1 2 DR. ALICIA TOON (Orcid ID : 0000-0002-1517-2601) 3 4 5 Article type : Invited Review 6 7 8 Insect pollination of cycads 9 10 Alicia Toon1, L. Irene Terry2, William Tang3, Gimme H. Walter1, and Lyn G. Cook1 11 12 1The University of Queensland, School of Biological Sciences, Brisbane, Qld, 4072, 13 Australia 2 14 University of Utah, School of Biological Sciences, Salt Lake City, UT 84112, USA 15 3 USDA APHIS PPQ South Florida, P.O.Box 660520, Miami, FL 33266, USA 16 17 Corresponding author: Alicia Toon 18
[email protected] Ph: +61 (0) 411954179 19 Goddard Building, The University of Queensland, School of Biological Sciences, Brisbane, 20 Qld, 4072, Australia. 21 22 23 24 25 26 27 28 29 30 Manuscript Author 31 This is the author manuscript accepted for publication and has undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: 10.1111/AEC.12925 This article is protected by copyright. All rights reserved 32 33 Acknowledgements 34 We would like to thank Dean Brookes for discussions about genetic structure in cycad 35 pollinating thrips populations. Also, thanks to Mike Crisp for discussions about plant 36 diversification and Paul Forster for information on Australian cycads. This work was funded 37 by ARC Discovery Grant DP160102806. 38 39 Abstract 40 Most cycads have intimate associations with their insect pollinators that parallel those of 41 well-known flowering plants, such as sexually-deceptive orchids and the male wasps and 42 bees they deceive.