A Taxonomic Revision of the Genera Crowea, Eriostemon and Phebalium

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SJNAJNOC SUMMARY

A taxonomic treatment is presented ofthe genera Croweq,Eriostemon, and Phebulium (Rutaceae, tribe Boronieae, subtdbe Edostemoninae), which except for one New Zealand species are endemic to Australia. These genera, and several others in the Rutaceae, were included undet Eriostemonby F.v.Mueller (1859. etc.) and by some later authors, or under Crowea (Barllon, 1873). They ire hire kipt distinct but with a modified circumscription. One speciespre- viously plactd in Eriostemon has been shown to be a Pityrodia (Yerbenaceae) and another Xo be a Boronia (Rutaceae). The following five sections are described as r.ew:.-Eriostemon sect. Cory' nonema, Gymnanihos, Nigrostipulse, and Osmsnthos; Phebalium sect. Gonio' clados. Fifteen new species and sixteen new infraspecific taxa are described;in addition two new specific combinations and seventeennew infraspecific combinations are made. ',, uoxrlJ,, lE rpou3d aq1ur flluenbesqnspeqsqqnd se .ro (t961) g 'pa '1d runJorJ?qJeHxepul ur esorll 01 puodsouoc suorllr:ltc eql ur pesn I 'pe^loûr suorlErâ.rqq"eqJ "x"t eq] Jo uorldrJJsJp aq] rJUe petrc u00q s"rl perpnls sueurrcadsrunlr€q.rorl eql Jo uor odold pus r,{1uo ,{ eruJoN 'tuoql qlr.r cgrcodsuot pe.roprsuoauoeq o^sq lsud eq1 ur pue sel"ls urals?e eql ur souo 01 pel?lar ,{losols or€ sabads uzllertrsny urolsal& ,reJ " q8noqll€ uorlnqulsrp l"lueurluoc-sueJle s"q EIJUJBJJlql Jql ur sJrJedsaLlt Jo ouoN 'sequl lueraJrp ur r:aua3 lueJaJrp 01 3uo[3q lceJ ur [\q uowaisot,lg ur pepnlcur uaeq el"q €ruopaFJ ,\\eN pu? ecr4v qlnos luo{ paqrJcsep sercadsIereAeS asuos lculs aql tlr e"eruoJog eql Jo sreqrueutIIa o1 serldde ,(lqrqord srr1lpue (puepeT ,tr.eNur r&r{rga?d Jo serceds ouo ruo{ gede) r:rueurstl pu1?"rlsl}snv ol crru3pueel€ EJeueBeelql IIV 'r;:eo.rddesrqr lu€rru,r ol uorlrunlslp Ier6oloqdroru 1rlotJSns,\\oqso1 reedde,{eql se ?reue8 el€Jedes s" luerll SurureleJ lue I sJelJ?r?qJlueroJrp uo slsruuloq ,{q pereld srs?qdruoaql ol SofproJaepeue^ s€q serceds,(urru3ouorlrsod eq1tun11or1at14pue oaowalso!.tg'Da$o,tJ "Jeue8 eql urqlrl& ,{ul,{rloufs 8rn^Juelc pu" sdrrlsuorl?lel lno Sunurod puo,4aqueql ssncsrp ol ,{r"ssacaull lleJ lou el?q I uos"al slql JoJ pu? 'pol?cJ?urep ,{F€elc are faql qJrq,u ,(q sralcereqc e^"rl 'ereq peJeprs -uoc eaJql eql ol pal"ler,{lesolc q8norllle '€Jaue8Jer{lo as3qJ eseruorog eqtr 'luowalsotlg JO SJaqrrletrtJ3q1O l€re^3s Jo uorsrllcur eql qlr.r seurleuros Jepvr. ,(ll?nsn)cueuoStloc paroprsuoc ue3q e^"q se^lasueql"Jaue8 eql -ro'JJq]ouEol "Jeue8 erll euo uro{ peJJeJsuerlueoq ^tluonbe:.1e,tzq sercedspepnlcur erJJ ',{Jntuec Jo ql6I ,(lJeepuB rl18I el€I eql ur,{reôcsrp Jrerll ecursuoq"rcosse .rroq}Jo esnef,aqJoqloSol pal€ajl JJ?:eded srql ur peqdz.rSouour€.reua8 oalq] eql

NOIJJNOO'I{JNI HISTORICAL SURVEY

Crowea i[ This -relevantse[us was describedas a member of the Rutaceaeby J. E. Smith 1798, the specimensbeing receivedfrom John White, -M.D. of Port Jackiol and raisedfrom seedin l790. It was not the custom of Smith to give monotypic genera (" a specific diff'erencefor sDecific epilhets when describrog " a solitarv speciesbeilA e!idently a most glaring absurdity (1808))' it was therefore leit to H. A;drews (1800) to ptovide the lirst specificepithet (C. saligna). A second species, C. ongustifolia, was described.by Smith (1808) fi oi a plent collecledin WeslernAustralia; howeverthe publicationwas passed tunnoticedby botlnists and the samespecies was describedas new in 1849by Turczaninow who used the same specific epithet. In 1855F. Mueller describeda third species,C. exqlqta"from a Victorian olant. and in 1863 G. Bentham described a fourth from a plant collected in kestern Australia. Other names have been publishedbased on matedal ir cultivation, but these are all forms of the New South Wales plant, C. sctlignq- In 1862 Mneller m ade Crowea a seclion ofthe genusEriostenon and \Liled the two eastern states speciesunder the \ame E crowei, This generic union was followed by H. E.-Baitlon (p. 387, 1873)but subsequentlyin the same volume (o. 463j he united Eriostinon and Phebqlium under Crouea. Apart from differencesofopinion among taxonomistsas to whetherone ol two species should be recosnise.din both eastand west Austrllia there ha\e been no sub- seque[t new moves involving the taxonomy of the genus

Eriostemon This genus was describedby J. E. Smith in 1798 as a meruber of the Rutaceae. He appearsto have had two speciesin mind at the time, one of ( Diosmq uitiflora') came from South Africa. However, for the origin of which ", the genus he gives'" Australasia a name which has not- appareltly b-een takei to include South Africa. C. H. Persoon named and described the first two speciesin 1805, an Australian plant rvhich he called E. austrulasium, and a Soutlr African planl E. capense (a nomerr novunr fot- Diosnw unifora L.). Smith (1809)laier includedsix speciesin the genus:ofthese. two.are of South African origin and are norv ilcluded in the genusAdenandra, one (E patadoxa) is now plactd in the geous Boronia, and another (-E.salsoliJblia) in the genus Philothica. A. Jussieu(1825) limited the genusto the Australian plants and in his circumscription,which is acceptedin this paper,he tecognizedsix species (two then unpublished). In 1859 Mueller included iu Eriostemon the gerera Phebalium, Microcybe, Asterclasiq, atd Geleznowia,and subsequently (1862) Chorilaena and Crowea. Baillon (1873) similarly lumped sevetal genera \Ldet Eriostemon, but later in the same wor'k r.rnitedit witli Crowea. Sntce that time few basic charges have been made in the circumscription of these genera wlich were considered distinct by both Bentham (1863)and Engler (1896and 1931). Several soecies from New Caledonia have at one time been included in Eriostemotl; these however differ markedly in appearancefrom the Australian species and (according to published descliptions)-contain only one ovule in each carpel. They are now usually placed in either Myrtopsis or HalJbtdia, which genela should probably be excluded from the tribe Boronieae.

Phebalium This genus (based on P. squamulosum)was described by E. T. Ventenat in 1805 from mateiial collected in New South Wales. He considered the genus to be related to Baeckea a\d Leptospermum in the family Myrtaceae. J. E. Smith (1814) transferred it to the iamily Rutaceaeand recognisedthree species, 'lsru€toq JJL]lo^ue i(q pe$olloJ u$q lou seq uorlcs s\rll $q oatfo.tD snurS eqt ar unu7qaqd pareld uopeg '(I€6t igggl) pue 96g l ) ralEug ,{q pr,rolloJ se,t uoqducirunirrc' sr\j .:pu joqaqTnj 'lJes 'l3as sel unuoqaqd ptle (stsdouaD\pqJ 1r ur Supnlcut) Dl,uauotaTsljol4ces s.rallantrtr 01 tr petrlu{ pu? snua8" s? ,rnlpqaqd peurc]€t (EggI) ru?qfueg .g 'unupqaqd Jo uorlJrs ls.rgsrq ur nerssnf,{q papnlcur sarcedso,trl aql peceld (7991) relel aq.(.aoLuatsoLtI uorlJJs E s?) qcrqa otur DlsoloejsY snue8 oql pelEeJc Jo 'wnuDqaqd ,Dwauola1 peq 0q (gggI) Ja\IJEA ptJe ,slsdouaoluoqTsuorFos e.ra,r o$ql: pe^lo^ur sercadssnou"^ eql peJuld eq qrrqm. olur strorlJesleJJAes peleelc (2981) relEl pae uowatso!.tg o1 snua8 eq1 perreJsuelt (69gI) rellanN .C

,(q snueEeq] Jo lunocr-eerll qrr,, pelu*nbceuneq or prr""dd" x:lfl.:*;f '.wnawsotp .,. g s:rcads Jl8uls Jql peur?luoc erqnp ,dg,, .uorlses p.ILl] or{l :lwnsoryLuDnbs'd pve 'wnlpJlu8Daola .d ,sdazuo .Ualp.tDlllq-d ,unlptolps 'd) d selceds eAU peuleluoc uollJas puoces oql l(sndreco:n :) orsolotatsv snuai erlt ur paield,rou orc qcq,r (rarlotadoxaq.d pve w lplaoaue.4.) selcads o,\\l poureluoc uollcos lsJg eq1 :(seu"u e^I3 plp eq qcrq,lrio1).suoqces eerql 'soneds lou olur ureql Surprlrp g-paqJl.sap snue8 oq1 qdei8ouoiu e irr (eg7gf) 'V 'uowa4sottg 3o nelssnl sttua? aql u\ snaLupnbs.!rpour€tal pu? (.CC sdatub :4 puB '4.) .snue8 wnsolnLuonbs sercads o,$1 ,{Iuo pesluSoca: eqt 3o lunoccr s.qllus-Jo are,rsun ,{ltuor"ddE s€/ or{-r ,(y7gy) elopue3 ep .d .y .Clllq€.I .4 . ,tunsopurbibi . snawDnbs uoutalsotlq uo peseq) analualn pue umlbluap a a ERIOSTEMON, PHEBALIUM, AND CROWEA AND RELATIONSHIPS WITH OTHER GENERA

Eriostemon and Phebalium as delimited here exhibit much greater intra- generic morphological diversity than is found in the other genera of the Boro- nieae. and their representativesections sometimesappear to have greater amlritv with these genera that with the other sections of their own genus (e.g. Pheba'liun sect. Phihaliun to Microc),be, P. secl. Gonioclqdoslo Chorilaena, zrd Eriostemon sect. Nigrostipulae 1o Philothecq), However, I have retained the commonly accepted divisions because the morphological diverse sections are linked through intermediates (e.9. P. sect. Phebalium through sect. Gorlo- clados lo secl. Leionema), because the segregated genera possesscharacters which do clearly demarcate them, and because cytological evidence suggests that this is a natural grouping (S. Smith-White,1954). These other genera have been included by Engler in the subtribes Erio- stemoninae,Boroniinae, and Nematolepidinae. The last sub-tribewas separated on the b.rsis of the stamens being ligulate, a character which appears to seDarategenera related to each other and independently, to specresof Phebctliunt sect. Goioclados a\d sect-Eriostemoides.

Philotheca. This genus is morphogically similar to Eriostemon sect. Nrg/o- stipulae w\th which it sharesthe same type of anthers, seedand stipular excres- cencies. It is distinct in having monadelphous stamens and in chromosome number (n : 14, fide Smith-White, 1954).

Drummonilita is usually included in the folmer genus o[ account of its mona- delphy but it has flowers atrd vegetative characters-which are otheru ise quite disainct. The anthersare presenton only 5 ofthe filamentsand are dorsifixed and ilr.rrnobile(not versatile wjth a slender attachment); this type of attachment is also found in the genus Crott'ea bul there do not otherwise appear to be close morphological similarities between the two gelera.

Microcybe. This genusis obviously closely related to Phebalium sect.Phebalium with which it sharesstaminal and seedcharacters, and a lepidote indumentum. It diflers in the compact heads of flowers, i[ the free or almost free sepals,in the glabrous petals (or almost so), ald in the reduced carpel number.

Geleznowia, a monotypic Western Australian genus, has seedswhich are the same as those fou.\d ir\ Eriostemon sect. Nigrostipulae. However in staminal charactels it is different as the anthers lack a white apiculum aud in fact have a retuse apex. Its afinities are uncertail. ljrocarpus. In this genus the seed is similar to that found in Phebalium sect. Eriostcmoideswhich sectionit also resemblesin anlher morphology. I seeno reason to separate from it the genera Pleurqndropsis and Asterclasia, It \s characterized by having flowers developing in successionin clusters, by the minute calyx, by the partially united carpels,and by the massivereflexed stigma,

Chorilaena. This genus shows strong affinilies to Phebalium secl. Gonioclodos. The calvx is patelliform with valvate lobes, the petals are valvate with stellate hairs on the outside, and the anthers have a small acute apex, In all these characters, and in the seed, the relationship is with P. anceps. lt has been placed in a separate sub-tribe because of the presence of a thickening (ligule) near the base of the staminal filament. This thickening is little different from the tuft of stellate pubescencefound in P. squqmeumwhich speciesappears to offer in several of its characters a connecting link with some of the ligulate genera' lo II ((zn6DzLt69 reN pl^'s{lta H f Jc) *

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STIPULES. These organs are often present i\ Eriostenon seat.Nigtostipulde but are absent in the other sectionsof Erio;temon and are not obiervablein Phebalium and Crov,eq. - Wh-en presentthe stipuleson the young branchletsare at first delicate, muttrcellularstructureb about 0.2 mm in length on either side of thepetiole; !hey_maybear a few unicellular hairs. As they get older the basal cells Lecome filled with a dark-brown or black, brittle resin-iikesubstarce which enlarses and often forms a hemisphericalexcrescence on either side of the leaf-base] philotheca . These stipular excrescencesare also found inthe getus (which is closely related to Eriostemon sect.N igrostipulat) but apparently ar€ lrot present in other generaof the Rutaceaeu hich as a farnily hli always-been coniidered to be exstipulate.

CALYX. In Eriostemon and Croweq the sepals are distincl and imbricate and the calyx is clearly demarcatedfrom the pedicelapex. In phebalium Ihe sepalsare free (and valvate or open| or rutiaedjDto a patelliform to hem! sp^hericalcalyx which (except in sect. Phebqliuftr) merges into the fleshy apex of the pedicel.

P_fl9 A" intrastaninal rirrg of fleshy tissue is found in Eriostamon speaes. This is usually convex and bloader than tlte ovary into which it may merge or it may be -distinct. owing _to different texture or colour. The carpels aie somewhat embeddedin the disc and in this portion are joined together. In Phebalium tlre disc is much less distinct and i\ sect. phebalium is eiiher absent orinsignificant. In P. rude it forms a short base to the ovary and it p. anceps it bulgesslightly around the base of the ovary. In p. squincum the disc-is approrimatelythe samediameter as tlte ovary but demarcatedfrom it. In the secl.Leionena a disc as such is not presentbut is replacedby a gynophore which is prominent in somespecies bufentirely absentjn othersie.s.. p. ro'tunt]i- fo.lium), ^ There does not _app_earto be il sharp demarcatio[ betwein the gynophore of sect. Leionenta.the disc of sect. Gonioclados,and the apparent abiince of disc or gynophore h sect. Phebalium

OVARY. The typically 5 carpels are shortly urited at the base bur are orner- wise free. In one species (E. trachyphyllus) they are joined (except at apex) by a peripheral membrane (epicarp) but the radial walls remain free. The style is always solitary and is shortly.5-rayed at the base where a branch passes to each ofthe placentas. The carpels are always biovulate, and are apparently so throughout the Boronieae, a distinguishing character from thbse New Caledonian species which have sometimes 'fhe been included rn Eriostemon or Boroniq. funicles arise collaterally on the axile placanta; however the ovulesare sub-collateralor almost superimposedpossi6ly owing to the sltape ofthe loculus. See Fig. 1, 1 2. The apex of the carpel is usualJysolid and sterile (not so in Cronea and E- trachycarpus) and in fruit forms a more or less prominent apiculum or IOStrUm.

OVULES. These are anatropous, 2-integumented, and the micropyles of both of those presentin eachloculus are directedupwards. Engler 11890and l93l) describedand illustratedthe funiclesand ovulesas being"superimposed with the micropyle_of the lower ovule directed upwards and of the upper ovule irected downwards; this is not correct. Normally only one seed develops in each carpel. 12 €t

't '8IC ees 'spesul ^q uo[lnql4srp paas uI onl?^ aq ol Jo 'lces ,{la{Ilun sI lr :lu"cUIuSIsuI pu€ ,{rp fll"nsn sI 1ng (sorl|rnuso ptde Dwauollg uowatsoyg 3'e) suorpes aql auos ur punoJ sI (alclunJ el{lJo uolsu?dxe u? 'e'r) -aDaluoutalsoltgJo Iu€ enJl y arl] Jo peas eqt o] peqJEDESuIJq sB sJollo,t\ euos ,4q pouoquaru ,. IIrE,, agtr sluasoJder,{ygegord drecopue I€luoceld eg1, 'DWaUOla1'lte9 UfiUpqaqd pvz 'oa$o,tc 'pulauoul)o) pes uowalso!,tg 'peosPu1? 'lxos uoluatsol.tEqll?'(\ esec eqtr sI uoDlpuoc Jell"l sIqI eq1 uo luelsls -Jed ,{ ?nsn are rlloq puB ru.rg osl€ sr lueluqc"]l? r€lnclunJ eql )cql sl 1l JJ 'p€s lnq pacsqop 6ql luo{ peqcElep s3uocoq uoos snouBJqrueu Jr dr€aopue 'elcrunJ I?}u$"ld srql egtr dq pses eql 01 paqcBttr" sumrusJ lnq dJ€3opue -crls?le eql uro{ uJol seuocoq aauocsrqepSuunp pue (sur3r€u erll 1? lsBel 19) 'Iccot urql sul"ttreJ uo4rod plueceld l?rx€p? eql allq,!\ 3ql uJo{ sPaJSJdrJ Jq} Su odord JoJ rusru€r{ceu e Sururroj 'snouEelqrec pu" clls?le euomq snFcol oql sll?,r puB eql :seJnpnrls o/t\trolur s3l?Iluo]aglp drPcopue Jo 'e8pa l?Ix?qe lerel?l eq1 IeIx" eql Suole secsqep 1r :(sladr?. Surrnoqq8rau qtl.{\ lc?luoc ur ,{lsnor^aJdoreqa qloortrs suEruer lr slle,r I?rp?J orll EuoF) esocnJJe^Jelnpu"lS ,tlensn pue ,{poorrrJo snoea€rJocsI Jeru;ogeqa drecopua pu" dJ"clde eq1'sJe["[ 'al?cu" o,r\1olur p5l?r?dos s0uroSaqorerrJeo eqJ lp elxoceg Jo lceJe uElu3J 'JIn)Id ,(oqt :aseq eldls er uro{ et"rBdes pun eEJ?lue'eytgeg;r'qed.rtc aqJ Rutaceae tribe Boronieae subtribes .Eriostemoninaeand Nematolepidinae

KrY To GENTRA 1. No ligule at baseof staminal-filaments. 2. Stamensunited for more than half their length. 3. Stamensall perfect Philotheca 3. Stamenswith 5 reducedto staminodes Drummondita 2. Stamensfree o! slightly united at their base. 4. Calyx inconspicuous;petals induplicate-valvate,tomentose outsidej flowers maturing rn successlonln ctusrers Urocarpus 4. Calyx distinct, or if small then petalsimbricate. 5. Calyx shorter than petals. 6. Anthersvith a sterilewhite apiculum.(sometimes.minute ): stamenserect or inflexed G\cept L. d?oerti\.filamenrs u!ually ciliaLe wilh simplehalrs. 7. Anthers pilose Crowea 7. Anthers glabrous Eriostemon 6. Anthels\ ith no sterilewhite apiculum (som_etimes \ ith a lerminal'glandr, sramens Spreaolng.glaorous (or wlln SlellaLehatrs at ba)e). 8. Ovary bicarpellary; flowers small, sessiiein terminal heads; petals glabrous or nearly so Microcybe 8. Ova-ry-s-carpellaty (or if fewet then flowers not in heads),petals lepidote outside or glabrous phebalium 5. Calyx longer than petals Geleznowia l. Hairy ligule at baseof staminal-filaments. 9. Flowers in densereflexed heads Chorilaena 9. Flowers not in heads. 10. Sepalsunited into a hemisphericalcalyx, petals denselylepidote outside Nematolepis euphemiae 10. Sepalsfree, imbricate, petalsglabrous or almost so. 11. Branchletsaqd lower surlaceol leaveslepidote; petalsvalvate, connate Nematolepis phebalioides 11. Branchletsand leavessparsely villousj petalsimbdcate, free Muiriantha

,4 " qal€xo 'J '€ .rn,nredsol oleaunJ-Ssolqo '(seull 't se eel :elerel + ro r€ln8ue jlesnlqo ,rdoJl?uw lseel lE) snolnroqndslalqauerg 'z J[drltr :(osEq q."a EuEIIes'J + JeaI'snolqelts ]Bel IrIo{ }uermaapelSuE al3uls e) po8u d,{lMorreu ro telnsue Flrtnre slrlqf,uerg € 's8ulrd 'alpue Jo Jred13 se luelJncop1ou seêe-I I El€luep Je^ €lloJlNn8ue'J qI ellq \ xlensn sJo.{\ou: l}eu'al€ôqo o} trldlllo saêe'I Z E![oJpsn8u€r€^ €lloJpsn8ue.J aI 'Suolqo_,1 {uld I ensn sJel\og'.{Jp uaq \ pe,lrnagrEJBISUI oJreu ol leoull saêe-I Z 'urels 'etel1lJras Suole s8ur,delelnclluep-aorreu o,^[]se lrlerrncaposeq eql se^Be-I I '€IJolcIA pue S3IBI\ 'sol3eos qlnos ,{\0N luo-r.} o,\\1 pu€ ?rT€Jlsnv ulalsei& luo.IJ auo ee.Irll 'Jpuv 'a8qDS 'J :serJads ed,{I '(1561)eltq6-qrnus epg '6t : u ou euosouorqJ 'lqBI€Jls 'I '9 :g-t 'I 3Il ees 'rouedns eluPBr 'xoûoc-ou"ld suopel,{1oc'pa^.I[c Îlrl8{s 'e1ere}ofrqure :snoldoc ujodsopuo :l"s"q ?zPI?qJ 'l?seq '^[uo ?1sot]olno u,rorq aql fq pereôc '.{qsau erlder leqdur o1 (peas jo xedr .reeu)e1,(dorcrur uro{ Sulpuo}xa ur8l"Lu 3uolt e,loor8 ,lrol1eqs 'ellluqI€IXap" '{clql ? unlrq :al?8[.Lroc ,{l]€ln8e.ur .ro qloorus pu€ )celq ?lstlol3lcs 'u,roJq 'lqBI"Jls :snoec€soc + Blso] Jelno :8uol ruru g t-Z utSjtu leuepe '(oqd?] 'ul8.reru qlr,{\ ruroJrual + ol lxeu pue la,\\ol le,(qseu SuluoJeq IBIxspa 3uole lrre reeurl t Sutuuo3 elolunJ 3ql) elisses p33S snoueJqrueru sulSl€ut pull raddn orll "(qseg drecopua peqc"lls sql Jo lJ"d I"llu3c la'{\ol eql I"Jelul'lualsrsred puE peq3"tlE ,{lurrg d:ecopue go uorpod 1e1uect1dqlr,\\ pees petslq 'uou,rdlso!.tE 'secrdt -aq ur se eruecsrqep pepuno.r qll\\ r?tot lcere g o1 dn 1o '.telnqolS '[edr"c lrnrl eru8rts qc€a Jo uIS]Etrr l"tx?pu Jo xed" r€eu palrosul 'uoqs el,(ts sprz,tdn palca;rp seJ,{dorclluaq1 qll^\ pesoduuedns + soltlô :,(lleJel€llo, Sursue pue :Uoqs ,fto^ so[trunJ 'ellx€ uo4elueceld :slelnôJq 'xede elue1sou qtr,,r\'rsrpur p3ppaqura'(esBqtr" petlun,{pq311s)eer;'9 slodr€c 'csrp 'ymtraqdsrureq 'snoJqelS',{JEô lsnorqelS eôqe urlu 9.0 €c ,{JEÔ punore ur8rEur peqoT ,rorl€u r SuruJlog '[eulru"ls€]1ul cstq 3uo1 utut g-7 e8epueddeelue;s 'esold 'e1tloecur1tolut sp;e,tdn pecnpo.rdpuu esold ,{1as:tds 'xede-lueu"[g etcaql ',{llzurpnlr8uol Surasqep 'esJo.I}uI 'o}€Ploc-Suolqo erll Crowea angustifolia Sm. J. E. Snith in Rees,Cyclo. 10:n.2. (1808). T]pification:'FoundbyMr.Menzies.nearKingCeorge'sSouncl,onthewestcoastofNew rlolLlnc, l\oe not seen. C.ângus(f-o^lia"turcz.Bull. Soc. Nat. Mosc. 2212:13 (1849); Walpers, (.1852); Ann. Bot. Syst. ?:249 Bentham, Fl. Austral..lJ-30. (461); J. Muelt.,'Fragm. S,1OO(lSiSJ; 49.9.(".-f,, .Bo-t.,Mag. t28:r.7870 (1902); Heal, Card. Chron, rn. og:S_-ij tlSZt); i,"1lo6, wildfl. w.A. 36 (1921). Typificatio!: "Nova Hollandia. " Swan River. Drum. coll. III. n. 12.,, Ho]otypus: KW (photo seen); iso.: K 12. Drumnond, Rilgr" (King 9ryql ..DrurninondGeoLge SoLLnd aic'ording to Hooker f., 1.c.)(photo seen),MEL 4548, NSw 69928 i2. i8q5 -. zriostemon turczanitloviiF. Muell., Pl. Indig. Col. Vict. 1:120(1862), (based on C. ingustifolia -lutcz.). Woody perennial or urdershrub, 0.3-3 m high. Branches slender, glab_ rcus, smoolh, terete but with tarow, denticr-rlate wings decurrent from-e]ther side of the leaf base. Leaves chaflaceous (rarely irb-coriaceous), smooth glabrous, sessile with a broad base, linear to broadiy elliptic or obovate, apex 1ou.r]dgdJq acute,^midribprominenr belorv. Floners axiilary, usuallysoiitiry. Pedicel glabrous, 2-5 mm long, rrarrow at the base but fhick ind fleshy towards the apex. Bracteoles2 4, basal or sub-basal,ovate to lanceolate. 0 . 5 1 . 5 mm lorg. Mature bud narrow-ovoid, ca. 8 ml.r long. Calyx liemispherical; sepals free, imbricate, cor.iaceous,sub-orbicular, 1:5_2 min iong, glabrous, ciliolate.Petals persistent, thin, ovrte, cr. 12 mm long, white to pinf,, Secoming darker with age. Stamenserect, ca. 7 mm long; filamentslinear, glandulai \errucoselouards the aper. sparsely pilo\e on rhemargins and adaxiaisurlace; a,ntner.thecae2 mm long. pilose.abaxially towards the apex; sterileapiculum lnrn. prrose..ttnear-tanceolale.uhile. ca. I .5 mm long. Dj.c purple.forming a narrow.,undulare.sloping margin lo_lhe o\ar). Orarl ca.0.7 mm high:sr"yle + equal to stamens.pilose; srjgma globular, + twica style width. Coici erlct rounded at summlt, ca. 3 5 mm long. _Seedsubreniform,2 2.5 mm long, outer testa co aceous,reddish-brown, sclerotesta smooth.

la. var. anglstifolia

Leaves linear to larrow-oblong, margin denticulate and recuryed when dry. Flowers usually pink.

DrsrRrBUTtoN:Western Australia, in the extreme south-\vest.-Map 1. WESTERNAusrRALrA: South Bindoon, Barmere 151.(MELJ; Kalgan, M. Davis 526 (AD); Two-P€olles*Bay, C,.!\. cardner 3318.(PEBTH);King Ceoigi SEL,nO,Laa.i,rin, :ufv fsjd .]\4_EL);Bow_Riv_e-r,-H. T._Saw, Jan. 1925 (PERTH);4 niites east of OysterHarboui, Albiny, E. Wittwer255 (KINGS PARK).

lb. var. dentata (Bentham) P. G. Wilson, comb. nov. C.denttlt.t R. Br- exBenth.,fl.Austral,.l :330 (1863) [dentatus:toothed]; Gardner, W. Austra I Wild (1951); wildfl. fl. ed.8. " 70 Cardner, w. Au,lral.r.7s (1959,. Typification: W. Australia-.I(ing "fvcz, ceorge's.Sound, Baxter (Hb. il. Brown),'. Holot'?us: K (photoseen). C. angustifolia \ar. de tata(Be\\th.)Blacka1l, How to Know W.-Aus- ra]. Wildfl.258 (1954), comb. invalid (without full citationof basioivm). C.angustifolia Turcz. Benth.,1.c.: F. Muell., (1875). " .\ar,.platlphylla Fragm." 9:106 Typificatior: Franklin.river,.Maxvel1." Isotypus: MEL 4549 On the siopinghills near the entranceof the FranklinRiver ". Leaves elliptic to obovate, flat. Flowers usually white. Chromosome no. n:19, from material collected at Manjimup, Pemberton, fide Smith-White(1954). DrsrRrBUTroN:Westem Australia, extreme south-west.-Map L WESTERNAusrRAuA: Nornalup, A.M. Ashby_6t.(AD); Walpote,A.M. Baird, 24.xi.1947 (IIWQi !!efr9y_"__f*!,Poronsurup Range, B.G. Briss!, 10.\.1960 (NSW); Alba;y, J. Ctark, 9.ix.1923(PLRTHJ; Big Brook. Pemberlon. M. Koch2239 (MEL). 16 '(8i8t)08Il:tpuEO uuv'uouol^I:(I€8t) ZJ.,f ,.fff oeC'uoc C \. sriqpo:rttltolltDlJ 'GL8I) luollrc{ gzn J qns /8t:t Id lsrH (s,rr\orpuv)srrt{,s 3' '(€96I) "le 6t€ tsrc p.{sId'cse^ qpu€H la elp€og :(906I) seler\\ 'uoxrc :(t68I) seLE/A qpu€H 'oqJleg 10eroo^l 0S S N Id 'SurtrlsSt S N T4 '3'Du8ltDs') :(/88T) '90T: II salelA S N .oS uur-I rord :(DlDlDxa) pLe'Dlto.lltot uo pes"q) Bellrurou'etd d'(298I) 6Il:I t3r^ IoJ Srpul ld'IenN C !a^o.t)uoualsoltg '{1ua1slxe_uouueur -r.eds) q"t:ed.,(J rEe,{aql uI'speestuo{ posreJ puelloH r\eN : uorlEJU|dAJ 6/ ..06rI ,, '7 'rrcrr)'(I€6t)iol, +^/ r:rr^ rr lrp^\.r:arc

'k optaap ,,leqlrda eql pesno.roJeroql e^€q 'LuS I ollo.f!$n8uD J 01 peJJeJSuB.rtjr (ru,(uouroq .Iele1e Suteq) el?uJntSelll eq plno,r 111uz1d srqt loJ :leqtrde l"tJrJ?^ lsorTreeerll sI ,, "ll,{ttdf1"ld,, "llq,^(\ 'tuetuuoJr^.ua01 asuodsarJrd,{louerld " ,{luo eq ,{erueJnlxelJ?el eqlpu" 'Dtoiuap IEIrol€Gr aBu"J JeleeJB rl1r.\\ u,rop srelcErErlc tsa\I J Jo se^Bel Jo 'snoecuuoc IBoJq esnlqo eql lsurEBEs" ss^"sl alnce urrp eq1 ,{q s3rJadsJe1l€l aql 'Dtotuap') ruo{ 1r paqsm8ulsrp Iu"rllueg 01 p3Jr3JeJ,{luoruuoc 1ul1d pe,leel -peorq eq1 sI 1I:lle,\\xEJAI,{q,,re^I)I uII{UEJC,Jotll 1e pe]cellocl€ual"tu uo pesEq sB,\\ 't]iLuegDt\qdfiDtd i€^ zcrnJ DllotlsilSup l,rrlo.rJ olu€u eql 'J \ eLt pLnoqss:d,{toutrld lueJegrp eqtrsn1r1s cgrcedsr.rjur ro cgrcadseqt uo oprcep shrub to 1.5 m high. Brarches smooth, glabrous, with a narrow wing decurrer]t from each leaf base. Leaves chartaceous, sessile, entire, smoofi, glabrous, flat or slightly recurved at margins when dry, narrowly (to broadly) elliptic, 30 x 4 mm to 60 x 13 mm;apei acute to obiuse,apicuiaie; base narrowly _cuneate;_midrib prominent beiow. Flowers solitary,ixillary; pedicellerete, 5-grooved,5 13 mm long, with 2 pairs of basal,blu;tly deltoid bracteoles ca. I mm_long, Calyx hemispherical; sepalsfree, imbricate, broadly ovate to sub-orbicular,3 3.5 x 2 5-3 llrm, coriaceouswith a thin ciliolati margin, glabrous. Petals chartaceous when dry, elliptic, 12 20 x 4 7 mm, ciliolate otherwise glabrous, pink (rarely white) to purple, apex f ilflexed to cucullate. Stamens pyramidally arranged and cohering by their promilentlv pilose margins, 4 5 mm high (artisepalousslightly longer). apicesspreading during anthesis;filaments narrow-obloug; anthers ca. 1.5 mm long. StylE short atrd thick, ca.0.3 mm long, pilose; stigma massive,subglo6ular ca. 0 7 mn wide. Cocci + erect in fruit. ca. 7 mm high. Seedovoid-renifor.m, 3 8-4 5 mm long; sclerotestairregularly rugulose. : - _ Chromosomeno. n 19, from material collectedat Kuringai, Berowra, fide Smith-White (1954). DISTRIBUTToN:New South Wales,Port Jacksondistrict.-Map 2, NEw SourHWaLEs: Killara (Sydneysuburb), E. Ashb),25.ix.1915(AD); BaLgoCreek from Igl{rLbogf, Blakelyand Shiress,Oct. 1953(NSW); Teffe1 HiiL, B.c. Brrggs,10.vi.1911 l\q\ ): PoflJack.on. Roben Bro$n, 18025 t\S\ l; SouihKincumber. Cl-Chiooendale- l.\ii Iq5l(NS$):Ro.e\illeChaie.in:and\lone \crub-fore.l. l.A.S. iohn.on 484iNSU/. VrcroRh: PineMountaio (neat Walwa), Upper Murray, C_ Walter, Oct. 1891 (MEL). ^ This speciesshows only slight variation in flower sizeand colour (a rvhite form is reported) and in leaf shape,the latter no doubt partly inflr.rencedby environnrentalfactors. It is probably fonnd only in the area around poit Jackson. The single collection from Pine Mountain (near Walwa), Victoria, on tbe upper Murray was recorded by Ewart atrd Tovey, Proc. Roy. Soc. Vict. 32: 193 (1920),and cited by McBarron, Contr. N.S.WalesNat. Herb. 2: l9t (1951). The specimenwas correclly deternined but I think its natural occurrence in that area is very unlikely. C- saligna has frequently been unit€d, under various names. with C. exa.lata. ll may be distinguished by its glabrous branchlets,pedicels, and sepals, and. by the silgle _narr_orv-wingdecurrent from each leaf base. In C. saiigna tlre_lea_vesare usually elliptic and acute, whereasin C. exalata they are spatuiate with the apex obtuse to rounded. As foted previously, the genus Croweq was named after James Crowe. a surgeon who was also a keen botanist and especially ilterested in the genus Sa/r.rc(the willows). According_to_J. F. Smith (1808)the epithet ,,salilna', (: willowJike) commemorated this interest of Crowe's. However- this specific,epithet was.aclually provided by.H. C. Andrews who called " " the plant the Willow-leaved Crowea from which the scientific name is most likely derived.

3. Crowea exalata F. Muell. f. \4ueller.lrcns. Phil. Soc. Vicl. l:lt (1855r.lex alarus: $ingedl: ( _$irhouf. Benrham, Il..{qulql l :J2s L86l):f..Mueller. Fragm. 9:106 ( 1875): woo s,proc. Linn. Soi. N.S.Wates 11:929(1886); (1931.). " Ewart,Fl. Vict.704 JVpification: On the ro€kytops of Mount M'Farlane,about 5,000 feet above the sea;on the gravellybanks of the Mitta Livingstone Mitta and Rivet" to\rardsLake Omeo, and olr the BoggyCr€ek in cipps' Land." Lectotypus:MEL 4410 Mitta Mitta, ad ripar; ghriostm 18 'roIooH "llenl,l{- td ueD 1oruBqtuag t"rou ,,rrrr"ir,,, lrr^ loJ Srpq Id d :GSsI) p:1 ursel jJeqoo11:(OtSt u:9 r:qrr1po1:(1€8T) llr. do'rouslalI 1gi ;Jeis'reyuegti 14 09:Z'qreds :OieD vf t uep csbl-14 ittistt 8i€:z 3?A teN tsrH l(I€8I) z6l.:I 'lsrH ueD'uo( sued sn'^{LU?t 'nrrssnf :{}Z8l) 0Zl:I poJd 'ellopreJ C:(928I) I8t:ZI IEN tsrH I ep :,. urnulelsouE,,([Z8I) 081;lI loleJ- doe'I pB.VeDV e^oN sntlrElIle sooN : ojdd ig1 px3'saegui qirtus:(tO8I) pnrd'raqroeD 99t:I ld 'u,tg'ucioiJei GoSI) €SI:g C C:(908I) : [ueirefs: uorueli1oo,r : uoue](86t1) Izz:t cos usrl sue4'rllltus E I .UIS NOI\ISISOIUf,

'lrn{ uI 'lnoloc ueer8 ol oBu?qJpu€ ,ro.rJurrur?ruer sl€led eq] ?rJolcIA ulalsed\ uI altq,t ur u,rorq rlsrppo.reuoceq o] r?edd"'lEu3l€!u unu?gJeq ruo{'pu€ 1In4 uI rdpeorq eruoieq qcrq,n s1e1ede^"q sluPId srlei6 rllnos ,,lrJN,pr.tr,{op-ro8rul stll x,,(l?Jpue sleJlpedsnorqrlS lsourlepuE tlrJlssnoLnlJqnd ,{l!sr€ds e,{luo suq osle'b,tnioy1 pue-er,ro51 uao,\\leq punoJ rrrJoJl€lsuoc " :3suocseqndlq8rls ,fte,r ,{yuo pu" so,\Ee[e8rrl seq lr sel3lN rl]nos ,{\aN Jo lsEs-q ou l"J eq} uI 'sno1n:eqnd,{lesuop orour sr Illels eq} pu? 'lsBepue rllJou eql ur esorll u€ql loll€rus eJe sre,rou: se^"al eq] set€ld qlnos ,t\aN pu"lul uleqlnos pu€ l?rlolcl^ uI 'lunlueunpurPue ur pus azrsle^\ou pu" J33luI ^lqBJepls[ocsauE,\ selte(]s sIqJ 'lc 'l.tlrslp '{.un€qol;El 'p,.J 'pt-ltuo:J6'I"Jl'i"8tl;tj;$ o;rpuog : ('rEI',I)696I t^ tI-€I'ullsoJ f pue tqtuoqlu€I 'tfil roo,rtoq:(gI M) 996I qeJ { V lll^\oH lt{ ru : vrtor3l^ '(,1ASN) '.elqslsuoJ ')ooJJ.{tuBSIf : (/ASN) 096I I^ TI CarudpaJo N ltu 9 -[ uel 'leeq) g 'I.nf8uu.rlgru 'ln '\\erFuv :(A\SN) rrr'62'uEujroog 'I Z16I'uepg IJEIS'., 9T6l ru',{eFuItW :(IASN)nl6l 8nV'ucLuroog'I I q:iLqlJ t-Z Jo lueLo llel V !! re^tu 'qeeruanj '8nV 'u€turoog 'I ',. qBIq slrlels,{u?tu saaioag : (mdNl soef I ' { Z-Tlnoqe Jo Sur.ror6rr]rot uoslref uod eqt ueqtlueld telqsnqq.nu V,, e,\\oH: s3'1vl grnos ,4\eN '7dr1,,q'uuolcrrlu.rels€apueleJluac'selEi6qlnoS,\\eNuJe1s?E:Nollnalx'rslc 'ePLt\ 'ou84os uI s" peas pu" :lllljg ulru 9.0 3 '{cql 'uoqs 'ec 'sno:qr18 'l"lnqol8 "lrl8rls:osolld'8uot ultu ?J el,{ls 9.0']eU 'ueor8 ',{r€Ao otrur8r€Lu peqof,\\orr?u e SuIuuoJ csl( ,'\\olle,{,uellod Irep 'elr]urolnc" 'Suolqo :8uol ruur ec e8upueddc eql.Ie1s'6uo1 urlu eo e€ceql 7 I '?uol -,rorlEu sJ3qlu? :{ord 'uqtl'\\ po" suISJEulerll uo esolrd ruul e-9.2 '(urqlr,r xe,tuocsn6lidesairr;'paue]]eu sluaulelU:6ulp"elds ,{llunlue^3seclde eluels eql 'etB3uqrur^llq8IIS suaulElS (]tn:j ul uee;8 saoIllsulos)e^r"tu '1In4 el?d ol )turd '8uol lllru (S) 'elE^o p?olq ol ,\\oJ-reu uI lepeolq pue ZI 0t 'snoJq"l8 siroecegeqcSururbceq 'urqt 'elecr:qur ,{FuoJts sleled 3:}"lorlrc o} 'el"c snoyrreqnd'8uo[ruul g flpeorq o1 .relncrqro-qns qrul ,{l3uo]]s Z Z'011]^0 'xed" sledos :letueqdsrltlarlx^lEJ snoF.reqnd,(l3sr"ds arll spr€,\\ol fqseu '8uol ruLUt S lecrpedlseloelce;q ll"tus g Z 01se^?ol3q1pu€ (ssJl lo Suol I'rlrul) elcunped:l.roqs" o] pecnpo.Itooqs erll Jo se^?Jllucrd€-qns,\\oJ " tstlu"eqlooqs ,{.rt1irxugoqs t

Woody _perennials,shrubs, or small trees. Branchletsangular or rerere, sometimes channelled, glandu)ar punctate, often with glandular verrucosrties containingan aromatic oil, glabrous,glaucous, or with sirnpleor stellateindu_ rnentum. _Lea\esalternate. simple, exstipulate (or, in sect.NigrostipL ae, \\ith minute subulate stipules uhiclr form dark excrescences),shortly petiolate or sessile;lamiua terete to flat, fleshy, coriaceous or chartaceous.glandular punctate,often glandular verrucose,glabrous or with simple or stellalehairs. Inflorescenceaxillary or terlrinal, ioose or compact, in-umbels, cymes, or racemes,or with solitaryflowers pedicellate or subsessile;bracteoles prominent (and then usually persistent),or minute ald caducous,basal or subtendingthe flower. Flowers pentamerous(rarely tetramerous);sepals free, f imbricate {a,l lerst a( the brse), persrstent;petals free, imbricate (rarely valvate),ovate, elliptical, or obovate, thin or 6rm, with or withoLrta prominent midrib, t glandular punctate,glabrous or with simple, stellateor lepidote indumenium on one or both sjdes,flgquently also papillosewitbin, u hite to blue, pink, red or mauve. Stamens l0 (8) the antisepalousslightly longer than ihe anti_ pctrlous,,usulrlly erect or pyramidally arianged and incurved over the ovary (aprces becoming recurled during anthesis);filaments usually firm and i flattened, ciliate or pilose (glabrous in E. Jitzgeraldii, glabrouj and f tereG and.-spreading.inE. deserti); anthers versatile, f cordate, glabrous, with a small or promirrentglabrous white apiculum, dehiscencelongitudinal introrse (the antisepalousfirst). Intrastaminal disc ring-like, or lobid, fleshy, rarely obscure. Ovary + embeddedin disc, pyramidal to hemispher.ical,carpels (3 4) 5, free or slightly united at base (n E. trachyphyllus uniied to near apex by a peripheral membrane), with a solid, sterile apex (absent in E. traihl,_ p/ry1fus),unilocular, placentation axillary, biovrrlar, the fturjcles very short and arising collaterally on the placenta; ovules f superimposerl,anatropous, 2-integumented,micropyles directed upwards; style solitary, often short at first but lengtheningduring anthesis,divided at the baseinto brancheswhich are lnsertedabout the middle of the adaxialmargin of eachcarpel, pilose, stellate, or .glabrous; stigma capitate or shortly 5-lobed. Fruit of up to five cocci which split.alorg adaxialand apicaledges; rhe cocci erector spielding, usually with an apiculum or beak formed from the srerileapex of the carpelfpericarp of two _separatelayers, the epicarp coriaceousor wbody and often verrucose, the endocarp mostly cartilaginousand elastic with a small membranous or fleshyportion around the placenta;dehiscence is due to endocarpejecting the usually solitary seed and itself from the carpel. Funicle shoit and thick. Seed _sub-reniform,flattened or pyriforn, sessil-, or subsessile,at lirst with the membranous placental portion of endocarp attached, or the placental endocarp thick and persistent; the funicle sometimes forming an aril bn adaxial margin of seed,this aril when pr.esentlinear, thin and membranous or fleshy,cJn_ tinuous.atbase of seedwith raphe; testawilh outer layer thin, membranousto sub-coriaceous, inner layer (sclerotesta) thick, crustaceous, black, smooth or sculptured; micropyle oval or linear, towards apex of adaxial margin; hilum in centre ofadaxial margin, linear to narrow-elliptical or deltoid, superficial or sunken; raphe massiye and fleshy or_shrunken and insignificant, superficial or in a prominent hollow, covered by the outer testa or s-ometimesby a thin or thick crustaceouslayer; chalazaol the lower adaxial margin oi basat; endo_ sperm fleshy;.embryo terete, t axial, slightly curved, cotyle"donspluno_"onu"", radicle superior, snuElqlEs 'dsqns sfuroJlp 'fi 'qL 'clldrlle /trql 'leg 'II ( o puE ia s N) SuoI ruur / ol trIfeorq 01 Suolqo se ee'I sJuuoJlp'dsqnssluxoJlp 'g 'el ( ' ' SuoI wur '.{ôleq polesueqr A'elojelqns IA S N O) 8-t Jo o^?auoc'aôqexoûoa Jo leu + ro ploquroqr ol eleô' qsouseêo'l II '(solEls urolseg) (^\oleclpalouueqa Jo aûruoa uoql elelol-cltlsJl) eslAueqloso^€o'I 6 snl€lmJds 'g 'tI eleln. 'luocserq€J8 -rde-1ce1qs..lee1 :serruerrnaopJeol lnoqtr^\ ro snorqelS slelqtuerg 0I 'E '0J IJauprBB olEtnsrdelou so,\Eel:serruerrnrep J?aI snorq€lF uee,{\}oq soulL snolnreqnd peorq ql!/1islelqJuPlg 0I '( 'oltêl. 'elelel-qns'elerel V 1(\) ,l\oleqpapunor ro so^€o-I 6 'ssalroBuolrutugslEled:leupueuLlllro'rlEôqoolrlrrrlput'{LLseUseêe_I-8 '9 snlJoJlclrag ( l S N) Suol lutrl 6 e3 sle}od:lllncl.e se^€o'I 8 '(ur8reLlrspre,rol ]sBolrle) eptslno luerseqnclspted / '3seqt3 secuorserrxo.relndrls lreJq Jo .tr€de qlJ^\ sa^"el 9 'leulurrol sre^\oll I 'g sn ,{qd,tqred t ^\oJeqeêouoa'Suolqo sel€eJ:;uolluLu I er'elrqa sJ€loo:lcelesJeA\oTg € 'g suqnu s per oled sre.^lou:^\olgq xeûoc pue eôqe peuetleg'elerol-qnssa^ml :punoi llc rsolnsolld ulels t 'g '€ sneullJoJ JJB olrq.r sro^\ol{ I snaulrJoJ g '€ peJ sJeA\oJg s 'aolrq 't oûcuor + ro leg'Suolqo se^Bel:solcuerlncop-Jml usordleq srull uI esolnsolrdLUeIS 'auol lulu 0I-/ q?led:ellq.{ ol par lue}r1u sre,$olJ € '( 'Foolqo 'Iclql V lA) eôqEpeu3lleu ; eler.l qns JI .lo soêoI Z sFeaull'g 'z ( N puE Suol nrtu S e.'ellq^\ sleled :lrero sle^\og :ela.ra]se,\ee'I Z /A S V S) 'IJEIITX€ SJO,\\Oll I lvlndllsosDrN z oll"s snlsele4snB'g 'I $rreds,{LIIO NOn:IrSOtX3 I OllCeS

eetndlrsol8lN.I TollJJq ' : io sr iilo 1'u:js:qiio qr' * urLlr ,{rElllxe JI to leulrural sre^\ou luesqt luasard -9 seouecser:r_xeJElndlls :souJ[leu]pnilSuol uI Jo petoD?Js(lLr0soJd Jl) urols uo sJIeH BUTAUOIIq' _ t' otlJJs 1:ip's1nosnolnprdsrq {lesrEds ]o) snolqDlSslBlod,:s.tLl{t '{lEIIIxe ul ulels uo slrEH .ro,,{ll]ll lxe sJe,$ou: luesqe se3ue3soj.xo.te lndlls : pel3lttrcs(loesercl Jl) 9 'Ieseq'elnulur'seloel3eJ[II 'olecuqur'eSrtlseloal'ela soqluEurso t oll.es (vI ) t€s€q-qns I 't uocseqnd slrlealeL[] ueql seurlleurpnlr8uol LrrJl ro'sJleq perelt]].sqll.{\ lo snolqElStuels t '(lBurlujel rurauou{roJ s oll.es snorqelSsleled :,{l€}rlos seLllgetuoslo) FrEIlrx€ s:a,,aogienisrl tuern.ap posreruee,\\leq saull lEurpnll8uol ueluns ul srrEll Lll!^\ltlols t 'per lo )iuld ol e1!q^\sl€1od : sJe,{\ouleultuJel esool ^\eJ € ol I lo ro {JEIIIX€sJuarseJouuI ! s,{urelqJouElJ ortras ( V lN) )ttro ol anlq 'oiq1 9 'sno.r!!1t slileit :JurecerInururre] ! ro (relsnlc ro) ptoq l.edtuoc e o'ueoseroguJ € 'sl?q eldluls 'saloolaslq qtr.r ro snorq?[8 sI?lad :lestq ueql llleserd Jr ro luoulluord lnoqll'\\ Iecrped Z -t eplslno olopldel uouelsolrg orrras 'lueuruord -eta1a1s'xpsop ile1Jd :se1oe1cerc1el9rrrqur papunor l€le^as qll^\ lecrpad z 'snorqqS ro ,{rl"q lue F :l(1|ptD.t)Ez!!!) lrEdluo. 'pe8ut!e sueLuels pue t.ero uaql sno.rqelSJI rol tlnslrq ,{llEprllrer,{dpue pa^rnculro lrore I sonruuuur,{.} / orlJJs' 'Surplorcls'repuols sirorqelBluptd :relnpoelSepue snorq€I8 sueLuEls I sNollJSs or &11{

'(trueustl 3tIIpnlcur) eIl"rlsnv :Nol.rnalxtsl( 'st?d sntstllll$no uoutdJsol'!3: selteds €d'{J 'sreqlulr pul? pees roJ S t sBrC eas (tEOt) eltLllrt' '(sn6plDJ$nD -qtltrls epll S u] /l) 8Z lo tI '= LI ieqlunll eulosolLloJrlJ petals 7. '' glabrousoutside. :"""1,",;,'"1i"1,::fflu;lktlf,"Llru';h:tiX1$;"r"-,*",16:t.t,,qili,l:: 'i; ,.":':l and^,epdr. " $,rhour3 shnd_tir.eJpjcr,Jr,n_. ifll'i:',fl':il"*:1,'*.Lltil*:.ffil"3,.w:X,luu:;..,1ea,apextoa 13' staminal lilanentse\,.nry atfenuate, E, angustiforius grabrousot sparserycjl-i?t 6 Leateswith no stipurarexcr:esccnces ar i6I t.) ,e. base. r-u*", rcrat\ gtabroL. oLlls,Je. J). tsranchlet\ I Duber j:::l "1,,o$, Ijrer j re. r"."". _i,-.tirl,"i;:I:j:,::, r( ); rea\ e rerete. ^-aptc /ate: \epal\li :uO_orbicrrrar I N,5.\i';,."'J'i]'il,,iil , j ';" ;;;ftf;:"r'.""111i.li::1:i:""':""" rr or ;::ff];::,: ::l;:;.!'jffi;; lf,Ti[.,;],"""i,:1,:,:jl f-5i;*1rul..it"ii;Tn:till t, frBi:i;I.fiTJ: ;;*ti:t outsicie(ati::;Fr reasr,",,*;. #;;j,,r"":;g !,,;.;t; ifi3,,,.ry.Xt*, branchreispuber_ lg. Branchletspuberr - rlous in nar.row lines, densely minate vetrucose; sepalsacu- llj Branchlets E. ratcaius evenlv puberulou. all .ound, Ia. I eare. bmooth ." """,-lt rninutcJ]tomentot. _.''-rharlerruco.e. 19 Lcalesglabrou. or glabre.cenr 15' E lomcntellus 20. LeavesaDd seDa , ;;;#;$:i:*Xt;ft?;*,",. 1s.E romentei,us ,J;.1.*".. ;T:l:hl#lYHri,;f5f marsinonry. hirc$ i,h broad 2L^. pefals "Fu denselyserjceolrs oursjde, .r"nt1 pul" p;,n"t ,Tr$:i:ll,:;:

Sectio 3. ERToNEMA t to narrow-oblo/rg L:llJ:ft" (or nauow-ovate),concave aboveor with inarginstighrry 2. SLemglabro s {or minulel),tellale.$hen young in J. brem glaucous;pedrcel f. bt.ut,i). se\sjlewiih severalO"*l f.*OV 1""t", imbricate bracteoles 3. Slem ne1glnug6u5; pedicel shortly pedunculatewith a whorl or."r"ii"r; ,?;*kt 2 stem hispidulous;pedicel shortty peduncutate toli.,P; myoporoides wirh a,vhorl of .n'r, t "u'pticor obovate, ffion"o oftenbroadiy so, if naro' tn",,',",rn.."r1*0"::]l;lij 4. Plant glabfous. ' l;S:::'"lfJJ:1'11"::iJ$i':i'"1ilJ$illn#,l.sho.rvro),,w,thawho,r orbracteores ar i iffi ;1,11',fJ;ij: J,"lli:::i,llfil ; ;,", ",::;; ". E verucosus pedicelalsessile nr]o#. 5. withseveral small inlbricatebasal Oru","o,.r, ,.uu.rtliu.";"il1::of,lt,r; t ts:itt.l^t' ovet3 cm lons;carpels urited krnta at peripheryjffowers s-merous obovare)under2cm r""*i';"':;"'1""'1"9'"1'5 .. $#:il:"i,l33lih:ifJ:JJ n""iT'id. Lea\es - lhl. bfoadlv oborate or obcordate. apex noL mucronale 8. Leaf-shapevarious, apex mucronate. 22. E. obovalis 22 rJllIonI^I'c :(€68I) st sepasN lc pu"H'oqclo8 lo oroo}I :(6/8I) I/:I 1c.l^ld l€N : (;;8ii : uCa-'uo[ (vzll ozL: I pord'ellop 6i I :6:ub;irriiii"i,t 'git: t f rit rlzol: 1 lsis 9,'J 'E -nej-5p-:116g1)617 I i itn-:ztrietain4rurruos1esnqrlcnrl eq tetJlrata!)sntDloa)uol 'pre\puE trnrJ rql prqrrJsrpFlqetunsord eq qcrq^\Luo{ uort3altoc .$tuB{ qdosol ()1sre]ar (/08I) req oe9'C t:qifus E.I {9]J-P?1l11ej qqrasod'ueas :ed.{J ?lseler}snvuI qeII- :uollB'grcl^J lir:61eu rio:l' paqrrcs:ep^ iou 'erstlersiivl.( 'uoosrod 1i!pniny':o; etueLl.{lrze uE (s081)s9t: I lueld u{S '[ 'sJed snlssl€rlsnE uorualsolrf

'egdeJueluns Jql o] SlruaôJallllJq'prEq " sI oleqtr '^lleJllrqlod,(q 'peûop qxrq,\\ lr aDryd4so;3rg u punoj :di1 aqt LLIoU lcjs 'lces eq plnoc lnq eEeruo.rog eql ur enbrun s uowatsolry Jo p33s erlJ- ',esolcsl dlqsuolleJe.Iarlt fllecneuoSol,{qdleqr:lqlssod sr ]I PUctrrlo/o€r Jo sJltodsAuElu o1 ,{1t.tt1tuussd\oqs 1r slelceJeqcJSJII} u[ {Soloqdlour A]eôlue l"uttu"ls aql puE 'tunluaompurel"llels '[ecLpJd3]EloJlJErq JLll Aq pJqslnbtlllslpsr pu€ uoqtes Jarllo .'(ut o1 ,{1lugr: asolJ oll sEq u Jldllouotu sI UollJJsslgJ 'uo&rad .wrsr1rrls,lrl gr :salsads ed,{I 'pees eseq aôqE luul I I l]3 erld?l Jo pue Je,!\ol lE Jo '{JrLLt rzeFrlc iSurre,locpedlqs piollap 'JDlr.Lq € ^q {ll"urr}\J puE else}orelcs erJl ,{q ,{li"ur3}ul pcptllloJJns 3uo1 urur E. L EJ lsuu"rlJ palJJ]rp '\lprl?'\\ur!\oP 'epr,\\ " scrdnaro qcq,r oqdrr ,{qseu eql olur suedo unllq erll Jo elpplru cql 'ecal ur eJrJrJoue l"rxepDJo eJtLIaJut'pcderls-pto11ep Lunllq : qloorus ("lsel -oJelcs)leuul',{ssol8'{tEIq'sroocnrJoJ = EtsalJJlno :JJPJl?r\r1pE Jo Jlluec ur 'ped?qs-prol1ep',{r1seg-'1Furs '3uo1 'turoJ IIl" :J\l?JuoJJJrJ IPI\l?pl, ruu 9 EJ -iL'e: ;slrsias peeg' 'sur6:?ur slloulliqu3lu qllrr 1cql (pe3s tle:tsrsrecl Ielal€l'xod" pecsrqopuo) drucopual"luectld el"llsoje 1€ popullol ptl€ pBorq'lJeJo icco3' "e:ltirtlec r:ri:rir1i:xocle-ilrrelsou q:ll\\ sledr"J :el"lnalde-ellq^\ ,{puou -rur&cl 'esocnr-ro,txede '51er1tc,{lloo,\\ sluou.lDIIJ'pa8ueJJE,{lepturv-t'{d s;erpie 'snoactuoJ siaruels leprslno e1oprde1e1u1le1s'elsJtlqlul s1e1ed:eltctrqur 'prdledas 'peieilrcs 'sl'ro-lolunu spdes isaloil'cr.rq etzci.,q-l {11,lt.31sses IecIPod :,{lplllx"',{.r€lrTossio,{\olg'alElndllsxs sa^?el:el?llels ^lalnulru slalqtu"Jg uouafsolrg i orlces

'z€ ,{IuO llJrsJp g sorrads soElNvNN^D I O93eS x,{lerols€l 'dsqnssnroBlpou 'g qle enlq sl€]ed 'suol 'plollop-,ucreu lesolrd:elitiats iullt Z ec sledas : Lo€lp urc A I-I sp€aq-re'{\olJ z 'g 'eI€ sruoglpou'dsqns snroglpou lold ol onlq slP]ecl: esolrd-rltllrls llesrDds 'tuoJ'iutu ol plotLrp sorreu : tuDrp toJ [-z {peeq-re'lroll z 9 f'ola1oo5ir:e1:eauq ledes '(ro^o sr Sorra\\oueroJoq lno 8ur^\or8 sotulleuos slxe eql) pegq l€ululel lJEdulol e uI sle^{olg I 'leultuJal snlDJlds'! '0€ etllotE.lsseu"eJ e ul sIe'\\oJJ I s,{l{v_IFIJoNY,rJ 9OIlCeS

'esornJJe^ 'g '62 ( snorqelSsueuels :esnlqo = lElnpualSse^t6-I T tJpt€rc84g V I\) 'ollq,{\ 'ele^le^ 'Z suaEmd 'g '82 ( l\ puE V S) Pnq ut {uld ro sF}ail seplould ( V'lA) per ol?.rrqtul slEl6d Z ! lZ 'qlootus 'esolrdsueLu€ls :aleuojcnLll so^?3_I I vt{!NoN^{oJ I O9.3S '92 ,{luo lernrq'g sol]oals soHlNvhlso t ollJes snlnpldslq'g 'IZ 'eleôqo e1nlo,lei ialeue,rc-:e1npuel3 uJSreLo ,{l,^ôJlEuJo e]€aunrqo ,{lA\oJJeuJee-I 6 'esooluJo,\ snllollxnq 'g '02 pe^lnaai,{llueuruold }ou ul8letu 'r\oqp rel€ôqo tcq$rluos ro qtooLrts e\EJuor '([]q8lls.ro teg ol e]eô p€orq soÊ0-1 6 i'Hq.:}i;:jl#iiit,j,j,lff,;lJ:::fi;pti:'tnti.il,"yrt?,:'.fi::Br.itj6.ii,ilt; Srnitl.in^ Rec... !,.Y,/!li/:i Cj_crop.ij: no. I ,180o,:jiilF,.l,}lRLrJLe, Ifa,r.. r,:,1t:nn. S lL[,:ishiiril'i :ll'"ff i{Jrir!:i,:r i:riiii'il ^."],:*,,n 3,1,'*i1,':ji;;dl9*,j,:;:1,["trisl.rff\]!'l: wares,by John whire, M.n. who

i1,J,',jltgf1$lli-;:iill;l*riTfir:i.irrjj"jjl;i,iii-i l;" ;l:; ::,ii:.o?:f r i", fl:f :"" JI :i.'ol ll l:.):'; Ijll.;,j ;1 i,",'fl,t' r i:::; ,,.,r",uii""?iill ellipticor i.i:}:i".i,,i,\i;fi:"ii;;;]' ij: ;;,'.:,;' .iilfllJni,x"il:?J obcvate),flf,r,.acure,30 x 3 5 70i'8 (, il;_(o iii!ill,,, no.rorrry ;il#tr'!1,tr1,1il:#:'''ilh?:{"f':':{{'ii*i'i:;:}illiiT*rir'srigh'lli' lli$h*$,tJlffi ,11,?,"jli.'i;.51?l,l{1xhJr".i",,l,qt{1",}ll,T+i!il;:l",}*ll:*l**lnli.:!ii:,:iiiitil'ir*;l".',',rr,il,**iffi outwardlyfalcate'in lower half, tereteana stLaigfifaloue, ""jip".i"'fL, "". + _,, *rf"l'lt€jx'$;Ji!:1i{:.:',y,T:li ;#.1*ltk*lr}liit;: "l:u'u: #,"jlfrffL*.*qd:sJrl{::'l"ub[k+i;i;iuJ,',,f ca. 9 mm hish. ff#'{,$;

H3l::ffi','.TfliliiiLlfl andsL eueencrd,la. subsp.australasius 6ili.i*'ut,* lb. subsp.banksii la. subsp. australasius

Leavesnarrow-oblong to elliptic, faintly neryeil. Chromosome : no. n 17. from mlterjal collectedat Kurringai, fide Smith-Whiie (1954). Warrah, DrsTRrBUTtoN:p_ur, "our, of New South Walesand eueenslandfrorl lllawarra north to Fraser Island and in the border mountains nea,lStaoif,oip._,Uup :. QUEENSLAND:Pyranrid Mt.. ca.30 mi fronlSrrntborps., H. lrv,i\, 6 ^j.tSlt 1Bntl; F.u.". B]3ei;,ru:,f*fli3,i&,1?Z-^!3*l,Ly^, K.N. shla rre 1BRrl; irn -an riay, wide Bay eo,\4F j\er Niir?::1,,\^:';;.T.:[;b)juy;$itrLhelsr ',lu r, r \ansR Ricr- ,r sur: Mr Burralii;;e;i."ri:-c,i "u"ef $"il;;;*i [i:,"T["ft tliifl lTj,l:1"",i.::jJrtJffj;ti{.r},:'"0n.""c Da!i.r|,Nsw,,toniol,i[_'ca :o,i Sor lb. E. australasiussubsp. p. banksii(A. Cunn. ex Endl.) G. Wilson, stat.nov. E. banksiiA. Cunn.ex Endl. in Erdt :,,r]:;Fg.rll ?t. Huegel15.(18J7) lnarned afrer Sir 'T:it.Xfilji.*o"i;,i?J;,i'il;.J,i,lii.'fi,L'liTlf,To,F,l;;'idi-i'3.tird63j; !',"f,'"l or EndeavourRi\,er N.s.w. fl:1,;$.i1"U"13:tll,.shore 1a.Cunningha,n Juri 181e.),,. .2 L:aves broadly elliptic,to 3.5 x I cm. [ 5-nerved,apex rounded. DrsrnmulloN: North-east coast of Queensland-Map 3. RBiili'-fJ,?;;'il;,:.HlTill:l,'f,ir::i li"i.lf,flXi,1i.,#lfiR.ii, 1770 (MEL and 24 'ec€Jlns lerx"Pa eq:l uo pees eql Jo 'lElcgledns eseqeql spJ€,rolpunoJ st pue ulq]'aqd€J eqt pue ,{lr,tecuocou sr qJrq^\ u snaPlaqwo|.t lll punoJ\t tu-loJJLllJ.ll\a uv ulEl"qJ Jlll 8ul rjJql J 'Jrldrlle-,\\olreu -rncsqo,{lelsq'utql sr Surro,r.oceql puE '^\ollErlssr ^lt^12JuoJ;Lll 'urql_e sr urnlrq Jqt'stptouawoid,f.t4t u[ 8uuo,\oJ snoecelroc f,luo seq E 'g lnq ,ttiiiruor reltuts e serdnllo eqder or{1 stUPtuaLuoi u1 @luuo!1p 7 ur sr iere,toc flelelduroc lou) sur8rtur I€lalel pu€ le,t\ol eql ,Lrore,{?l snoec eql -ilsnri lcrqt e ,{q pera,loc sI puu tll€eueq ,{:}Il?cuoc daep " sordnccosrldul ieere pr611epE sI unlrq eLIll.DLlplD8'I uI Suuaôc snoutrqueu ulql t; ,{luo '(ra,{el pur ericier-lercgr6ansurq:l r qlr,t stlJoJ 01 snosc?lsnlc'{culq'PJErlu,{q fero,roc eqder ueluns,(ldaap e surltlcrtl'v') stlutollip'g ul punoJ:l"q1utolJ e8uu.r il.rrqnp:rsoreq BIlull\nVu.rJl(JM ur punoJ.llJJdsJtll salPlStllJl'Pl f,qlJo saDedseql ol {L.-'LnJIlJEd cJllddD J\oqe uJ^lUpJJs JqlJo uollollJsepeqf '"qr?Jlsnv 'solseds uretss^\ uI IT pue "Il"JlsnV rl]alsEeuI S 9I ',qrÊJ epISle,,dol uo "z"lsqJ :SuLIeAoJprollap erll pulqoq ,{ll^€c Jo '{r"lq 'pedtqs-pto11ep'p"o.Iq deap e 3ur11goqd"r :ra,{"I ûoq ! dq pepunor '-lElnJllc'JIBtLls -rns ,11znsnpuc eczj sJlueJJaou unllq :eltSnlJoc lerxepeJo 'ulql ,(TesJa^su?r1ro 3so8nJ ,{TJ"ln33-Ilrlo qlootus €lseloielas :lueledsuell ,{is,\elset Je]no:luesq" Ir? :?uol ruru 7 7'(,{letlue8uPl ruJoJl.l,{d)uloJruei + 'JLrssJS 'snousJqurJru pJJS JtIq,\\ d-recopue1e1uace14 Sutpvards ,{IIISITS Jo 'olerlsor 'prlos l.rjro = sllJJo.l xode elrrels " rl]I^\ sladreJ:el"lncldB-3]Iq,\\ Joqlue :esolld 'peue]l"U slu€ul€lu :st€lad u"_rll lel.loqs ,{ll?nsn lo 'el€clJqrul _'pe^.rnf,ut paie sueruels :urqtr,t osolld"d Jo snolnroqnd sp:led :as€q lE eluc 'esEq '?llsses -[q!ur 'e]{ slBdos 1Bel€loeF€.lq]q ^lelnultll :l€ullulel + '{aelq 'll"tus laclPed .ro ,fue11rxesre,uolg selndrls telnqolS tllr,'!\ ue]Jo s3,\83-I 'lpua xo uu11J V ltu olIp uoa)tsol)g i edll

'n,.rrroyu", '€octuerqureuletllrgJelcl Bchecopue :€ItuJoJllld Ilellue8ueleuollJos u! = ou,lul! 'selueSJo^IpJelr^el Ipero '|ltrlso.l rccoJ €lelnarclB_oql€el0qluE :"ljntrul elae.re le^ 'sr1rss3s+ JeA euruelg :snlejoaicr:rqrqelnurtu rseq snllealped elElndrlselnu!lrl adeese^J3d eIIoC

'etcl -otrel ot se ]ou d (298T)IZf: I ].rA loJ Srpul ld'llanl l C o*"uor.,, ,""" ,o*"r"?u?.|] '{r"lq lelndr]s : ?lndlts - re8rul '4 €Aou orlJes uosll/& D eelndpsorSlp 7 ollcag

',(aupfs Jeeu cl pu?lsueano q ou u-ro-rjeurlcodo] snonuDuoJ 9 peru.IoJecuo ,(lqeunse;d puc (adrot{luels leau ruolJ uorFalloc " Jo Lrorldecxeaq] q1,tl) lseoc eql l€ou punoJ rlloq eJB ,{aq1 (pe,rlo,tureJB srclcuJeric Joqlo wql el€clpur f€Lu IEI.IeleruJelleq :alue:}s '4$luDq .ro pnq ur Ip dsqns Jo suollcelloc ,t\oJ? .{luo ueas 3^"q i) od?tls J€€l ur ,{luo .raJrp ,{lluoJsdde pu" t?lIutIS '{ra^,(lleolJ or" serJedsqnso,tt] aqI 'eul?u perlslTqno^lprl?^ s.uoosled erougr ot s:lsrrl€loq Jeqto pa1 e,teq flut 1! lnq ellopu"J op Jo ose, erll uI os ,{lqeqord se,r qcrq,t'a:}ou ,{lrlBcols,qlrurs ulo4 pe,\uap euo perlslTqndunuE 3q o} urnrsel?rlsne,,leqlrda eql paunss? oq,\\ (t981) tu?qlua{ fq pelou sB.llstql '... 'g snnp url ropull .{ru,(uou,{sLrr ..'tLrS wnlsDlD.tlsnD ,, pery (rz8l) IlJeED ta !r e11opue3op V (OOgt)qlprs ,{q popr^orc auo ra1u1oq1 io leqltde s,reu -re"D reqlre p3sn alerl s]slus]oq ,{lluenbesqns ocue.leJeJ€ rl}Lt,tIInJ uI srsou 'uoDducssp -8rrp cgrceds s,[oos]ed eA33 ptl€ s.uoos.IedruoJJ patl-top flqeurns ',. -erd urnleloecuel ,, torlllde culcads ,lleu e pept,ro-rd(108I) leulJeeC C ) ',, 'loJ srsognrqnssrt"loecu"l ,, srsouStlpculcads l€rloIllpp€ eql rlrr'\\(86/ t) qIIUS go leqt slredo: setcedseql ro.J(908T) uoosrad ,{q ue,tl3 uotldrjcsepeq1 2. Eriostemon linearis A. Cunn. ex Endl. $;,i!{-{5ifgl.it?!,: #;,*ti:i[i;i:,ii!,!'?i;;]tlT:iii ""lilti1'l yil?:i;: i6l6ffiif,l'!:'Pi'Jillli,l,i5lTii,jliilh]'"i,)f;y,.!::tll;%6ii,oi*l"lpr '.r s w,r.. il'i:l;"ji:li;,"*eii,ii"if."';;,:1,J.".i."[:,;J"-':'i)';];,,1r1;"iy?[i"ili],r"r"ji?',lil-' F. Muell.,Linnaea 2s:376 (18s3): f .^11,(fri{iir,- c. Muellerin watpers,Ann. Bot. syst. Itij:?f ?iil# j,:':ffl,ll?lr'#j,,$;i'J\x',i'i.ilii:;lilgt *&:l',i,rnErders-,an ge Small shrub. Branchlets densely -wittrglan<1ular_ve[ucoscJ glabrous or puberulous y-oung_. llinytely rvhen teaules sn,att,!ful-f.,tiofri,lur.tlprf.r; tamlna seml-terete,frorn 0. 8 cm long and fleshy with ioulae,l-apei up to 2 cm t-l:c^.ii^d \\ ith acuteapex, imooth oigrardut", ;;;;;, glandrilar punfiate. gra.broas,:1:lq:l upper surface -! flalteled and canaliculate,lower surface roulded Flouers axilJary.solitary; perlicel4-6 -,.,., Iong,-'tl,i"i.. to*u.A, ape\. lnrnutety puberulous or. clabrous,-minutely bracteolat;'i.i"r, at base. Sepals, .j,^t-.TT fli:hy, glabrous, rj1':";p:L*ti::* ]olet "ii-lui". ,iJ?:,'.T,"-#l"-airow_euiptic, lno.re J,l*f:iT'ffi fr"Ftdenscl)50"ii,l!i!lrii'".?]:"iilini?''fi to$urdsbase: antlrer,.uborbiculrr cl. I mm loi wtnte,apicLrtite,p"tr." i..p' "i"rge. Disc narorv, slightlyurS;#i" ii..tti iiii'l;'.;+iil.rll"#il#if#;i,T,T,,'f*J,rl:?,?#'*iil,,Ttiii pronrirrentlylobed. Cocci sprea o'o'"'"n'11rosttate. See.r,ubreniform.-ir,i.r,",-'u, l;;;:rLi.if':1"il'loJ!nt

,?Ti:'?iJi'J,if :fli"H:'#l ft:r,"t.^:l-1";f,"'i^9;il1:,#'$:.ltl - eastwardsto Wilcannia-Map 12. lfI ror )iilSiilil'yifii-*:111.11i13.1i,tf 6:yffi r tNswl'lhacka'inga, r' E.carne ;%ltii;iiilTY;;,Yl6,J:-".:XlT$:l'.1'.,i: q. Houard.Li\ ,ADi H,.lili5.1l,r),:;J,liix,i,i,,^.ilfflif,. iliiiir!6irf;l rco5 vi. rrrr,iir,".t"","ia-R;l;'i:B.'afi;;l: Tlris species_exhibjts comparatively-littleregional variation. The speci_ i!'d,i:t?$ixi'J, iii,il."*i#:'tllll.r:k*-nf .*rjrffkJ'fl:i :t"i],:?',i,*"i:-"?lf1""'ii€#:l,X?:,sL*tr?lt'".1:i'bolii.tion.r,on' , ,9 ljl*:y maybe distinguished from E ai11r,iii, *iti- iiiii it wasunrted P",Y.'iili,ji',1 ?T; j ;.l, il :lT l J li and by the Ji.. il"T :Iij:ilx'Jl f *:n : l Ll l"li slenderstaminal filaments. It.is the only -..U"i ii'La. tvig__ stipulaefoLrnd in the EasternStat( otirercharacteii, iG;. ;lL;;";;u'i.i'n +;llll,f"l;ff ill,:r,li';l''f coccitleus,E. nutans and E. pachl,phyllu.r.

3. Eriostemon coccineusC. A. Gardn. C. A. pl. cardner,Hooker's Ic. 34: 1.3378(1939) lcoccineus : scarlet]. EJ.iH?:""J;.'Jr?':!1,1\,i"-g]i,,ifiiiifieDistrict, near Koorarawarvee, 80mi west or

, Unde-rshrub, Branchlets glandular tuberculate, pilosulose 3;:ffi',::i,"*""*fif, '?ff;*"ff ;xff ,*ltr;*rrt?.".*iil;.i;;ii: ir grooves 26 --l

'alq,r '3uoT .]JJ]J ,JtEJuqulr pu? urqlr,r asolnsolrd -* 9 n, sl€lacl :Brlol ',{rlsau'prollJp .{r1sag ;Buol IIILUI €c.'snolq?I8 slEdes leq,râuos urur g 7 'snoJqEli 'selo,ep?rq .lraja ,,&€1rlos,,ft"lltxB Ieseq elnurlu I?.reêsrllr,\\ Iocrped sre,{\olJ sur8JEruelellnq ,{lluauruoJd ',{qseu pu? ,{}r,Lzcuoc,ro]J"u € rllr,r aceJ.rnsJa,rol :qlootus'xeûoJ f11q311sece;.rns leddn:elelnpun ut8leul ,esnlqo 'urui 'Suolqo .,(qseg ."c es"q pu" xad€ Z-9. I x g-t ,41ttoqs "uru"l :8uol luur I elolled lJoqs € qlr,r snoJq"TA sJ^"a-I .oBB rl1r,r soiuejsercxe ,{ryoc ,1ol;1q 'l"urpnlr8uol '.r€l[pu"l8 Surdoleêp ,{lesruds.sercue.r.rncep-jea1 ura,nlaq se,loorS ,\\orr?u ur esolnsolrd ,(1e1nunuslelqcurlg .rl8rq ru y lrc qntqsrepun .dllllqd ).l uollurege\ '\Erqtsrq ^ t hi .tt0ts90Zq6t Zl urEldpue,ur qF.qLu S l qnrqs uo rrprenlooJlo \\ rru0z D8J rr) lszta6cv t.nd(toloH s elnJ -roe-oqle else]ruelu srnuol lllut .ec \urLLluE,sllPurl JsuJp snqtut3ffLLL.e8uot ru(I 9-0 isn 9.9_t elenueDe-neeulletuelllBlq : e8uol ruul g eJ.Eqle Eleled: snBuolrrlu C-4 ln xn rssarisnllalrped :ipere 'alrelllos 'salEJlrxeseJoTJ esolrJDarelrlelol€J ,e,\eauoc 'pxeûoc 'sD?[npun etsn8ue elor_leJLrll?ul8ed rolrôl ororrodns.€ur3€d snqrul8rrLlr;osntqoocrde.tunr 7_9.1 i 9ig:rBuo1t1o l0lrAelq,'e

'lqnoP ul ull?Luallsnu uoxel srrll snlels etll €are srql ur auop sr SutlJellocierll]nJ 'eplslno Jo '-{.r€llrxe lllun snorq€l8qns pue urqtr,r snorq?l8 + sFled a1rr1,rqtr.r ']u-elnu 'g o.IBsJo,{\ou eql'8uol ruur g ,{1uo1nq sn.rLrlJJo) o1 edzqs reltuns t jo sea"els"rl lr :((IV'qrsH'etlE uoslr/\\)uotseJco euo uo fluo palcailocr-roaq srq erpJ"SlooJ rllnos uro4 eq1 .stlllttldltpod.!. .setJeds e s€ Jo 'luar"d 1rrald lcurlsrp tr pel"ejt e^"rl e^r:lelnd reqto eql 01 elqe lue esn"ceq puB 'uos€orsrqllolI lseSSns lou I ,{1rlzco1srq1ur luos3jdsr tr Jo ,4:}lunruruoc" t€q:} s:}ss88ns qJrq,r suors€ccoo,r1 uo pelcellocu3eq serl erpl€8looJ ulo.rJ erll te^e^\oH 'unuEqreq Jo lso,r lu"ld eql ur paprcepoq louu€c srqlluoDrzrprrq,{q qBnolLlt lo uotl?losrpulr uorlelnu rlbnoJillJeqlra ,snaut))o).!. ruorJ pol?ui3uoe^?rl rlloq elqrqord 'rltnos lEr]l sl ]I IUJ luo{ Jeqlo er.l)pue erpJe8looJ rur ruo{ euo '?ole 06 Jo lsJr 0Z srp.rBilooJ eql ur pelcelloJ ueeq osl" e,teq s1ur1dJelrurs lorl:l"l o,ul 'sLtoinu g pul- saploLltd Suraq sreqlo 'sle,\\ou !. oql pet qlrr',\uowalsorJg Jo sercedseatql eql Jo euo sr s[JJ sueuE]s o]"nuell€-r"ourl eqt pu" uoDlsod ro,roU,{lBIIrxr eql qlr,r soJ"rls sE 'sl"tpauTi{ lurld lrql }r sel3odssel!:ls uretsee eql ()1 pol?lel ,{[lueJ?dd€ sr sercedssrql

'((Iy) uoslrii|d 'ssor3ureqlnos tuI :(CV) fqqsv 'ssorJ 8t9t 'ulldv 'SurprsJo lseo Z9 0t6 V uroqlnosJo lsea:(HIIAd) €96t H I I TEBITD:vr-rvusnv Nrrgrs?I\ 'y dr1,q erpruSloo3 lsa^r sqleeq-puEserll luo{ ,{luo u,rou{ lrrl€rlsr]y urJ1s31Â:Norrnsrd-Lsro Jo 'uoes 'Jeup]€D lou peas ( c epu) a8pe Je]no uo elEJlso.r 'xade [ ,{1poqs te popunor .{11q311srcco3 Surprerds .{11q8r1sseqol eru8rls 'suelllels :esolrddlosffds ol + el,(ls :esolrd,{lesuep elirals Jeddn 'etsJtsoJlBnbe 'q8rq 'ec :lurl',{Jrô ur spJE,rlno Surpeerds rlltu T sled.r"J u"qt repr.\\ 'peo.rq 'lrg S. 'elepcrde-e1rq.u l?q,reoros csrq o8u€ro eled ua1lod f,luoqs ,{re,r '3uol 'ec '3rrol 'e1?nuo11€-l€ourl ruru I srorllur :,{lloo,r ,{lesuapurSr?Lu urur 0I €J sluourlrlg l€urrul?ls rrr8JBureql spJe,\\oleprslrlo os flesreds pu" urqlr,r asollu -uourol 'peJ 'pe1ee>1'3uo1 ruut ot'8uolqo-pzorq sFled l ururp ruru y ec xedr 'lucr-rpor1,{c 0I 'snorq€[8 'rqnpue13 1e Surpeerds €lloroJ e1€lnJrde-)r?lqxede 'ul8reur 'urqt '3rrol 'prollep e1rlor1rc rltr,\ lnq ,{qseu luru E.I "J ,'(lesnlqo sledes xad" pepunol q:lr^\ + png es"q:}s e]3loe}c"rqdl3llurur 'snorq"l3 '8uol 'pe^JnJerlscupurl,{c ',{J?lrlos'^l?llrxe ruu I g learpod:luulnrl sro,rolC 'el?lnJrdi? pu" esnlqo xed" 'eôq" ot"Jncrl"ual ,'(l,roJ.r"u 's,r\oJ{J€lq,{letrnurlu 'srro.rqel8'mLu pue l"U o,rt ur ,\\oleq elelln q-:u1npuul3 ,{lpe)Irzur S. I x 0I towardsmargin on outside,keel prominent; staminalfilaments llnear, attenuate towardsapex,4,5.5 mm lo[g, marginsdense]y $oolly; anthersca.0.6 mm long. w_itha prominent white-apiculum 0..1 015 rnm iong; o.,ury pyramidal ca. .1 5 min high. apex pilose; style alnost glabrous, _g-iq,.ral io'ltamens. rrult and seednot seen. DrsrRrBUTroN:West of Coolgardie Map 4. WrsrERNAusrRALr,\: N. of Bullabulling,in sanclieath,l. S. Beard3327 (PERTH). This speciesis related to E.,coccineltsand may be oI hybrid or.igin. Tle two collections seen do suggestholveyer, that it is'stabilized in the sindheatir aboLtt 30 km west of Coolgardie.

5. Eriostemon nutans P, G. Wilson, sp.nov. [nutans : noc]dingl Fruticulus ramosissinus;ran,li teretes,granduroso-verrucosi, miDute hirtelli. Foria clavato- teretta.obru,a.,ca.,/l ntm longa.glabra. gla n du lo,or errucor,r- 5tbularum ercrescennb magna.ntgTd. glot,u/afH. | /ofe\ \o/tlaflL a\i//aref. nul:lnrer,.pcdlce/lur graci/r',, ca. g ntm Jongus,sep_ala Jate ovara. ca. 2-j mnt Jonga,corncea, corol/a laie cl jinclrjci, ca. jI x 7 mm, petalis pallido-rubris,pilosulis: stamina erecta, lllamentts jineari-linceolatis. dense|anatoj gi{ra,ris:ova,riurn ca. 1.5mff aftum.dense piiosum, carpejlis patentjbus apicitu promrnenre, ol|oDrs;sryrus rongUs, sparse prtosus * " Holotypus:C. A. cardner 12030, Sandyffats 17.viii. 1953,Ninghan (ca. 320 km NNE of Perth),Wesrem Australia (PERTH).

. Undershrub, densely branched. Branchletsterete, glandular verrucose, minutely hirtellousall round. Leaveswith a pair of prorr.rinentblack stipuloid excrescences;lamina ciavate-terete,obtLtse, ca. ll mrn long,-u glablous,-lower surface ,roundedand.rirh larye glandulur \errucosiLie\. p-persur.face nattenec. tto\4ers50tr{ar)- a\rllar). nutanl: pedicelslender- clr. g mnr lorrg- recuryed,thickened towards apex, minutely braiteolateat base. Sepalsbroadiy ov,qtei.cil:2,5 mnr lorg..coriaceous, minutely puberulous; corolia broatlly cylrndrrcal..(not spreilding),cl. ll x 7 mm, petals imbricate, broadly ovate, pale red, pilosuloseon both sideb;statnens ereit, equul to or slightlyeiceeding petals,filaments linear-atteluate, densely woolly ciiiate, antheis ca. 1.4 mm lorrB,very.minuteiy.white-apiculate; ovary broad, l. 1rulcatein outline,ca. I ), mm rlgh. denselyprlose. carpels spreadingand with a pair of prominent rlol]ow lermtnai .tobes: styte long (even in bud), equal to stamens,sparsely prlose;sltgma lobe\ 5preading. flrrit rnd secdnot reen. DrsrRrBUrroN Map 4. This speciesis oniy known lrorn collectiors at the type iocality. It has obvious afinities with E. coccineusand E. pachyphy/lus with which'species it shares the solitary a.ril)ary flow,ers, tlrc corolla colour (in E. cocctircus)-and the lo-ng pu bescent.style. It is peculiar.in having the carpels terminating rn a patr of large spreading hollow lobes which presumably give a characteristic shape to,the frurt. Sone other speciesof Erlostezon have carpels which are slightly-eacir bilobed (but not spreading);these however, are not closely related to other or to E. fiutafis.

6. Eriostemon ericifolius A. Cunn. ex Benth.

{. Cunn.-exBenth., FL A!stral.,1.:335 (1863) [E{ca, a ieath, and folium : leaf]; Moore et Betche,Handb. Fl. N.S.Wales46 (1893);Dixon, Pl. N.S.Wales50 (1906). " Typification: Skirts of Liverpool Plains, A. Cunninghain.', Holotypus: K (photo see!), leg. A. Cunningham No. 13. May 1825.

Shrub l-.2. m high, much branched and wide spreadirg. Branchlets sparsel5glandular-\errucose. puberulous all round. Learer ,hirtly petiolare wrth small. black. acicular to globular stipules; lamina acicular, 4_g mm 1ong, ,{llq8lls'q1ooruscsrp :8uol -- r'o 1"" Ile uI'e:}"lncrd€-atlrl^\,{llroqs'etltô Joql[e :JI€q.reddn ur esold Surtuoceq(11oo,{ utgruor'xedE el"lnqns " o} Jôq! '8uol 'Suolqo-^\o.u€u 'seprs pe,roJ-reu rutrl E. g-g . t Euetu"llj l"ultus:ls qloq uo esolnluaurolesr.rrerTto'qupru rElnpu€13 snorqelS-qns lo snolq€13'{clrll'}uou -rorordrl1r,r 'srseqluE :1" pu? pnq uI ellq,\\'3uo1uru g '1ecpd111e-3uo1qos1e1ed I ',{rlseu :e]"lorlrc ur8r€Lu'(snolnJeqnd ,{lesJ"dsro) snolq"lS 'esocllJ.rs^r?lnpu"l8 '3lloI 'pro11ep 'snoln]aqnd '3sEq ruur 1 r:c ,{1es[lqo sl?des 3uo1 ruut €-Z 1" el"l -oelcerq-rq,41e1rrurur allssas uI sJe,,!\olC el"lJund locrpad:srels&lc l?urulJel'(1"U '8uolt I 'f,rldrTla rz1npuzl8 1c€lq ^latnurul pu? papuno.Ixedp uiru 4 ?c -p€orq ()18[olqo 'snoec"r.rocJo)',roleq pelleuuBtlcol 3^Bcuoc'eôqe xeûoc 'aprs 'snolqel8'ruul reqtra uo seqrsoJlure^ i?lnpuelS rl]r^\ ur8leru I x 8 Bc 'ruur 'proquoqrZ I ',(qseg pu" elerol-qnsol Z x t rc ro el€ô ,{[JBInAeJrr eururll :selndrlsJ€lnqoJA ol Jelncrc"'{c?lq'[l']LUs qll\\ a:]tlottedXlyoqs se,t?a'l punoJ '3soonjla^ 1e snolnreqnd + J€[npu"l8 ,{les-redss1e1qcur,;rg qnJqsJepu0

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7a. subsp. difformis

Leaves fleshy, terete to ovate or rhomboid, concave or channelled below. Drsnteurrol:- Queensland,south-east region; New South Wales; Victoria, north-west-MaD 5- lig],,b'*li1,3:ii\,pllfilx;J.I,.'iti?ii*"i Lazaridesi.:,ldj[{"?ij}y,.:,"pffi'.i:ti,:,1,?l",i.i& Srory 52 IAD). \w SourH WArI s: Cobar. | . Abrahams212 | \SWJ; \] magee.R. H. Cambar rirral76e,;1,*,Mootwjngie,J. B. Cleland:Derri,!ons.n,.,,"ni:l:.llljj:Jli.,1!T,isi;|tll[x'f"',i]fi:8';l: 5.ix.1962 ( VrcroRrA:Sandhurst, R. Thorn,33(MEL); Walpeup,H. B. Williamson,Sept. 1913 (MEL).

7b. subsp. smithianus (Bentham) p. G. Wilson stat. nov. Benth.,Fl. Ausrrat.1:3i5 (1863) !;W1fifrrfiir:f:*:f.n&r [sn,)ithianus,a manuscript epithet Typification:"-QueeniJand. WideBay, W. Hiil;,nearBrisbane, Henne, N.S.Wales, Macleay iilfi:t","*i'il, ,hi.,i"1"!3',"Y8[uing;t;,"8*:.il:;;;i] wid'ji];;, ii""ill""a;G parv(olius E. R." Br.e\ Benlh..t L Auslral.I :315( t86l). ryprncatron: Queensland.Shoalwarer Bay, R. Brdwn (He.b. -shoalwater passase, R.Bf.).,, Holotypus:K Bay mountains.',(phi,i" .ee;j; i;.'iRibtiXi,, rrrgi 401& NSw

oblongto broad-ellipric. . . ., !.ur.. ca. 7.mm Iong.flal (or slighrlyrecuned.l. conaceous; margtn sparsel) glandular-undulate to glandular_cienare. DlsrnrnurroN: Queensland, near the south-east coast; New South Wales, near the north-east coast-Map 5. QUEENSLAND:Watwick, F. M. Baiiey,D_ec...18j5-(B_Rl)j Inglewood, (NSsw);Dunethan J. L. Boorman,Mar. t9II Rock,J. calbrai;li 16.vri.1962(Biij;B-u;;L;il., ii fri*'ji'i, C. r. wr,iti NEw SourHWALES: Macleay R., Dr B€ckler(MEL).

.E. diformis is.superficiallysimilar to E. ctngustifolius, especia llv to lhe sub_ spectesmontanusi rt dlllers liom that speciesin the scatteredstem- p u bescence, in the recurved leaf margin, and in the petals being wfrlie anJ1om'entulose on tne outsrde- d:{o!y.,t, subsp.smithianus diffen.from ^^, ,L-.- rhe rlpe subspeciesin having rar rea\esand ln lhe petalsbeing slightly less romentulose on the outsidel Ihe type subspecresls an inland plant. E, diformis subsp.Jmllr?aras is found nearer l.hecoasl: plants Nhich are intermediarein leaf fbrm betweenthe luo "* found berween.Inglewood ::l:p:.].: and Warwick in south_easteueens_ land.r.e. In a geographrcally'nlermediate locality. Benrham - distingui_shedthe speciesE. parvifoliusby leaf shapeand absence or^aconsplcuous mldfl b : howeverlhese features are r ariableanddo not pror ide infallible characters. He also stated " " .. that. t^hepetats were gtabroii'in E. porri_ foliusand usualtygtabrous in E. difrorhis,ur. iriTiiinii iin tactthe netatsare tomentuloseourside in.the.rypes of both of these.although less conspicuouslyso than in subspeciesdiformis, 30 IE '906 III^ sluuld '"rlerlsnvlrlenqueqee.rt N c €96I sz're^I{ 'ruqnjrs eprspeor 'a8ued ,{Uo'J uN .re,\\ol qlnos'gll9lt96 CV:snd,{loloH 'elellrc-o}euel 'Et"lnqns oldruqe snsre,{ulealdE ?lueluelg :erqelS Ia^ esolns -olld-otrd 'e.rqEl3'erqElS sll€lou 'slurloJ'll.qlun'srLeurtxrll'\lurloj'll.qLun'srleurLurll ElluecserolrllPrlurJ\lrou![ esndsrcrPds el}uer)ur Erlxo eleloil eLPlod t L| 'e8uol 'ojiulnrrde r:u'olnJe oJIde'ElqnrrLpueJ-elsn8u"Brdns 'erq€13'Esocnre^-osolnpqelS :€uelnalce:€ueln.Ice "uelnqolS"lelnqolS 'eJBIu'BAJBd ?ll{ocsoLxe tunJqn urtu_61-7'e1e,re1cLuru_or'-z'el?Aptlie,tt6,^ sorol eullueleuJtuel Ie^TeA 'Isoanl 'drts :erllssesqns:Prlrssrsqns"rIoCPItol rlnsofdrlnsoTld sllsnSuesJclns\rJln< uIur'rso)nrJo\-osolnpuelS rlnLueJ :snlnJInJl '(I€6I) 'uE^\A SO/ lcl,\ lC luleq€rD uot4 sqrruS'g '(8t6I) ')tc€t8 g6t Z pe tr'do : (tZ6I) Zt€ lelsnv S IJ'l.el8 lreg.Ilprdgwold]snlloll^ q'Z '(068I) tZ l"rlsnv S .rlerlxg IC qpueH'41etr [IelpuI'I uou] spuloglp'g 'petl' suorulaedsuErl"rlsnv urolso/ pue selE/ S N 'lcxe'etdd (tggl) 9€€:I lErlsnv l{'q}uod I qluag .dou] snlpllp)ai &^ slt oglp g ',roJJ€u LlEot: runrToJ : sntsnSuEl ',r.ou'ds 'uos1t16 'D 'd snIIoJIlsnEuE uorualsolrg '6

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Leal + lerete or clalate subsp.angustifolius Leaf obovate.concale above subsp.montanus

9a, subsp. angustifolius Lerves terete or clavate. 2-10 mm long, narlorvly canaliclllatc above, rounded below. Petalspilosulose within. ChrQmosone no. n 28, fiorn matcrial collectedat N'lt. Lolty, S.A., llde " ". Smith-White (195,1)as E. blcvilolius DrsrRrBr-rrroN:soLrtlrern South Australia,south-east Queenslarrd, westcrn New South Wales (?), westernVictoria VIap 5. QUEENSLA\D:Sw of Kogan,B. G. Bfiggs, ,l.vi.1961 (NSw); 50 mi w ofDalby,Dar.ling Dowrls Distfict,S. L. Everist2161 (BRI). VrcroRrA:20ni S oiNhill,J. J. Ackland l6 (MEL);l2 mi NNEof Bendigo.H. L Aston30 (MEL);Pafish of Catiabrim,Liltle Dcscrl, IIicks, l1.xii.l950 (herb. tsIicks). Sollr ALisrrl^LrA:Middle River,Kangaroo Is.. E. Ashby 112(NSW) North Tusk Gammon Ranges,Hj. hichlef 12741(AD); Ml. Serle,R. Hill l4ll (AD); Hd. of Senior.D. Hunt ll82 (AD); r1r.8lackSprings, T). N. Kfaehenbuchl892 (AD); l5 Mile Creek,Yankaninna Rangc, T. R. N. Lothian207E (AD).

9b. subsp. mon{anus P. C. Wilson, subsp. nov.

hrontanLrs rnontane.] Folia obovata,ca, 4 x 2 mm, crassa:supra concava, levia, subtus conlcxa, manifeste gland- ulosobullata. Holotyplrs:MEL 4507,Victoria, Grampians, Nw slopesol'Mt. Dinlcult in watcrshedof Dcad BullockCk., " spreadingshrub 3 ft. lrigh,buds red, llowefs pink to *'hite. On almost bar-e,gently sloping rock-face ", 12.x.1962,T. B. MLrir2647.

Leaves obovlrte, cil. ,1x 2 nrm, thick; uppet surface concave, smooth, lower surface covex, strorrgly glandular bullate. Pelals glabrous (or glabrescent) rvithin.

DrsrRrBUTroN: Grampians of Victoria Map 5. VrcronlA:T. L. Mitchell265, 18.vii.1936(MEL); BlackRarge, P. R. H. St. John,6.x.l929 (MEL);nr. Mt. Zero, C. wilhelni (MEL)i Mt. Byron, H. B. willianson, Oct. 1927(MEL). 32 €t ',{€g 'rnoqreH '(HI{3d) t6L,{aq^\aN) eu.{eqJ }eoB:(680t'Ifl l,) ',. sdlllrqds-Iu€q fol€ls !o" :('IaN) Ila.^\!e1,,qreÎd ptued2ift-*lirili5 eqi .rieu ie,tr1 '.6roeD,-,v-'adroq}slle^€u xei,l 'p.re1g:1w'1oepiirse.u )ioorg.,(1ro{ :(HluEd) 996r iiea utnrc jb,r NNui,rqi rr'r.,oqsil^t:(-lthl).toz oN lloJ rlrgpuu 99 oNpuoL r'!! v/d' lrriiriaalEt'qzqrfovHat'rruerrdsaJols€ettDJ99?c'llIHIf,InAoH:vnvlJsnvNrarsl/lt '6 del,,\tr-8ul) e{"'I spJ€,\\ol quou pu€'3cu"J3dsE puo'{aq o1 '€ll€llsllv 1sea,{ugeu,{eq3 uro.rJ ]sl?oJ qlnos eql iteu o]elseiN:Nollnslulslo '^[tlo s]set urqt qll.\\ 33J11plollap lelluec 'lJrql 'eso8nJ tl 8ur^"ol urBrtur pue eseq }e le,{e1olllJq 3 ^q eqd"i 'lJelq ^p3leôc {1-rulrr8arirur.Jro.rJlJ\ SrtolLu Lu S Z Z paa5 al3ucratno" tto :lelnrtrle p uru ot:tratot.l.:o3 Sutpeoldrtrllq8tl' er'u8rts tit.,o,ls ie aterL",.,iradc'qirq I '(lrLuLuns 'tnoL<]uigJl{ls :JlrrJts Jql YJoElil pJpunol slJdiPJ- le JlEtl-JJddn'tlSttl J.oln\olrd,(1:tnuu.to) \no.tq?13 LLulS Z Pl'lEplLuEJAo\-l12\o ::r\nPul '^\oll?u ',{.re,loto ulpr,n ()1TenbJ I cslp :(snolEdesl}u€erll ul os AllueulurolqJ lelalcnurue-8uotqo,{il\oJleu'JSIP a1er1,r':fluo1,.,,u z eql ,llolaqdlal"rpeunur DJlrJstt| sluJLLtEltlI€ulLuPls :ltlJultuo.td Puu lJllll qll-plul tllLlll\\ JsoJnluJtrlol lluriireruspremoi asoiurutot {lutilt.lo} JPlslno sn-otqc13 :ttqn'3llol Luulq uJ 'snolq"l8 'ut8reut 'ate,lo stuted :pepunol ol Jsnlqo xlce aleloqlc ulql qll{ 'sno-rqe18'8uol ,{qs:u'8irot *,u o.'protlaP ^lPcorq sleda5 ruur 7 1 lectped i 'l?urLuia} :lueiqe alrunprd :X.tl]]rlos,{lLcnsn sre,\\olC puelb l"ululle:} ll?urs ':,roqu e utr,* pjpunbr xade Jlell.lclll?ucc{1,mor;tu pue l"U "tol€q pepuno'l 'r1c1lrnd 'elelellues sno'.riy,;1d .te1rrpurr13'8uoluru 8-q Eururel:elelndlls {c?lq ,{tetniuLu'atetorlaci ,(pbqs sa."r:41 salJueJlncopJ"3l uee,lrleq'{{loo Suruiot -jq '.5'ruol LitrieuaqsLro.rqclS lnq snop.Ieqnd slelqcu"rg qnrqs.repun '.900! '(HIUEd) tdes'cln8unluerrof ru rouprtC V J :snd{loloH 6€6t 'tunjle '(oinsolldalnurtu rrrdc l5^) turrqPlB urLUS [ ?c'eFplur"rtrd etprtrr'!.lpd. :etpnL:tte eFuolqool'nBue BluJLuelg:lluJUlLuold Lunue\o :r.lqet3qn.r:,r 'elnrsnrrqel; 'eguol '?]Eô '?ql€ essi.o etsor ieltsjueuiot snlui [e^ BJqelSel]xe uul 9 er "r"ira ,.ittr.r ruur7-1 .n11:rriod :ruetllci' eaSet tolcurruret ':toll oleuro llPurLUJalopuPIS 'lninornro.,tioo:nr-nriurliueS re errild i;.ron. Pl€pulilor snlqn' Erq!13cSuol lutu 8 s'elnJ\nll -JrJl :etp,ndris-or3ruJlnLr!u erloj llqPlS''lloJ qns lJ\ llnr!qno llntxer ::{tlllrJns cu Upl ' 'erd '- ':qlueg ii 1 I qlueq uou] snuolto)at're^ sluttoglP 7 .t0e-626ru?,r€.usnv ur.lse^N rolsru,,,:.{ rl3$,x"Borli5f;:"?"yu,?,#].fir ,,

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ll. Eriostemonfalcatus P. G. Wilson. so. nov. ffalcatus : sickle-shaped] Fruticulus;ramuli densevertucosi, sulcis angustis r]linute puberulis. Folia exstipulata, graciliterteretia, falcala, ca.6 mm longa,glanduloso-verrucosa, glabra, apice rotundato. Floressolitarii terminalesi pedicellus ca.2 mm longus,minute puberulus; sepala anguste triangularia,ca. 3 mm longa,glabra, apice tenuj, carnoso, acuminato; petala elliptica usql€ 7 Nn1 longa,extra glabra, intra puberula;filamenta anguste obloDgo-acuminata, ca.4 mm longa,pjlosa; ovarium anguste pyranridale, apicem versus pilosum. Holotypns:W. E. Blackall9i7,Oct. 1931, Yellowdine, Western Australia (PERTH). Small much-branchedshrub. Bralchletsdensely glandular verrucose and minutely puberulous in nattow lines. Leavesexstipnlate, shofily petiolate; lamina slender-terete,falcate, ca. 6 mm long, glandular verrucose,glabrous, apex rounded. Flowers solilary, terminal. Pedicelca. 2 mm long, minutely puberulous. Sepalsnarrowly triangular ca. 3 mm long, with a lleshy, subterete apex, somewhatglanduJar-verrucose, glabrous; petalseliiptic, to 7 mnl long, glabrous outside (or minutely puberulous on the margins), puberulous within, the midrib not prominent; stamilal filamentsca. 4 mm long, narrowly oblong-acuminate,pilose, anthers millulely white-apiculate;ovary narrowly pyramidal, the upper two-thirds pilose. Fmit not seen.-Map 6. This speciesis known only from the typc colleclion made near Soutltern Cross. In its sepalshape and leafcharactcrsit is quite distinctfrom any other species of Eriostemon. From E, thryptomenoides,which also has long sepals, it is distinguishedby the verrucosestem, long leaves,and by the absenceof a broad dark band on the petals.

12. Eriostemonrhornboideus P. G. Wilson, sp. ltov. [rhonbos - a top, oides : like, after the leaf:shape] Ramuli sparsiveducosi, glabri. Folia exstipulata,crassa, laie ellipticavel rhomboidea,2-4 mm longa,gianduloso-bullata, glabra. Floresl-3, terminales:pedicellus 2-3 mm longus, glaber;sepala obtuse deltoidea, ca. 1.5 nnn longa;petala ovata, ca. 5 nlrnlonga, glabra, alba, costaprominenti; filamenta liDeafia, sursum attenuata, 2-2.5 mm longa,apice sparse piloso; ovariumpyramidale, ca. I mm altum,apice sparse piloso. Holoiypls: AD 96445067,Westem Australia, I km N of LakeKing township,bush l-3 ft. high,flower whitish-pink, 16.ix.1964, P. Wilson3228. Small undershrub. Branchletssparsely glandular', glabrous, without leaf- decurrencies,erupting ireguiarly in black corky excrescencieswith age. Leaves exstipulate,sl.rortly petiolate; Jamina thick, broad-ellipticto obovoid or rhon.r- boidal, 2-4 mm long, margin - undulate,apex rounded,flat on both sidesor roundedand glandularbullate below, glabrous. Flowers1 3 in sessileterminal umbels;pedicel2 3 mm long, glabrous,with severalminute caducousbracteoles at base. Sepals obtusely deltoid, fleshy, almost smooth to slightly glsndular -T

'snSsol 'snsspra '€r3ru urlu 9l-90 snll$rpad seleulqlolt-l soroll eltuouluord equeJ 'seraxe :?lqnrrde-o.13['(ele rq os@clserdns elqelS'€soinro^-osoFpuelS 'Ele^plclqnJglndrts 'selarelle^) olsn?ueBrloJ rlnraqndosreds Ia^ rrqq8 IInuEl{ sn}l€ru I ec re}nlJns [patlrod ,{l orls - sntslncld€] '^ou ds 'uoslrl!\ 'g '4 snlelncltlu uoll|alsolrg 'tI

'sa1ndr1sjo acuesqearll fq puv 'suert ,{Iroc SulleuJall" qll,il slol 'slqed {ceJq 'sel?ol 'stuols -Llcu?rqerll uo sduls tue.uncep uoer8 oqt fq Jo ePISltlopu" snorqep eql ,{g s1ue1dr€l[urs uro{ perlsrn8urtrslpoq,{"(Il sJltads srqJ -rerrro^dN:(Hruad)0l,q?nor.r'v(uosqrc iutqnlAr^,(HllttrXi*l;1&'ri3i{'{l^f;c^;1;; ol lr.{lellnlÎ:(XIVd SDNI') 98SZprBog S I Jo aN lrt O€:YnYdrsnv Nnrlsar,\A '1 deYg 8ut,toc,,!\o3 'Erl"rtsnv o:l qlnos (uolpl"Je9 Jo lsPe) e,relltrtrAlruo{ oielsei& :No[nsltlrsr(I 'uees 'q8lq tou peespu? lrnJj €ul8rls :,uoleq asold ol,4ls:snoJq"l8 ll?urs ',(J"ô 'ulqt Luur I ?J leprruer,{d ,{1pto-rq,{:e,to :e^n"u Sutpeeoxaf,llq?Ils cslp :3uol urlu "c unlncld" ellrl,'t\" qtl,( 3uo1 tuut [ "c.lerlluE snoleledtlue '8uol 2.0 luru y.6 tc unlnctdz ellq^\ luaurluord e qtt,t 3uo1 tlul S.0 €c Jeqlu€ snol€dosrlue:xed" 1? pJunl-aso1dsnoledesque eq:l puu ^lloo,\\,{lesuep3uo1 urur 'ec 'spr?,\\dnat?nue]]" dllenptr8 Suolqo-,{oJt"ustueulslu ' S.E + 'eptslno l"u[u€ls :tueuruord quprru 'etElorlrc 'urqlr^\ snolnleqnd ,{lesuep snorqtlS '(Iurd '3uol 'e}rl.uqLUr'sleled 'e]3lol[]. qlr,r pe8u4) e1rq,\\ rutu 8 9 cndllle 'sno-rqEI8'r"lnpu?13'3uol 'elecl.lqlur'a:.t1 urtuZ E. I r"lnclqroql1s 01o1?ô sltdog '3s"q '8uol 'snorqBlS ls s3loolc?lq snocnpec elnultu Jo JItd € rJll'\\ ruut S-l 'esocnue,\-.rElnpuBf :l?uruJral sre,lAolj ^\oleq pepunoJ 'aôq" lecrpod'esnlqo 'esocnr.re^f t 'sno.Iq"l8 '3uol elelncrl"uec pu" peuall?U Ieppult 13 tuur E-t 'ploôqo-,rorr€u 01 aleJel-rures'alrssesqns'al"lndIlsxase^"e'I '^{lot euloceq uoos qrrrl,r sduls elrrparuja1ur u,\\olq puE selJueJ.IrlcopJEal ueelS lueuluo.Id 'snorq?I3 'asocnrJe^ qtr^\ rplnpuelS stelqcusJg qnlrlslepun lceJe llttuls : osr '020rqco) xeli,r '106r 'rdes 'errersnv urelse/d'suT*Jflgrt'ttffttff.,l.t,tyff?S3; 'unlqElS'unlle ultu I pJ'elEplulpr{drtEI Lunueô:slsolld unrtl€deslue snqrcrde'BsouBIasuep 'el€ulurnce-o8uolqoEtueloplg:lluaururold etsoJ'EInJoqnd €Jlul 'erqelS 'eql? 'D8uol '€crldrlle 'euelrcIqroqns Ertxe urLu 8-9 €lelod :Erq"l3'eBuoJ rrrlu Z-9 I Bt"ô "ledrs -sn8uolultu 'reqrlS snllecFocl:sel€urturel'€-I sololC elepunlor "4uI 'elelnarieueala^ S-I 'esnlqo '€8uol le "incsnruBld"Jdns !u(I 9-€'eeploôqo etsn8ueIe,{ ellejel-Iluos erJoC snqrluaJrncepsrrlo] qe snqrluerr^ srrula€llunt 'JJqelS'rsotnJJe^ Ilnuei :snlncunjC

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',{cuel.rncap JBel ,{uE ecuasqE 'sa^€el 'sl"ted Jo eqr ,{q pu? proqLuoqr peorq eqt ,{q erll Jo aprsur eqt ro3 tdecxo snorq"F, lelaldruoc 3uraq,(q seuo po1tlo: ruor3 peqsrnBur:lsrpsr sercedsslqJ '(HJUgd) €rlt erIou ( t{'8ul) e)"'I:(HINad) 868.{eq.{aN)'Surql?fN ror : (HlXAd) a?rooC y'at"3ap.\eN rur :(HJXad) gSrZraupr€C Jo'urBunqleiA N 0I €82 S '99 Jo E / J V J Jo ]so.l^:OASN) tt8l ou lloc puz puoulrunrc f : vl'Iyarsov Nurrs:rlA 'L d"W-lse,r-qlnos'elerlsnv uJe:lsed\: Norrnsrursro 'UJJSlou pJJs :atelnJrd?,{lpre,\\tno 'q8rq rulu g.Z '€c 'r"ln8u€rp€nb 'lJe.ro lccoC €ru8qs :snoJq?I8 al^ls 'xad? spj€,rol esolrd,{lesrBds 'qEtq 'lEprlueJ,{dIl?urs 'e^neul ',{rBAo 'uqlcsrp ruur I eJ ,{Je^o :elslnpun u"q1rapr,u.tr1}q8qs 'elelncrde-e1rq,Lr '8uol 'er :e8usJo ol€d ue11od ,{le1nurur ruur I Jorllu" :xed? spr?^\ol osolrd-,{lloo,n ,{les.r€ds'8uoJ ruul g.Z-Z 'sprs,rdn et€nuell" 'rBeurl '(uqll. sluerrr€lgl€ultuEls :lueururoJd quprur snoFreqnd ,{lesredsro) snorqelS '1urd '3uo1 'ele,ro 'esocnJJe^ eled o1 e1rq,t rulll 9-g.t spled :8uol urul g. I lra glaber. Sepalacrassa, anguste deltoidea, 1.5-2 mm longa,nigro-apiculata, ciliolata, aliter glabra;petala anguste oblonga, ca. 6 mm longa,intra pubescentia, extra glabra sed margineol versuspubescentia; iiamenta lineariattenuata,3-4 mm longa,in dimidioinferiore lanata, superiorehirsuta, antheris ca. 0.8 mn] longis,brevitet albo-apiculatis. Ilqlotyplls: Norseman,shrub to 120cm, ffowerswhite, on basaliiclavas, 17 Sept.,1965, J. Bale18s (PERTH).

Undershrub ca. I m high. Branchesterete (not sulcate),glabrous or sparsely puberulous. Stipular excrcscenceslarge, black. Leaves narrow- clavate,4 8 nm long, slightlyfalcate, somewhat convex above, rounded bclow, glandular verrucose (when dry), glabrous (or sparselyhirtellous above), prominently black-apiculate. Flowers 1 4 in a terminal cluster surrounded by foliage leavesand frequertly overtoppedby )ateralbranchlets. Peclicelthick, 0 51'5 mm long, glabrous, minutely bracteolateat base. Sepalsnarrow- deltoid, 1.5-2 mm long, fleshy,glabrous but ciliolate,black apiculate;petals narrou-oblong, ca. 6 mm long, pribescentwithin and towards margin oLrtside, otherwise glabrous, white (to pink?). Stamens linear, attcnuate upwards, 3-4 mm long, woolly in lowel half, hirsutc in upper; anthersca.0.8 mm long, shortly white-apiculale;disc nalrow; ovary narrow-pyramidal, ca. 2.5 mm high, apcx sparselyhirsutc: style shortly exsertedfrom oviuy, terete,glabroLrs. DISTRIBUTION:Western Australia, near Norseman. Peninsula,N.E. of Nomenun,ultrabasic rocks, 30 Aug. 1967, P. J. tsarnes7159 (PERTH).

This speciesis known only fron the Norsemanarea where it is apparently confined to oularops of ultrabasic rocks.

15. Eriostemon tomentellus Diel. Dielsin Dielset Pritzel,Bot. Jahrb.36:120 tab. 39 c J (190,t)[tomentellus .= miiutely ronr- entosel;Engler', Pflanzenfanr. ed,.2. 19a 258, t 1ll (1931);cardner, Enum. pl. Austral. (1931). Occ.70 " Typification: Hab. in distl. Austin a Menziesmeridem versus in arenosisllor. m. Oct. (D. 5164a)."Type not seen. E. stowaftliiS. Moore, J. Linn.Soc. 45:166 (1920). Tlp;fication:"Trryning; Stoward,29l. Nungarin, Id., 794. " Syntype:MEL 4547fsto\\,ard 29t).

Undershrub,minutely tomcntosewith slellateand simple hairs or rarely glabrescent. Branchlets slightly glandular verrucose. Lcaves exstipulatc. shortly petiolate; lamina clavate to subterete,ca. 3 mn long, f flat above, rounded below, glandular verrLrcose,sparsely to denselyminutely tomentose, apex rounded and lninutely black-apiculate. Flowers 1 4 in a sessileterntinal cluster. Pediccl l 5 3 mm long, minutely bracteolateat base. Sepalsfree, imbricateat base.broadly deltoid to suborbicular,i mm long, slightlyglandular, puberuloi-rs,ciliolate, minutely black-apiculate,petals ovatc, 4 5.5 mm long, white with pale red centreoutside, puberulous except for outside of promineni, thickenedmidrib; staminalfilaments flattened, linear-attenuate, sparsely pilose, ca,2 5 mm long; antltefsca. I n.rmlong, with a small, white apiculum, pollen bright orange;disc slightlyexceeding width of ovary; ovary narrow pyramidal, ca. I mm high, denselytomentose, upper half of carpelssterile; cocci ca. 4 rnm high,.truncale, shofily rostrate; seedblack, ca.2.5 mm long, smooth; raplre massive,sunken, coveredby a thin layer of testa only.

DISTRTBUTTON:Western Australia, widespread, Corvcowing to Kalgoorlie north lo about 28' Lat. l\4ap{.. WEsrlRNAusrRALlA: Nalkain, T. E. H. Aplin 511(pERTHtl Walsoolan. W. E. Blackall 4008(PERTH);Southem Cross, R. Erjckson.July 1952 (M EL); nr. Sa-ndstone, C.A. cardner 2:0?,(PEB:III);Conet Vale,C. A. cardner11113 (PERTH); Ninghan, C. A. cardner 12005(PERTH); Victoria Rocks SW ol Cootgardie,A. S.Ceorge 423E(pEirTH). _\

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'uoIlEIl?A sI Sullcalloc leuqc 3o ,41r1rqtssodeql {Jrre1co} ^J€sslcJu 'rnornn13 peuaricg ueqtla3uo1 .,aurrn.r-".n ,-uor.:tiipue';t6{e leq^\JLuos'leulou qrnt.u se^EJl eq] elpJEBJooJpuB ol"A loluoJ J"eu urorJ lBllelerrr Jaqlo uI :!;i;^;J; jqr 'ornl"..' ueq'$ sueo.rcsds i jo'"okl ol sp-uodsar.r6iur:o3 rprq,tr sledes pu? se^B€[o1 urnln'Id" iiiros uo air>iczy lsourp st urhlueurnpur eq1 . 3ql fq'Suue^oc esolu€rlo] dlslnulru IcElq eql fci pu'e'so,tie1 i1e.'r.elcfproqs sir r,i sirceds-pereyarruo:; peqstn8ullslp eg d?ul sryPiualuoi uoualsoltg Undershrub. Blanchlets moderately puberulous all over without leaf dequrrencies. Leaves exstipulate,petiole ca. 0.5 mm long f appressed1o stem; lamina fleshy, elliptic to obovate l 5 3 mm long, flat above, rounded below, glandular punctate, glabrous or with ciliolate margin, apex rounded. Flowers terminal, usually solitary; pedicelvery short ca. 0.5 mm long, densely puberulous. Sepalsbroadly deltoid to suborbicularwith a fleshycentre, and abruptly narrowed to a prominent green fleshyapex, in all 2.5 3 mm long, thick, smooth or slightly glandular, glabrous, ciliolate; petals ovate, ca. 10 mm long, white to pink, densely sericeouson both sides; staminal filaments united for ca. 1 mm at the broad base, flattened, attenuate upwards, 5-6 mm long, denselypilosulose; anther ca. 1.2 mm long, shortly white-apiculate,pollen white; disc glabrous, thick, wider than ovary, undulate; ovary narrow-pyramidal, ca. 2 mm high, densely pilosulose, upper-half sterile; style sparsely pilosulose;stigma small. Cocci with long erect beak. Seed not seen. DISTRIBUTTON:Western Australia, from near Geraldton east to near Sand- stone.-Map 7. WESTERNAusrRALLA: Mt. Tallering,J. calbraith 425 (PERTH); 24 mi W of Sandstone, C. A. Gardner2514 (PERTH); 33 mi N of Muliewa,J. W. creen 1560(PERTH); MurchisoD, R. Oldfield(MEL); 54 mi NW of Cue,on top of breakaway,T. E. H. Aplin 2524(PERTH). Eriosteno sericeuslesembles E. thryptomenoidesin vegetative morphology and in some floral characters; it diflers from that speciesin having large flowers with petals unifonnly white to pink and completely sericeous outside. It is unique in the genusin having the staminalfilaments shortly united at the base. In this characterit is similar to the easternstates representatives of thc genus Philotheca which it resemblcs also in vegetative morphology.

Sectio 3. Erionema F. Muell. F. Muell.,Pl. Indig.Col. Vict. I rl21 (1862)lerion : wool,neDa : threadl. Branchesglabrous or with scatteredsimplc hairs, sometimesglaucous. Leaves small or large, usually coriaceous, exslipula(e. Flowers axillary, solitary or in cymes. Pedicelpedunculate or rarely sessile,with two or more basal bracteoles. Sepals free, imbricate, coriaceous. Petals imbricate, glabrous(occasionally sparsely hispidulous outside), minutely papillosewithin. Stamens incurved, shorter than petals; filaments flattened, abruptly narrowed in upper third, pilose;,antherglandular- beneath the white apiculum, pollen orange. Carpels with a solid sterile apex. Cocci oblong, erect or slightly spreading rostrate. Seed flattened-ellipsoidal,3.5 5 mm long; placental endocarp (on dehiscedseed) membranous, easily detached; aril a narrow, fleshy cord along adaxial margin; outel testa somewhat coriaceous,sr.nooth brown to black, glossy; sclerotestasnrooth; hilum a prominent linear groove on adaxial edgc from micropyle to seedbase where it broadensat comntencc- ment of raphe; raphe basal fleshy,prominent, covered by a thin black crustaceouslayer; chalazabasal near abaxial sidc. Lectotypus: E. mvoporoides DC. Eight speciesof which sevenale found in easternAustralia and one in Tasmania. The speciesof thjs sectionhave a uniform appearancein stem, leaf and flower morphology and ciiffer from sect. Nigrostipulae most noticeably in the seed. This is flattened,rvilh a linear bilum and corjaceousouter testa; in sect. Nigrostipulae it is thick, with a deltoid shapcd hilum and thin outer 1esta.

18. Erioslemonmyoporoidcs DC, A. P. de Candolle,Prod. I :?20(1824) [Myoporum, a genusof treesand shrubs, ojdes - ]ikel; Sprengel,Syst. yeg. 2.321(1825); C. Don, cen. Hist. l:792(1831); Delessert, Ic. Sel.Pl. 3 t27.tab. 47 (183'7r|. Hooker, Bot. Mag. 59: 1ab. 3180 (1832); F. Muell.,Pl. Col. Vict. I :122 (1862);Bentham, Fl. Austral.1 :313 (1863); F. Muell.,Nat. Pl. Vict. 1:71 (1879); F. Muell., Fragm,9:110(1875)l F. Muell.,a.' Srrt V]"J Pl. l:143(1887);Stirling, P.oc. Linn. Soc. _:

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'90t 'ltoHee^oN polleqel'1t€t (06It ') :sed.{}osl.iI9H ,. t""f-i:jjj oN :no9Ig,,'uoq 1gt\l 'ls{s' l19\l snltoluaug '(is8riieZ: illsrH 5 :Qigti wt:i/ilaet\ 3u ds xe qe$ '(uaas oloqd) .. ZZSl lro tS oN Lueq8uluunJ 'v,, ') :snd.{loloH .. reqo}.O ul sre.uolg:;ell1 s.xoj tnrg {IJolL' : uoll€rgr.d'{l -rdsn.,, (zt8l) -1o{ da{'sredl€lA:(1.€8I]t1:I1!l"l}9q:9jltl'{:(It8I)-z6r:I rtroa-bt09:r'rsrs lp1erg 'uImJ g 'isrr1tied :Gz8l) Iat sep6 5 1.1liielq r3oa9 ul v snvptdell 'lu.rlo '(seesoloqd) su€J 5p snl I ')(r-D ., lZ8I€^oN "i;;;,ii??1t.ii.& €lgJ lloH ^noN,, pelleqel :snd.{rotoH,.IeruetroIIoH :3iS:S,;'".S,jqH:li,;t""'i,",'i,?s,;3f!frZt'rirt.i'.y*Tii''*"l}1ff";311""J;f;:ri .l;'s*;g;{lffil'sfl"$il:i'".r.,:ryr,'$#".;i,s"J"Tl}J.ill:a;fij;ldi;TJ[xTEe$

I 3. Leaves chartaceous,oblong-obcuneate, (2)2.5-3.5 cm long, apex rounded; peduncle short, 1-flowered(Glasshouse Mts, Q.) 18f. subsp.leichhardtii 2. IJaves + concave(at least when dry),2-4 cm long; peduncleshort, 1-3 floweied. 4. Leavesobloog-elliptic, acute to obtuse,to 4 cm long; pedunclel-3 flowered; stanlinal filamentsciliate and pilose towards apex (centralN.S.W.) lEb. subsp,acutus 4. Leavesobovate, to 3 cm long, apex rounded; pedunclevery short, usually l-flowered; staminal filaments very shortly ciliate and sparselypilose (or epilose)at apex (NE of N.S.W. and adjacentarca in Queensld) 18c, subsp.epilosus 1. Stem almost smooth. 5. Leaves coriaceous,elliptic, conduplicate and falcate when dry; peduncle thick, short, 1-4'flowered(NE of N.S.W. and adjacentarea in Queensld) IEd, subsp.conduplicatus 5. Leaveschartaceous, narrow-elliptic to narrow-obovaie,1.5-2.5 cm long, deeply concave above; peduncleslender, short, l-flowered (coastalSE Qoeensld) 18e. sulrsp.qu€enslandicus

18a. subsp. myoporoides

Stem moderately to faintly verrucose; leaves oblong to broad-elliptic or broadly obovate, apex rounded to acute, 2 l l cm long. Peduncle 0 5-20 nm long witl.r (2) 3 8 pedicels; staminal filaments ciliate or woolly-ciliate, slightly -Dilose lowards aDex. Chromosome no. n 14, from material collected at the Nepean R., fide Smith-White ( 1954).

DrsrRrBUTroN: Queensland, extreme south-east, far eastern New South Wales; eastern Victoria*Map 10. QUEENSLAND:Mt. Barney, 2,500-4,700ft., S. L. Everist 1390(BRl); Mt. Lindesay,3,000fi., N. Michael 2067(BRI). NEw SourH WALESI Burragorang, T. V. Alkin (NSW); nr. Picton, C. Barnard, Sept. l94l (CANB); Cox River, W. F. Blakely, Oct. 1899 (AD); Sugar Loaf Mt., Braidwood, W. F. Blakely, Oct. 1890(NSW) jTinderry Range, E. of Michelago, N. T. Burbidge 6658 (CANB); Bargo R., Tahmoor, C. Burgess,3l.viii.1963 (CBG); GroweeCk., w of Rylstone,H. S.McKee, Sept. 1952 (NSW); Happy Jack R. gorge, nr. Junction Sbaft, 3,800 ft., Snowy Mts., I. (i. Filmer.26.ii.1957 (NSW). A.C.T.: Upper CotterDam, E. Cauba,14.x.1958 (CBC). Vrcronr.r: Bogong Ranges,nr. head of West Kiewa River, J. H. Willis, 12.1.1947(MEL)i Mt. St. Bernard, C. Waller, Jan. 1887 (MEL); high mountains nr. Omeo, F. Mueller, 1854 (MEL); nr. Port Phillip,F. Reader2 (MEL); Mt. Hotham,E. Gauba,9.ii.1955 (GAUBA).

18b.subsp. acutus (Blakely) P. G. Wilsonstat. nov. E. mJoporoidesvar- acutus Blakely, Contr. N.S.Wales Nat. Herb. l:124 (1941) lacutus : acute,referring to the leaf shapel. Typification: " Shr'ub 2-3 feet high and about the same across", l0 ni north of Crenfell, R. H. Canbage,1.5.1900." Holotypus:NSW 68728. E. attii.r Sprague,Gard, Chron. 1II. 33:307 (1903). Typification: " Thisnew Eriostemonhas beencultivated at Kewforsome considerabletime. .". Holotypus: K (photo seen). E. mlopotoi(1es\ar. mi or lnon Benth.l Benth., FI. Austral. l:333 (1863), p.pte. (Lachlan Rjver specimen).

Branchlets densely verrucose; leaves narrow-elliptrc, 2'7 x 4 nm to 40 x 6 mm, f concave above when dly, apex acute to obtuse, muc.onate; pedu[cle 0 5 mm long wiLh l-4 pedicels I 10 mm long; staminal filaments ciliate (sometimes sparsely so) somewhat pilose irt apex; fruiting cocci spreading with a long, acuminate rostrum.

DISTRIBUTION: Central New South Wales-Map 3. NEw SorIrE WALES:Kamarah, W. R. A. Baker, 17.x.1917(MEL): Crong Grong Range, F. S. Bames, 16.ix.1952(NSW); BeelangraSlopes nr. Grimth, N. T. Burbidge 6424(CANB); Rankins Spring, R. Jordan, Aug. 1953(AD). 40 It 'OAS$ coo 'reBls 'f "d €sooN :(IUB) 116l 'q8noroqspue'I'q8norc o{"'I : (Ng) g'J 'suorlEnlls'sttollEnltsuedouodo uIor ol s,{ex I'€qrr&q}ooJ tll€ 'tAPrEqqnH'"{J'')J { a ) ') qe,riieaigue ueaq :1ixgjj(rrq) 616I .5atra-u-er;:(.eat'srtuer'{.eC'srtuelJ fl(I ,Â oopuooloopuooJoopuor : OUg)(Iug) 9t6I ldeslots'su5ul{Cldes'su.EelC S WsrW 'Ien Ig) 616-1616I 'slell'sleula 13 'sselg'sserg 'f 'I'€lpur.I'I'€rpunolEJ '€rpunoFJ qNYqNvlsNasoo Ieg sears€J urTurtr L :fsNvl): (ANVJ)(ANYJ) t€61tf6T lco urnlle/AlllnlelArllnlelA uo f : rrNdrilro 'II 'pu€lsuoano'pue1 II d?I^{-lculslpd?Jl{-lculsrp ,(eg ept6 lsee-qtnos: NoIlnaIursI(I '(el?rtr 'pera -sor ^l oqs o1) olelncrda f,punlq rccoc SuIlIn{ :xede spr?.&otrssolld + ',(re111os -3uo1 :elerirc-,{iyoo,trsluaurelg 3uo1 uru: lacrped 13uo1 lauruelg 'esocnJts^9 E Intu(g)t-g.0 'repuelssslcunped 'JôqEe^BJuoJ Allqirls Jo qloous 'elnce 'esnlqo 'ulur 'slBôqo-/doJJBu 'etreuorcntx fllroqs o] € 9.I x gz-q I_ o1 cr1d51e-roi;eu sea"al :osocnJJel ,{lelnultu pu? ,{lesreds slolqcuerg '9g€II0 sndftoloH sJe.tô$'qmqsqnsJo gruqs,tol) lsn8nv'(sro^\ou IlIg : .. 's1zuOlurd 'trpunole3te€6I :-eiiij igiis rsue,tg- T s ruhne/ rpues uo uoururocl.re,l ,, : uoqecglal{r

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']I^l '(ASN) plogrelrcJ 9I pl€g Jo eseq : (1gI'{) 106I }rO 'uErlroog Jo N ILU .I f'e)erc iOASN) gI s.{\olpuv ) A llejil^\o( suosllid :$rrY/l\ Hroos,$:rN '8srl.oH 'eq.roqI,{C. {t{S) 196I ( C :(IIS) 'ueaog'I -I'I'etuoqluEls qNv'rsNrroo rx J'suertuv:(IASN) t06I IlnI'seluroo{ : 956I IZ 'I I dBn-puelsueeno pu" selE^\ qlnos ,{\3N u3e,l\lJq edlorllu"ls lsJ[ sulElunou .lepjog :NolJ,naluJslo ' 'xed? sll?q ^[eJ e qlr,!\ Jo asolrde 'sallstJq glls oqs lE '3uo1 qlr,\\ eleJlrc ,{1as:edsslueurryg :8uol ruur urur 7 1 leurlttEls 67 'po^JnceJlectped 'PePunoJ elcunpad :^r?llJos f,ltuzurtuopa:d sJe,\\olC sleuolcmu ol 'llllu, esnlqo xed" x 0Z €c'el?Aoqo se\eJI : esoJnJrs^^lesuap slelqcuBJg '906I 'puelsueen6'erre;u?llErll\ 'ttrun :snd,{losl u€ruroog 'I s ôN îltl./,.8&rt8il 'orqelSoreJ oalde 'epldq esrEdselleurtllels Pluaur€lu :snSuol ruul snlla.rped:snSuol lrlul snlncunped:IIPlios enburnloldselolC oleuorcnlll t-Z 'olepuntor Z-I 'ele 'o.\mcej Ie.\osnlqo ocld€ urtu t x 0z ea oqo €IIoJ:rsocnre ossepITnruEu 'sJI"g [sluetu"lu Ieurur€ls eql ot SuuJeJeJ lnoqlra : snsollde] 'ôu 'uoslll 't8I dsqns D 4 snsogila dsqns 18f. subsp.leichhardtii (Benth.) P. G. Wilson, stat. et comb. nov. E. trachyphlllusvar. leichhardtiiBenlh., Fl. Austral.1:333 (1863) [after Ludwig Leichhardt, the exDlorcrl." 'Fronl Tlpifiiation: Brroa' (N.S.wales?),Leichhardt." [: "Mt. Beer\tah,Classhouse ". M6., Queeoslaodl.Holotypus: K (photoseen); iso. : MEL 4536 Brroa,2.Septbr.43 glasshousiensis (1926). E. " Domin,Bibl. Bot. 89:286 Typificatiotr: Queenlland;Glasshouse Mts., C. T. White 1909." Isotypus?:BRI 042441 "_GlossbouseMountains (Slopes of Mt. Coonowrin)C. T. White,F.N.C. Excursion.Sept. 1909." E. myopotoidesvat. mtTor [non Benth.] Benth., Fl. Austral.1:333 (1863) p.pte., as to Queens- land specimetrcited. -E rcarel lnon Paxton]Benth., op. cit. 334,p.pte. (Queensland speoimen); Bailey, Queensld Fl. 1:91(1899). Branchlets f denselyyerrucose; leaves oblong-obcuneate, 20 x 6 35 x 8 mm, flat or somewhatincurved (when dry), apex roundedto truncate,abruptly apiculate. Flowers sparse;peduncle 0-2 mm long; pedicelsolitary, slender, to 9 mm long. Staminal filaments rvoolly-ciliate,conspicuously long-pilose towards apex. Cocci erect, prominently rostrate.

D$TRTBUTTON:Found rear the summitsof the GlassboirseMountains. Queens- land-Map 1l. QUEENSLAND,Glasshouse Mts.: Mt. Ngun-gun,S. T. Blake 18765(BRI); Cruickneck,on upperportion ofmountain, D. A. Coy,24.v.1935(BRI); Mt. Beerwah,D. A. Coy59 (BRI); Mt. Coonowrin,alt.c.600 ft., C. E. Hubbard4129 (BRI). Eriostemon iiioporoides may be distinguishcd from related speciesby its being completelyglabrous, by the leavesending in an almost cuspidatepoirlt, and by the presence of axillary peduncles (sometimes very reduced) bearing at the apex up to 5 (sonetines more) flowers. It may be divided into several fairly distincL morphological groups which occupy separategeographical areas (except at one point where two ol them may overlap). Thesemorphological groups I have distinguishedas the following subspecies: Subspeciesmyoporoides. This subspeciesis extremely variable, however the variability is largely confined to the foliar morphology and, from study of herbarium material, often appears to gradd from one form to anol.her. The type form is found in New So[th Wales from near Denman in the Noflh to Batemans Bay in the South; it has long, narrowly oblong-elliptic leaves and prominent peduncleswhich bear severalflowers. In the Blue Mountains of New South Wales, in the A.C.T., and in the mountains of easternVictoria, is found a form with coriaceous, broadly elliptic leaves; this mountain forrr could be given infraspecificrecognition; however the flowers are very similar 'mountain to those of the type form. On this form' were based the species E. cuspidatusA. Cunn. (described from the Blue Mountains) ar.d E- lancilblius F. Muell. (from the Victodan Alps). In eastern Victoria this mountain plant sometimes has thick, very broadly elliptic leaveswith the edgespurple tinged; a sterile collection of this form from the Upper Genoa (eastern Victoria) made by C. Walter was described by Mueller as a new species,E. amplifolius. At Mt. Lindsay and Mt. Barney on the New South Wales-Queensland border, is found a form with broadly obovate leaves,rounded at the summit, with the staminal filaments only shortly ciliate and sparselypilose. This is the only forn.r of subsp, myoporoitles found in Queensland. Although normally considered as a small to medium sized bush it may, " accordingto J. H. Willis, Austral. Encycl. 3:400 (1958), attain tree srze in Victorian mountain sullies ". Subspecies acutus, This subspecies,which is found in inland New South Wales, differs from the type subspeciesin having much smaller, usually oblong- elliptic leaves which in the dded state are frequently concave above. The peduncle is usually very short and I 3 flowered. lt is very sirnilar to subsp. 42 \

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I 19a. subsp. scaber Stem smooth, hispidulous; leaves subterete due to conduplicate margin, hispidulous. DrsrRrBUTroN:New South Wales, east coast, near Port Jackson-Maps 12, 13. Nsw SourHWALES: Como, Georges R., J. L. Boorman,Sept. 1916 (NSW); Oatley,J. H. Camfield,Aug. 1893(AD); Panamia PicnicPt., L. A. S.Johnson 806 (Nsw); nr. Liverpool, L. L€ichhardt(MEL);Kurnell, J. L. de Souza,2.x.1950(NE).

I8b. subsp. latifolius P. G. Wilson, subsp. nov. flatus : broad, folium : leaf] Ramuliprominenter verrucosi, minute hispiduli (vel glabri); folia codacea,leviter verrucosa, oblongo-elliptica,ca. 12x 3 mm, sup.aco.cava, glabra (vel spa.se hispidula). Holotypus:NSW 68808,Bundanoon, dry sclerophyllforest,27.ix.1957, J. C. R. Holford259. Stem markedly verrucose, minutely hispidulous (to glabrous); leaves coriaceous, slightly verrucose, oblong-elliptic, ca. 72 x 3 mm, corcave above, glabrous (or sparsely hispidulous). DIsrRrBUTroN:New South Wales, south-east, from Nowra south to Nerriga- Maps 12, 13. NEw SourHWALES: Clyde Mts., W. Baeuerlen33 (MEL); Bundanoon,E. Cheel,14.xi.1925 (NSW); Mt. Bulee,T mi NE of Neriga, E. F. Constable,28.x.1957(AD).

E. myoporoides-E. scaber subsp. Iatifolius NEwSourH WALES: Nerriga, J. L. Boorman,Jan. 1915 (NSW); 3 mi S of Sassaf.as,E. F. Con- stable,20.ix.1961(NSW); Mt. Bulee,E. F. Constable,28.x.1957(NSW); Conjola, W. Heron 34 rNSW). The type subspecieshybridizes wtth E. buxifolius where the two grow together between Botauy Bay and Waterfall. Further south, near Bulli, have been collected plants which match the p\tative E. buxifolius x scqber cross and, although I have seen no herbarium material of either parent speciesfrom this area, I considerit possiblethat the Bulli plants are of the sameorigin. The subspecies latifolius apparently grades rnto E. myoporotules in lhe Pigeon House Range, and near Jervis Bay with E. buxifolius. Near Nowra some collections I have seen suggest that the two subspeciesof scaber grade into each other, the typical subspeciesdeveloping to the north and subsp. latifolius' to the south. In its glabrous form subsp. latifolius resembles E. myoporoides subsp. queenslantlicuswhich also has narrow concave leaves and solitary flowers. The latter may be distinguished by its smooth stems and densely pilose staminal filaments:lhe lwo taxa occurin quile distinctregions. The subspecies/atfo/riis n.raybe distinguished from the hybtid. E. buxifolius slbsp. buxifolius x scabersubsp. scaDerby the smooth leavesof subsp. /alrrfollas which are narrowed towards the base,the lesshispidulous stem, and the sparsely oilose' staminal filaments. I was at one time of the opinion that the name E. dolabratus Reichenb. applied to this species. However I am now confident that it does not and that the plant described under tl'rat name is most likely E. vetucosus. All herbarium material determined by me before 1967 as E. dolabratus should therefore be referred lo E. scaber Paxlon.

20. Eriostemon buxifolius Sm. J. E. Smithin Rees,Cycl. 13: (1809) [Buxus : thebox tree, folium : leaf];A. P. deC-andolle, Prod,1:720 (1824); Sprengel, Syst. Veg. 2:321 (1825); G. Don, Gen,Hist. 1:792(1831); Spach,Hist. Nat. Veg. Phan.2:138 (1834); Delessert,Ic. Sel. Pl.3:27. tab.45 (1837); Hooker, D;r. Mag.7x,4rol i1844); Bentham, Fl. Ar.rstral.1:333 (1863)j F. Muell.,Fralm. 9:11i 44 st snolrtptdsrq{lasreds 'eôq€ poluepul Ierrpad ,{\oleq esocnrrea-r€lnpu€I3^pu" 'aleeunc.lu"ul-9{. -{]11- ,{lSuorls ur8reur "{ou"u ssuq :el"uoJcnlu or-rorur: ateueLt-esocnrr:,t 'ul!u pui p"pt'no.t xade :snolnPldsrq^lesl?ds V Y ZI'8" el€ôqo sel?e'I 'por4'a11opue3 'eôqeeos uopecgltj.{l roC (tz8l) 0ZZ:I ap " N^ s loIlxnq'g pu€uos).efod,"euel,qa.,o^qpelco;:TUi"'i'^*.";ij:qiiiiili.'ili'i1:'".1!r!,!i ': '5seq Ere\oqorrc ienealrqi.. qr!q^\ur '(latre^t:ll.lu l'.t1]lYE klrunlq oqi te troridii ''' ',.-tt 0p ^q re^B i6 u"'iis uotiouiop eqruo peseqj1ai11s1s1111 sntlolxnq ? roJ 0ltopLLEJ otltpie id5iodseqt loi u:^rasl .. 5tru !! q'1"'S': reqtJo uortll.uert e sr'uortdrrrsap :uorlsJuroÎ?:l:l:+tj oqi .jaô*bq':"rlonoio,ii:r.ier 5qt .rri.;pa11j 1e9eteu"r Jo Lrenl3acuor:1ot oN 'lslH g '(lt8]i) Z6Ltl ue9'so( D s'torcqo'&^ sulloJtnlq 'C 'd 'q0Z ',\ou '1"1s 'uoslrid (uoo C) snlEôqo dsqns 'etqelsuo] '0{SN) 096M'82 C g 'IEq 'J€lnJlpurdlrd 'ouolspues'eos III puelP"oqpesoctxe uo shr'f lulod :L dSN) l96Ml^ 9 'uo.u!oJ 'fueliieoq p:sodi::qep'roSleg rd plorqoq:1lU^8) (t ueuls

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runlqqr^\ ,rauueur o."es .{ro^ oql ul poqslo*;,{11noo,,"""offi!i::^i{*t.:'.".{i:!li,ill.3^"o.!i l;1".,i'i'1",:nruru,g{":.:iil,ff'1"#ff.'"[:;$i:$]T;it'';3iilill'J:::::hditil '(t96I) rlsl( p,{SId rs€A qpueH."lE10 rtPlr{:(9161) 9II snsuaC ,eq.reg 0Z€ 'uoxrc rsnlll la uepteli v;6(1j l! sep6 5 4 id :(5061)0t q-,!-!-t, l{ llrsnv 'ie'p"rlrosp"i irrnng ui nertirg :Ggr)it"solelNs Nili qpueg'oqrteg It rroot{ :G/8I) DISTnrsurroN:New SouthWales, east coast, near the north sideof Broken Bay-Maps 13,14. \rw SourH warl\:-Wondabyne. F. W Blaket)and D. W. C. Shires..l0.vi.lq2l (\S\ l; Calga.T).O.Cro\s.28.-viii-tc45rADJ:tlladua.R.\4.Cambage40b7{NSW}:SofWoy\4oy; M. Foote,19.ix.1952 (NSW); FrenchsForest, Sandstone heath, M. Tindale,7.x.1946(NSWI:

E. buxifolius-E.scaber '.- NEw SoUTH.WALE_S:]\lelv.-t_I_gl!enq l', Banks and Solander,1770 (MEL); Nationalpark, !. !: Co]'sq!le,??rLx.t.g:i-(Nq!V); waterfall Road, J. J. Fletcher,l.viii.t 888(NSw); Bulli; A. Morris,25.ix.l927(ADW); Huskisson,Jervis Bay, F. A. Rodway 44-6(NSW).. As lectotype of E. buxifolius I have chosen the unnamed variety described ,,var. by Smith as having ellipticaileaves; this correspondsto de Candolleis 6', and to G. Dot's var. elliptictts.the descriptionin eachcuse beiDg bused on that given by Smith. _ In its typical forrn it is found from just south;f Broken Bay to just south of Port Hacking-. Around JervisBay a form occr"rrswith longei Ieavesand flowers and with hispidulouspedicels. To tlre north of Broken Bay E. buxifolius has narrower leaves and forms the subspeciesoboyetus. Thrs subspeciesappears to representa cline between E. buxifolius sLtbsp.buxi;folius in tl're south and, E. hispidulou; ir the rortlr. Thc subsp.ohoyatus is thereforevery variablc atd could equally well be col- sidered as a subspecicsof E. hispidulous, or the nirme of ihe " tlant could be expressed by a formula, e.g. E. buxlfulius-E. hispiJulus". Tlrjs latter alter- native,although representing 1lte situation involved,is unsatisfactoryfrom the practicalstand point, and in the lield the plant evidentlyexhibits considerable constancyof morphological featuresover a wide area (fide L. A. S. Johnson irr rched.). . Around Botany Bay ald Port Hacking E- huxifoliusgrades into E. scaber. and atJervis Bay irrto l-. sr:a6er subsp. latifolius, the form taken by the planl being dependenton the morpl'tologyofihe parentsand thc degreeoi introgres- sion. Considerablevariability is thereforepresent in herbar.ilm material.

Efiostemon buxifolius Sm. x myoporoiclesDC. '! E. intetmediusHort ex W. J. Hooker,Bot. Mag.75: tab.4439 (1849); Marnock, Floricult. Mag.and Misc.Card_.4:_tab. 49.f.3 (jA:Dr. 1840) (with-out an analysisor-a description); plan- chon,Fl. derS^e.res_5:t.443 (March 1849)(a copy ofBot. Mag. tab.4439);Witperi, ann. (l851). Bot.Syst. 2:249 " Typification: from Robert Barclay,Esq., of Knotts Creen,Leyton,'. Hoiotypus:K {ohoto seen). E. -yoporoirt",,^,. mrirorBenthan, Fl. Austral.1:133 (1863), based on E intermediusHorI., not as to specinenscited.

Hooker consideredthat E. inrtnnediusmjght be a hlbrid between E. buxifolius and E. myopor:oides. The illustration ippears to bc that of a small- leaved, single-flowered form of tlte lalter species; however the branchlets are describedas being pubescentand this points to a hybrid origin cs suggested by Hooker. I have secn no herbarium specimcls fiom the ;ild whicli'agree wilh both Hooker's illustration and descriotion. The hybrid E. bu,rifolius r myupuroidesis reporl ed by BeadIe er al., H andb. Vasc.PI. Syd.Dist. 320(l963), 10 occurin rle lower Haukesbury area. I have not seenany herbadum materialfronr that area u'hich suggests;hybrid origin, but, sir'lceboth E.. bu.xiJbliussubsp. ol'olrzras and E, mltiporoictrs occur.thire, such hybrids conld be expected. "8. The specimenscited by Bentlram under nl,oporoid,s vat. minor " are referable either to -E. nyoporoidcs subsp. leichhardtil ot to E. mloporoides suDsp.Qcrlr.t. q.v. 46 Lb se.{^'uorducsep pyy "9ry{$ .,rose snppdslr? qlr.r prrq{q e ^rluophe'oprJq{q 'rrrrlalSuv{iq'!:-'1!jii a",t i:drbrj'uo-nebiLojit5un red nue;qo un euuror .p nteJl!Eê 't:aaD selsu'p,' :sollergld'{r iijl ,EiSiii'g )":i"irnqr'"rn ui: eidn6nv l l ep serres '",;Yf:lfJC;%tTft:"t1'}"; srodl€,lA'ro[onr,{ J :o98r)96€ 'q€} :t olstrnolcu,p,", "t 'all aJ eq 01 l€adde aseql :palcelloJ ueeq aêrl 'roourl"J spuq,{rl 'roqleSo] $,otB saptotodottu'g_p\tE sn\p!dslll^ I JJJtl,\\ 1" pu" 'dsqns '!' rapun popnlcul sirielunory-en1g eqt \I snpôqo - se rylplxllq 11 re,lo ,(1JI"J-,{uct8o1or1d;otu sl"lr eculs le'ê'lroH e,riq 1 eiie ept,tr.e luulsuo5 'g 'sar""di o^r aili uea.trteqourTc e Jo 3d .{nueplâ s r:snloJixnq Puv snlnp -rdsr4 'g uee,Lr.jeqelslperuJelul sl qcrq,u 1ueld E punoJ sl{€q ueloJg Jo rllJou 'uolllrqlllslP o1 noslcel l.!od luolg ul l"lsEoJ,eloru al? qJlLl'^l|'taqDs'g puu srylolrxncl'!' er" u?ql pu€lul JallllnJ purroJ AII€rauJi sr sJrJeossIqJ ..Bouquorrne:(.rv) rdes'uot'u,EHo '"*",']1?5,1'!l6Jz;i:fu'fll$lSJ 3 | r16l o sqnrq< 'lobiquogg :r r zo6l \'9z sse€JnBJ'-. 'Jlrrds o$l rql qrr^\uollBlJosse ul lerJ^rs 9g:'ttiU Hrnos \\]N .rjrrq,ti Leinte[.. suolrdel :{ IASN ) Z96t r^ Z':503rnB JlooLuqeL:sarvi 'E saPro'rodo{ut x srynqldstq

'(IASN) lI qllus V',{lJaJ sueBlesly' (rlnd pU :(tS\) rro :rA\S\) IS6t\r'0Z uo

DrsrRrBUrroN: South Australia (northern Mt. Lofty Range), Victoria and Tasmania-Map 15. TaSMANTA:Fingal Ck., A. Simson1162 (MEL): SwanPort, C. Stuart694 (MEL); Mt. Amos, FreycinetPeninsula, M. E. Phillips,31.i.1962 (AD). VrcroRr^:Between Hayfield and Seaton, Gippsland, W. Cane,19.iii.1956 (MEL); Murderer's Hi1l,Dunolly, M. E. Phillips,20.ix.1961 (AD); SpringCully Reservoir,S of BendigoT. B. Muir 2555(MEL); Mt. Dificult Rd., Grampians,T. B. Muir 2603(MEL). SourHAusrRALrA: Black Springs, J. W. Wilkinson,12.x.1893 (ADW): Tothill Range,D. N. Kraehenbuehl1091 (AD).

This species shows only slight n.rorphological variation throughout its area of distribution. In Tasmania the leaves normally tend to be narrower than found on the Australian nainland. However on Freycinet Peninsula a plant with broad, imbricate leaves(to 12 x 9 mm) is found, considerably larger than in the mainland form. E. yerrucosusis normally a low bush although in Gippsland it is recorded as growing to a height of eight feet. In the past this specieshas been confused \\ith E. oboyqtus,a plant found only in the Blue Mountains. Its afflnitieslie, however,more with E. nryoporoides as is shown by the stem, leaf, and inflorescence characters, and the similar anthers. I have not been able to trace any specimen seen by Reichenbach on which he could have basedhis speciesE. dolabratus. The H. G. L. Reichen- bach collection in Dresden was partly destroyed in the revolution of 1848-1849, and the remainder, housed in the Institut fiir Botanik der TechnischenUniver- 48 'OASN) xI "I 6t6I ZZ'uosuqof'S'^la{elg V 'uostrtA s33ug 'g'dED ue8lonA:UASN) IZ6t !^ / {'lA ll t :(,{A.SN)096l lx €I '5qJ1.gC 'qlgrqllrelg 'srrvlA 'ere,i-?;in:err'ru I (IdSN) €88I eunf :slN onLg Hroos 'aIIIN 'sul€lunotr '91 dtyl-sep16 l{lnos ,{\eN { enlg :NotrosIuJsICI '(ts6I) elH,& 'tl -ql$rs apg 'ql€eq{c?lq le polce oc ur9{ : u ou aurosoruolqo lt?lrel"ut'saplo.todo{ut !, uI 1"q} 01 .rellluls pees 'eelsoa 'urnlnclde 'popunoJ' oqs E ur Sutleurrural oq-q1lid.buol urlu 9 t Pluoz 'uoqs 'ec jcoia jepr,t .{pq8rp Eu8qs :sno1qBl3 f,ra^ elf,ls rcio3 e1,{1srieqt 'el"lcund e1.ri1s'fepunor e qir,t sladre.r :q8rq,urur I Er-'qlootus ixede lroris ',&e1ledrec-(7) -.re1npu613'luprLueL,{d-pvo:q ot ';cotraqdsrLueq t ^rEô:qlp!lr 'urqt 'ellq,lr ur xjeno SLirliecxa I1ig31s cslp :unlnude el€lound r"tn puelS 'ol"pJoc 'Jlnurur E '3uoT tutu 8c Jaqlu? :pllqtr .I3ddn ur p3l.ou?u ulrd\ !.0 '3uo; '3uo1qo-,tro.r.tuu {1tdn.rqe'1ur3-reur uo Xllercadiateso;rd tutu g [- uJ 'snorqelS '$".r"11'ng :feualciqr qlrprur- qllll pe8up ilirerurlg leuiurils -'ale.rlrqurr :)tuld 'snoJq"l8 jlrqa,r '6"uo1ruur ec ;ct1dq1e slelJd :elelolllJ dlJlnulur f- 'JSEq 'Jpl\\ 'r{3rq uu 'plollrp AIpEorqslEdrS^ l" stTorlJErq t. l-l I t.0 '3uol xrs'qlr,r"irio:qe13-'tpri:trdn relrrqt Suturoceq tuLu 6-7 lertpad l1itls - 'f,rsl1xe 'ut8:eru iiluesqt eljunpad : .{ieirlos srorlolC snoln:eqnd ,{DqBIs 'eôq? 'snoet"uoc e qrua ro sn6rqel3 ',,to1eq osolncnJJo^ ,(pqSlls qlootus 'urrir'91x 'ec 'dleprocqo-oleauncf,peorq e1e1ot1ed-f,1lloqssa^"o-I esocnJJal I 'esoln:luourol 'elejel 'qbrll flocr€cs Jo lou EelqcueJ{ rul o} qnJgs ll"tus ., ZZSI lco 9t oN'tlrBq8uluun] V'selel^ qlnos.{\eN ,surelunol\I uoureti6u3,,p:1eqe1 '94IlI0 : sed,{losl enlg ssMelu,roqo 'u415 11e'7-gEy11w txg :1ueei61oq6j; b-nor1o1o1fi suretunollanlg s.;u:3aa:qt Jo r8rr1,, :uotlecgld'{J ' 'G961OZ,E.usrc p,{S Id tse^ qpu€H te le elp"e{:(S16I)tl€:ot sotej{S N cos uuI'I cord'uo}lllueH:(gqql] q! s9J"4{_SN ld'uoxlo :irngit 'jte,tS qPuEHlqJle8 lr rroo,\ :prlrJueurJsdc lpreqqJre I ol 'E(9/81) 'dier3Ei 'r511inr.f lJ oi i; i1 i j:ara:a tfqgliy66:1 teJrcrrv ll tuBqlurg:- tunreôqo..a1[8]) Lunu lcrHuoD uo6 oqrolBurrr{'r ijleaong;l 1i:q:rlpu'x1:i:5,rurl:{lt8pZ6L:t D:lrdeqsJeJl'plell irlcieL{s-dir-:srp,ro : qol (szsl) l€€ selelAs N rlelÎ JSoeD uI sun3'V 'uunJ 'Y s.rlEôqououalsolrg '€z

'uotrxed taqo)s g 01 perJeJe.l3q plnoqs otueu.loruJoJeql lapun eloJeq eru ,{g ll" eJoJ ,961 ',taqns'8"Peulru.tolep leue}elu -irar{t pue lco:rocul st ur?lrec ^\ou rrre I sIqJ se u,lroul ALuourruoJ serce^dseq1 o1 perld ile iwiuqolop !' eru€u erll l"ql slqeqo.rd1r 1q3noq1 1 snue8 s1q1 Jo l€ual"ru unu€qJeq uo sleq"l ltl?ulruJelop Sutceld se,anI erull oql lY 'pel"quelsqns 0q louuec,{1r1uepr eql qclq^\ loJ-eqr eu?u laITJ"e u? ecnpollur u?ql reql€.I olnl"lc -ueurou pildicce ,{lubiuuroc peulel3r on€q I sorc€dseruus 3q1 ol fldd€ 'g allqla sntoJqopp' pug snso)nJDt 3 soluEueql l?ql elq?qold 1I .lsplsuocI an8€A f,1erue:1xeeJe,{ slu"Id u"ll€Jlsllv roJ setuaJeJer^lllecol ,{lntuer q:}6I ,(l-rEeeql uI sB eou?Jglu8rs olllll Jo ^lq?qord sI srlll reê.{\oH 'pe'le,nocsipueoq ueql peq lu"ld eqt qctq,{ ur ectld fpo oql sP,{\"Iusurs?J st lcerJocur ueeq e^"q eull 1"q] 1€ pll1o,\\ ?tleJlsnv pu"lulBlu ol Suul{JJ JI ',. 'ecuacserouul €rpu"lloH €AoN A ,, ul8l.roJo ,{Jlunoc aqJ eql Jo sllelep Jsllq 3ql op s€ stlsornt.tat'!r Jo 1?q1qllll lle,lr sealS€Jeel eql Jo uollducsep aqJ Erpu"lloH "ôN g- "urlllrurls .t'etlurpnltul '?iuol_llloJlxnq 'ql 'E slTlr'uJqrl ulnuJp sr]ou unjog eJlsEqsle 't JJlrcJlJelrsqo slnBrttloc stu8utu srsocnl;e,l euJelxe'sl{e€l euJe}ul ?JIOC 'qqca stsocn;ra,t euJ3trxasnqluloJljq€loP-opcrldnpuoc srrloJ :.r'?l,rg -Dlop uoualsoltg :Jnlepl^ slTlqBlourolu selcods ?dou slleuei lurxoJd,! -:s,^(\.olloJsE sI pu? ,, q3IS ?ueqcequ3qcleu scaqlollqd,, Jo l?ql raue uo senurluot qc"queqclell ,{q ua^6 uo4ducseP erll pesnoq sI snlJg rlc"q 'UJIAA'turusnW punoJ -uoqrreu Jo ruuJ?qJeq eql a.laq,{\ saqJsuolslqJnl"N eql ul aq fpoc ueuuceds lu"ôleJ oN 9y61 3o Surquoq aql ul peôrlsap s€,tl'lnlls E. obovulis,in seedcharacters, shows evident affinitjes with other members of the section Erionema. Itis probably most closely related to 'E rilgalrr which has a sinilar anther and ovary shape and also has a reduced number of carpels. It shows very little rnorphological variation and does not appear to jntergrade with any other species. For sone time the speciesr. oboralls had been confused with,E tlerrucosus and its own identity lost. For this reasonMueller (1875)loc. cit., regardeda specin.rencollected by Leichhardr from the Blue Mountains as a pubescent variety of -C.o6orrilr,r, whereas it was in fact the true species. It may be clearly distinguished from E. verrucosttsby its smooth, pubescent branches and by the absenceof a peduncleto its solitary flowers.

24. Eriostemon virgatus Hook. f. Hookerf., J. Bot.Hooker 2:417 (1840) lvirgatus : lwiggy];walpers, Rep- Bot. Syst. 1:504 (1842);Hooker f.. Fl. Tasm.1:64 (1855); Bentham, Fl. Austial.1:332 (1863); F. Muell., Fragm.9:111(1875): Rodway, Tasn, Fl.24 (1903);Curtis, Stud. Fl. Tasm.103 (1956). Typification:" Rocky Mr. Gunn (n.485,1837) New Holland,Mr. Cunningham." " Cape, [" New Holland wasi]1 error, the corect localityis MacquarieHarbour, Tasnunia, 6de Hookerf., Fl. Tasm.I:65 (1855).] Lectotypus: Cunn no- 485 (1837), K (photoseen); iso.? NSW. Syntypus:A. Cunningham," rockyshores of MacquarieHarbour'", K (photoseen). Slender woody perennial. Young branchlets blr"rntlytriangular, becoming terete, sparsely to densely verruculose, glabrous. Leaves sessile, larrou. obcuneate to oblong-elliptic or narrow-obovate, l0 x 2 mrn to 20 x 4 mm, subcoriaceous,glabrous, obtuse to rounded or truncate, mucronulate,nalgin slightly glandular crenate and slightly recurved, upper surface glandular punctate, lower surface snooth to slightly glandular verucLrlose, midrrerve somewhat prominent below. Flowers solitary in the axils of the terminal lea\,es,tctramerous; peduncle absent; pedicel slender, 4 6 mm long with 4 or more small basal bracteoles. Sepals f semiorbicular,ca. I mm higl.rand wide,fleshy with thirrciliolate margin, glabrous: pelals broad elliplicca. 5 5 mm Iong, minutely and sparselyglandular, white or pale pink, glabrous; staminal filamentsflattened, oblong-elliptic, ca.3 mm long, ciliate and sparselypilose llear apex; anthers cordate, ca. I mm long, apiculum rounded, fleshy and minutely glandular-punctate; pollen orange; disc slightly exceeding and + continuouswith ovary; ovary hemispherical,ca. I mm high, smooth, glabrous, carpels narrowed at apex to a shofi stedle apicuhun. Cocci slightly spreading ca. 5 mm high, apex rounded to truncate, shortly rostrate. Seed similar to tlrat in E. myoporoides, ca. 3 mm long. DlsrRrBUTroN:Tasmania, near the coast-Map 33. TASMANTA:Port Davey,J. Bufton,1893 (MEL); nr. RenTine,W. V. Fitzgerald,Nov. 1893 (NSW);Rocky Cape, R. Gunn485 (NSW): Coles Bay, H. J.King, 16.ix.1961 (MEL); behveen ArlhursR. and Circular Head, F. Muelle.,1875 (MEL); Strahan, F. A. Rodway1155 (NSW). This speciesis differelt from a]l other rnembers of the genus Eriostemon in having a consistently tetramerous flower. lts anther is also peculiar in the apiculum being fleshy and minutely gland-dotted. It shows closest amnities to E. obovaliswhich has similar anthersand has a reducednumber of carpels.

25. Eriostemon trachyphyllus F. Muell. F. Muell.,Trans. Phil. Soc. Vict. 1:99 (1855) ltrachys : .ough,phyllon : ]eafl;F. Muell., Pl-Col. Vict. 1:121 (1862); Bentham, FI. Austral.l:332 (1863); F. v. Muell.,Fragm.9:lI0 (1875);F. Mue]l.,Key Sysi. Vict. P1.1:142 (1887): Sfirling, Proc. Linn. Soc.N.S.Wates 1l:1057(1887); Moore et Betche,Handb. FI. N.S.Waies45 (1893); Maiden et Betche,proc. Linn.Soc. N.S.Wales 24:640 (1899); Dixon, Pl. N.S.Wales50 (1906):Ewart, Fl. Vid. 706 (1931). "On Typincatjon: the mountainsat the Snowj River,near the PinchRange, on rocks.', Holotlpu.: MFL 45ll: i.o.: K (pholo \een). 50 erT"Jlsnv uJel$l6 ur puno3 'sercedseu6 'Fs€q €zEleqs:Jo,(BIsnoet"lsnJc qloous ulql e f,q osle (pua ',{qseg'les"q pzepqc 1e),(pred put €lsol relno ,{q pePuno:.Ins eqder :l€Icgredns 'JeaurTurnlrq :osoanJro^ "lsetorolcs 'urq1 ?lsel telno :$eJ Suop proc 'Iclql 'eleJlsoJlztxepz '13ale ,(qseg ',trorreu e :ruroJruoJ-,$ou?u + peas ,{luoqs Iu" 'peaJn3rrl + Ictot :e13lncrd€-31lq,\\flalnuru sJ3qlu? suerusls :J"lnclgJoqns 'el"cugurr 'ol€cuqrul sl€das :es€q eql reeu seloap?Jq elulo ,{lpeorq I"JeAes ',tu€lrTos 'snolnJeqnd-al€llots ql$ elrssesIecrped :. relllxe sJe,,!\old ol€lndqsxe ro snorqrlS ileurs se.real :snolrueqnd-ol€llels :o snocnelS seqcuerg 'llrnl J /rJr.rgJ:.dr(f -ntruuouas rl'rlsorrorr^erq 'rlcele + rcco) *,',"",0"11",iiflitX1,t..?g:J.il.S1t,o,uTf'd'J '€,{rnaur€srtuels:snE^o alel'spetrrqrul slJoelae.rq 'sllssas snflec]pocl :soIelllle 'luelr seJol{ 'elelndtsxe'Blnreqnd-ol€Ilots 'e^red los Io^ erqBIS eIIoJ: llrueqnd-ol"nelsle^ linelS llnured 'ecuEJ8s4 [re,trog : soqlu" : eurso] 'd €AOu ollces uosl/d I sollluBltrso t oqces

'eprs Jeqlre uo 'e^€q Je,{\oU" rllrr!\ qcu"rq e 'lrx€ eluss aql uI ol elqlssod sI }I es?3slql uI 'a^rl€le8a^ Jeel 'sJe,$ou eq ^?u (ouo I€rluoc aq1,(geurou) pnq euo lo olul doleêPII3 ,{Etu eseqt :IX" rlc"e ur ulroJ spnq g ,(1ltturou :sercedsslqt o1 rellnced sr J"eI 'secrdB tuerua8u?Jle Iocrp3d oqJ (elelnJld" lou) papunoJ Jleql uI pu? sladlec palrun ,{1yer1redSurêrl ur sercadsuou]elsolJA Jeqlo e uro{ sJaJIp lng'Itlnluetu -npur Jo ecuesq" ur pu" srel3?r?qcIuels pu€ Jeel ur sapto,toaotw'g 01 sallluu" s,rorls elues eqt uo ,{l?^ pu? uorlnqulsrp 01 pel€laJun er? luoprê 11 1ue1d 'a8u"J slIX? uI sre,{\ouJo Jequnu Jo ed€qs J€al uI secuoloJlp 1rl3IS I€cIqd"JB -oe8 s1r lnoq8no.rql ,{Soloqdrou sll w lu?lsuoc lsotule sI selceds strll '(.Igt 'slnll!\ 'lt8!q l) Zg6tlx O€ H f lJ 8 lnoq" seqsnqSulpBerds'epl^d .{ra^ ' reJ'selelAqlnos ^\eNqll^\ roproqsqopoo8 uo'6uIr€uIJ ly{ lJ 000't + lle'puEIsddID'eqneD'E lsee 'ueqtnrg :(DgJ) i96I rx 6T ru'u oqLu€l,:(iASN)Jerl.lelc f f'ooruo:vNorJIA 'rra\oDsueÊ:(osr) 0e6r'rr^ rr 'sso'rns )'uolel'ursr(o*t"35ttri"4-]iftrUft;1':"frt 'ueuroog 'I ',, stttl eql ur s{eer. slu€q eq} uo {Irol qJrr uI SuI^\orB'lJ slueld f 'uenIErV Jo llos 'uepg ZI-8 Jo luoJselnJj ilel,, :(ÂSN) tZ o rôznog op ,{4. V^ lA : sa-IvlA Hrnos ,^^rN '€ oetrI uolslnurs s? quou JeJ s" sel€I[ qlnos ,^^3N Jo ls?o3 ls€e eqtr JBeu pu" €uolcf^ lsa3-qlnos :NoIrnsIuJsI( '8uol 'l." 'sapnrodo(ut ullu !r uI lerlt ot J"lrurs 'al€lnord" 'Surpaords E paas lou (1tq8rls pue gede 3ut111dsJol?l rccot aql 'qfrq 'ec 'pcueqds urru l?q,\\auos lsJU l" elnsd€J qlpr,t ur + eluStls 9 Ienbe'sl[B,\\ oql qll,l,\.snojq€I3 el.41s:eleFcrd€ tou tnq pllos prlql Flpar 'dJBcued lEunu.Iel Suole pu? lruums 1e ee{ Je,{€lurql e ,{q u€rrur rolno rreqg3uo1e Jo '(qefuec osl" pue eseq 1e pelun syed:ec:popunoJ xede uoe,ûog etluet 'snorqelS 'qBIq 'lecueqdsrtueqPUBJo uI al€tsoc .{yrq311s)qloours ruru 9.1-1 ,4re'ro :,{r"Ao SurpeeJxedpq8rls pur rl}r,r snonuuuoc'urql csrp :unlntrde allq,t'lunlq 'elnurur '3uo1 'elep]oo E puu puel3 l?urulelgns " qll& xode uur I Ec lsqtu? :esolrd 'Euol IIIru uI ol?nuellB ,41pnper8 'r€aull 'peuen€U S.9 e ll"q-reddn 'snorge18 EueurBlg :Ioe{ lueuruord e lnoqll,{\ lnq qu Suole J?lnpuelS '3uo[ lBurru€ls'.rldl[e 'snorq?I8 'ur8relu rutll t-9 sl€lod :et"loqrc ulgtr B qtl^\ reppuelS pue ,{qsou 'opr,r pu? q8rq urur 'ec 'plotlep ,(peorq qedeg 's3loap?Jq 'protlep 'll€rus 'snorgEl8I 're1npue18'(]rnlJ I?SEq Isr3ôs q1t,r.1. lII re8uol fllqSlTs) Suol rutrl 'repuols slecped :]uesg" el.unpod :t srelsnlr uI ro .{rE}IIos ZI-9 Jo ',t\ol3q ',{re rx? sJe,r{olC 'el"lluoJcnu 'pepunoJ 01 elnce xede lueuluojd .{pq8qs quprur 'snoJq€lE 'sarlrso3nJJe^ ol enp 'osoln8nJ 'pa^JnceJ,{lltl8lTs 'uru 'es€qIlelus 'etr3ôqo-,\\oJJ"u ul8Jelu aql qlr^\ l?g 6 x Eg o1 tB po,,r\oJj"u, lenper8 ro c4dr11eo1 cr1dr11e-3uo1qo'snoece eqt 'ellssassea?e-I snoJq€I8 'snotnelE 'asocnlJo^ 'aleral 'q8lq l?q,^leluos slelqru"Jg ur ,-Z oeJl IIBtrrsJo qnJqs This section has some of the characters of sect. Eriosteno[, e.g. stellate indumentum ard imbricate bracteoles, but diflers markcdly in seed and ovary morphology. It is also superficially similar lo sect. Erionema in which section it was originally included by F. Mueller, but apart from the characters roted above it differs also in the althers which in sect. Erionema are glandular beneath the white apiculum.

26. Eriostemon brucei F. Muell. F. Muell., Fragm.7:38 (1869)INamed in honour of Lt. Col. John Brucel; Engler,Nat. Pflanzenfam.4:139 (1896); Diels et Pritzel,Bot. Jahrb. 35:320. fis. P-F(1904);EDgler, Nat. Pflanzenfam.ed.2. 19a: 258 fig. 113 P-F (1931); Gardner, Enum. Pl. Austral. Occ. 69 (1931). Typiilcation:" In vicinialacus Barleei; J. Forrest." Holotypus:MEL 4533. Undershrub to 2 m high. Branchlets erect closely glandular verrucose. Leaves sessile,(obovoid ca.2 mm long, minutely stellateto) slender-tetete, acute, to 16 mm long, recurved towards apex, canaliculate to concave above, rounded below, glandular verrucose, glabrous, glaucous, (to narrow oblong, 18 mn long smooth and concaveabove rounded and verrucosebelow, densely stellate-puberulous). Pedicel 0 5 4 nm long, glabrous to stellate-tomentose. Sepalsca. 1 mm long, fleshy,glabrous, with scarious,ciliolate margins; petals broadly ovate, ca. 4 5 mm long, white to pink, glabrous; staminal filaments flattened, rarrow-oblong, gradually to abruptly attenuate towards apex, 1 5- 3 mm long, subglabrous to pilose and woolly ciliate; anthers suborbicular', 0.6-1 mm long, disc thick, fleshy, nuch broader than base of ovary, pale mauve; ovary pyramidal, ca. I mm high, glabrous or sparselypilose at the short sterile tip; style glabrous, ca. 0 5 mm long, stigma lobcs slightly spreading. Cocci, oblong, ca. 6 mm long, almost smooth, shortly divaricatc-rostrate; funicle forming a linear attachmcnt to seed along adaxial margin (aril), which becomes fleshy llcar altachment to raphe. Seed thick ca. 4 5 mm loog. 1. L€avessub-terete to narrow-oblong,5-18 mm long, 2. Leavessub'terete, glabaous (at leasi when mature) 26a. subsp.brucei 2. Leavesoblong, concaveabove, densely stellate-tonentose 26c, subsp.cinereus 1. Leaves sub-orbicular (sometimescaducous) io 1-5 mm long, denselystellate-tonentose 26b, subsp.brevifotriN

26a, subsp. brucei Leaves sllb-terete, acute, 5 18 mnt long, glabrous. DrsrRtBUTroN: Western Australia, between lat- 27" and 31' S. fron.r about 50 to 500 mi east o[ the west coast Map 16. WESTERNAusrRALrA: N of Bullabulling,Sandheath, J. S. Beard3315 (KINGS PARK); l5 mi W of Coolgardie:B. G. Briggs29.i\. i 960 aNSW); Menzies, W. D. CampbeilAug. 1899 (PERTH):nr. LakeBarlee, J. Forrest1869 (MEL); Mt. SinBleton,C. A. Cardner12138 apERTH)icibraltar, A. S. George2698 (PERTH);54 ni N of Mullewa,J. W. Green1570 (PERTH).

26b, sub.p. brerifolius P. C. Wilson. sub\p. nov. [brevis : short. folium : leaf] E. brucei'/ar. dpr),11r.r Blackall, nom. invalid. How to knowW- Austral.Wildfl. 258 (1954), wirhoura rdl,nJ;agnori. or cilalionof Lype Ramuli et folia densesiellato-puberuli ; folia inlerdum caduca,late elliptica vel suborbiculata usquead 1.5 mm longa et lata, supra concava. Flores brevissimepedicellati, gemmis palido- purpurels. " 34 mi E of Mt. Magnet, erect shrub 50 cm high, flowers HolotvDus: J.". W. Creen 1618, btuish:while27.viii.tq57 tPER I Hr 52 t9 'Icql 'PexaUuI '.ult{111 fltdnrqe ro lcore sluoLuEIUleurtur?ls :snorqelS osollld"d 'errl :eseq 'alrssJs xi:rnur'u,sieijo isterlqu''ro stedrs le e1u;oelcerqIlJlnurLu i,4.,nt'1ds ro 'olelndtls r"5'p",i :rJui*r"r ,{reilrxu t.,r^o13 rlerrl .reaurl -xo selee.1 selcueJJnceP-J?o[uea,tlaq s,torrn3 ul esolnsolrd s]rlqcu?Jg 'IPut'I sua&nd g ted{L 'J^Jiq JlE!ull Lunllq :5url'lclJd'snJJclloJ'11? euitueS rlPllsol Joll^olq -qns eelqeceld-sndreaopue:elelrts rolll€orpnlrBuol.ro1|11lqns - tesserr'e'olld -rre^rDrJJo'r elernJrde-oqre:tnu',.u e.r:iiue :otilnrriib 5tonrqeor=lde es.teds ;'r;iL;;;*rrl;i! ,nrqeld'ereir.'n.oq1',indotnuriu irlur Elelod.:srlss:s'nllrrlped:l!Jelllo' 'sipdruriel 'llnsolrd srJLns$ le,l seriilxe seiolg er:eeurf 1:^ Ellolsl elloJ sllsn;ue llnured 'qnlJ lpearq] Eureu ru^ror] 'd 'S ?Aou oucos uoslliA D EruauoudroJ olcos

'"IT?JlsnV ulelseid Jo uo€e: u'tou1 e111q'{1p -crueloq srql tro r?ol?elqu.tapLsuoc i :e,to ,{1eco1:ncco o1 pepedxe eq eloJaJaqtr rdur iurjrd jui "tnts iitt tb l.ru.l srLltqBnir.rqt p:.r:ttecs a:e teql sfu'trelea:q JlluJlEl uo epnu :La^ pire sarredsqnsJeqlo Jql JoJploJJJ ^ltlcJol AueJo quou sJlu not JJAo Luoll JJB ll lo uJes ene'111 suo,lcollor {;uo eq1 sJlJJdsqns Lurrlurunpur oair attijo ladeqs16u 1nq)a'zrs.1eag aqr qttli. sn!Pfllarq'dsqnsJo e1r11els,(lesuep eq1 saulquoc 111"q] ul SullsJleJulsr s'?2ra'rlJsJrJJdsqns erlJ 'peulurexa ou sI eJeq] eleq l lerriledr unlleqrJq 3ql^ul uorlsJ8Sns srtillo:o,\\l:ql uJe\UJquotlepuj8latut etec'pur ,{utu 3ul}JJlloJ roqtrnl pue '{1;e -5r'uiroqpare8irsa,rur rlLian

eJou qlr^l stuJoJ erll ur punoJ Sulaq ,41tso[d- raleorS eq1 pLre larrped ra3uo1 put eql qloq'salft^ oqe guaurelg leu'uinls arh.;6 ,{1rso1dJo oar8opaql lectped Jsntqo puE le)JrLllelE Iaql luPld snolnJJqndJql ul JTrq'tt ro uiSuor:ut 'Jlnre 'reP(rels- ir.,o1 tnbrqi]a:t1 trid,i aLltre'i5unc PuE aq ol purl sr\Erl eqt '."(srusldiunok erit uo) inolnieqnd A1:suapot snorqelS 't1a1:ldutocSutoq urbrr a8uer aseqt sol"el pue tuJls uo runluerunpul Jo lunou? eq] ul uoll€Ilel a-o"s s, ar"q1 qbioqle:iue:eedde uI uJoJIun,{FIeJ sr selcadsqnsed'q oqJ .(HItAd) 'uls $pua^el €t€ uor8sr)ta I oleleg (HIUa4) 596l l^ 8I :

'Id 'leu8"t'{ '(HIXAd) 8I9I-: uoorC I }N Jo g Iur t€ j (HJ{Ad) rZqJll^,tz-'reuprBo V J 'ln,5iipws (lirrEga) 8aiz-uldv H-a J'pul{ seu{edJo /A }ru 0z:YrN{rsnV Nrsrsald '91 de1,q-ecuu.ord u"aElueJg uJolsol\ 'auolspues o:l puIC Fu(ed'"IT?JlsnV uJa]s3lA : NolJ-nalxlslo 'pnq ur aAn?ru r:11o:oc'alelectped f,proqs fron sra'lro1g ']?FclqJoqns 'snocnp€c serurleruos'o,loqt !,lecuoc'?PI,Y\ pu" 8uo[ ruut 9 I 01 or c11dq1e,{piro:q sa,reel : snolruoqnd-el€llels ^losuep se^"a[ pue uI01S apiculateat _apex.glabrous or pilose:anrlrers nrilutely white_apiculare, Cocci dl\ancate.shortly ro\lrate. Seed.somewhalren_iform. plump. 2_4 mm long. u.ith,rhe auached..{placental)endocrrp sub-coriaceousand per5r\Lent;resra DracK.tongtludtna y stflalc.not ec\ily separableinto tr,rolaycrs, sclerolesta Oornrnanl:htlum sltorl. arrou-oblong_in centreof ldarial face:aril ab.ent; raphefleshy, past ing from hilum to seed-baseand corered wi[r a glossycrusra_ ceouslayer; chalazasub-basal towards adaxill margio (seedof El pinoiclesnot knorvn), Three.species,two fron.r Western Australia, one from South Australia and Victoria. . The,threespecies included in thi. sccrionshow morpltologicaldiversity In se!cra.lchilrrcrers but rlte)have l generrlsimilcrity to eachorher qhich i! grcaterlhan thal to specicsin ln; other seclion. . . Tlre t$o species.6. fit.grraldii antTE. pungenshave both at one tlme been lncludedrn Ihegen u\ Ph.l,alitnr.rlrc forrrer on accountof itsgla brous filamcnts {ano irpparenltynot)-tprcUlalc anlhcr.). and the li ler beciru:eof itS \alvate petals. l1 lsapparenl that singlccharlclers are rrotsufficient to assignspecie, to.gcnera and cenitinlyljte characler of \ill\ate peLalsin 6. /4rr?gcr?r;rrsi ha\e arlsenqrllle lndependenlj) 1u lhat lrrundit pheloliurysect. I eionemaas tlteTuo sectrons have otherwlbc uo close molphological sin-rilarity. Chromosome counts for the three speciesar.e unknown.

27. Eriostemonpinoides P. G. Wilson. sp. nov. lPinus, the pine, oides : Iike] t:r curn.ulcisnubcr'r,lr. Inrer sulcos verrucL,\i er gtabrr. l':y]],1:_:_r i]] -olund3ra tolia aL.icutrri3, !Ll^l-.,t']9!!1tc\rter tort?.,,e\rr. gtabrJ. sub_lus \el rriang.Llarra.supra canatj- ',9i!'-l-! peJrceu\.4__- nrrn tongu.. Lrasi rninure bfacLeotir:: sepata rmDflc3rJ.il'jji-.-' tale oblu{o]elrJlntle\: ClellotLlea.J.que ad 1.5 mnr l,rnga.carnosa. glabfa;pelald Obovara cx, o,) nrrn rongr gtabrdt 3rro-rubra. htamelrr petJlin.rdirridio rnferio'e ov.Lto.\upe,.:ore lereti, .Il longa: nlar)'enla .eppl,.ra d; nidio in'eriorc apptrnarn. dirnidro :u|refrore sru(curo.i9:-l.j cfa* cLTt dL.obuscJ\t- laleral.l.. proJ-und;..ci. 4,5 ntm long.rtJnll err l.ilc untonga, ca- I mm longa, minute apiculata. Holotypus: C. A. Gardner 9408, Surnrnit of Mt. peron, Western (PERTH). Australia, 26.\iii.1g19

Small undershrub0.3 0 6 m high. Branchletsrnitutelv glandular verru_ cosc in slrongly dernarca(edglrbrorr. leaf decurrencies,jLrEerulous in rhe rn{ermedtatesurlke.n btrjps. Leavesshortly petiolate; lamina acicular,some_ \rhat pungent. c]. I crn.long.slightly twisted, smooth, glabrous,f rouirdetlto rrangular Dctow,ccnatrcuttte above. Flowers I 3, terminal; pedicel4-7 mm lllgt C]1!l:],1,.seprts.broadly and obrusely_ dcttoia, to r-:'ilnirong, Rcsrry, grabrortb:p.-tals rnrblcate. obo!ate,ca. 6 mm long, dark red;staminalfilamenii drssrmrtar..lheal1ttpctslous o\ate belowand + tereteabove, ca. 3.3 mm long, sparselypilosrrlose arrd abruptely , apiculate,the antisepalousflattened belo#, Decomrngtnlcker c bo\ e !\ ttJrtu,o deeplateral concavitiesand a ventral mediaj 4.5 mm.Jong,,pilose I'dgl :". above; antipetalousanthers broarlly oblong, ca. I rnm tong. tntisepclous.dcltoid,pollen pale yellow; disc very-thill arid con,ttuent\ r1h o\aryl ovarybluntly pyrlmidal ca. 2 nrm high.carpels rugulo>e On Dac[.nitrfoucd InlO a sr]'tooth.5lefile ape\: style,lender.glabrouS . equal to stamens; stigma small, no wider thar siyle. Fruit and sied nor seen.

DrsrRrBUTroN:Westerl Australia: west coast, between Coorow and thc Hill River Map 6. WESTERNAusrR,\LrA: ldi'i?;3.,;i1ji:,ll".i#;:"'i#'121 mi W of Winchesleron Eneabba road, A. C. Beauglehole12121 !"'it"fi:l: Ri"e"" c;;ne'ross iiilrui; 2j .i '9t269 t SN: os! IHJUEd :snd,{loloH .. €06I reqotoo ur oau?redsAJo quou suleld PIr€suo Je/ og ul sebodsslql punoJ L, : uollecgld.{f '(It6I) 'reupleC 69 t O lErlsnv Id runug : lpl"raBzlll 'A l\ €{e poul"Nl (i06I .{et{) /t:I ou cos lsIH l€N'lerlsnv d|I's^\OrpuY d U ) 'JpuV 'd '62 U 3 lppraBzlg uouralsolrg 'el"crJ?,\rperE rcsoc3q1 qloq uI pue'sdq pexegul ,{l8uor1s qtr^\ f,qsag a:e qtoq Jo slelrd aq1 x:de rrrql l" IrIq} eI? qcrq,\\ slueurBlg aql rlo srteq J?lnp[813 :,teq qloq r.tpTola8,zt !j[ J pu? suaSundz '!!ppfi?z|{ lBurru€ls 'g ? Jo }€ql o} lellluls sr }I .reêld.oq:saptoald qll\\ selceds slq} Jo paos eql ereduoc ol elq" u33q lou a^€rl I lltlrJ pu? sueurels JBIIIUISe,req rqctiqtrsaptourd 'g pua tlplDotzl{'!' qll^\ pec"Id }saq sI pue trcuuslp ellnb sI 'poos 'uels 'sJa:lc€leqc 'wnUoqaqd 1l pu" IsJoU :€ol Jeqlo lJ" uI'Jeaa,\\oH Jo DwauoDT uorl:,asaql ur esoql solqueseJ$Icads slql sl€lod ol€^le^ SuIAsquI '((V) u/ôlroprogol uJo}seiA:(OV) x'It 'ptrElalJ .{Uo'I t0€ lunH d le[{ 'rereJ €t6I { f'o8ueu ;unoiruirttns iLteus.roH:(1ACV) 096t ! l€ C A'eFsuluod elio1 reao'I llolueq :(ot) il 'pueie13g 'elnsulueder,{g '€tttJ 01 I"B ,{Iee.r}S:YnYursov Hrnos /s6I €Z I rrellonl '(.IAW) uosuelllr/Ag 'u,{\otedoH:('Ia},,I) €S8I ôN l ^{ 'uoo8rnts 089 l 'qlroN l] I pu€lduqv tW : (CV) 6t pueplrv f uolecl^JesJo N lur ZZ: vrlorJl .7 de1,rtr uor8er 'el.Iolcr^ I"rluec pu? uralsa,{\ : uorltes u]eq]nos'elTBJlsnV qlnos :NoIJ"nsluJsIO 'eso8n:'snoecer:ocdJecopue paqc?lle 'suorl€SnJJoc :qlooLus Jeuur arl] lsulpnll8uol eug pue esJ"oc qll^^ Jelno erl] 'sla,{e1 'uuoJlueJ 'Surrrleru o,tr.13o elsetoralcs :8uo[ tuut t g.Z ,41,no:leupees 'el"lnord€ '8uol 'I"]uozuoq -l g ro 1 ,{1uo ,{fiensn ruu 9 t pu" ol?Jtl"ÎP f,18uor1srcco3 e1,Qsueql :apeorq .(pq311seru8rls :srsoqlu€Suunp 8uru,e_ql8ual 'aluols 'popunol snorqep e1,(1i:lser ulo{ pelscleutop pue p-Inll reddu aill 'snoJq"[8 'qlootus sledr"c lq8rq utu ec pprue;.{d ,(1esnlqod]eô :,tJ€ô u?ql 1 'unlnclde ropr,r 'ueeJa 'qloorus 'lueu[uord cslp :,\\ollo,{ ueflod popuno.r lr?p '3j€nbs 'pel-{l?p 'roqs " q1r.r Suol ullu "c + raqluE :os€q sP.Iarto:tal?lolll3 tlrruuns I ,{1as:eds 1e s.rreqre1npue13 qlr^\ seulleuos pu€ elulnsl}cund .r"Fpu"I8 'ileyncrde f,pdn:qe pu" popunor xede '(3ôq€ ele.Iel SuruIoceq snotedos + '(aptslno -Bui) iseq spr"^rot peueg€U '{cql slu3ul€lg :pexegur d,p I"ulur€ls '{L1sag'6uo1 sdrl elrq,4{'uorl€ue,\_ snol^qo to qup[u tnoqllt ruur re lurd) 'f,qseu '3uol7 'ei 'e1e,rb '51r.tie.r sl"led :xed" pepunor qlr,tr 'a1e1ot1tc'sno.rqe18 'el"ô '"c ruur eo ,(lp"oJq sledeg 3uo1urtu I seloep"Jgel"Ioeau€l'[es€q 8.0 '8uol 'slalqcu"Jq I"raês q1r,r'snojqelS ruut S-€ latrped :,4lelllos ,{ auJou erll ?uo1e qr8iral euros rog '^J?lpxe sJo..itoli qtrptut lueurulord ,fuel qlr,t lueserd 'snolnpllq€cs er"Ins Je,\\ol 'qtoours 'l€U ecejrns raddn ';e1npuz13,{lelnulu Jo snorqzlS '8uo1 LuuJ(SZ) 'lue8und 'J€lnctce€uIL!t?[ :e]elolled [luoqs so^?a1 ZI-8 'sot3ualJncap 'snolnprrq"cs Jo snolnJsqnd sduls el"tperulalul Jrct snolq"lS 'qBIq '"r + pe)r"ur ,4t8uor1srllr^\ slelqcu"rg tuc 09 ol qnJqsrePull '(8n6T) pE do')JeT8:aIi6|) lJrA rreaa:(tZ6I) 'Icelg:(€98I)86t:Z 1r, lq1 lg Ztt l€rtsnv S ll 8t[:I lersnv H'Lucq]ue{({5lpo!-I) sua8u d unllDpqd 'acJ :snd.{roroH ,n 'n,,o,.',.f;u0,'ifir$""i'djf,}t"rR t""{ :..eqclrw ,rr1 r, 'qsnq ',(qqnrJs flT.i;.f,"qsg''uotualsollg rirnttu:eo-o1oe1e 3o e:u;reedde eql qanrrlqttM futds E SulLoloJ ;d selcodsneu ejqei.reurere oslepunoJ e^{ ledoH u l e l] lltutunss]I uO,, :uollErgld.{J '(96et) p : (f srle/l^ qpuPH'eqrlaEle oroo'\l 97 sa1e165 ]d'uoxrC 'rleJ:f6l8l)68T)tt S N LJ 'lleny{ :(068I)tZ l€rlsnv S rleJlxgIJ qpuEH Z1:I lrr^ LiI']eN d :(9r,8I) g:(zq8t)lZtit ts{S log de6 Orf.A':ur6ejg'neirlrt's.redie,\ rrt,t toJ Ld'ltrnl^t J:(tt8l)€28:Z : tlue3snd: sueSundl(8€8I) 99I:z IersnY E padxaoerqJ,'[oq.]It{ ur,(elpul'I '1pur1 suaiund uouolsolrd '82

'u,\\ou{un sI 'pre8er stql ul 'pees erour dleq lq8ru qcq,tr eq1 ,(1a1eun1roJu1] s:olc€r"rlc leuluels eql '{q pue f,Soloqdroru JBel pur urols eql fq polse88nsst rtplota8ztg[ 4 P.uEsua8und '7 o1 drqsuorlelar sll sl€led eqtr Jo lnoloc eq] uI pu? sluaru?lg Ieullu"ls eql Jo urJoJ oql ur qloq uowatsoltg snue8 eql ur anbun sr selcadssIqJ E. aprrcas Diels" in Dielset Pritzel,Bot. Jahrb. 35:321 (6.x.1904): cardnef. I.c. Typification: Hab. in distr.Coolgardie pr. Gilmores. . . . (i.5267)." Isotypus:PERTH. Phebaliutnapricum (Diels) Ewait et Rees,Proc. Roy. Soc.Vict. ser.2. 25:ll I (1912). E. tr'rrlro.li./rLuehmann ex Ewart.Proc. Roy. Soc. Vict. ser.2. 20:79(1907). Tlpificari,.n:"Nor.eman.u. A..J. BJ . l8q_. Holol)pLr\:Ml L4719.

Snrall compact undershrub 30 60 cm high. Young branchlets with stronglydcnarcated, glabrous, r'errucose leaf decurrelcies,minutely puberulous in the narrow, sunken, intermediate strips which become corliy- with age. Leavesshortly petiolate;lamina seufterelc,3.5 5 x t mm, glabrous,r.oundid and verrucose below, llattened, smooth, and sometimeschannelled above. apex obtuse 1o be rounded. Florversterminal or axillary, solitary; pedjcels 1 2 mnr long, glabrous,bracteoles ca. 6, basal,triangular, ca. 0.5 mm long, glabrous.. Sepalsobtusely deltoid, ca. I mm long, coriaceous,glabrous, cili6- late: petals thick, slightly imbricate, ovatc, ca. 4 mm long, without obvious venation or_middb, white, apex inflexed;staminal filamenti erect,contiguous, fleshy,angular terete,ca. 2 mm long, becomingthicker towards apex,trutcate and shortly-apiculatejustbelow the anther,sparsely glandular puncfate, glabrous or wiLh a few terminal.glandular hairs; anther + square,ia.0.5 rnrl long, very minutely white-a.piculate; disc exterdirg slightly beyond base of carpel-i and not confluentwilh them; ovary bluntly pyranridalca. I nrm high; carfels wjth the upper third narrowed into a rounded sterile apex, glabious; siyle thick- f. equal to stamens,glabrous, broadenedat apex into a flat, slighily lobed stigma. Cocci strongly divaricate,ellipsoidal, ca. 3.5 mm loog, aprcu- late. Seednarrowly reniform, ca. 2 mm long; testt with one la1eronly apparent (the sclerotesta),1olrgitudinally strjate; rrrrthed endocarp smooth. somewhat conaceous.

DISTRTBUTtoN:Western Australia; betweenEsperance and Norseman-Map 9. WESTERNAusrRALrA: 20 mi N of Salmoncums, W. E. Blackall994 (PERTH); Or sandy ridges_belweenEyres Reliel sandhunmrocks and thc Bank, Maxwell(MEL); 40 km W of Daniell,P. Wilson3167 (AD).

This species is peculiar in having glabrous petals and thick, glabrous, staminal filanrents which differ from those found,ln phebalium in thEir shape and in their erect, not spreading, habit. In these characters, and in the fruit and seed, it comes norphologically closest to E. putlgens. The la er species has valvate petals; in E. fitzgeraklii they are slightly imbricate.

Sectio 6. Cyanochlamys (Bartl. ex F. Muell.) F. Muell F. Muell.,Fragm. 9:110 (1875). Eriostemo s]bgen.C)i.anochlamyr Barrl ex F. Muell., Pl. Indig. CoL Vict. 1:119(1862) lcyano\ oarKDrue. cnrantls Inanel. ClanochlamtsB^ttling in Lehmann,Pl. Preiss.I :l7l (1845),nomen provisoriurt. Branchlets with stellate hairs. Inflorescencea terminal cluster or raceme: pedicelwith (or $irltout)a prir of basalbracteoles. petals irnbricate. thin, glabrous; staminal filarrents flattened,pilose; arther white-apiculate; disc a narrow ring around base of ovary. Cocci erect, apiculate or ;hortly rostrate. Serd sub-reniforn, abaxial margir conrex, adaxial margin straight; attached (placental)endocarp_ thin, white, caducous;aril slender,linear, along adaxial face, firmly attached to placental-endocarp and easily separaied fr6m seed; outer testa membranous; sclerotesta smooth; hilum superficial, linear; raphe small,,sub-basr1.c9v9r5{ by ouly a thin integument; chalaza near base of adaxralmargln and \rstblethrough raphe. )o LS .(ueos 'tg6€ -raw 'rtz69 oloqd) ) ,{dS,N osr :(ueos oloqd),\At :sncl.{}oloH u Lunr0 erpuelloH eAoN,, :uo!]e.glo^tr : 'GngI)..t6 l\I 'zcrnJ hI.Z/ZZ .sol I l?N cos ll11g st?4tnlr, I' '., 6€8T'puotulunro'loÎU uB,uS,, (SNVJ uoes oloqd) gCJ :snd{lotoH (uoLoJ rr^ld uu^\S txorC :uolleJgld^l 'trrorroi JJo lErlsnvld r!nul .reunrP.):rlt6lr8sz:e6l z pJ'lrldo'lrl8ul:(q681) 6g1iyJ tuelu:zueg4rel8u3 :(s/8li 0ll:o LuSEU'llJnh J :at98l)9ft: I lErlsnv.ll rriirriiu6g:(il-ttgt)-trt:-t ssrojJ1a'uueL qo'I or SutDreS:(zt8l) t6g:I ls{S log dad 'sraci1e,11 'epou ',{elpul'I : lfeie,trog : snrou € .= snpoul (6€8I) II^x ddv 8au log .spJ€^\pa 'Ipurl snroulpou uoruolsoFg '[€ '(selcads 'sueurcedsuJerllnos eqlur ,{ eclpelods sll.Iccoosl€ lelt"l€qc .rell?l sIqJ 'qrrprur ;eq1o ur se su€;eur eql spJ€,!\ollou eql 3uo1e)sFlod eql Jo aplslno 3ql uo ssaurTloo,r-alEllelstrq8rls,toqs sJe^\oueuos pu? qlnos erll ruo{ osoql uErll s3^Be[rop€o]q e^"rl ()1puel uo4nqrllslp Jo s:]lurrIuJeq ou eql tuo{ suollcelloJ 'a3uei pirqderSoe8 sll ]troq8noJql uotlet.rt,l lq8qs tr1uos,toqs setcedssrqa 'C{SN) x 's38lrg 'olsrlemleNedeJ :(CV) ttlgl relqJla fH 'dootrel:(.IAr{) 096I tt O I IZ0Zssrord'qrad Ju }I uB,\\S:(HIdAd)9tS€ IIE).?tg g Id rnanseltnt 'ru',{lln3 lleqsollcoJ:(,{\S$ 8Z6l ,{lnf'sEcn'I S H V'uotplereD :vr.lv{lsnv Nn:lrss^\ 'g dulÎ*ro.trg 1yg eql se q ou J?,Js" ls"oc lso,t-qlnos eql leeu'sll"llsnY ulelseaa:Nolrn{ xrslo .(t96I) ellql(-qlrus 'q '{r?d '82 apg eturdot eliq,r '3uo1ruut g.t ?c 'JLldlllJpeorq sle1ad:JlerlrJ.'^lloo^\,(l3s.I?ds 'relnpue;3'3uo1tuut eJ'PlollJp'JleJllqutl'J34 sledJS (lu;:qPJseql ro) S. I 'Japuels JSEql€ JtEloalJEJqrq,(lJlnulut ^rJA'3uol lulu / ?c loJlped:snocnp"c 'aleirey '3uol '€c .{lesizds urru t cIldIJIs-p"oJq slt"rq :el€u?l (,{lesuepol) flesreds '3uo1 'snolq?18 (erbur ro) ruc ol euec€r aldurrs Iuultural ? ecuecseJogul 'elepundSI 'eôq€ 'etrnc? '3uol o] olr?u€l ,{les.rEds ru1npuel8 e^€tuoc lulu 0Z 9 'crldrga-,Lr.oirruol reeulT 'ellssos se,ttel snorqelS ro e1eue1e1e11e1s,{lesreds 'q8rq 'l"IuuoJed 'lcaJo 'lopuols qloorus slelqcuerg ruc 09-0€ ,{poo^\ .(u3?s '., *"",, oloqd)l ) snd.{loloH t6 u llo) lreqtrC sIIE}uopIJJo?rpuElloH ,,?:*ii?#+ '.{\ouluEz]JnJ 'ls{s loa uuv 'sredlEr\\:(6t81) tltz/zz.sot[ lBN'los llng urnsga g 'ueestou ed{I .. (le8nH)punos s,a8roaC 8uIX,, :uoIlEJgId.{I ":":';:":'.1T":L;;:"#,11^#.,:'^:,',:1i,:ilil%;1 'srodl€iA "le t09: I ts.{S log dod :(a€8I) sI le;enH Id .unoA It r1ut ut ,put,,,o*#!}'} 'ueos lou ed,(I .

'snue8 aql ur suouJes roqlo uo.IJ uoller€des r"elc " pu€ salJads o,,(l esoql uee,$.leq lql esolc € 8ullse83ns ul sJepEJstlc leclSologdroru reqlo uoddns pu" Ir;crluepl trsorul€ eJe sntpruJs g pLre sruollpou'A Jo spJas eql '€rT€Jlsnv ujalsed! ruo{ qloq'serceds o,ldJ 'qclx V 'srlrr?ds !' :ed,t Undershrub; branches erect, smooth, sparsely stellate-sericeous when youlrg, Leaves sessile, slender and semiterete, 7-15 mm long, glabrous or sparsely pilose when young, acute to obtuse, + concave above, rounded below, glandular-punctate. Flowers in conpact terminal heads 1-3 cm diam. Bracts foliar and herbaceous, or scarious, suborbicular and ca. 2 mm long; pedicel 2 4 mm long, stellate-pilose; bracteoles 2, basal, naviculiform, 2 4 mm long, scarious, stellate-pilose; sepals free, imbricate, narrow deltoid to linearlance- olate,.acute, 24 mm long, stellate-pilose; petals broad-elliptic to broad- obovoid, 5 6 5 mm long, thin, glabrous, blue to pink, apex slightly thickened and sometimes cucullate; staminal filaments linear-attenuate, 4-5 mm long; anther suborbiqular, minutely white-apiculate, ca. 1 mm long, pollen pale orange; oiary pyramidal, 1-1 5 mm high with a short, sterile, apiial portion, stellate-pilosealong axial margin otherwise glabrous; style short, giabrous; strgma + truncate.slightly uider than style. Cocci shortly rostrate,in all to ca. 3 mm long. Seedsubreniform, ca. 2 mm long. Flowerheads2-3 cln diam.:sepals linear lanceolate, 3-4 mm lons, snarsellpiloseoorneflrous Flowerheadsl-1.5 cm diam.;sepals narrow-deltoid, ca. 2 mm long,stellate pilose 3lb, subsp,lasiocalyx

31a. subso. nodiflorus E. nodiflorusyar. :ubglahiforu:tDomin, Mem. Soc.Sci. Bohdme2154 (1923). Type nol indicated. Flowers in terminal heads which often become lateral due to the continued growth of the branchlet,2-3 cm diameter. Pedicels(l .5) 24 mm long. Sepals either linear-lanceolate, ca. 4 mm long and sparsely stellate-pilose, or narrow-deltoid, ca. 3 mm long and glabrescent; petals 6-6.5 mm long, blue to pinkl carpelsalmost glabrous. DrsrRrBUrroN: Western Australia, Darling Range-Map 17. WESTERNAusrRALrA: 70 mi N of Perthon Geraldtonroad, A. Ashby459 (AD); Lyndhurst, Wagin,C. A. Gardner6493 (PERTH); Toodyay to Bindoon,C. A. Gardner8707 (PERTH).

31b. subsp. lasiocalyx (Domin) P. G. Wilson, stat. nov. \tar. : E. nodifrorus" lasiocalyxDomln,l.c. lasios woolly]. Typification: Cranbrookto Warrungup,sandy plainsj Mt. Toolbtunup,A. A. Dorrien- Smith." Holotypus:K (not seen). Flowers in compact terminal heads l-l .5 cm diameter. Pedicels very slrort, 1 2 mm long. Sepals narrow-deltoid ca. 2 mm long, stellate-pilose; petals ca. 5 mm long, blue; carpels densely stellate-pilose on adaxial margin. DrsrRrBUTroN:Western Australia: rlear the south coast-MaD 17. WEsrExNAusrRALrA: Ml Toolbrunup.Stirling Range,T. E. H. Apiin 2051(PERTH); West Mt. Barren,C. A. Cardner,Sept. 1939 (PERTH); 12 km N of Albany,R. HilI 1460 (4D); nr. CapeArid, Maxwell1875 (MEL); Manjimup,R. D. Royce2362 (PERTH); Cape Naturaliste,l. C. Wiburd,Sept., 1904 (NSW). E- nodiflorus was descdbed from material collected by Drummond in 1839 probably from near the Swan River. The photograph of the holotype shows the plant to have large flowerheads through which the branches have grown. The specimenslrom the Swan River area which I have seencorrespond to this photograph in habit and have much larger and longer pedicellate flowers than those from further south or south-east; they also have a less pilose calyx and an almost glabrous ovary. Bentham (1863) commented on the variation in flower size but considered the extremes to be connected by intermediates. Domin (1923) aiso noted the variation and described two varieties, the one 58 'esocnrJaa '3uol J€lnpu€I8 ruo t-Z (9.0) olelnqns ,rorr?u saêa'I ,{IJoJ Suruoceguoos pu? u,uoJqsduls el€rperuJelur 'sercueJJncep reynpue13f,11ure3 'q8lq 3ee1 ueer8 qlv'r 3uno,{ ueq,r s}eJrlcunrg ru (€) Z-I qnrqsJapunpoJA '('ISI^J) u s uruorJ '(/ASN "IEI '(IV) 3nv '89TJ ou qco){ : sed,{tu,{S 'uruojJ ssr}\t '.,{o{e'I '.I I t06I 'V/A } t 'q3ot ..'3ur,ro3^\oJ,, €68I pue U poo.A\Icelgraddll ueaa\loq :t06I jl :uortmgrd,(J '(6'8I) ro)ooH '(L06D 'cos '!ler'l'1 uou 36lll ulou 0r..6I Z ros l.r^ .{oX cord snlpautatu! uoua[olrE '(026I) ros uurl 'aroow 'saeU 99I:E loS I S:(Z16I)III:SZ Z res trr^ ros',{oU rord to uEr,\A(]F4rrd) luasapunllDqaqd -taw'(uaasoroqc!) '0rr0sr96 :snd.{rosr(898 cco nLgn'(u ) cv .. ;gSf: S$d8&ilfff rog srsouarssnecrln4 ur ssoD sreqlnos rd arpre8looJrlsrp u|qeH !, :uorle3grd,{J qel892:e6I pe ureJsezu'gdreldug:(806I) res ôo "oru:!lf,[! €II Z '02€:9€I€I:02'z '.rn.o5 :llresopeql Jo : llrosopl(i06t) C-y 6t qel qrqel 1oglozlrra te qarc ur Ieztrrd![ 'zlrJd .z€ .qrasapuoulaFo.rrfl 'ecu?c -gru8rs crleuaAol,{rld eq f"lu ecuBlqueseJ LIarLIt{,(snaploqwoqJ ur punoJ Jo'3'a) 'l}as !r uuoJ ourojlxo oql 1e aDlnaJSo.rSlN uowaJsoLrg01 J"Furs sr lr eJerl 'urq1 :?z?l?qc eql Suuncsqo lou ,{JeAsr aqder eq1 pu€ urn11q oqs f,rea. l"rxep€ 'sr3]c"r"qc " s?q pees eql Jeqlo slr ]soru ur sJeglp 1l uoqcas.qcrq,{\ 'laes lsJolj Jo 'snorqe18 woJJ'oaauoQj urupqaqd Jo lecld,(lar? qarq,r'suerurtrsSuryrerds tsur,\eq ur uolaatsoug Jo suorlces Jeqlo IIE rrro{ s.reJrp uorpes srql 'Erl"Jlsnv u.relsed! ur puno3'serceds auo ÎUO 'elsel Jalno pue aqde:3o Suuaôr urql q8norql elqrsr,rpue ace; Je,\\ol uo €z"l"rlc :Jed?l snouEJqueu urql € ,{[uo ,{q pe;e,roc IErx€p" Jo Jl"g 'pcrldllla pue urql ,{;e,t eqder lacz.; uo ,{}rêcuoc lq8qs go erluec ur '(8uol l?rx"pe 'snou€lquou IxuI €.0 Ec) oqs fra^ ulnlq :qloous ?lsoloralcs "lsel Jolno :tussqe : elrq,tA'urqt drecopue (leluece1d) peqc"llE 'luJoJrual pees 'leu 'p€orq,{reÎr.r" 'snorq"l8 'Surpeerds + csrp :et€lnord? otrq^{,{latnuru srsqlue sueuels :snorqelS 'el"cuqurr sFled :elecugrur {llq8rTs 'eorJ sl€dos 'elrsses ',4rqyxu 'elryndqsxe lecped l,fue1qos sre,tiolg seêel :snoJq?lE lu?ld

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.xtlnols0l .dsqns sntoEflpou .g 'u€?A,\ Jo oouolrncco u,\\ou{ oql Jo quou eft se4rFcol asegl y l?ou tuo{ euroc uees e^Brl I u.roJ snoJqelS aql Jo suorlcolloa luacar ,t1uo egl :q ed Jo q ou ru)t tnoq" luo4 osl€ pu€ Jo^rl{ ue,\\Segl uo ,{?,{pooJ t€au uro4 sJ€o^ 0ZI 'et?trs luac3r ur popelloc uaeq s?q ruroJ ed,{} eqJ perrp eql ur ,{ltrsour '(8uol '?r) {urd er" qcq,r\ sre,{\oge^?q qiog urur € sl€dossnorq"lSgns ro sno.rqe13 'paseq 'g qlg\ (relauerp un Z'ea) sp"erl Je,\\og Jell?us seq se^\ DaDtlD) au?u oql qcrq,{\ uo 'urjoJ reqlo oq1 i(3uo1 ruur y 'ec) spdas osolrd .4lasr?ds 'al€loeJu?l-,rorreu qlr,r ( ur?Ip rur g-Z) spuarl-Jeaa.oge8rey seq ruro; ed,{1 'ue3s 3qI e^sq I suorlcelloc ,{\eJeql IUo{ ,u-ol1?peJtsJelulJo uoll€crpu} ou sl oJoql qclq,r ueo,4{laqsurjoJ o,r1 sercedsqnsed^l erD urqlr,^ papnlcur o^?rl I 'saloadsqnsedfl eq1 ss erues eq1 sr uorldr:c 'sueurceds -sop eqt rrlo{ q.rq,r lnq uorl€trc tl:.og]]t (sruol[uqDlSqns re^) '?oJ? Jo 'JB^) Jarllo eril pu€ oBu?U Sulljrls eq] luo4 IErJetEruuo peseq (xtrlDotsol narrowly canaliculateabove, glabrous. Flower: pedicel 0 5 5 5 mnt long with scveralsmall, basal bracteoles;sepals deltoid-orbicular, ca. I mm long, fleshywith broad scariousmargin, minutely ciliolate, glabrous; petals spreading, ovate, ca. 3 mm long, thin with thjckened midrib, whitc; staminal filaments, narrow-linear becoming terete upwards, ca. 2 5 mm long; anthers sub-orbicular, ca. 0 7 mru long, minutely white-apiculate,pollen pale orange; ovary somewhat embeddedin disc, glabrous,broadly pyramidal, at llrst small but enlarging duling anthesisto 0 5 mm high (above disc); style glabrous, ca. I 3 nln' long, stigma slightly broader than style. Cocci + erect, ca. 2 5 mm long, trurcale arld shortly apiculate. Seedplump, reniform, ca. 2 mm long. DISTRIBUTIoN:Wcstern Austraiia, fiom Kalgoorlie lorth-west to the Yalgoo area-Map 9. WEsrr,RNAusIR^LrA: 30 mi NE of Wubin,J. S.Beard 2587 (KINGS PARK); Dromedarie, w. E. Blackall34,+0 (PERTH); Chooli, C. A. Galdner2078 (PERTH): S of Welbungin, C. A. Gardnef2747 (PERTH): 15 knr E of SouthernCross, R. H. Kuchel2104 (AD); Nun- garin,J. J. Veitch4 (MEL). This spccicsis peculiar in the genusin having slenderglabrous staminal filamentswhich are spreading(not crect) at anthesis. On the basis of these slaminal characters Ervart referred it to the genus Pltabalium. It shows however a slronget resemblanceto the gcnus Eriosteflonl it has free, imbricate, sepals,a broad prominent disc, ard minutely white-apiculateanthcrs. In the flower position, preserccof bracteoles,and in the externalstem morphology it shorvscharactcrs found in Etiosteorcnbut not in the genusPhebalium. lt thus appearsto be a rather anomalousspecies showing no closeaflinity rvitl.t any other section of Eriostcnon or to any sectioi of Phebqliun.

PHEBALIUM Vcn1. Ventenat,Jard- Malm. 2: I02 ( 1805),[fronl phibaleon, a name used by some Greek poets lbr the myrIle(.Mytus can t rrr L.)l; J. E. Smithin Rees,Cycl.27: (1814); Nees et Martius,Nova ActaAcad. Leop.-Carol. ll:l8l (1823);de Candolle,Prodr. l:719 (1824); Jussieu, Mono- graphiedu genrePhebalium, Me|n. Soc. Hist. Nat. Paris 2:125-137, t.l0-12 (1825); Jussieu, M6In.Mus. Hist. Nat. Paris l2:479 (1825); Poiret, Dict. Sc. Nat.39:463-465 (1826); G. Don, cen. Hist.1:791 (1831); Meisner, Pl. Vesc. Gen. 1:14 (1837), op. cit. 2:60 (1837); Endlicher, Gen.Pl. I I55 (1840),p.pte.; Hookef i., Fl. Nov.Zc\. 1 :4,1 (1852); Hooker f., Fl. Tnsm.l:63 (1855);Bentham et Hooker,Gen. Pl. l:292 (1862);Bcnthan, Fl. Austral-1:336 (1863); Engler,Pflanzenfanr, 3,/4:140 (1896);Black, Fl. S.Auslml. 342 (1924)i Engler, op. cit. ed. 2. 19a:259(1931); Ewart. Fl. Vicl.706 (1931); Curtis, Stud. Fl. Tasn.103 (1956); Beadle et al., Handb.Vasc. P. Syd.Dist. 319 (1963). Eriostemottlnon Smith sensu stricto] Poiret, Dict. Sc. Nat. 15:198(1819); F. Muell.p.pte., Pl.Indig. Col. Vict. l:125-132 (iE62). Croweasect. Phebuliurr (Ventenat) Baillon, Dict. Boi. I I :277(18E6). Undershrubs,shrubs, or small trees. Leavcsalternate, exstipulate, simple, terete or - 1lat,{leshy to corjaceoLlsor cha aceous,glandular punctateand often with proninent verrucositics,glabrous or with simplc or stellatehairs or lepidote. Infloresccnceaxillary or telmiral, of umbelsor cymes,or the flowers solitary; llorvcls sessileor pedicellate:bracteoles small, obscure or $anting, persistentor caducous. Flowerspentamerous (to 8-merousin P. nottii). Calyx entire and hemisphelicalor t lobed, or of distinct (not imbricatc) sepals, frcquently merging into thc thickened pedicel apex. Petals free (or rarely united), valvate or imbricate,thin or thick, glabrous or the outsidelepidote, white, yellorv,or various shadesof rcd or green,the tip frequentlythickened and iollcxcd. Stamens10, free, spreading;lllaments subulate or filanentous, glabrons (stellateJepidotetowards base in P. squanawn); anthen versatileor basifixed,cordate, introrse, dehiscing longitudinally, with or without a terminal gland. Intrastan'rinaldisc nirrrorv, co|ltinuous with or distinct from ovary, sometines forming a gynophore. or inconspicuottsor absent. Gynoccium sphericalto pyramidal,apex sterileand solid;carpels (2) 5 (8), free or slightly 6U 19 esocnr!^ ,,,els s '8uol unloplilol 'rB^ I{n}opldal 'd 'sL eôq€ e}eclns,{llelpoul luur-02-0l J€o'I 9 'd '8 qnq (ulu I :hxrler:ilopid:1ira"|slrrpod:rôqeuoar8-llnpPu€e^€tuoc{llq8lls01}eusoêo'r'8llpuourumrp - rq*ôqo 'r"^ urnlopldit 'd ql qalq pue xoûot urur 7 ic x,tler : eloiirdel snourBn:roj{tdeap leclpad:e oqe.{ssol3 JBa'I 8 'Pa^lncal {Fuorls lou J€o-I I

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9. Leaf channelledabove, obloDg to cuneate-oblong,over 8 mm long. 10. Calyx short (to 1.5 mn] high), smooth; leaf narrowly cuneate-oblong,nrargin scarcelyrecuNed 9. P. brachycalyx 10. Calyx large (over 3 mm high), verrucose; leaf broadly cuneate-oblong,ma.gin recurved 6c. P, tuberculosumsubsp. megaphyllum 9. Leaf not channelledabove, oblong or deltoid, to 7 mm long, margin strongly recurved 6. P. luberculosum

South Australian, Eastern Australian, and Tasmanian Species 1. Calyxlobed. 2. Leaf closelyrcvolute and + terete,smooth, calyx with prominent deltoid lobesca. 2 mm long 11. P. lowanense 2. Leaf not closelyrevolute (oa if revolutethen surfaceverucose), 3. Leaf with midrib impressed(oflen deeplysulcate). 4. Leaf linear-clLneateto narrowly oblong-cuneate. 5. Leafca. 5 mm long, linear-cuneate,margin glandularcrenate; calyx almost palelliforn 12b, P, glandulosumsubsp. angustifolium 5. Leaf over l0 mm long, narrowly oblong-cuneate. 6. Upper surfaceof leaf smooth; calyx hemispherical. 7. Leaf flat or somewhatconduplicate, margin bu1late,.", *""*O 1i.l.f:$Jlrr. ?. Leaf flat or slightly convexabove, margin entire and recurved,apex enurginate or shortly bilobed (T.) 14. P. daviesii 6. Upper surface of leaf + glandular, convexj calyx hemispherical1() cupular or turbinate l2a. P. glandulosumsubsp. glandulosum 4. Leaf + elliptic, over 2 cm long- chartaceousor subcoriaceous. 8. Leaf smooth,without prominent glands. 9. Calyx large (5 rnrr dianr. or more), cuplrlar, (5) 6-8-lobed; petals pink to purple (inland plant) 17. P. nottii 9. Calyx 2-3 nm diam., s-lobed; petalswhiie (to pink). 10. Calyx lobesbroad-deltoid, ca. 0 5 nm high 13c. P. squamulosumsubsp. longifolium 10. Cal)\ lobc.nirro$ JelloiJ,c.. l 5 nrnrhigh 14. P, woombye 8. Leaveswith prominent glandsabove; branchesglandular-tuberculate 13g. P. squamulosumsubsp. verrucosum 3. Lea[ with no apparent midrib. 11. Leaf narrowly oblong-cuneate.convex above 12. P. glandulosum 11. Leaf obovateto orbicular to 10lnm long, nargin + recurved;calyx henispherical, lruncate or with shallow deltoid lobes 13k. P. squamulosumsubsp. ozofhamnoides l Calyx truncate or undulate on margrn, 12. Leaf linear, linear-cuneate,narrow-oblong, or closely revolute and + terere. Il. Midribsrrongl) inrpres5ed Jhn\e. 14. Leaf linear, 10-25mm long, entire, smooth above 13f. P, squamulosumsubsp. lineare 14. Leaf narrow-oblongor linear-cuneate,5 l0 mm long, margin glandular undulate. 15. Leaf margin sliglrtly recurved; leal ]inear-cuneate;calyx very shortly hernispherical (almost palellifornr), slighdy lobed 12b. P. glanilulosumvar. angustifolium 15. Leaf margin not recurved; Leafnarrow oblongj calyx shortly hemispherical,truncate l3e. P. squamulosumsubsp. parvifolium 'eaf 13. Midfih 1ot apf\arenrabore; |evnlureand lerere 19. P. st'enophyllum 12. Leaf narrow-to broad-elliptic,obovate, obcordate, or suborbicular. 16. Leaf obovate,obcordate, or suborbicular,+ coriaceous,midrib not imprcssedabove. 17. Leaf 2-3.5 nrm long, apexrounded to truncateor obcotdate; petalsca. 2.5 nm long 18. P, obcordatum 17. Leaf 7-ll mm long,apex rounded; petals ca. zl mm long 13k. P. squamulosumsubsp. ozothamnoides 16. Leaf narow to broad-ellipticor oblong-elliptic- 18. Leaf narrowly oblong-e1lipticor narrow-elliptic,6-20 x I 2 5 mm. 19. Leaf oblong-elliptic,to 12 mm long, apex rounded, nrargin slightly lecurved,thinly lepidote above when young (Vict. Alps) I3i. P. squamulosumsubsp. alpinum 62 E9 'd '8t ,3uol ,olldlllo .IT IlxoJ elniE xede urc ,-S seABo.I '3uol 'snooreueqc 'ftln8ue LLrLU4-9.2 s.Êol :snoJq€I8 ,{l8uorlsslolqcu€rg .0I 'xed€ elrJolsou qlra{ sledJ€a:qteoueq snoJqEIS seêe1 6 tunetusolp'd '62 xed€ olrrets,elorel-qns.iuoJ e qlr^\ sloclreJ:peeqla€dluo3leururel e st sr6.,'{ou: qleeuoq etellets,{lesreo:r ro esolrdsoÊe-I .6 'e]€llets 'esolrd'snorqelS ,{lesreooto JEelJo eprs.topun 9 unleluep 'd '92 snoJqelS,{reô:rapuals seLu.{c '8uo[ 'e1eôqo-,ro!eu .1 dreilr)ie :olelncrtuep urBrDLu Llrr g t oi iuolqo-.uoi:eri se,leal saproe€uqrel'd '82 ' ( ,& S N sll\i onlg) rede eluets Euol € qtr^\ sled.rer.snolqelS .proô ,{l^\or€u lr"Ô .g UnIoJIJII,{qd'd 'tZ ( A puE slr{ ,(^rous)(snorqel; ,{leftr) snolnroqnd-elellotsolrll \ ,F.ueqds ,{rBÔ .8 'leurLLuel-qns '}trBdruocpue '8uol 'Lrr8retrl ,{l}soLu llerlls saurl3 ,{lellrxe :orgue luur lt-8 ;tnioêr eqt oi enp eteror-qnsb1crjilrlle-8uo1qo sa"ea 1 I 'snolnreqnd-elellels .9 .lrq^\ ,tlelno(u se^€olJo oprsrepull 'rollellls lneJrxaloue lou oseq seÊe'I 's '6€ '8uol 'o8rel .E sualqtu€'d lnmrx6ldure aseq rur 0I 9 soêe-I 'slEtod .Auol 'I uEqt loSuol Iltq8rls ro ol lrnbe + suauels :BorpaoJds+ ruru ot slelod 'd S urruoulplr!^ '!,t esoluatrlol-01?ne}s,{lesolc '8uol ,arrdrllo ,ele.lteÎp lrlels :peqolrq xede rur t Z er Suolqo ,{1sio1.rqoJou se.leal .9 'It IlsraqlnrrE d ,8uol -Er .reln8u€ .t osolrdluols :osntqo xode Luc I } + eleirre^rp seêe-I 'Jlejel re

'd 'tz urnrtojngro C[SN) peror\og-(€)Iecuecserogur:peuoDeg,{l8uo4s otur Surppr8 '3uo[ lerrped:tualsrslrd'sl€des'otEô 'Z pue $A\oU,rAoloq lsnf pellosor ruul S.I-l saloelJeJq:snorqElS slBlad .Z 'gE 'ds '02 'eloprdol .Z uneuEnbs mlJBqoqd poro,{ou-g Z eruorserogul sle}od 'rloprdrl FrP\O I 'd '02 unauBnbs polo,rou oJoruJo Z,{llensn oju6asoJouur :lerrpede]eJat+eqlJoJI€qre^\oJur,{lpnsn'snornp"c'alnulrusaloolcercl:snolqelS,{J?Ô.I saploru?lsorrg'zouces unsoFurBnbs'dsqns unsolnuenbs 'd etl .6oleqaloprdel-snou -tSnJJeJol ,&eÎrs'eôq€ papunor Jo leg 'snoeaelr:qr ol snoeJerror-qnsseêe_I 'ZZ unoJBIroJ 'dsqns tunsolntuEnbs'd 'qgl (,{[ 'str,,{ elSunqLunrreld)salers snotrlr poret}ecs 'tegS N 'snoocEuor qlr/ .rolaq otoprdol-.{ra^lrs ro e}errtdnpuoc ,{llq8rJs sa^Be.I ZZ 'etoprdelsnour8nrreJ 01 ,{ro^lrsqBled pue \,{ler 'e}nc€ 'snooceljoc '8uno^ ol popunoJxede 01 snoe3"ueqc ueq,r eôq€ snotqeJ8so Ba.I IZ 'dsqns 'r€t urnalue8Je umsolnuBnbs d ( lA S N IEls€or)eloprdel ,{re^lrs sleled pue x.{lql :popunor xode 'snoojBtrEqr'3Lrno,{ ueqr^ oôqB etoprdal-rlellols soûe.I 'IZ '8uol ruLuS Z qstad'relFws sto,^\oll 0Z 'd 'SI !0tn (raproq I S N O) Sool turu 6 ec slElod'o8rel sre^\olC OZ 'Euol ruru 0/-01 (1,)ildrlle seÊe.I 8T 'dsqns 'pel aIlJrJB unsolnuenbs d ( plsuaano gS pue pueiur)3unof, uaq^\ oôqp snorqtlS'pe^rnrer ur8ieul'oltlcE .^dSN '8uol 'rrtdrlla-,rorrpu tou 01 popunor xode ruru 0Z-9 ro crldtle-Suolqo ,{l^\orleu J€o-I 6I ll. Leavesnato\t-elliptic to spatulate,2'5 5 cm lorg, apex obtusefo ?..lfif.rrrrrn 10. Bmnchlets + terete,leaves less than 3 crn long. 12. I-eavesterete. fleshy: flowe$ on shod pedicelsin upper leaf artt,rll.)|?. _on *unr 12 Leavesneithea terete nor fleshy. 13. Leavescoriaceous, oblong to oblong-elliptic,to l0 cm long, entite; flowers in + compact terminal heads. 14. Flowers solitary within the leaf or bract; pedicelfleshy (T.) 25. p, oldfieldii 14. Flowers mostly in cluste.sof 2-3 on an axilla.y pedunclejpedicel slender (Snowy Mts.) 27. P. phylicifolium 13. Leaveschartaceous or if corjaceousthen not oblong-ellipticor entire 15. Leaves narowly to broadly obovate, flat, chartaceous,l 5-3 cm long, apex obtuseto rounded 30. P. elatius 15. Leaf shapevariable but not as above- 16. Leaves oblong-elliptic, ca. 2 cm long, flat, apex rounded, margin minutely oenulate all round 40. P. nudum 16. kaves smaller or with recurvedmargin or + truncate apex. 17. Branchlets pilosulose in lines betwe€n decurrent ribs, verruculose; leaves elliptic, ca. 5 mm long, apex rounded (Q.) 32. P. gacile 17. Branchletsstellate-hairy, sometimes minutely so, smooth or verucose. 18. Leaf-apex + truncate or bilobed; carpels2-3(4). 19. Leaf entire, slightly scabridulousabove; antherspink (SA.) p. hill€brandii 19. Leaf denticulateor serrate,srnooth above; anthersyellow (V: UJ.lU.Or_

18. Leaf-apexacute to rounded; carpels5. 20. Stem smooth, stellate+ all round or glabrous. 21. Leaves broadly obovate to orbicular, 6-10 mm long, flat, glabrous, + inbricate when dry 37. P, rohmdifolium 21. Leaves ovate to sub-cordate,3-5 mnr 1ong,ffat or convex above, often scabridulous(S.A. and V.) 35. P. brachyphyllum 20. Stem verucose, pubescentin longitudinal lines; leaveselliptic to broadly obovateor sub-orbicular,entire or minutely erose(V. and N.S.W.) 36. P. lamEophyllum

Sectio 1. Phebalium PhebaliumseEtto Etiosternoides Endl., Gen. Pl. 1156(1840), p.pte., not as to lectotype; Endl., Enchir. Bot. 612 (1841). Phebaliumsectio Lepidoltu Reichenbach,Repert. Herb. sive Nomencl. Gen. Pl. 1:197 (1841), p.ple., nomen. Eriostemo secti,oPhebalium (yenter\at) F. Muell., Pl. Indig. Col. Vict. 1:129 (1862),p.p|e. Phebaliumsectlo Euphebalium Bentham, Fl. Austral. 1:337 (1863),p.pte., nom. illeg.: Engler, Pflanzenfam.3 /4.141 (1896),ed. 2. 19a:260(1931). Leaves lepidote (at least on lower surface when young), Inflorescence a terminal umbel (or ralely flowers solitary); bracteoles normally absent or insignificant. Calyx truncate to deeply lobed; petals imbricate, lepidote outside, glabrous within; stamens + equal to or longer than petals, filaments + filiform, glabrous, anther glandular apiculate, yellow, basifixed; disc not apparent; ovary lepidote (rarely glabrous), with or without a small sterile apex; stigma capitate with slightly spreading lobes. Cocci erect. Seed oblong- reniform, usually thicker at the base,2 3 5 mur long, adaxial margin f straight; attached endocarp thin, translucent, caducous; aril linear, short, thin; outer testa very thin and inconspicuous; sclerotestalinely or coarsely longitudinally rugulose; hilum linear, superficial; raphe small, shrunken, covered only by membranous layer; chalaza towards base of adaxial face and visible through raphe. 'fype: P. squamulosumYent, 64 99r20-t|t 5g '(HJUga) '8uI) IZtt ecxou C U eIe'I 'uep,{H reupJeD Jo N [u I :(HlX3d) ttlt earoeo S V 'JolleHJo E ]tu Sg-:(HItl3d) 6€tZ V 3 ieplaot !(Hj;att/9zl ller.elg g lA lt{ ol edoH are"I :vllv{rsnv Nxarsty'A 'SZ dEI eere 'trsa,\\-qtnos'ell€llsnv aJ€JD e{"1-nurlF^\l?O eql uI uJ3lsay'^ :NoIrnsIdlsIO 'a1e8nr.roc,{11eutpn16uo1 'q8rq ,tllr{8rls '8uol turu ?J pees pepunol xedt lutu I f "c lccoJ xad€ S Z 'elnutur al,{ls ot qlpr,t uI €LuBIls:snorqelS elfls :lrluulns 13 pap[noJ lEnba 'elopldel puB sladre: oL11qlt,lr 'Jlcllediec (t-) snoulSnJJ3Jol ,fue^lls lJf,re f '3uo1 'Bc 'l?lnclqJoqns iqBrq urtu cr '1ectr:L1ds-qnsfu"ô :,\\olle'( ruru 1 'uItu I 'er '1e;r1dr;1a sjailtt,c::pi.tno :ioprdel snoutBnr.reJot ^JJ\lrs 8. | \ 9 6 ',\\ollr{ 'qlooLus 'lJrq1'Jr€^le^ tsoutlE JlEdsleled :Jplslno JlopldJl snourbnJlal '1roqs 'siqol prollep,{1pto:q qtr.ll ept^\ utur put q8rq uu 9 9'ec'l?.ueqd 'er 9.'lecl.reqds-qns t -srlueq ,{luoqs ,{:e,l x,41uc:rJ8lq tuul g pnq Jrnlelu :saêel 'slelqcu?lq edeqoj ,{q papuelqns 'eltssas lroqs Surleuluuel ,{re1t1os lera,tas 'snolnplrq€cs 'snorqt18 s.ra,rtojg' aloprdai-elef1a1s:re;rns J3,tlol o1 qloorus 'esnlqo 'ultr SurLuoJJqSuno^ uaq,t\aloprdel ^lesJ?ds ace.;rns roddn 8.0 x 9-t 'alirer pu" atnloâl ,{lesolc ?ururel !e1e1ot1ad,{luoqs sJêe'I elopldel + 'rltoolus snour8nr:eg o1 ,{ra,t1ts slelqcu"lg qBIq tuc 0E ol qnJqslapun

:snd,{lost'0261 ^oN 'lII uostrlApue ueqraH sel Bluo}sol le u!etoo*;,, , tfo"liJ:;1F,ff '(ZZe 'ueqreH,. L,8 1e.r1sny6 .oS Iod I V (I DtuA[lun'te^oloLlDnod aqbur]I'tl 'urnueqraqeql ul reuplBC{q ed,{]sE palcelos uaulreds oql '(HfUAd) 'llel.€I8 :od(lolre'I leuple5 'slle.Iln4ur nplllegelxnl uo^v rlslpuI:/981 l€61 laorllrH .. llBlcelg s.re]tEHrd :/€61lle)aEl8 I y' rulp\\olLrAlo ilrupreD lJOr'LuU slsolnl srsoue:eur 'srqerrE] IeqraH v q D.tuAu re^ DloL[lrnDdtq,boD]n ,, :uone.Uld,{l 'ltnpqnop: snnsrqLusl(zf6l) u8l:/z ierlsnv i Jos 10u t'reupreDV J 'upreg 'J 'I V urnn8tque unllcqaqd

'uorpnporlul 'seeq luecoJJo osl" ^3uoq ^q pelequnJ ueeq aêq ,{Eurqcrq.r sru.re,msptrq{q jo uotlcnpord eq} pa8e:nocue ecusqJn}sIp puel lereua8 t"q1 slsa?Snssrql uouluoc elou qJnu ,{leÎl"leJ eJe,|\spuq^q lueredde3o suorlcalloJ,(Jntuac WoZ,{llBe puE g16l elel eql ut:s3t3adslue.red e^4"lnd eqt s,(e,nlz,{1.Ieeu ele,t "tlEJ:}snVulelse/A uI opeu suolpelloc ,{1J"3 Jo 'uorleulllod-ssolJ eql l"ql sI lulod SurlseJsluluV lcesul Jelep ,{€ru lnoloJ 'unw\)uDuD) 'sesec re.\\ou lcuflslp 2,'3'e 3ut,t1o,tur leqlo ul :sa(ull 8uue,'tou (3urddelre,r.olnq) luareJrp ,{q pe}ueârd ,{11e4rudsr uorl€zrPuq^q (unlo!1{'4 pue wnsofiuaqn '3e) sercedsasaql euros uaa,\\]o8 unll€qjell d Jo I€uelBu'ses"c Jo ^pnts aql uolJ ecEld ue{"} a^cq ol paunss" sr 1r sleqlo uI auros sl lceJ e sr uoD"zrplrq,{q pauluuoa aêq suolls,uosqo plalC eeJB eules aql ul l?ql 'r'f/rJ .rnJJo asaql eJotu ro o,Âlalaq^\ putloJ uelJo el€ stuJoJel"rpeuuelul 'd Jo 'sJrJJds -ttpoJq o! runlDq)qDuD)dr uErlBJlsnvuJJlsaÂ ueês eql uI 'elqrssod se,r tse,r pun lsee 'suollnqulsrP11r1s uae,rleq serceds3o p€ards uaq,\\ elult E l? os plp pu? PelIuIIl alou eâq qcrq^\ 'suorlJes Jeqlo eql ueql lelT.IeesJ3]c€leqc e,\llJullslp sll pe,\loâ 'sel"ls ,{lqrssodlr selels lls ur sa^rl€luaserder8ur.rr,:q uotFes,{luo eq} sl slqlsV urelsBa eqt ur punoJ,{ue ot u€ql sercedsuellEllsnv ulalsoiA euros01 pel"lel 'lu€ld 'uao dlesolc arour aq o1 sreaddv uerle.Itsnv q]nos 2 nq d qAnorlll€ '?rlsrisny lsa,r puE ts€eqloq ur punoJsI uollcessrql ut selJedseql Jo ouoN 'lequrnu ledJ"c pscnper € pu" sre,ôu sp?aq olrssas8urêq ur ,{lledrJuud slaglp qJtq^\ agtr)onlJ t snue8 et{l ol Jo 'reê,\\oq pot€lar flasolc sr tJ snuagerll ur suoqcesraqlo erll Jo {u€ o} S3I}asolc ou qtr,r soroedsJo a8?lquess" snoSolouoq E sr wnlpqaqd uollses erll 's31"ls uJ31S"Eeql pU? ?IuEtus"I '€rl€rlsnv qlnos uro{ uot puB sll?llsnv uJelseld luo{ sutu 'sercadsuaeleulN Phebqlium ambiguum may be distinguished from all other species of Phebalium by its solitary sessileflowers and strongly revolute, sub-tereteleaves. It is peculiaiin having a reduced number of carpels.normally 3 but occasionally 4: in the latter case one is reduced in size. The almost valvate petals appear to be an extreme form of the + imbricate condition found in the sect Phebalium. This species shows only slight morphological variation and no hybrids between it and any other specieshave been observed. From Miooiybe, in which genus it was first placed, it may be distinguished by the solitary flowers, the more prominent calyx, and the lepidote petals.

2. Phebalium claYatum C. A. Gardn. C.A. Gardner.J. Ro\.Soc. w. Au.lral.27:l8l (1942, lcla\alu. - clubbed] 'Hab propeWidgiemooltha, ilor. m Typification: 'in drstr.Coolgardie, in arenosolutosis, Norenrber.Cardner. Holollpus:PLRTH. Bush to l 3 m high. Branchlets terete, glandular verrucose with pale.red slands.closelv silverv lepidote. Leavesclustered on short branchlets;petiole ierete l 5-2 inm lorig.closely silvery lepidote: lamina broadly.peltate. {sub- spherical),ca. l 5 mm diameter, containing a large spherical_gland;.apex aimost truncate,smooth and becomingglabrous: sidessmooth. closelysilvery lepidote, and passirg into the petiole. Flowers solitary,-terminal on short bianchlets, sessile,subtended by two lepidote linear bracteoles ca. I mm long; mature bud snb-spherical, ca. 3 5 mm long, apex truncate; calyx broadly obturbinate2 5-3 mm high includingthe deltoid-acuminateteeth ca. l 5 mm lons. closelv silverv Iepid-ote:petali obovate to broadl) elliptical. 4 r 2 5 5 ix 3'7 mm, white, siiverylepidote outside; stamens slightly exceeding petals, anthers oblong, ca. I mm long, pale yellow; ovary ca. l 3 mm high, densely silverylepidote, carpels slightly spreading with a smallapical boss; style exceeding siamins, glabrous; stigma minute, equrl in width to style.- Cocci erect, 2."5-3 mm high, apex rounded. Seed (immature) slightly longitudinally corrugate. f)IsrRrBUrloN: Western Australia, Coolgardie district Map 25. wEsrERNAusrRALlA: Scahill Sandalwood Reserve nr' Londonerry,J M. Frank, 3.viii.i929 (PERTH)i 15 knr south-eastof Londonderry,nr. Coolgardie,bush to 4 ft., P. Wilson3126 (AD). This speciesis the most distinct ol the Western Australian s-peciesin the section Ph;balium. The morphological uppersurfaceof the leaf is probably representedby its apparent truncate apex, while the solitary gland.occuptes much of the interior. The flower is eculiar in being solitary and sessileand in constantly possessingtwo li,.car bracteoles.

3. Phebalium canaliculatum (F. Muell. et Tate) Wiltis J. H. \villis,Vict. Nat.74:169 (1958). Etiostemoncanaliul(rlrs F. Muell. et Tate.Trans. Rov. Soc. S. Ausirai. l6:337 (1896), in note t\nderE. tuberculosut[canaliculatus : with a longitudinalgroove] TvDificaiion:" This Dtrlicularplant with almostteaete and channelledleaves, was collected du;ingCiles' tbird e\peditiontowards Ularing. Drumnond'sspecimen has some of the leavesfully oneinch long." 1sqls11DU5lWillis. Ioc.cir.t: BetweenVlcloria Spring\ and Ularing.7-9 Ocl l875 Leg. JessYouns(Ciles'4thErpedjtionr, MEL46ll. {AIthislimeCiles\astrarellingbetweenthe Mulgabbie Salt Lake to the south, and Lake Rebeccato the nortb.) Syntypus:Western Aus_ tralia,J. Drummond(MEL 4632). PhebaliurntubercutosrLzr [non (F. Muell.) Benthan sensu]ectotvpical Diels et Pritzel' Bot iahrb.35:l2l (1904). 'ed,{lol3el Jo ecloqc s,,.1p* u otln 1".,d-,rlu | ,stllrlA peut€lel eÊrl I .reuueLusnon8rqlu? u? uI uetJuA\se,tr e13I pu€ Jollan1I ,{q slJI ? egl aculs 'wnwfDlpuoJ jo ed,{1o1ca1eql sB slllllA ,{q uesoqc sE^\ qtlrl,'ll 7 'lou dtrl rro pol3allor ueurcads eql qcl€u seop 1! lng st lt IJP,JuI qclqâ ,sa|9'N -tg .. €9 runrc eq lsnu srql ,, .rellentr'!^q palElouu" uaaq seq (Z€9t l,{) uerurcads puourunrc perequnuun ue l"rll s,\\oqs unllEq.Ieq 3urnoqlej{ aq} 01 eauereJed ur?qlueg,{q pellc oN eql 3q ol paunss",{3ql qcrq,{'runIJBq f9 'co[ 'el?I -lerl lroql ur uerurcedspuoruulnrc " ot leJeJ tIf, Pus JellsnhJ 'pes?q SE,\{ 'dr,! uollolJcsep rulnsolnJtaqnl JO 'J s.uprllueg leql ueutceds lell€l srql uo fluturu sgt\\ 1I x[lDtupDtq qc]Elu ol punoj pue selceds ql!,tr e{"lg lO ,{q pertduroc l"Jolas Jo IBUoI€uI I S '(€cIoA} u33q s€rl 'tuBqluoB,{q pelrc osls'e9'oN uolFalloc puoulturlq oqJ -o1celnsues) urls(In)taqnt d enll Jql IJEJur sr lr:ueqluJ8 ^q IIJJSs",\\ qclq,{ '1\'S -IaN) r!o{ (91I t runrreqreqau-rnoqlet{ eql ur euo eql ,{lqeqord ..'rlsnv'Lu€qluag ,, sr ,{q pelrc Je^tx plereSzllC eq} ulo{ uollcelloc lla,txej,{ erlJ, 'uorlpadxe .salrC uo pe1ce11oc1utld eql ed,{1o1ce1se u3soqJ s?rl put setceds,\\eue eqlJcsep01 pepuelul .{3qt IJP,JuI lBql paLunss"s"q '1!J 'col slTlry'i\'reê,\\oq :(.. se^?elpe[euu"qc pu€ ele]al lsourl" qlr,r tu"Jd -rtqncrgrdsrqJ,,) lu"JS ,(laura:1xese,t 1uuldeql Jo uollduf,saprreql 'eru?u ,reu e Surpr,tord ,{luo era,\\ ,(erll lstl} pounsse ,{eql ecut5 (,,8utrt1n sple,rot uorlrpedxg pJIqJ .selIC,,) ttorlrpJd\1 qutlol ,sJlrO uo pelcellot ueurrcadse puu uollcelloJpuourunrC peleqtunuunuE ale,{\ eleJ pu" lallanh[ Aq 'uteqluag ueas sueurrcsdso,'\l a\ll 'snQry)llouDJ uoLuaisotrySulaq stueu slql Jo wnsop).taqu J peunss? eql roJ au"u ,teu e peptnord eroJelarll e]"J pue re11en6 ureqtueg,{q uaas s1ur1d eq1s? sarceds etuss eql ol pa8uolsq3lrl€u 3lnd o] peqsr^\ ,nou ,(eqt rlclq,\ o] suorlcelloc o,lll 3tl1 lcql petunss" puP'(stlsofi) -)aqnl uoLua$oLq st) satcadseql peqllJsop lslg erJ ueq,t purul ul p"q Jellenl^l 1rq1 tueld eq1 ot ^ldd" lou prp uorlducsaps.ru"qtua8 lerlt pe^resqo(968I) alef pu" reltant{ 't9 'ou puourunJq pu? lellg plere8zltg uor; ueullceds € uo peszq uorldrrcsep " paqsrlqnd tullsoT)raqnJ wn!lDqaq{ eu"u aql lle,rx€t! 'lJ

'(CV) uostrl ureqlnos trII ldes 'uotreÊt 6tst d'ssorJ'serê(I 'erpr€8looJJoA Z9:(ASN) 606I 'ueprEl,\tH [ :(Hldad) Z96l'Y!'l { J Jo ,^dS]tu 0€ s)Jcou qBnol 'SuIIInqEllng :(SNVJ) x 'It^ -g -9 erropr^ ru:(Hldgd) ttJ 'sffiu{t V 'etpre8Joo]JoS li6l / Icorg '6lEAterro3:(t SN) 096I xr'62 I g Jo /d rtu 9I :vl-rvusnv Nlarsgl\t\ '81 d€t{-(8uI.IdS "uotcl^ 'ElF,,JlsnV uaana))orlroogle) Jo tss3 ot sllrH ue8uol ulelse/A :Nolrnsldlslo 'uoes los p33s pu" :ltnrC seqol Supvards ,(l oqs 'lloqs a,lg qtr,n rtu8rls :snoJq€I3 el,{ls :xed" prlos E o1 3,\oq€ pa,\I\oxPu{rcô :,rolle,{ '8uol ulru g.l 'EJ 'Suolqo 'pJ axa sJaq]u?:o^nelu eled sluolllBlu [e^\ 'e^n€ur-al"d :urrllr,& snorqelS 'eprslno eloptdel ,{.re,t1ts o1 rlutd->1:ep l€urur"ls 'eplslno 'uur g x g-y ec'1ecr1drl1o'aletlrqrul{l}q8rls sl?led :ulqll\\ snorqelS Z '3uol 'sc eloprdel ,(Ieso1c rulu 9.0 "c $qol prollep eq] Surpnlcutq8q ruru I '11eurs x,{1e3 lecuerlds pnq :3uol ruur 41 slectpad :lequrn ellsses l"ulutlot E ecuocsarouul eôq3 al?lncrT"u"c '(a,toqe sno;qe13-qns 3ultuooeq ,{lerer) 'esocnJ]e,l runluounpur asoru?nbs,{.IaAJIS esola e qlr,'t\JeAo llB SuIulqs relnputl3 'Lrlru 'e,\oqE 'elEcl?J I x SZ €J psuell"U dFqAITsJo alarel dllq8tls :o lq8terls uuruel lelelorled dl oqs,{ra,r se^se-l alopldel snout8nllel pue,{Je^lIS ,{Iasolc 'sorltsocn.rJe,\ l"lnpu€13 IIErussnorournu qll^\ slelqcu"rg qgrq !u Z ol qsng Phebalium canaliculatum differs from P. fliJblium in having the leaves canaliculateabove, and in the flowershaving pink petalsand spreiding stigma arms. The scalesof the indumentum have a more silvery appearancetha-n is found in P. .filifoliun. . ^ In the areaaround Coolgardieand westto Yellowdine plantsintermediate fn lorm between P. canuliLulatum and P. tuberculosurzoccur. These show all gradationsbetween the two speciesand may be recognisedby the leaf shape (narrow-oblong,flat to recurved,frequently lepidote above), the calyx si2e, and the pale pink flowers. About 70 miles north of Coolsardie at Coniet Vale and 240 miles to the north-west betweenSandstone aud Anketell are found similar plants which appearto show a morphologicalrange between these two specres;however, I haveseen no true P. tuberculosumfrom this area. putative hybrids betweenP. canqliculatum and P. maxwellii yar. obovatum are represenled _by]l{. T, Burbidge 2627 (CANB), 22 mi S of Coolgardie and C. L. Webster (NSW 69577) from Coolgardie. These specimenshave a leaf shape intermediatebetween the two speciesbut with the silveryindumentum of P, canali- culatunt, and, with flower characterc similar to those of p. maxwellii yar. obo- Natum. The most westerly record lor P. cqnaliculatwn ^ " is one made by M. Koch from Cowcouing". His mtterial matchesthat collectedfrom'near Cool- g.ardie- I have not seenany other collections from this area and it is possible that the localitv is incorrecf.

4. Phebalium filifolium Turcz- Turcz.,Bull. Soc Nat. Moscou25 /2:259 (1852) [filunr : thread,folium : leaf; Benthxn, FI. Austral.t :344(1863); Diels et Pritzel,Bot. Jahib. 35 :323 ( 1904i:Eneler. Nar. pilanzenfani. ed.2. l9a:260(| 931) "sob. Typification:Drunlmond n.206". Holotypus:KW (photoseen); iso.; K..Swan Riverto CapeRiche, (5th coUn.) Drummond 206 'i (photoseenl. MEL 410i. EtiostemonfliJbltus ('furcz.) F. Muell.,Fragm. 9:108 (18?5). Rounded shrub to 1.3 m high. Branchlets slender, smooth, closely lepidote.. Leavessubsessile; lamina slender,sub-terete (bluntly triangutar in cross-(ection),ca. l5 r I mm: uppersurface smooth. glossy. glandulur puneLure, glabrou\:lower surllce with prominentmidrib. closely silvery to ferruginous lepidote. Inflorescencea 3-8 floweredterminal umbelj pediceica. 7 mm"long, mature flower-bud sub-spherical,ca. 3 5 mm high. Calyx-ferruginousJepidote short, I 1.5 mi high.including the.drlroid lobes of ca. 0.6 rnml closely outside,glabrous rvithin; prrals broadly elliptical,3.5-5:5 x 2.5--3.5 mni, pale to bflght^yellow(rarely white). ferruginous-lepidoteoutside, glabrous rvithin; stamrnat.fitamentsshorrly exceedingpetals; anthers broad-oblong, ca. 1.5 mm long..yeJlow;_ovarysub-spherical, I mm high, silveryand ferruginouslepidote, carpelsslighrly spreading uith a small apicalbosi: sryle dqual ro'petals, glabrous;sligma capitate. slighrly brolder than sryle. Coccierect,2 2.5 mm hrgh, apex rounded with a small sub-apicalboss. Ripe seednot seen. DISTRTBUTTON:Western Australia, south-west-Map 36. WESTERNAusrRALrA I nr. Lake Lefro), E. Cronin, t 893(MEL); 80 km W of Daniell,p. Wilson p. j2q1{AD); l0 km E ol LakeKins township, Wilson3233 (AD); 25 km NE ot Dumbleyung,-S.pj. P. -wilsol ?399(AD); BooraanSiding, A. ,q.snUy928 (Ab); l(aremb;en,E. ,q.shty, 1930(ADW)i WonganHills, E. H. kin; 104(ADi.

Phebal ium f lifol ium-P. mqxv.eI Ii i WESTERNAusrRALtA: Esperance Bay to FraserRange, Dempster, 1876 (MEL); Coolgardie, C. Walter(NSW); l7 mi S of Norsenan,N. T. Burbidge2698 (BRI). 68 69 'lJ€lq'rl€SnJroJ 'q8tq ,(ll"urpnlr8uol paeg 'e13ue relno uo unlncrd? ue tlll^\ pepunor xede '31"1Id8 rulu '"c 'lcaJe 'eT,{tsu?ql :apeorq ,{ltqEqs su8lls :snoJqel8 € IccoJ 'eloP el,4ts :ssoq E qlr^\ pu? papunor xod? '3urp?ards .{llqElls sledrec ll?rus'q8rq 'ec '€c 'ePlslno -rdo1snour8nrre3 Luur 1 frt,ro :^\olle,{ Suol tutu I sJeqlu? eloprdel snourSn:re3',1r.o11e,4 '3uol turu g. g 'ec 'pctldl11esl?}od : ulqll^\ snoJqBIS 'eprslno 'rlloours 'urut eloprdel snour8nueJ flesolc 9.0 et Jo qleel plollap eqt Surpnlcur q8rq urru 'Bc 'll?urs xfle3 6uo1 ruur 'repuels I 'ellsses 'l?uluJel9-g loclpad :lBcuoqds pnq ornleu :teqrun pale^\ou 9-€ ? oauorselou -ul 'aloprdel ,{lesolJ 'rlloorus ec€JJns]e,,!'\ol :quptui passeJdlu! uE lnoq]Ld 'snorqelS 'qtoorus + papunoJ eJEJJnsladdn :papunoJ xedE : elslnueJJ ,{llq8!ls 'ur!u 'Suolqo 'snoac?uoa pue elnlo^or y t-€ €uruel :el€lotlad ,{llroqs 'ele3ue^rp S.l'aloprdel 'qlootus se^Ee'I ,{lesolo slelqcue.lg qnlqs IIEIUS

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'dn peJeelJ eru sercedsJoqlo qlr,l.r suorl"le.r olqrssod aql pus ap?tu ere suorlcelloo .IequnJ lllun paprJJp oq louuec sIqI 'uorlruSoce: 'selcads 'd luepuedepur e,rrasep,{eur lr Jo reqlous pu" unlpJll! '(HIUEd '0902 'tg8 u€^\teq ep"rSrelur ue luesa:doi ,{?ur eseqJ Ils ecfod 'ZgLZ.leup.reD) I]plrar-l'd sur8reur pe,rrnca: ,{lSuolls qll{\ sa,leel3uo1qo-,troueu 'qloorus Eur^"q 1ue1de punol sr (u^\oJg e1u1 quou) uordueJ l€eN 3o 'alqsue ,4.rs,toq uec dnorA srqt;o ,48o1oqd.lorueqt punoJ ern Sursso.rc-1ceq3o saperS ocurs urln) r"eu pJpalloc uaeq e^"q xIln 'd pue unuolll{'d lle 'sso.rJ 1totq uee^\leq sJ3qlo pu€ u.reqlnos Ja3upur7 urporral l J€eupalsalloc ueoq "êq 'd wry1&ldn8aw rB^ wnsoln)Dqu pue urulolll{ d uee,,!\leqseper8retul 'rurr,ts plrqfq e^rl€lnd " ruo{ ep"ru uorpalloc el8urs e,(1uo ql1.,ir Surleopfylensn sr auo eJeq,\\lunrJ?qreq rql uI alenJBêol llnculp sI sIqI sueur -rcadselerparu.relur JO seuosxalduoc e uroJ 01 sB os sezlpuqxq ,{llp"or lI osaqt Jo ellros rlu,r pae unrlDqald lces Jo srequau u?ll€Jlsnv uJelse^\ roqlo qll^r uorl€rJoss" ur aSuer slr qcnru reô sJncao Lunlplllg 'd lecrqderSoe8 1o 'x,4pc l1eus e Sur,req 'wnsoln)DqnJ 'uorlrpp" 'se^"a[ ur d ruo-rJ ur pu" ur:ojrlg Sur,teq ur sJeglp "lt dnoJg Luryfqdo,Drlu'd eqt sraquletu rerllo uroJC 'slBled ^\oJIa,{eql (t) '€ru8rtrs 'sea.eeJJo 'et"lncreqnl 'qloorus puB lFrus eq1 (g) pue luets 1ou aqg (g) 'seêal eql ace;rns reddn eq] uo oeur?nbs puE " ecuosqe Jo 'd Jo IeuuEqoI"IpeuJ Jo IEJaueAeql (1) lq wn1o1n:>goun uroJJ peqsm8ugs1psr selcads srqJ qlr^\;ur^\ori) ssolJ uroqrnos Joa .lj,rsr :(Hruaa vo(rYll,,tirtJ lTl',Sr'$*(*5$ #i,ii: :((V) 9Z610r^ It'puEIelJ'g f'(sercodsluored qtr^\ 8ur,uol3) urper-raAl :yrrvxrsnv Nrl?rsa^\ utn11,(tltlo8aru te^ unsoln)taqu x wnlp/ll! .J

lueredqrr^\ 'ur^\o.,") iiun^otqunc JoaN .ur 8z :(ov) ,ro, ;!$,oio.Ti.:',jl'#ts"r$trtff Suraor8)ururpso)JoNru{ZI:(CV)/0Zguoslri'd'(sor.edslueredqlr^\3uL\or8)neru€(fJo '(serceds n tul 08 :(HJUad I9 ,{qqsv V luerpd q}}{{ Srrr.ro.i;) urtn: :vnv&snv Nuarsa^\ unsoln)raqnJ x tunyoltgf '4 DrsrRrBUTroN:Western Australia. south-west-Map 2. WESTERNAusrRALrA: Borden to Pingrup,R. Hill 1463(AD); 15 km NE of Kukerin,R H Kuchel2018 (AD); 14km W of Pingrup,P. Wilson 3283 (AD). Pheb.rlium microphyllum resemblesP. .filfoliun in flower characters and in the lack of glandular tubercles on stem and leaves: in leaf shape it comes closest to P,-tuberculosumfrom which it differs rnarkedly in calyx size and in having a smooth stem. It has a restricted distribution and, from the few colleciions seen, appears to intergrade with some other species in the section Phebalium. Furthii collections and field data are required to confirm this. The correct application of the name P- microphyllum has -bee-nconfused over the Dast 100 viars: this was partly due to the fact that the Drummond soecimen cited bv turczaninow did not have a number and it has only recently been possible to verify its identity. A fllrther cause for confu-sion was that Turczaninow cited another Drummond collection (no. 208) ls being possibly a variety ofP. microphyllum; this is however a fruiting specimen(and the type) of P. lepidotum. In'1d62 Mueller published the name Eriostemon tuberculosus' "itsivins -rept.t.nt a short description. Thjs namewas taken up by Bentham who assumed to a new species(although his conceptionof the speciesdiflered it would appear that Mueller intendedit to from thit of Mueller's); however " " be a nomenclaturalsynonym of P. microphyllum(the epithet microphyllum beioe oreoccuoiedin"the gerrusEtiosteionl, for in a later publication ( frans Ror]Soc. S. Austral.l6 (189b)33?), he uith R. Tateuhen discussingE. tuber- calosas wrote:-" This ii the ltue Phebalium microphyllwn of Turczaninow, as is clear from the original description,but the specificnarne given by him wasoreoccuDied in theqinus EriosrimonlortheP.brar'hyplt;//lrriofBentham". aciepted by Diels and Pritzel, Bot. Jahrb. 35: 323 (1904)' This'synonymywas ") placed E. rubuculosusF. Muell. (" tuberculatus in synonymy under who " P. microphltllum,-canaliculatum). but kept Preralium tubetculosum Benth." as a distinct species(F. ln both casesthey applied the names in the sense adopted by Mueller, not in the originat sense(for P. microphyllum) nor of my lectotrpification llor P. tuberculosum). I-lrave seen a photograph of the holotype of P microphyllum, and I have seen probable isotypesin herb. MEL which agree with Turczaninow'sdes- criotion. I have dlso seen material which Mueller determined as E, tubet' culi.riis and I have come to the conclusion that they should be treated as distinct species. In order to avoid creating new names I have acceptedBentham's assumotionthat Mueller intended -E. tuberculosusto be a new species(and there ii nothing in Muetler's original publication to indicateotherwise) and I have chosenas-lectotype a specimenso determinedby Mueller, and which is probably one of the speiimens cited by Bentham under P tu.berculosunt. also " i a. not sure whether E. tuberculosLrsvar. laeris" was intended by Mueller to be taken as a validly publishedvariety; the narne was applied by Mueller to an unnumberedDrunrmond collection ard he has so annotated two herbarium sheets. The specimens are however P. mictophyllum sensu stricto and may be isotypes of ihat speciesas they appear to be identical rvith the holotype at Kiev.

6. Phebalium tuberculosum (F. Muett.) Benth Bentham,Fl. Austral.1:343 (1863), not as to description. Eiostemontuberculosus F- Muell..Pl. Indig. Col. Vict. l:130 (1862)[tuberculLrm : a little tuber.an allusionto the wartystem and leavesl. Tvoificatioir:" Soulh-WesternAustralia again futuishes an alnost conspecilicrcpresentative iri Eriostemontuberculosus . . " No specimenscited. In the MelbourneherbariLrn there aresix sheets which corrld have been seen by Muellerat thetime of its publication,and which have beenso determinedby him;theseare fbur unnun,beredJ- Drunmond"S.W. collections, Drummondno. 97, and a Maxwellspecimel Lectotypus:MEL 5115, Austr." Ieg. G. Maxwell. P. microphyllumlnon Turcz.] Diels et Pritzel, Bot. Jahrb. 35:323(1904). '70 'sep"qs 1'*nr1ugi14)-r"pun selou osle ees) el'IPaur -Jelur\loqs sprJqlq aqI oltqntt tt-uilso7l'raqnt'd ut puemo;1:'4 tqBtlq {11ensn :ilotoj'tnrio ,,, ion,,oit'tit,l ul u! '.rnjro osleo8ueqr e :dnq:3e01PUE \AIEJ lr:qtr8bt "q1 Lu"J1"1ind! ^,roLB.{iql e:oq,tl punoJ 3q '{elu s3rcadso'tt1 erp uje,nr:o'saper8r:lu! pull lnJJo lelad\oq oP spuq,{H sJelc€J"ql a^IpuJlslp .,'"qru'nt"-, (llctr\n o.\ l:qt {q,trsurcldro,{-1q1'sod qctq'lr'rolllEJ srJ$oU lr lnq q},^\uol}etJoss" "l *!1Il;;lJ;,",:s;j iniolii{ a .'J:i,#,,_l'ji;K;d, ru, \eq : x,{1e.rlloqs {la,r e qtt n \lJ^\oU rrolla{ pue s,uolsqlootus Luryl'qdoJJltu.J -srjnou ,"o1:( pus 's\oqP :trrlns {1daaplsEJl lE ro JleJrldnpuoJ{llqSrls 'D./g \lJ\\oU p.Lre'J^oqe Jl€lnJ 1e1 aie:urr5-re;urle seq xAIDJ'(4 J luld -'leriec teet Jlaral JJpuJl\ N

'8uol 'elBounc-Buolqo uql,{qd€8elu'dsqns't9 po^rncor,{llq8lls luur ST / se^€01 urn ,{qddqtsrq 'dsqns 'q9 AuoI LUtuS.Z et'allsses se^eo_I 'eteloqed umsolnrDqq 'dsqns '€9 Suol ulut l-' ,{llloqs so^€e'I '5lnlo,^0.I AIesoJf, so^"a_[ '3uo1 'ee '{.?lq 'sle8nrroc ,tl1uurpn1r8uolfptln8ent tuut 9 7 p5puno:'t:lra IJJoJ sJqolSuJpe:rds paas q8rq uu (.g:er llururnslE - 'q8lq rdec-ctu8rtq : rseq sprB,\\ol:lellels ,(lrsredsJI{ls :rloPtdrl kiru?trt ,jri^i"'r"rtr",tdsr etel 'slelad nj ru:q ,{reio-:no;1a{s.raqtue SurprJJ\J^lltlStls t',iuir'' I imol1e(:1ud 'rt1dt1;o ,uarnrs:tpttino etbprdt;snourBnr.ral ot :ttqn '{lpeorq ':sornllS,n 'peqo1-protlep ';ectraqdsrueq s1elod : :to pidal-snou,dn.,ja1 {1de:p *irn-l auoi du-, 9 1 ;:trfed :laqLunpaLrmog-g ; e :c-uarsarogul oloptdal {15so1r':sornlrrr .ru;npue13(;luluru-ro-td slelqJuErB tlbrl.lul Z ol qnlqS 6b. subsp.brachyphyllum P. G. Wilson, subsp.nov. [brachys : short, phyllon : leal Eriostetuontuberculostr lnon F. Muell. sensustricto] F. Muell. et Tate, Trans, Roy. Soc. S. Austral.16:337 (1896). Folia sessilia,cuneata, revoluta, ca. 2.5 x 1.5 mm, supra glabn et fere levia. Umbellae I-3-florae; pedicelli2-3 mm longi. Calyx ca. 2.5 mm altus,paofunde dentatus, intus stellato- leprdotus, Holotypus: AD96350150,Vjctoria Desert,camp 59, near Victoria Spring (ca. 190km ENE of Kalgoorlie),Western Australia,22.ix.1891, R. Heims.Iso.: MEL4811.NSW 69582. Leaves sessile,cuneate, closely revolute, ca. 2.5 x 1.5 mm, glabrous and almost smooth above, not medially sulcate. Umbel l-3-flowered; pedicel 2-3 mm long. Calyx ca. 2.5 mm high, deeply dentate, ferruginous-lepidote outside, stellate-lepidote within.

DISTRIBUTToN: Western Australia, only known from the type locality NE of Kalgoorlie-Map 19. WESTERNAusrRALrA: Vicinity ol Queen Victoria Spring, A. R. Main,25.viii.1960 (PERTH).

6c. subsp. megaphyllum (Ewart) P. G. Wilson, comb. et stat. nov. Eriostemon.tuberculosusvar._ me-gaphyllus Ewa , Proc. Roy. Soc. Vict. ser. 2. 19:39 (1907) lmegas: lalge,phyllon : leall. " "M. Typification: Cowcowing,1904". Holotypus:MEL 4545p.pte. Koch no. 1330',. Iso.: AD 96350166,NSW 69580.

_. _ Leaves shortly petiolate, oblong-cuneate, 7 15 x 24 mm! apex obcordate, slightly recurved, base cuneatet upper surface slightly roundad, smooth to glandular scabridulous, medially sulcate. glabrous leven when young), margin tecurved and glandular-crenulate, lower surface with a prominent glandular- verrucosemidrib. closely silvery lepidote. Umbel (l) 3 6-flowered: mature bud spherical:ca.4 mm high: pedicel4-6 mm long. Calyr hemispherical, 3_.5-4.5 mm high including the prorninent deltoid lobes 2 2 5 mm long, glandular verrucose, silvery to ferruginous-lepidote outside, silvery stellate- lepidote within; petalswhite, broadly elliptical,5-6 x 3 mm; antheis oblong, l-1 6 mm long.

Chromosome no. n : 32, from material collected belween Mullewa and Geraldton, fide Smith-White (1954) as " P. drummondii ".

D$TRTBUTToN: Western Australia, Cowcowing-Southern Cross area-Map 19. y*Tul}_4j].Tgt11_l<9orda, !. E. Blackall,Aue. 1924(pERTHr; Bultabulting,w. E. Blackall4062(PERTH); Merredin, J. B. Cteland,Jl .viii.I926(AD); yijgarn,nr. Mtl-Moore, H._1,_{ilg & De CourcyLefroy, 1889 (MEl);.sourhern Crdss fd Ctr-ooti,e. Satisbury6j {PERTH).

Max Koch collected this subspeciestwice from Cowcowing in 1904, nos. 1083 and 1330. The separateoriginal (?) labels are attach;d to two speclmensmounted on thesame sheet in rheMelbourne herbarium (MEL 4545) of- which the specimen aumber 1330 corresponds to Ewart's description; the other specimen,-n^o,1083, being a form (meniioned by Ewart) which appears to grade into P. flifolium. I have therefore taken the Melbourne specimen with no. attachedas the holotype. Also in .1330 " " Melbourneis anothei sheet(MEL 4544)\ irh a labej M. Koch 1063 conrainingborh the tvDeDlant and the intermediate form. In Sydney a sheet (NSW 69580) bears nirrnbir 1083 wbich cont_ainsonly the type plant, and similarly in Adelaide (AD 96350166). In Perth herbarium the same number is attached to the intermediate form. It is therefore uncertain to which plants the two numbers originally belonged. (coar Jerel '(DaJeqrunuunss,{{ u"-,ir-d, "d,tl "qr "sn?cag{lued) o,tlll't tlstla'r ) 'ou 'ur?qluag i'oii#s .rarrnraui paluasarde:967 uerunedsaql l€ql Perunssg pu? urroJ3 eq ol prroP]:.t1:]-e,1usq^\ Z98t ul rejlan4 qrofi -"in11tt1dot.t1w^'41o ll ^ outuEZJln I AO Delou All€ulitllo se,n 13g7 puouurnJcl uelulJJds ed^l JqI :-*itio'litrt'a :4 suorlz;ndodqtrn'r '111eco1 ap:r8 1)ue' nsolntiiii 3o 'x^lec -raturor sreaddepue:deqs-;ea1 ul alqetJE^sl ll sleladaltq/ eql pIlE ri"tua",ir;"^tq" ireclnsrigeipaiu 5.,ri q5rq^ i:,r'ea1Suolqoileg eq1 '{q setrads '&^ unrloqattd iiele1:r tuorJ paqsrn8urlstPaq luw wn1op1da1 unrcptdal '(HIXgd) tZ ,{e^^l.org A O 'g 'ueuesroN NurrsilA\ 'sunC uotules : (Hlt{gd) Itl,I uIdY H J Jo E It! 86 : vrlvtrrsnv unsolnJJaqv' d -wntopdal wn!lqqatld

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runlEôqo'lE^ 'qL so^"e'l Buolruriis.z I.I sloJlped:aôqe olEclns lou'luLLl S IxI €3)clql'olEôqo urmoordar'.re,{ 'El auol .iru,'g-1 .rapu!1ss1::rp:d :5ôqE rlelnJlleuetr tulu t r SI-0I ea Buolqo-^lolreu seêe-l 'ue;s lou PJJS snlnJld€ 'q8rq p Lllt,r p:puno.r uul EJ'lJ:le IJJoJ llluuns lP ssoq lterus lluluns le f 'qBtq e qlrr sledrel atoptdal-snout8nrl:1 tuu-t| €J {lE\o:eplslno ortos tteui 'Lu'Lu 'l?uldlllo snoo'in.,rai'a1,q,t x '{lP€orqsl€lad :ulqlltr iibpr.ilr 9:g-g '8uolI t snorqeia 'optilno !1opide1snou€n::eJ uru I-r'0 seqol PIolleP qlltt. peJJ\\oU-9-[P u8ruLuLu c.z -c t Y^le) lectlaqdspnq JlnlEu :lJqun luulujJJl arujcso.,o-uul eloprd;1fuorqis ecelrns:a,to1 :snolqu13':.e\uoJ Jo leu eJqlns :e1e,ro"qo 'snoec?uoc ."4a" '-iu t x g o1 Sucilqo "uIuBI :elelolled z sr-i i 'qloouls ,{l oqs sa^€e'I 'e1oprde1 ,41aso1c slalqcu€lg qnJqs II"tuS .(t€6I) 'relSug 092:e6TZ po lxeJuezoeu

'(M 'tz6l 'Pu€lelJ 'g 'f loJlt!'d rlll^\) urParrel,'{1€ lll^ I! Aqpm (xtlpetqtorq dqlla) uISEpuf,J ru €Stf uosll1A D d ^q ^lll"col aruus aqt'tuo-,1pelc:jlo. stieurieds io srrl:s {q pJlEJlsnllrst \E'Lun!pJ!l{ d qll.,"' (1tu:reddt pin xtli:"titl'rri dr qlr$ srptrirrlui ll rl?Jlns flluautuord atotu ,(1,$ol'l"u lut e,roqe esocnJ.Ielalottr sr 1r sut.roJleqlo tll J,\oqE alelnJllsu€t i.ronDus rlooLu. lsottrltare qcrq* ierea;req:d{r eq1 a8uel lectqdelSoaS s1r lnoqSnbrql uoueuer lect3oloqdroruelqeraptsuoc s.ntoqs ,tarJ?^ sIqI Turczaninow based a new species,Boronia lepidota, on this same collection but did not mention its former inclusion under P. microphyllum. This last paper was published too late for Bentham to include the names in the Rutaceae section of Flora Australiensis and possibly for this reason it has been subsequently ignored. About 30 to 50 km north of Esperance is found a form with broad, recurved leaves, strongly glandular-repand at the margin. This plant may warrant specific or infraspecific recognition and I have seen no specimens to suggest that it intergrades with any other form or species; however as I have seen only two collections of the plant (Wilson 3006, and N. H. Brittan, 11.viii.1951),and as in flower charactersit agreeswith P. lepidotum,I have not given it separate status.

7b, var. obovatum P. G. Wilson, var. nov. [ob - inversed, ovatus : egg-shaped] Foliaobovata, crassa, ca.5 x l5 mm, integra,apice rotundato; pagina superior leviter convexa,levis, nitida, nec sulcata;pagina inferior convexa,costa prominenti. Pedicelli brevi,1 5 2.5 mm longi. Calyxobturbinatus, ca. 2 2.5 mm altus,dentibus deltoideis I mm longisincludentibus. Holotypus:MEL 4801" BetweenIsraelite-Bay and Pt. Culver". G. Maxwell. Leaves narrow-obovate(to obovate or eUiptic),thick, ca.5 x 1.5 mm, entire, apex rounded; upper surface conyex, smooth, glossy (without midrib): lower surfaceconvexwith prominentmidrjb. Pedicetsshort, 1.5 2.5mm long. Calyx obturbinate,ca.2 2.5 mm high includingthe deltoid teeth of ca. 1 mm. D$TRTBUTToN:Western Australia, towards the western extremity of the Great Australian Bight-Map 36. " WESTERNAusrRALrA: Near the Westernextremity of the GteatBighi ", leg.Carey, 1877 (MEL); betweenEsperance Bay and FrasersRange, leg- Dempster, 1876 (?) (MEL). I have included this plant as a variety of P. lepitlotum as it is similar in flower charact€{ and as the two varieties appear to grade into each other, It is also similar to P. drummondii but may be distinguished by the leavesbeing glossy and convex above (not dull and concave), by the shofi, ferruginous- lepidote pedicel, and by the larger calyx. As its status and affinities are uncertain I am considering it at only varietal rank. Further collecting is required in the area where it occurs.

8. Phebalium drurnmondii Benth (1863) Bentham,Fl. "W.Arstral. l:343 lnamedafter the collector, James Drummond]. Typification: Australia,Drummond, n. 13". Holotypus:K, " S\ranR." (photoseen). Eriostemonbenthamii F. Muell.,Fragm. 9:108 (1875). Branchlets smooth, closely lepidote. Leaves shortly petiolate; lamina coriaceous, narrow to broad-elliptic or obovate, 3-5 x l-2 mm, entire, apex obtuse to rounded, base obtuse, upper surface flat or slightly concave, dull green (when dry), smooth, without midrib, glabrous, lower surface flat, smooth, closely silvery lepidote. Inflorescencea terminal 3-6-flowered umbel; pedicels 2-5 mm long; mature bud spherical, ca. 3 mm high. Calyx shortly hemispherical,.ca. I mm high including the shofl deltoid lobes ca. 0.5 mm long, smooth, silvery- to ferruginousJepidote outside; petals elliptical, ca. 4 x 2 5 mm, bright yellow, silvery- to felruginousJepidote outside; ovary ca. I mm high, rufous lepidote, carpels with a sn,all subapical boss; stigma minute, + equal in width to style apex. Fruit arrd ;e(d not seen. 14 u?auol6 otll J?euulo.IJ pelcollor eJe,\\os1: {eql l€ql slqeqold T,"f?di!tT 'ra'i1o!aror" .re.^,ogp-ue jtel ur ueql o:l l"rlluepl ls,ourle,s.lsepeds slql Jo puz 'solou,{lrt?col ul id^r eqtrini,ldoH tnotllr4\:rP'(6119glls 13lN) JurnoqlaW "roqt p-ie '(Eg ou) ru€rllue{trq pJ}rJuJlurJJds puoururrrro aqJ lu"i"ri 'qloollls sr XAIEJeql lsoLule puE llPtus Ala^ 'JPUE 'rlEJtns ,{pa>1:eu,':1err1dnp.uocro :}?U q xllor[tp0tq ul ,{llErprur l!i]'alEJIns :!l puz '"io"nt"i' rlauorls ptiu :e8uo1sr x.{1ccaql epq^\ ^lFrpaurlou 'adEqs u! papunbl'e1n1oier st;te1 aq1^leu€^ le.ll?leql u[ X-AIE-opu? J€e[ e,loqr 'aul€soql eq ilsnbriq6 f.ot wniolncnilitl rE^ulnsor)DqnJ d uro{ srsJIPu aq :xklocrqtotq ;' 1i"aleLutllr'lr,'e{€l8 1 5 :q {q "tey 1e oi ltreqr^punoJ 1o Juo o.rrpdruoJuaj,t Itpt"tl sgtl(t9 pllouulnJC)ulq ^q p)llJ \lJquntt Jql Jo :7 s! pun *rto1ntr,,i1r1 qlr,t sarr:ds srql pasn3uoJur€qluog 1"ql luepr^e lI 'rui1o11nq:4 repuls ,4;e,tosle sr lJ relceruqc u[ slalc?Jvtlc 01 ')e^..runs.oln)taqfilJ€el 'd qclq,tr olur) MnlltqdD8aw Jsel"o1 pue ruols ut'(eftla ,te,u i1 'd u1 utn trqdo'ntw d o] pellll s! pue t.ra1"nroi1i-ie.trog ;1.rOi :^!!::l:t)'tq '4 'ufpeJretr pu" 'ssolJ u]".1lno5 1c a-peutu::q arct\'wn4oly{ l "J q.s,'suolltslloc pue xtli)tqcoiq uea,lrleq elerpeutrJlul srJlJErlqJ 3 ur$o - snoue^ loqlo lp"ri"lloi u-aq e,req selceds e3rql ""ql 1?:^J:9..'^:P?'rbr3lurqll'^d uollsz ururolror Due ultn) uoe,lrliq .ro-lwnsoln).laqni'd pua unloJtl4 d swadde "prr?i'5^lrt".ta6rii' ut ueo stiut xt1btQltotq unlpqa1d 1eq1 1I

-uer rur er:rct, szorltt\ s 'BurslH. '--t1,i.r,.',,rilrl,tosl'IruorJro r . l:t "orJrir5qino5pueurlucxtr:rmlrq:trtv!rsrv\drI\r\1::lll' :::.'g[u-'v,i^rr.[|l,X"Jlt"*;}5li 'ellt .g1 de'\I-eare ssorJ uleqlnos s11r11 ue8uo16 rlsnV ulalsoiA:NollnsIulsIO 'ue;\ 'qBIq '€J ]ou PoJS pJpunor xJdP luLu E :elopldel 'lcara rcJoJ el,{ls uEqt repeo.lq trlecrers-uLublls,'sno-rQc,{-3l{1s 'qat! -nj !:prslno eloprdal snour8nr'ro1o1 {renlts ,i-",a..ia" ru- i f.re,to 'splslllo ')rr,/t\'rlLU z \ b Pf,'l?JllolllJ,nllteolqtiitod :ulqli\\ 'noJqnlS JlnpldJJ ii.Lr, ',iro6.,i''siio1 "iui s s ric seqolptoll3p ;#i;;"J 'lleLus 'l€clleqds:ql 1:,'-p,-"tl"lpnq 'rr8ni ruur i uc 'turollt:lecl IJe^ x,{l€J q8rq ruul S Z ec g17 '.reptrols :lequtn qar:',\1-:H:9-[" atuac oinieur :aioL urLu leclprd 'qtrprutl€u1u]4 -salouureloplclel snournnllel u1 ir"nlts esocnrle'lluel[uord t r1]1'u ',;iorqnra':"'ntint {;tqdrls ,:;;""''.,";;' t;lerpaLutliu:urLuorrl :lerrldrrpLror -':lelrrrtoil-aruilnq pJpunoJ \JdE ,.1"," ,a"ir"t .re..ldn urareur:oleut8leLu: ol "..1 ':leaunc-Suolqo se^"01 '-* i cl oI ,{I,\\ol.l?u€ulru?l :3}1]lo]:}3dAluoqs :qnJqs 'etop,d:i '"4'l,io5n.r.to,rre1npuul8 lleu's snoieurnuqlln^ sl3lt{Juerq

'u^\ortuo{ satrluiz :r 'slllHLr!3uo^A 'u,:Hto"""tg]#.niil:;$""*D' i tt'3lili"""i; ro^erf 'slSuol urlll S 0 er sreProlleP

:,"'31.illjl.,ljl,l.;$T'J';l';:iH'l:1\Y,;ll;?o-r.i '"',XXJ31#'i.:*oj'U::i3,':'X'J:1|iX*"';f1'Jl'o-1e-ul3r-eute. """i,r'r",rrdn\ eur8ca:olalnuo:r-o^re11nq eur8r?tu le^ olepunlor ,lri^Jr 'cteeunr-.,Fuo1{o rutrssrsornlol'llopldal ,.ian',ii, s.l \ sl-01 elsnauinilog llntued ' uoul unsoln))dqt 'etdd (t981)[it: I ler]snv lJ q]uo8lqluaa ( llenw l) J 'Suua 'lloqs :{qreLql lx (1c.r roc x(t?J 'uoslr,1yD '6 '^ou ds d xdluf,'{qr€rq unlltqaqd

'ederls ^lPeu€ix sreup- 1I sulels 1eal uI o,\\l eseql ruo.IJ 'd rlloouLSpu" sec,{p:r qir.t sri,tog rtllurts,e'teq "::l! 11eus .Il:.::!ttldu)!wLunlDqaq4 iue Lunito[ttrf'2 o1 palelar ,{1:solc eq ol sr"edd€ llpaowl'!ruP oruuospue uua^\oc :(Hrusd) oo' '"l.[*oA"l[lf i"]'J;:3s"t';ili:fi]"i$;lli'X '€rlellsnv urolseid :Nolrnsl{Islcl 81 de] I-1se^\-qlnos 10. Phebaliumbultatum BIack Black, Trans. Roy. Soc. S. Austral.40:460.^ra_b.47.(1916) j:311,[-iihi]iJ.h,1ig,ilill,'tih..'dtilil*.,'"1".#*'r;;;:,..;[bullatus : blisrered];Biack, Fl. S tlif;ii,i.!'rrepr.dorus.F.v.M..var. '): \liiq11-rJr,,tgbe;i amiefotius "r]lr-[:.y:,f( ra,i"f;;ba;r;.;;;t,"r. J.Cudse, 3,T!:"";o't::li,[iJ" i"fli."ff ;I,'i'" I",,.;,';i*jliirl,;i: +!.x ue 13j,,,#r;il;i#,'#3,"911,1'1-.i,ll;,,1i"'J,'.,;;l",li:5:r"y"llf:Lr,';imi:i,n glandulosum P. yar.bu)latum (Bla;k) Court Vict. Nat. 73:174|lg57\. p:.,itidr:,r' ffi"':i:?; fititf#".djnit$tu,.lrTatl pl,6i'.y#','t'-J,i!:Hi, M"'"' ni"' it*iioi tr'i w,mmeraanoni ADe65r400r {* Mil f}"Jli}#!.1*l'"1' E. lepidotusInon SprengellTate, Handb. Fl. Exrrarr.S. Austral.24 (1890). ,.;f?i(i!:rin.b." HookerlBeDtham, Ft. Austrat.I :342(1861) pro pte.,as to s. Austraiian g:::ifiit.":r::1'tl:,1ii*iJiil'iii"Ju.*,'fJ,',il'i,f; strarghlor cur\ed. 3,:f",lT_li';ti?ff; narrowlyoblong_obcuneate.. 6_l2 rnmfong, I .l_Z mm wide i!{:ffi n*:n:1:H351f,-ffi',"1U::m-*l:;*-*ru:;,:1}:",f,' Jff il*it:::*li :m#:i*:l: :i$HrTll"'J'"i:4:1,""':;.1',ff IXj*y*Ui iili::i{:]_,f-i":{!it?H:*Utl;1,i[T"t]'t;,{nmt*#*lii ii:# i{sffi,l';";'t ; lll,fill;;:i",,iilll;",*il:,:;,intil,1;:,,,$fi 3J';i&":fu!fiXll"'#*,'il',ff'"'i,"ilifi "ffi' no"o*tisubi-renirorm' i,iE'::f.ni'.1?,"# f ffit ,*d?,ttffllr1t3#lliifd i z s -- ib'l' testan#rv ir,,,*lt?ri?,J",fi,jftf,."f{"'ljff i.#f,i::;l:t,:H,};y}if J,l?iilti districtsrrom Evre Peninsura 3f;:i"Y;:?i'i'#*i.'l*i:; !.**rn east- VrctoRr{:Fllam. W. R. A. Baker.I2.\.t9t2 tM_EL); ?:';:-ilLlsi1ui,i,;i*i;*1,:::,,,i;rl:"1.*j;:fl;tlil;l'jj'i:,i\l'Tilll: SW of LakeHindmarsh f'li,li,lf Lrveristr.TTe {PRr);Kinchina. r w.creen r54 lfii,i i;?iJi}':lilT:'ftJ' 25 i" 1893rADl: ll mi N of Koonibba. l,enrnsura.L). F. slnlon. lg.ix.tq59(A i'|r3v Eyre s:d]il]ii.*'ti:#Ti:i.H "'""flf','3"ff;. #iil:lil il'B:ilf; ii:l,'"",,:, '{'i;;.;ifiltili!#:!il'l*'#ifli!tl'.ir';:lt*uia**,ix,y';i!ii1ff],'"1,:t'i1T,#i:ii:i:il:il,1t*e*rl, 1li:,11t[i"ffi::fr il:ll

g a ronduplicate leaf with a deep' natrow, medial channel, and no recurved ti;;"3j1Xt" mindw]ren p. ""*#,X''j?'.f I describin g se dif o rum. ;;;;{'p:;;;#Z:,# ii.fJilff "TTililiSlllv-in Uffi ;;'*ffi i"ig,:iXl.'!i'*;: :.wn.t{u(l rouolrErllqnd aqr raue lurrrleLu p"''""11 {1uo Ilt!:ll9j^" ^11i:-':i ?v i,,,i;-i qir^sa,raai s,,1i pes'ruoc {c"rs q,,i"Hi]$li,X.fifj!:"T;Jl "," Surls:'tolut spaeseq1 reqteq.{\ ro acnldse1e1 llqs roll€ulllod ra!P:ti- 'srseqlue seqle'lrou }^"-:'1.:]eq-l lell€ (oseq "i1 p1.,o^1y' 3ul^\olloJui,le pue-peuedo su,n.uaial{j' aqr rn-!l.;.t J9 t"l"tdt ;;"il;^;;;'il;i,d;..iHoT:"uuj",';i*i,r3il,'"Hi:{",rri',J'J'":lffi1oYr-,^."ir're"u n*g,rr'"qi qrr.'n)bet'oc :f i4:.sj.,'/-d J-eqlool pe]€ler uortsaSSnsou uJose^?q dnotawnsopttuon's J Jql Jo slequelu. l tllq^l::a.lI qclq^\ ul ipiiepr,raqanoqrtv uorlrr.re,rFcrSoloqdroru *.]^19 ,','6:1cr.,1 ol pelclrlser:y sr sorcaossIqJ "iinriri.v qrnod bue 1o ee'r' ll"uls " '(aV)ZrOt PerrEqS t ) l'l :ix.::#i'iiiJffli,"f;?s"B'i(;"iilH,tf.aqJeJ'."q!IT]JtrTl'il;1gX1"'.WlY";t

,. ^";Sit48l,ti;lH'",f.,r;;lll":ffil#J'*ll1tr,i",lllii'i"';l"ifiT$',!1;$1l1^ .82 d?trÎ -erl€.rlsnv qlnos ulels?e ul ?3J?luoc"fp€ ptlE "llolcr^ uJalse/(\ : NoIrnsIdlsI(I 'ele8n:-toc 'qBIq 'papunor xede flesra'tsutrl'ur ultrl e 9 Z snout8 'tcale rrJoJ "ttlut- i-3tlr. :sprlqro'llr re'no1 elopldel-elt11e1s 'Suol 'srsJqtde Surureue:pue pnq -;.-J ,{i"su"ep Luur6- ic rarlu put Sutrnpôs '{1p1qi 'ti3rq urru- ec ur pelrnceJ pu" poloJ "1nt, t"lt,p'it1 fuailrs llio'i: 1, lecueqds --4"""ii".o :iuo1'r* 'auoibo ''':qtut-::.prstno otoprdel ,^"11Jx q r''z i €J pue{realts :no11at tu r"u S Z \ S nJ'Jlnôqoslsled :Httol Lu ur Z "riou,3n.,t4 'eseq '{re^lls 'prollep'r8.rel srqoL 'urlll!,\\ sno:qv16.;xod-e tP'!^91-l:t,I:l 1" '5ioilr,i.r"rn1!tt '.rlatq 'a1tu'qrn1q" xed"eql 01 rLrrrqi u,r'l"cueqdsi-! {tp,":lq"llf:c- rliriir a'urqcea.rsicioj x,{1ec aqr o:t.Ieplg:dll:,1]l^:orq.pnqernl"ru :i uo1 ru ui' r e,rn,lriF"r q"d, p,"J":$.",H?; '';f q 'e1optde1a1e11a1soJ"Jrns la,'llolr il"ff iH,lJ,i""F"::i""Y,iJ 'asnlqo'urur elnloâ'I -,iqiis iptlaip ro qlo"orusecej;ns iedrln t x zl-t'sut?rtur li'.l,l;[,:Ji.,'Hli,tii:,".u':$i;,;il['*3&]r'1lion1";';',;l:;1,,;".1:o'0" -lf '080€I€96(V : osl :'btf t^l :\nd(loLoH ' uoprt'rjos]o r{udrsallu-f I ,ecuoj qlno5 eqt suole sllrLlfu!s erlLpLuuo tr"..o n'touno, lnoqe _rep:o{"u"ui1e'ainy ,, .",""o. oqr.req..^,ue^\or ro.{runor ue,ror.rA aqrr*il;il,]#r.ri,i:Tili;"il:H Irl

"'";:"-ptl"iii?i uorlnqrrlslpFcrqdusoa' 'arll 3.rar{,ty'!t"'1"!4.-?:: }.?-'"'""t1' 'e8uel u,,1lse8dns'{uii Scuesqeeq} Pu" l"Jlqd"l8 reqlo rllli uollBzrplJq^qJo 1o'satr:ds -oJa ra,toocue:eedd? tuloJrun s1t uerlul]snv ol'uos "1'oi1^sti -ulalsoiAplno'lrslqf o1 ,{1r.r'eyurrsi1r 'er1e.t1sny qlnos ur uorsur}\r^ll:]':^'-:l'.^li:11:e 'pesrec'ireieq 'oql"l!i-\-.1']^1111 ssort" uoll e'ioprn 1r:er9pue ure14 ^eql -icruntur.uoc-ro1uieroJoq €rl€rlsnv qlnos o] pelerleuad J" urroJauo ]:]1"1-i:l] eu?s 3ql ri;;;qipi " "un11,trlaobati'dsqisrunsoln:t'taqnl d s" ,1colsl€lsacu? girLLiro.r'ur8i.,o ue,1er1sr''y,urelrcy'A 1o st uwo11nq,--'4 rro-rr . seped su;Hii:y,H1]i1r$, ^]I:113!14^111.1.1 >1w{t,1 w4 p fi t1d.}3e su ! i:l]:.::i:: l]1ffi11?; -relul Pu" uJloJ JoA\OUpU" JBOJ 'xi1o)irpn'tq 'i ot oste sJolc"i€rlJ 1ei1 ur pue utn !(1'lDiatu dsqns wnsolnctaqnl ;{fffuT','""i',Tly"rt"fn+U'"i:liil,i$l#1",1$1,"t1;{W" nn 8urfdor " sr -aruoslaqlrde leleirt,r pJrisrlqndui ui ". saldliui" :, 19-1o^tl" .,i,r.,,aur?u lducsnuuLu EsE {JElg '{-qPrl!J 3u!llrds-IIJ rallenhl ,,sn11olaiub ''llenlltr 'g'wingppas t3.1fl:^t eql auo ,tq jiri or luas dieq 8ur^eqrI -1ctJ6 Jo 'I'^{tf '{q {l-tpaltc ueurrcadseq1 ililtsio"o si ., re.tn1r1rairg-,, ruii:j pt. 2 of the second edition of his Flo_ra (the pa containing the Rutaceae). ,,neir Tierefore.clthough the descriptionand loiality reference Bordertown,, grven under f. stenophyllumcould equalll apply to p. lotvancnse,there is no evrdenceto suggestthat he had the latter speciesin mind. I have seenno herbarium record for the occunence of p, stenophyllumin South Australia,

12. Phebalium glandulosum Hook. HookerinJ\4itchell, Joum. Exped. Int. Trop.Austral 199^(1848) : with glands]; Benthanl, lelandulosus FL-Ausrral.1:342(j8$); Walpeis,Ann. eot. sy.t. z,)30 lr Ssijfbaitev,eieeniidl FL^l.tlcJtl89qr: fngler- Nal. Pllanzenfim. S 4:t4t (t8q'6).eng.r, op.iir. et. Z.'lSu,:OO Fl.vicr|0{lelt).Blacq.n.S.A {rat.i44(re2;r.biJck.op.cir.ed.2:s00 ll;jl],t""" Typification:collected on l6th June1846, ca..,Sub-tropical ll milessouth of Mt. Owenneai the Mananoa River. Holotypus:T. L. Mitchell331 New Holiincl,.tony'."uin..,,, f (photoseen); iso.: MEL 4751. Etiottemonlepidotus var. glanduloras (Hook.) F. Muell.,Fragm. 9:107 (1g75) comb. illeg. P.:ediflorunF. Muell..lranc. Vicr. In\I. l:10(1855) pro ple. \ensu lecrot\oica. Typification:_"In the Matee" Scrub, at tbe M.urrayn1,er, I_a[i iif6.ii,'i[a iute fyr.eff.,, Lectotypus:MEL 4720 Lake Lalbert and Lai

Calyxdeeply hemisphe.ical, silvery lepjdote, teeth shortly acuminale subsp.gtandutosum l2a. subsp. glandulosum lu.inu narrowly oblong-cuneate -,-, 5 x 1.5 to 30 x 1.5_2.5 mm (to broadly oblong-cuneateand ca. 3 mm wide), apex truncate to retuse or bbcordate, margin undulate, recurved to revoiutei upper surface smootlL or rugulose between prominent hemispherical verrucosiiies, stellate when young, ptine to 78 6L slJol eltu'ls plJe eleq uo lueld tlo_l-" ,09969 ,, Lo6r'662'tzLtop oaiq'rie3" ",'.t:"l"tHi,lr?*:tiillXgplfi* iAsN:snd.{roroA '7 'lEN '{(elelg unsilipuilti ier'unsolnpuolS ,lou : (lt6t) qrog sel€IAS N rluoJ el tzl: I 'rZI 'uoslriN'D 'dsqns 'Aou 1€1s d (^le{"lg) unsolnpuelSo

-rn'r 'orqelsuo:)I a'peo)i "'";!ti\I"'3tl'-t3 ':?13i^t;%Jl$li :oAsN)8t6r'x, zl 'rz dEtN qtnos'^âN : Nolrnflurs(o -"3te 3eq?lJ3)'I-de9 s,ro3-oqulnJrnl^l : sel?rÂ '.. 'set:pdsqnrsrql ,{lq€qold sl JuolsduC '9I : u ott JurosouorqJ 'lln.,, (t961) ellqÂ-qllurs,{q pe;rc uoq5e1loeegr 'aloDttlll'(rorlts'esoJnlle\ tluotls 'q3rq tuul PJ (LuroJlllJledilerrr:Lldsrr-uaq :e1npue1dJpr^\ tuul z I lo'"ti"Pu"lt p:ss:rdtut srf,\\ol-l ouJl iu*-7 s1irrpr6 ,{rr,r x(1er :llEtls '(llq3!ls pue JsoJnrlJ^ '"fi1i'un1a'inj-'n' '"tidn-'p"^rnlal 'rlerunr {rdoap :rrau-pttu 'esnlet \-8 utEretu puj"to'u"n.,lxade'utu l-s 0 s "i"i"i"" ,tr"p.i"ra'"rorn.,,tn slelqru?ra -j;:;!t-;;;,r!"'t -rn1npire1a'{lasreds putr flalnuru ;:ll4iiil *",*û.,;,,.+rs";ls:'$l ,J,'-:"::ii.;,;Y,;!"'i9,1;;i9!''1""'J x'r tsltttqbcvr-1.1#3':'?'.4l{l:i :sndf,loloH ^:N ::qerrexor oqrunrrn!\ 'ltl'j;li";; ;z;t8i ^^sN "o'u"*:'"li'JJll',l;llllfi,hi'{i{i{:jtj",jllUtfiliilfi:}i:UX.ft*lifu,,n*l le oibcun:lecrde'ruttt I E-0x 8 ''elt : snlsn'ue] Lleel .= runrroJ "ro.,'r'u 'uosl^d 'q11 'ôu 'dsqns D d unlloJ!]snBu€ dsqns '3{Jnog 'uo!}els Jo qlnos eql ul lEql 01 l€lulou ne,no31n141B ueaqseLl'selSunqunlJ"iN PunoJ PelJJLloJ 'tcltlutstEq t\rL]:: tunlot'lniA ";r;;-rj'"r,. L,8u',.{ro,s:te:1 ql'^'1ue1d Y.,., ol) aq1,"1q-3i!i si""ilt"' i:qjo " i.q'i-tl-:-,(15 I x lz qrnos ,{\eNureq}nos '! rl-:I,:ualsunqrumr€lN t'ii"ji "ar"i qlr^\ ruroJ puno;sr (srlEi&tltnos ^\3N) esrreLa'iiei" tou op qrrq*.rnrJo selPrpJtll Jqt ul JlqrssoduorlElrLurlrp sl?led rn*'tt'oj "p:ss:rdruro'n1 esaqi Surp€erds -lelul leê,\\oLl :uollruioJer lut'Iln'il '"toqn ro {la>rers ,{1de:p"i1- "qi 11u.-le-lott't"l"lA ,ui';"; i;";r"iri;'tiq 'zrio1rt1 uralsa'tr r".lnjr "i1l ntreirs,iy qlnoi ut pue ll]|"!î:N r]l,it jltu'qrniqo st,xtr1erJtll :J-^rJ,u-plLupesserq u.roqlnosuI rJole slsleo aq1prie '3uo1 suq satradsqnssrr'1l puElstteano ul -;'ii;$pi, q1,'tr1ee1 ,tro.ueu e (lljol uolel{oH :(OV) '(CV' (lnf uotlc S TBuPE llA tRrl 51'o1 uesio-rprv:vtrvtrsnv runos 088IIII^.82 pue 6t8I ôN'"oo"1 i:iiiji1n'u''J6 .(,usN)r06 r,uo, Lv,c ôra rs'erooqu'c "':,-]:::_ :li^!d;d,ffi '' ;:A ,ru;ffi+,i;iii'.,1".,J,13i^?;J&'i'J53,1I,il'ijl?::ii ;,q:,"'Jii'uo\a''-r;e'Hil(i''"r:.'[r;l{"'1'o" 'o iiti*-l$;-,- ls,l*'l';'* i:*:l'i:u;l lrru; 'H-r,eo.,e oqluer '"",'!'o."i".1',gt;:i,il1J'ii;$t,;'#"3'itil.f i';[l'"'$)'tf"$1i3]#8 'tZ 'eIrolcrA ul3ls"e dEn "llI?llsnV qlnos -urltllnos rlsuo;n') urJqln os : Nolrnsrufslcl euerlxe pu€ uialsa^\ : sel"/A qtnos \\aN : Pu 'ssocnlJo^'qleol alerlturncu 'ePI$ ruu qlr,t ro rlelnpun ulSr"ru *t S I I-8 0 -proltop lueurueird '!n.,"u-pt'-'ra'to-z-z^ \g]:lpesserdurr '{1deep 'i;"r'r'*:1oiil.f ao""p or "rnutq'iiqo'"i16 rJfl,:*,.,111,:.y^ rr'errucose, 3" .r]i:_1"iTs; r-;";,,";;:r-ctosetyii"i,1.?,i"li,j'il,',i$;ferruginous_repidote. k:l:::l:::lllt :ll.^l*:::1.^1..:."",.;.; ;; i; ;;d";;;;il';i;; j,i i I i:sijld,lll'.Ln' :l:.^' rbi:lp.!_: impressed:,^d;;!,h* ;;;i;;;';"i",".i',.fl3"ir,. il,';:iff ca.3mmlll'.; Calyrshonty r.0*a Tr l.gnq hemisphericat..ca.o.g ln; ;tg;. lli'mm \ margin,"" uirh.broad. deIoid,,.lobes. rcle, l_:l],,,9i.rg. detroidiou.r. .roi.ij-rir,;.;;;;"i.;;i;i";,i,Closell silrery and ferruginous ilj1l,jriJ;liDrdoLe: broadllelliptical' ca ['Jil;,:t;'rt'T.t: 3'mmt,i"e. srii'v ?"i r.,r,iginori DISTRTBUTTON: nofth-eastNew South Wales_Map 24. *otu., torringron,J. L. Boorman,ocr. 1911(AD);ibid., s"ffoF;ut" R. Thomas,16.x.1932 This subspeciesis characterized I rts smooth branchlets, short leaves {\^ hich ;h ha\e no.nir.impres,.a,oia-.lut,ir pr.rr.J ,niO-iil1. ":l_.have a# c;',il?:li,:f5:f::h :xill ffi"..,: ^";;;i'il;#h;lil,llllfi.i']11"''3J;-1J.1?il,lni,illi,""l::;1,:,',?:o:':),1"^g:';b;twe€n.ti,i,,uusp'."i., Lio'","n"" J,iii-r"uu.o ;"":l_"j^l;:!::!:i!?:,*,,b,p.;;;;;;'#"d;";tiil"lJt"i,ir:;?ijffT*".^y*l_ill.rmediate :"11 srnarro6l,oig_cuneate - -.^'** reaveswlich ;; sLdruErv butli: :1"."i1:i:::1,.:ts.are prominentlyglandular ;;".",ii,iiti;""'jrecurveo ;; flow5n are small and_the calyi rruncate. rt is nâ

13. Phebaliumsquamulosum Venr. ,jVentenat. _r'$'a".i.ut.".il:?,19,'Jard, Malm. 2:tab. 102I(l Ifg;l, tly3I1 qr.2:-164 tt82i): C. Don, [::i:'i,?di' i l8'i r],^'d.'":^1"j, li:i ;r;.\9e._+r60rrt ti:llitieil'""tsil11':;:pflanzenfam. Cin rr8ebr:Baitey. r*,';l;i5"illiil"lir'irf:ilruil$,.}i:;i't'tf lJi"fil; tr r ngter.\ar. 3 4:l4l eueen\td.Fr.t: tel,l8'ee jr i;G. ;;;.:j.-j:op.crr.ed.2. fr?irul t9a:260.ir"',il;g:i;il:il![nj,i'tab. (l9jl).tl'; **n 5 ' qlleinairide ra-N;u\;6"diiiil. "i *o,..un, les,:{oJ:il.ll::'.1.!.,i1y.,-$-.i-b:1"":0.:1, monlagnes." t'torletteCitlei:'"il :fi:i:pi:j!tij'Hlii".,:,t.*j.ni'*'tj;.1;lti,:*u: Type not sien. i'i:r{i,T:#X:!,i.'.';,:E:,il-v,.:li:,Y6i,i;liiEriosremon lepiclorus .rfi,y i;lii,i'i,',l.illt,"'l'.fi; F. Muell.,Fras;. : 9:107 ttgzil,ii;! i,F GGiri* onII r r. gry$intjri'if"yp;{ii**iqis#fi:,:tiriril**#F#l?r.x"": "i?'J*t"*r; trfiiirf:!"'sirll'.^4i;"El':* ";.t.#,.'iilifi $ldii:ir:t E. elaeagnifolius(A. Jussieu)Baillor, Hist.pl. 4:388(1873). Shrub to 3 m hish (to slendertree 7 m high). .Rranchletsterruginous :"f;il:i:'n:ili11,f;'33i".i,i'o:[","i-'91-'ar;rv Jparsetv',"0 'i",t.rv i"o.i- tt; j",.*:*[it $:ff J;il{ I,ig';:ilT,".",1i',1fl1:i.T,:i?!rT.fi,; l5rl:.1.:_b:;:.;!^.';;ii";i;j;':':_.!,,l1lj,J;jl$.,il:i,''ll.T,",t:l ffij,;li ffi:,;',:ffffi, j:, j',I,iil" ji'#ffi :H,ll.jl,*":fl betou'tnnoieice"'i''i'"*i[ *::X*n:rut,it qi:!T,rrt;y;fll:t[Iff,:.,'."'i'iiil';fJ:l;::l'H'#:,''il"t:.'i;i,},'irfr ft$ti,!",::ilii:|,-:ri:; ;T:i't",trt"i.,"S,f.#;.1$i*i[*li"Wl;Si outside,1i![iid,,i:i'?glabrous and yellow ! ;ti to crearninside;;i,;tr;il;i""1,"i:i mm irigh, I8 '02 o€hl -?rJolJrA 'sel"A\ .puBlsueanb uJelsEo qtnos ,raN ulelspo ES :No[nslurst( '(t96I) oltq,\[-rttluIsopg'sel?A\ r{lnos,reN 'qteaqlcelg 'rE8uun) '91 .ou tB petcelloc J€rJaleLuuo{ - u euosouolqJ 'qloorus 'el?Junrl ,(snolnp - + x{;e3 asocn[ar telnpueJt ]ou -uq?cs,{lerBJ)r{lootus eceJJns radddn'asnlet lo JtEcunJl'Jsnlqo ol elnce xede 'snoeceuoo-qns '3rldrlle ro snoac?!€qc'8uol utru 0g-SI o1 tuolqo se,real umsoFrusflbs'dsqns .EgI unsolnurEnbs.dsqns'e€I '(snoeJEuoc-qns ele.unrl x,{l€r'olncEo} pepunorxede ro) sno$eI€q. so^€a-I OI unaJerJoJ'dsqns 'q€I (o8ueualSunqurnllEld) alBlnpun 'ssntqo'Jrldrlle-Suolqo 'snoaJEuor ,(pueurwordx,{lec : e}etrtdnpuocro teu soê{ Ol .eleFpun .Jrtdrllr-peoJq ot elerunlt x,{leJ ol ^\olrEusoÊ:.I 6 urnlloJlSuol'dsqns'q€I 'osnlqo'crldr (puElsueeno)eleluop ^llsauruord x,{lei'8uol urc l-Z e-^roreuse Be.I .6 .pnqur uoêaôqe snorqel8 .snoec€uoJ ol snooJeUeqcse^pe.I ./ runsolnruenbs'dsqns'eet aloprdolsnourBnrroj + ^llBnsn'asnler ro atecunllol esntqo,Suolqo-^\oreq seÊa-I .g umalueBr€'dsqns 'J€[ 'pepunor 'olpôqo e]oprdolXre^lrs sre^rogpue se eel Jo aprsrapun xade qllr' ol artdrllesoêg-I .g '3uno,tuoq^\ lseol l€ eôq€aloprdel-elpllols .snooc€leqc seêo-I .l 'qloorus(x,{pJ,{llensn pue) seÊel puB tuals .9 urnsoJnJrel'dsqns'3€I esoJnrJo^x,{lBc pue .seêal.uots .9 'tllru Z x SI leô dllensn ritdtllo,(lpEorq ro Jrldrtlesaêo.I Z eIIJerB'dsqns 'pgl otErunrt x{lEJ:prssordlur'.ug quprLu.3uno[ '3uo[ + uoq.a^snorqelS e.eJrns reddn rIIw El-Z rr]dtllr-^lolleLl Jo Suotqo-,rbJreuse,teal .9 unuldle'dsqns 't€I atelnpun xxlEa '8uno^ uaq^\ eloprdal Iturql eJeJJnsrrddn 'posserdur (ltqFrls qrrprlu 'pe '3uol 'p;punor rncor,{l}q8ts ruur(Zl) Ol-/ rrde'Jr}drljo-Suolqoro Suolqosa^m-I 't 'apr^\ 'agdlllo-A,\oll€u '€ oJolu ro LuLuZ Jo Ssolqo saêe-I unlloJr^JEd'dsqns'etl !8uo[ 'Suolqo-,{1oJJeu .t esocnrre^,{lluauruoJd rou 9 etr saêo-l 'dsqns 'erlue '3uol .lEeul .t e.IEeurr J€I qloorus ruru 9Z-OI s3êo1 'epl^\ 'Ssolqo-A\ores tlrur I Bc Jo teeurl so^€e.I .t u,or(sl) or ol dn pu. .rrdrla-^\oreuor Buorqoro'Buo' *";t:':: etf!'f"'lj!";;'3";t""j] , '8uol 'orldrlle ,snoocallBqr ruru0, S to Suolqool JEourl ol snoeaerJocsoêe.I I saprourrrEqlozo'dsqns '{tI .^{oleq 'pa^rnool 'oteôqo-pEolq aloprda!-olEllels,{losool ul8reLu'8uoluttu lf ot ol eleôqo-Suolqo.snod)eiror so,{eo-I .l 'suorlelndod erll ernlou eql ol .uorl?apug,{q ol 'J3tc"J"rlc Jo lBurlr tnq anp lou ur alerpaturelurtErl,reuros oq p1no,r s1ue1d3o raqurnu aAJBIe taqt ueeu osle plno,r tr ]nq ,{lrlrq"ue^ ssel pa,roqs uaql $rceds eql ueeur 'laêJ tsrll plnoâ srql sercedsorll 01 sercedsqnsoqt asr"t 01 aq plno,r oJnpacoJd earl"uJell" uy _'se,lrlrlueserdeJsalEls uJelsee eq] Jo lsout 3o uorlerue8leue l€nlueê elqrssodeql qlrl uoncas eql ut serJedstaqlo Jo uoqelaprsuoceql elelrssaJouosl" ,{llEcrSolp1no,r,r dels eql pue ruJoJ esJe,\rp,{leueJlxe Jo slueld urBluoc ueql plno,r lr sB elBudorddE urees saop oJnpacold srql .a!(Euoo,$ '; lou oslu flqrssod pue LunsoqpuDlSwnsolnpuDlS d oprllcur ol s" os ]r e8.re1uee8telue o1 etqrsealolqrseoJ eq .uoneu?^'uoneu?A plno,r 1r pu" seuoJ pel"leJ uo{ peleledespel ,{1djeqs1oulou serceds€ sr lI lecrSoloqdrour elquJeprsuoc s,roqs pelrurlep aJe\I sE unsoptuonbs :4' 'ale8nrloc ,{ll"urpnlr8uol ,{celq ,Auol tutu .?J pees 'xedz 'q8lq ."c ,pere e .elejrdz6 1r pepunor ,{llq8rls ro elecunll rutu g. t rcco3 .pnq ,(eseq Jo elnurur Erubrts :sueu€ls u"rll JelJorls l€q,re(uos ur lqSreJls sprE,,!\ololoprd.l-elallats ,{ler?r ro) snorqelE elfls :eloprdol snojm ol ,4r'a,r1ri QUEENSLAND:Kingaroy, A. W. Long,Mar. 1933(MEL); Mt. Ballow,Moleton District, alt. 3,600ft., shrub2-.! m on top of nrt.,C. T. White lll0l (BRl); Amiens,nr. Stanthorpe,L. Pedley1488a (BRI). NEw SoorEWALES: Torrington, J. L. Boornlan,Oct. 19ll (NSw); ClarenceR., Dr Beckler (MEL 5OlO);Peak Hrll. Poiro Bay, W. F. Blakelyand D. W. C. Shiress,7.viii.1927(NSW); EarringtonTops, L. R. Fraseret a|.,7.i.1934(SYD);National Park, Pt. t{ackingDist., 10 ft. bushy.E. F. Constable,3l.viii.l948(NSW);Ulladulla, W. Baeuerlen,1883 (MEL). " ", VrcroRiA:Howe Range, extreme SE Victoria, a slendertree to 20ft. hjgh J. H. wiliis and N. A. Wakefield,24.x.1948(MEL); Baw Baw Rd., J. Calbraith,20.ix.l958(MEL); wilsons Promontory,leg. Pitcher, Lx.l9l4 (MEL). This subspeciesas hele delimited, includes several different forms which may warrant recognition, but as some of these forms are known only from single collections it appears unwise at the moment to further fragment the species. There are two principal forms which are fairly widespread in New South Wales and are well represented in herbaria. The fir'st, which includes the type of P. squamulosum,has leaves elliptic, acute to entire, chartaceous or sub-coriaceous, and glabrous above. It occurs near the New South Wales coast from Botany Bay north to Patonga. The second form includes the types of P. aureum and P. elaeagnifoliutfl, the leaves are thin, oblong, with the apex bluntly retuse, the margin slightly undulate, and the upper surface often has small scattered glandular tubercles. It is found east of the Great Dividing Range from Jervis Bay north to near Castlereagh; from field notes on some herbarium sheets it would appear that this form is frequently found near streams. In Victoria is found a form sirnilar to the second one and with the leaf slightly lepidote above. Near Stanthorpe, on the Queensland.Neli Sotlth Wales border is found a forn similar to the type form found near Sydney, but with the leaves oblong and obtuse. This plant matches P. whitei in appeararce but is t.tuch smaller in all its parts, and since the two are found in the same general region it seems fikely that P. vhitei evolved from such a form ofP. squamulosum. The style is normally glabrous; however in specimensfrom near Yarra Junction, Victoria, it is stellate-lepidotetowards the base. The calyx, truncate in the typical form, is also not constant in shape: at Bairnsdale on the south-east coast of Victoria, it is broadly deltoid-toothed as in subsp. longiJolium (from which it markedly differs in leaf-shapeand in the stigma); it is otherwise similar to the Victodan forms of the type subspecies. Most herbarium notes on habit refer to this plant as being a shrub. How- " ever in the mountains of south-eastVictoria it developsinto a slender graceful " tree to 20 ft high (fide J. H. Willis in sched.).

l3b. subsp. coriaceum P. G. Wilson, subsp. nov. [coriaceous : leathery, referring to the leaf texture] Folia coriacea,late oblongo-elliptica, ca. 2 5 x 0 5 cm, planavel parumconduplicata (haud recurvata),supra levia et pallido-virida,apice obtuso vel rotundo. Floresnunrerosi; calyx undulatus:corolla flava, extra argenteo et rufo-lepidota. Holotypus:AD 96425099(dupl. ofCBC 005299)between Camp Pincham and Camp Wombe- long,Warrumbungle Mts., New South Wales,28.viii.l96l,leg. M. E. Phillips. Leaves coriaceous, broadly oblong-elliptic, ca. 2'5 x 0 5 cm, flat or tend- ing to conduplicate (not recurved), obtuse to rounded, smooth and drying pale green above, closely silvery lepidote below. Flowers numerous in pedunculate umbels; calyx undulate, corolla silvery and rufous lepidole outside, bright yellow within. DlsrRrBUTroN:New South Wales, Warrumbungle Range-Map 22. NEwsourH wALEs: Belougery Spl't Rock,warrumbungle Range, alt. 20001t.,shrub 120 cm, B. G. Briggs,31.iii.l96l (NSw); GooraweenahCk., WurrambungleMts., 1800ft., basalt, bushyshrub 3 ft., L. A. S. Johnsonand E. Constable,18.iv.1952 (NSW). 82 T

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's3lB/\\qlnos ,{\eN'e8ueu '1ue1d aql ur puno1 sl snoec":ll"tlJ" tlll^\ lnq -rultutlsy .lttLtrqlD J"Jl 'x,{1uc 'eleclldnp Iul1o.rb:nti,11jr 1,1ar'rq 3ql uI put elelnpun eq1 ur -uol ^llr{8rls3ur:q tr ur'os1eput jtal e,1rjo JdDqsJql u unsul,nuonbsdsgns ruo4 sJeJIpll eSuBUolSunqrunJ.lEA\ Jtll ol PalJlJlsJlst sJlJJdsqnsslrll Stem glandular verrucose. Leaves narrow-oblong to sub-terete, 6 x I- 10 x 0.7 mm, margin entire or glandularcrenate, apex rounded,upper surface rounded, glossy, glandular verrucose, deeply sulcate, lower surface smooth and f plane to convex or roundeC. Calyx truncate,0 T x 1 5 mm, closely brown-lepidote. D$rRrBUrloN: Central New South Wales Map 21. NEw SourHWALES: Yalgoglin, " a plant of 2 3 ft., denselyfoliaged to the ground", J. L. Boorman,Sept. 1918 (NSW); Cobar,I. M. Curran10 (MEL): nr. Beckom,ca. 30 mi from WestWyalong, A. Mitchell,Sept. 1959 (NSW). This subspeciesapproaches some forms of P. glandulosuxtin having large glands on the leaves. However in that speciesthe leaves are recurved and the calyx lobed. Compared with the typical subspeciesthis plant would appear to warrant specilic recognition; there is not however any large morphological discon- tinuity between the two subspecies.P. squamulosumsubsp. parvifolium is allied to subsp. Iineare (but occupies a district geographical region and does not inlergrade); this merges into subsp. gratile whrch in its extreme eastern form is similar to subsp. squamulosum.

13f. subsp. lineare P. G. Wilson, subsp. nov. flinearis : linear] Ramulileves. Folia linearia, 10-25 x I mn,,integra, supra levia, profunde sulcata; ttuncata vel obtusa. Floresnumerosi, graciliter pedicellati; calyx brevissin,us,truncatus vel undulatus, rufo-leDidotus. Holotypus:NSW 69862, Scone. N.S.W., 31.viii,1907, R. H. Cambage1671 ; iso.: CANB, SYD. Branchlets smooth. Leaves linear. 10-25 x I mm. entire. smooth and strongly channelledabove, apex truncate to obtuse. Flowers numerous on slender pedicels; calyx very short, truncate or undulate, rufous lepidote. D$TRTBUTToN:eastern New South Wales, from Denman north to Murrurundi- Map 21. NEwSouTH WALES: South of Murrurundi,sandstone conglometate, R. H. Cambage,1.x.1907 (SYD);Shoalhaven R. aboveBadgerys Crossing, W. Forsytheand A. A. Hamilton,Sept. 1899 (NSW).

13g. subsp. yerrucosum P. G, Wilson, subsp. nov. fverrucosus : warty] Ramulivearucosi, folia chartacea,plana vel marginibusleviter recurvatis, oblongo-elliptica, 15-25x.2'5 30 x 6 mm, apicerotundo vel truncatovel leviterretuso, pagina superior ver- ruculosa,in statu juvenile stellato. Calyx profundehemisphericus, manifeste dentatus, vertucosus. Holotypus:NSW 69872,Tia Falls,New SouthWales, Oct. 1900,W. Forsyth. Branchlets glandular-verrucose. Leaves chartaceous, flat or with slightly recurved margins, oblong-elliptic, 15-25 x 2.5 mm to 30 x 6 mm; apex rounded to truncate or slightly retuse; upper surface stellate when young with numerous small glandular verrucosities, lower surface smooth. Calyx deeply hemispherical, prominently and acutely dentate, glandular-verrucose, silvery lepidote; petalssilvery lepidote outside. DNTRTBUTIoN:north-east New South Wales-Map 21. NEw SourHWALES: Walcha, sources of the Namoi R., A. R. Crawford,92(MEL); WoI- lomombiGorge, low shrub,J. B. Williams962 (NSW). 84 s8 tttff 'Er8unqoJ 'uretd 't";f'1$)"ltit6,l 'uosuerllr/Ag I{ :('Ial^t) 6t0€ IInI t I J euldl€ :(tN, s33ug 'U qnJqs'otEJnBltl 'rueleJrlB-l :{ll,\) uolsv ZS6|l!\'lZ'epr,u D B {t Z ',{elle^ 8bl 'I uqo11 tJ Z pue q8rq U T qnrqs'ssIPIdqBrH 8uo8og I)coU rEeu:vrtro.Ltl^ '02 dBlN-(sal€rtA rllnos ,t^eN lsea-rllnos ptr?) €I]olcIA ulalsse :NoIJnslxJsI(I 'eplEno oloprdel snour8n.ueJ 'a1optde1 '.rolle,{ ol e1q,r sl€]od aleluep ,{1o1nutrupu" 3}€lnpun ut8ruut -snour6n.rr:1'lBJueqdsruteq ,{luotls x,{l"J ur_ru 'pale,l^ou-S€} _'3uo1 5 T lectped '(8uno,t ':lrssei s1aq,.un eloptdal snour8nrreJ ot ^ro^lls flasolc eit;jrns Je,llol 'popunor u5(,t elopiael dlurqt) ,{sso13'qloorus aoeJrnsreddn ol esnlqo xede 'sno3c?tjoJ 'ruru g.Z x x 'a.ll1ue'crldJlle-8uo1qo o1 3uo1qo se^?e'l ZI-S.I I 'q8lq 'ec 'alEJlsoJd ,(lluenbe.rJs3qiu?-lq Je,|.\ol Lu I qnJqs lceduoc 11uurg

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'wnsoppuDlS 'dsqts wnsolnpuolS'd 01 slotu puodser.rocsJelc"Jrrqc lcrog eq] :^3olor{dJoru J"el Jo srs?q eql uo unsorywDn "t d uI }I papnlcul e^Eq uox€l € s" ptraprsuoc aq plnorls lJ 1"q1 s1se33nssrql :pepJocal I lruqsrp 'd uaaq s€q Lunso)ntDt dsqns unsolntuon.t rlJlq,{\ uI ?ele aql J"eu luo{ 'wnsolnwonDs SXE1eSeqlJO Jaqlre JO SUOIICoIIOC OU UeeS errerJ I JeAA,\\oH osqns unsorywonbs'J put. MnsopputtlS'J uee.{\:loqal?rpettItalul{ll€c]Solor{dJoru eq ol 'x^l€c sr?edd€ 1I crlsuoFereqt osF sI sel?Js snout8nrrel Jo a3uasqe erlJ paqol 'l?Juaqdsrrueq ,{1daapoql .{q pus se^Bol pu? Iuels eqt uo sarlrsocnlJe,\ -:e1npuelE;o ecuaserdoql {q peqsln8urlsrpeq,4eu satcadsqnsstql P. squamulosum stbsp. alpinum- subsp. ozothamnoicles NEw Sourrr WALES:Three Mile Dam, Snowy Mts., K. M. Cooper, Nov. 1964(NSW); Mt. Coree, P. J. Darbyshire 530 (NE). A.C.T.: Snowy Flats nr. Mt. Gingera, 5,500ft., R. Pullen 2338(CANB). This subspeciesin its typical form is restricted to Victoria; in the Snowy Mts. of New South Wales is found a plant intermediate in character between it and subsp. ozothamnoides,with which it appears to intergrade. I have normally referred the intermediate form to subsp.ozoliannoldes sinceit appears to be geographically isolated from the typical form of sobsp. alpinum. A form of P- squamulosum subsp. graclle found between Dubbo and Narrabri in New South Wales is very similar to subsp.a/pin an. However the leaf in the former has a prominent filiform depression over the midrib in which the epidermal cells are different; it is glabrous above even when young, and the calyx is smaller and almost truocate. The character of the upper leaf surface of subsp. alpinum (non differentia- tion of epidermal cells over midrib and pubescent upper surface of young leaf) is shared also with subsp, ozothamnoidesand subsp. argenteum with which taxa subsp. alpinum is probably most ciosely related. l3k. subsp.ozothamnoides (F. Muell.) P. G. Wjlson, comb. et stat. nov. P. ozoth.lmoides F. Muell.,Trans. Vict. Inst. l:31 (1855),lozothamnus, a genusof Aster- aceae,oides : likel; Bentham,Fl. Austral.I :342(1863); F. Muell.,Key Syst.Vict. Pl. l:141 (1887);Ewart, Fl. Vict.709 (1931); Beadle et al.,Handb. Vasc. Pl. Syd.Dist. 319 (1963). Typification:"On the gravellybanks of the Mitta Mitta and LivingstoneRiver." Lecto- typus:MEL4876 " UpperMitta Mitta,4,000 ft.", F. Mueller;iso.: K (photoseen). E. ozothamnoides(F. Muell.) F. Muell.,Fragm. 1 :103 (1859); F. Muell.,op. cit. 9:107 (1875); Stirling,Proc. Linn. Soc. N.S.Wales 1l:1055 (1887); Moore et Betche,Handb. Fl. N.S.Wales 44(1893); Dixon, Pl. N.S.Wales 49 (1906). Compact shrub 1-2 m high. Leaves coriaceous, oblong-obovate to broadly obovate,cuneate, or suborbicular,1l x 3.5 to 7 x 6 to 9 x 7 mm, entire, apex rounded, margin recurved to revolutet upper surface sparsely stellate to glabrous (always stellate when young), smooth or scabridulous, midrib not impressed; lower surface smooth, stellate. Inflorescence a sessile umbel of 2 5 flowers,pedicel 4-5 mm long. Calyx hemisphericalca. 1 x 2 mm, silvery or silvery and ferruginous lepidote outside, margin truncate to shortly lobed; petals silvery- to ferruginous-lepidote, yellow. Chromosome no. n : 32, from material collected at Mt. York, New South Wales,fide Smith-White(1954).

DISTRIBUTION:north-eastern Victoda, eastern New South Wales-Map 20, NEw SourHWALES: Macdonald R., Ingulba,nr. Walcha,on slate,2+ ft. compactshrub, B. G. Brisgs,23.xii.1953 (NE); HassonsWalls nr. Lithgow,C. Burgess,8.x.1961 (AD, CBG); KiandraDist., E. Betche,Feb. 1897 (NSW). A.C.T.: Top of Mt. Coree,E. D'Amey 243(CANB). VrcroRrA:Cobungra, H. \49lgan,Nov. 1927(MEL); alongmargins of Mitta Mitta and Big Rivers,up to 3,000ft., J. Stirling127 (MEL). This subspecieshas leaves which are stellate above, at least when young, and which have an inconspicuous midrib. These characters it sharesto some extent with the subspeciesalpinum and orgenteum, It shows a considerable degree of variation over its geographical range; in Victoria the calyx is truncate, in the Snowy Mountains it becomes more or less lobed, in the Blue Mountains it is again truncatel hovrcver farther north near Backwater (north-east of Guyra) the calyx is deeply lobed, obturbinate in outline, and approaches the southern form of P. woozrbye (which however has the leaves glabrous above). From near Guyra also comes a specimen of 86 l8 'el,4.ls ()1qlpl^\ ur lenba eu8rls:snorqelB J['{ls:e}nJe 'eetu?nbs e qr'^ 'q8rq urLu I fuPô :'l.rolJel stedreJ ,4re,n1ts3o'3utra,lo5 lrrql z-S 'Jl€q '3uo,*, c'c.z 'SuotqosletlluE : lletlladdn ur etoprdelsnottt8n:la1 le^\ol ',tbit:,4 iruLu I EJ '(lEJrlleuu^sPeuo\) JIP\oqo t,r riop'd:i d:itrs rq8uq 6 3 '(os :ror'iia'ri" '"toixdat iriour8nuej put tue^lls^lesolr'urru s € x s z .r,l sreljd 'r!'i-lc1o 'lcrrrJtldsrLu Jq x,{p3 '3uo1uIIu pnq eJn}eur tsorulz -roi-aturun,r] 9 'iirssrs'lBurturl 'lJ!Lli 'rioLi'. :lrqtuE prrr\\og-(t)t-, '3uoi *- t-z ierrpad 'snoJqcl8'qlooLus u ecu:lsarogul :no1aq eloprdel {.tor1ts{14;o1l aJEJJns 'paptrnol t:lelnrletrel 'sulSlgur reddn 'olearinc'flpeoiq'estq rlde Al'lloJJeu le 'clldllJe-Suolqo Dan.rn:ajleqmrLuos'a.r,1u:'u,r L x 9 01 €ultu"l:olEloll3d '{J oqs iare:1 ':toprdol snotttBrr;01 pue ,fua^lls ,{lesolc slstqcueJq :qnJqS '88C :snd{! '€q.l)q{\\..-.I t0 tUA -oloH (1ôllr{rqffuq sr;$oU 'qnrq\' :C)S uunD N \l!\ luo!}etuld{a .. Zt6llar;q,1 un\olnluDnb\'d ltiitrl os,oi pl'u;enOros {o5 rorJ r'J uttttob[trotq )e^ 'olrt{/A 'f prtuEu] [IlauB^ aql Jo roqln! eql J rJUE 'ôu 'ruou 'uoslri[ 'sI lBls le c d laltqa unll€qaqd

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'sueurceds LunlJ"qr3q pectld illerurou e,leq 1 sercedsqns1"q] qlJ^\ sl tr pu? olut seper8slql punoJ sr tutlLtldlDdsqns 'ttyo' s3rJedsqnslauIJoJ oql ']Ji{ Pu? -Lltqqlozo'dsqt'suee,\\l3q elelpeulelul lu?ld e,e3l€ o{slnJso) 0ri1ul '1nu1d stt11.;o sdlqsuorl"leJeql ^Juelc 'saPnuLu'qlozo 'osql].s ()1 pep$u sr "er? uI Surlcelloo JatIlJ\11 p$e wn;ol lEql 'J\oqE 'sa^"el -nLuonbsdsqns rlloq qI,\\ setltug,es1sa33rts ]Eql alEIlJlsALrsl"ds 'rlqElsuoJ 'd a1n1o,rer'3ub1qo-,t6:jeu qli,t (wOt n VZ J A) wnsofuDnbs DISTRIBUTIoN:south-east Queensland, border mountains-Map 23, QUEENSLAND:Wyberba,3 fr., F. D. Hocking,196f (BRI); Lyra, Stanthorpe District, W. R. Petrie,June 1921 (BRI); Wallangara border mounrcins. c. Wcrd,4.rx.t96?i(AD). This specieshas the largest (single) flowers of any Phebalium. It resembles P. squamulosum,and a form of the latter found at Stanthorpe has a similar but smallerle_af, thus.suggesting a possibleconnection between the two species. .. The petals vary in shape according to their position in the bud. Thiy are elliptical when on the outside and rre then entirely lepidote; if overlappedat b^olhmargins they are obovatewirh lhe marginsforrning glabrous uings: ifoverlapped on one side only they are asymmetrical, only one margin extendiirg as a glaDrousw1ng.

16. Phebalium woombye (Bailey) Domin Domin,Feddes Repert. 12:133 (1913). rourn3:28r (r 'x'18e8) Inamed arte' the tvpe il:{:,yt|fi{tf:{g:,3*iL']i3l!i!!l1" frltc Typification:Woombye, N. CoastRailwaj. W. French.' lsotypus:K (photoseen). "A..woombyc \ar. interh.LliaBailel. Queensld. Agric. Journ. l6:189 tl905j. l)pi6carion: MounlCoolam. W. Fa$celr-Slor). lsotlpus:K (phoro.een). ,. A. woo.mb!e-\ar.parrrifo-lia_Bailey, 1or. cit. Typification: Mount Coolam,W. Fawcett- Story."[: Mt. Coo]uml Isotypus:K (photoseen). . .Shrub to 2m .high; stem stellate-lepidote. Leaves shortly petiolate; lamioa chartaceous(rarely coriaceous). oblong-elliptic to broad-ellipti;,l5 x 2_ 60 x 9-15 x Il mm. entire.apex obluse [o rounded.base narrow cuneate [o rounded. up_persurface fat (or rarely convex and then with recurved margins), smooth,. glabrous, _canaliculate,lower surface smooth with prominent midrib, silvery lepidote. Inflorescenceof terminal 4-lO-flowered-umbels: oediceli 1-1S -+ long; mature bud shortly.cylindrical to obovoid, ca. 5 mm long. Calyx obturbinate, ca. 3 mm long, including the narrow-deltoid teeth of ci. l-5-mm-, silvery to silvery and ferruginous lepidote on both sides; petals obovatd, to 6 x 2 5 mm, white (to pink in inland form.1,ferruginous lepidote outside: stamensequal.to petals, inclined to one side of flouer.anthers oblong. 1.2 2 mm long, yellow; ovary sub-s_pherical,ca. 2 mm high linctudingindumdntum), styleglabrous or very sparselylepidote at base;stigma peltate, ca. twice as wide as -style. Cocci slightly spreading, ca. 4 mm high, apex rounded. Seed ca. 2.2 mm long, finely longitudinally corrugate, black. DrsrRrBUTroN:near the south-eastcoast of Queensland-Map 23. QU€ENSLAND:N^oosa H,eads, sandj soil, in wallumscrub, _bushy shrub 3 ft. highand 2 ft. wjde, HelenI. Aston-98_(BRI, MEL): Pr. Cartwrishr, Jean Calbraiah, t4.viii.t962itranl);Cooiuni B€ach,E. N. McKie and R. S. McNab.20.vii.1935 (BRI); FiaserIs., on sananitini. sii, q. E. !.pp-q 11!g (PB]); BdbieIs., shrubro 6 ft., teavesaromatic (rither like crnnamon), C. E. Hubbard2577 (BRT)

P- woombye, approachlng P. nottii QUEENSLAND:Mt. Wheeler,Rockhampton, Thozer, Nov. 1869(MEL); Biggenden,fls. pink, C.T. White7332 (BRI); Glenbarvia Tiaro, petals whire, S. F. KajewskiI52lii{EL); c;a;;h; BackCreek, Canungra, flowers at firstwhite turning pink, C. T.'White7799 (BRI'; Mile;, R;: pink, D. M. Gordon(BRI). P. woombye shows considerable morphological yariation over its geographical range. In the typical form the leaves are smooth and glabious above, and the petals are white, This form is found at or near the coast of south-east Queensland from Bustard Head, Middle Island, in the north to Morton Island in the south. Somewhatinland, from Rockhampton in the north to Biggenden,Gundiah, Glenbar and Canungra in the south,ls found a form in which the upper surface of the leaves is wrinkled lwhen drv) and stellate (when young), the lower surface is stellate rather than lepidote, and the petals EE 68 .r r,r.. .sa'ue,ê^rEH rlrH read :('rah{ '"t) ;[?tt\:3"LiSY;'J%'9rs;Jffi:P]] r,j sudrr"nal: qSiq te ruotspues q8nor uI punoJ,, lslnquellldoJ leddll : s!.rvlA l-Ilnos .^âN '(INg) 'I ''U ]suo^3 s pu" 99tz uosoqol /A ! € ol qnlqs .,'a8plr ,{uol\ pue (puesuo \u!\o(l EIurqnPBJo \\ lltl tl :18\VJ} ttts r8prqlng L \ -.'U S 9l qnrq.ir:do,, (o.re8ur)lu rleûapl:rA\SN){5llEB l,\ l effuEUuorlrPrdYl:o\vls\llrO '€Z d"r\-sel"y'i\ qlnos ,^âN lsea-qlJotl puP puElsueeno ulolsea : Nollnslursl(l '1re1q':teBn:lo: '8uol 'x,(l€J ,{llEurpnlr8uol ruur ot speas pe?l"lue eq.}u1 pasolcue pue xode 1u ''q8r( 'er t 'tcera 'suelu"ls 'pnq p5puiior uru S lrroJ 11eusttu8tls ol Pnbe + ur'tqBrerls :sLrorqtlSlsoLulE Jo ,to1eq eloprdal-a1e11a1se1dls :al?lledJeJ-8 I 'l?Jrleqds 'iropideqsnour8nr.roj 'qdrq uur fuEô :^\olla,{'8uol turu g Z-Z Z 9.1 'l,e.\\oU 'Suolqo 'lasxe ,{ytq8l1sro papnlcur slerllue epls euo spl",\\ol peurlcul '(,t"lc Jo 'qt-ni surruuts:urqll^\ sLtoJqeLB uo ldacxe)3plslno eloprdsl snoulSnl.I3J i1i:3e 'uttu 'iicrirnLt ot elnre qlr\\ Jttq^\ ol 8uIp"J) e,\n?tudeeP o1 lutd tr-g 'qt8uel x plad o1 I ,{EIr Suol.lo ue q11,{elelncrn8un ro alslnleds 71-g 7l 1 'g-gt/ '(secu,ecseJcxe ol Jre-"noqo-r.cir:eu1oi.re sJ€led:1tn{ ut ?ut8rulua elopldal r,DlA\sJLullJrLtos) rplslllo pu" eplsul elopldefel"llals snouISnJJ3Jol ,{lo^lls '(riru go rlla:l reln8ut-'trt,{1'nor:-eu,o1 prollaP_ 8 I eql Sulpnlcul)tutu 9 x 8 6-7ieirlndnc 'pnq 'q8tq '(uuoJ o1 E xi xf1e3 aqt Sutdola.tuax,{pc aq1 tulu g "c urerllnos ur xedt pauelltg lo pepunol qll./t\ ploô ()1) pnq arnleu 'allsses'lEultulol Iecuerlds Lurug-7 sloctp:d :slaqun pele,,'\ou-g-l ecssf,selouul :8uol 'qlooulsJo 'luauruord J^JJu-prLu'Jtoprdel-elEllols ,{re,l1ts ,{1eso1c oJ€JJnsre^\ol :sno.rqe13'pJIlJuueqJ,{1,No:teu'rllooutsec€JJns leddn :elB3unco1 asnlqoeseq 'pan.rn::-r 'tuul 'p:punor xade {l}q8r;slo l"U'slrlua 6l0S o1€l x SEol t x 0Z 'ir1dt11eo1 cr1

'uoq"lruniep cglcadse4ullultr.rE^\ ol rzaddelou seop'paseqeJe,\\ sallsll?^ o^\1 'serJJds Jql lrlJeJEqJ qJrql\ uo slql ut puno3 edeqs uI uoll€ll?^ eqJ '3 Jeal 'xIler go durqol pue :deqs eLuSIlseql ur e (rluou eql ol leqgnJ rlcnu puno; sr (rrriml wntpltSuol dsqnstunsolnaonbs d ol ,{lIuUlEauos aq o1 srueddu oslu lllq1 'Eur8rlsJlElLJd eq] uI puc x,{lBJeql Lunlueunpul ptulalul eq} 'dJo ul alqeJJlfou'!!ttou'd ol fpuapr,reere aIqwoolll se11lugelsesolc eql Jo 'l(St8t) r0t :6 tu8erCl 1lttoLt d ol ,, snpPldal aoLua$or.q,, ulo{ qc€olddt uE pa,\\oqs ll lsql polou orl,r Jallan/{,{q uees sB,tl 3ôqe peuorluaul llollcolloc uoldueq{cod 3qJ 'lueld st8unuEJ eql 01 srelceJ"qc lellluls pu€ x,{I"c psqol E s"rl (sueqdal5 ld) uox4 stql J?e[ 'dsgrrs lo proJJJ lsoLuur5qpou eql : unauaStD unsopwDnbs 4 o1 qceo.rdde ue i1se33nssrql utqlr\\ rloPrdJl,{losr€ds fluo st xflur eql eJeqesn?caq 8ul1sa -Jelur osle ere (pJocar ,{1r1eco1 3q1)"l8unu"J tuo{ sselulteds eql lsouureqlnos '!!|ou 'd 'sur3-rrura1nlo,ro: ol peNnJoJ qllt\ Suolqo-,totJBL|e'r Jo tuloJ sal?iA qlnos.{\oN eql ol Jsllturs seÊel q]I.\\ lnq selJeds oû eql uee,t]eq lelce.Isqc Je,\\oUu! al€rp3urrolurru.roJ " punoJ sI (pustsuaano ls€e-qlnos) salII l J"eN 'soraeds o,\\t aql uaa,rleq au11codo1e 3o JuEuruor^lJalsBe un sr urJoJslq] leql elqlssodsl ll puEl/lloa d qll'\\ palelcoss? sJelc?rBqcoJ? sJe^\oglutd pue 3ealjo aJEJJns-rrdd n JlEIlJlS { uld 01 ellq,{\ 3.I? P. nottii ts characterizedby its deeplycupulate calyx, with up to 8 lobes, acconpanied by an equivalentnumber of petals,stamens and carpels. The petalsare narrow, sometimesclawed, and are pink to mauve in colour. In the presenceof internal indumentum to the calyx, and in the petal colour,it slrowsamnities with P. v,oomh1e,rvhich species diliers most obviously in having a smaller and turbinate calyx. Another featlrrenot apparent on herbarium rnaterial,but noted by two collectors,is that the stamensbend to one side, a ciraracteralso sharedwith at least some forms of P. woontbye. P. r.)ttil inhabits moLlntainouscountry and appearsto have a rather disjunct distribution. Over its geographicalrange it showsconsiderable morphologicalvariation both in leafandflower form, with generallyspeaking an increase in flower sizefrom soutlr to north. With this goesa changein petalshape from narrow-obovateto unguiculate,and a changein bud shape from ellipsoidal with flattened apex in tlte south to sphericalin the north. The number of floral parts varies between different florvers on the same plant and it is difficult to make accuratecoLlnts on herbarium specimens.However in the south( near Peak Hill) the llowers appearto be predominantly5- or 6-nerous while in the north (Expedition Range)tl.rey are normally 7- or 8-merous. The typical form of P. nollil is tbund in mountain ranges100-200 mi from tlle sea,$'hereas P. ttoontbyeis coastal. llt the interveningcountty in southern Queenslandare found plants intermediatebetween the two species;these may representremnants of a cline which is at presentknown fiom onlv a few collections, c.B. fron Gundiah, Biggenden ind Mites (see rlso notes under p- Woombye). The type localityof P. aol/ll wasgiven by Mueller.asbcing in tl.reNewcastle Range; the typc specimenbears a collectorslabel with the infornration" Dev- ", ranges Flinders & Burdekin possibly indicating the ranges between the upper reachesof thesetwo rivers.

lB. Phebaliumobcordatum A. Cunr. ex Benth.

A. Cunn.ex Benth.,Fl. Austral.1t342 (1863) {ob .- inversely,cordatus : heart-shapedli-' Ewartet Tovey, Proc. Roy. Soc. Vict. ser.2.32:202 (1920);Ewart, Fl. Vict.709(1911). 'SW Typilication: d StCeorge's Range, A. Cunningham."Holotypus: K " A. Cunninghanr No..164,June 1817." (pholos-een).,1!o.: M'EL 47'10. The "Sr. Ceorge,sRange,'iJpro- bablyone of the rangesnorth of Grifiith,New South Wales. Etit\rcno notto ii ts.Muell., Fragm.9:108 (1875); Moore et Betche,Handb. Fl N.S.Wales 448(1891):Di\on.Pl.N.S.Wales49(1S06),basedonP.obcorclarr{mlnonE.obcordatusHook.

Undershrub 0 7-l .3 m high. Branchletssmooth (or slightly glandular verrucose),closely ferruginous lepidote. Leaves shortly pe[iolaue;ramrna broadly obovateto obcordate,2x 1.5 3 5 x 2.5 mm. apexrounded ro rruncare and retuseor obcordatc, base cuneate,upper surfaceeitl.rer flat and slightly sulcate_inmiddle (no definitemidrib) with 2-8 henisphericalglandular pustules, or slightly convex (not medially sulcate)and glandular.punctate, glabrous. Iower surfacedensely silvery lepidote, margin entire. Inflorescencebf small sessileterminal umbels,frequently on short lateral branchlets;pedicel 2 3 mm 1ong. Calyx shortly hemispherical,0.5 I mm l.righ,1-2 mm wide, snooth, closely silvery to ferruginous lepidote, margin undulate to tru[cate; peta]s elliptic,to 2.5 x 1 5 mm, white to pale yellorv,silvery and ferruginouslepidote outside; anthels broadly oblong, ca. I mnr lolg, yellow; ovary subsplterical ca. I nm high, silvery lepidote; styie slightly shorter than stamens,glabrous or.with a 1'ewbasal scales;stigma srnall. Cocci erect or slightly spieading, 3-4 mm.high, verrucose!apex rounded. SeeLlJ mm long, blacli, iiregularl-y longitudinally corrugate. 90 'uleqr ueo,nnleq""n1o r"J1l, uoltezlpyq^q ro uol1?p€.,3relull3rll tsa8Snso1 Istroleru ,{ue ueeslou el"q I (uloJ J"al ur ,{lledlcutrd sleJlp lr rlslql\ uot11 unsoliuuonbs dr o:l p€lelal ,41eso1c,{1luopr,te st setcodsstql rl8noqlJv 'e)j€'t 'q r 'q'rq lr / qn^rsr.ere es.reos stllqpues Suuelnpun ertrtrAlJo ,f;tt..try :Tq"i*[ 'e1oro9 (,46sN) a-pli:tar'9 dsdtL'.rei5 16 I 3o3N ru tll zz.uortv.a'llltlN {o!q rs'elooqlur(r :ifst,l Avt iZ putl).V I t'lJt

:uo'or^r,nL Jo-r r"1 .n,.,o,,'1 L11..rriir'pffi,.!.,iil,,;t'l;llli'i'.ll'ill;".',l3lIjl#' 're^r'r xqrclld[ sri l"uerotuc]ctanrqv lrlai irist 1it{ :snd,{loFe'I-.. rollent\lg '{errntl e{1 si,.re--,no1trrrutrbc e:e'rtel eqr1o uasaippui. solelunon u€IdluelDeqt q ', : u'|lit:Yl:f:L uorriqcr'^\or.,Pu: souelsl (€e8r) €t€:T .,,"#'jl#:zqr;ji,,j1#j1d;1jf :':,;Jj'jjjY:l:";1, 'ue^\t 'Icelg:(9161) :(tz6l) tit IErtsnVS IJ 9II ld sol€y'AS N snsue)'eqrlegle uapIEIA '6I eqctag te ueprc11 ( qlrreg) un1,{qdouals runtteqaqd

'dBFeô uollnqtllslp Jo s"el" elsq^\ ueê sJlJedsJeq:lo q1l,t s€ selceds aceld se1e1uoq"pclSlelul ]?ql 1se33ns01 ecueplla ou uoasêdsq I 3'a) tcuuttp i se li dutureta: *e'I QunudP dsqns purz wry1oftttttd^'dsqts unsuptwonhs d prtz wnsolnfuDlt d Jo sLtlJoJeuos ol IEIIUIIS qtsnoqllY 'sa1e5suno.,q qsippr-r na1 [ {[uo rl]lA\eloptdal-,{:e,t1ls,{lureur eJ",{eql sels,ld laptslno qlriog ,ta5i ur-i!i.req.^r aloptdelsnoulSnrJoJ eJ€ €lloroo PUB x,{l€c Jql PuolJrAul luJJ+tp oslE(t sellE o11lrql tlt unluJutnpulJqf sJlnl\nd Le1npue13-1ûcri:qdsrr-ueL1o3-rt1'rie.1 e rprt oleclns ,411erpeuecP,Jrns Jeddn "ql 'JlEpioJqoJJc sJÊJlJrll sJler^dqlno5 ,treg uI sJJld" pepunol qll{\'al€Fund 'seêol .,e1npue13pue a,noqex:'ruor {1}r13t1s el€ôqo aêq suolulcodsu"uolcl^ eql ?loqqo] ()1 ls"J-rl ou ",{urg ulorJ sal"lA qlnos nd.eNuJels€e-qlnos uilace1d piri,rls ulo{ pue'(su?tdtu"rg eql ruorj flltglqnop osl€ pus) ?uolcl1 ui oariuag reeu sorJrltcol ,'lreJ" tuo{ popJotal ueeq s?q selceds slgJ

'sdrlrqd '^lloung ;ffi I96I rl9l g l,{ ru'l3lrrsurtd:('IEtl) t16I xl-9,UUtu" ^ t sscroluelD :YIl{orJIA lrrtsdiq\\'oSrpura ru :129t (l:Jt/,l) 8Z6l 1.O'plu^\oH AIsslll !r " '":liij&iV a\oqE.' oo/ pr !u!,r\puE\ ue,u!,\r,r 'u'* ; l;ti,l:"I';ti';Tirl,'i'"',it sui HLnos ^1{\ .1 eroqqoj:1,6511) 8la rBEqurPJ H U qFiq sl Pr'8urlP(\\:sllv\\ 'E€ 'Euo}cl^ delr'{ seleid qtnos ,,\\eN lErluac rllnos u.Ielse,\\: Nol.lnalulslo ^ The,fornr of P. ttenophvllunfound nearthe Warr.umbungleRange in New South Wales cJoselyresembles that from westernVictoria bu1 specimensfrom the.formerlocality haveleaves slightly glandular pustulate and a-calyxwhich is undulate (not truncate); the flowers are also slightly larger. - In some collections (e.g. from near Dimboola, western Victoria) the leaves are . narrow-oblong with only moderately recurved margins. Thcse approach in appearancesome Victorian forms of p. squonulosuntv,rbsp- squamuloutm, which horvever are found nuch farther to th; east and differ in midrib indentation and in indumenrum. J. M. Black in his Flora of S. Australia ed. 2 (1948),recorded this species from near Bordertown, but I have seen no specimensso unnotated. The lectotype,collected by J. Dallachy, is statedto haveconre fro|r Mt. Abrupt in the Grampians;this is the only collectionI have seenwhich is unequivocallv stated to haye come fronr this Range. " " The two other collections labelled Grampians are without collector'sname or date, while in addition, one of them is on a sheetconsisting of mixed material.

Sectio 2. EriostemoidesEndl. Endl.,Gen. Pl 1156(18,10); Endl., Enchir. Bot.6t2 (1841). Phebeliumsectto Lepidotiu Reichenbach, Repeft. Herb. sive Nomencl. cen. pl. l:197 (1841), p,pte.,nomen. Branche.und lerrrc'lepid.,lc. Innorescenceof arillaryc) mcs(sometirnc\ bearirtgonll_onc llouert: pedicelhibracteolare near rriddieor rowrrdsaper, passinginto.fleshy baseofculyx. . Sepalsfree, slightly imbricatein bud; petals rmbncate, glabrous, \\'hite; sttminal filaments somewhat flattened. slabrous or steliate near base: arrther suborbicular, slightly retuse; disc prominent; ovary with a short, solid, sterileapex; stigma _ equal in width to style, not lobed. Seed_broadly cllipsoidal to sub-ienilorm,'2-2.5mm long,'adaxial margin shaight; attached (placenlal) endocarp thin, translucent,caducous; aril, meinbrtnous. lincrr, along rdaxial face; outer testathin, smooth, glossy, dark-broun: sclcroteslr.moorh; Ir ilr rnr sLrperFcial. narrowlv eliiotic. ca. ll5 rnm Iong; raphc small, thin. covered by outer testaonly: chaiazuon adaxinl face torvardsbase, slightly obscuredby raphe. Lectotypus: Eriostenon squameusLabill. Two species,from the easter[ stalesand Tasmania. 'TIre phcbalium _ relationshipsof tlris section are with scct. Gymnot.lotlosr,nd (probably more. directly) \\ith Nemqtolepis phebalioides which is technically separatedgenerically on the brLsrscf having ljgulate staminalfilaments. It is fnterert'ngthat tt P. s1.uameumtl)e positionoccupied by the ligtle in N. phebali- orde.rrs trKen by a stellatetutt ol harrs,again suggesting a relationshipbetween the t!vo.

20. Phebaliumsquameum (Labill.) Engler Engler,Nat. Pnanzenfan.3/i4:1,{l (1896); Druce, Bot. Exch.Club Rep. l9l6:619(1917): Ewart, Fl. Vict. 708(1931): Curtis. Srud. Fl. Tasm.104(1956); Beadle et al., Handb.Vasc, PLSyd. Dist. 319 (1963). '!':.,it#fi:,::{_:!f:\HfliJi"..,!.'"t'l;l'r;,i'i...'"','i?1,'n'n',',19"rl,rT,i.ifi",,.i.,i"i,'!l: '.:i'.('8?iJ; (1887); ii#,1:'lfii:il, l.ili.'.!iTLi. i,#ifJ!,i'..il",lii: ,l.l,3JL:t?i.,i,.",'j, Moore" er Betche,Hanib. I-1.N.S.wates ++ tttijr; CoO"ly,-i"i'-.'ft. Z+(rsOf). Typification:Habitat in capireVan-Diemen.. llotypes (?): MEi 5025and 5026. cvclop 2Trn3 (1814)as to nameonlv' rom illes (r' sq'lam"'ls i;f:ii:li11it'il.l,iLRees' 92 .,aau, 'st s!êc rr^,t^l,u"t* ]ne.I sul):(tal ) -$zJ '(oÊ'lt'\t . pue poo^r'r(a)qJ. poo"ti-"o]1n1". se ir'rou1(i1eco1 " { sz oorl Doos-8o61 . . qrrrqs rLU :vrNvnsvf pr-.g'i e i..t't:rgrtc i-ap,qriraI N: u I ' ))ôc Jo N t ,;;:l"i .u '.1].,0,92'.1 '('t3N)6"u']sv I ual'H'q8rq 6."-1 lle'uosnErtl "rs'rc"id,i6 '13t"l uellv ru:vlt{orJl1 pre lttH sre^€tuee^\req :(tean ) r'{ I'looqureurrEid ru :l\ASN) 'r^^SNTr oqJliS poo$fiullds :l I ll ) l8Z ue\sLurrs'uo\'lrEflro-d ttRI r;Yutro I r'..utoo*' cl'rEJ.rnolor(rrnd ro i$.,}*iisr\ J;J',r ti"'!'.'.,"r'jo.t":'tridI-os{l; lirlo su-rqrJIrrErpLu;lt qarq ru8nrreq.p)p^\orJ u)q* olrl lc,r*u,q*-pudiii.ro'fq6nrq.:J ;qnrqi 'speoH pao^\J :1]"gi"iir i l"c"i",r :illgj v sra>r :epuals ;1; ffi":"J:: -lA'e8uL'Uuosreqd.el{'IoolqBuucls :(0l.669,IASN) peols A I'sl urono:cNv-IsNaano '97 del^-utueutstl : EuolcIA : Nolrnalurslo : selc/ qlnos ,{AoNJo lssoJ eql l€au : puclsuaano uJe:}ste-rllnos 'poo'{\auuds-,le,pu"'€Iuaru '(tE6l) 'selEy'i\tllnos ^\eN alqiN-qlrurs ePu 'ou 'l'rii r'uo'u'tl u auosoLuorqJ *sr ieuotiip-arp ur pa1ce11ocl"rreluu '(snorq"l3 ro slnlJels'{lssuep 01 '{l3sr€csstueruulll l€u[u?]s ,lel"r) Jleq la,$ol uI elnsllq 'wa^Z'Z 'tudrlle sau^J '"l"u,o,n'"e oj asnlqo x 0I 01 :p3re,rou-oz S(€) 'rBlnBue 'qBlq ;ino""oliriti iiicel 'q1oo,us slalqcuerg u zl 01 eerJ unaut€nbs 'dsqns 'e67

:asnrel 'dsqns-;-J" 'coz psla^\og-(9)t-Zsallrfc unsntor 'si61 ra^-rEInPuelS II'sJ.,ds slelqtuErg fi;;;;4 ur g ol'snoar*lrtqc seâal:esoru 'dsqns q0zu19)t uncJEllor ol so^€''r ^r a(r) saulrr:pepunot,.ede'snoe'elror iuol Lurz 'aOZ unauenbs'dsqns 'orsulunr€ 'sno1T::€:i iei",,"g-67-e-ls1 .0.,% oresnrqo ":H"_::'"d l;::.::':1":i]""," 'leprosdrlle '8uol ruu g Z Z ^lpeorq prJS JsclttJrrlJ;^rc1npuc13^'.'13ue .te1no uo elelirctde^lslnulur pus poPunoJ asopd tjrq?,1.r"in'q8rq tuuig cj Burpea.rds{pqirls rrioj :scqaql.spre*ot {r:\rudsAlJrBl ro snolq€|tssuJ.!els ol lunbo :1{1s:sttolqelS'JEllrpuElS'q3!tl iler'-,aqdsqns t>lqlp!'lt ru,u er o1 lcptutur,{d,{lpvorq '{rt;ao :(i) 3^neiu,'Areô 1 'qsr,l jirp:3uo1turu,8,g ''r-o-t11Y€'snorqel? ui 1ni,u"-1 u'rug.g i5'1crqr 'e"roqr::? :o 41"11"1s)tlestiilp oi '{lcsruds ela:e]Sutruocaq ro rreu.ro^\otur etns.LlLl 'ruauit 'slutrd ;;'i'":";";l ; o],i'spioLoxtriiarl' stuaurelg or tllSurlur lEnbe "u,'qi'3,'o1 'rel :-':;.;;;'; ;;i"t pi'nra -u' E-trleritdrlla slErad :(norqElS :3uo1tutu '1eprosd111erpunlq -; p;;Li-'illqi'di';1 *r'1 "r'p,o1"p s1udeg 9-7 '8uol snocnpec-elnutur tlll'n^ urur s p,iq'c.i "'rn1,i* :;1nq ia,*o1 u, salo:1cziq z-0 'J)f,loJJuEl isnornpcc'\\olJqrloptdJl ireilrs'Juo1 tutu L I sl)Blq tarroad 'rPInBuu '6rou':iioprdet,{railrs {1asop 'lJJrpJdpuu elJUnpJd:PJJJ\\oU-02-[ ':tu'{r csoolro aJurrsog ir[jn"r :o qrau:17f 1 ot 67I xrel;txe lrPdLuoJJo -ur 'asnre.IJo 'papunol osnlqool aleiii-nct xed? "ldolaqlueunuold PUB pue ,î,onl"ii".,J.ut i,.ip,* eloprdajf.re'tlrs '{1:solc eceJins r3^\ol'uaar8 IssolS .;l;":';";;;;o;;'!j,irrn,."'dhn"".,nui rvu'iuc z z x 0l 8'0 x 8 l'l"clldllle 'rnoolnlrt:q" toc ?urtu€l :el?loqed '{l}roqs i,;;"'.'.d" .l';t;;.,rnu" o1 sn-oacut 'qnrqs rllo"'t' "i"r"r ro relnSucslrLqcu?rg q8rq ru 71 ol eerl ro '(uoesoloqd) ):snd'(loloH ., loo/ SurldsJo'{llulcr^ eql sJedl€a:llt8l) ur \rrt!^!ir!pp:peq\ ur..:uorlrJUrd{f rziet,sos-t l'^s loB d:u- l6l:l l";tl"bo j65 SooD'pleu uI LLreqBuIuunJv untDla 'J ;;,; 9 :(9;311 silcfi s it tuav't '9€l 'nalssnf 'tDtt ollq'd q$ u,{sold'uetuou llc do V xe reqols saPloudD2olad '(0etl) : (€981) tt tOanSprrql H plsuaenou'{S.'Itlteg fbe: I 'tueqlu.g reliioH-17v311Egg:lsxs log cleu'srecllel re:lsnv Il : i9EgLl:t1; 5!: i:irjiil ,::Xdg-;;illiî;1;l;3;9,;;.;.";;;,rlTt;,Ji$N 1gi 1 ,iotsrisi-r,zriori'uoc roti 16l:I rstH Urc D :{lz8ll t9li .:"iiii lr,oil '.. rii"tprnrrls,. iiiSl)t€l:7 s.Pa IPN lslH Jos Llrr[^l'nerssnfv ualPrDlllqd 20b. subsp. coriaceumP. G. Wilson, subsp. nov.

[coriaceous : leathery] E.ovatifotius Ir\on (F. Muell.)F. Muelt.lF. Muell.,Pl. Col. Vict. 1 :131 (1862) p.pte.; F. Muell., Nat.Pl. Vict. l:75 (1879);F. Muell.,Key Syst. Vjct. Pl. 1:l4l (1887). P. ovatdoliumbron F. Mueil.lBenth., F,. Austral.l:345 (1863) p.pte.; Ewart, F'. Vict.709 (1931)p.pte. Fruticulusramulis levibus. Folia coracea, late ovata usque ad lateelliptica, usque 1 8 x 0.8 cm,apice rotundato. Cymaebreves, (l)2 4(6)-fiorae;filamenla staminales glabra. Holotypus:MEL 4833" BetweenHardinger Range and Mount Wellington ", Marchl86l, F- Muellea;Iso.: K (photoseen). Shrub with smooth branchlets. Leaves coriaceous,broadly ovate to broadly elliptic, to 18 x 0.8 cm, apex rounded. Cymes short, (1)2-4(6) flowered: staminal filaments glabrous. DISTRTBUTTON:known only from the mountains near the head ol'the Macallister River in easter;rVictoria-Map 26. VICToRIA:" Source of theMacallister R.", F. Mueller(MEL 483,t);" On quartzrocks towarcis thehead of theE. branchof theMacallister R. MuellerfMEL 4835).

20c. subsp.retusum (Hooker) P. G. Wilson, stat. et comb. nov. P. relusumHooker, Journ. Bot- Hooker I :254(1834) Iretusus : blunted(notched)l; Hooker, Conp. Bo1.Mag. 1:2?6 (1836); Hooker, Jouin. Bot. Hooker2:418 (1840); Walpe6, Rep. Bot.Syst. 1:505 (1842). Typification:" Dr. Scott.Mr. Lawrence,1831 ". Syntypes:K (photoseen). P. billardierivar. rrr?ru,z (Hook.) Hook. f., Fl. Tasm.1:63 (1855). Shrub to 4 m high. Branchlets glandular-verrucose. Leaveschartaceous, oblong-ovate,to 3 x 0 8 cm, apex rounded to truncate and retuse. Cyme short,2-3(5)-flowered, staminal filaments glabrous or sparselystellate near base. DISTRIBUTION:north-eastern Tasmania Map 26. TASMANTA:Prosser R., shrub6 8 ft., Dr. Story(MEL); Rockyplaces on the southEsk, Launceston,S. Hdnnaford31 (MEL); FingalCk., A. Simson1176 (MEL); Mt. Barrow, M. E. Phillips,16.xi.1960 (AD).

The geographicalrange of the typical subspeciesis liom southern Tas- mania to the Macpherson Rangesin southern Queensland. Throughout its Cistribution jt exhibits little morphological variation. Compared with the mainland plant, in the southern Tasmanianform the leavesale smaller and lessacute, and the cymesshorter; the staminalfilaments also changefrom being sparselystellate or glabrousin Tasmaniato denselylong-hirsute in " " " Queensland. In Tasmaniait is known as Lancewood or Satinwood" and in the north- west of that State grows to a height of 12 m and provides timber for telegraph poles (fide W. M. Curtis, op. cit.). P, squameumsubsp. rel?./Jr./rlis fourrd in north-east Tasmania. The type has no indication with it as to place of origin in that State; however material from Fingal Creek matches it very we1l. It evidently grades into the typical sub- specles. P. squameumstbsp. co aceum is superficially similar to P. ovatifolium, It differs from that speciesin the absenceof sub-floral bracteolesand in the glabrous (not lepidote) ovary. It was on this subspeciesthat Mueller, Ben- tham, and Ewart, all based their records for the occurrence of P. ottatifolium in Victoria. 94 'tsuol '1np,o,au1" Luru7.7:1, '{lpeorq ro uroJrser-qns papunol E-ql!'\\puB elecunrl flpeorq peeg el8u" lelno uo runlnrtdc lJoq,s 'snorq"l8 ,';ugril rud g ric '1co.rercco3 eif16ol qlprnt u1 lenbo^f ELuSrls pg(1s.i:de e1ua1.11vLu. E Llll\\\lJdlBJ'atoptdol ,{.rlr1rs'qBrq r"uut I PJ'lEJluoJ ,{r:srrrcooun-,(touolq (lcro:lrtteu'(Ju\o ol qlpl\\ut lnnbJ'qdlquu E 0 eJ 'rade 'r"lnJlqro jt'p ttrut g rr 1e osnler ,{11t131sl-o poPunol )]r,rtt :3uii1 6-uut11 'snorq"l8 'oleJe}-qns'lepuels -irop:ot r.,"ulon 's1e1:d -r:yoqs ,{11q8r1s 'sno.rqe1'3'1pnq 'elepund ,u"ti,..1i : p""lgu, ,.1r1 ur efrsin6 :iurc) clrq't re1npue18 :ct '1err1dt11a':1elt.,quri '3uot ruru 9 s1e1ed:a1r:lcund r"ln-pu€13 :(alopldel ,{ EUuOlrJUo)sno.rqe13 3uo1 ultu z plL,llJp-JseqlE JlDJIlqLul\llqblls lJJJJ-qns '15rqr tuLtl EJ sled:5 1r.rrp:dqlrn 5nollulluoJpun {tl'JU)seq l:rrou:buol t '1iprbsdr1le,(11,rn1q pnq-reôg eJnleu :sledoseql olul ep"ri eseql_'ra'l.rou3ql Jo tiorjq tiirt \JloJlJr-'rqrPLrrur< Z puE 3uo1 ru ut I l I slJerqele\o-;uolqo Z T (l8uolts ur,^ i6uoi LuLu1 T riqriSorp:uatleg PUElJrql lJJrPedpue slJunPed :ijur(r pa.r:r.o11-(t^lJ]er) I ftPlIrxe polsa8uoJJo eJuecseJogul popunoJaseq 'pJpunor ol asnlqo rctlu:a1optdal ,{:e,t1rs{lesuep puz tllôoursec€JJns, le^\ol :kssorS'tonqur ua^J)snoJq€ti utoorus;olelcund relnpuef e3€JlnsJsddn:]"U '".r'1u!'uir ilec4dr11e 'snoec"uoc i t I t-l " g1-9 r 6 ^lp?orq ol e1"ô.'{lp"orq ':1elortod,{llior1s ie\EJ-l- JloprdJl'noiiiSn-r-ro1-a1ud (lJSolJ'esoJnrra^ JElnpuElS (tJsJ[d. 'Jlara] ol .rt'lnduu{1tqBr1s stalqcuclg qnlqs lJPdtuoJ \\o-I suelllrssdsu?uo}.I^ Furpnpxa IIE:(f€61) 092:e6l Z pe ]r' do 'relFug ruoxl(l LuiJuezuEud :(9061)6i, sale/'\ S'N Id :(968I)l?l:t/t ''osl"N'r€lnuE:(t681) tt saraM< N-r.r'qpuPHeqctog r:cLLr,-'rru:i7g8i)gsoi:1I sote/ASN uul.I'ord'Surlrlls ,tisdlirf rst5 t5;' e/: l 1?r^ c.:(ze8I)l€l: i trii 1j 1.rL4 1|en1r1t^:(6t8I) 14,1".!.t.'ll:_14" uow)tsô!]v .rjj",n-tioS 1ant1 g:(6q911€0t:l tu8"ll'llant',i J (IIen!{ J) tunlplunô '(uaos sulelunoit Fun,(unltsnqrluoLtl q ...'t 1..rellanl^l proi oloqd).. 9Suel 'zt8t 'o€8€-!z8t '.,11su'e1,ino14 trje,tu'ila - 8ts69 ,^^sN lJl\ :srd'{lu^s.-^..sulElunot\l g.iirurjw aqiio st:Bdjurtite.qr u1., *r;^rlltrMouSpue xe:rn1,l1 'sd1yue,le.rtsny':-q'i,ii;5i:"i1:l;.,:l.,iiil,]ti.l SZSIiSl'l:sndilolrr-l e.p1o iei,nbs-cq, re i.J,,i:if:}Y.1",{t qt:8 r'l'\ \\rl lo8 : .td ordrr98l' stt: I lPrr\nvl-{_urfqlue8 :...tunIuJrlc\o.. lqs8l) -suera' f '11:n1,1g:i1ej1 l iunrli,Jp:deqs-Fir - snleôl(sS8l)66:11'IA ros lrqd ll'nn J 'lloill 'Iz J I unlloJllBô urnllEqaqd

'JEL 'Lutlsnl)'t tunawonbs dr :luxlcl utluelus"J oql selqueseJ ,{t:sot::.rol:req} puE s;'\ral esn}JlSuIA?tl ur uollcelJoJleuloJ Jtll tuorJ sla$Ip '2981 'H isel€/ qlnos ,4\aN tiqf tttioS ,LriNt) SOottI o ou o6"q!ri"J N 'uEJq^) ru ol1tq{trrutl lrl pJ}JJllosueaq seq (aluels)troullc3os lelltuls V 't1tr,t 1leepXltLol.rejstlus Jq u€J srLllaroieq portnbersr ecuaJse.roguIJoezrs-irl pui'edeqslee[,ursletJd aloptdelSur'req I"rJa]€ure.roJll 'sl€t;d 'sledas irt s.l"gtp 1r'um4t1!1o'to'2iuiorg r.rero puE rloptdel 8ut'ltq ui serids teqt ursepal;the true sepalis then very reducedin size. The lowestsepal (i.e. the one following in the spiral phyllotaxy of bracts and bracteoles)is lepidote on the outside, whereasthe others are glabrous. This specieshas only been recordedfrom the Snowy Mountains in New South Wales, all collectionsbeing from within a distanceof 12 miles of Mt. Kosciusko. The Victorian localities given in several works for P. ctvatifolinn all refer to tlre similar alpine plant P. squaneum subsp. c:oriateum;this differs from P. ovati/oliuzi in having no sub-floral bracteoles, a more slender cyme, and in the slabrous ovarv.

Sectio 3. GoniocladosP. G. Wilson sectio nova [gonia angled, clados : branch] Ramuli et folia lepidota. Florescymosi vel solitareset axillares;pedicellus in dimidio inferiorebibracteoiatus. Calyx palelliformis vel hemisphaericus.undulatus; petala valvata, extra lepidota,alba; filamentaslaminum applanata, glabE; antheraalba, acuta; discus brevis;ovarium apice sterili brevi. Coccierecti, obtusi vel breviterrostrati. Sentenleve, latereadaxiali nec concavo; hilum anguste ellipticum. Type: P. ancepsDC. Branchesand leavessilvery lepidote. Flowers in cymes or solitary and axillary; pedicel bibracteolate in lower half. Calyx patelliform or hemispherical,undulately lobed; petals valvate, lepidote outside, white; staminal filamentsflattened, glabrous; antherswhite, obtuseand apiculatej disc short; ovary with a short sterileapex. Cocci erect,blunt or shortly roslrate. Seed narrow-reniformor bluntly ellipsoidalwith a straightadaxial malgin, attached (placental)endocarp thin and translucent;aril linear, fleshy,situated between two cartilaginous strands, easily detached; outer testa thin, dark-brown; sclerotestasmooth; hilum superficial,n arrow-elliptical ; raphe shrunken, sub- basal,covered by outer testa only: chalazasub-basal, + obscuredby raphe. Trvo species,both endemicto Western Australia. This section is evidently related to sect. Eriostemoide[ from which it may be separatedby severalfloral characters and by the seed. This seedis peculiar in having two persistent cartilaginous strands on the adaxial face between which the aril is attached. Similar seeds are found in the genus Chorilaen.t and in Nematolepiseuphemiae. both of which in other chaActers also show evidentrelationship (e.g. in the hemisphericalcalyx, lepidotepetals, and acute antners).

22. Phebalium anceps DC. deCandolle, Prodr. l:719 (1824) [anceps : t\\'o-headed,referring to theangular branchlets]; Jussieu,Mem, Soc. Hist. Nat. Paris 2:133. tab. 12.I.2 (1825);Poiret. Dict. Sci. Nat. 391464 (1826);Sprengel, Syst. Veg. 4/2:164 (1827); G. Don,Gen. Hist. l:791 (1831); Bartling in Lehmann.Pl. Preiss. l:l7l (184+45). " Typification:in Nova-Holl.orientali." Holotypus:G-DC (photo seen), " NouvelleHol- lande,cote orientale, Mus. de Par- 121." IJussieu,Mem. Soc. Hist. Nat. Paris 2:134 (1825). givesthe followingreference to rhe type (?) specimen" Vidi sp. sicc.in herb. Mus. Par., ex Nova-Hollandia occidentale(Porl du Roi Georges)."1 96 L6 'turoJlllrl€d '3uo1ruru (1ep;os uo plnpun'q8rq LutuS l-/ 0 xtlEJ 4-9 5 -drtte pnq ".,n1eui !'"r1nr 'tirlJq '{Luoqs u:ljo pue lecrpad ,{Dunlq 01) l?cl.rJtlos 'luelsrsred'?uol 'rseull jloq'r"obit ui itsureile io elnoddo-qns ruul E 0 jo 'eloprd:1 '8uol 'r"lngu",I i"1611"^nrq: alelirelcerq-@)6 '{r!'r'1s Lutu 0I-€ leclped i,{.re11*ru 3q} olut Surpur8 ueql pu? el"aunc :de1r1os sre,toig . e1o4edlroqs pJPunol tllr^\ p]qoltq ol esnlJJro p'PunoJ {r,toi-reu ot flipuno.r aseq'sJqol 'qlootus '3unb,(uailn atoprd:1'(rr\lrs {lurql snoer rlJo-:-orqnia a"rrtroreq 'aleprooqo 'al"^oqo -o"r" ;ai gg :o (o,l-fteull) 0rl1u0 i 6r urui 1:rzlncrqroqni 'J"ln8ue 'qnlqs 'snoecerroo oj snooce!eqc se^1le'I slelq'ueJg llBtuS posea)(sl'r)80r:6'rrr do'tant{,,(urr,,.0,'l]lio'9'T!'rr!"X:t'I:f#;13':'0";{";i& '616t -fEI l :ed,{}osl ..6[02 oN ssrerd .qroH0'8r'^oN 6roue)) srrrrgilrslqrl)iforor;l,TTff";:iiiy:::,"r.i:'Ti,'{li#;f;#l} 'O9bt'l8bt -llN :sedlro-'I .,s602:u,ssre:deraH.iJ,!il??I,Xfi',i::,$rib:ti_r"r_,Ldd:?.il:t jlSli#-r3yi :,g^,Jtl"gr:5i;-,Tj$"'i'll$i.;:;gt'"r.,,i,,-iJ?,:T$l?',$q;fL!';,ffi,?; ffiij.fil1*x,"li '€z '[ €B apnI utnllEqaqd

slr oqror Bur,do .,ruerd rers,rs,, ",:!1ii!ti[!ii:l:'{#f#dl d?s lcere qnlrls ? sl 1l '1seoceq1 'i"q1oi,o.rj .,e,3morb 1ou s:op pu e sdute lrs prte sut-8tcLUlaelJ Jo ',"ic"as iu*i qrt^\pteu eqr ir1ezlplqrq b-1, pu€ esue, elllll sltqlr{xe-.,:-",,9.q' sdaJu'-]-9-tl.::t1.1 Lunuo@qd leciqonraoda slr tnoqaiolqt uollEIJ?^ .erepurer t :(^^s N) r06 r'ô N'pr"ro'zrr,t nn''-"ryya1ap$2lstif;ffi f9,p€ft i ,,l]jiri"lt:l*lx;#,1"iiiil.i:S Jt+r;li r"aq'l;:rJ:.3'?9\'lflll*t, 'o,." ,., ,^ gZ d?1,'{-ls3,tA-qlnos'3rl€llsnVuJelselN :Nol'lnslulslCl 'u^\orq'qloours''uorluursz "i'l"gf",:$lll"StitilRii:i".J',?,?il';Hl'T:;i: Surpeerdsl.roqs € lnoqll,^ lo qll, j*"tl'*11".';'i::l'l: ;tm*":lian,' i"ijTi i:":,i',?Î^l'"T:,i:xlfi" ;"d;'"t';;i;'li;p'aai x'ren-1,sqs1{ uû i-1' ltc1uoc-(lptorq x,l.l",t"',1:iJ;'Hl,i"J:,.J#:lffiô1..1erueqds*ueq','.3i *j:"*ti:l*"i*;';fi :::, *1, 'rsEq le peorq r,i' nt "iopror-8uo1qosroqtuc :spue16e.tettdrls ""J n 7. 1 '3uol .1']i erp ue,ql 'el€urutnce'Fq;;;i;;-;1.,ito"'ol"loecu"l ,l1,lro'r.,tusluaiuelg tuLu g 5 el.'s1e1ed 'op,rslnoeroptde-l ;;;q. rto..'rn sua,uE;.:u'tllr^ snorqelS 'allLl,lt 'urut 'tc '1zcr1dq1: snolq€18 ^ra^rrs xlesolc g r 9 '(ipeorq s1e1ed:ult{lltl 's3qol peplÎp :.p,sino "lop'.1"1X,rr,.'1rs'qtootus proltep ^lp"olq olq ^"'\\-llEq 'l"cueqqs,,u"q 'l"cueqds .rrgru u.rtu c.T "o ipiotjt x't1ej auol tulu 9-t in",i seioalcdrqlue1s1sred.- 'e1nu1ut lo rpd p'L;i";il i.,i1n* 'ilppt"i -jq1 'xede '8uol i qi,^ ao61Luru 71-7 sfeciped :snocnpuc spietrolazls ul Sulsearcep 'eloptda1 r"ln8u" seqcu?rq eql qll'l.r *,ii 71 91'"1tt"n-'i1er,.oslie:q '{ra't1s Pu€ pue {le11rre3'rJJuaJSJJoUUI JlslnuolJnul u3Uo Suorljtr-c t sJLu,(JleulutJJl 'ls-rg ;;; ;;.; 3t ".ntqd xidr: sno.rqelSiurr"uocaq te aloprdelfrarlts.{1utt1t 'snooiegefc :3]"lolled f,luoqs Iot"-i,.'Lr'g "'zt-t r " / Irjcirc{rtle euturtl i 'qBIq qruqs repuels rJne"1 'qqoo--i re1€rie ,{13uor1sslelqcuerg ru g .l1

,",t.':i'll$,0,:;':j'"fJ,'l':;iJ3.i'J'':,:j:.ruTlrT.!,HffJliltiiiii'itiiq'L.ffx p!rirqrrursl {ttol).0/ \Jo l'o-ti1.-i.oiiu"ro..iots ui.,."j r"t 1v'qvlsn'utonb". l lerrcnv i*ui;ili,,alfu.ll.*eqlli;;:i"J,1'i?:;:?l,ll,il,i;i'!",t",l3xiilH$"jil3r1 ' :(698r)€0r: r '(u3€rc' rlanl,{ J :\sz8t)zzt z "n r"s ';(i'ld8Jl9ilf;X:'::;"8:'#'E margin or with short deltoid lobes; petals elliptical 4.5 x 2.5-6.5 x 3 mm, white; stamensshorter than petals, 3-5.5 mm long, filaments flattened, narrowly elliptical to oblong and shortly subulate at apex, anthers cordate-orbiculai 0.7-1 mm long, minutely apiculate with glandular tissue, white; disc equal in width to and + continuous with ovary, very short, ca. 0.1-0.3 mm thick, dark green (?); ovary 1-l .5 mm high, rounded with the carpels appressedto style base, glabrous or silvery lepidote, style short (0.5-l mm), glabrous; stigmastrgma + equal in width to style. Cocci erect ca. 5 mm high, almost smooth, apex truncate to rounded, blunt to shortly apiculate. Seedbluntly ellipsoidal, ca. 3 mm long, smooth, dark-brown. Ovary lepidote; branchletsprominently angular; Ieavesnarrowly obcordateto sub-orbicular 23a. subsp. rude Ovary glabrous. Leavescuneate-obcordate to broadly obcordate 23b. subsp, amblycarpum Leaveslinear, apex tounded 23c, subsp. lineare

23a. subsp.rude Branchletsprominently angular when young; leavesobcordate to sub- orbicular7 x 5 35x l7-10 x 10mm; ovarylepidote;cocci often shortly rostrate. DtsrnlBurroN: near the south coast of WesternAustralia betweenpt. Irwin and Cape Arid-Map 27. wrsrERN _AusnALrA: Barker Inler, straggling shrub 2 ft. high, J. S. Beard 2291 (KINGS PARK); Mt. Croper, N. H. Brittan. 1.xi.19581 (UWA);(UWA): Summits of RavensthorneRavensthorpeRanse " Range, C. A. Gardner.Gardner,Cardner. Nov.Nov. 1935 (PtRTH):(PER(PERTH): BrBaldlsland. l, MaxwellMa\welt (MEL(MtL 4963);496J):.. Gooseisd. Baj', It:. MiddleMrdote tsrand, lsland,KectrercheRecherche Archipelago],Archrpejagol, R. Brown,Bro\^n, (MfL'4,(MEL " ",'. 16.i.1802l6.i.l8O2 4977);Miles Island forntPoinl on MiddleMiclclleTs., ls., den.edensebushesbusheq 1oto 5 ft.fl. highhiph J. H.H. Willis,Willis. 22.xi.I95022.xi.1950 (MEL 49761.

23b. subsp. amblycarpum (F. Muell.) P. G. Wilson, stat. et comb. nov. Eriostemonamblycarpw F. Muell., Fragm. 1:102 (1859) [amblys : blunt, carpos : fruit]; C. Muellerin Walpers.Ann. 8ol.S)sr. 7:520 (t868). Tlpincation:'' " ln locisapricis ad llumenFilrgerald River.' Holor)puq(?j: MEL 4556, 935Filrgerald R.. I ro 2 ft.". iso.: K (Max\ ell specimen,photo seenl. PhebaliumambQcarpr.r? (F. Mue[.) Benth., Fl. Austral. 1:345 (1863); cardner, Enum. pl- Austral.Occ. 70 fI931).

Branchlets slightly to prominently angular when young. Leaves narrow- cuneate to broadly obcordate 15 x 4 17 x 9 mm. Ovary glabrous; apex of cocci rounded, sometimesbluntly rostrate. DISTRIBUoN: South-westof WesternAustralia, from Nyabing eastto Cape Arid-Map 27. 'WESTFRN AusTRAr-ta:west River.C. Andre\ s. Oct. l90l (NSWj;20 mi E of Jerramungup.- ' shrub2li. f. Lullfit,,J6ls fKINCS PARK); nr. CapeArid. Max"ell, I875(\4ELr.

23c. subsp.lineare (C. A. Gardn.) P. G. Wilson, stat. et comb. nov. P-:lineare.C. A. G_ardn.,J. Roy. Soc. W. Ausiral. 27:180(1942) linearis : linearl. Tlpificalion: " Hab. in dislr. Flre. ad'. monrernRagged propi lsraelireBay, i; lapido.i. frulucetic.flor.. m. Ocl. Cardner2864 Holol]pus:PFR IH.

Branchlets angular wten young; leaves thick, linear, narrowing slightly towards the base,ca. 20 x 1-1.5 mm, apex rounded: cocci 1 truncate, minirtely apiculate. D$TRTBUTToN:Western Australia, near Israelite Bay-Map 27, WlsrERNAusrRALrA: Mt. Ragged,W. E. Blackal 1154(PERTH). 98 'tuser',{e^\pod:(sr8l)60t:6rx;"rg'tpnt{ c:(91.81)91.:tt8I **l€gg3.iqiiq:;rht;l+ sreded'llonIII:(298I)6zI:I tJrA Io) SrpulId'llentr,t C ( )ooH)srrDtaou uowatsoltg '(S98I) 'relooH '€82 ou setr se^rsrolooH C f €9:I rus€I IJ uI sB I r\\,{q lir]rJ'(ltJrrroru! serl ou uunDoqt lEqt olq€qold sluees 1I (€t8T'€82ou uunC) (€ZZu)'uun9 rl,.\l eeseql]o oqloôqe (0T0t IASN:(o.rs otoqd)) rsecl{tu,{S.. 'eauel^\e'I IeêI io.l OOg't.louolleôle uB te surelurlolllurelsanA erll uo :(IZt u) rJ,{,, :uon"agtd.{J '(9961) 'srlrnJ 'ret8ua:(968I) tol rus€ftjl pnts :(It6I) 092:"6IZ pe lrJ.do otl:t/t ruBJuezuegdleN'rat8ug:(t98t) 8t€:l lerlsnv li'tueqluog:(Sl8l) €9:I tus€J 'lj r'J :(Zi8I) deU'srodlEnA :(0t81) re)iooHloa ro{ooFl 're)iooH909:t ls{S 1og 8It:Z '}og ['relooH'lelooH : (rf 8I) 691: I Id .I : [euE]uoul: snualuoLul(t€81) 9SZ: t re{ooH I '{ooH unuE}uoru luml8qaqd 'tz

'uoLualso!Jgsnue8 eql sleg 'eJn}xal Jo -rueu q1r.tt,{lsnor,nqo olg serlluUe sll ',{uto:}?u1lpu€ ed"qs ul lueJe$Ip e.rese^lesuaql sl€led arll ellq,\\'tcullsrPellnb sI tI'pees s1tur pu€'sJelc"r?qc 'Sleled 'laes I"JOU lArllo Ur JeA3,,\\Oq eleAJeAsll Jo lunotc" Lto otuduolal ul saa8unduoLualsotq pepnlcur ru€rltueg pu? lellal.IjÎ qtog 33u?Jgltl8tsJo ]ou 'saqol ere leêl srql le qcrq,r sJe:lcEllltlc x,(I"c 3uo1 pue utnluaurnput qSnor s1r Jo sfsuq erlt uo s1s(louaD[toq)uoltJes eql pa1"3a]3es,{lluervddt ::11en1r1 'luojeJrp,{ra^,{lpcr8o1oqd:oruare se^lesueqtspJos aql sosm asaqt u nouatso!.g suor:laesaLuos uI punoJ oslz ert sd-recopue1e1ueceld 1c1q1 'lecrd,(tJo 'snue8.I3q}o qBnoUll€JoJ sr drecopue1e1uece1d,(qsou ot{J fue qll'd ro tlorloqaqd Jo uorlces Jeqlo ûe qlr,& selltuuE esoJcou sr\olls uollces slql 'puPleez,\\eN pue 'rruetusul 'selels uJ3tsBg '?rl?llsnv qlnos uro4 seJJedseuo-flu3,tJ 'lprr I wnqoltq unlDqJqd :snd(rorJJl 'eqd€i 'lssuq-qns ,{q pe-mcsqo lo I€sEq ezepqc iSur:e,toc ,{sso13 snoec€lsnrc oJ snoeJEuoc u[l] € qll{\ Jo ,(rlseu eqd"r :clldrlls I"sEq-qns l€s€q',{ssol3 'uql -r\\olJr?u ol J€eurTurnpq :qloouls "lsotolalcs :qlooLus lo llnp 'sur8r€ur Elsel Jolno :tuasq? lu€ :luelsrs.rad l€JelBI uqt qll^\ lnq as"g spla,{T -o1 snoecegoc pue 'dlscopuo (le1uace1d)peqa€ut :lrlSIEJls :l: ut8;eur '8uol {crql 'rujoJlueJ-qns l"ry€p? ruur E ?o pees olnurur €ul8lls :pnq olnl?u ur pedoleêpllo,r el,(ls:eluels xede,fuelo l(luasqe ro cstp " SunxloJ sautDaruos ro) sled:ec sE Suol s" o:l urq] railoqs '.{.IeAou€qt Jo,^.!olJ€ueJoqdou,{8 :osnlel 'elaJel 'slgled ,{ldaap sleqlus :srorqelS Eueul€lg uEql Je6uol ,{ll?nsn sueur€ls 'ateÎ"^ :snoiq"l3 ,{llBnsn sl"led :xed" Iaclpad fqsou oql qll{\ snonutluoc pu€ es"q eql p4run .{ltqAIIS 'uedo ro etr?Ale^sl€das es"q aql spr?.'!\ol l? 'snouel seloalce.rqgo :red c q1,^dfllensn Iecrpod :osour,{c ,{ltuenbe:3 ecu3c -serouul 'sr€q elEII3ls .ro eldurs qlr,tr Jo snoJq"lS sal?ol pu" sleplJu"Jg ',. soploall.{qduodelsolrg sB 'ssnf rTtI 'do ''llenw 'J 'laes !, 'V unausolp untpqaqJ \io pespq llr slsdouaoluoq) uoa2lso!)E .lpEerql : eluau'q100rus:oro (2981)szt:T1.IA'loJ Slpul ld'llanll Joazuolal'p.s uoaalsoyE 'rotaus 9grgt'fflqlti; 'r'" clo 'ret;ug:(968r) o1:t/t u,"Juezu*gd : (t98t) rte r rctJ 'qlueg 'c) ('[enhl Bluouolaa t orlcas

'euqc snonu4uoS ir ruroJ ,{\ou {eq} wq} lsaSSnslou op suoltraallot$^at\oql.wnd)D)tlgt?tr dsqns 'uollnqulslp Jo ruJoJeuo.Ilxo ue ,{luo eq fewr atoaull selcadsqnseql Jo seer" eleredes ,{dncco osle ,(eqJ ',{re,to oloptdel ro sno;qe13u :eq1te luecefpe 1nq 'Jeel qlr,r edeqsjeel;o uoll?ltoss" eql UIE]ol ,{ll?laue8 lng sJ3}c?nqc 1tn{ pu? 'qcu"rq ur raqlo qcee olur eper8 LundJD){lqLuDPUB aPrr soJJedsqnseqJ Small compact shrub. Branchlets terete, stellate-puberulousin lires between the prominent, somewhat decurrent leaf-bases. Leaves crowded; petiole short, flattened,ca.0.5ca. 0 5 rnm long; lamina fleshy, terete to I cm long.long, + flattened and somewhathat sulcate above.above, rounded below. smooth.smooth, glandularelandular punctate, apex obtusobtuse. Flowerslowers solitary in the axils of the uppermost leaves, pedicelI53mmIpeolcel l.) J mm long, fleshynesny andano swollen beneath thet calyx, glabrous or sparsely stellate-puberulous; bracteoles 2 3, close to the calyx, sepaloid (when 3 then the lowest sub-foliar); mature flower bud bluntly obovoid, 3 4 mm long. Calyx lobes deltoid ca. 0 7 mm long, fleshy, glabrous; petals elliptic, ca. 4 mm long, wlite, glabrous, tip prominently inflexed; stamens slightly exceeding petals, filaments slender-terete; anther sub-orbicular ca. 0.8 mm long; gynophore short (ca. 0 2 mm high) narrower than ovary, purple (?); ovary sub- spherical, ca. I mm high, irregularly corrugate, glabrous; carpels with upper third sterile; style * equal to stamens, glabrous. Cocci erect, ca. 4 mm high, apex rounded and shortly rostrate on outer angle. Seed ca. 2-3 mm long. D.IsrRtBurloN: Tasmania, mountains of the north-east region-Map 28. TASMANTa:Westem Mountains alt.3,500 feet, R. C. Gunn283,20.xii.1843 (NSW, HO); SummitofBen Lomond," low-thick-shrub", Dr. Milligan1056 (MEL); Mt. Barrow,plateau at summit,H. M. R. Rupp38 (NSW). This species has evident afinities with P. oldfeldii which has similar sculpturing on the ovary, and a similar white, stellate-puberulousindumentun; in this latter character it also resemblesP. phl,licifolium and P. dentatun. lt differs from these and all other speciesin the section Zeioaeruain having terete leaves.

25. Phebalium oldfieldii (F. Muell.) F. Muell. ex Benth. F. Muell.ex Benth. Fl. Austral.1:340 (1863); Curtis, St. Fl. Tasm.104 (1956). Eiostemonoldfeldii F. Muell.,Fragm. l:3 (1858)[named after the collector,Augustus Old- fieldliF. Muell.,Pl. Col.Vict. 1:125 (1862); C. Muellerin Walpe.s,Ann. Bot. Syst. 7:520 (1868);F. Muell.,Fragm.9:109 (1875); Rodway, Tasm. Fl.24 (1903). Typification:" Mount Laperouse,Tasmaniae. Oldfield et Stuart." Holotypus:MEL 4822, Oldfieldand Stuart1875, " baseof Mt. de la Perouse" Feb.27,1857]' iso.: K (photoseen). Compact shrub ca. I m high: branchlets terete or slightly angular when young, stellate-puberulous all round. Leaves shortly petiolate, Iamina coriaceous, oblong to oblong-cuneate 7 x 2 5-15 x 6 mm, flat or with slightly recurved margin, apex rounded or slightly retuse, base cuneate, rnargin entire or minutely crenulate towards the apex, glabrous, glossy, midrib impressed above and slightly prominent below. Inflorescencea compact terminal cluster; pedicel fleshy, glabrous, glossy, ca.2 5 mm long, with two short glabrous bracteoles (ca. 0.5 mm long) inserted towards the base; mature flowerbud ellipsoidal, ca. 4 mm long, with rounded apex. Calyx fleshy, glabrous, lobes deltoid ca. 0 5 mm high, petals; elliptical,ellrpucal, ca. 4 mm long, pale pinkplnK to white,wnlte, slightlysliehtlvsllgn y glandutar.glandular,slandular. tiotlptip inflexed:lnnexeo;; stamens sliehtlvslrgnslightly y exceedinsexceeorngexceeding Detals.petals, filamentsnlamenrs slender-terete, anthers orbicular-cordate, ca. 0'6 mm long; gynophore sho , ca.0.3 mm high, ovary sub-spherical,ca. 1.2 mm high, corrugate,glabrous, carpels with upper third sterile; style glabrous, + equal to stamens. Cocci slightly spreading ca. 2.5 mm high, rounded above and shortly outwardly rostrate; seednot seen. DIsrRrBUTroN:Tasmania-Map 28. TASMANTa:Near Mt. La Perouse,F. A. Rodway,Dec. 1898(NSW); St. Clair, C. Elliott, 2.i.1947(HO); HounslowHeath, Cradle Mt. " In densedwarf shrubbery on exposedmoor, ca. 4,800feet, small erect branched bushy shrub under 1 foot ", H, D. Gordon,12.i.1942 (HO). 100 I0l eplslepun 'Ja8uol 8ur^"q ur setcodslall"l uro{ sJaJIP 1I sa^€al-eql, J-o -aql 'ellq'$ pualumur J3rllo "qf "o -"luj-ripur' stloln:eqnd-c1e1lels qllrA selceds 'd wryotuap untlDqaqd iliio egi s1 rqcrr4r'^"wir11o!5ryn1dol pal€ler llasolc sr '(,{AsN)8s6l x s :151'0"0.1iqlii:r.$",)if"'b'liiT't,3,[';;'*i,,''1i!id:'i..';l"ii]'l'liB i'Qfr!i"Xi'; :i;s"NJ9!oi',''stub.ur]5115!'t'uztqn-i't'ttel':?olsruc't'pue'xlroruo'rapriqquo\ :$ii;,*t';,j.,ii:llx;:.Xg'J;8,;j,,-:i''s,,ll'.1'r,,1*,1'l'11T,:i ,llx.l1q',1 '62 d€I{-et"ls lqlrou ']d e8u"u JellEjqrc aql. oqE eq1 ol eqt Jo tsee-q ou oq1 uI 'sel€ia -ul,sueqdals qrii,j, jqr ui e.rre.â.e111ruo:.; rs'oc oqt .r"eu qlnos '\\eN :Norrnarulst(I '(iS6I) ollq1\-qlluts ePg '91 : 'q"Jjeld "lI eôJ au"'l 1€ pelcelloc l€I.Iel?u ruoU u ou tulosoruoJtl3 'ec 'uloJ 'u\\orq qsrppJrlrep'3uo1 utru S Z xede -rrar poes altlnctde ,{lpre,$lno Ilelirce pun ^!'"q: ^'":1--31.-" ;1cera ,l-'punor'snorq"l8 el'{ls €uo1 iuui g.g ec t""o3 s,i"uln1t ur:q1 ialoqs Ipq8r ts, 'snoJqel8'qloorus,'qblq uu '€t :xedi: e1:a1i-ir"i"'ipepunor goqs " qlr'lr slJd.r€J^',{rEAo I qcnru e:oqdou'{8 ;t""j-qd; :qsrg Luri g [ ?c usql'ro^\or]Bu 's1e1ed 'eo 'atBploc-lElnctqJo sJeqluE 'olelel-repuels slueurelg !?rrol ruul T 'snorqulS'elrqa\ 'prllop rrir"qi tu"ri,n1.:pi""gi' dtl ol $ollrI elPd b"'ij.*" 'cridrxj '^qsag'q8rq-uu -pul1a'auoi t,il g.6 e-r '1et5d:sno:qu18 c 0'Er pnq prollirpllpeblq soqol r,{1e.; 3uo1uru 6 elx:du pepunorqlr^\1r!oôqo ':jleq 'alnurur pu" snocnP?c l,in1bla i"^b1ur ieloelce:q luelsts:ed o',ttl lc?rq ,rpui1' :3uo1turu il";;; q'i;3uo1 *',u y - larrpad 8-'z'snolrueqnd-rlellrls 'rElndue "1,Lnp"d :q'ue.rq setu'{l ,{trs.rEdsro snorqElts 'qgp1ut :!] .ii1l;'.pa8trct:e'snolnleqnd fue1pxe pere,r,rog-g1'"ttq^ n"io ""u""d"ro1iu1 ru,eulgord-ql^r'tl. -"riii"tt 'fr16orusectgrns re,*ollqgplur posserdurl3uol3 elBl[3ls ^telnuru 'qtoours ol el"cun-rl .lo li.o.rqnia fssbl3 ect'4rns reddn :al"luaprq p"n. 'elnlo^JJ ?11r"] x xedr :elilicrluap ro 51t1uep oJ pa^lnJal ur8letu:tulu,S,x 08 8 St (ol JEJull' -7 x ib tl x oc) JIE\oqo-,lAolluulo rttdtll:-nolleuol 6uolqo-\\oJlEu ueq^\ 'r:.;":;);;,rr!'J,;;1 :auo1 *1tl 7 ec olorldaaeuell?u qlr/t se^?e-I, 'eieral 'q8lq ro-'8uno'{ qnrqs etelats .{lainuru a slolqcuerg ru 9 ol eel} lletus (zs8r)ogtt z / sz'rrc do' z nJ czf,rnI) Tr,r::j:;:r; u6esrou odrr ,.uosuaqd6l!_,! 'eqr: '(f6RI) seF,{A qpul?H ffSii#i3::.:;Llsifl 6";?:"q1iH"il?tii#:j?3i3^* rm'is?;il.,U.ei!i,:ru.i",1$X,gK:;l!;i'".ri:t#i,;if€t S N IC :i'J),y:;,;:,:E 'ueac lou :d{f .. uo')iJB[ nuod e 'JEd\nN qJequ.rrrsdsrplA-..:uo!e4ld'':I.";T^8'l:P{ l";:,s.f9,r,"r"r';;lJ:i;l!t"'l ;::',H.iiPriira:l,?',jif sli'1?':;"i ,If i.)"' ;;r':;;*jn"i.Liir'iv,"iirov"'r'- i lg';' 'ules lou rd(! .. bsl llrque-l I V '{q sn o1 pelmtunuruo.puE 'puDIroH ./$eN Jo lred oruos tuo{ esoJD *o ,o t1?l?ii;i;liil.ati"rlt 't*'!t 'td JSEA'tE roilpplq.lll!9- y:jlglr.sop 01 s" (I€6I)801. r'l^. tt .l?ti,o,j r';3ffii"'t8't';::.1;lî"','*i!'i,;l,Jgilq,!,l,s 's r :i;f l"t,li' ;luE,i;i,13$'iirJf,t"J/ '92 'uIS unl€luop un.rpqaqd

',totta,(st lt wn4o[tJlll.ttdJ ur sEJJrq,\\lpnq Jql ul sdrl >1urdqtt'u) itll ; tie,rb -qr uo SuunldlnJSJql ur put;'(sJuql ul Pue euu,r sr JnoloJr'e"otl 'ed?qs 'unluaunPu! eiiinounpoa ;ou1 sri'to11 {re11,xe,4I?ITos eql ul J?ol ur uroJ eqiuori lcde'sarr:ds jeql urorJsJeJIp 1I o{slncso)'lt{ r"eu PunoJ 'M 'd wn!DqaAd snorqelf oql o1 ,{1et::dsa PJir$qd or rBlrwrs sr tlpPwa dentate, leaves, much larger axillary cymes, and a glabrous ovary. lt is apparently restrictcd in its distribution to near Sydney on the east coast of New South Wales (with one record from further north in the Gibrattar Rarge). Mueller (1862) treated P. phylicifoliun as being conspecific with P. dentatun, but from his description it appears that he regarded orly the short leaved plant with a puberulous ovary (i.e. P. phylicifulium) as being found in Victoria. However in t}le Second Syst. Census, Mueller lists the two speciesseparately and records both frorn Victoria. I have seen no specimens of P. dentqtum, on which Mueller could have basedthis record. Ewart, in Fl. Vict. (1931) 708, correctly described the plant but gives as its distribution the Grampians; again there are no specimensin Herb. MEL to support this statement. I have seen two herbarium specimenspurported to have come from Victorja: one is labelled " " " Port Fairy (Herb. NSW) ald the other East Cippsland (189?)" (Herb. MEL). Both records I consider doubtfully authentic unless from cultiyation. The solitary collection from the Gibraltar Rangeis intriguing and, assuming the details are correct, representsan interesting case of disjunct distribr.rtion. Penfold, 1.c. records the odour given o1Tby the crushed leaves " as being similar to that of P. co-tll a pleasantpassion-fruit odour ".

27. Phebalium phylicifolium F. Muell. F. Muell.,-Trans-Vict. Inst. 1:32 (1855) lPhylica : a genusof Rhamnaceae,fo]ium . leafl; Bentham,Fl- Austral.1:339 (1863); Ewart, Fl. Vict.707 (1931). Typification:" the highestpeaks on of the" CobborasMountains. and on the sourceso[ {he Mitta Mitta." Lectotyplls:MEL 4888, MunyangMountains 4 5,000feet also on rhe SnowyRiver, Jan. 55. Dr. Ferd Mueller", * "; anotherlabel "Inon samesheet Mitta Mitta 4.000 feet ;so.:K (photoseen). Syntypes: MEL 4886 rupjbusallissimis monris Cobra 6,000feet. Frutex anple cxpansLrs2 ft I ft. alt. Febr. 1854.Dr. Ferd. Mueller". NSW 69938. Etiostemotlphrli(ifolias (F. Mue1l.)F. Muell.,Fragm. 1i105 (t859); F. Muell.,Cens s 11 (1882):F. Mltell.,Key S)'st. Vict. Pi. 141 (1887)t Stirling, Proc. Linn. Soc. N.S.Wales 11r1055 (1887);Moole et Betche,Handb. Fl. N.S.Wales43 (1893);Dixon, pl. N.S.Waleszt8 (1906). E. pfu,licoide:[non (Spreng.) F. Mue]l,lF. Muell.,Sec. Cen. Rep. l0 (1854). E. unbelkt.us[!on Turcz.]F. Muell..Pl. Cot.Vict. 1:t2E (1862) p.pre.; F. Mue ., Nat.pt. Vict.1:73 (1879). A compact shrub to I 5 rn high. Branchlets slightly angular when youl.lg, stellate-puberulousin broad strips betweendecurrent leaf-bases. LeavCswitlr erect,flattened petiole 0.5 1 mm long; lanina + spreading,coriaceous, erltire, oblong-ellipticto subteretedue to the siiglrtly recurvcd to revolutc mlrginb, 8 x 2.5-17 x 1.5 mm, upper surfacesmooth, glabrous,lower surfacesmootb, white stellate-puberulous(or rarely glabrous). Inflorescencea corrrpacr shortly cylindrical terminal cluster, tlte short pedunclesin the axils of the uppermost leavesand bearing l-3 flowers; peduncleglabrous 1-3 nm long; pedicelsglabrescenl, f lleshy, I 3 rnm long witlt an ovate,ciliolate caducous bract ca. 0.4 mm long and a pair of minute sub-basalbracteoles; mature flower-bud obovoid, ca. 3 mm long, apex rounded. Calyx obturbinate,lleshy, glabrous, Jobesdeltoid, ciliolate 0 5 mm high; petals narrow-elliptic,3.5 4.mm long, gland-dotted,pale yellow; stamcnsabout 1+ times length of petals, filamentsfilamentose, antlrers cordate-orbicular, 0.7-l mm long; gynophore inconspicuous,ca.0.2 mm high; ovary obovoid to sub-spherical,ia. i mrr.t high, white stellate-puberulousol mrely glabrous, carpels compact rvith a short,solid, roundedapex;style :l equtlto stameus,recurved in bud, glabrous; stigma minutc. Cocci suberect,ca. 3 mn high, apex rounded, outwardly very shodly roslrtte. Seed(not ripe) sub-reniformca. 2.5 rnm long. DISTRIBUTION:Mountains of easlcrnVictoria and of the extremesouth-east of New South Wales-Map l3. NEwSourH WALES : Tuross R., MissM. A. Arnott,Feb. 1928 (NSW);junction of Thredbo and LitlleThredbo Rivers, A. B. Cosrin,Aug. 1949(NSW); Mt. Kosciusko,J. Mcluckie, Feb.1928 aADl. 102 'sul.tunol 'u, onfit ,"rn^ qlnos ,êN l enlg u'$ou){ :NolJnalurslo eql ul ql€arPlselq 1t 3uuet11u3ed'(1 aq1 urorl dluo 's'rorqe1a o^' "t"f,tl tou peespu" llnrg 3lnururturi.t: q'{}:,i-elial:^:ry1}' , 'sirorqelS '3uo1 urrir tc 'plo'r'o-llo:rtu '{re'lo lq8lq ,qr ip,^ ôr.rni, spcfuec i 'JlEproJ-JPLnclqJo ruLu BJ 'Ieqlue *- i.g e" lecupuq,{c l.loqs csrP :8uol :$ft?i;"'"1;'i,l1.:,"J'i*L'1"''J"'JlI .rtilt,'*t';:,'i;!',:,*t:\'";]itil'aool 'ec 'plorlop 'xod€ .tnl.i 'siorqeia urur5 seqol ;;il;;';;;l';-ôtrn,t :aseq pue ]q.ld ;qf orur Sutssed,x,{1e.; rqr lE srlo-rlJ'rqio rtcd e iR*,Llx[r[r'pil j$,t,ilt'.,"";:."l;tt;lif J5,il;iH,i,iit]":,:",?Jf iixl l:tn'ru (p3rncsqo ,t:;i"r., .'r;;;iJ" r"du'sn6lnreqnd-erz11e1t"rrqI 'ruur 'Llloours isprundn PYn,llo?-YspJAlnJ.'{l}tt8rTs ,{1e,nin) ece,;rni ra,ltol eciyls :addn '€cI 'snolelroJ EurLuel:i,uol ruul t"g e""ef"i"f lsoutF PuBelnlorel {1aso1c I 'poue11€U shorqelS SLtruroroq uo,os,-'snolnlsqnd e1orled lJeJo rllt,\\ s3ÊJ1 'qnrqs -J1n11"1s"1tq^ '8uno,{ neq,t .rulnau" ' llq8lls slelqcuerg IFI ' uou] snplDttItld'g '(El8l) 60T:6IIjBeJJ llent\l C l'Jlenhl C ( IItnW d) ':'u"])':|*;1*#*";ff"8 'qpueH'erooN xe I (tu'q8uruunJo, "*;;"ill,3l,q:l!; IrenAI 'etd'd Z8tt pue old d l8tt'IgN '(""*."SNiEl\"3..Y,;,iff'13.1,1"T";iii;;i'';Ti:liX'i,H:''';ii|*fiff *11'rt"lii*?l (916r)9I|lil :l:$i:,1J,.'"i::iB;;l:',',1i""lili"#'iqIgl'1,!X:li'ji';S;5:.i".i,X!;Li#: '::"'itii3'{-!i! 5;::ir,uxi"l't;illi,i;gii** Fi;:,*^"d';is,s-#pii#xTtl"#1* ;T1.jjg '82 'uunJ 'v sap.roaEuqJE[runllBqoqd

'selcadso.t1 3q1 .IePsn selou uI prssn'slp l3q nJ sI slqJ 'sel?^a qlnos ,\\eN sulsJunoj l inlg'tq1 ttto'; iutoc I1e e'rer101 lq8noql Jo ed'{:} qcrg,\\ pu" ueos p€q t"11",,11 q"1i1-^$ploaoiqttl d Jo l?yel?ur " " sB,!\ oqr "ql ot "np fur€d ^tq"qord satr' wntDgap d qlt'lr uolsnJuoc i1p"s

,r,rat,r e 'cuPe

'9t '- 'ou auosoluolrlJ -qlps ePu'elotql"eH 1" p3l3olloc IBIJel"u uro{ u 30a. subsp.elatius Small shrub ca. 2 m high. Branchlets glandular-verrucose.Leaves narrow-obovate,ca. 15x 3 mm, apexobtuse to rounded(rarely slightly retuse). Mature flower bud obovoid, ca. 3 mm long. Ovary sub-spherical,compact, ca. 1 mm high, rugulose;carpels with only a small apicalsterile region. Cocci erostrate,slightly spreading. DrsrRrBUTIoN:New South Wales,extreme north-east-Map 31. of river in gravelly deposits, small shrubs NEw SourH WALES: Upper Copmanhu.st, "on side ", +6 ft., J. L. Boorman, Oct. 1909(NSW); New England anon. (C. Stuart) (MEL 4702).

30b. subsp.beckleri (F. Muell.) P. G. Wilson, comb. et stat. nov. Eriostemonbeckleri F, Muell., Fragm. 9:109 (1875) lnamed after the collector H. Beckler]; Moore et Betche,Handb. Fl. N.S.Wales43 (1893);Dixon, Pl. N.S.Wales48 (1903). "Prope l'iuminis River; Dr. H. Beckle.." Holotypus: MEL Typification: originen Clarence " " 4589 " Mclrnnan's Cteek, ClarenceRiver, Beckler." with the name Edostenon Beckleri written over the name " Edostemon erosus"; iso.: NSW 69929. P. beckleri (F. Muell.) Engler, Nat. Pflanzenfam-3/4:l4l (1890); Maiden et Betche,Census N.S.WalesPl. 116(1916); Engler, op. cit. ed.2. 19a:260(1931). Eriostemonerosar F. Muell., nom. illeg. (proposedin anticipation of its future acceptance) Pl. Indig. Col. Vict. 1:121 (1862). Typification: "InDr. Beckler'scollection ofplants from a plant. If this plant should prove referableto Erios- the ClarenceRiver occurs Rutaceous ". teino il may rcceive the name t. eros4r The specimen refened to is the type ol E. beclc- /eri F. Muell.

Shrub to 5 m high. Branchletsalmost smooth. Leavesobovate to broad- obovate 30 x 6-20 x 10 mm, apex rounded and + retuse. Mature flower bud ellipsoidal, ca. 4 mm long. Ovary sub-hemispherical,ca. l'5 mm high, rugulose towards apex; carpels with a broad, flat outer face, the upper third sterile. Cocci diraricate, shortly rostrate.

DrsrRIBUTroN: New South Wales near the north-east coast and in the extreme south-east of Queensland Map 31. QUEENSLAND:Springbrook, Macpherson Range, 300 ft. C. E. Hubba.d4225 (BRI); Dave's Creek country, Binna Burm, fem forest,erect shrub up to 6 ft., sweetscent, R. Jones241 (BRI); Mt. Lindesay,summit, alt. 4,100ft. R. Stewart,10.x.1932 (BRI). NEw SourHWALESi Brunswick Heads, J. C. Wiburd,Sept. 1937 (NSW); Dorrigo, W. Heron. Oct. 1931(NSW): Comboyne, E. C. Chisholn,June 1934 (NSW); Port Stephens,J. L. Boor- man,May 1912(NSW).

Phebalium elatius slrbsp. elatius rs olJ'ly represented in herbaria by a few collections, two made by Charles Stuart near Tenterfield, and a collection from near Copmanhurst. Collections made in the Lamington National Park it morphologically and suggestthat the type collection represents approach 'fhis the extreme form of the more widespread plant subsp. beckleri. latter sub- speciesI have separated arbitrarily from the typical at a point where there is an apparent break in the cline betweenthe two. It is possible that after further collecting, particularly in the Clarence River-Tenterfield area, there will be no disjunction and the arbitrary division will then break down. However, for the present the subspecificnomenclature is useful for distinguishing morphological extremes. P. elqtius is evidently closely related to P. obtusifolium and.P. gracile which also are found in the extreme south-eastof Queensland. From theseit differs in the leaf shaDeand the almost terete stem. The nature of the distribution of these three species,and the varjation in form of P. elatius, suggeststhat they ave only recently evolved through geographic isolation from a common stock. 106 rlrur s . 0 "c,p'rpu 1lf,c,. .{rrr ous -JlpproJ sJJtllun :snol_q"lb'esolu:."*:l-":ru;,u,.liittT,J; J" ? .ili,'f,l ;[',:".'I;a',t#*,""flllilil;:ll:;;$Jiil'","11,1:':;li!i";"",'{:ii! lil$pl$uH:î:ll:;;d*îrli:i{+",H-f'Ei,*ii:i$i;:{ +:*l-*i5,-*ru**il"r:-i**1"*:i*]l#i$t',il'1ffii,i;:ii"1Ji;ii:$lEr"^li':^L1i'-iq::i"*.Ft':',:":ii lit;Hii;ti:?fi *- *,:"fF,:,1*#t*"#li,i,'di:"'li'Llli;'":i;l;'il'i' iiHig i3i?':;' 'r8€II0 I-{s''l9ll!{ili.. j]ii 'sJIIr Îtor uo t ::,jriffi {ifii:,siJ:"'fr::,Jsl;q";:$}l l.'66 oN'ellq,î\. J J 'e]lqÂ '[ropuels : sr]rcEril (6t6!) 69:05 plsueero troS fou 'ord l, J 'J 'ZC etlq,\\ 3 c11cel3 utnllEqaqd

,?t"#;; ot xade-eluetsolnulLu r ,{luo 8uraequt Pun arll """'.r.rq,413 'ecT.'"dt!:-ln^:l--yl^'t"t' eql 'olllsod iiJir.* rti-", 3ur:q i,, rotrut ell iubrj ecseiogul lo s3lcedsrcurr,oJ arll uIoU "ir u, pun ino.rqef ,{1;1:ldurorSuroq ,ur sls.Ulp1! 1."i,,i)"J'p,',ni,,j,1,,,1'-g-"11.:/iTl'",rii""j;1",,1i':Ho'iJl ,"|":H;iiilo* lculsrq uole.rohtr elll Jo c3re ll"G ,0,"^{5"11,;'!1,;[]';j;1"^;i.s,"x,;i.,";+ti:it.l:ls'i'::$l$s*"fJ"l\;l;$ ' euollx3 :Nollnslu-lsl'l I t dc11-pur:lsuoenf lsE€-qlnos

'arnurru 'sueur?'sQr a1'{ts'"t,.,"'i:";"1?1".f,!"1i"3t::ttlliiti eru'rts:srorq"lfl le.be'q3'q 'r'?J '(r"^o .rrr,iiiari, ?i".ijqnlB-l"to1na.,rplni.'tu- rut-tiI le-r'rrqds-qns '1Ljr-rpu uLLul'relnclqro :iatq -* E.6 uc'snorqep r1fc .{lgo tis a.rotldou{3:(i ).rllq^\,'8-uot -"!op',o"'.,,qlue tfy".l-,j:I,[S;s;Ty:jilUl1ff],fi,1i1"r#,,']i"'; :r1q1\-qsr\\oLlJ^'pJllop-pu"lb'il

'rnindli. t*fi*'il*rn*l$l$,li*lli:r"rffiro qttl^lgl ur sr3'ltoii +ffi :src.tros"eiou 0LJo "r"unb"o 0z L:i!Y]:1.e. i.,li.?,it:i.]*fifi' ;Jil'l J"l':#l';"T'i-l,T''::ff ';$d,i:1i"r'',"'tlTfi! **:l,.$ii1$i-**#:.'{rli:',1r.;"'''":;nil};.{iT$T;i,'ilr*; .s'u,rrorl (1 .c 'qE 1 erriv selpeol - :l5i"B1'.ly_;,il"1i.i#J;i*Hl:;H'?i",,':H JO N ILu I lnoq€ lJ Jo ,*"u"-s::'J';l'.11i:ii$;;:'x'l3fi "';J,;*ti;lF'u;'i'rii;l3'i'."iriilr:lfrh'id,l*lliii!'iii:rff:iil,!l{:ii:!i.::l;'}1:i:1i$i}ll"ilxri"':e :$i:*^x.],1*"l;;sl';i;;" ",,pi1ru "fiffi .;;q;is .i!iqn:ea5r" ."orri,T" .l,.Hr"fi:il,rl'",d"] : 'd 'd 'tt '^ou ds 'uosll1yD utnlloJlsnlqo high, red; ovary cylindrical ca. 1.5 mm high, glabrous, apex + truncate, carpels with the upper third sterile; style + equal to stamens,glabrous. Cocci divaricate, ca.4 mm long, rugulose, rounded at apex, outwardly short rostrats. Seedbroadly reniform, ca.3 mm long. DrsrRrBUTroN:extreme soulh-east Queensland, apparently confined to a few neighbouring mountains Map 31. QUEENSLAND:Mt. Greville,S. L. Everist7243 (BRI); rbid.,"It growsin creviccsof cliff rocksand also", in crevicesof fiat rockson the top of the mountain. It is a ihin bushfrom 2 4Jt. high E. J. Snlith,20.iv.1938 (BRl); M1. Moon " on theTeviot a maintributary to the lagoon", B. Scortechini441 (MEL). This specieswas lirst recognisedas being a distinct taxon by F. Mueller who made the following note in Melbourne Chemistand Druggist 7: 64 (Dec. 1884) (under Eriostemon coxii):-" The Rev. B. Scortechini, now zealously engagedin the phytologic exploration of Peral, gathered on Mount Moon, in Queensland, an Erio;temon allied to E. eletior, bLrt with narrower leaves, revolute at the margir'r,with mostly solitary pedicelsand abbreviated peduncles. As a speciesor variety, it should bear his name." The collection referred to by Mucllcr is Scortechinino. 441, collectedin 1883(MEL 4783).

33. Phebalium bilobum Lindl. Lindleyin Mitchell,Three Exped. 2:177 (1838) (not seen) [bilobus : 2lobed]iLindley, op. cit.ed.2.2.178 (1839);Walpers, Rep. Bo1. Syst.2:823 (1843); Bentham, Fl. Austrat.-1:340 (1863)excl. South Australian specinens cited; Engler, Pflanzenfam. 3/4:140 (1896); Ewart, Fl. Vict.708 (1931); Engler, PilanzenfaD. ed. 2. 19a:260 (1931); Curtis. Stud. Fl. Tasm.104 (1956); (1957). Willis," Vict. Nat. 73:195 Typification: We foundupon the rnountain [i.e. Mt. Wi]liamla mostremarkable species of Phebalionwith hollyJikeleaves and brightred flowers,resembling those of a Boronia. It wasrclated to P. prylicifoliun,but quite distinct." Jul] 15,1816. Holotypus:CGE (phoro seen( A\B) Mrr.nl\\rliarnno.24q ;j\o.: K (phoLo,een).\4LL. Eriostemohillebratdii F. Mue]I.,Trans. Phil Soc.vict. l tl0 tl855),r,otn. ilIci.lp. bilob m Lindl citedas a synonyml;F. Muell.,Journ. Bot. Ke$ Misc.8:17 (t 856); F.-Muell., pl. Col.Vict. 1:127 (1862); F. Muell.,Fragm. 9:109 (1875); Rodway, Tasm. Fl. 24(1903). P, hilebrandii(F. Muell.)Engler, op. cit. ed.2. 19a:260(1931) pro syn.s:ub p. bitobum. E. hillebrandii B. q'lgtfoltlts F. Muell., Trans. Phil. Soc. Vrct. l:10 (1855.) "var. ". ' F. Mue ., Journ. Bot. Kew.Misc. q:37 (1856) longilolius T)prfication: On rhe rocky summit ol Mount William, 5,000 feet above the level of the sea". Syntypest MEL 4608 and 4616 " Anthers yellow (pale), Nov. 53 "; MEL 461'7. P. truncatutllHooket l, Fl. Tasm.1:64 tab. 9; (24.x. 1855);Hooker f., op. cit. 2:158(1860)i " i.Ha6. C Muellerin Walpers,Ann. Bot. Syst.7:522 l.1868). Tlpificatron: lCunn, 1947)', FlindersIsland, in Bass'Streits, and Schouren [s]and. East Cor\r, Gunn." Syntypus:Flinders Island, R. C. Gunn 1947(HO). E. serrulatusF. Mu€U.,]iagln. 1:4 (1858);Hooker f., Ft. Tasm.2:358 (1859); F.v.Mueller, Pl. Col.Vict. 1:128(1862);C. walpers, "In Muellerin Ann. Bot.Slsr. 7:521 (1868). Typification: vallibus silvaticisad originemflu\|l BunipBunip Creek." Holorypus: N.EL 4620 iso.: K (photo seen),MEL 4622p.pte.

Small shrub ca. I n.r high (to small tree 4 m high). Branchlets slender, terete or slightly angular when youlg, smooth, minutely or coarsely stellate. Leaves rvith a short erect petiole, lamina spreading, chartaceous, shorily ovate- oblong and constdcted below apex (obpandurate) (in the Grampians and Tasmania) or narrorvly elliptic-oblong or obiong (in Gippsland), 12 x 4- 20 x 8 55 x 10 mm, flat, or with slightly recurved rnargins lClarnpians), slightly to strongly serrate, trulcate to strongly retuse, ninutely glandular. pul]ctate, sparscly stellate to glabrous, midrib slightly impressed above, promineirt below. Inflorescence a terminal compact cyme with the flow€rs in the axits of small terminal bracts or in short pedunculate clusters; pedicel slender, 4 8 mnr long, red (Grarnpiars), nlinutely stellate, minutely bibracteolate in lower half. Matuie bud narrow-obovoid, ca. 4 mm lolg, red (Grarnpians) or Dale srecn to white (Gippsland and Tasmania). Calyi sparsely siellate, lobes leltoici, 0.5 108 601 'taqLun ecuecsoJou (9t) (7),eqr Jo Jur^c € ur lEururl;} ,{tTensnsje,,!\ou OI-t 'rn(.,,no'.lqnrt ol snolqclE qloouJS -u; snolqelS eJlllns le,\\ol '(latnutu Pue '{lturnbaLlur8rELu'rsnlrr 'ro prqolrq "";rrn. r"iJ;'arrtie'1tup ui,1^r paijnj:L 'Luur x z x,-g^(s^2.x €) (r€ln" ;;;i";; ;;;i;u;;t1- pt* olnctinil xrdB z Lr s 'snoJJellsqJ lo aleto ol :lttouncqo-JlEPloJIllJotls -roto-qns{lJrEt) duolqo :ate11:is{losteds pttr-' F1'nuttu ;;i;;l ;"1;;"r;r":iJ"-1o,t"arp'Ju''t"int1 l"]ir"r"-,"p,i"t.sralqruer5i qarq ui 99 0€l€ruuarrd ^0""" -'"0'".1:,,:;;;t, ',.^ 'r]; ';,i'*w:f{# i#'JT{l'1pJ;'11Hffi:lgff?;ffi :i {ii#';l" ;ili}:'"'iff :iy?:K 'tJ"ilt1"i9t;1ilt"";+"[":Xw;l: 'rerlsnvs rc ')cers: o1d oId trrill oor't(to,"u"'l'f$o ;3 '(0681)iZ Prlsnv S I}eIxE 'llenl{ 4llpuuqaulAA 'lJ qpueH 'elel : Selll ruou'eld .ooe'ord (298I). LZI:I 1'l'{IoJl'I "111?,,'J'3Jr /6:L;ttsqJqi' l?T;JfjlIS i.: i3$ll$ ^',j3{ jft ii,1t!'oil"'1"]fi';"ii"l!lf"^il rd'i"";q {;';'-;q'' "d";,;t';1"$lfi ;: ;;:.;lt':",fi#;:i,:ilid'ltl;i,,;;iuv,;19,qt;ll"^'*"'i'*i1 ''*""'*"' 'lpserqellH luleqltiA r( rel]e petrlBul(196I) 961:[r'lBN ]f,14'sIllIlA H f 't 'tt slll,^N H lpuslqa[lq unlpqoqd

3ql uo,lu-Jpuedap'41tled '^rErJlEltoJastie s{erllu sJlJrurUJql Jf,uts'rol)eJJJEds i,n.'u"o"isoqi u'^salnno1o ug1111o-o eqrr€ql ;i:;,;i;it"d;'ii"antr Juo a-re.{:qt-te.frotr'{rellodlel sreaddtl:.ro.J:raqt ll lJqlo"qr "t8qn'x1"ia'1i PJlenlrs 'qcrrl€lloJ) i.,n salnno:qt {rellodreclquaq16 -rrpenbrr ,p,, nq "pit f"'e1ti ui,lra,Jr Lailcunirjqi or Buip.ro'eos:'err ffi:;"ii 1o Jiil''3r1""Jil'^3lll^",.,

:ortq* *j""T""d"",1i$,t",ft ;:iJ"',Hii"*';":,1T:,i,.i"X"ji1;'"1#,,'"TiJ[,"lJt"*il'il""-iii'f-torj u""i "ieti'1'suji"c:ds"aq1 "q "l tl.n:19:,(::: ur'to-1.suerduler9eql ^\{ etll ttrolJI -'no1u"fnq-i3"og !qr.'o't:t^oq urnteqJJtt ul-1J^l!tl$JJe spnq spuJt JoEq\-J Eal t1ilrunto;un'tt d] PruEtusel pr€\\ol JJour 'e1e'ies-J- 'raauolq5.'ir ere se^!e1- lsnvrltas ;;;;""qr t; 1eg rpiio'rls -allqrlnl sa^€ol ur ispnq re"aogqsrpper pue s1 frunliatitg 'uotls .t"F,li su€rdtuBrD-P::j::: Jq L punoJ Jl€rJJsxLlrldrl('pcoJq tY,11s'sin4oitq Jl luPld ,l "jrj rjiiii;rpilnJioloqitro* i, uonnqurslp;ooseer"r,i:j"&r""iltp";tJ:r:qr- 'ezls 'se^tel el"ljes sl[ r3tsr"l runlncrde ecuesqE'sleqlue '\\ollo'{ uo ldnlq€ Jo r)qunu pe'np?J s1 ^q sercadsl€ql ruo.IJ p"qtinaultiil eq xeu 11 td ser'[ wnqollq.Fdr€c d n ,n.1 osie qclltt ttptnJqallt1 tillnn s"ltluse luepr^r '(.$SN)8S61 3nV s33ugD:g :i:l!'JE?'i'i#?i",T::"N;gJ,,wiJi3r,3?.'"Js:idi#,;"'ii,l'"#;fr"[x"']f:"!J':iI (oH)^e..^povl,'ior*i}ti f"',9] x u"8I[Iy{ I"que "g] ff,trSTt).f i?iF,',::ffi}.,,X'R ftSl ZZ'619'€t "19t,y1,f::^ls€ot lsee eql uo d"I {-ll3rls sseg-put'suerdurers aql ul Tu'l'l "ru',us'r:i,hn1sdd'e rs",iur *t "lJL"l"jf r:"J'"1i.'j;",'"c .Buol q11,t,pue xedz-1?-pe-tor1-e-1.'8uot uluI s lulu I'rcco3 €c urnJlsor IEuruuol," 'uot1s,u '\l) 'alzrire,ttp xede olu3ls 'lunlci ot,"! q,t1oi1:1..1""r" t-( urtu 6 I-I .iloln" :(ntin"i*r pue put1sdd19.1rii:-ri r

Pheb(tliumhillebrandii rs closely related to p. bilobun (\.\ith which it shares a lednced carpel.nrrmber). P. bror hypltyllun- p. lamprnphl:llunt, and p. t.otundi_ Jottunt. tt lna) be dtsttngUjshedlfotn P. bilobum.witlt ulriclt il uls at onerrme united, by the pink anlhers and the leaf shape, and from other related species also by the reducedcarpel number. - The Kangrroo lslrnd fornr {of \rhich I hare seenonly one specimenin flower) has small, broadly orbicll.r,r leaves and hls lol-apiculatc iarpels. tt p. may be more closelyrelated to hrochyphl.llnrbut I have not included it in that speciesas it has a reducedcarpel number and retusc leaves,characters found ilr P. hillebrqndii.

35. Phebalium brachyphyllurn Benth. Bentllan,FI. Austral.1 1341 (1863) lbrachy^s - short,phyllon : ledf];Black, t-t. S. Austral. 144(la:4)r J. M. Btack,rnns Ro1.soc. s. eustrat.si:l+o BlackFl. (l9481 d93t;;j;i:l)i'(1e31);J. M. S.Austral " ed. 2.499 Typification: EncounterBay and nearCoftin Bc,v,,F. Mueller.... S),Dtypes:K .,Spencer,s and St. Vincentsculf on the low coasrranges..- lptoLo .."n1, UEi ,45iA.i ln montibus hunlilibuscalcafiis DroDe Comn-ba\, febr. 52;, 1eg. t. WittretmianJi eniounrer-bay. Sepr.27" (?) leg. H. Bihrt. Sih. tiflj:ri#':t't:?:::ii!.! (Benthan)rate' I{andb Fl Extratr's Austral 24 (r8e0),without

E.^nticrophyllus_F. Muell. lnon p. nicrop!1l\ry Turcz.(]852)1, Trans. phil. (1855);F. Mue Soc.Vict. l:99 ., J. Bor.Kew Misc.8:37(185b); r. vueti.,i,t. -oi. Vi"i."riizS F. MLrell.,Ffagnr.9:109 (1875 ). trSOz); tou coasr|anges of Spencer,sancl sr. vincenr,sculf.,, i::'f."t,ti:i,: ;;?,i,,:!e syntypesas

Snall undershrub. Branchletsterete, smooth, mirutely stellate-llrrr (rarely glabrous). Leares.preading. rery shortll periolate-coriacedul or whcn ory wlln recur\cd or Incurved margi[s, minutely stellate_scabridulousor gt-1b..r.1:1.:l]ir,":"larc to \cry,broildlyovate or sub_coittate, q xZ'i il 3 x 4 mm, afex oDruse lo rouncled. l1rllorescencaa small terminal cluster with the flowers.axillary to 1he terminal leavesor bracts,o, ,o,"aii-ar-i_j on a short i.."_l-!2,4 mrnlong._mi,nurety bibracreolare: ,nrrure bud btunrtl ooovoto.n1li,]:]., 9.9 ] J 4 rnm tolg. Caly\ tlesh).glabrous. lobes delroid, ca.0.5 mm c .4,nrmlong. minurety gtrlldrrlur. r,,hire ringcd wlrn:3,1$,.,T1",': ptn Illr?y-..,b9)119 K In Dud.Itl\ sttgh y lnfle\cd: slitInens equal to petal.;hla ment. slend-er_ rerere:anlner cordate-orbtcular.cr, 0.9 tnm lorrg.pinl,: gynophoreca. I mm suD-sphccal to olord. ca..l_ mm high. glcbrous:carpel uirh Iq"j 9].1{ ( a ape\. occj spreading.c". 4 mm high. apcr ::::,_,1,"11:,.irounceo _l':lll: ..tishrt) and tplcltltte on olrtcrangle.,.Seed thicl. reniform.ca.3 im long. u,.,or.o,o#l"o,"nrH;i"t,?.rq,H^T,|;f 'euo1sl,(aulo{ eutlcodol." ,.r*..: are11 -ar '4 srql ur pslltutl?p sY "iqflrrn itirrrirr"" ii un11'{4t1-otthuol nded :$14.]r$s6lrr ii,'"3Jrlg*;**s3'H),'"igJi:fr #t.:iJ#,!.s,T':"#r';:i1.W"'r'(,ÂSN) 'Jo'{ld '1 'a8o!X ElloqEprrr8: tt6l fEI l J,'J V '(-iar^,r)/r.8rê$:19:191{.,iilfq;i,,;,t:I*i";f,'!fr]":r,6,1,;i;i;H". i#:f ;t,i.i:1y1*;f,t'ij'i:Ail.Jli.?fi'ili";l l;ru*$ ;l;fi*t i:,i.! surprlrc aq16uo1e qlrou errorcirt u-tels'ei3b sureltrnour"1r;:t:::t"::; : :i,1,';"1:ii3$'$,Xl',t"'"^';?.ilTflrj. t'.'^,f"r:iiiiy:i"i:ft.i#:!',']:i ':' ;l ;;'X jlln ".,1+ti;lI f ;Pll Jilt;;fi :-,::$l :;.i1"';' i1J^;"ii,'';;;,{;,' i,g!l'"?ii,iJf:l:U"';"":l*,[';ilu"llil,il;Ji$.:ll: ";tt'it ;i.?;,ifFlifi iil;i"ll$#il*jj':f -:frlljjilu :lt*,:tur :l?:i,^r# 9*rt ?ii;!-"J,l"Jti;;:1;;-oiû'ir,e,*og'a.,nrr,'.rrrq re'trol

,pmt*nt':::*;r""{"'',.::','+r1i+'"-"lpi---$ii;ffi\ 0,,l5u,n J :HiHlJ;;it'a1l,noqo ;*i n:l*ri,?''fj il-'i:il:,^:tr'snoeJE rT]i,li! i" €ulul€l :;;;;;;. ,tlpebrq or crldr[o EtlJ o] snoaJ"rrorqns ;:'l;il#d^,r;;ii; ii;:,*::i. g S:"T.j,._" ur) .lo'sJlJuJJJnJJp-JEJI Sno:lq^El^ l:l#"";"1:1lli""J;ili,* Ji"y:. [ll,luint*l:,!r*l"tlsUllil,'"di'J,i,"I 1'11"il:i,l;3;;;ii"* 'n r'tn'of{i}^:#11+ii reuosrede uo pes"q rrq€unsorg-:.-r-:s,p r"no11" {"f"1 l.{t,;i;*,,t,,':'ltn'fulii"iri*ii""!*.q,";f8ig'ft:,1fl1:l,lii'rti:i;;*i:..^i :.;i,;fi;;D'lr;iiJrJ,.ittv.n!qrro"''oiunourauro'loslLuLUnst"Y,niliiSo*,"^.n,0 .Tli9;ll":J ei:it'Jj,;: i ''lqsiiq-lroiuLuell i,["i,"];\"r'^t6E8|)€t:z l)la J( i,,""-iiili{r,nil,l#;l )r:riif; '1314 'lrB^\g 'ruetl}ueg '(lt6l) 80r 1l :(t981)0t€: I l€rlsnv l{ '9€ 'q1uog ('1en61 q) urnll'{qdordurElurnlpq?t{d

eJou,"11111,",1.,:l-:1.."*.r::l.l$j;Ti$,'mu{,h:i,:';$$iiTf'.ffJi uJo.tJLunllILFo {dLurl d Jo luloJ l#i'.;is'$ii:Ti}.iiiililitt fflf::i"'".1,T-s,t#i{:{:;!#bl,' iJ; '{Lrsotr sr rr'?,,//.t{d"l"t"^jr|., sr"g,p t', git'.lit woil ttpuotqallttl J o] pel"lrr o*, ;i#tty*ffl;f :xtfrtii;+1"#t ;i." l {$,1i ;Ti{jf **isiil'il#llHriii'€Irolcr6 'sPr?'t '7E de1ntr-tsa.t re1 '€ll€J}snv qlnos :NoIrnsIxJslO -ls"a €lnsurued e;,4E uorS uolSar ureqlnos In the type (a fruiting specirneD)thc steln is prominently verrucoseand pubescentin lines,the leavesare tbin. elliptic ald aiute, rnd the fruit strongly rostrate. In southernNew South Wales the stem characteris similar but t'iri leavesare tlticker and obovate,and the fruit only minutelv arriculate. In the form lrom Lithgow 10 Rylstoneand district the stem is st;llaie all round. and the leaves sub-orbicular; the fruii is not kl]own to me but from the ovarv morphology it is probably similar to that from soulhern New South Waled. The Rylstone form closely resemblesP. brachyphyllum, but differs from that species,and. agrees with the lype_ofP. lamVroThyllun,in haring a verrucose stem.:mooth.glabrous leares. lnd a_shorrgynophore. Owirrgro rhisapparent cline I am retailing theselorms under the one speciesand hare not atGinpted to .create arbilrary infraspecific eltitjes. The rnorphologicrl approacf to P. brachyphyllwnin the northern forn is, I consider.onlv iuoerlicial.

37. Phebaliumrotundifolium (A. Cunn. ex Endl.) Benth. Benthafl,Fl. Austral.1r341 (1863): Bailey, eucensld. Fl. l:193(t899); Cray, Contr. N.S. WalesNat. Heib. l:44 al96l) Etiostemonrotuntlifoliu.s A. Cunn.ex Endl. in Endl-er al., Enum. pl. Hueget 15 (t817) l'.otuldu:,, .r9ql9,foll! ry_ ^^ _ls"ll i Watpers,Rep. Bor. Syst. l:504(r842); vcioreet eltche, Handb.I L N.5.wJle\4l (1891);Di\on.Pl. N 5.$Jlc)18 rl90b'. T)pificrtion: Hurrtef. Ri\ e r lA. Cunninghdm,1827)." Holotypus: K,.,A. Cunninghamno. 1827. MountDangar, Hunrils River, New Souti'Wales',,ipt,oto seeil; iso.f: MELfJ,_l.lqg.J 21954.

,, Densc bushy _shrubto 2 m high. Branchietsterete, smooth, minutely slellrte. Lea\es shortly_petiolate, erect al]d solncwhatinrbricate (at least iir or'1eostate), tamtna sub-corlaceous,broadly obovate to orbicular, 6 x 6 l0,r 6 mm, ^er)tireor minutely erose towards the rounded apex, smooth, graorous. Innorescencea termlnal compact, globular raceme, the flowers usuallysolitary- in the uppermostleaves or bracts-;pedicel short, Iieshytowards \llmnrir.ca. 1.5 (r'arel1 mnr.long .witha peduncle3 mm longand I t3t-pedicelsl. n]|nulelyand spcrrel)slella(e. wrlh I\\o sub-basalcaducou. bracteolcs; rnature flor.terbud blurrrlSobovoid. 4 5 mm long. Callr obcorrical.pas.ing inro fleshypedicel irpex- l-obes brorJly delroid.ia.0.5'nrm lriglr.glabrous; petars narrowlyellipric. 4.5.b mm long. uhire to pale-ycllou.glarrdular punctare, trp rnilexed;stamen^s shortly exceedingpetals, filaments slender_terete; anthers brord oblor)9.(l) 2 mm long._5ellow:glnophore absenr: orarl cllindrical, ca. r. mm Itgu. gtablous-c:lrpejs ur l aprcali/2.2 J sre|ilc:sryle slender equll to slcmens...Cocci^sprelding uherr rnalure.4 6 mnr high.shortll ros- (rate. Seedbroldly renilormca. J.3 mm long. DISTRIBUTTON: extreme[o h-eastNew South Wales and South-easteueens_ land Map 32. QUEENSLAND:BailandeaD, dense growing shr ub to 2m high, on hjlls amongst granjte boulders, c. T. wnib-92125_(B_Rt);nr. Stanlhorpel r. M. whajie .i0j w;iiii,iiii", r. L. eo;il man,July l9l2 (BRI). irswj; NEw SourHWALIS: "in Torrington,. crevicesof acidgranrte orturop.', J. B. Williarns, 9::i.i.f::1-,1Dit Howc . rargeburhy ptanr.-or.l_o ri. itr, i"a .i rii";ili widrh.usuatrr gro$rngIn crumps . J. L. tsoorman.Aug. lc)05 (NS\ ) Tlre type of ,E/roJtemonrotundifoliunr is recorded as having been collected [ear.the HuDter River lnorth easternNew_south 'IS Wales). I haie not seelrany s_pecrmensot spectestromthatarea. Howevermaterialcollected from near the New South Wales-Queenslandborder compares well with the holotype ai Kew.(phoroseen). wilh a pos\ibleisotype in Velbourne,rnd also wiih rhe descrrplron_gi\enby Benthamwhich was ba:ed on the rype. All marerial sludied,differs frorn rhe original descriprionin having iiellare_puberulou. Drancnlet\(nol gtabrous).,b!t lhts.charccter_ma; hrte becn oterlookedby tndlrcher. lt ls possiblethat Cunninghamcollected the plant near the Darling €II 'rulod 'r"q.rya,6i"ii':it91t]:{lliir?:d{i 0r-8lu"rd .{rre^o

..ls4npt otolfisro4) lln{ uotssedsnorJsnl (;eso1c{-t:r qcrqtrilnopo,{trnr1 E lrLuJ^lrpEJJ alll lo leql sJLquesJJ ln3rq8rlep"cl 'sje6u,r eill u3:,n1:qSurqsn:r uo so^"el oqJ,,:I?ql selou ploJued')I V 's",1:"dorci yn.,1"1perirelqrlnp,r e,neqo1 ri11an1r1dq lqSnoq] lsru lB s€'lr 1l rtllue u"ql -.re.ldt:uotna.r liiq.t ro3-rnopo FuqsrP B r^ELl sa^Erl s11 lq8t:q srql JIsq ssel 1sa33nssrolcollor .rerllo lnq l3 97 Sutqreer se lr sploJJl 9881 oI ueljane"g '1rq-eq strJJds sIqJ leprue.rdd pue oderlsJ?ol sll ol 8ul,\\o snonJldstloJ sI

'laloc'ueluroog pplujeJ.{d 'q;lq 1 f ..!Iq"q uI Pu?l.eclluoJ 'ueLtuoog }J 8-9.rt "u"1d(ltJ?'ffi oBlp'i.. uttr ieoi reansJo trLuuns:(A\S\r 600l rsl,{ l- f .' rnopolJsllqlp irri'nlo oi'.]rkoo trarq:iirtrioqe'rrLuLur.. tr:i:qtuo {;uopunol.. uoxrllrNru ttlt\ ep'{lJ iuip:i5eg Hl ros rr{\ :(n Sli ;8Sl re6 \A poo^\prergru uln JPoI rEBnS:sr'lvl '62 d"tr l-Poo^ prPlg JEauulElunotu JlSurs " Jo lrulutns eql 01 pelrull f,lluarEdd? re^rt oP,{lc :sel"ld qlnos ,\\aN :Nolilnalul-slo 'eleJlsol '3uo1ruru y ol'uj.IoJtuel ,{lpJE,\\lnoxede'q8rq ruru9'ec'elu8nr 'e1ue1sP€S 'Mol"u -roc 'icare tcco3 pu€ l"cupurldc ,41,torwu jler1 Jeddn lou 'jioln8nr 'qalq 'LuroJl?lop 'rcar: sr:drer :sno.rqi18 ,41e1nulu tuu Z "c fl€ô :uBrututu c.n er'tc5rjpurt(r-lJoqsJsrp:8uo1tulu 9 1 18uo;qo^lpEolq sJJqluE 'trriuelu ilere..luEql rJBuol .{ltqEr1s ot relroq' trllqBIlssurLuEls ,ir"rlr.,epi,"is 'p3llop :o:x:uur x:du ettq,t t-tstnolla,{ol ^LuPJIJ puelH'buolLu ul E eJ JIEAo 'iriorqeia 'rcq} ,{i,,,rorieus1n1:d qlirq urul 8.0 €5 plotlep .{lpeorq saqol xIl€J 'iade paprrno.rqtrn 8uo1ru t-u s l:J'pro\oqopllq e.tnlE!u-:rsEq.JqlspiE'Mol atelo '8uoi -"1"e.rqt.irye1t.r.ti.it rur.ui tc lijrped :snolnrrqnd {psreds ro snorqelS pue paueueu ,{l8uor}s seqJuclq Jql sle,\\ou 1o qur,{roc lP,ultlllel " ecussse]ou 'JlnJ" 0f-0I',$oleq iu1 sl"oJnc ^l'ttou€u JSsq xad" luouluroJd pue eloq€ ueluns 'xode 'luc q'ip,ur ':noqe ^sso[8'sno]qel3 3ql sple,ltol al"F.u3^s.'qloouts I x / ji.î 'crldr 'snorqP[3'r€lnpuBl3,(lJlnunu x S.S 11e,{ymo:reu snostEt.ruqr srêol 'rerriaui ifaiojiS sjaplcut:g q3lll ur (49)f I qnrqs IEprurerAdloerE 't8l0r ou uelrenees-'6t9t-13I :snd{loloH uelroneE8,\A ,4dSN: osl :66I 'eprp I .! : eoseqt .to 1e,161eqj e,loqaleeJ 969'9 lnole eq1.Jo serrnos eql lv,' :uoIlD'LJrcl'(I '(.LZ6I)L7 qel 0i€-l€t:09selPn S N .oS U I'pioJued":(906[)6t selP^dS N ld 'uorr6 :[f,€up{S.Jo.xoJ rC rrlJe :rg6[1ify s:te^\S N -F qpuPHrqJ]r8 le Jroo^ "llrn'\ ) f p:tueu11-9_331 JJCI l7a:LlsrSSnrC pue l\rursqJuersPlerlsnV I r/rn' ltoa)tro!t:l 'rqrlog '(9T61)9lT sluEldsJlPy' S N snsueJ to uepJel '8t oqcleg lJ urprPt^l ( llrnl l) llxoJ urnll8qaqd

'ecuocsolou 'edr:qs erll ur puE 'JJoqdouIS eJ-uJ-sq? led.rtc 'd Jo 'd -ur lreduol':deqs 3ea1eql ut unll[tldolLltuDl pue wnll(iqdA|lptq LUotJ 'ura1s v wnljtqdotdwnl luo.IJ sraJlp setcadssq1 prie qlooLus e-Buirerl d 'prrJncco 'Je^Q{ (ed,{1 selMsco[ Jo uolsnJuoJ ]Eq] pue lalung oql J€eu s€,tr eq ueq,tr rrll uo ur^r8 Jtep rqt) /z8I lsn8ny ut paurnler lsn! P€q aq gcnl,& urorJ su'^doo Shrub to 2.5 m high. Brarchlets terete or slightly angular, smooth, glabrous, leaves sessile, chartaceous, amplexicaul, broadly oblong-elliptic, 6 x 2.5-10 x 4 cm, flat, smooth, glandular punctate, glabrous, apex obtuse to acute, base with lounded lobes margin serrulate, midrib prominent below. Inflorescencea terminal compound cyme,sometimes glaucous, 20-200 or more flowered; pedicel angular, ca. 2 mr1] long, minutely bibracteolate towards the base; bract ca. 1 mm lorrg, caducous;malure flower bud broadly ellipsoidal, ca. 3.5 mm long, apexrounded. Calyx fleshysmooth, glabrous, lobes broadly deltoid, ca.0.5 mm long, minutely ciliolate; petals narrowly obovate, ca. 4.5 mm long, white, tip irflexed; stamens+ equalto petals;filaments slender- terete; anthers oblong-cordate, 1 5-2 mm long; disc short (ca. 0 2 nrm high), narower than ovary; ovary subspherical,ca. I .2 mm high, smooth, glabrous; carpelserect, the upper third stcrile. Cocci + erect,ca. 3 mm high, rounded at apex and very shortly rostrate on outel angle. Seedreniform ca. 2.5 mm long, smooth, black.

DISTRIBUTToN:extreme lorth-east New South Wales al]d adiacent area in Queensland. Apparently confined to mountainouscountry-Map 15. QUEENSLAND:Mt. Normannr. Wailanga.ra,ca. 4,500 ft., M. S. Clemens,10.xi.1944 (BRI): BaldMt., 4,100ft. bushyshrub 4 fr., E. F. Constable9.v.1961 (NSW); Wyberba,Mrs N. Gunn,SeDt I932 (BRI). NEw SourHWALES I Bismuth n.. Deepwater,A. McNutt, Aug. l9l3 (NSW). This is the only speciesof Phebaliumwith an amplexicaulleaf. It has a restdcted area of dislribution and within it shorvspractically no morphological variation.

40. Phebalium nudum Hook. Hooker,Ic. Pl. 6: tab.568 (1843) [nudus : bare];Walpers, Rep. Bot. Syst. 2:823 (1843): Hookerf., Fl. Nov.Zel. 1:44 (1852); Hooker f., Handb.N. Zeal.Fl. 39 (1864);Kirk, Stud. Fl.N. Zeal.85 (1899); Cheeseman, Man. N- Zeal.Fl.93 (1906);Cheesernan. lll. Fl. N. Zeal. l: (1914); (1931); tab.26 "NewEneler, Nat. Pllanzenfam. 19a:260 Allan,Fl. N. Zeal.l:424(t96tj. Typification: Zealand;Owae, on the eastcoast of the norlhernIsland, nr. Colenso. 1838(n.56). Hokianga,Edgerley." Syntype: K, Colenso56 (photoseen). Accor.ding to Allan, loc. cit., the Colensospecimen came from Owainear Whangaroa, North lsland. Eriostemonu(hlt (.Hook.) F. Mucll.,Fragn- t:181 (1859); F. Muell.,Pl- Inclig. Col. Vict. l:12'1(1862). Shrub or small tree, branchlets slightly angular when young, minutely stellate-puberulous.Leaves with flattened petiole ca. 2 mm long; lamina solnewhat coriaceous, oblong-elliptic, 15 x 5 25 x 6 rnm, flat, glabrous or glabrescent,smooth and without a prominent midrib, glandulal punctate, margin minutely crenulate, apex rounded, base broadly cuneate. Iufloresceuce a terminal cyme of 12-50 (or nore) flowers, the branchesminutely stellate- puberulous; flower-bracts narrowly ovate, l-2 mm long, caducor.rs,pedicels slender, 3-10 mm long, minutely bibracteolate;mature flower bud narrow- obovoid, ca. 4.5 mm long, apex obtuse. Calyx somewhat fleshy, minutely stellate-puberulous,lobes deltoid, ca. 0.5 mm 1ong,ciliolate; pctals oblong- elliptic, ca. 5 mm long, white, tip inflexed; antipetalousstamens equal to corolla, antisepalousto ll'times as long; anthers suborbicular-cordate,ca. 0.7 mm long; gynophore shortly cylindrical, ca.0.4 mm high; ovary sub- spherical,ca. I mm high, smooth, glabrous;carpels without an obvioussterile apex. Fruit and seed not seen. DlsrRlBUrIoN: New Zealand, nofih-west region of North lsland (according to Allan, loc. cit., fron lat. 35" 10 37" 30', in forests,especially marginal). NEwZEALAND, NoRrE ISLAND: Kiwiriki, T. Kirk 83(AD); Little BarrierIsland. Miss Shake- sp94!(AD); Huia, Auckland,alt. 900ft., K. Wood 28480(NSW); Marakan Hlrbour, anon. (NSW). 9I I lceduoc g ecuecselouul l\olaq lueuluroJd pu" edoq? pessetdurr sr qctq,t qupru 8uol" srt?q ele[els,roJ " JoJ ldeJro snorq?l8'rllooLus'el?lspor]p oql 'ut ur (elnloê.r.Io)pe,rrncer'xede peqo|q ,{1lq8t1s:o3 ldecxa eJllue 'snooc"tr"qc8.0 'ellsses 9.0 x g-9. 'eseq o] pa^\oJJeu,{ll"np"J8 pu€ et"aoqo ^ld\ouEu Z 'rJtoours 'l€ln8u" 'qnlrls se,reel snorqelS d13uo:1s slolqcutrg II€IUS (uoosotoqd) )'9t6t'Igt{ : osl:!t6t'13ti,t :snd,{}oloH ,, snsr6Îeg ploJo^\lllrnors reÎU €IE,{oA ueurnge}rnfslcpluer8 snQIdN uL,;;::E^Xlgfrt 'Id 'uoxrc qpuBH'orp}og oroolN:G88I) q"l :Z do :(!68I) tt sot€I S N lC 1o 9I r'llenhlllc 'ITenII'J:(/881)0tt:I-Idtrr,\']s,(S,{e)'llent{.I:(t/81)t/:l rcIA ld ttN J :luolsleu' o rnorroqrn peLuEu](098I)]I q"] I0I:Z u3lrll'lentr\L ! lluolslDI uoua$oLLq 'GI6I) 9II ld sole/AS N snsuoJ'eqclog}i uaprElr'{:(t98I) 6t€:I lD4snY[i'LuEqluog 'q1ueg 'Zt ( 11en11g) 1go1s1urun.rleqoqd

',{1rFco1euo aql (Uo{ '0681-/88I poued erll Suunp palceiloc sB,\\ ueos o^.?q I selcedssql Jo IeIJel"tlI IIV 'wnto1{tplttt 'J esorll selquesa: .(3o1oqd:orupue .rnoloc rlaq} uI tl:}oqsueu"ls Jo 'f,reao 'selqulosoj pu? sl€lod arIJ 'sJetc?r€r{c Ja,rou slr ur oql JoJ ldocxa 'd 'J osl€ 1r qcrq,\\ wnausolp 01 um]ueunpul l€lllurs E srlll llsJaalfi,uD) '62 Osl l-?ArlrOW r€e(I €el€ llurus ? ol pepulser ,{fuar?dd? 'putrsrp Je^rU op^ll 'soleid qlnos ,{\eN :NoIrnsNrsIO

-€lsnrcdlurql'^ssol; € ,{q pere,toceqde.r :3uo1 utu.t t o} )cltl} o*,.,i{ll;:;i.i 'el8uB 'lru-Itllns 'esoonJ.Ie,\ lelno l€ elnu-roJrq .{l}roqs 1" pepunoJ J3lnputl3 'q3rq 'BJ 'tcora Luur rJcoJ sueu"ls u"{} Jouotls al,{ls :[oI od le3rdu 'esnler S.g 'oluals 'qlooujs 'snolq€I8 ,{11q8rls IFUIS " rltl\\ slJdl€f,'papunot xede 'q8rq urur ec '1ucupur1,{c-poqs,{l€^.o :sledJ€cuoo,uleq pe,norrnJ ,41}ueururord ',{.r"ao 1 '8uol '?3 'Suolqo 01 qlpr,r ur l?nbe + tutu 9 0 "J csrp:3uol tulu 8.I -aleproc ,{1,ror.reusreq:}uB :snojq?l8 'per 'esotueru€lg stueru?lu :slsted lrep 'r"eull 'pexegur oql se Suol s€ ecr,ul + so:ru€ls :1ueurluoJd quplul ,{11q8t1s 'ueeJ8-qst,{o11e,{'.tt1uput13 'el€crldnpuoc xedr 'eso1rd ,{losJ€ds lerl,t\eulos '3uol urur g.g 'ec 'cr1dr11o-Suolqo,{1,\\olleu sle1edlseqol reln8ueul dl,lrouuu 'el?^lE^ '8uol o.lur pepr^rp sprlql o,{\l reddn 'eplslno esold ^lesr?ds '3uot 'l?oruocqo 'urqt 'osnlqo 'lEplosdlTle rlltu g rJ xf,p3 Suol ruur 8 et pnq le,{\ou '3uo1 'seloelcBJq 'esolld ojnleor :xed€ eq1 ;r uru.I g "t r"eulT Jo JI"d t rlll\\ 'esold '3uol ,{1:sreds uru 9-6 lecrped :sp?rq to s3^eel l€utluJel erll Jo sllxll eql ur sre,rou ':lu"}nu 'l€urlu]el ecuecserouul elnlo^eJ ol pe^lncaJ 'e]"p.roc0I t Jo 'asnlqo 'asolrd ur8r?ur ,{prlSrls Jo elecunJl + oseq xodu ,{losreds 'relrpurl3 'qloorus 'eJque '-- 'J"ln8ueul g-S t x OI ,(lpeo.Iq 01 fl,t\o]r"u 'snoecEuoc '€c 'peuell"U'lceJe EurruBI :3uol luur I eloled :3]"cu"i.Ip se^"e'I 'srrcg 'aleral eldurrs qI^\ osolrd stalrlcu"rg q8lq trl I ()1 qurls Iteus '.,9 '068I '€Jnrol (uoasoloqd) X uolrane{lu€ttllld ounf ,, 0t9t :6t9t.IgI,'t : osr :t9S uelrenegrrle!l[,,\A '/88t ,{lnl 'e,{ruow,, tAII :sod.{lu.{s '5p^l) 8[9t'IAW :snd,{}o}coI ., uelranaEglA'puel}seoJ,tpues uo eqtreeu'€,{urolN lV ,, :uotetgrdlJ qpueH'eq3}es q;lin"o.utlfri:t$;!"\i.Jf;ii;X!"r;T"utgl'lit serer{'sreqlnrreJs N rc }eerooru :Iso*^ ' 't f Jo rnouoqsr peueul (068I)9t:l leN lct^ ']anl tlsL2qti.t.tD.uowatso!.tg 'o16l) 'oqrleg 9II Id solelAs N snsue3 te urPreN 'c) 'It eqcle{ ]e ueprel J ulonw llsraqlurJ lunll8qaqd

'eruetusEJ '"rT?JlsnV 'euolcr^ 'selelA puB ql[os ls?e-qlnos qlnos,reN uJels"a'pu€lsuoen| lsee-q1noseueJlxe eqt ur punoJ osr^\retllosr 'DwaaoDT uorlces sIlJ lces arll raqurorll € sI lI salcadsMnlpqaqd Jo l?ard,{l 'sluerus"J Jeqlo ,{uB rl1r,r dlrugt 3uor1s ou s,{oqs lr qinoqllv Jo "Il€Jl -sny Jo aprslno punoJ snuaS srql go serceds,(luo eqt sI unpnu wnlpqaqd terminal cyme of 4-7 flowers on a recurved peduncle, glabrous or glabrescent; bracts caducous (not seen): pedicel fleshy, ca. 3 mm long, with two linear bracteoles ca, I nm long towards the base; mature flower bud cylindrical. Calyx hemispherical, fleshy, glabrous, lobes deltoid, ca. I mm long; petals oblong-ellipticca. 8 mm long, pale green,1ip slightly inflexed: stamensmuch exceedingpetals to l6 nm long, lilamentsgreen, filamentose, glabrous, anthers elliptic-cordateca. 2 mm long; gynophoreshort, ca. 0.3 mm high; ovary shortly cylindrical, slightly swollen at the base ca. 1.7 mm high, glabrous; carpeli rounded at summit with a short sterilc apex; style + equal to stamens. Cocci erect, 4-5 mm high, rounded at summit and apiculale at outer angle. Seed nol seen. DISTRIBUTToN:New South Walcs, Twofold Bay district Map 34. " NEw SourEWALIS: Wyndhan, Plantsgrow to about2ft. high,much branchcd and twiggy growths,growing on thesides of hillsamidsr litrge loose'wood) stones where a sliRhtexudatioi-t. J. L. Boorman,Aug. 1915 (NSW, SYD); Pcnrbuja. shrub2-3 fr., diy stonyplace',, J. L. Boorman,Aug. 1915 (NSW). Phebaliuru rctlstonii is known only from tltree collections which may have all come fi'om the same locality, the last known collection o[ the speciei bcing in 1915. Referencesby both Benthamand Mueller to a Leichhardtspecimen. and by Maiden to a Forsyth one, apply to a distinct species,P. iritliflorun. _ The related specres,P. viridiflorum ,tnd P. sympetolum,may be distinguished from P. rctlstonii by the stellate ir'ldumentum of the first xnd the utited Detals of the second. All three species are characterized by having a sinilar leaf- shape, while the yellow-green corolla and long stamens are also characrersnot found eisewherein the genus. The three speciesare each confined to a snall isolatedgeographical area and show no prominent morphologicalvariation.

43. Phebaliumsympetalum P. G. Wilson, sp. nov. [sympetalus: having united petals] Ramulianguiares, glabrescentes. Folia chaftacea, obc-uneato-eiliptica, 1.5 3.5x0.+0.gcm, glabra. Infforescentia_terminalis, saepe nutans, 1 3-flora. Caltx herrisphericus,lobis lat6 deltoideisca. 1.mm longis; corolla syDpetala,cylindrica, ad 15 rnm ionga,virido-flava, stanrinibusbreviter exsertis. Holotypus:AD 96434202,New South Welcs, near Olinda, Rylsrone I)istriet,2,700 ft., sand- :rgn9^fjgc:.6.8fr. high. foser. rodding.green. pedjcels rcd. fitamenr. green. enrhers lellou, 2.i\.lq5l, L. A. S. lohn.on. Lorlnu.: NS$ hooq5.

_ Shrub 2-3 m high. Branchlets anguiar, smooth, stellate-puberulous when young,.soonbecomilg glabrous. Leavesshortly petiolate,chartaceous, obcuneate-elliptic,1.5 3.5 x 0.4 0.8 crn, smooth, minutely puberulousalong midrib otherwiseglabrous, margin slightlyrecurved lwhen cli.ylantl 1 serrurare towards the retuse apex, midrib itnpressedabove, prominent below. In- florescencea terminnl cluster of 1-3 flowers, often nutant; peduncle and pedicel slender, s-parselystellate luberulous, red, peduncle ca. 4 mm long, pedicel ca..7mm long with two minute oppositebracteoles; mature flower bud o6tusely cylild_rical_ca._ 12 mm long. Ca11r,fleshy, hcmispherical,glabrous, lobei broadly deltoid,ca. I mm long;corolJasympetalous, cylindricrl, to l5 mir loDg, greenish-yellow, separating towards the apex into erect acute lobes, minuteiy glandularpunctate; stameos shortly exceeding the corolla; lilamentsfiramenrose, glabro_us,green;- anther oblong-cordate, ca. 2 mm long, yellow; gynophore very short, ca. 0.2 mm high; orary doleiform, ca. 2 mrn high, iiregirlarly corrugate;carpels rounded at summir. the upper third srerile:iLyle gla1rou6, + equal to stamens. Cocci erect, ca. 4 mm high, roundcd at summit and shortly rostrate on outer angle. Seed not seen. Chromosolne no. n : 16, from material collected at Rylstone. " fide Smith- While t1954)as P. ral5tonii\ar.". 116 Lll .,. gI 'cl qcerl 'sul€lunou ,{{cor alls?C aql Jo 1oo-I eql lv " V gt {"W ul u"- s'peai'.iir,{^-yiqniplig'e sieeqlPr"qqcl3-I ,{q pe}cellgt-:t!hl..'11"'l

;l*,1,y;;:illlt"mj13-unqurn-rLb6 :.ll*uu;r:":mi'*"",11i,""?'iiij"ffiTilxli srrradss{qI ii 'ijair"u rql ol prlJrrlsarrq ol srEJddE '(.lAI,i{) slloolA 8el 'reÎU i,l,":ii,::i.,j$i);l1f^''on'!S1*,';11,"31"'llii3i rf:''::,.1'":4i":'l';iffi'"'i"i'il;i,'' ,1".ltx liiliuli"'trul +ruH":;11"^'l:Ho11il.'^1"9i['3J''1"'*'^l,li,iJl'ii"'selBy'A '?€ dEl\I-aBu?U elEunqrunr:e16 qlnos '\\eN :Nolrnslursl'f 'uees 'albue lou PaeS Joltlo uo puz pepunor xede'al"8nr.Ioc,{lesre'\sul?rl'5311 alurlsor ,{llueurruord 35 ?^-:3Li ili';*r;":'i""i:,8i;Ti"'.i:;tJ"illb"ig,{"&ilTJ'l['i:sfi"JH"iff'esolueurelgslueru?lg 's1eled "'l*tglg* n", ;]"p;;::s";lqd ,r!qr,rn i.orq.Jti 'eplslno sr: 6uor se ecl,tl ol sueurcls:pexegut ,{llueuturoJd dI} snolnJaqnd l"t"n"il 'r""ia:;trr"r ;,irnri""aitlnput131ee1 Insrop qlr'' suol linJ ln?yl!-n elellels -,u'11 'ec 'eleloecuel-8uo1qo,{1,to-rreu spl:d : saplsqroq uo snolnrrqnd '8uol 'prollep "{qsag '1urlaqdslur-etf SuoI urlu Z e. n1"1,,"e s"qoi -x'{lBJ 'einlqo 'p1ir,lti-,Ltor.l?upnq-le,trog ernl"utlelpplul ol{l JP'eusaloelt€Jq -- 6 n" slcsrq o^r qtr.tr^3ubiruur 3-y slicrped:snornp"c:3Y91---*,!1^^jn"ull 11"-r" 'esorueuror-jiellalsruroJlllsqrun-qns ro esotudc'luelnu iirog :pera.tbg-71-i ;itoqe 'e-sol ""uoioiogul tolaq tuiuruord passerdurr inu,lo,rir 'lurrs:rqel3 are.Jrns-qlrpltltreddn -'ua,iro1-a1n1101,; eizlins ro,lrot t ann"ri r;1'1!l- 'etreunc .'f uJqr^)alnloêj Jo es[q'xedepeqolq,{llroqs Jrll JoJld-JJxe^JJr}tle'(^]p 'e1e1oqed :q1oo-i '.ite * :rqo'11"a';rq' pjitn"". + t.o-t i f7 :l-9i:,")1::tc(Z)t qnJqS kproqsse,rzej asoluourol-el€ll3]s'elalJlslrlqJuelg q3tqu '6t6t 'lal?{ : osl ::*$'s:B;*qilgl.6î:g1";'ifl'#1Til[?iH 1*"1*';i*:.'illi"'fl;^ffi"] l{P3?;5'l[ ",lf,lT:J.'t$![o,q-1t1"i.ii.,^:li:i:$lii{+",#; "1,nauo,n,p,,,n-op,'.[]f,"ill,53',:,'fJ8 ..;ri'.gtrtlli. ifir;".:*::',H"t"9:'"';'.'q'j."1i$r:i:ii,1y"i,l:ii$r"r$.i1:'i#rri.iiT'; '(206r) ii:fffii:i't;iiqal;ffxiffi ''!$i$"*l;ll'#:;i;iiTi,#fi ,T* 'uJJJd stPr'rrr] IP::amog snloU 'tt 'ôu 'ds uosll,1y5 d lunrogpl4^ ulnllvqoqd

's.reqlut oqrpue 'rnolor pu" rd€qs€lloror eql uasx3lF^ "t{l lg il:?"Ji^"i '4 puz'truotslot b1palelor ^t:rl3llo".#l{31'fffif; si'si wn,og4i,ut,," 7 ;i-"" 'uolsnJ ol ue LuoJluollJullslp sLllllsl JJJql JseJ qJlq/$ ul {lEpuoJJ( l? snotutadode :l/EOl s'plo enp'srrclerodur{: srqt ltrc+t D toJ t,]""!-"!1._"1^::..|-qeqord -uooautl idatotpual,/ sf,lJJds Jtll pue D'rJJoJ 5nuJa eq1 sJlquresJJ ll JJIJPIEqJ siqr'ui ,ti"r5o patiun seq qrt\t^ untloqaqdJo sarrrd\Aluo rql sr srqJ qiii,#,F$,H,il''#ii"xtifrT;i :q,'l,#g;,#";tsi1"^',",.;i;y.,;;ffi 'sal?A\ '7g de4-auo1s1,{11 .r?ou qlnos '&aN :Nolrnslursto " duplicate in herb. BRI has a field(?) label reading Rocky creek at the foot of the castle." They correspondto collectionsfrom the WarrumbungleRanees. but I can find no data to confum that Leichhardt was there at the date ciied. Severalcollections, including Leichhardt's,mention that the platrt was $owing on trachlte. - P. viridiflo-rurni.. glorqy related to both P. rulstonii and p. sympetalum,- but may be distinguished by its stellate-tomentoseindumentum and terete branchlets.

1r8 6II '0.6 'trc 'clo 'urujn"lllnc osl" a3s)(tE6l) 961 z pa s3err lsaro'{ '(4ot) -loN^ 'u]unelllnD -.,,n6 'slcuprc apg so'z s1re4 ts{5 leiffiy 'tuotr do 'llrnI y.rapa.ry o!floJlDtt 3:l1r (Sq8l IJO) tt lrJ 'J 1jrrr"o11')''a -. ,iiyt,,ltp otpiolirlI ,sqet iityr g:g:ui6t-r3 110n91I !!lptDqqrtrl .{ 6t) urrunrrnD (Jluol/\)YJrril'Y Lurt\'rerznor]tlol tr)popual'a olptoJjng = (098I) I6t:01 11uolf i5a'pury 'cls sndtoto tll '- (gSSt) wnlaSlnuonbs 'qlrLr{ o.r,tEllnLuonbst)tsolotalsY = (8t8Il'4oog 'g t{ooH) 'u8erg 'llenl't[ lnlootl ii,lilttlj uo-paseq 3411 Luoui69g1) yg1:f c 'ds stttlto:tot11 : (€9SI) '4:?y .qlues (.{ooH) D.!oL[!puD.tBDtsrlpla$v = (8t81) '7 .uta;d-'ll;"o'rt" -("::{!puottsnto{rpuotB unlttDqaqd uo possq tgEsrl soL i- c i'4oog) 'JcseP3 'ds 'l 'g qulpg 'uIeqBrC allrDt:l l)raL ollrld: (?€81) 9ll:9t urnot lllld'$aN 'zarnl DSoJuJatorllouzaP1 = [(gt8I)^ 'g 'ur8lrg '11en11 g rtltouza|aS'n 'qcru y,trsotr./1a,r uoul (oEtij rdlr '^ou 'uoslliN 'D Ertroro{ quor d Gtela) saprcrslqtJ 'r :l;{ ile;}lrtle slelc uI s9tcr saplouPlqo} = (n061)I T x6t 1'azt:gt qrq"t '(9581) ,trtg x3 turullrd saPrrl '3a1[ ' tlt:)uowlunrpg -lpqaryl snd.lorotn = tuou (gSSL)SOt'l ru3"i1 llen] l I '(soot)st 'lJ lezN 'rlulA JpU 'ltlEA {-rJlsrol D IJ U'f) rt?/''1l'f.Fnd:D1o'DlJ urTrlrJ\JJqJ :t 'rlloJ snlPtuJ|J iluo,r,l,.,r.op"enb\quttzSlr 6g 'r'u:1ndrg slrlroH 'ds sndtoto.tll : (gSSr) qlLreL Duq/lxl1q'' Dls0lorals'l = :uraerg 'J llwvtt8ulaun'1 [(oogt)u,s trtt,,tti,i'i-ii,ii tsit iiioi'e i1en1"q 'ufl1€lllnO (IIrqE:r)^runriis DS oq w,Oo:t sLsdo t t I hl 'g -t '69 ' wnsoqurr'tot = Gc-vz8 8s *n"'"up"1o"i'o'f*i 1itq.o1 'ds sttdtotot2 (;ssn1)otloJia.uot DtsDlo DtsY = (sz8tl ":'\,'-y y!!t1.["'otto" .= rilueg ''tt:ntri:s ^ srulo{l torg tunuiqn,t)io piinri'1[9g11991:1' LtrBcr4 (narssnfv) 'u",\\C 'tP^ i^tuo{llfw ( ll3ntrl C) saPtorDtPnq 'il""ili sapi'tot1int1 'teL snlolldDrg arlt:tonrlq= (q/81) ror,o-:.,ig;l i 'zaJnL'DtodDnDd aqhol)tlI : (EqSL) l srptoLDDltloq) --u"J lLrnIN' 'J sniDtltln'g ,rrr1rr",,rE p-"rnq'":3"11,,i,oti (oss'rlso i'-i:u'airj 1!n1'ri 'otlfM ( -1, D'tul{tttnDtfttQuaPv '- 1 'uoosta4 'g ''3e g9}: u'{5 asuadot Dto{tutlDLttsot o uo pes"q I 'n.,u Gdsij 1 1i ul ']JrrrocuL rq oJrJrql ^.Pruprocer '{lll?col "l] '9 s"al!:1ryj1:1Y-ln'1o"J snueBslgl Jale'troH Suurnrlo sB";qielbpda.ldiq3 ,(q pjprocer uaeq lou :peinuaq!",1; rtl-l-r-l edf,l ;;;;; ;t 1o.i Dryot'(ttd .'1ua3' "-Ilrlrsnv^punql "qr go'"i seieei pue ruils oq1 uo salersrrellad:q1 ll]rlueJ '}W - *eu-sue{lelJ ^q ur (,, sropumg ,, I3q!l 1:urarro ei11)repuog "tuo4 1unol'^tr q5un'q'"1""it u poqr'rlsapse't*ltlad. DJlrrrroJue.q a^eLlot ltLdnoql 'd pur ir,i;":i'';i;J:;iil;;v...'1t'otipla'1 Lt d' DtP"lrttd' {0b81)lb )prfsnV lcog '.{oA 'a1e1 sn)lldlo uowalsoug 6i:*g1 i s'uerf la llrntr4 C e sel"clpul : (ut,(uou,{sprnlelcueulou ? s3}uclpul = :ulfuours clutouoxB] )

S1WYN OAOO.IJXA E. linearifolium DC., Prod. l:720 (1824) = Geijeru pamifora'tar. crassifolia Benth. (1863) = G. linearfolia (DC.) Black (1924). E. marginata (L.f .) J. E. Smith in Rees,Cyclop. 13: n.5 (1809), basedon Diosma marginata L.f. (1781) = Adenqndrq marginatq (L.f.) Roem. et Schult. (1819).

E. no1lr,s(Benth.) F. Muell., Syst.Census Austral. Pl. 1l (1882)nom. illeg. = Asterolqsia mollis Benth. (1863) nom. rlIeg. = A. correifolia yat. mollis Maiden et Betche (1902), based on Phebqlium hexapetalumA. Juss. (1825) = A. hexapetala (A. Juss.) Druce (1917) : Urocarpus sp. E. noyae-caledoniaePancher ct Sebert, Not. bois. Nouv. Caled. 271 (1874) non vidi, nomen nudum fide Guillaumin, Bull. Soc. Bot. France 67:64 (1920) pro syn. sub Mytopsis macrocarpa Engl- E. oblongfolius (Hook.) G. Don in Sweei, Hort. Bdt. ed. 3. \29 (1839); Jamboli- fera oblongifolia (Hook.) Steud., op. crt.796;,based.on Cyminosmaoblongi- folia Hook. (1834) : Aoorrychia /aevrsForst. et f . (1776). E. pallidus (Befih.) F. Muell., Frcgm. 7:22 (1869) = Asterolasiapallida Benth. (1863) : Urocarpus sp. E. pollidus R. Schlechter, Bot. Jal.rrb. 39:142 (1906) nom. illeg. non F. Muell. (1869). Type from New Caledonia. Probably best placed n Myrtopsis, certainly not a species of Eriostemon. E. paradoxus J. E. Smith in Rees, Cyclop. 13:n.6 (1809) = Boronia paradoxa (Sm.) DC. (1824) : Boronia sp. E. pleurandrotulesF. Muell., Fragm. l:106 (1859)nom. illeg.,based o\ Astero- - lasia phebalioidesF. Muell. (1855) A. pleurandroidesBenth. (1863) fnon A. phebalioides (Harv.) Benth. (1863)l : Pleurandropsis phebalioides (F. Muell.) Baillon (1872) : Urocarpus sp. E. salsolfolius J. E. Smith in Rees, Cyclop. 13:n.3 (1809) = Philotheca austrelis Rudge (1815) nom. illeg. (superfluous epithet) = P. salsolifolia (Sm.) Druce (1917).

E. sandfordii F. Muell., Fragm. l:107 (1859) fnon E. calycinus Turcz. (1849)] based on Sanfordia calycina Drumm. ex Harv. (1855) : Geleznowiu calycina (Harv.) Benth. (1863).

E. spathulifoliusGandoger, Bull. Soc. Bot. France 60:458 (1913) "Hab.: Australia, Victoria ad Esmerald [sic] (Macl-ennan.)." I have not been able to obtain a photograph of the type of this species; however the des- cdption agreeswith a form of Asterolssia found at Emerald in the Dande- nong Ranges of Victoria. This form was included :under " Asterolasia muelleri Benth." by Ewart, Fl. Vict. 703 (1931) [- A. asteriscophora (F. Muell.) Drucel, but it may deserve recognition as a distinct species under Urocarpus.

E. trinerve Hooker, J. Bot. Hooker l:254 (1834) type from Tasmania, non Leptospermum trinerye DC. (1828), : L. glaucescensSchau. (1841), fide J. H. Willis, Muelleria | /3:136 (1967). E. trymalioides (F. Muell.) F. Muell., Fragm. I :106 (1859),based on Asterolqsie trymalioides F. Muell. (1855) : Pleurandropsis trymalioides (F. Muell.) Ewart (1931) : Urocarpus sp.

E. unifora (Linnaeus) Smith in Rees, Cyclop. 13:n.4 (1809) : Adensndra uniflora (L.) Willd. (1809). 120 'urou 'pnu (I I6I) 'loc 'uuY 'urun"lllnc xa reqcuEdsttlo|rund'1 0II:6'Z'r3S ellosr"Itr snni '(f,luo eJnllncJo sll€laP 'xnrg '^eU 'sraqAes '! qlrm) 'pnu ulou (t68I) 16:02 e8lag lroq wnlPf!ry l 'pe 'll.rq 'uoH 'leer,\S volrY suaJsarnolS'g 'pnu urou(6Eg1) 99, : € ur uod D xe 'pnu LunplawlD'g 'uou,(6EgI)6ZI :g'pa'lug uoH'laa.',!\suI uoc'D xe,{e1celni '(i68I) '( slIErrPqt!,,|\) pnu 'urou /6:02 sellsxnrg Iuo arnltnr Jo rsrrqtsts 'erq3uu19 1a i61aq e:nlpctgoq.l rp Jn^rll s"ol'sll'p uouatso!'tg V(ION VNII^ION

'ds sndtocotll 'trll|mou. : 'qlreg ({ooH) otaBqruuonbsolsDlota$V = (6t8I) (898I) j8 - 'prii,1l j r)1oof, iot uafidttE : 777' t (8t8I) Id JI rrl oo11wru az11ruuonbs 4 'flanhtr 1rI ( qluag) 's!11otluowalsoltg ees .t"N rurelrl 'nelssnf wryoladoxaq (gz8l) I'J II'1 I€I:z slred i;fi:cog v 4 '(698I)'[ant't'C C{ooH) 'ld'cI 'JoIooH sn.n7frpuDt|uowalsou7 oes (8t8I) vzL'l:g wntoglipwtB 4 'ltalxoq d '(€98I) '.{ repsn elou oos 'snuaapullslp " Je^e,totlfDuergddv llenru )iinti* srdalotrtutaltr7ir'(ur,{uorstq Jo. uoll€clpul 1ca:tpuruc '{po 'runirj 'reupr?g (llontrI 'g) aorutaqdna qri^) treOr) Ol'ccA-'le:1sny 16 4 ',, '(6S8I) 'llenw 'C ('ssnt) snllollattoc uowaJsoltE'0s unlloJe"eroc ', :gi1.z '1e51 '...etr 'nalssnf 'v wrulollt]o' 'd riTgii oir irrei 1s111tos tr 'cos 'uul-I 'f 'erooy{ 'ds aq,lco.tcryu: (OZOI)99I:9t ioq S unlDlldo|d (It6l) flel€lg (AP1e1g)lrutot1 'tersnv ',{lerletg 'd -Ba!4rnqolsDpatsy = ((,16], 9tz:0l 1EN uwoqBuDpnq 'snua8 e se uotlluSocelsa\JJsJp pu" (selJedsedft )ql)'zJJnJ lcutlstp'jy '(€981^'rdV) saprcryoqaqd' i1tr,'rrr-reu:3uo.r lou st setledssrqf 6tl:t lili"j, "ir"ni,nr: -q aoiwa4dnalv' : . rrrlxed.. (9681) rrl8u3 ( ttluog) uatxof, stit)1oiiwu1'1: (€t8I IBt{ 0€) 9t€:l I?rlsnY I{'tueqluag ualxDq'd 'J) 'ds sndtocotI : (ftOt) ecnrq ('1en14 otoLldo?s 'qluog, -l@tso'V = (eu?u snongredns)Soll[ urou (e981) Ya anu olsDl ''[ent{ 'J oqaqd -ota$y 7 (SS8t)tg:t'lllsul lcl^ su"JI umtoLldo?slblsDum '1ces 'sndn)o.tn: (otSI) 99ll Id'uoo'Ipu] saploaoJ unlDqaqd ACKNOWLEDGMENTS

My thanks are due to the ofrcers in charge and the staffs of the various herbariafrom whom specimenswere borrowed, and also to the directorof the BotanicalInstitute of the Academyof Sciencesof the Ukrainan S.S.R.,Kiew, for sendingphotogaphs of the typesof severalspecies described by Turczani- now. I also wish to thank Dr. S. T. Blake for the trouble he took, while botanical liaison offcer at Kew, in comparing several specimensheld there. I am gratefulto Dr. N. T. Burbidgefor allowing me to read her unpublished notes on the relative dates of publication of the generic names Urocarpusand Asterolqsia, I should particularly like to thank Dr. Hj. Eichler foi photo- gaphing the collectionof typesin the relevantRutaceae genera held af Kew, and for bis encouragementand adviceover several years.

1.22 'wtsoquonbs 'g 'lYNsrNsA .zol'qet tZ uoslerlrle1l1el ep uIp&I alllPqa'ld (S08I) J 'bl tsol IEN JoS lunJEllnuuouulnl'lleds snqtuoltduJsrpsllr'fpe tltz/zz IlnB "^\o\rNvzjunL *iioii'r-l.li uou-5nbs-nrnqtunretueld urnjlu:8 €lres sEJtc {6t81) N '€0€-L8Z:Z log { eq) uo Suuerq 'leJlsn\ 'Erogu"tl€llsnv eql u! eqrrraul ro lueurdolo\op^reuollnlo^r 'duHla-Hul{s :ttiosocuorq3 {ts6[l r"i ii-p'"" i"ni""lnir.l ezjruoio6 e{i u; sriqurnu s '/Z (tI8I) Io^ elpeedolc'{J .saed rutllPqaql A I'IIuIÎS '€I (608I) 'I{ll''{S lo^ Epe€dolJ'{] 'saad uoaalsoltg A -f '0I (808I) g lo^ €lpe€dol'f3 ,ste>J Dahu) t'Hll''{S 'vz-nzz:9 'slueld (86tI) 'g 'IIII^S '.oS uul'I su?rtr Jo ?leue8 ^\au ,{lue'Alol Aur}Blersrop€leq) f I [o^ ru €]ueld slsdoufs (908I) H J'Noosusd .'€I 3II:'{411:lrri I eeecElntl (298I) No^ J l €Lrolcr^ Jo xuoloJ eql o] snoue8lpul sluEd eq.L 'I0I-Z0I: (698I) No^ 'f,a11aolÎ I eEIIe4snV eeqd"lSol^qd €luewBEJd C '.1tl^'cos 'llqd II-6:I susrl, (998D No C'{d'I.Iany{ 'zI-oI qel 'LEr-9ZliZ 'unlpqa\tr '(ecz8l) ao 'oflssof srred teN lsrH .os tuql I eluo8 np qdeJSosow v 't8n-rwizr ad 'nflssnl srred leN tsrH snr{ tuqhl soFetnll sop ednolS el ms eJlourgl l GZ8I) V 'zurelÎ '€9 3I 'l) 'reu-qrer{ ssld\ €9:t961 "p!lY !9y, !?::'Ep!l€xo)('s6I) 'c'1ruYH otogqnt sloii piin (ei6elia,f iiooels iaii3 ui'r erleloour'4sopnrsdrr( '0/-69 sllEluepl.O eele4snv ulru€luelil oll€toqmua (lt6l) V'C'{aNc{Y{]} 'pe 'do 'teec?lnl{ 'rra'lcNg .s9z-6lz,e6l'z llc 1lserd le ret8Eg e€oluolog C€6D V ' rrttf,LweJ '.€ecslnd9t l-lEIt '068I) I / E 'lalDNg -uezuegd uoqcqrot€N 610 llllerd lt JeISuA eeeruoro8 V 'uxlpqaqd .02/-6Il:I s{srn}eN sl}€uetsrs snuo.Ipord (tz8l) sa'd'v'qrrooNYc '692'o '{'a^fto pu€ 'g 'la '1'l\.t3v',Ig srea\ouplrl\ uelellsnv uJelsodl ?!\ou){o},{\oH (656I)'f 'Zl€-I0€: 'O 'r{YltrNsg I slsuelleJlsnv erold e€tJelnu (€98I d€I/'l 0€) '088I) 'A 'H 'No,rtrv{ 'l,lZ:II enbluelo{ eP orl€ulro4clo 't'Io^ setu€la sop eno$IH (€t8D a H'No'rrYg '(008I) 'H'snArf,oNv '6L q'Jl ta,(lollsode{ s}sru€loE

(sJ8ed ,fuolJnporlul ur pelrr s{lo^[)

SAJNEUACAU EXPLANATION OF FIGURES

Fig. l. Eriostemondeserl/: (l) L.S. of carpel showingplacentation of ovules,(2) Attachment of ovules to pl^cetta. Crowea angustifolia seed: (3) Side view, (4) Adaxial face, (5) L.S. through base. E. australasius seed: (6) L.S. showing embryo, (7) Adaxial face, (8) Adaxial lace with attachednlacental endocarn- (1-2 from Wilson 3514,the rest from cultivatedmaterial)

Fig.2. E. mloporoidesseed: (l) Side view, (2) Adaxial face, (3) L.S. through base. -8. difformis seed: (4) lA.daxial face, (5) Side view, (6) L.S. E. gardneri seed: (7) Adaxial face, (8) L.S. E, tornentellusseed: (9) Adaxial face, (10) L.S. (l-3-from R. Hill990; +6 from cultivatedmaterials; 7-8 from A. ceorge s.n.; 9-10 A. ceorge 4239')

Fig.3. E. thtyptotteror'rle,r seed: (1) Adaxiat face, (2) Side view. E. pungensseed: (3) Adaxial face, (4) Side yie\at: E, spicatus seed: (5) Adaxial face, (6) Side view. E. desefii seed. (1, Adaxial face, (8) Side tlew. P. brachyphyllun seed: (9) Adaxial face, (10) Side view. P. fade s€ed: (11) Adaxial face, (12) Side view. (1-2 f-romBlackall 3490; 3 4 from Shaffad 409; 5,6 ftom cardner 2702; 7-8 from Voitch 4; 9-10 from D. Hnnt 2222;11-12 from A. ceorge 7263)

Fig.4.-P. squameumseed: (l) Adaxial face, (2) Side view. P. bullatum seg (3) Adaxialtace, (4) Side view. P. brachyphyllum: (5) Endocarp with seed enclosed, (6) Adaxial face with placental endocarp attached, (7) Elastic endocarp aftet dehiscence. (1-2 from Macfarlane s.n.; 3-4 from Wilson 1002; 5,7 from D. I{runt 2222')

Abbreviationsused in illustrationsof seed- c : clalaza, m : micropyle,h : hilum, r : raphe,hr : hard covedngto raphe,tr : thin covering to raphe, s : scletotesta.

Fig. 5.-Anthers (abaxialface exceptwhere indicated). l_,.Ctoweaangustifolia, (a) Aba"\ial, (b) lateral (median) view. 2, E. australasius-

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S ern8ld t4ap I C. angustifoliavar. angusrifolia O var. dentata X r. mrcrophy um /

Map 2 C. exalata O C. salilna l, E. brevifolius. E. pungens X I€I

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ii I Map 4 E. coccineus O E. pach/ph/llus A E. nutans X

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Map 5 E. angustifoliussubsp, anSustifolius O lubsp. montanus X E. difformissubsp. difformis A subsp.smithianus a 't 32 cct

sneploquroqi'l V 'l v snafllas 6 ,a,iqsi / d?N

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Map 8 E. spicatus O E. thryptomenoidesX

Map 9 F .1.<6ri E. fitzgenldii X E. Sardneri A

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OO !oo$o t- E. scabe. subsp,scaber X subsp.latifolius O E. ericifolius r\ E. linearis ;

I Map 13 E. hispidulus X E. buxifolius O P. phylicifoliumA 136 LEI

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Hap 15 E. obovalis X E. verrucosus O P. ambiens A

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Map 16 E. brucei subsp. brucei a subsp. brevifoliusa subsp, cinereus A

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O xllitolsrl dsqns '3 a snroutPou dsqns snroglPou /l drN l'4aPl9 P. tuberculosumsubsp. tuberculosum o subsp.megaphyllum A subsp.brach).phyllum a

-^ I Map 20 P. squamulosumsubsp. squamulosum o subsp.alpinum X subsp.ozothamnoideg v

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P. squamulosumsubsp. argenteum o subsp.coriaceum subsp.lonSifolium A tz d€N

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o Map 25 P. anceps X P. ambiSuum C P. clavatum L

..4 Map 26 P. ovatifolium X IFfl P. squameumsubsp. squameum O Vo t subsp.coriaceum A qed suDsP.retuslm a

144 aff\ (' \ / ool /^o ovv \ \:,*-\ I V !!PIeUPlo'd Fr )f l)

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Map 29 P. carruthersiiX r. cox | ) P. dentatum V

Map 30 P. daviesii a P. diosmeum O P. lachnaeoidei A l 146 L'I

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o eov9.-r"- ^-'- -'- ."ab' Map 33 E. virgatus X P. bilobum O P. hillebrandiiA l l l l

0 Map 34 ,I,o. P. ralstonii X $"- P, sympetalum O P, viridiflorum A 148 6rl

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Xf X f l ',rt ?YVi ll. l x n XX INDEX TO NAMES

New nanes are in bold type, accepted names in Roman, others ln italics. Numbets refer to the page

AdendFJrcwilld. . .. Lnilora (L) willd. .. 119,t20 Asterolasial. Muell. ' 10 a.taieco?hor (f. Muell ) Druce ... 121 Arl.klr?grdn?il f Blakely) Brrkel) .... l2l b tilllia Benth. ... 119 chotilocnoidP\ f . \,4rell. 119 rolr y'i1ia (Jus..) Eenlh. .... r 19 \rr. ,ro1l^ Maid, er Berche .... 120 glaaily'/ola t Hook.t Benrh. .... 119 fit rappral,r(Ju.s.) Drttce ... 120 mollt'Benth. .... 120 muelle,i Benth. ... 121 pallida Benth. .... 120 phtbalnide' fHar\.) Benth. .... 119 phPbalioide' | . \4uell. .... t20 dc findroide\ Benth. .... 120 .,quon ligcro (Hool.\ Ber'th. ... 119 rrymalioitlt" | . Mrcll. .. 120 woonble F. M. Bailey .... 88 tar. i,tr.rm,dia f. \4. Baile) 88 \ar. p,r,'fy'b1ld f. l\L Baile) .... 8t, lJoronia J. l-. Snr. fxbianoides(Dielsr P. G. Wil.. ' rls Lpidrta Tutc/. pa.adoxa (Sm.) DC. .... 120 Chorilaenatndl...... l0 ungu:tifolia f . Muell. .... 104 Crouea J. E. Sm...... 15,8 secl. f//o'reno, (Snr.) Ba'll. ... 20 lecr. lra,'ort?a Baill, 15 5ecr.PrPrar'l,''? (\ enl.JBaill. .. 60 angu.tiloliaSn.. Map I .... 16 \ar.denrala (Benrh.) P. C. wjl'. t6 aueu,tilolia furcz. 16 \ xt. plat fph ! l la Benth. 16 dcntata Benlh, ... 16 exdlatrL M|lell.,Map 2 .... 18 lariJblill C. Don ... l1 nncrantha Hor\, .... 18 /ldlrii (F. Muell.) Baill. .... 89 .alignaAnd".. Map 2 ...... 1'7 major Hott. .... 18 s/riardHorl, ... 18 sraDlaR. Crah. Dio,rna L. maryitnto L. f. ... rzo .... 119 Dnrmmondira .... 10 tlaeocafpusL denlaru

hil lebrandi i F. NIuell. \ar. brcvifoliusF. Muell. . . .var.lo gifoliusF. Muell...... _.. nrsproulusSpren€.. Maps. l.J. 14 ...... nooKertL l4uell. intermedius F\ a intcrmedIu,Matnock AendackMontr- lamprophyllns F. Muell.. lanceolatusK. F. Cae n. lancifoliusF. Muell. lci.hhardtii F. Muetl. lcpidotus Spreng,. var. Ila duluru| F. \lucll- linearisEndl.. MaD 12 lincarifolius DC. . . liniloliu5 Seghers marginatur\L.f ) sfi. naxw?llii F. Muetl. ..._ nicrophrllu! F. Muell. mol/rr'(Benrh.)F. Muell. .... montanus(Hool.) F. MuelI. morronii F. Uuell. myoporo;desDC.. Map t0 ..._ var. .r.alrs Blakel; subsp.acutus (pralery) !. c. Wii;,,rraip: ...... subsp.conduplicatus p C. Wils..Map p. -] ...... subsp.epitosus C. Wils..Map II .. sltbsp.leichiardlii .... {Benrtr.tf. C. Wits.-Vap t| ... vaf. mlnor Benth. .... 40,4 {C. I. whire) e. b. w;ti.. Vaf |l ncftuouus.__..,.::I:1.-qu""nJundicus spfeng. nodillorusLindl.. Vap t7 r ar. lasiorclyx Donin slbsp. p. lasiocalla{ Dontin) C. W jls. ... . \ at. su b g tabr ifi or ur l) omin ,o/r F. MuelL. novae-ealedoniac panchir er ieberr nrdas(Hook.) F. Muell. nutansP. C. Wils..MaD 4 ' obcodatu: Hook. . oblongifolius\Hook.l C. Don oboralisA. CUnn..Vao l5 ' oldfeldii f . Muett. ovatilolius\F. Muell.)F. Muelt- ozothamnoidet\t. Muell.)F. Mucll. pachyphyllusP. C. Wits..Man a pallidus(Benth] f. Muell. palr,y'//rR. SchlechL paradoxu' Sm. pareifoliu5 Benrrl ph),licifoli! tF. Vuett.t f. Vuilt. .. var. lachna?oidcsMoore .. . phllircidcs tsprene.) F. Muell. pinoidesP. C. Wilc..MaD 6 pkurandroides F. Muell. pulchellus Lemaire Du clattryGxill pungensLindl.. Mao 2 qurcnslandicusC. T. Whirc quercifolius(fndl.) F. Mucll. racemosustndl- ralstonii F. Muell. .. rhombeusLindl rhomboideusP. C. wit;.: Va; i rotundiloliusLndl. .... :alicifolius Sm. ra16'nrr(Andr.) Baill. ,nlo'1;L1;,. a^, 152 99r29-(Z)

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