Northern Ovampoland“ [=n Reg.] NAB: Ohangwena, Oshikoto 27 nimrod [Olivier in litt.] (Fabricius, 1793) Carabus w, c AF: BEN, GAM, GBI, Regions] MLI, MRE, NIG, NRE, RCA, =intempestiva Péringuey, 1892a Anthia “Ovampoland [=n NAB: SEN, TCH, TOG Ohangwena, Omusati, ssp. nimrod s. str. “Senegalia“ [=SEN]; BEN, Oshana, Oshikoto Regions]; GAM, GBI, MLI, MRE, NIG, Damaraland [c NAB: Erongo, NRE, TCH, TOG Kunene, Otjozondjupa =errans Olivier, 1795 Carabus SEN Regions]; Upper Limpopo [=ne ssp. bouvieri (Sternberg, 1906a) Anthia “Congo francais: Haute Kemo, RSA: Limpopo Prov.]; British Zuflufs des mittleren Ubangi“ Bechuanaland“ [=Nortehrn [=sw RCA: Kémo Prefecture] Cape, North West Provinces] =lemoulti Kuntzen, 1913a Anthia “Fort Sibut im =vagans Péringuey, 1892a Anthia “Ovampoland“ [=n NAB: Nordscharitschadgebiet“ [=s Ohangwena, Omusati, Oshana, RCA: Kémo Pref.] Oshikoto Regions] 28 omoplatum [Dupont in litt.] (Lequien, 1832) Anthia s AF: ANG, BOW, MOS, =modesta Obst, 1901 Anthia “ Ngami-See“ [=nw BOW: NAB, RSA, SWA, TAN, ZAM, Ngami Lake] ZIM =grandis Sternberg, 1906b Anthia “D.-Südwest-Afrika: ssp. omoplatum s. str. “Cap de Bonne-Espérance“ Okahandja, Windhuk“ [=c [=RSA]: Free State, Gauteng, NAB: Khomas, Okahandja KwaZulu-Natal, Limpopo, Regions] Northern Cape, North West, ssp. incolatum (Péringuey, 1892a) Anthia “Delagoa Bay“ [=s MOS: Western Cape Provinces; s Maputo Prov.]; e RSA: ANG: Cunene Prov., BOW, KwaZulu-Natal, Mpumalanga s MOS: Gaza, Inhambane, Provinces; SWA Maputo Provinces; NAB: ssp. parvum (Obst, 1901b) Anthia “Mikindani (Deutsch- Caprivi, Erongo, Khomas, Ostafrika)“ [=se TAN: Mtwara Kunene, Ohangwena, Reg.]; Lindi, Ruvuma Regions; Okahandja, Okavango, MAW, n MOS: Cabo Delgado, Omaheke, Omusati, Oshana, Manica, Niassa Provinces Oshikoto, Otjozondjupa 29 revoili (Lucas, 1881) sn. Fairmaire 1882b Anthia e AF: “nord du pays des Regions; ZAM, ZIM Somalis: montagnes des Gonzalis-Ouarsanguèles,

288 289 vallée du Darror et Karkar“ Cape Provinces; s ANG: [=e SOM: Bari Reg: Buuraha Cuando Cubango, Cunene Kar Kar Mts.: Darror Valley]; Provinces; BOW, MOS: Bay, Mudug Regions; se ETI: Inhambane, Maputo, Sofala Somali Reg. Provinces; NAB: Caprivi, =schatzmayri Straneo, 1942 Anthia “El Bur, Somalia“ [=c SOM: Erongo, Okavango, Omaheke, Mudug Reg.] Otjozondjupa Regions; TAN: 30 duodecimguttatum (Bonnelli, 1813) Anthia “Arabie“ [=ne AF: EGY; sw Dar Es Saalam, Dodoma, AS: IRN, IRQ, JOR, KAT, Lindi, Mtwara Regions; ZAM, KUW, OMA, SAU, UAE, ZIM YEM] =minima Bertoloni, 1857 Anthia “in ripis fluminis Manárra 31 venator (Fabricius, 1792) Carabus n, w AF: “Senegalia“ [=GAM, provinciae Inhambanensis MRI, SEN]; ALG, EGY, LIB, Mozambici“ [=se MOS: MOR, NIG, NRE, SUD, TCH, Inhambane Prov.] TUN, WSA; sw AS: SYR =baucis Dohrn, 1881 Anthia “ Holub‘schen Ausbeute: =cursor Olivier, 1795 Carabus SEN Zambese“ [=BOW, ZIM] =curstrix Jacobson, 1906 in error Anthia =sansibarica Kolbe, 1898 “Daressaalam“ [=e TAN: =venatrix Jacobson, 1906 in error Anthia Dar Es Saalam Reg.]; Lindi, Mtwara Regions sulcatum species (Häckel sn. Obst 1901b) =perturbatus Basilewsky, 1980 “Maputa; Delagoa Bay; 32 sulcatum [Olivier in litt.](Fabricius, 1792) Carabus w AF: BFO, GBI, MLI, MRI, Laurenço Marques; Rikatla RCA, SEN, SSD, TOG [=s MOS: Maputo Prov.]; ssp. sulcatum s. str. “Senegalia“ [=SEN]; GBI, w Zululand; Zulu; Natal; MLI, MRI Hulukluwe; Ndumu [=s RSA: =angustata Dejean, 1831 Anthia SEN KwaZulu-Natal Prov.] ssp. adelphum (Thomson, 1859b) Anthia “Région du Nil Blanc“ [=SSD]; BFO, CAM, GHA, s MLI, CYPHOLOBA Chaudoir, 1850; T: Anthia alveolata De Brême, 1844 NRE, RCA, TCH, TOG (species: 57) =simplex Obst, 1901b Anthia TOG: “Hinterland“ =Thermophila Bertoloni, 1849a; T: Thermophila ranzanii Bertoloni, 1849a =variabilis Obst, 1901b Anthia TOG: “Hinterland“ =Polyhirma Chaudoir, 1850; T: Carabus macilentus Olivier, 1795 =voltae Basilewsky, 1946 “Haute Volta: Pundu“ [=BFO] =Diabatus Gistel, 1857; T: Carabus macilentus Olivier, 1795 et Anthia gracilis =notata Basilewsky, 1946 U.N. “Haute Volta: Pundu“ [=BFO] Dejean, 1831

CYPHOLOBINAStrohmayer, 1928 (genera: 5 species: 70) alstoni species (Häckel sn. Basilewsky in Kleinfeld & Puchner 2012) 1 alstoni Péringuey, 1892a s AF: “British Bechuanaland“ ATRACTONOTUSPerroud, 1846; T: Atractonotus mulsantii Perroud, 1846 [=n RSA: n Northern Cape, (species: 1) sw North West Provinces]; =Atractonota Lacordaire, 1854 in error BOW; n NAB: Ohangwena, Omaheke, Omusati, Oshana, 1 mulsantii Perroud, 1846 s AF: “ľ intérieur de Natal“ Oshikoto Regions [=RSA: n KwaZulu-Natal]; Free State, Limpopo, Mpumalanga, Western

290 291 caillaudi group (Häckel sn. Basilewsky in Kleinfeld & Puchner 2012) =transversefasciata Müller, 1938 “tra Bohotle e Berbera, 2 alveolata (De Brême, 1844) Anthia s AF: BOW, MAW, s MOS, e Somalia inglese“ [=n SOM: RSA, se ZAM, ZIM Woqooyi Galbeed Reg.] ssp. alveolata s. str. “Port Natal“ [=RSA: KwaZulu- 5 bihamata (Gerstaecker, 1867) Polyhirma e AF: sw KEN, n TAN, UGA Natal]; Eastern Cape, Gauteng, ssp. bihamata s. str. “ad lacum Jipe“ [=ne TAN: KwaZulu-Natal, Limpopo, Kilimanjaro Reg.]; Dar Mpumalanga, North West Es Saalam, Pwani, Tanga, Provinces; e BOW, se ZAM, Zanzibar Isl. Regions; s KEN: ZIM s Coast, s Eastern Provinces; e =consobrina Péringuey, 1896 Polyhirma “Transvaal (Middelburg)“ [=ne UGA RSA: Mpumalanga Prov.] =lesnei Sternberg, 1907c Polyhirma “Luitpold Kette (Afrique ssp. plantii (Chaudoir, 1861) Polyhirma s MOS: “baie de Delagoa“ orientale allemande)“ [=s KEN: [=Maputo Prov.: Baía Maputo]; s Rift Valley Prov.] MAW, ne RSA: n Kwa-Zulu =silene Strohmeyer, 1928 “Ouganda, parages du Lac Natal, Mpumalanga Provinces Rudolphe“ [=e UGA] =algoensis Péringuey, 1896 Polyhirma “Transvaal (Barberton)“ [=ne ssp. grotei Kuntzen, 1913b “Eldamariverstation“ [=w RSA: Mpumalanga Prov.]; KEN: Rift Valley Prov.: Mozambique (Laurenco- Eldama Ravine]; w Eastern Marquez, Rikatla) [=s MOS: Prov. Maputo Prov.] =albini Bénard, 1921b Polyhirma “Afrique Orientale: Kedong ssp. ranzanii (Bertoloni, 1849a) Thermophila “in provincia Inhambanensi Valley“ [=w KEN: Rift Valley Mozambici“ [MOS: Prov.] Inhambane Prov.]; Manica =kedongiana Bénard, 1922b Polyhirma “Afrique Orientale anglaise: Prov.; ZIM Kedong Valley“ [=w KEN: =minima Bertoloni, 1857 Anthia “in ripis fluminis Manárra Rift Valley Prov.] provinciae Inhambanensis =loitana Strohmeyer, 1928 “Ostafrika, nördl. Massailand, Mozambici“ [=se MOS: Loitakette“ [=sw KEN: Rift Inhambane Prov.] Valley Prov.: Loita Hills] =dohrni [sn. Chaudoir 1850] Sternberg, 1907b Polyhirma 6 biloba (Fairmaire, 1882b) Polyhirma e AF: “pays Somalis, 3 amatonga Péringuey, 1892b se AF: “Amatongaland [=se Oursangueli“ [=s SOM: Bay RSA: n KwaZulu-Natal: Reg.: Uar Tugar] Tongaland]; Delagoa Bay“ [=s 7 bouvieri (Sternberg, 1907a) Polyhirma e AF: “Envir. de Harar, MOS: Maputo Prov.: Baía de vallée de Herer: Bou Sidimo“ Maputo]; ne RSA: Gauteng, [=e ETI: Harari Reg.]; Limpopo Provinces; ZIM Oromiya, Southern Nations, =babyana Strohmeyer, 1928 “Simbabye“ [=n ZIM: Harare, Nationalities, and People‘s Mashonaland East Provinces Regions 4 apicata (Fairmaire, 1887) Polyhirma e AF: “Sômalis“ [=e ETI: =wayuana Strohmeyer, 1928 “Nordost-Afrika, Nordgalla, Somali Reg.; SOM: Banaadir, westl. von Suaisee“ [=sw =piaggiae Gestro, 1881 Polyhirma (partim) Mudug; Shabeellaha Dhexe, ETI: Southern Nations, sn. Müller 1938 Shabeellaha Hoose, Woqooyi Nationalities, and People‘s Galbeed Regions] Reg.]

292 293 8 caillaudi (Laporte de Castelnau, 1835) Anthia e, s AF: DJI, ERI, ETI, KEN, Reg.]; Ubena-Langenburg, MOS, NAB, SOM, SSD, SUD, nördlich vom Nyassasee TAN, ZAM, ZIM [=Mbeya Reg.]; Mpapwa“ ssp. caillaudi s. str. “Sennaar“ [=se SUD: Sennar]; [=Dodoma Reg.: Mpwapwa] DJI, ERI, ETI: Harari, =massaica Kuntzen, 1913b Polyhirma “Massaigebiet: Käsebog; Oromiya, Somali, Tigray Werther“ [c, n TAN: Arusha, Regions; n KEN, SOM: Dodoma , Morogoro Regions] Mudug, Woqooyi Galbeed =ukerewensis Kuntzen, 1913b Polyhirma “Insel Ukerewe des Regions; SSD Viktoriasees“ [nw TAN: Mara =polioloma Chaudoir, 1848 Anthia type locality not stated, [“côtes Reg.] orientales de ľAfrique“ =KEN, =usagarensis Kuntzen, 1913b Polyhirma “Usagara“ [=n TAN: Mwanza TAN sn. Chaudoir 1848] Reg.] =roeschkei Sternberg, 1907b Polyhirma “Abyssinie (Colonie Erytrée)“ =punctata Bénard, 1921c “Afrique Orientale (Lemek [=ERI] Valley, Nyanza Province et =dohrni Sternberg, 1908 Polyhirma “Harar; [=ETI: Harari Reg.]; Amala River, Sotik)“ [=sw JH of Sternberg 1907b Dadal [=n Oromiya Reg.: KEN: Nyanza, Rift Valley Dada]; Adi; Halin“ [=Tigray Provinces] Reg.] =glauningi Strohmeyer, 1928 “D.-O.-Afr., Ngorongoro- =harrarensis Strohmeyer, 1928 Ejassisee“ [=ne TAN: Arusha nn for Polyhirma dohrni Sternberg, 1908 Reg.] =erythreensis Bénard, 1924 Polyhirma “ľErythrée italienne“ [=ERI] =guruana Strohmeyer, 1928 “D.-O.-Afr., westlich und südl. =dilagala Basilewsky, 1980 “Berbera; Somalie. British v. Gurui-Vulkan [=c TAN: Somaliland; Hargeisa [=n Morogoro Reg.]; Iramba- SOM: Woqooyi Galbeed Reg.]; Issansu, Mkalane“ [=Singida Mudugh, 5 km nw of Marcegur Reg.] [=Mudug Reg.]; Ethiopie =holtzi Strohmeyer, 1928 “Mtukulla“ [=s TAN: Ruvuma (Ogaden). Galadi; 3 mls n of Reg.] Shahade“ [=se ETI: Somali =neumanniana Strohmeyer, 1928 “Ost-Victoria-See, Ngoroine- Reg.] Mukenge“ [=n TAN: Arusha, ssp. biscutellata (Fairmaire, 1887) Polyhirma cw TAN: Tabora [Reg.]; Mara Regions] Arusha, Dodoma, Iringa, Mara, =saramica Strohmeyer, 1928 “Daressalam“ [=e TAN: Dar Es Mbeya, Morogoro, Mwanza, Saalam Reg.] Rukwa, Ruvuma, Singida =zenckei Strohmeyer, 1928 “D.-O.-Afr., Bez. Tanga Regions; s KEN: Rift Bismarckburg“ [=sw TAN: Valley Prov. Rukwa Reg.: Kasanga District] =kolbei Sternebrg1907b Polyhirma “Deutsch Ost-Arika (Mpuapua, ssp. boucardi (Chaudoir, 1883) Polyhirma “Transvaal“ [=ne RSA: Ugogo)“ [=c TAN: Dodoma Mpumalanga, Limpopo, North Reg.] West Provinces]; MOS: Gaza, =clausi Kuntzen, 1913b Polyhirma “Kilimatinde (Deutsch Inhambane, Manica, Maputo, Ostafrika)“ [=c TAN: Singida Sofala, Tete Provinces; ne Reg.] NAB: Caprivi Reg.; e TAN: =goetzei Kuntzen, 1913b Polyhirma “Ugogo [=c TAN: Dodoma Lindi, Mbeya, Mtwara Reg.]; Uhehe-Iringa [=Iringa Regions; ZAM, ZIM

294 295 =septemcostata Dohrn, 1883 Anthia “ Holub‘schen Ausbeute: ssp. seruana Strohmeyer, 1928 “Süd-Afr., Ost-Bechuanaland; Zambese“ [=MOS, ZAM, ZIM] Sogosse; Serue“ [=e BOW: =littoralis Kuntzen, 1913b Polyhirma “ Chifumbasi (Mozambique)“ Central Distr.]; c RSA: Eastern [=nw MOS: Tete Prov.] Cape, Free State, North West =clathratipennis Bénard, 1923 Polyhirma “Lindi“ [Reg. =se TAN] Provinces; ZIM =ledouxi Strohmeyer, 1928 “Süd-Afrika, Geluque Hills“ [= 12 prolixa (Fairmaire, 1891b) Polyhirma e AF: e ETI, SOM, e KEN RSA] ssp. prolixa s. str. “ľAfrique orientale vers =rhodesiana Strohmeyer, 1928 “Südwest-Rhodesia“ [=sw premier degré de latitude nord“ ZIM] [=e KEN: North Eastern Prov.; =seineriana Strohmeyer, 1928 “Südafrika, Livingstone s SOM: Shabeellaha Hoose [=s ZAM: Southern Prov.: Reg.]; Jubbada Dhexe, Jubbada Maramba]; Ost-Sambesigebiet Hoose Regions [=se ZAM]; Sumpfgeb. von ssp. somalica (Gestro, 1892) Polyhirma “Uorandi; Uebi“ [=e ETI: Maschi, Sambala-Ranga“ [=ne Somali Reg.: Uebi Scebeli NAB: Caprivi Reg.] =(Wabi) Shebelle River]; c =teteana Strohmeyer, 1928 “Port. O.-Afrika, Tete“ [=w SOM: Mudug, Nugaal Regions MOS: Tete Prov.] =tiesleri Strohmeyer, 1928 “Port. Ost-Afrika, Chifumbasi“ 13 trilineata Strohmeyer, 1928 sw AF: ec ANG: Bailundo [=w MOS: Tete Prov.] [=Huambo Prov]. ssp. bozasi (Sternberg, 1907a) Polyhirma “Pays Tourkouana, á ľouest du lac Rodolphe, dans ľAfrique graphipteroides group (Häckel sn. Basilewsky in Kleinfeld & Puchner 2012) orientale anglaise“ [=nw 14 cardiodera (Fairmaire, 1887) Polyhirma e AF: “Makdischu“ [=se SOM: KEN: n Rift Valley Prov.: w of Banaadir Reg.] Turkana Lake]; s ETI: Southern 15 chanleri (Linell, 1896) Polyhirma e AF: “Tana River“ [=c, Nations, Nationalities and e KEN: Coast, Eastern People's Reg. Provinces]; se ETI: Somali =neumanni Sternberg, 1908 Polyhirma “Djala; Uba“ [=s ETI: Southern Reg.; sw SOM: Gedo, Jubbada Nations, Nationalities and Dhexe, Jubbada Hoose People's Reg.] Regions; ne TAN: Pwani Reg. =mirei Basilewsky, 1963 “Mare ďOrtobi“ [=SSD] =tetracha Bénard, 1926 Polyhirma “bassin du fleuve Juba; Ikuta, ssp. livingstoniana Basilewsky, 1983 s ZAM: 24 miles n of Afrique orientale“ [=sw SOM: Livingstone [=Southern Prov.: Gedo, Jubbada Dhexe, Jubbada Maramba] Hoose Regions] 9 griseostriata (Fairmaire, 1884) Polyhirma e AF: “Makdischu“ [=se SOM: =chauleri Csiki, 1929 in error Banaadir Reg.]; Shabeellaha 16 divisa (Boheman, 1860) Polyhirma s, sw AF: “prope lacum Hoose Reg. N´gami“ [=nw BOW: 10 grisescens (Fairmaire, 1884) Polyhirma e AF: “Makdischu“ [=se SOM: Ngami Lake env.]; s ANG: Banaadir Reg.]; Mudugh Reg. Cuando-Cubango Prov.; n =serifoveata Fairmaire, 1887 Polyhirma “Sômalis“ [=SOM] NAB: Caprivi, Ohangwena, 11 oberthueri (Sternberg, 1907b) Polyhirma s AF: BOW, RSA, ZIM Okavango, Omusati, Oshana, ssp. oberthueri s. str. “Motoppo Hills (Rhodesia)“ Oshikoto, Regions; s ZAM, [=s ZIM: Bulawayo Prov.: ZIM Matobo Hills]; ne RSA: =circumcincta Chaudoir, 1861 Polyhirma “ lac N´gami“ [=nw BOW: Limpopo Prov. Ngami Lake]

296 297 =amboensis Péringuey, 1896 Polyhirma type locality not stated, ssp. intrusa Basilewsky, 1980 c TAN: Kilossa [=Morogoro [“Amboland“ sn. Péringuey Reg.]; Tabora [Reg.]; Dodoma 1926 (=n NAB: NAB: [Reg.]; Mamboia [=Ruvuma Ohangwena, Omusati, Oshana, Reg.; nw MOS: Niassa Prov.] Oshikoto Regions)] ssp. kigonserana Strohmeyer, 1928 “Kigonsera“ [=s TAN: Ruvuma =circumscripta Roussseau, 1906 Polyhirma in error Reg.] =elegans Bénard, 1922c Polyhirma “Angola: (Luiana, Maniomba, ssp. longefasciata Basilewsky, 1980 c MOS: “Gorongoza; Prov. de Luankunda)“ [=se ANG: Gorongoza, Tendos du Songué Cuango-Cubango Prov.] et riv. Vunduzi; Inhacoro, env. 17 graphipteroides (Guérin-Méneville, 1845) e, s, se AF: ANG, BOW, DOC, de Chemba; Nova Choupanga, ERI, ETI, KEN, MAW, MOS, près de Chemba [=Sofala NAB, RSA, SSD, SWA, ZAM, Prov.]; Chimoio; Vila Pery; ZIM Revoué, Zemba; Buzi, sur le ssp. graphipteroides s. str. “reg. Massilicatzi“ [=se MOS: Revoué“ [=Manica Prov.] Gaza, Maputo Provinces]; ssp. lukuledicola Strohmeyer, 1928 “D.-O.-Afr., Massassi; Lindi- BOW, RSA: Eastern Cape, Massassi; Namupa [=s TAN: Gauteng, KwaZulu-Natal, Lindi Reg.]; Lukuledi; Ndanda Limpopo, Mpumalanga [=Mtwara Reg.]; Wandonde- Provinces; SWA, ZAM, ZIM Gebiet, am Rovuma [=Ruvuma =suturata Perroud, 1846 Anthia “ ľ intérieur de Natal“ [=se Reg.]; MAW, nw MOS: Niassa RSA: KwaZulu-Natal Prov.] Prov. ssp. alternata (Raffray, 1885) Polyhirma “plateaux inférieurs des ssp. methneri Strohmeyer, 1928 “Trockenwald bei Mtotohovu; Bogos“ [=n ETI: Tigray Reg.]; Unterer Umba“ [=ne TAN: ERI, e KEN: Coast, Eastern Tanga Reg.]; se KEN: Coast Provinces; SSD Prov. ssp. bembana Strohmeyer, 1928 “Zentralafr., Ubemba“ [=se ssp. monardi Burgeon, 1935b Osi [=w ANG: ]; Caluquembe, DOC: n Katanga Prov.: Cassinga, Mucoti, Sangueve, Ubembe]; BOW, ZAM Cubango Provinces ssp. bilunata (Boheman, 1860) Polyhirma “juxta lacum N´gami“ [=nw ssp. nyika Strohmeyer, 1928 “Nordnjassasee, Unyika“ [=sw BOW: Ngami Lake env.]; TAN: s Mbeya Reg.]; se DOC: MAW, w MOS: Sofala, Tete, Katanga Prov. Zambezia Provinces; ZAM, ssp. oskari Strohmeyer, 1928 “Ostvictoriasee, Ngoroine- ZIM Mukenge“ [=n TAN: Arusha, =tiesleriana Strohmeyer, 1928 “Port. Ostafr., mittl. Sambesi“ Mara Regions]; s KEN: Rift [=w MOS: Sofala, Tete Valley Prov.; UGA Provinces] ssp. rukwana Strohmeyer, 1928 “Südost-Tanganjikasee, =tschifumbasica Strohmeyer, 1928 “Port. Ostafrika, Chifumbasi“ Rukwa-See-Steppe“ [=sw [=nw MOS: Tete Prov.] TAN: Rukwa Reg.] ssp. hehana Strohmeyer, 1928 “D.-O.-Afr., Kidugalla; Fuß ssp. udschungwana Strohmeyer, 1928 “D.-O.-Afr., Iringa-Mgololo; des Mageberges; Uhehe- Udzungwa-Gebirge“; 1450m; Iringa; Tosamagonga; Iringa; [=c, e TAN: Iringa, Morogoro Mitteluhehe“; 1500-1700m Regions]; Dar Es Saalam, [=wc TAN: Iringa Reg.] Lindi, Pwani, Tanga Regions

298 299 =fischeri Strohmeyer, 1928 “Daressalam Hinterland, ssp. neavei Burgeon, 1929 “Rhodésie du Nord, Muneka; Puguberge und Kisserewe; Kapema à Kipaila [=Njoni Mittel-Rufidji [=ne TAN: Dar Kapema]; Katumba; Mbiliwa Es Saalam, Pwani Regions]; à Wantu [=c ZAM: Northern Daressalem, Pangani and Prov.]; Katanga [Prov. =se Hinterland; Tanga [=Tanga DOC]: Kambove; Mufungwa- Reg.]; Kilwa [=Lindi Reg.]; Ambwe; Kundelungu; Morogoro; Uluguru [=c, e Elisabethville et Kapiri; TAN: Morogoro Reg.] Kiambi; Lomami =stuhlmanni Strohmeyer, 1928 “Vitsonha?; D.-O.-Afr.; ssp. pluricostata (Fairmaire, 1882a) Polyhirma “Zanguebar“ [=BUR, wc Daressalam“ [=ne TAN: Dar TAN]: Iringa, Mbeya, Es Saalam, Pwani Regions] Morogoro, Rukwa, Singida, ssp. ufiomina Strohmeyer, 1928 “D.-O.-Afr., Ufiomi“ [=nc Tabora Regions TAN: n Dodoma, s Manyara =kilimatindica Strohmeyer, 1928 “Kilimatinde [=c TAN: Regions: Tarangire National Singida Reg.]; Nguruberge“ Park] [=Morogoro Reg.] 18 grimaudi (Bénard, 1922a) Polyhirma s AF: “Haut Bassin de =niggmanniana Strohmeyer, 1928 “D.-O.-Afr., Uhehe Udzungwa- Zambèze: Loenginga“ Dabaga; Uhehe Iringa und [=w ZAM: Western Prov.: Umgeb.; Tosamagonga [=c, sw Luanguinga]; e ANG: Moxico TAN: Iringa Reg.]; Madibira“ Prov. [=Mbeya Reg.] 19 kavanaughi Basilewsky, 1983 sw AF: c, s ANG: “Cuche, =uhligiana Strohmeyer, 1928 “D.-O.-Afr., Massai Ngotiek“ within 100 km radius (North)“ [=c, ne TAN: Arusha Reg.] [=Cuando-Cubango Prov: n of 22 semibrunnea Strohmeyer, 1928 e AF: “D.-O.-Afr., Mikindani; Cuchi]; Huambo Prov. Lindi; Makonde-Hochland, 20 leucospilota (Bertoloni, 1849a) Thermophila se AF: s MOS, e RSA Ntschitschira und Nanjamba; ssp. leucospilota s. str. “Inhambanensis“ [=se MOS: Namupa; Donde; Barikiwa; Inhambane Prov.]; Gaza Prov. Mittelrowuma“ [=se TAN: ssp. semilaevis (Chaudoir, 1861) Polyhirma “baie de Delagoa“ [=s MOS: Lindi, Mtwara, Ruvuma Maputo Prov.: Baía de Regions]; n MOS: Cabo Maputo]; se RSA: KwaZulu- Delgado Prov. Natal Prov. 23 somereni (Bénard, 1930) Polyhirma e AF: “Afrique orientale 21 perspicillaris (Chaudoir, 1878) Polyhirma e, s AF: BUR, DOC, KEN, anglaise: Kiu“ [=c KEN: Rift TAN, UGA, ZAM Valley Prov.]; Eastern Prov.; ssp. perspicillaris s. str. “montagnes du Schimba“ [=e ETI: Oromiya Reg.; ne TAN: TAN: Pwani Reg.: Shimba Arusha Reg.; UGA Hills]; Arusha, Dar Es Saalam, =linelli Strohmeyer, 1928 “Brit.-Betschuanaland“ Tanga, Zanzibar Isl. Regions; probably error KEN: Coast, Eastern Provinces 24 tetrastigma (Chaudoir, 1848) Anthia e AF: DJI, ERI, ETI, KEN, ssp. luluensis Burgeon, 1935a “Congo belge: Kafakumba; SOM, TAN, TCH, NIG, UGA Kapanga; Sandoa; Kayoyo“ ssp. tetrastigma s. str. “côtes orientales de ľAfrique“ [=se DOC: Katanga Prov.] [=e KEN: Coast, Eastern, North Eastern Provinces; ne TAN:

300 301 Kilimanjaro Reg.]; n ETI: Brit. O.-Afr., Eldama River Dire Dawa, Harari, Oromiya, Station [=s KEN: Rift Valley Somali, Southern Nations, Prov.]; Escarpment; 6500- Nationalities and People´s, 9000ft Tigray Regions =wahehophila Strohmeyer, 1928 “D.-O.-Afr., Uhehe-Iringa =gallinieri Reiche, 1847 Anthia “province du Tigré“ [=n ETI: [=c, sw TAN: Iringa Reg.]; Tigray Reg.] Madibira; Uhehe-Ubena; Tosa“ =dimidiata Roth, 1851 Anthia “Tigré in N. Abyssinien“ [=n [=Iringa, Mbeya Regions] ETI: Tigray Reg.] ssp. amala Basilewsky, 1980 sw KEN: “Amala River, Sotik; =quadriplagiata Gerstaecker, 1867 Polyhirma “ad lacum Jipe“ [=ne TAN: Guaso Nyiro (sud) Sotik; Kilimanjaro Reg.] Lomek Valley, Nyanza Prov.; =wituana Strohmeyer, 1928 “Witu-Tanafluß; Witu- Narossera River, Loita Plains Pocomonie“ [=e KEN: Coast [=s Rift Valley Prov.]; n. w. Prov.] Tanzanie, Utegi“ [=n TAN: ssp. conjuncta Mateu, 1966 e, ne TCH: Ennedi Reg.: “la Mara Reg.] mare de Ullmann; Mare de ssp. smiliophora (Fairmaire, 1887) Polyhirma w TAN: Tabora [Reg.]; Kagera, Damas; Mare de Bodoué; Manyara, Mara, Rukwa Mare d‘Orromé; Ouadai, Iriba Regions; BUR [Ouaddai Reg.]; se NIG: rég. =eggeliana Strohmeyer, 1928 “Viktoria-Njansa, Bukoba“ de Maine Soroa“ [Diffa Reg.] [=nw TAN: Kagera Reg.] ssp. subfasciata Müller, 1938 “Giumbo alla face del Giuba, =insulana Strohmeyer, 1928 “Viktoria-Njansa, Insel Somalia Italiana“ [=s SOM: Ukerewe“ [=n TAN: Mara Jubbada Hoose Reg.]; Banaadir Reg.] Reg. =mangatiana Strohmeyer, 1928 “Mangati, norwestl. von 25 trilunata (Gestaecker, 1884) Polyhirma e AF: BUR, s KEN, ne TAN Irangi“ [=n TAN: Manyara ssp. trilunata s. str. “Kilimandjaro“ [=ne TAN: Reg.] Kilimanjaro Reg.]; Arusha, =pontina Strohmeyer, 1928 “Tanganjika, Karema“ [=sw Dodoma, Iringa, Manyara, TAN: Rukwa Reg.] Mbeya Regions; s KEN: =trefurthiana Strohmeyer, 1928 “D.-O.-Afr.“ [=TAN] Eastern, Rift Valley Provinces =fossicola Strohmeyer, 1928 “D.-O.-Afr., Hohenlohegraben“ intermedia group (Häckel sn. Basilewsky in Kleinfeld & Puchner 2012) [=ne TAN: Manyara Reg.: 26 bottegoi Müller, 1938 e AF: “Somalia anglica: Yaida Swamp] Lafarugh-Aberi [=se ETI: =jaegeri Strohmeyer, 1928 “D.-O.-Afr., Iraku-Ejassisee“ Somali Reg.]; Abyssinia: [=ne TAN: Arusha Reg.: Eyasi Arussi-Galla; Kaka; Scioa“ [=s Lake] ETI: Oromiya Reg.]; Tigray =sagarica Strohmeyer, 1928 “O-Asagara“ [=ne TAN: Reg.; DJI, n SOM: Togdheer, Dodoma Reg.: Sagara] Woqooyi Galbeed Regions =vagans Strohmeyer, 1928 “D.-O.-Afr., Massai-Ngaruka =piaggiae Gestro, 1881 Polyhirma (partim) “Scoperta a Tull-Harré, fra i [=Engaruka]; Ngotiek; Somali Isa, a Kakak de nelle Massaisteppe; Kitumbeni- sn. Müller, 1938 vicinanze del lago di Caraba“ Vulkan; 1500m [=Kitumbeine [=n SOM: Togdheer Reg.: Ali Mt. =ne TAN: Arusha Reg.]; Isa]

302 303 27 cinereocincta (Fairmaire, 1887) Polyhrima e AF: e ETI, KEN, s SOM =schroederi Strohmeyer, 1928 “Kihuiro; Kwakijembe“ [=ne ssp. cinereocincta s. str. “Ouébi“ [=e ETI: Somali Reg.: TAN: Kilimanjaro Reg.] Uebi Scebeli =(Wabi) Shebelle =wituensis Strohmeyer, 1928 “Witu-Kipini; Station River]; se SOM: Shabeellaha Congwanamanashoh bei Dhexe; Shabellaha Hoose Kipini“ [=e KEN: Coast Prov.] Regions ssp. djubana Strohmeyer, 1928 “westl. des Ganale Djuba“ [=s =subparallela Fairmaire, 1887 Polyhirma “Ouébi“ [=e ETI: Somali Reg.: ETI: Somali Reg., s SOM: Uebi Scebeli =(Wabi) Shebelle Gedo Reg.] River] ssp. saganicola Strohmeyer, 1928 “Nordost-Afrika, Nordgalla =denekana Strohmeyer, 1928 “Denek“ [=se ETI: Somali Saganfluß bei Dembole“ Reg.; se SOM: Shabeellaha [=s ETI: Southern Nations, Dhexe Reg.] Nationalities and People´s ssp. alluaudi Sternberg, 1907b “Tavéta (sö. von Kilima-Njaro, Reg.] Brit. O. Afr.)“ [=ne KEN: Coast ssp. sambara Strohmeyer, 1928 “D.-O.-Afr., Forhani-Ukami Prov.]; ne TAN: Kilimanjaro [=c TAN: Iringa Reg.]; Reg. Pangani; Tanga [Reg.]; ssp. jubae Basilewsky, 1980 “Somalie (Basso et Altro Daressalem“ [=Dar Es Saalam Giubba). Barbera“ [=n SOM: Reg.] Woqooyi Galbeed Reg.]; ssp. semidorsata (Fairmaire, 1893) Polyhirma “Sômalis“ [=s ETI: Oromiya, Alessandra [=Jubbada Dhexe Somali Regions; s SOM: Reg.]; Gedo, Jubbada Hoose Jubbada Hoose Reg.] Regions; KEN: Jubaland; Lagh 30 intermedia (Boheman, 1848) Anthia ne AF: “Sennaar ad Bahr el Bogol“ [=North-Eastern Prov.] Abiad“ [=se SUD: Sennar]; ssp. rothschildi (Sternberg, 1907c) Polyhirma “Afrique orientale anglaise, ERI, e ETI: Afar, Dire Dawa, au sud du lac Rodolphe“ [=nw Harari, Oromiya, Somali, KEN: nc Rift Valley Prov.: Tigray Regions; e TAN: s of Turkana Lake]; s ETI: Morogoro Reg. Oromiya, Southern Nations, =ferreti Reiche, 1847 “province de Tigré“ [=n ETI: Nationalities, and People‘s Tigray Reg.] Regions =leucomelana Roth, 1851 “Tigré in N. Abyssinien“ [=n 28 dartevellei Basilewsky, 1967 sw AF: sw ANG: “Senhora del ETI: Tigray Reg.] Monte Sâ de Bandeira“ [=Huila =dauana Strohmeyer, 1928 “Südgalla, bei Dolo am Prov.] Endfluß des Daua in den 29 hamifera (Harold, 1881) Polyhirma e AF: s ETI, w KEN, nw TAN, Ganale“ s SOM [=s ETI: Somali Reg., s SOM: ssp. hamifera s. str. w KEN: Taita District [=Rift Gedo Reg.] Valley Prov.]; Ukamba [=nw 31 interrupta (Fairmaire, 1887) Polyhirma e AF: “Sômalis-Sud“ [=s TAN: Shinyanga Reg.]; e KEN: SOM]: Banaadir, Jubbada Coast, Eastern Provinces; ne Hoose, Shabeellaha Dhexe TAN: Arusha, Kilimanjaro Regions Regions =grisescens Fairmaire, 1887 “Makdischu“ [=se SOM: =karoliana Sternberg, 1907c “mont Karoli, pays de Rendilé Banaadir Reg.: Mogadishu] (Afrique orientale anglaise)“ =fallaciosa Müller, 1938 “Somalia italica, apud vicum [=c KEN: Rift Valley Prov.] Duca degli Abruzzi, prope

304 305 Mogadiscio [=se SOM: Dondegebiet; Barikiwa“ Shabeellaha Dhexe Reg.]; [=Lindi Reg.] prope Margherita ad flumen 36 spathulata (Gestaecker, 1867) Polyhirma e AF: “Aruscha“ [=n TAN: Juba“ [=Jubbada Hoose Reg.] Arusha Reg.]; Dar Es Saalam, 32 intricata (Dohrn, 1882) Anthia e, se AF: MAW, MOS, TAN, Iringa, Kilimanjaro, Lindi, ZAM Mtwara Regions; e KEN: Coast ssp. intricata s. str. “Montes Nguru, Sansibar“ Prov. [=c TAN: Morogoro, Zanzibar =indecorata Sternebrg1907b Polyhirma “Deutsch Ost-Afrika“ [=TAN] Isl. Regions]; Dar Es Saalam, 37 suturella (Chaudoir, 1866) Polyhirma se AF: “Des bords du Singida, Tabora Regions Zambèze“ [=MOS: Zambézia =neonympha Dohrn, 1882 “Montes Nguru, Sansibar“ [=c Prov.]; se KEN: Coast Prov.; TAN: Morogoro, Zanzibar Isl. MAW, TAN: Dodoma Reg. Regions] =suturalis Kolbe, 1898 “Daressaalam“ [=e TAN: Dar macilenta group (Häckel sn. Basilewsky in Kleinfeld & Puchner 2012) Es Saalam Reg.] 38 chaudoiri (Péringuey, 1892b) Polyhirma s AF: “Transvaal (Rustenburg)“ =bergeri Strohmeyer, 1928 “Kilimatinde“ [=c TAN: [=n RSA: North West Prov.]; Singida Reg.] BOW, ZAM, ZIM ssp. gaza (Péringuey, 1896) Polyhirma type locality not given =ellenbergeri Bénard, 1925b Polyhirma “Bechuanaland, Gaberones“ [=s MOS: Gaza Prov. sn. [=se BOW: South-East Prov.: Basilewsky 1980]; Sofala, Tete Gaborone] Provinces; MAW, ZAM 39 edax (Péringuey, 1892) Polyhirma sw AF: “Amboland“ [=n NAB: =marqueza Péringuey, 1896 Polyhirma “ Mozambique (Gazaland)“ [=s Ohangwena, Omusati, Oshana, MOS: Gaza Prov.] Oshikoto Regions]; sw ANG: =forstmanni Strohmeyer, 1928 “Brit. Njassaland, zu Zomba u. Huila Prov. Blantyre“ [=s MAW: Southern 40 elegantula (Fairmaire, 1887) Polyhirma e AF: e DOC, TAN Reg.] ssp. elegantula s. str. “Kipalapala, prés du Lac =sambesica Strohmeyer, 1928 “Port. Ostafr., mittl. Sambesi“ Tanganyika“ [=wc TAN: [=w MOS: Sofala, Tete Tabora Reg.]; Arusha, Dar Provinces] Es Saalam, Dodoma, Iringa, 33 obtusata (Fairmaire, 1887) Polyhirma e AF: “Somâlis“ [=e ETI, Kagera, Mbeya Regions; e SOM]: e ETI: Somali Reg.; c DOC: Nord Kivu Prov. SOM: Bari Reg. =dysmoeca Strohmeyer, 1928 “Victoria-See, Bukoba“ [=n 34 posticalis (Fairmaire, 1887) Polyhirma e AF: SOM TAN: Kagera Reg.] ssp. posticalis s. str. “Sômalis“ [=c, s SOM]: =iringana Strohmeyer, 1928 “D.-O.-Afr., Ubena- Banaadir, Jubbada Hoose Langenburg; Madibira Regions [=Mbeya Reg.]; Daressalam ssp. virgulifer (Sternberg, 1907a) Polyhirma “Pays des Somalis: Ferrad“ [=Dar Es Saalam Reg.]; [=n SOM]; Mudugh, Nugaal Udzungwa-Dabaga; Uhehe, Regions Iringa u. Umgb.; Iringa, 35 schenklingi (Sternberg, 1907b) Polyhirma e AF: “Tabora [Reg.] (D. O. Mogololo; Uhehe, Tosa; Afrika)“ [=wc TAN]; MAW Mpangwe; Mittel Uhehe =amaculata Strohmeyer, 1928 “Njassa [=s TAN: Ruvuma [=Iringa Reg.] Reg.]; Kilwa; Donde; =obsti Strohmeyer, 1928 “Landschaft Irangi, Jkome“ [=e DOC: Nord Kivu Prov.]

306 307 ssp. conradsi (Sternberg, 1907b) Polyhirma “Insel Ukerewe (im sö. =gallemaertsi Rousseau, 1905 Atractonota “ Tschinde“ [=e MOS: Victoria-Nyanza)“ [=nw TAN: Zambézia Prov.: Chinde Distr.] Mwanza Reg.] ssp. seineri Strohmeyer, 1928 “O. Betschuanaland, Sogosse; ssp. heoeca Strohmeyer, 1928 “D.-O.-Afr., Nordwest-Massai, Mague [=e BOW: Central Sossian Ngoroine“ [=ne TAN: Distr.]; Rhodesia, Matoppo Arusha Reg.] Hills“ [=s ZIM: Bulawayo ssp. lukugana Basilewsky, 1980 “Zaïre. Tanganyka-Moëro, Prov.: Matobo Hills]; Nyunzu; bassin de la Lukuga“ NAB: Caprivi, Kavango, [=se DOC: Katanga Prov.] Otjozondjupa Regions; ne ssp. moschiensis Strohmeyer, 1928 “Moschi“ [=ne TAN: RSA: Limpopo, North West Kilimanjaro Reg.] Provinces; ZAM 41 fritschi (Chaudoir, 1883) Polyhirma sw AF: “Kuruman dans ľ =damara Strohmeyer, 1928 “D.-Südw.-Afr., Okahandja“ Afrique australe“ [=n RSA: [=e NAB: Otjozondjupa Reg.] Northern Cape Prov.]; BOW, ssp. zuluana Basilewsky, 1948 “Zululand“ [=se RSA: e NAB: Omaheke Reg.; ZAM, KwaZulu-Natal Prov.] ZIM 43 macilenta (Olivier, 1795) Carabus s AF: “Cap de Bonne- 42 gracilis (Dejean, 1831) Anthia se, sw AF: BOW, LES, MOS, Espérance“ [=s RSA]: Eastern NAB, RSA, ZAM, ZIM Cape, Free State, Gauteng, ssp. gracilis s. str. “Cap de Bonne-Espérance“ [=s KwaZulu-Natal, Limpopo, RSA]: Eastern Cape, Gauteng, Mpumalanga, Northern Cape, Free State, KwaZulu-Natal, North West, Western Cape Limpopo, Mpumalanga, Provinces; BOW, LES, s MOS: Northern Cape, North West Maputo Prov.; NAB: Khomas Provinces; s ANG: Cunene, Reg.; ZIM Huila Provinces; BOW; LES, =foveata Perroud, 1846 Anthia “ľ intérieur de Natal“ [=se NAB: Erongo, Ohangwena, RSA: KwaZulu-Natal Prov.]; Omusati, Oshana, Oshikoto, LES, ZIM Otjozondjupa Regions =exarata Boheman, 1848 Anthia “Caffraria“ [=RSA, ZIM] =clathrata Klug, 1885 Anthia “Cap de Bonne-Espérance“ [=s =dregei Guérin-Méneville, 1849 Anthia “Natal“ [=se RSA: KwaZulu- RSA] Natal Prov.] =fossulata Perroud, 1846 Anthia “ ľ intérieur de Natal“ [=se =bennettii Marshall, 1902 Polyhrima “Somerset West, Cape Colony“ RSA: KwaZulu-Natal Prov.] [=s RSA: Western Cape Prov.] =simbabyana Strohmeyer, 1928 “Mashonaland, Simbabye“ [=n 44 notata (Perroud, 1846) Anthia s AF: “ľ intérieur de Natal“ ZIM: Harare, Mashonaland [=se RSA: KwaZulu-Natal East Provinces Prov.]; Free State, Gauteng, ssp. laurentina Strohmeyer, 1928 “Delagoabai; Rikatla; Laurenco Limpopo, Mpumalanga, Marques“ [=s MOS: Maputo Northern Cape, North West City and Province]; ZIM Provinces; s MOS: Maputo ssp. scrobiculata (Bertoloni, 1847) Anthia “in provincia Inhambanensi Prov.; SWA, ZIM Mozambici“ [MOS: Inhambane =amabilis Boheman, 1848 Anthia “Montes Makkalienses“ [=c Prov.]; Gaza, Maputo, RSA: Gauteng Prov.] Zambézia Provinces; ne RSA: =spuria Boheman, 1848 Anthia “Boschjesmans Rand in Limpopo Prov.; BOW, ZIM Caffraria orientali“ [=sc RSA: Northern Cape Prov. ]

308 309 =tembena Péringuey, 1896 Polyhirma “Mozambique (Tembe)“ [=s RCA, RSA, TAN, TCH, UGA, MOS: Maputo Prov.] ZAM, ZIM; sw AS: SAU 45 opulenta (Boheman, 1860) Polyhirma s, sw AF: “juxta fluvii ssp. tenuicollis s. str. “montagnes de Schimba sur Svakop“ [=c NAB: Erongo la côte ďAfrique vis à vis de Reg.: Swakop River]; Hardap, Zanzibar“ [=e TAN: Pwani Khomas, Kunene, Omaheke, Reg.: Shimba Hills]; Arusha, Otjozondjupa Regions; ANG: Dodoma, Kilimanjaro, Tanga Huambo Prov.; BOW; s MOS: Regions; c, e KEN: Central, Maputo Prov.; RSA: Eastern Coast, Rift Valley Provinces Cape, KwaZulu-Natal, North =brevitarsis Müller, 1938 “Fort Hall, Africa orientale West Provinces; ZIM inglese“ [=c KEN: Central =scutellaris Chaudoir, 1861b Polyhirma “ lac N´gami“ [=nw BOW: Prov.] Ngami Lake] ssp. aenigma (Dohrn, 1881) Anthia type locality not stated, 46 rutata (Péringuey, 1892b) Polyhirma s AF: “Zambeze; ?Transvaal“ [probably “Zambese“ sn. [=ZAM, ZIM; e RSA]: Dohrn 1881 (=BOW, ZIM sn. KwaZulu-Natal, Limpopo Péringuey, 1896)]; s MOS: Provinces; BOW, MAW, s Maputo Prov.; RSA: KwaZulu MOS: Gaza, Maputo Provinces Natal, Limpopo Provinces 47 sambesina (Péringuey, 1908) Polyhirma se AF: “Zambezi River (Feira)“ ssp. banghaasi Strohmeyer, 1928 “D.-O.-Afr., Tanganjika“ [=s ZAM: e Lusaka Prov.: [=TAN]: Shinyanga Reg. Luangwa] =bulungwana Strohmeyer, 1928 “Njembe-Bulungwa“ [=nw 48 semisuturata (Chaudoir, 1866) Polyhirma se, sw AF: MOS, NAB, ZAM, TAN: Shinyanga Reg.] ZIM ssp. bifasciata (Bénard, 1926) Polyhirma ssp. semisuturata s. str. “Zambèze“ [=w MOS]; Gaza, ssp. circuligera (Fairmaire, 1887) Polyhirma Tabora [Reg. =w TAN]; Manica, Maputo, Zambézia Kagera, Kigoma, Singida Provinces; ZAM, ZIM Regions; UGA =neutra Dohrn, 1883 Anthia “ Holub‘schen Ausbeute: ssp. crampelina Strohmeyer, 1928 “Franz. Kongo: Scharigebiet: Zambese“ [=ZAM, ZIM] Fort-Crampel“ [=c RCA: ssp. commista (Péringuey, 1904) Polyhirma “North-West Damaraland Nana Grébizi Prefecture: Kaga [=n NAB: Kunene Reg.]; Bandoro] Ovampoland [=Ohangwena, ssp. eldamana Strohmeyer, 1928 “Eldama-River Station“ Omusati, Oshana, Oshikoto, [=w KEN: Rift Valley Prov.: Regions]; Southern Rhodesia“ Eldama Ravine] [=ZIM]; n NAB: Erongo, ssp. errata Strohmeyer, 1928 “Nordwest-Rhodesia“ [=w Otjozondjupa Regions; ZAM ZAM]; MAW; nw TAN: =damarina Basilewsky, 1959a “South West Africa: Damara“ Kigoma Reg. [=n NAB: Kunene Reg.] =bemba Strohmeyer, 1928 “Nordwesttanganjika; ssp. gambosana Strohmeyer, 1928 sw ANG: Gambos [=Huila Ubemba“ [=nw TAN: Kigoma Prov.] Reg.] ssp. vassei (Sternberg, 1907a) Polyhirma e MOS: “vallée du Pungoué: ssp. guluana Strohmeyer, 1928 n UGA: Gulu [Dist.] Guengère“ [=Sofala Prov.] ssp. heha Strohmeyer, 1928 “D.-O.-Afr., Uhehe; Uhehe- 49 tenuicollis (Chaudoir, 1878) c, e, s AF: BFO, BOW, CAM, Kidugala; Mitteluhehe; Uhehe CON, DOC, ERI, ETI, KEN, Tosmagonga; Süduhehe, MAW, MLI, MOS, NIG, NRE, Iringa-Mogologo; [=sw TAN:

310 311 Iringa Reg.]; Nord-Njassa, ssp. pangwana Strohmeyer, 1928 “D.-O.-Afr., Mpangwe; Ubena-Langenburg; Madibira; Bihawara“ [=sw TAN: Ruvuma [=Mbeya Reg.]; Rukwasee- Reg.]; Singida Reg. Steppe? [=Rukwa Reg.] =vicina Strohmeyer, 1928 “Grenzgebiet zw. Ugogo und ssp. hemigraphis Strohmeyer, 1928 “Insel Ukerewe [=n TAN: Turu, Saranda-Sawa“ [=c TAN: Mara Reg.]; Südviktoriasee: Singida Reg.] Muansa“ [=n TAN: Mwanza ssp. peregrina Strohmeyer, 1928 “Nordwest-Massai, Sossian- Reg.]; Arusha, Kagera Regions Ngoroine“ [=ne TAN: Arusha =circuligeroides Strohmeyer, 1928 “Ost.-Victoria-See: Ngoroine- Reg.] Mukenge“ [=n TAN: Mara, ssp. reflexicauda (Bénard, 1925c) Polyhirma “Congo français: Haute- Arusha Regions] Sangha, environs de Carnot“ =ochthodysma Strohmeyer, 1928 “S.-W.Victoria-See, Bukoba“ [=sw RCA: Mambéré-Kadéï [=n TAN: Kagera Reg.] Prefecture] ssp. horni (Sternberg, 1907b) Polyhirma “Kassanga (Tanganjika) [=sw ssp. salamensis (Kolbe, 1898) Polyhirma “D.-O-Afr. Kaiserl. TAN: Rukwa Reg.: Kasanga]; Gouvernement“ [=TAN]: Dar Langenburg (N. Tanganjika)“ Es Saalam, Dodoma, Lindi, [=s TAN: Mbeya Reg.: Manyara, Morogoro, Mtwara, Tukuyu] Pwani, Rovuma, Sinoida, =fipana Strohmeyer, 1928 “S-O Tanganjika: Ufipa- Tanga Regions; n MAW, n Plateau; Kassanga; MOS: Niassa Prov.; ZAM Ufipa: Msamwia; 1850m; =kondana Strohmeyer, 1928 “Irangi, Kondoa bis Irangi Bismarkburg“ [=Kasanga, sw [=sw TAN: Morogoro Reg.]; TAN: Rukwa Reg.] Iramba, Issansu Mkalama ssp. humilis (Guérin-Méneville, 1849) “Wood Medina en Arabie“ [=w [=Singida Reg.]; Ufiomi, Grofs SAU: Medína]; ERI, ETI: Afar Ufiomi“ [=Dodoma, Manyara Reg. Regions] =anchora Raffray, 1885 Polyhirma “Abrechoho, dans la vellée =makondana Strohmeyer, 1928 “D.-O.-Afr., Mikindani; duh aut Anseba“ [=n ETI: Afar Lindi; Makonde-Hochland, Reg.] Lukuledi; Ndanda; Suiva; ssp. kafuena (Péringuey, 1908) Polyhirma “Kafue River (Northern Kionga; Unterer Rovuma“ [=se Rhodesia)“ [=s ZAM: e TAN: Lindi, Mtwara, Ruvuma Southern Prov.]; se DOC: Regions]; Katanga Prov. =sambarana Strohmeyer, 1928 “D.-O.-Afr., Pangani-Tanga; ssp. kilimana Strohmeyer, 1928 “Kilimandjaro; Moschi“ [=ne Tanga Umg.; Tanga-Magila“ TAN: Kilimanjaro Reg.] [=ne TAN: Tanga Reg.] ssp. kiwirana Strohmeyer, 1928 “Massewe-Kiwira-Flufs; ssp. schirensis Strohmeyer, 1928 “Brit. Njassaland zw. Zomba Njasa“ [=sw TAN: Mbeya, und Blantyre“ [=s MAW: Iringa Regions] Southern Reg.] ssp. miskelli Basilewsky, 1980 wc KEN: Londiani; Naivasha; ssp. uelensis Burgeon, 1929 “Haut-Uele: Api; Mauda; Uasa Nyrio River, Mukogodo Dungu“ [=ne DOC: Orientale Ranch, nw Nanyuki [=Rift Prov.] Valley Prov.] ssp. ugandana Basilewsky, 1980 s UGA: “Tororo; btw. Jinja and ssp. ngona Strohmeyer, 1928 “Kigonsera“ [=sw TAN: Muria or Mbwago´s; Bugisu; Ruvuma Reg.]

312 313 Elgon; 6200ft [=Eastern 55 rohani (Bénard, 1921d) Polyhirma sw AF: “district de Huilla, Reg.]; Karamoja distr., Mt. Menongué“ [=sw ANG: Huila Kadam; Kotido; Nakwai Hills; Prov.] Turkana Plain near Nepal 56 zambeziana (Bénard, 1922c) Polyhirma s AF: “Haut Bassin de Pass [=Northeastern Reg.]; Zambèze: Loèti [=w ZAM: Nimuli to Murchinson Falls Western Prov.: Lungwebungu]; [=Northern, Western Regions]; Loenginga“ [=Luanguinga] w KEN: 40 mls e of Mt. Elgon; = rhodesiana (Bénard, 1925a) Polyhirma Kacheliba [=w Rift Valley NN for Polyhirma zambeziana Bénard, 1922c Prov.]; Mumias, Kisumu Road“ =benardi Basilewsky, 1931 [=Western Prov.]; SSD? nn for Polyhirma zambeziana Bénard, 1922c ssp. uhligi Strohmeyer, 1928 “Massai-Ngaruka [Engaruka]; Massai-Ngotiek“ =ne TAN: species incertae sedis: Arusha Reg.] 57 tatumana (White, 1846) Anthia unknown mouffletii group (Häckel sn. Basilewsky in Kleinfeld & Puchner, 2012) ECCOPTOPTERAChaudoir, 1878; T: Eccoptoptera cupricollis Chaudoir, 50 brevivittis (Chaudoir, 1866) Polyhirma sw AF: “Benguéla“ [=w ANG: 1878 (species: 2) Benguela Prov.]; Namibe Prov. =brevicollis Csiki, 1929 U.N. 1 cupricollis Chaudoir, 1878 e AF: BUR, DOC, ETI, KEN, 51 kassaica (Bénard, 1927) Polyhirma c AF: “Congo Belge; Bassin MAW, RCA, TAN, UGA, ZIM du Kassaï“ [=DOC: Kasai- ssp. cupricollis s. str. “montagnes du Schimba“ [=e Occidental, Kasai-Oriental TAN: Pwani Reg.: Shimba Provinces]; DOC: Bandundu, Hills]; Dar Es Saalam, Bas-Congo, Kinshasa Morogoro, Tanga, Zanzibar Isl. Provinces; n ANG: Zaire, Uige Regions; s KEN: Eastern Prov. Provinces =goetzei Strohmeyer, 1928 “D.-O.-Afr., Uluguruberge“ [=e 52 lundana Basilewsky, 1964b sw AF: e ANG: “Lunda“ TAN: Morogoro Reg.: Uluguru [=Lunda Norte Prov.] Mts.] 53 moufletii (Chaudoir, 1866) Polyhirma sw AF: “Benguéla“ [=w ANG: =methneri Strohmeyer, 1928 “Kwakijembe; Trockenwald Benguela Prov.] bei Mtohovu; unt. Umba“ [=ne =subhamata Oberthür, 1884 TAN: Tanga Reg.] 54 overlaeti Burgeon, 1935a c AF: DOC =salamensis Strohmeyer, 1928 “D.-O.-Afr., Daressalam; ssp. overlaeti s. str. “Congo belge: Kinda“ [=s Daressalam, Puguberge- DOC: Katanga Prov.] Kisserawe“ [=e TAN: Dar Es ssp. kwangoana Basilewsky, 1963 “Kwango, Kianza terr. de Saalam Reg.] Feshi“ [=sw DOC: Bandundu ssp. albertina Basilewsky, 1980 “Zaïre: Niarembe; Mahagi; Prov.] Mahagi-Niarembe; Région ssp. lualabana Basilewsky, 1955 Lulua: Limdji Riv.; Kapanga; ďAbok; [=ne DOC: Orientale Tshibamba; Sandoa; Lunkinda Prov.]; w UGA: Butiaba, rive Riv.; Kasai-Lunene Riv.; orientale du lac Albert“ [=right Kapelekese Riv.; Kafakumba bank of Albert Lake] [=s DOC: w Katanga Prov.] ssp. alluaudi Sternberg, 1906c “Kibwezi (Wa-Kamba, östlich vom Kilima-Ndjaro Britisch Ost-Afrika)“ [=s

314 315 KEN: s Eastern Prov]; Rift [=Iringa Reg.]; Mpangwe“ Valley Prov.; ne TAN: Arusha, [=Singida Reg.]; Kagera, Kilimanjaro, Tanga Regions Rukwa, Ruvuma, Tabora =betheliana Strohmeyer, 1928 “Usambara Neubethel“ [=ne Regions TAN: Tanga Reg.] =nyika Strohmeyer, 1928 “Nord-Njassa, Konde-Unjika =eldamae Strohmeyer, 1928 “Eldamariver-Station [=w [=Rukwa Reg.]; Neuhelgoland; KEN: Rift Valley Prov.: Moko“ [=Ruvuma Reg.] Eldama Ravine]; nördl. =taborana Strohmeyer, 1928 Tabora [Reg.]; Ugogo bis Massailand“ [=ne TAN: Arusha Unjamwesi; Unjanjembe; Reg.] Kakoma; Turu [=Kagera Reg.]; =montium Strohmeyer, 1928 “West-Usambara“ [=ne TAN: Kilimatinde; Iramba, Issansu, Tanga Reg.] Mkalama [=Singida Reg.] ssp. burundina Basilewsky, 1980 BUR: “Makoronkwe, dans le ssp. meridiana Strohmeyer, 1928 “Brit. Njassaland zw. Zomba Mosso; Bujumbura“ und Blantyre“ [=s MAW: ssp. chalcodera (Harold, 1881) Polyhirma w KEN: Taita District [=Rift Southern Reg.] Valley Prov.]; Ukamba [=nw ssp. pufirana Strohmeyer, 1928 “S-O Tanganjika: Kassanga; TAN: Kilimanjaro Reg.] Ufipa-Plateau; Tanganjikasee“ ssp. clarkei Basilewsky, 1980 s ETI: “Sidamo Prov., 38- [=sw TAN: Rukwa Reg.] 40 km e of Neghelli; 65 km ssp. ukerewensis Strohmeyer, 1928 “Insel Ukerewe“ [=n TAN: e of Neghelli; Wadera“ [=n Mwanza Reg.] Southern Nations, Nationalities ssp. viri Strohmeyer, 1928 “Neu-Kamerun; Uam.-Gebiet, and Peoples; Oromiya Regions] Bosum“ [=w RCA: Ouham- ssp. gallensis Strohmeyer, 1928 “Nordgalla, Gambo; Guda Pendé Prefecture]; Kémo, [=c ETI: Oromiya Reg.]; Lobaye, Nana-Grébizi, Nana- Abessinien 70 km nw v. Mambéré, Mambéré-Kadéï, Harrar“ [=Somali Reg.] Ombella-M´poko, Ouham, ssp. harrarensis Basilewsky, 1980 ne ETI: “Harar; Besetino Sangha-Mambéré Prefectures [=Harari Reg.]; Dire Dawa“ =crampelina Strohmeyer, 1928 “Fort-Crampel“ [=c RCA: [Reg.] Nana Grébizi Prefecture: Kaga ssp. katangana Basilewsky, 1953 se DOC: Katanga Prov.: Bandoro] Upemba N.P. ssp. wituana Strohmeyer, 1928 “Witu-Kipiui“ [=se KEN: ssp. kigonserana Strohmeyer, 1928 Kigonsera [=sw TAN: Ruvuma Coast Prov.] Reg.] 2 mutilloides (Bertoloni, 1857) Anthia e AF: DOC, KEN, MAW, ssp. lukeledina Strohmeyer, 1928 “Lukeledi [=Lukuledi, se MOS, TAN TAN: Mtwara Reg.]; Lindi; ssp. mutilloides s. str. “in ripis fluminis Manárra Makonde-Hochland“ [=Lindi provinciae Inhambanensis Reg.] Mozambici“ [=se MOS: ssp. madibirana Strohmeyer, 1928 “Madibira; Ubena-Langenburg; Inhambane Prov.]; Maputo City Kilossa-Ssinger; [=Mbeya and Province Reg.]; Bihawana; SO Usagara =labrata Fairmaire, 1891a MOS [=Dodoma Reg.]; Mitteluhehe; ssp. adonia Kolbe, 1896 “Ostafrika“ [=TAN] 1000-1700m; Pawagga Bez. ssp. fontium Strohmeyer, 1928 “Kigonsera; Ost-Ungoni; Iringa; 800m; Lusulwepass Kipenhere“ [=sw TAN: Ruvuma Reg.]

316 317 ssp. lagenula (Gerstaecker, 1867) Polyhirma “Mombas“ [=se KEN: Coast Tendaguru-Lindi; Makonde- Prov.: Mombasa Distr.]; Hochland; Namupa [=Lindi Rift Valley Prov.; ne TAN: Reg.]; Brit. Njassaland zw. Kilimanjaro, Tanga Regions; s Zomba und Blantyre“ [=s UGA MAW: Southern Reg.] =roeschkei Sternberg, 1906c “Ost-Usambara (Deutsch Ost- Aafrika)“ [=ne TAN: n Tanga GONOGENIAChaudoir, 1844; T: Carabus tabidus Fabricius, 1792 (species: 7) Reg.] =Microlesta Chaudoir, 1850; T: Microlesta spinipennis Chaudoir, 1850 =intima Strohmeyer, 1928 “Eldamariver“ [=w KEN: Rift 1 atrata (Boheman, 1848) Anthia s AF: NAB, RSA Valley Prov.: Eldama Ravine] ssp. atrata s. str. “Caffraria interior“ [=n RSA]; ssp. mela Strohmeyer, 1928 “Rombo [=Kilimanjaro RSA: Free State, Northern Reg.]; Massai-Ngaruka; Cape, North West Provinces; Nord-Massai Sossian- ne NAB: Caprivi Reg. Ngoroine [=Arusha Reg.]; ssp. nigrina (Boheman,1860) Polyhirma “juxta fluvios Svakop et Uhehe-Iringa; [=Iringa Reg.]; Nolagi“ [=c NAB: Erongo Bukoba-Victoriasee [=Kagera Reg.: Swakop, Nolagi Rivers] Reg.]; Ufiomi; Issansu bezw. ssp. schultzei (Kuntzen, 1919) Polyhirma “Kubub“ [=s NAB: Karas Hohenlohegraben, Mkalame- Prov.] Jaida-Sumpf [=Manyara Reg.]; 2 endroedyi Basilewsky, 1980 s AF: “Afrique du Sud Usuafua, Bayoberge; Utengele; (Little Namaqualand). Ubena; Kilossa; Langenburg Honderkliepbaai [=Hondeklip [=Mbeya Reg.]; Insel Ukerewe Bay]; Rietfontein; Alexander [=Mwanza Reg.]; Unika- Bay; Holgat Mouth; Mbosi; Kundi; Nkila-Ukimbu; Oogarabies; Namaqualand Pt. Kilimatinde; Kilimatinde- Noloth; Buffeis riv. Valley; Tabora; Kilimatinde-Turu; Quaggafontein; Outspan Iramba, Issansu Mkalama; Farm; Gemsbok vlagte Farm; Irangi-Ikome; Grenzgebiet, Strandfontein Farm; Kwass zwischen Ugogo und Turu, Farm; Namaqualand“ [=w Ssuna-Ssingida; Mtotea-Ssuna; RSA: Northern Cape Province] Landschaft Bassi, Salanka 3 immerita (Boheman, 1860) Polyhirma sw AF: ANG, NAB, RSA [=Singida Reg.]; Kigoma Reg.; ssp. immerita s. str. “vicinitas fluvii Svakop“ [=c BUR, se DOC: Katanga Prov.; NAB: Erongo Reg.: Swakop ZAM Riv.]; Hardap, Khomas, =kamica Strohmeyer, 1928 “West Ukami“ [=sc TAN: Otjozondjupa Regions Iringa Reg.] ssp. cribricollis (Chaudoir, 1861) Polyhirma “Benguela“ [=w ANG]; NAB =vinza Strohmeyer, 1928 “Ost-Tanganjika, Uwinsa- ssp. kalaharica Basilewsky, 1980 RSA: Kalahari Gemsbok Mlagarassiflufs; Ost- National Park; Mata Mata; Tanganjika, Könsi-Uwinsa“ 25 km s of Mata Mata; Twee [=w TAN: Kigoma Reg.] Rivieren [=Northern Cape ssp. selenolepta Strohmeyer, 1928 “O.-Afr. Lukuledi [=se Prov.]; “South –West Africa. TAN: Mtwara Reg.]; Lindi; Duineveld near Stempriet [=c NAB: Hardap Reg.];

318 319 Wildheim Est, 384; Katzies, Mpumalanga, North West Keetmanshoop distr.“ [=Karas Provinces]; Manicaland“ [Prov. Reg.] =e ZIM]; ZAM 4 oxygona (Chaudoir, 1844) Anthia s AF: “Cap de Bonne- 2 malangana Strohmeyer, 1928 sw AF: n ANG, s DOC Espérance“ [=s RSA]: Northern ssp. malangana s. str. “Malange W.-Afr.“ [=n ANG: Cape Prov. Malanje Prov.] 5 rugosopunctata (Thunberg, 1806) Anthia s AF: “Caput bonae spei“ [=s =buchneri [Kolbe in litt.] sn. Strohmeyer, 1928 “ Malange W.-Afr.“ [=n ANG: RSA]: Eastern Cape, Free Malanje Prov.] State, KwaZulu-Natal, North ssp. kwangoana Basilewsky, 1962 “Kwango“ [=s DOC: West Provinces Bandundu Prov.] =rubiginosa Perroud, 1846 Anthia “ ľ intérieur de Natal“ [=se 3 vethi Bates, 1889 sw AF: “Benguella“ [=w ANG] RSA: KwaZulu-Natal Prov.] 6 spinipennis (Chaudoir, 1850) Microlesta s AF: “Cap de Bonne- Espérance“ [=s RSA]: Eastern, HELLUONINI Hope, 1838 (subtribes: 2; genera: 26; species: 182) Western Cape Provinces 7 tabida [Olivier in litt.] (Fabricius, 1792) Carabus s AF: “Cap de Bonne- HELLUONINAHope, 1838 genera: 14; species: 40) Espérance“ [=s RSA: Western Cape Prov.]; North West Prov. AENIGMANewman, 1836; T: Aenigma iris Newman, 1836 (species: 1) =spinosus Linnaeus, 1767 Carabus “caput bonae spei“ [=s RSA] =obtusa Chaudoir, 1869a Microlesta “ľAfrique australe“ [=s RSA] 1 iris Newman, 1836 AU: “New Holland“ [=AUS]: e coastal: e Qld, e Nsw NETRODERA Chaudoir, 1850; T: Anthia formicaria Erichson, 1843 =newmani Laporte de Castelnau, 1867 Nsw: Cooks River (species: 3) =splendens Laporte de Castelnau, 1867 Qld: Port Denison

1 formicaria (Erichson, 1843) Anthia sw AF: ANG, BOW, NAB, AMETROGLOSSUS Sloane, 1914; T: Gigadema atrum W. J. Macleay, RSA, ZAM, ZIM 1887 (species: 1) ssp. formicaria s. str. ANG; s RSA: Free State, KwaZulu-Natal Provinces 1 ater (W. J. Macleay, 1887) Gigadema AU: ne AUS: Qld: Russell ssp. damarensis (Kuntzen, 1919) Atractonotus “Windhuk [=c NAB: Khomas River, Cairns District; ne Reg.: Windhoek]; Okahandja coastal [=Otjozondupa Reg.]; Karibib; Omaruru [=w NAB: Erongo DICRANOGLOSSUSChaudoir, 1872; T: Helluosoma resplendens Laporte Reg.]; Reheboth“ [=Hardap de Castelnau, 1867 (species: 1) Reg.] ssp. kalahara Basilewsky, 1980 c BOW: Kaotwe; Gomodimo 1 resplendens (Laporte de Castelnau, 1867) AU: ne AUS: Qld: Port [=Ghanzi Prov.: Central Helluosoma Denison; n Nte, n Wau Kalahari Game Reserve]; =parvulum W. J. Macleay, 1873 Aenigma “New Holland“ [=AUS] “South West Africa. Kungveld, Mangetti Forest, betw. Tsumkwe and Chassie“ [=e NAB: Omaheke Reg.] ssp. peringueyi Basilewsky, 1948 “Transvaal [=e, nw RSA: KwaZulu-Natal, Limpopo,

320 321 EPIMICODEMASloane, 1914; T: Helluosoma mastersii W. J. Macleay, Range; s Nte, Wau 1871 (species: 1) =longius Blackburn, 1901b “Central Australia“: MacDonnell Ranges [=s Nte] 1 mastersii (W. J. Macleay, 1871) Helluosoma AU: ne AUS: e coastal: se Qld: 9 nocte (Newman, 1842) Enigma AU: e AUS: Nsw: Woodside, Gayndah; Nsw near Sydney [probably uncertain]; s Qld, e Sau GIGADEMAThomson, 1859; T: Gigadema titanum Thomson, 1859 =titana Thomson, 1859a “Morton-Bay, Nouvelle- =Gigadema nocte (Newman, 1842) (species: 13) Hollande“ [=ne AUS: Qld: Port =Aenigma Newman, 1836; T: Aenigma iris Newman, 1836 Denison] =Enigma Newman, 1842 in error 10 obscurum Sloane, 1914 AU: e AUS: s Qld: Cunnamulla =Penichrodema Gestro, 1875; T: Gigadema minuta, G. thomsoni, G. paroensis all Laporte de Castelnau, 1867 sulcatum group (sn. Sloane 1914): 11 biordinatum Sloane, 1914 AU: ne AUS: Qld: Torres grande group (sn. Sloane 1914): Straits: Prince of Wales Is. 1 grande (W. J. MacLeay, 1864) Helluo AU: ne AUS: Qld: Port 12 froggatti W. J. Macleay, 1888 AU: n AUS: n Wau: King Denison; ne coastal Sound; Kimberley Reg.; Qld: =prominens Sloane, 1914 U.N. Qld: Coen ne coastal 2 gulare Sloane, 1914 AU: “Central Australia“ [=n 13 sulcatum (W. J. Macleay, 1864) Helluo AU: e, n, w AUS: Qld: Port AUS: s Nte:]: Tennant Creek; Denison; Nsw; Nte, e Sau, Qld Wau: w coastal 3 maxillare Sloane, 1914 AU: ne AUS: Qld: Cooktown; =paroensis Laporte de Castelnau, 1867 Nsw: Paroo River Kuranda; Princess Charlotte =minuta Laporte de Castelnau, 1867 Qld: Ipswich Bay; Townsville; New Guinea =thomsoni Laporte de Castelnau, 1867 Qld: Port Denison Is.: s PNG: Western Prov.; =damelii W. J. Macleay, 1873 Cape York [=ne AUS: n Qld] Torres Straits: Mona Is. =grandis sn. Gestro 1875 Qld: Night Is.; Port Denison; HELLUARCHUSSloane, 1914; T: Helluarchus robustus Sloane, 1914 Somerset (Cape York) (species: 2) 4 rugaticolle Blackburn, 1901a AU: “N. W. Australia“ [=n Wau]: Kimberley Reg. 1 robustus Sloane, 1914 AU: wc AUS: Wau: Anketell, near Cue lopngipenne group (sn. Sloane 1914): 2 whitei Lea, 1914 AU: c AUS: se Nte:Finke River 5 longipenne (Germar, 1848) Helluo AU: s, w AUS: Sau: Adelaide; s Qld, Nsw, Vic, Wau =politulum W. J. Macleay, 1871 Gayndah [=ne AUS: se Qld] HELLUAPTERUSSloane, 1914; T: Helluapterus niger Sloane, 1914 (species: 1) nocte group (sn. Sloane 1914): 6 bostocki Laporte de Castelnau, 1867 AU: c, nw, s, w AUS: n Wau; 1 niger Sloane, 1914 AU: wc AUS: Wau: Murchison Nsw; s, w Nte; Sau; Vic District =intermedia Gestro, 1875 Nsw: Lachlan River =punctata Sloane, 1914 U.N. Wau: Shark Bay =eremita Blackburn, 1892 Wau: Fraser Range =longicolle Blackburn, 1901b “Central Australia“: Oodnadatta [=n Sau] 7 dux Blackburn, 1901b AU: w AUS: Wau 8 mandibulare Blackburn, 1892 AU: c, w AUS: Sau: Everard

322 323 HELLUOBonnelli, 1813; T: Helluo costatus Bonnelli, 1813 (species: 2) 5 laevifrons Darlington, 1968 AU: e New Guinea Is.: “N-E. N. G., Toricelli Mts., Mokai 1 costatus Bonnelli, 1813 AU: e AUS: ?Vic; Act, Qld, village“; 750m [=nw PNG: Nsw Sandaun Prov.] =carinatus Chaudoir, 1848 “Melbourne, sur les bords de la 6 latipennis (W. J. Macleay, 1887) Helluosoma AU: ne AUS: n Qld: Cairns Rivière des Cygnes; Adelaide“ 7 latipes Darlington, 1968 AU: w New Guinea Is.: [=s, se AUS: Sau, Vic] “West N. G., Rattan Camp“; 2 insignis Sloane, 1890 AU: se AUS: Murray-Darling 1200m [=West Papua (IDN): basin: Nsw: Darling River Jayawijaya Dist.] 8 politus Darlington, 1968 AU: c, e New Guinea Is.: “N- HELLUODEMALaporte de Castelnau, 1867; T: Aenigma unicolor Hope, E. N. G., Morobe Distr., Maba 1842 (species: 2) Vy., Menyama [=ne PNG: =Simoglossus Chaudoir, 1872; T: Simoglossus niger Chaudoir, 1872 (=unicolor Morobe Prov.]; Maprik [=East Hope, 1842) Sepik Prov.]; Oriomo River“; 6m [=Western Prov.] 1 brunneum Sloane, 1917 AU: n AUS: Nte 2 unicolor (Hope, 1842) Aenigma AU: “New Holland“ [=AUS]: HELLUOPAPUA Darlington, 1968; T: Helluopapua toxopei Darlington, Nsw, Qld; s New Guinea Is.: 1968 (species: 2) PNG: Western Prov.; s West Papua (IDN): Merauke Dist. 1 cheesmani Darlington, 1971 AU: Waigeo Is.: “West N. G., =batesii Thomson, 1857a ?Helluomorpha “Ega, Amazone supér.“ [=s Mt. Nok, Camp 2“ [=West Papua AM: BRA] (in error sn. (IDN): Raja Ampat Dist.] Laporte de Castelnau 1867 2 toxopei Darlington, 1968 AU: w New Guinea Is.: AUS: Qld) “West N. G., Rattan Camp“; =niger Chaudoir, 1872 Simoglossus AUS 1200m [=West Papua (IDN): =castelnaui Chaudoir, 1872 Gigadema AUS Jayawijaya Dist.]

HELLUONIDIUS Chaudoir, 1872; T: Aenigma cyanipennis Hope, 1842 HELLUOSOMALaporte de Castelnau, 1867; T: Helluosoma atrum Laporte (species: 8) de Castelnau, 1867 (species: 4)

1 aterrimus (W. J. Macleay, 1873) Helluosoma AU: ne AUS: Qld: Cape York; 1 bouchardi Baehr, 2005a AU: n AUS: nw Nte: 20,5 km ne coastal nne Bullita, Gregory Nat Pk; 2 chrysocomes Maindron, 1908 AU: e New Guinea Is.: 15.57ºS, 130.29ºE “Sentani“ [=West Papua 2 hangayi Baehr, 2005a AU: ne AUS: c Qld: Wild (IDN): Jayapura Distr.]; PNG: Horse Creek Central, Eastern Highlands, 3 longicolle W. J. Macleay, 1888 AU: n AUS: n Wau: King Madang Provinces Sound; 3 cyaneus (Laporte de Castelnau, 1867) Helluosoma AU: n, e AUS:e Qld: 4 atrum Laporte de Castelnau, 1867 AU: ne AUS: Qld: Rockhampton; e Nsw: Clarence Rockhampton; e Nte; New River; n Nte Guinea Is.: s PNG 4 cyanipennis (Hope, 1842) Aenigma AU: n AUS [=“Port Essington“ sn. Sloane 1914]; n, ne coastal =viridipenne W. J. Macleay, 1887 Helluosoma Cairns [=ne AUS: n Qld]

324 325 NEOHELLUOSloane, 1914; T: Neohelluo angulicollis Sloane, 1914 (species: 1) 2 binoculus Bates, 1892 s AS: “Bhamò“ [=n MYA: s Kachin state]; e IND: Ass, 1 angulicollis Sloane, 1914 AU: ne AUS: “Tropical Mhy; [LAO sn. Andrewes 1930] Queensland, near Cairns; 3 bisignata Landin, 1955 s AS: ne MYA: Shan States: Townsville“ [=ne Qld] Pekkong; 900m 4 distacta (Wiedemann, 1823) Helluo se AS: w IDN: Java Is.; PLATYHELLUOBaehr, 2005a; T: Platyhelluo weiri Baehr, 2005 (species: 1) Sumatra Is.; SIG 5 hamaticollis Bates, 1892 se AS: n Borneo Is.: Sarawak 1 weiri Baehr, 2005a AU: n AUS: nw Nte: 8 km e by [Prov. =e MAL] n Jammam, Keep River Nat. 6 labrosa Nietner, 1857 s, se AS: “Ceylon“ [=SRL]; w Pk; 15.45ºS, 129.109ºE IDN: Java Is., s IND: Ker, Krt, Tmd; MYA, n VIE, THA; AU: OMPHRINAJedlička, 1941 (genera: 12; species: 142) ne AUS: Qld; New Guinea Is.: =Helluomorphina Reichardt, 1974 West Papua (IDN), s PNG =piceus Schaum, 1863 Acanthogenius COLFAXAndrewes, 1920; T: Colfax stevensi Andrewes, 1920 (species: 1) nn for Helluo labrosus Nietner, 1857b =affinis Gestro, 1875 “Bangkok“ [=sc THA: Krung 1 stevensi Andrewes, 1920 e, s, se AS: Sikkim: Thep] Gopaldhara; British Sikkim: 7 lineola Andrewes, 1926b se AS: w IDN: Sumatra [Is.]: w Pashok; 2500´ [=IND: SiD]; MAL: Selangor Assam: Sadiya [=Ass]; Bengal: 8 rubrothorax Louwerens, 1949 se AS: c Borneo Is.: Chapra [=Ben]; Bombay: “Muara Tewe (Zuider- en Bassein Fort [=Mhr]; Burma: Oosterafdeling van Borneo)“ Tenasserim, Mergui [=s MYA: [=c IDN: Kalimantan Tengah Tanintharyi Reg.]; Indo- Prov.] China: Tonkin, Hoabinh; 9 wilsonii (Laporte de Castelnau, 1867) Pseudohelluo AU: ne AUS: Qld: Brisbane; Yen Bai [Prov. =n VIE]; [se AS: w IDN: Java Is. sn. Annam: Cuarao; Vinh [=s Andrewes 1930] VIE: Nghệ An Prov.]; LAO: Houei Ko; Natung; Muong DAILODONTUSReiche, 1843; T: Helluo cayennensis Dejean, 1826 (species: 2) Sai; Sop Choun; Vientiane; [=Khammouane, Oudomxai, 1 clandestinus (Klug, 1834) Polistichus s AM: Pra [=n BRA]; Ama, Phongsali, Savannaket, Bhi, Mgr, Spo; ARG: Chc, Crr, Vientiane Provinces]; se CHI: Eri, Sfe, Sge; PAR, URU Fuj; NEP; VIE =rufipes Brullé, 1838 Helluo “dans la province de Corrientes et sur les bords du Rio de CREAGRISNietner, 1857; T: Creagris labrosa Nietner, 1857 (species: 9) Coronda près Santa Fé =Pseudohelluo Laporte de Castelnau, 1867; T: Pseudohelluo wilsonii Laporte de (republique Argentine)“ [=n Castelnau, 1867 ARG: Cor, Sfe] 1 bigemmis Andrewes, 1931 se AS: s Sumatra[Is.]: =erythropus Chaudoir, 1843 Helluo Lampongs, Wai Lima [=w NN for Helluo rufipes Brullé, 1838 IDN: Lampung Prov.]; Perak: =intermedius Chaudoir, 1872 Polystichus not stated [“Haute Amazone, Batang Pedang, Tor Camp; Serpa“ sn. Reichardt 1974] 1800feet [=w MAL]

326 327 2 cayennensis (Dejean, 1826) Helluo s AM: “Cayenne“ [=FRG]; lacordairei group (sn. Reichardt 1974): BRA: Goi, Pra, Spo; VEN 8 ares Reichardt, 1974 s AM: se BRA: Spo: Barueri; Atibaia; Ipiranga; EREPHOGNATHUSAlluaud, 1932; T: Erephognathus xiphogenius Paranapiacaba; Piracicaba; Alluaud, 1932 =Erephognathus coerulescens (Fairmaire, 1903) (species: 2) Pirassununga; Santo Amaro; Sorocaba; Goi: Leopoldo 1 coerulescens (Fairmaire, 1903) Glycia e AF: e, n, nw, MAD: Bulhoões; Msg: Araxá; Ankarahitra [=Antananarivo Camanducaia, Vila Monte Prov.: Betsioka Reg.: Verde; Pedra Azul; Poços de Maevatanana Dist.]; Caldas, Morro do Ferro; Pna: Fianarantsoa Prov. Arapoti; Cachoeirinha; 600m; =xiphogenius Alluaud, 1932 Ananalava [=nw MAD: Jaguariaiva; Ponta Grossa; Sac: Mahajanga Prov.] Joaçaba; Nova Teutônia 2 margarithrix Alluaud, 1936 e AF: ce MAD: “Périnet“ 9 anthracinus (Klug, 1834) Helluo s AM: “Brasilien“ [=wc, se [=Toamasina Prov.: Andasibe- BRA]: DtF, Goi, Pna, Spo Mantadia National Park: 10 argus Reichardt, 1974 s AM: se BRA: Spo: Barueri; Analamazoatra] Piracicaba 11 bechynei Reichardt, 1974 s AM: wc BRA: Mgr: Chapada HELLUOBROCHUS Reichhardt, 1974; T: Helluobrochus ares Reichardt, dos Guimarães 1974 (species: 26) 12 birai Reichardt, 1974 s AM: se BRA: Spo: Itu 13 collaris (Chaudoir, 1877) Pleuracanthus s AM: “Bahia (Brésil)“ [=ne brasiliensis group (sn. Reichardt 1974): BRA: Bhi] 1 bacchus (Reichardt, 1972) Pleuracanthus s AM: se BRA: Spo: Itu, Fazenda 14 cribricollis (Chaudoir, 1872) Pleuracanthus s AM: “province de Mines au Pau ďAlho; Msg, Sac; ARG: Brésil“ [=wc BRA: Msg]; Mgr, Chc, Mns; BOL, PAR, PER Spo; ARG: Crr, Mns; n PAR 2 brasiliensis (Dejean, 1825) Helluo s AM: “Brésil“ [=se BRA]: 15 darlingtoni Reichardt, 1974 s AM: c, n PAR: Concepción Eps, Rdj, Sac; ne ARG: Mns [Dept.]: near Horqueta; 3 brevicollis (Dejean, 1831) Helluo s AM: “Cayenne“ [=FRG]; n, Amambay [Dept.]: Amambay; ne BRA: Ama, Amp, Mha, Pra; Guaiará [Dept.]: Paso Yobay; PER Villarica; wc BRA: Mgr: =ebeninus Bates, 1871 Pleuracanthus “Ega“ [=n BRA: Ama: Tefé] Rondonópolis; Msg: Viçosa; 4 capixaba Reichardt, 1974 s AM: se BRA: Eps: Santa Spo: Botucatu; Ribeirão Preto; Teresa FRG: Roches de Kourou 5 cribratus (Reiche, 1842) Pleuracanthus c, s AM: VEN; n BRA: Ama, 16 horqueta Reichardt, 1974 s AM: n PAR: Concepción Amp, Bhi; COL, COS, FRG, [Dept.]: near Horqueta GUY, MEX: Ver; NIC, PER, 17 inconspicuus (Chaudoir, 1848) Pleuracanthus s AM: “Brésil mérid.“ [=wc SAL, SUR BRA]: Mgr; ne ARG: Chc, 6 luctuosus (Chaudoir, 1872) Pleuracanthus s AM: “Brésil“ [=s BRA]: Pna, Crd, Crr, Mns; c PAR Rgs, Sac, Spo; BOL 18 lacordairei (Dejean, 1831) Helluo s AM: c, n ARG: Tuc; 7 oculatus Reichardt, 1974 s AM: se BRA: Eps: Córrego “particulièrement dans le Sierra do Itá de Cordoba“ [=Crd]; Lrj, Sat, Sge

328 329 19 negrei Reichardt, 1974 s AM: n ARG: Sge: Choya; Brasília; Mgr: Rio Verde; Fernandez; Lundo Coral; w Msg: Araguari; Betim; ‚Minas, PAR: “Chaco“ Parzud‘; Moro da Garça; 20 petrus Reichardt, 1974 s AM: wc BRA: Mgr: Utiariti, Unaí, Fazenda Bolívia; Spo: Rio Papagaio; Rio Verde; Avanhandava; Botucatu; Piraju; 400m; Goi: Jataí; Msg: Pirassununga, Emas; ne ARG: Uberaba Mns; c, e BOL: Ichilo [Prov.]: 21 sanguinolentus (Klug, 1834) Helluo s AM: “Brasilien“ [=BRA]: Buenavista; Santa Cruz [Prov.]: Goi, Mgr, Msg; n PAR Montero; n PAR: Concepción [Dept.]: near Horqueta; oopselaphus group (sn. Reichardt 1974): 2 eulinae Reichardt, 1974 s AM: wc, e BRA: Spo: 22 bucki Reichhardt, 1974 s AM: se BRA: Msg: Serra São José do Rio Preto; do Caraça; Pna: Guarapuava; Botucatu; Piracicaba; Piraju; Rgs: Serro Azul; c PAR: Guairá Pirassununga; Bhi; Goi: [Dept.]: Paso Yobay; 280m; ne Cabeceiras, Lagoa Formosa; ARG: Mns Jataí; Leopoldo Bulhões; 23 oopselaphus Reichardt, 1974 s AM: ne BRA: Pnb: Recife; Vianópolis; Mgr: Rio Verde; Paulista, Maria Farinha Msg: Araxá; Rgn: Natal; 24 pilipalpis Reichardt, 1974 qs AM: BOL: Santa Cruz Spo: Botucatu; Piracicaba; [Prov.]: Montero Piraju; Pirassununga; n PAR: Concepción [Dept.]: near subrostratus group (sn. Reichardt 1974): Horqueta 25 linearis (Bates, 1871) Helluomorpha s AM: “Ega“ [=n BRA: Ama: 3 heros (Gory, 1833) Helluo s AM: “Brésil“ [=wc, e BRA]: Tefé] Goi, Mgr, Msg, Pna, Rgn; PAR 26 subrostratus (Bates, 1871) Helluomorpha s AM: “Ega“ [=BRA: Ama: =bellicosa Laporte de Castelnau, 1834 “Brésil“ [=BRA] Tefé]; Mgr; Pna, Rgs; n ARG: Mns, Sat, Tuc; ECU, n PAR, macroptera species (sn. Reichardt 1974): PER 4 macroptera Chaudoir, 1850 c, s AM: “venant de Brésil“ =sinuatus Liebke, 1935 Pleuracanthus qn ARG: Mns, Sat, Tuc; se [=cw, se BRA]: Eps, Mgr, Msg, BRA: Rgs: Porto Alegre Rdj, Spo; ECU, PAN, PAR, PER HELLUOMORPHA Laporte de Castelnau, 1834; T: Helluo heros Gory, =chabrillaci Thomson, 1857b Herinnis “Rio Janeiro“ [=se BRA: Rdj] 1833 (species: 4) =obscuricornis Chevrolat, 1858 “ le Brésil intérieur: Lagoa =Herinnis Thompson, 1857b; T: Herinnis chabrillacii Thompson, 1857b Santa“ [=se BRA: Msg] =Erinnys Gemminger & Harold, 1868 in error (emendation in catalog) HELLUOMORPHOIDESBall, 1951; T: Helluomorpha texana LeConte, heros group (sn. Reichardt 1974): 1853 (species: 24) 1 araujoi Reichardt, 1974 s AM: wc, se BRA: Goi: Cabeceiras, Lagoa Nearctic groups: praeustus group (Häckel sn. Reiche 1843): Formosa; Corumbá, Fazenda 1 clairvillei (Dejean, 1831) Helluo “ľAmérique septentrionale“ Monjolinho; Fazenda Nova [=n AM: s USA: se Mis: Orlândia; Goiânia, Campinas; George County: Lucedale sn. Jataí; Vianópolis; DtF: Lindroth 1955]: Aba, Flo, Gor, Mis, Nca, Nje, Sca

330 331 2 ferrugineus (LeConte, 1853) Helluomorpha c, n AM: s USA: s Tex 9 texanus (LeConte, 1853) Helluomorpha c, n AM: s USA: e Tex [Smith [Cameron County: Brownsville County: Tyler sn. Bousquet sn Ball 1956]; Aba, Ari, Ark, 2012]; Aba, Ark, Cor, Gor, Gor, Kan, Nca, Nje, Nme, Okl, Iow, Kan, Ken, Mis, Mso, Neb, Sca, Tex, Uta, Vir; ?Flo, Nyo; n Nme, Okl,Ten, ; ?Ari; ne MEX: MEX: Chh, Coh, Dur, Sin, Tam Tam =languida Caey, 1913 Helluomorpha “ Texas (Brownswille)“ [=s neotropical groups: USA: s Tex: Cameron County] femoratus group (sn. Reichardt 1974): 3 latitarsis (Casey, 1913) Helluomorpha n AM: sw USA: Ari [Pima 10 femoratus (Dejean, 1831) Helluo s AM: “Sierra de Cordoba“ County: Baboquivari Mts. sn. [=c, n ARG: Crd]; Tuc; Crd, Ball 1956]; Nme, Tex; ?Okl; n Mdz, Mns, Snl; BOL, BRA: MEX: Chh, Coh, Dur, Sin Pna, Mgr, Msg, Sac 4 longicollis (Bates, 1883) Helluomorpha c AM: GUA: Capetillo =sparsus Brullé, 1838 Helluo “province de Santa Cruz de 5 mexicanus (Chaudoir, 1872) Helluomorpha c AM: MEX: Yuc; Ver la Sierra“ [=BOL: Santa Cruz 6 nigripennis (Dejean, 1831) Helluo “ľAmérique septentrionale“ Prov.] [=n AM: e, s USA: s Gor sn. 11 nigerrimus (Klug, 1834) Helluo s AM: “Brasilien“ [=wc, s Lindroth 1955; Nca: Wake BRA]: Goi, Msg, Pna, Rdj, County: Raleigh sn. Lindroth Rgs, Sac; ne ARG: Mns 1955]: Aba, Ark, DiC, Gor, Ili, =coracina Mannerheim, 1837 Helluomorpha “Brasilia interiore“ [=BRA] Lou, Mar, Mas, Mic, Mis, Mso, =melanaria Reiche, 1843 Helluomorpha BRA Nca, Nje, Nyo, Okl, Sca, Tex, 12 rubricollis (Schaum, 1863) Helluomorpha s AM: n ARG: “Rosario am Vir La Plata“ [=Sfe]; Bua, Eri, Sat; 7 papago (Casey, 1913) Helluomorpha c, n AM: sw USA: “Ari wc, se BRA: Goi, Mgr, Spo; (probably southern)“ [=Cochise URU County: Ramsey Canyon, 15 13 squiresi (Chaudoir, 1872) Helluomorpha s AM: se BRA: Msg; Rdj; Pna, miles south Sierra Vista sn. Spo; BOL Bousquet 2012]; Tex; n MEX: Chh, Dur, Son ritae group (sn. Reichardt 1974): 8 praeustus (Dejean, 1825) Helluo n AM: e, s USA 14 io Reichardt, 1974 s AM: n BRA: Ama: Manaus ssp. praeustus s. str. “ľAmérique septentrionale“ [=e 15 ritae Reichardt, 1974 s AM: wc BRA: Goiás: Río USA]: Aba, Mis, Nje; ?Flo Verde ssp. bicolor (Harris, 1828) Zuphium ne USA: n Mas: vicinity of Salem [=Essex County]; Ark, unicolor group (sn. Reichardt 1974): Con, Cor, DiC, Gor, Ili, Ind, unicolor subgroup: Iow, Kan, Mar, Mas, Mic, Min, 16 brunneus (Putzeys, 1846) Helluo s AM: “Brésil“ [=se BRA]: Mso, Nda, Neb, Nhm, Nje, Bhi, Rdj, Sac, Spo Nyo, Ohi, Okl, Pen, Sda, Ten, 17 gigantops Reichardt, 1974 s AM: se BRA: Spo: Barueri; Vir, Wis, Wvi, Wyo; ?Aba BRA: Bhi: Candeúba; Goi: =laticornis Dejean, 1831 Helluo “ľAmérique septentrionale“ [=e Jataí; Mgr: Cáceres; Msg: USA] Passa Quatro =pubescens Klug, 1834 Helluo type locality not stated 18 oculeus (Bates, 1871) Helluomorpha s AM: “Ega, Amazons“ [=n ssp. floridanus Ball, 1956 se USA: Flo: Volusia Co[unty]: BRA: Ama: Tefé]; Amp, Pra Ormond =janus Bates, 1871 Helluomorpha “Ega“ [=n BRA: Ama: Tefé]

332 333 19 unicolor (Brullé, 1838) Helluomorpha s AM: “provincie de Yungas, =tripustulata [=Dejean in litt.] “Pondichéri; monts Bolivia“ [BOL: La Paz Dept.]; Guérin-Menéville, 1843 Helluo Neelgheries“ [=se IND: Pud, BRA: Ama; ECU, PER Tmd] balli species: =quadripustulatus Schmidt-Goebel, 1846 “Birma“ [=MYA] 20 balli Reichardt, 1974 c AM: s MEX: Oax: 34,5 mi n =infuscatus Bates, 1892 Acanthogenius Bhamò [=n MYA: s Kachin Pochutla, route 175; 4,700 ft; state]; ne IND: Ass 22,4 mi n Puerto Escondido; 3 bicolor Andrewes, 1920 s AS: “Bombay: Belgaum 2100ft district, Kaoeqaoe“ [=w IND: glabratus subgroup: Krt]; Mhy 21 diana Reichardt, 1974 s AM: c PER: Jauja: Satipo 4 binotatus Andrewes, 1931 se AS: Sumatra [Is.]: Tapanoeli, 22 glabratus (Bates, 1871) Helluomorpha s AM: “Pebas, Upper Groot Mendeling [=w IDN: Amazons“ [=ne BRA: Pui]; Sumatera Utara Prov.: Tapanuli Ama, Pra; BOL, COL, FRG, Selatan]; Manna [Riv. =s PER Bengkulu Prov.] =gagatina Chaudoir, 1872 Helluomorpha “Cayenne“ [=FRG] 5 chaudoiri Andrewes, 1919 s, se AS: BAN, CAB, CHI: 23 juno Reichardt, 1974 s AM: BRA: Amp: Serra do nn for M. trimaculatus Chaudoir, 1872 Gad; w IDN: Sumatra Is.; Navijo e IND: Ben, Jhr, Ori; LAO, MYA, PHI, VIE species incertae sedis: =trimaculatus Chaudoir, 1872 Acanthogenius “Deccan“ [=BAN, e IND] 24 agathyrsus (Buquet, 1835) Helluo s AM: “ľintérieur de la JH of Olivier, 1790 Guyane“ [=FRG] =ruficollis Heller, 1923 “Luzon [Is.], Laguna Province, Los Baños“ [=PHI: Calabarzon M aCROCHEILUSHope, 1838; T: Macrocheilus bensoni Hope, 1838 Reg.] (species: 58) 6 cheni Zhao & Tian, 2010 AS: s CHI: Gax: Tengxian =Apistomus [Schönherr in litt.] sn. Dejean 1831 7 deuvie[sic!] Zhao & Tian, 2012 se AS: PHI [Central Visayas =Macrochilus Hope, 1838 in error Prov.:] Bohol [Is.] =Acanthogenius Reiche, 1843; T: Helluo impictus Wiedemann, 1823 8 fuscipennis Zhao & Tian, 2010 AS: s CHI: Gax: Napo = Macrocheilidius Jeannel, 1949 (Sg); T: Macrocheilus ferruginipes Fairmaire, 1892b 9 gigas Zhao & Tian, 2010 AS: se CHI: Gad: Zhanjiang 10 immanis Andrewes, 1920 s AS: “Burma: Taung-ngu“ [=s Palearctic / Oriental species ungrouped: MYA: Bago Reg.: Toungoo] 1 asteriscus (White, 1844) Helluo e, s, se AS: se CHI: Hgk; Hai, 11 impictus (Wiedemann, 1823) Helluo s, se AS: “Bengalia“ [=BAN, Gad; w IDN: Java Is.; ne IND: e, n IND]: Ben, Jhr, Ker, Krt, Ass, Tmd; LAO, MYA; c VIE Hmp, Mhp, Mhr, Ori, Pud, =crucifer Redtenbacher, 1867 Planetes Hgk Tmd; w IDN: Java Is. 2 bensoni Hope, 1838 s, se AS: “Madras“ [=se IND: 12 lindemannae Jedlička, 1963b s AS: “West Pakistan: Umg. Tmd: Chennai]; Ass, Pud; s, se Rawalpindi“ [ne PAK: Punjab CHI: Fuj, Jia, Gad, Gax, Gui, Reg.: Rawalpindi District] Hai, Hgk, Yun; LAO, MYA, 13 macromaculatus Louwerens, 1949 se AS: nc Java Is.: Gunung SRL, VIE Muria, Tjolo; 700-1000m [=w =trimaculatus Olivier, 1790 Carabus type locality not stated IDN: n Jawa Tengah Prov.] =quadrimaculata Guérin-Menéville, 1840 Helluo “Indes orientales: plateau des 15 niger Andrewes, 1920 s AS: “Nilgiri Hills: Neelgheries“ [=se IND: Tmd] Ouchterlony Valley; 3000- 3500´; Malabar, Coimbatore;

334 335 Madras: Vizagapatam [=se 26 ferruginipes Fairmaire, 1892b e AF: n, ne MAD: “Diego IND: Tmd]; Bengal: Giridih Suarez“ [=Antsiranana Prov.]; [=Jhr]; Bengal: Pusa [=Bih]; Toamasina Prov. Bombay: N. Kanara, Castle 27 madagascariensis Basilewsky, 1953b e AF: MAD Rock; Central Provinces: 28 perrieri Fairmaire, 1899b e AF: MAD Nagpur; 1000´ [=Mhr]; United ssp. perrieri s. str. MAD: “Vallée de la Betsiboka“ Provinces: W. Dehra Dun, [=n MAD: Mahajanga Prov.] Jhajra“ [=Utk]; Adh, Ker, Krt, ssp. unicolor Alluaud, 1935 s MAD: “Behara“ [=Toliara Ori, Utp; SRL Prov.] 16 nigrotibialis Heller, 1900 se AS: “Celebes septentrionalis, inter pagum Afrotropical species ungrouped: Panot et Matinang montes“ [=n 29 allardi Basilewsky, 1957 s AF: ?MOS Sulawesi Is.: c IDN: Sulawesi 30 alluaudi [Liebke in litt.] Burgeon, 1935a c, w AF: “Tolo“ [=w DOC: Tengah Prov.: Matinan env.]; e Kasai-Occidental Prov.]; MAL: Sabah; w IDN: Sumatra Bandundu, Bas Congo Is. Provinces 17 parvimaculatus Zhao & Tian, 2010 AS: s CHI: Gax: Liuzhou: 31 angustatus Basilewsky, 1949 c AF: “Lualaba: Kapanga“ Luzhai [=s, se DOC: Katanga 18 quadratus Zhao & Tian, 2010 AS: s CHI: Gax: Cangwu Prov.]; Orientale, Nord-Kivu 19 saulcyi (Chevrolat, 1854) Helluo sw AS: “Saïda“ [=LEB: Saida Provinces; RWA (Sidon)]; ISR, SYR, TUR, 32 basilewskyi Serrano, 2000 sw AF: nw ANG: Luanda UAE? [Prov.]; Lucira [Namibe Prov.]; 20 sinuatilabris Zhao & Tian, 2010 AS: s CHI: Gax: c NAB: Khomas, Otjozondjupa Fenghuangcheng Regions 21 solidipalpis Zhao & Tian, 2010 AS: s CHI: Gax: Dibei 33 biguttatus (Gory, 1832) Helluo w AF: SEN; ETI, GUI, DOC: 22 tripustulatus (Dejean, 1825) Helluo se AS: w IDN: Java [Is.]; MYA Bas-Congo, Katanga, Orientale 23 vanharteni Felix & Muilwijk, 2007 sw AS: YEM: Seyun, in the Provinces; ZIM Wadi Hadramauth; 15°57‘N, 34 bimaculatus (Dejean, 1831) Helluo w AF: SEN; ANG; e DOC: 48°46‘E; 630m; Al Kowd, near Katanga, Orientale Provinces; Zinjibar; 13°05‘N, 45°22‘E; ETI: Oromiya Reg.; GBI; 20m KEN: Coast Prov. 24 vitalisi Andrewes, 1920 se AS: s CHI; “E Borneo: =bisignatus Reiche, 1843 Acanthogenius Moorjawa, Sanga Sanga NN for Macrocheilus bimaculatus Dejean, 1831 [=c IDN: Kalimantan Timur 35 biplagiatus (Boheman, 1848) Acanthogenius s AF: “Caffraria interior“ [=n Prov.]; ne LAO: Ban Houei RSA, ZAM, ZIM]; ANG: [=Xiangkhoang Prov.]; Tonkin: Lunda Norte Prov.; BUR, Hoabinh“ [Prov. =n VIE] DOC: Kasai-Oriental, Katanga, Orientale Provinces; SOM ferruginipes species group (sn. Basilewsky 1960): =cordicollis Müller, 1949 “Aethiopia meridionalis: 25 cribrarius Fairmaire, 1901 e AF: c MAD: [Antananarivo Caschei“ [s SOM: Jubbada Prov.: Andrangoloaka sn. Dhexe Reg.: Caschei] Jeannel 1949]; Fianarantsoa 36 burgeoni Basilewsky, 1967 c AF: “Congo: Kibali-Ituri: Prov. Moto; Niarembe“ [=ne DOC:

336 337 Orientale Prov.]; Katanga, Nord 48 ocellatus Basilewsky, 1953a c AF: “Congo Belge: Lualaba: Kivu, Sud Kivu Provinces; Parc. Nat. Upemba“ [=se DOC: RWA; sw TAN: Mbeya Reg. Katanga Prov.] 37 clasispilus Basilewsky, 1967 w AF: “Zaïre: Poko-Dili“ 49 overlaeti Burgeon, 1935a c AF: “Congo belge: Kapanga“ [=Orientale Prov.]; SEN [=se DOC: Katanga Prov.]; 38 crampeli Alluaud, 1916 c AF: “Congo français: Fort- Orientale Prov. Crampel, sur le haut Chari“ [=c 50 persimilis Basilewsky, 1970 w AF: RCI: Lamto; MLI: RCA: Kaga Bandoro Pref.]; n Bamako; RCA: Damraou; GUI, DOC: Équateur Prov. SEN 39 cruciatus (Marc, 1840) Helluo w AF: SEN; GUI, LIR, RCI 51 proximus Péringuey, 1896 s AF: “Natal (D‘Urban)“ [=se =royi Basilewsky, 1954 Nimba Mt.; 1200m [=GUI, RSA: KwaZulu-Natal Prov.] LIR, RCI] 52 quadrinotatus Burgeon, 1935a c AF: “Congo belge: Kapanga“ 40 diplospilus Basilewsky, 1967 c AF: “Congo [=DOC]: [=se DOC: Katanga Prov.] Bas-Congo [Prov.]: Mayidi; 53 scapularis Reiche, 1843 w AF: “Senegalia“ [=GAM, Stanleyville: Yangambi“ GBI, SEN]; n ETI: Tigray [=Orientale Prov.] Reg.; ne DOC: Orientale Prov. 41 dorsalis (Klug, 1834) Helluo s AS: “Kap“ [=s RSA: Western =boopis Roth, 1851 Polystichus “ Tigré in N. Abyssinien“ [=n Cape Prov.]; ETI: Tigray Reg.] 42 dorsiger (Chaudoir, 1876) Acanthogenius e, w AF: “Adowa“ [=n =fenestratus Raffray, 1885 “plateaux de ľHamacen“ [=n ETI: Tigray Reg.]; ne DOC: ETI: Tigray Reg.] Orientale Prov.; GBI 54 spectandus Péringuey, 1904 s AF: “Southern Rhodesia 43 elegantulus Burgeon, 1935a c AF: “Congo belge: Yebo; (Umtali)“ [=e ZIM: Beni; Watsa [=DOC: Orientale Manicaland Prov.: Mutare] Prov.]; riv. Kasai [=Kasai- 55 taedatus Basilewsky, 1960 c AF: “Congo Belge: Occidental Prov.]; Lusambo Kibali-Ituri: Bavungura; [=Kasai-Oriental Prov.]; Adranga; Arara, terr. Mahagi; Kaniama; Kapanga [=Katanga Dingila; Ibembo; Luma prės Prov.]; vieux Kasongo Djalasiga; Niarembe; Nioka; [=Maniema Prov.] région ďAbok; Uele Distr.; 44 hybridus Péringuey, 1896 s AF: “Natal (Escourt)“ [=se Stanleyville: Stanleyville RSA: KwaZulu-Natal Prov.]; [=ne DOC: Orientale Prov.]; ZIM Tschuapa: Boende [=Équateur 45 labrosus (Dejean, 1831) Helluo w AF: SLN; CAM, CON, n Prov.]; Sankuru: Komi DOC: Équateur, Orientale [=Kasai-Oriental Prov.]; Haut- Provinces; GBI, GUI, NRE, Katanga: Kundelungu; Manika; RCA, SEN Katanga; Lualaba: Parc. Nat. =dispar LaFerté-Sénectere, 1849 Helluo “ Sénegambie portuguaise“ Upemba, Lusinga; Kapanga; [=GBI] Kapiri [=Katanga Prov.]; =bimaculatus Murray, 1857 “Old Calabar“ [=s NRE] Ruanda: Gabiro; Rwinkwavu 46 longicollis Péringuey, 1904 s AF: “Southern Rhodesia [=RWA]; Tanganyika Territory: (Umtali)“ [=e ZIM: Malagarassi [=w TAN: Kigoma Manicaland Prov.: Mutare] Reg.]; Mara [Reg.]; Morogoro 47 moraisi Serrano, 2000 w AF: GBI: Quebo (Coli); [Reg.]; Uvinsa [=Rukwa Reg.]; CAM [Centre Prov.:] Yaoundé

338 339 Kigonsera [=Ruvuma Reg.]; MELADROMA Motschulsky, 1855; T: Anthia umbraculata Fabricius, 1801 UGA: Kampala-Kigoma; (species: 4) Masindi; Msisi; Mujenje; KEN: Rift Valley Prov. 1 angustata Basilewsky, 1970 c AF: DOC 56 varians Péringuey, 1904 s AF: “Southern Rhodesia 2 informicollis Liebke, 1928a w AF: “Joko, Kamerun“ [=c (Umtali)“ [=e ZIM: CAM: Central Prov.]; RCA, ne Manicaland Prov.: Mutare]; se DOC: Orientale Prov. DOC: Katanga Prov.; sw TAN: 3 katangensis Burgeon, 1935a c AF: “Congo belge: Mbeya, Rukwa Regions Elisabethville; Sandoa; =eugrammus Basilewsky, 1960 U.N. “Southern Rhodesia: Umtali Kapanga; Kafakumba; 18 miles [=e ZIM: Manicaland Prov.: sw Elisabethville“ [=se DOC: Mutare]; Sebakwe“ [Midlands Katanga Prov.]; Maniema Prov.] Prov.; ANG: Lunda Norte Prov. 57 viduatus Péringuey, 1899 c, s, w AF: ANG, DOC, KEN, 4 umbraculata (Fabricius, 1801) Anthia e, w AF: “Guinea“ [=Gulf of MOS, RSA, TAN: , ZAM, ZIM Guinea coast:] BEN, CAM, ssp. viduatus s. str. “Transvaal“ [=ne RSA: GAB, GAM, GBI, GHA, GUI, Gauteng, Limpopo, LIR, NRE, RCA, RCI, SEN, Mpumalanga, Nort West SLN, TOG, ?ANG: Huila Provinces]; ANG: Cunene Prov.; ne DOC: Orientale Prov.; Prov.; KEN: Coast Prov.?; MAW, TAN: Dar Es Saalam, MOS: Manica?, Niassa?, Tete? Lindi, Ruvuma Regions Provinces, TAN: Ruvuma =grandis Dejean, 1831 Helluo SEN Reg.?; ZAM, ZIM =opacus La Ferté-Sénectère, 1849 Helluo “Sénegambie portuguaise“ ssp. katanganus Basilewsky, 1960 “Congo-Belge: Elisabethville; [=GBI] Mufungwa-Sampwe; Parc. =brevipennis Liebke, 1928b “Joko, Kamerun“ [=c CAM: Nat. Upemba, Kaswabilenga; Central Prov.] 700m; Kabwe sur Muye; 1320m; Lulua, riv. Lunene; OMPHRA[Leach in litt.] Dejean, 1825; T: Helluo hirtus Fabricius, 1801 (species: 6) Sandoa; Albertville; Bassin de la Lukuga [=se DOC: Katanga 1 atrata (Klug, 1834) Helluo s AS: “Bengalen“ [=BAN, e Prov.]; N. Rhodesia: Abercorn“ IND]: Ass, Ben, Ker, Mhp, [=ne ZAM: Mbala District: Tmd Mbala] 2 complanata Reiche, 1843 s AS: “India orientali in 58 vinctus Basilewsky, 1960 c AF: “Oubangui-Chari: provincia Decan“ [=e IND]: Fort-Sibut“ [=RCA: Kémo Hmp; MYA, NEP Prefecture]; ANG: Cabinda, 3 hirta (Fabricius, 1801) Helluo s AS: “Indes Orientales“ Uige, Zaïre Provinces; CAM, [=BAN, e IND, MYA]: Ben, CON, GAB, GUI, ne DOC: Bih, Jhr, Hmp, Krt, Mhr, Ori, Bandundu, Bas-Congo, Pud, Tmd; SRL Équateur, Kasai-Occidental, =tristis [Leach in litt.] Dejean, 1825 “Indes Orientales“ [=BAN, e Kasai-Oriental, Kinshasa IND, MYA] Provinces; TOG =brevis Chaudoir, 1850 “Nord de ľHindostan (Simlâh)“ [=n IND: Hmp]

340 341 4 pilosa Klug, 1834 s AS: “India Or.“; SRL Zambezia [=MOS, ZAM, 5 rotundicollis Chaudoir, 1872 s AS: “Indes Orientales“ ZIM]; s DOC: Katanga Prov. [=BAN, IND, MYA]; SRL 3 congoensis Basilewsky, 1959b “Lukuga; Pweto-Kilwa- 6 rufipes (Klug, 1834) Helluo s AS: “India Or.“ [=SRL] Kasenge“ [=s DOC: Katanga Prov.] PLEURACANTHUS Gray, 1832; T: Pleuracanthus sulcipennis Gray, 1832 4 corpulentus Chaudoir, 1877 s AF: “République de (species: 4) Transwaal“ [=n RSA: Gauteng, =Ocypus Gistel, 1838 JH of Stephens Limpopo, Mpumalanga, North West Provinces]; Free State inca group (sn. Reichardt 1974): Prov.; n NAB: Ohangwena, 1 inca Reichhardt, 1974 s AM: sc PER: Cuzco: Omusati, Oshana, Oshikoto, Quincemil; 750m Otjozondjupa Regions; ZIM 2 psittacus Reichhardt, 1974 s AM: se BRA: Spo: 5 denticulatus Basilewsky, 1959b ANG: “Malange“ [=Malanje Descalvado; Rio Mogi; Guaçu; Prov.] Escamaruça; Msg 6 ferox (Erichson, 1843) Helluo sw AF: ANG sulcipennis group (sn. Reichardt 1974): 7 gerstaeckeri (Chaudoir, 1877) Meladroma s AF: “Afrique australe“ 3 sulcipennis Gray, 1832 s AM: wc, se BRA: Goi, Mgr, [=RSA] Pna, Rgn, Spo 8 kochi Basilewsky, 1964a e AF: “Environs de Balad, 2 4 tridens Reichhardt, 1974 s AM: cw BRA: Goi: Jataí, Km Sud de ľUebi-Scebeli; Fazenda Aceiro; n PAR: Gesira“ [=s SOM: Middle Concepción [Dept.]: near Shebelle Reg.] Horqueta 9 lugubrinus (Boheman, 1860) Polyhirma sw AF: “in regione fluvii Svakop“ [=c NAB: Erongo TRIAENOGENIUS Chaudoir, 1877; T: Acanthogenius sculpturatus Prov.: Swakop River]; Karas Gerstäcker, 1867 (species: 11) Reg.; ANG: Benguela Prov.; n RSA: Northern Cape Prov. TRIAENOGENIUS s. str. (10 species) =anthioides Chaudoir, 1872 Acanthogenius “Benguéla“ [=w ANG] 1 arabicus Gestro, 1889 sw AS: “Yemen meridionale: 10 sculpturatus (Gerstaecker, 1867) Acanthogenius e AF: KEN, SOM, ne TAN Tes“ [=sw YEM: Yemeni ssp. sculpturatus s. str. “ad lacum Jipe“ [=ne TAN: Highlands: Ta‘izz]; ne AF: DJI, Kilimanjaro Reg.]; Dar Es ETI, SOM Saalam, Mwanza, Tabora, =obockianus Fairmaire, 1892a Triaenogenius “Obock“ [=DJI: Obok] Zanzibar Isl. Regions =angustipennis Gestro, 1895 Meladroma “ Archeisa (Ogaden)“ [=se ETI: =helluonoides Ancey, 1881 Anthia “Uzagara“ [=nw TAN: Mwanza Somali Reg.] Reg.] 2 carinulatus (Fairmaire, 1887) Acantgogenius e, s, sw AF: MOS, NAB, RSA, ssp. infuscatus Basilewsky, 1943 “Afrique orientale“ [=w TAN: TAN, ZAM, ZIM Rukwa Reg.] ssp. carinulatus s. str. “Uzagara“ [=nw TAN: Mwanza ssp. patrizii Alluaud, 1926 “Afrique orientale anglaise: Reg.] Archer‘s Post“ [=KEN: Eastern ssp. agnatus (Péringuey, 1896) Meladroma “Ovampoland (Okavango Prov.] River)“ [=n NAB: Okavango ssp. rhodesianus Basilewsky, 1959b “N. W. Rhodesia: Ubemba, Reg.] près ďAbercorn, à ľextremité ssp. vicinus Péringuey, 1896 “Transvaal (Middleburg) [=e Sud du Lac Tanganika“ [=ne RSA: Mpumalanga Prov.]; ZAM: Northern Prov.: Mbala]

342 343 ssp. subcostatus Müller, 1943 “Somalia regione benadirense: P ogonoglossusChaudoir, 1862; T: Pogonoglossus validicornis Villaggio Duca degli Abruzzi“ Chaudoir, 1862 (35 species) [=se SOM: Shabeellaha Dhexe =Libresthis Schmidt-Goebel, 1846; T: Libresthis truncatus Schmidt-Goebel, 1846 Reg.] =Carpaulum Sloane, 1904; T: Carpaulum inflaticeps Sloane, 1904

ANDROCHAETUS Basilewsky, 1959b; T: Acanthogenius lugubris Schaum, 1863 (1 Oriental species group: species) 1 augustae Louwerens, 1949 se AS: e Java [Is]: Gunung Pandan (Res. Kediri) [=w IDN: 11 lugubris (Schaum,1863) Acanthognius s AF: BOW, MOS, NAB: Jawa Timur Prov. ] nn for Helluo grandis Boheman, 1848 Erongo, Khomas, Ohangwena, 2 carinipennis Bates, 1892 s AS: “Karin Chebá, alt. 900- Omusati, Oshana, Oshikoto 1100m“ [=se MYA: Kayah, Regions; RSA, TAN, ZAM, Kayin, Shan States: Karen Hills] ZIM 3 chaudoiri Gestro, 1875 se AS: “Cambodge“ [=CAB] =grandis Boheman, 1848 Helluo JH of Dejean, 1831 “Africa austr. inter. (Sveakop)“ 4 intermedius Bouchard, 1903 se AS: se Sumatra Is. “Forêt [=c NAB: Erongo Prov.: de Nirou“ [=w IDN: Sumatera Swakop River] Selatan Prov.] 5 latus Andrewes, 1937b se AS: se Sumatra Is. “Forêt de Nirou“ [=w IDN: Sumatera PHYSOCROTAPHINI Chaudoir, 1863 (genera: 5, species: 43) Selatan Prov.] =Helluodini Andrewes, 1929a 6 montanus Bouchard, 1903 se AS: se Sumatra Is.: Paggar Alam [=w IDN: Sumatera HELLUODES Westwood, 1846; T: Helluodes taprobanae Westwood, 1846 Selatan Prov.] (species: 3) 7 physoides Andrewes, 1937b se AS: Java [Is. =w IDN]: “Res. Preanger, Mt. 1 devagiriensis Sabu, Abhitha & Zhao, 2008 s AS: sw IND: Western Ghats: Tongkoeban Prahoe“; 4000- w sl. 5000ft [=West Java Prov.: 2 taprobanae Westwood, 1846 s AS: “Insulâ Taprobanæ“ Priangan] [=SRL] 8 sciurus Andrewes, 1937b se AS: Java [Is. =w IDN]: 3 westwoodi Chaudoir, 1869b s AS: “Decan“ [=BAN, e IND] 9 sumatrensis Gestro, 1875 se AS: Sumatra [Is. =w IDN]: Sumatera Selatan Prov.; Borneo HOLOPONERUSFairmaire, 1883 NN for Pamponerus Fairmaire, 1881; T: Is.: MAL: Sabah Pamponerus godeffroyi Fairmaire, 1881 (species: 1) 10 sylvaticus Bouchard, 1903 se AS: se Sumatra Is. “Forêt =Pamponerus Fairmaire, 1881 JH of Loew, 1849 de Nirou“ [=w IDN: Sumatera 1 godeffroyi (Fairmaire, 1881) Pamponerus AU: “I. du duc du York“ Selatan Prov.] [=PNG: East New Britain 11 tagalus Heller, 1916 se AS: PHI: Luzon [Is]: Mt. Prov.: Duke of York Isl. (type Makiling [=Calabarzon Reg.] destroyed in WW2)] 12 torvus Andrewes, 1937a se AS: Java [Is. =w IDN]: 13 truncatus (Schmidt-Goebel, 1846) Libresthis s AS: “Birma“ [=MYA]; IDN: PHYSOCROTAPHUS Parry, 1849; T: Physocrotaphus ceylonicus Parry, Borneo Is.; IND: Ass, LAO, VIE 1849 (species: 1) 14 validicornis Chaudoir, 1862 se AS: Java, [Is. =w IDN]; se Borneo, Sumatra Isl.; ne IND: 1 ceylonicus Parry, 1849 s AS: “Ceylon“ [=SRL] Ass; LAO; MYA; VIE

344 345 Australian species group: Jayapura Dist.]; Njau-limon, 15 danielsi Baehr, 2007 AU: se AUS: ec Nsw: 8 S. of Mt. Bougainville“, 300ft km S Mendooran; 31°53´S, [=Jayawijaya Dist.] 149°03´E 26 missai Baehr, 2005b AU: e New Guinea Is.: PNG: 16 giganteus Baehr, 2005b AU: e New Guinea Is.: PNG: Canopy mission, Madang Canopy mission, Madang Prov.: Baiteta province, Baiteta 27 obliquus Darlington, 1968 AU: c New Guinea Is.: “N-E. 17 glabricollis van Emden, 1937 AU: New Guinea Is.: “N-E. N. G., Eliptamin Vy.“; 1665- N. G.“ [=e PNG]: Eastern 2530m [=w PNG: Sandaun Highlands, Central Provinces; Prov.] West Papua (IDN): Manokwari, 28 papua Darlington, 1968 AU: New Guinea Is.: “Papua: Nabire Districts Dobodura [=e PNG: Oro 18 grossulus Darlington, 1968 AU: e New Guinea Is.: “vic. Prov.]; N-E. N. G., Lae; sea Zengaru, Vy. of Kua R., level; Busu R., E of Lae; Mongi Watershed, Huon pen.; 100m [=Morobe Prov.]; Neth. 800m; Lae [=ne PNG: Morobe N. G.; West N. G.: Hollandia, Prov.]; Papua: Kokoda, 1200ft; Sentani; 90m; Hol Maffin near [=Central Prov.]; Eastern Sarmi “[=West Papua (IDN): Highlands Prov. Jayapura Dist.] 19 horni Sloane, 1907 AU: “New Pomerania, Gazelle 29 parvus Darlington, 1968 AU: New Guinea Is.: “West Peninsula“ [=New Britain Is.: N. G.: vic. Hollandia [=West PNG: ne New Britain Prov.] Papua (IDN): Jayapura Dist.]; 20 inarmatus Baehr, 1989 AU: ne AUS: ne Qld: Cape Papua: Dobodura “[=e PNG: York peninsula Oro Prov.]; Morobe Prov.; ne 21 inflaticeps (Sloane, 1904) Carpaulum AU: e AUS: e Qld: Cairns AUS: n Qld: Cape York Pen. District: Kuranda 30 porosus (Sloane, 1904) Carpaulum AU: e AUS: e Qld: Kuranda; n 22 laevissimus Baehr, 1995 AU: w New Guinea Is.: “Irian Nte, n Wau Jaya [=West Papua (IDN)]: 31 punctatulus Louwerens, 1969 AU: PNG: New Ireland Prov.: Panai-Pr.[=Paniai Dist.], Bismarck Archipelago: Dyaul Nabire, Pemukiman“; 200m Is.: Sumura 23 latior Darlington, 1968 AU: w New Guinea Is.: “West 32 rufopiceus Baehr, 1993 AU: nc AUS: “N. T.“ [=Nte:] N. G.: Hollandia: Lower Mist Katherine, (ex cave) Camp, Snow Mts.; 1550m“ 33 schaumi Chaudoir, 1869b AU: “Mysol“ [=e IDN: W [=West Papua (IDN): Paniai Papua Prov.: Raja Ampat Isl: Dist.: Maoke Mts.] Misool Is.] 24 major Darlington, 1968 AU: e New Guinea Is.: “N-E. 34 taylori Darlington, 1968 AU: New Guinea Is.: “N-E. N. G: vic. Nadzab“ [=ne PNG: N. G.: Aiyura; 1900m; Okapa Morobe Prov.] [=ne PNG: Eastern Highlands 25 minor Darlington, 1968 AU: e New Guinea Is.: “N-E. Prov.]; Eliptamin Vy.“; 1665- N. G.: lower Busu R., Huon 2530m [=Sandaun Prov.]; West Pen.; Wau, 1200m [=ne PNG: Papua (IDN): e Manokwari Morobe Prov.]; West N. G.: Dist. Hollandia [=West Papua (IDN):

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The Annals and Magazine of Natural History (Series 9)18: 281. “Arfakgebg., Siwi “; 800m An d r e w e s H. E. 1929a: Coleoptera, Carabidae. Vol. I. Carabinae. In: The Fauna of British India, including [=West Papua (IDN): Ceylon and Burma. London: Taylor & Francis, xvii + 431 pp. + 1 pl. Manokwari Dist.: Arfak Mts.] An d r e w e s H. E. 1929b: Fauna Sumatrensis. Carabidae. Tijdschrift Voor Entomologie 72: 303-340. 2 drumonti Baehr, 2004 AU: e New Guinea Is.: PNG: An d r e w e s H. E. 1930: [Pp.: 31-32, 120, 136-147, 178, 206-208, 236-237, 282-283] In: Catalogue of Indian Canopy mission, Madang Insects. Part 18 -Carabidae. Calcutta: Government of India, Central Publication Branch, xxii + 389 pp. An d r e w e s H. E. 1931: Some keys to Sumatran Carabidae, together with descriptions of further new species. Prov.: Baiteta Zoologische Mededelingen. Uitgegeven door het Rijksmuseum van Natuurlijke Historie te Leiden 14: 54- 3 monstrosa Baehr, 2004 AU: w New Guinea Is.: West 78. Papua (IDN): Fakfak Dist..: An d r e w e s H. E. 1937a: On some new species of Carabidae, chiefly from Java (III). Treubia 16(1): 25-35. Mambuni-Buni An d r e w e s H. E. 1937b: On the species of Pogonoglossus found in Java and Sumatra (Col. Carabidae). Bulletin de la Société entomologique de France 42: 152-156. An t i c h t c h e n k o A. 2009: Anthia (Anthia) mannerheimi afghana ssp. n. – a new subspecies from Central Acknowledgements. We are obliged to all our friends who made their collections accessible to us and for Afghanistan (Coleoptera: Carabidae). Russian Entomological Journal 18(4): 273-275. their publication outcomes, without which our catalogues could never been written. 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