Aliso: A Journal of Systematic and Evolutionary Botany

Volume 18 | Issue 1 Article 12

1999 A floristic study of Sugarloaf Ridge State Park, Sonoma County, California Frederica S. Bowcutt Ecology Graduate Group

Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons

Recommended Citation Bowcutt, Frederica S. (1999) "A floristic study of Sugarloaf Ridge State Park, Sonoma County, California," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 18: Iss. 1, Article 12. Available at: http://scholarship.claremont.edu/aliso/vol18/iss1/12 Alisn. 18(1 ), pp. 19 .)4 C 1999. by The Rancho Santa Ana BoiJlllic Garden. Claremont. CA 91711-3157

A FLORISTIC STUDY OF SUGARLOAF RIDGE STATE PARK, SONOMA COUNTY, CALIFORNIA

FREDERICA S. Bowcvrr Ecology Graduate Group University of California Davis Davis, California 956161

ABSTRACT

A ~pecies list of 400 vascular plants was compiled for Sugarloaf Ridge St:ue Park, a 1142 hectare (2820 acre) park in the outer Nonh Coast Ranges within Sonoma County and adjacent western Napa County, California. Relatively undisturbed siJlllds of native perennial bunchgrasses occur a.~ the un­ derstory In ponions of both the oak woodland and the mixed evergreen forest. Grasslands on serpentine In the park are typically dominated by NasseiiLI. One of the sttlte's eastentmost coast redwood forests occurs in the park. Nonnative Centaurea solstitialis rapidly invades annual grasslands and other dis­ turbed anea~ in the park. The California Oepanment of Park ~ and Recreation (DPR) maintains an active prescribed burn progr.un m an effort to con110l Cenrournt solwtlalis populations. DPR alw Ul>C ~ prescribed burning to retard Pseudotsuga men1,iesii es tnblt~hment m oak woodlands. DPR biol­ ogms believe that fire ~uppression in this region results 1n an unn:uural type conversion of oak woodlands to milled evergreen foresL Key words: California state parks, Sonoma County, Sugarloaf Ridge State Parle. vascuJar flora.

lNTRODUCTlON The subject of edaph.ic endemism can be taught using ln this article I tist the vascular plants wh.icb occur the serpentine chaparral and grasslands in the park. within Sugarloaf Ridge State Park. Climate, geology, Due to the high oak diversity and resulting hybridiza­ and vegetation are aJso described. The park occurs in tion, the opportunity to discuss problems with the spe­ the North Coast Range of northwestern California, cies concept exists at Sugarloaf. The successional pat­ mostly in Sonoma County, but including a small area tern of Pseudotsuga menziesii invading mature oak in adjacent Napa County. Until approximately the mid woodlands. and invasion of grasslands by Centaurea 1800s. the Wappo tribe of indigenous people lived in solstltialis provides an avenue for discussing the influ­ a village at the headwaters of Sonoma Creek, with.in ence of fi re in ecosystems and land management. In current park boundaries (Breck Parkman, personal addition, using the C. solstitialis problem in the park, communications). The convergence of several differ­ discussion can easily focus on other issues of exotic ent plant communities makes th.is area species rich, an plant management, including biological control. G. ideal condition for subsistence Living. Since then. past Ledyard Stebbins (1998) recently tisted the park as one land uses have included limited conifer logging, char­ of the "outdoor floristic museums" in the greater San coal maJcing (from oaks}, and Livestock grazing, how­ Francisco Bay area which is "accessible to research ever, these activities ceased when the park was estab­ workers" helping to make Sonoma County "most fa­ lished in 1964 (California Department of Parks and vorable" for research opportunities. Recreation 1992). Given its proximity to urban areas, and many insti­ LOCATION AND PHYSICAL GEOGRAPHY tutions of higher learning, Sugarloaf Ridge State Park is an underutilized teach.ing resource both for second­ Encompassing 1142 hectares (2820 acres), Sugar­ ary and undergraduate education. The park is an ex­ loaf Ridge State Park ranges in elevation from 183 m cellent resource for teaching basic field plant ecology. (600 feet) to 832 m (2729 feet). The park is located North facing slopes support more mesic vegetation in the mountain range between the Sonoma and Napa than the outh facing slopes facilitating discussion of vaJleys. Mount Saint Helena lies to the north-north­ the impact a pect has on microclimatic conditions. west 24 km (15 air miles). The mouth of the Russian Supporting one of the easternmost Sequoia sempervi­ River at the Pacific Ocean is due west 55 km (34 air rens stands, the park provides a means to discuss the miles). Lake Berryessa is roughly 29 km (18 air miles) biogeography and autecology of a prominent species. to the northeast. Santa Rosa is 16 km (J 0 air miles) to the west. The small town of Kenwood lies 5 km (3 1 Current nddresl>: The Evergreen State College, 2700 Evergreen mjJes) to the southwest. The park encompasses Bald Parkway, NW, Lob 11. Olympia. Washington 98505. (email: Mountain, Red Mountain, the upper reaches of Adobe [email protected]) Canyon, and part of Sugarloaf Ridge (Fig. 1). Sonoma 20 Bowcutt AUSO

Legend

.. ~ roads fire road trails -..... "• .. foot-trails boundaries ,.---.. creeks

A campground

Fig. 1. Map of Sugarloaf Ridge State Park.

Creek bisects the park. Most of the headwaters of this This sequence is a melange of marine mudstone, silt­ creek are included within the park boundaries. Sonoma stone, sandstone, and conglomerate. Creek ultimately drains, via Sonoma Valley, into San Pablo Bay 32 km (20 air miles) south of the park. Also CLIMATE included in the park are portions of Bear Creek which Like most of northern California, Sugarloaf Ridge also drains into Sonoma Creek. Adobe Canyon Road State Park receives a majority of its precipitation dur­ provides access from Highway 12. Hood Mountain ing late fall , winter, and spring (Fig. 2). On average, County Park connects to the western boundary of Sug­ the total annual rainfall is 888.5 mrn (35 inches) in arloaf Ridge State Park. Santa Rosa, which, at 16 km (10 air miles) to the west of the park, is the closest weather station (NOAA GEOLOGY 1993). The average annual temperature in Santa Rosa is 14.7 C (58.4 F). Being at an inland location, the Sugarloaf Ridge State Park is relatively diverse geo­ park has a wide daily temperature range, particularly logically (Wagner and Bortugno 1982). Serpentinized in summer. During summer the park is hot and dry, ultramafic rocks (peridotite) of Jurassic age are best and as a result evapotranspiration rates are high (El­ represented in the northwestern portion of the park. ford 1964). The evapotranspiration rate is approxi­ Quaternary landslide deposits are found primarily at mated using the Thornthwaite method (Fig. 2). As this the toe of the south-facing slopes below outcrops of approach averages the high and low daily tempera­ serpentinized ultramafic rocks. This pattern is unsur­ tures, it underestimates the actual evapotranspiration prising given the instability of serpentinized material rate (Shelton 1978). The average growing season is when wet. The northeastern portion of the parlc is char­ roughly 200 days (Elford 1964). Winter storms bring acterized by the Franciscan Complex. This complex is strong southerly winds. Freezing conditions begin common in the North Coast Ranges and is a melange around mid-November and can occur as late as mid­ of sandstone, shale, conglomerate, chert, greenstone, April (Elford 1964). Snow does fall in winter occa­ and metagraywacke with inclusions of serpentinized sionally, but usually melts quickly. ultramafic rocks. Sonoma volcanics of Pliocene age characterize Sugarloaf Ridge. A relatively small out­ VEGETATION crop of Lower Cretaceous-Upper Jurassic Great Valley Eight vegetation types and associations occur in Sequence occurs in the eastern portion of the park. Sugarloaf Ridge State Park: annual grassland, peren- VOLUME 18, NUMBER I Sugarloaf Ridge State Park Flora 21

180

160

140

120 - -1•1-- precipitation (mm)

100 -D- potential evapotranspiration 80 (nun)

60 -~..--- temperature (C)

40

20

Ql > u § 0 Ql -.. z 0 Months Based on climatic data from the National Oceanic and Atmospheric Administration collected between 1961 and 1990.

Fig. 2. Thomthwaite diagram for Santa Rosa, California. nial grassland, cham.ise chaparral, serpentine chaparral, After two consecutive years of prescribed burning in white alder riparian woodland, oak woodland, mixed the park, the percent cover of native grassland species evergreen forest, and coast redwood forest. In general, increased. In addition, after the burns in July 1993 and plant community boundaries intergrade thus creating a 1994, C. solstitialis percent cover and the number of complex vegetation mosaic. This is particularly true of its seeds in the seed bank declined significantly (Has­ the oak woodland and mixed evergreen forest. Road­ tings and DiTomaso 1996). C. solstitialis continues to sides and other disturbed places serve as habitat for a plague many grassland areas considered impractical to variety of weedy species which are primarily nonna­ bum due to limited access for fire control crews. State tive. parks plans to continue herbicide use in these areas (Hastings and DiTomaso 1996). Annual Grassland Perennial Grassland Nonnative annual grasses and forbs from Europe dominate most of the grasslands in the park. Dominant Most of the perennial grasslands occur on serpentine species vary and include Avena barbata, Afatua, Bro­ substrates at the interface between annual grassland mus diandrus, B. hordeaceus, and Cemaurea solsti­ and serpentine chaparral. The perennial grasslands are tialis. Common associates include Aira caryophyllea, dominated by Nassella spp. Native wildflower diver­ Briza minor, Lolium multiflorum, L. perenne, Taen­ sity is higher in this plant community as compared to iatherum caput-medusae, Lathyrus cicera, Trifolium the nonnative grassland. Percent cover of vegetation is spp., and Vicia villosa. Livestock grazing of park lower than the nonnative grassland which approaches grasslands occurred between 1942 and 1964, and pos­ 100 percent. sibly earlier (California Department of Parks and Rec­ reation 1992). This past land use probably helped to Chamise Chaparral favor nonnatives over native species. Occasional na­ tive species within the annual grassland include Cal­ Cham.ise chaparral occurs primarily on south-facing ochortus luteus and Lupinus bicolor. Elymus glaucus, slopes. Species diversity is relatively low, with Aden­ a native perennial grass, often occurs where the grass­ ostoma fasciculatum forming a closed shrub canopy. lands border with oak woodland. Occasional shrub associates include Arctostaphylos Park staff bum the grasslands to increase native manzanita ssp. manzanita, Hereromeles arbutifo/ia, plant populations and control Cenraurea ~olstitialis. and Quercus berberidifolia. The sparse understory is 22 Bowcun ALISO made up primarily of Nassella cemua. During the first The oak woodlands may represent a disclimax created few years after bums and other forms of disturbance, by native people through burning (Breck Parkman, herbaceous species diversity increases. Post fire asso­ pers. comm.). Impetus existed to manage for oak ciates include Apiastrum angustifolium and Emmen­ acorns, which served as a staple food. Without burn­ anthe penduliflora. Lomatium repostum, which is on ing, portions of the oak woodlands in the park may the California Native Plant Society watch list (List 4), ultimately become mixed evergreen forest (Barnhart et occurs in this community. al. 1996). To maintain habitat and species diversity, state parks staff burn some of the oak woodlands to Serpentine Chaparral reduce P. menziesii populations (Marla Hastings, pers. comm.). Based on two 20 m X 50 m plots, one in the A distinctive chaparral community occurs on ser­ oak woodland and one in the mixed evergreen forest, pentine-derived soils (Hennecke Series in the park). there is some support for this management strategy, Ceanothus jepsonii var. jepsonii and Quercus durata albeit limited by the small sample size (Bowcutt un­ serve as indicator species. The native perennial bunch­ publ. data). The relative floristic similarity of the two grass, Melica torreyana, frequently dominates the plots was estimated by calculating the Jaccard com­ sparse understory. Other understory associates include munity coefficient (J), where J = (C/(A + B +C)) X 100 Galium porrigens var. tenue, Lessingia ramulosa, and with A total of number of species in stand A, B Malacothrix jloccifera. A healthy population of Cea­ = = total number of species in stand B, and C = total num­ nothus sonomensis occurs along Goodspeed Trail, on the south-facing slope west of Bear Creek. This spe­ ber of species in both stands A and B (Barbour et al. cies is considered rare statewide by the California Na­ 1987). My University of California at Santa Cruz Field tive Plant Society (Skinner and Pavlik 1994 ). Ecology students and 1 found a lower vascular plant species richness in the oak woodland (36 species vs. 46); however. the Jaccard similarity coefficient be­ White Alder Riparian Woodland tween the two communities is low (30). Based on this Alnus rhombifolia is the dominant tree along much coefficient value, relatively few species occur in both of Sonoma Creek. Associated riparian species include communities; therefore, the total species richness in Acer macrophyllum, Artemisia douglasiana, Umbel­ the park could decline if oak woodland succeeded to lularia califomica, and Urtica dioica ssp. holosericea. mixed evergreen forest. Barnhart et al. ( 1996) also cor­ Rhagadiolus stellatus, a nonnative herb not reported roborate this conclusion, stating, " P. menziesii inva­ from elsewhere in Sonoma County, has established in sions will result in reduction of biological diversity in several locations within the riparian corridor (Best et this region of California." al. 1996). In the lower stretches of the Sonoma Creek, white alder riparian woodland intergrades with coast Mixed Evergreen Forest redwood forest. Mixed evergreen forest is dominated by Pseudot­ Oak Woodland suga menziesii, Quercus spp., and Umbellularia cali­ Oak woodlands within the park are highly variable. jomica. Total tree canopy cover is roughly 85 percent. Quercus agrifolia dominates a majority of the oak Common to occasional tree associates include Litho­ woodlands in the park. The understory is sparse except carpus densijlorus and Arbutus men~iesii. The herba­ in tree gaps where a variety of herbs grows, including ceous understory is highly variable and species rich. Lathyrus vestitus, Lotus scoparius, and Madia gracilis. Common fern and herb associates include Dryopteris Shade-tolerant species in this community include San­ arguta, Melica spp., Stachys ajugoides var. rigida, icula crassicaulis and Toxicodendron diversilobum. Sanicula crassicaulis, Toxicodendron diversilobum, Stands of Q. kelloggii occur in scattered locations in and Trientalis latifolia. Mixed evergreen forest is pri­ the park, frequently with a dense understory of Fes­ marily restricted to the north-facing slope of Sugarloaf tuca califomica. On the eastern side of Red Mountain, Ridge on Sonoma volcanics and the Lower Creta­ oak woodland is dominated by Q. chrysolepis and Q. ceous-Upper Jurassic Great Valley Sequence. The soil kelloggii with Umbellularia califomica and Acer ma­ type mapped for the area is Rock land which is de­ crophyllum interspersed. Quercus garryana is abun­ scribed as being, " in rough mountainous areas where dant from near the confluence of Rattlesnake Creek there is little soil material" (U. S. Department of Ag­ and Sonoma Creek upstream to approximately the con­ riculture 1972). The county soil survey describes the fluence of Malm Fork Sonoma Creek and Sonoma vegetation on this soil as sparse with occasional stunt­ Creek. ed trees, which is not representative of the area under In the Sonoma Mountains, Pseudotsuga mem.iesii is study. Much of this vegetation in the park grows near invading mature oak woodlands (Barnhart et al . 1996). the toe of Sugarloaf Ridge, possibly because of ero- VOLUME 18, NUMBER I Sugarloaf llidge State Park Flora 23

Table I. Numbers of families, genera, and species summarized for Sugarload Ridge State Park, California.

Number of Percent of Number of Percent of Number of Percent of Taxonomic group families total famitics genera total genera species total species

Ferns and fern aUies 7 9.2 12 4.8 14 3.5 Gymnosperms 3 3.9 4 1.6 4 l.O Angiosperms 66 86.8 236 93.7 382 95.5 Dicotyledons 57 75.0 190 75.4 299 74.8 Monocotyledons 8 10.5 46 18.3 83 20.8 Total 76 100% 252 100% 400 100% sional material which accumulates, allowing the veg­ 1993 through 1997, and included winter, spring, sum­ etation to develop more fully. mer, and fall trips. Nearly all the park's trails were surveyed on foot, a couple were surveyed from a ve­ Coast Redwood Forest hicle. Coast redwood forest is restricted to the more mesic portions of Adobe Canyon, along Sonoma Creek, at FLORA lower elevations where the creek has deeply incised Documented plants totalled 400 within Sugarloaf the canyon. This stand of Sequoia sempervirens rep­ Ridge State Park, 75 percent of which are native (Table resents one of the easternmost in the state; another 1). The three largest families in the park are Astera­ stand occurs near the small town of Angwin approxi­ ceae, Poaceae, and Fabaceae, in descending order. As­ mately 15 km (9 air miles) to the north-northeast, also teraceae and Poaceae are the two largest families in in Sonoma County. Within the park, coast redwood the state, so their prominence is unsurprising. The high forest is a riparian community. On average, the percent number of leguminous plants is largely due to the high cover of Sequoia sempervirens within this community diversity of Trifolium. With 15 taxa, it is the best rep­ in the park is 75 percent. Based on seven tree cores, resented genus in the park. Asteraceae, Poaceae, and the age of the older coast redwood trees is roughly Fabaceae combined represent 35% of the flora. Further 120 years, thus the trees were logged circa 1875 (Bow­ floristic work in the park will no doubt add species to cutt unpubl.). Evidence of stump sprouting from trees the list; however, the majority are represented in this cut during this time is common. Lithocarpus densiflo­ publication. rus is a frequent tree associate, although percent cover Collections made as a part of this study added four is low at approximately 5 percent. The herbaceous taxa to the known flora of Sonoma County: Carex am­ cover is sparse with low species richness. Herb and plifolia, ]uncus nevadensis, Montia parvifolia, and fern associates include Adenocaulon bicolor, Dryop­ Rhagadiolus stellatus. Carex amplifolia and R. stella­ teris arguta, and Polystichum munitum. Based on geo­ tus were included in the county flora based on unpub­ logic and soil maps, coast redwood forest in the park lished findings of this floristic research. ]uncus neva­ occurs on Sonoma volcanics (Wagner and Bortugno densis, and M. parvifolia represent additions not in­ 1982) in either Rock land or Laniger Loam (U. S. cluded in Best et al. (1996). In addition, this work Department of Agriculture 1972). provides vouchers for species listed in Best et al. (1996) without the benefit of a voucher specimen; in­ HISTORY OF BOTANICAL RESEARCH IN THE PARK cluded in this category are the newly introduced Tri­ folium striatum not included in Hickman' (1993) and Three unpublished vascular plant species lists exist Claytonia parviflora. for the park: DiTomaso (1996), Stocking (s.d.), and Wright (1975). Vouchers appeared to be lacking except MANAGEMENT RECOMMENDATIONS for a limited number from DiTomaso's list, which was based primarily on vegetation research in the grass­ California Department of Parks and Recreation lands. Several of his specimens added taxa to the park manages the park for the preservation of native plant flora. A floristic study of Sonoma County was recently biodiversity. Three aggressively invasive exotics, Cy­ completed (Best et al. 1996). Specimens collected in tisus scoparius, Foeniculum vulgare, and Genista the park by a variety of individuals are cited in the monspessulana, are currently rare in the park, bow­ county flora and included in this flora when they add ever, a vigilant manual removal. program is recom­ taxa. Whatford (1994) created a vegetation map for mended to keep these species from becoming manage­ Sugarloaf Ridge State Park. He focused on areas being ment problems in the future. Continued efforts to con­ invaded by Centaurea solstitialis and Pseudotsuga trol Centaurea solstitialis and feral pigs are encour­ menziesii. My floristic surveys were condu~ted from aged. Recreational use of horses in the park presents 24 Bowcutt ALISO a means for introduction of nonnative species (e.g., ifomia (DAV) unless otherwise noted. Voucher spec­ Trifolium striatum). Given the recreational value of imens exist for 97% of the taxa represented in this horseback riding, 1 do not recommend prohibiting their floristic study. Nomenclature follows Hickman (1993). use. However, attention should be given to controlling Common names were gleaned from several sources: new introductions to keep them from becoming inva­ Abrams (1940; 1944; and 1951), Abrams and Ferris sive problems. This would include frequently moni­ (1960). Best et al. (1996), Hickman (1993), Mason toring trails used by the horses and around the corral. (1957), and Munz and Keck (1973). Herbarium abbre­ Future acquisitions are desired to connect, at least viations follow Holmgren et al. (1990). In addition the with trail corridors, several local parks to Sugarloaf following notations were used: * for nonnative species Ridge State Park including Annadel State Park, Bothe and + for unnaturalized garden rel icts not included in Napa State Park, and Jack London State Park (Bud the numerical analyses. Getty, pers. comm.). Sugarloaf Ridge State Park is al­ ready contiguous with Hood Mountain Regional Park, PTEROPHYTA--Ferns which is 608 hectares (1500 acres). RL.ECifNACEAE WooowARDIA 1-lMBRIATA Sm. in Recs.-Chain fern. Rare. Spring in FUTURE RESEARCH NEEDS oak woodland along Vista Trail. 8owcu11 1892. A thorough survey of 410 hectares (lOll acres) DENNSTAEOTIACEAE added to the park in 1997 was beyond the scope of this effort and is worthy of future attention. With this PraRIDIVM AQUIL!N\JM (L.) Kuhn var. PUBESCENS L. Underw.-Brack­ en fern. Common. Coast redwood f orest and mixed evergreen addition, the park's total area is 1552 hectares (3831 forest. Lnredo-Prendeville & Canon 59 ( DA V, NCC). acres). Voucherless taxa listed in the county flora as being from "Sugarloaf Ridge" are not included in this DR YOPTERIDACEAE study. Neither are voucherless taxa on the three un­ DRYOPTERIS ARGUTA (Kaulf.) Maxon-CoastaJ wood fern. Common. published checklists for the park (DiTomaso 1996; Coast redwood forest and mixed evergreen forest. Bowcuu. Ken­ Stocking s.d.; and Wright 1975). Voucher specimens nedy & Henricks 1754. are needed to confirm the presence of these and other POL.YSTICHUM MUNITUM (Kaulf) C. Presi-Sword fern. Common. taxa not included in this study. Coast redwood forest and ecotone between oak woodland and white alder riparian woodland. Botany /02 students /03 (DAV, NCC). ACKNOWLEDGMENTS POL YPODIACEAE I received much research support for which I am POLYPODIUM CALIFORNICUM Kaulf.-California polypody. Occasion­ grateful. Kate Mawdsley made significant contribu­ al. Mixed evergreen forest. Bowcurr 1755, Bowcutr 2145. tions by assisting with collecting, identification, and POLYPODIVM OLYCYRIUIIZA D . Eaton-Licorice fern. Occasional. manuscript editing. Bud Getty of the California De­ Mixed evergreen forest. Bowcu/1 1875, Lnredo-Prendeville & partment of Parks and Recreation and his staff provid­ Ctmon 57 (DAV, NCC). ed logistical support. My students in California Flo­ PTI!RIDACEAE ristics at the University of California at Davis collected specimens in spring 1993 and 1994. My students in ADIANTUM JORDANII C. Mueller-California maidenhair. Occasional. Mixed evergreen forest. Lnredo-Prendevilk & Canon 56 ( DA V. Ecology Field Quarter at University of California at NCC), Botany 102 studenJs 200. Santa Cruz helped to collect vegetation data in the park AsPIDOTIS DENSA (Brackenr.) Lellinger-Inclian's dream. Occasional. in May 1996. Dr. Ellen Dean and Jean Shepard gra­ Serpentine chaparral. Bowcu11 /939. ciously provided space to work at the University of CIIEU.ANTHES INTERTEXTA (Maxon) Maxon-Coastal lip fern. Rare. California Davis Herbarium. They also identified sev­ Rock outcrop in grassland along V ista Trail. Bowcutt /897b, Bow· CUlt 2082. Dr. eral grasses joyfully. Michael Vincent identified PELI..AEA ANDROMEDIFOLIA (Kaulf) Foo-Coffec fern. Occasional. the Trifolium striatum. Dr. Ann Howald gave a phone Rock outcrop in mixed evergreen forest. BowcuJt 1870 ( DAV. consultation. Karen Knoll provided field assistance in NCC), Bowcutt & Mawdsley 208/. Bury et al. 295 (DA V, NCC). July 1997. The Davis Herbaria Society helped defray PENTAORAMMA TRIA.'IGULARIS (Kaulf) G. Yatskievych, M.D. Wind· the cost of hiring a cartographer and paid for some ham & E. Wollenweber--Golden-back fern. Occasional. Mixed travel expenses. Dr. R. K. Benjamin and two anony­ evergreen forest. Lnredo-PrendeviUe & Canon 58 (DA V, NCC) mous reviewers contributed helpful criticisms on an SPHENOPHYT A--Horsetails earlier draft. EQUISET"ACEAE ANNOTATED CHECKLJST OF THE VASCULAR PLANTS EQUISE1"UM LAEVIGATUM A . Braun-smooth scouring rush. Occa­ sional. White alder riparian woodland. Bowcutr 1894, Lewis & Voucher specimens are housed in the University of Mutch 16, Lee 105. California Davis Herbarium at the University of Cal- EQIJISETUM TEI..EMATEIA Ehrb. ssp. BRAUNIJ (Milde) R. L. Hauk e- VOLUME 18, NUMBER 1 Sugarloaf Ridge State Park Flora 25

Giant horsetail. Occasional. White alder riparian woodland along chaparral. Bury & Her<.og 262 (DAV, NCC), Bowcutt & Keeney Sonoma Creek. Botany 102 students 80, Bowcutt & Knoll 2180. 1805, Bowcutt, Keeney, & Green 1806(DAV, NCC). LOMATIUM RF.POSTUM (Jepson) Mathias-Napa lomatium. Rare. LYCOPHYTA--Club-mosses Charnise chaparral. CNPS List4. Bowcutt /922. LOMATIUM UTRICULATUM (Torrey & A. Gray) J. Coulter & Rose­ SELAGINELLACEAE Common lomatium. Occasional. Annual grassland. North 33, Bury & Her<.og 261. SELAGINELLA WALLACEI Hieron.-Little club-moss, spike-moss. Rare. OENANTHE SARMENTOSA J. S. Presl-Pacific oenanthe. Occasional. Rock outcrops. Bowcutt 1897a. Coast redwood forest along Sonoma Creek. Bowcutt 1941. OSMORHIZA CHlLENSIS Hook. & Arn.-Mountain sweet-cicely. Com­ CON!FEROPHYTA--Conifers mon. Mixed evergreen fores1. Botany 102 students 29, Botany /02 studems 83, Adam & Weick 296. CUPRF.SSACEAE PERIDERIDIA KELLOGGU (A. Gray) Mathias- Kellogg's yampah. CUPRESSUS SARGE.NTII Jepson-Sargent cypress. Rare. Serpentine Grassland along Vista Trail. Bowcutt, Mawdsley, & Knoll 2189, chaparral. Bowcutt, Mawdsley, & Knoll 2187. Bowcutt, Mawdsley. & Knoll 2192. SANICULA BlPINNATlFlDA Hook. & Arn.-Purple sanicle, poison san­ Pl.NACEAE icle. Rare. Grassy openings in mixed evergreen forest and coast live oak woodland. Lewis & Mutch 1. PINUS SABI.NIANA Douglas-Gray pine. Locally common. Charnise SANICULA CRASSICAULIS DC.-Pacific sanicle, gamble weed. Com­ chaparral along Grdy Pine trail. 1916. mon. Mixed evergreen forest and white alder riparian woodland. PSEUDOTSUGA MENZIESO (Mirbel) Franco var. MSNZ!ESII-Douglas-fir. Lewis & Mutch s.n .. Common. Mixed evergreen forest. Loredo-Prendevil/e & Canon SANICULA LACtNrATA Hook. & Am.--Coast sanicle. Reported by 55, Adams et al. 260 ( DA V, NCC). Stocking (s.d.) and Wright ( 1975). Collected in Adobe Canyon and several other inland Sonoma County locations (Best et al. TAXODIACEAE 1996). Constance ( 1993) reports this species as occurring only SEQUOIA SEMPERVIRENS (D. Don) Endi.--Coast redwood. Very Jo­ along the north and central coast. ca:lized. Coast redwood forest, mixed evergreen forest. Loredo­ •So.NDIX PECTIN-VENERIS L.-Shepherd's needle, Venus' needle. Prendeville & Canon 54. Common. Oak woodland. Native to Mediterranean. Lewis & Mutch 14, Botany 102 students 88. *TORILIS ARVENSIS (Hudson) Link-Hedge-parsley. Common. Coast ANTHOPHYTA- Flowering plants/DICOTYLENDONEAE live oak woodland and mixed evergreen forest. Native to central and southern Europe. Bowcutt 1862. ACERACEAE. ACER MACROPHYLLUM Pursh-Big-leaf maple. Common. Mixed ev­ ARALIACEAE ergreen forest. Loredo-Prendeville & Canon 53 ( DA V, NCC,), ARALIA CALIFORNICA S. Watson-California spikenard, elk's clover. Botany 102 studems 290 (DA V, NCC). Occasional. Along Sonoma Creek in mixed evergreen forest. Bowcutt 1900 (DAV, NCC). ANACAROIACEAE

TOXICODENDRON DIVERSILOBUM (Torrey & A. Gray) E. Greene-Poi­ ARISTOLOCHIACEAE son-oak. Common. Coast redwood forest, oak woodland, and ARISTOLOCfllA CALIFORNICA Torrey-Pipevine. Occasional. Coast mixed evergreen forest. Repeatedly observed and avoided by the redwood forest and mixed evergreen forest. Collected near con­ author. fluence of Bear and Sonoma Creeks. BowcuJt, Kennedy, & Hen­ ricks 1762. APlACEAE ASTERACEAE. ANGELICA TOMENTOSA S. Watson-Angelica. Rare. Bank of ephem­ eral creek. Bowcutt 2204. AcmLLBA I\.1ILLEFOUUM L.-Yarrow. Common. Annual grassland, *ANTHRISCUS CAUCALIS M. Bieb.- Bur-cbervil. Rare. Recently oak woodland. and serpentine chaparral. North 52, Togioka and burned coast live oak woodland, grassland, and other disturbed Waraiclz 236. areas e.g., roadsides. From Eurasia. Bowcutt 1859, Bowcuu ACHYRACHAENA MOLLIS Schauer-Biow-wives. Oc<;asional. Annual 2137a. grassland. Bowcutt 2153b. APIASTRUM ANGUSTI'FOLfUM Nutt,-Wild celery. Rare. Recently cut ADENOCAULON BICOLOR Hook.-lndian guide, silver arrow. Com­ fire break in charnise chaparral. Bowcutt 2126. mon. Coast redwood forest and mixed evergreen forest. Bowcutt *CONIUM MACULATIJM L.-Poison hemlock. Occasional. Annual 1873. grassland. From Europe. Bowcutt /876. AGOSERIS APARGIOIDES (Less.) E. Greene var. APARGIOIDES-Agoser­ *FOENICULUM VULGARE Miller-Fennel. Rare. One plant along road­ is. Occasional. Oak woodland. North 53. side half way up Bald Mountain Road. Two plants along road to AGOSERIS GRANDIFLORA (Nutt.) E. Greene-Large-flowered agoseris. Bear Creek Ranch. Park staff has been working to prevent its Occasional. Coast live oak woodland. Bowcuu 1866. spread (Bud Getty, pees. comm.). Native to southern Europe. AGOSERIS HETEROPHYLLA (Nutt.) E. Greene- Annual agoseris. Rare. Bowcutt 2129. Rocky opening in serpentine chaparral. Bowcutt 1804. HERACLEUM LANATIJM Michaux--Cow parsnip. Occasional. White AGOSERIS R.ETRORSA (Benth.) E. Greene-Spear-leaved agoseris. Oc­ alder riparian woodland. Bowcutt & Green 1826 (DAV, NCC). casional. Mixed evergreen forest. North 54. LOMATfUM CALIFORNICUM (Torrey & A. Gray) Mathias & Con­ ANAPHALIS MARGARITACEA (L.) Benth. & Hook.-Pearly everlasting. stance-Lomatium. Rare. Annual grassland. Bowcutt 2154 (DAV, Occasional. Grasslands. Bowcutt 1905. NCC). * A.NTHEMIS COTIJLA L.-Mayweed. Rare. Grassy opening in charnise LoMAHUM DASYCARPUM (Torrey & A. Gray) J. Coulter & Rose var. chaparral. Native to Europe. Observed in flower by K. Mawdsley. DASYCARPUM-Woolly-fruited lomatium. Occasionl!l. Serpentine Bowcutt 1890. 26 Bowcutt ALISO

ARTEMISIA DOUOLASIANA Besser- Mugwort, wormwood. Abundant. MADIA ORACn.tS (Smith) Keck-Siender tarweed. Common. C oa~ t White alder riparian woodland. Bowcutt 1902. live oak woodland. Bowcutt 2160, Bowcutt & Kennedy 1832. BACCHARIS Pfl..VLAR!S DC.---Coyote brush. Locally common. Invad­ MADIA MADIOlt>E.S (Nutt.) E. Greene- Woodland madia. Common. ing annual grasslands in upper reaches of Adobe Canyon floor. Mixed evergreen forest. Botany 102 students 6, Botany 102 stu­ Bowcutt 2132. dents 48, Botany 102 students 205, Bowcutt, Kennedy. & Hen­ CAL.YCADENlA TRUNCATA DC.-Rosin weed. Common. Annual grass­ ricks 1758 (DAV. NCC). land. &wcutt, Mawd.fley, &: Knoll 2190. MADIA NUTANS (E. Greene) Keck-Nodding madia. Grassland.. Re­ *CAROUVS PYCNOCEPHAI..US L.-Italian thistle. AbundanL Annual ported by DiTomaso and Wright. Also collected in Adobe Canyon grassland and oak woodland. Native to Mediterranean. Botany and on Mt. Hood (Best et al. 1996). 102 students 43, Bowcmt, Henricks. &: Kennedy 1771. MADIA SATIVA Molina-Coast tarweed. Rare. Grassland. Bowcutt. *CENTAUReA MELITENSIS L.-Tocalote. OccasionaL Grassy opening Mawdsley, &: Knoll 2184, Bowcutt, Mawdsley, &: Knoll 2199. in chamise chaparral. Native to southern Europe. Bowcutt 1891. MALACOTHRJX FLOCCIFCRA (DC.) S. F. Blake-Wooly malacothrix. Bowcutt 2136 (DAV, NCC). Locally common. Serpentine chaparral. Bawcutt, Kennedy, &: *CENTAURI!A SOLSTITtALIS L.-Yellow star-thistle. AbundanL Annual Henricks 1750. grassland. Native to southern Europe. North 28. PS!LOCARPHUS TENELLUS Nun. var. TENELLUS-Woolly-marbles. Oc­ *CHAMOMILLA SUAVEOLE.NS (Pursh) Rydb.-Pineapple weed. Abun­ casional. Ruderal. Bowcutt 2142b. dant. Disturbed places. Native to northwestern North America and *RHAOADIOLUS STELLAnJS (L.) Gaertner. Locally abundant. Road­ northeastern Asia. Lewis & Mutch 9, Botany 102 students 23. sides in white alder riparian woodland. Native to Europe. Lewis *CICHORlUM INTYBUS L.---Chicory. Rare. Roadside in campground. & Mutch 25. Bowcutt, Henricks. & Kennedy 1778 ( DA V. NCC). Native to Europe. Bowcutt 2200. *SENECIO VULGARIS L.-Groundsel. Rare. Annual grassland and C IRSIUM OCCIDENTALE (Nun.) Jepson var. VENVSTUM (E. Greene) Jep­ opening in coast live oak woodland. Native to Eurasia. Bowcuu son- Venus thistle. Rare. Opening in mixed evergreen forest. 2077 (DA V. NCC). Bowcutt 1906. *Sa .. YBUM MARIANUM (L.) Gaertner- Milk thistle. Occasional. An­ *CJRSIUM VULGARE (Savi) Ten.-Bull thistle. Occasional. White al­ nual grassland and disturbed places. Native to the Mediterranean. der riparian woodland. Native to Europe. Bowcutt 1928. North 55. SOLIDAGO CALlFORNICA Nutt.--Califomia goldenrod. Rare. Along CONYZA CANADENSIS (L.) Cronq.-Horseweed. Opening in mixed ev­ ephemerally moist ditch in grassland on edge of riparian. Bow­ ergreen forest. Bowcutt 2064 ( DA V, NCC). cuff. Mawdsley, & Knoll 2197. ERIGERON BIOLETTII E. Greene-Bioletti's fleabane daisy. Collected •SONC1iUS ASPER (L.) HiJJ SSp. ASPER-Prick.ly SOW l.bistle. Rare. in Adobe Canyon and on Ml. Hood (Best et al. 1996). Opening in mixed evergreen forest. Native to Europe. Bowcutt ERJOE.RON l'Hn..ADELPHICUS L.-Philadelphia daisy. G rowing with 2171 (DAV, NCC), Bowcutt, Henricks, & Kem1edy 1772. Woodwardia jimbriata at a seep in oak woodland along Vista *SONCHUS OLERACEUS L.---Common sow thistle. Common. Dis­ Trail. Mawds/ey 43. turbed places. North 7. ERIOPHYLLUM CONFERTIFLORUM (DC.) A. Gray var. CONF'ERTTFLO­ STEPHANOMERJA EXIGUA NulL ssp. CORONARJA (E. Greene) Gottlieb-­ RUM-Golden-yarrow. Rare. Roadcuts in chamise chaparral. Bow­ Stephanomeria. Roadcut in coast live oak woodland and chamise 1919. 2128. cult Bowcuu chaparral trailside. Bowcutt 2062 (DAV, NCC). ERJOPHYLLUM LANA'TUM (Pursh) James Forbes var. ACHIU.AEOlDE.S .-rARAXACUM OFFIClNALE Wigg.-Dandelion. Rare. Mixed eve.rgreen (DC.) Jepson-Woolly sunflower. Common. Serpentine chaparral. forest. Native to Europe. Bowcutt, Henricks, &: Kennedy 1783. 2169, 45. Bowcutt Botany 102 students *TRAOOPOOON PORRJFOLrus L.--Salsify. Rare. Disturbed places. Na­ Fo.Aoo CAUFORNICA NutL-Herba impia. Common. Serpentine tive to Europe. North 36. chaparral Bowcwt, Keeney, & Green 1816. UROPAPPUS UNt>LEYI (DC.) Nutt.-Silver puffs. Occasional. Rocky *FJLAOO GALLICA L.- Filago. Common. Annual grassland. Native to opening in serpentine chaparral. Bowcutt 1820. Mediterranean. Bowcutt 1855. WYETHJA ANOUSTIFOUA (DC.) NulL-Narrow-leaved mule-ears. Oc· HELENIUM PUBERULUM DC.--Sneezeweed. Rare. White alder riparian casional. Oak scrub along Brushy Peak trail. BowG·utt 1828, Bow­ woodland. Bowcutt /930. cult 1925. HEMIZON'IA CONGE.STA DC. ssp. LUZUt.UFOUA (DC.) Babe. & H.M . WYETHIA GLABRA A. Gray-Mules ears. Rare. Oak woodland. Bow­ Hall- Hayfield tarweed. Common. Grassy opening in coast live cult 2130. oak woodland and serpentine chaparral. Bowcwt 1889. Bowcuu /938, Bowcuu 2060. BETULACE.AE. HIERACIUM ALB!Ft.ORUM Hook.-White-ftowered hawkweed. Occa­ sional. Mixed evergreen forest. North 49. ALNUS RHOMBIFOLIA NutL-Wrute alder. Locally dominant. White *HYPOCHOE.RJS OLABRA L.--Smooth cat's-ear. Common. Annual alder riparian woodland along Sonoma Creek. Bowcutt, Henricks. grassland. Native to Europe. North 26. & Kennedy 1773. *HYPOCHOERJS RADICATA L.-Rough cat's-ear. Common. Annual CORYLUS CORNUTA Marsh var. CALIFORNICA (A. DC.) W. Sharp-­ grassland. Native to Europe. North 41. HazelnuL Rare. Coa~t redwood forest. mixed evergreen forest. *LACTIJCA SERRIOLA L.-Wlld lettuce, prickly lettuce. Rare. Ob­ and white alder riparian woodland. Bowcutt, Henricks, &: Ken­ nedy 1767. served by the author in disturbed area along Gray Pine trail near junction with Vista trail. Native to Europe. LASTHENIA CALIFORNJCA Lindley--Goldfields. Locally common. Ser­ BORAGINACEAE pentine chaparral. North 30. AMSINCKIA MENZJESII (Lehm.) Nelson & J. F. Macbr. var. INI'ERMEOIA L ESS INOlA RAMVI.OSA A. Gray- Sonoma lessingia. Locally common. (Fischer & C. Meyer) Ganders-Fiddleneck. r.mcher's fire-weed. Serpentine chaparral. Bowcutt /937. Occasional. Open oak woodland. Botany 102 students 32. MADIA ELEGANS Lindley S.~. VERNALIS Keck.-Common madia. Rare CRYPTANTHA MVRICATA (Hook. & Am.) Nelson & J. F. Macbr.­ in park. Sunny opening in mixed evergreen forest. Bowcurt 1869, Jones' cryptantha. Collected in the park by Baker (BeSt et al. Bowcutt 2205. 1996). MADIA EXIOUA (Smith) A. Gray-ThreadStem madia. Common. Ser­ CYNOGLOSSUM GRANDE Lehm.- Hound's tongue. Rare. Oak wood­ pentine chaparral. Bowcutt 1819, Bowcutt 1885. land. Bowcutr 2090, Botany 102 studems 86. VOLUME 18, NUMBER I Sugarloaf Ridge State Park Flora 27

•Mvosons DISCOLOR Pers.-Forget-me·noL Occasional. Milled ev­ CARYOPHYLLACEAE ergreen forest Native to Europe. Bowcutt, Henricks, &: Kennedy 1795. "'C'ERAmUM GU>MERATIJM Thuiii.-Mouse-ear chickweed. Qcca. sional. Annual gra»land. Native to Europe. Lewis &: Mutch 13, Pl.AGIOBOTMRYS NOTHOFUt.VUS (A. Gray) A. Gray-Popcorn flower. Woroich 229, Daehler &: Lu 255. Occasional. Annual grassland. Botany /02 students 101. Botany 102 students 279 (DAV. NCC). MtNUARTIA DOUGI..ASII (Torrey & A. Gray) Mattf.-Douglas' sand­ won. Occasional. Serpentine chaparral. &wcuu &: Kennedy 181 I. BRASSICACEAE •PETRORHAGIA NANTEUIUI (Bernat) F. Ball & Heyw.-Proliferous AnfYSANUS PUSIU.US (Hook.) E. Greene--Dwarf athysanus. Rare. pink. Common. Annual grassland. opening in mixed evergreen Annual grassland. Bowcutt & Mawdsley 2085. forest. and roadsides. Native to southwestern Europe. Loredo­ BAR.BAREA ORTHOCERAS Ledeb.-American winter cress. Rare. An­ Prendeville & Canon 67 (DA V. NCC). Botany 102 students 224, nual grassland in campground. Bowcutt &: Mawdsley 2068 ( DAV. Botony 102 .ftudents 285. NCC). Srt.ENE CALIFOilNICA Durand- Indian pinlc. Occasional. Recently *CAPSEU..A BURSA PASTORIS (L.) Medikus-Shepberd's purse. Occa­ burned coast live oak woodland. Bowcull 1858. sional. Annual grassland. Native to Europe. Bowcutt 2070. •StLENE OALt.ICA L.-Common catchfly. Common. Annual grass­ CARDAMIN!! CALIFORNJCA (Torrey & A . Gray) E. Greene- Milk land, serpentine chaparral, and roadsides. Native to E urope. 18/4 maids, tooth wort. Occasional. Oak woodland. Bowcull 2089. (DAV, NCC). Waraich & Togioktl 240. Bullock & Beever 241. CARDAMTN£ OLIGOSPERMA Torrey & A. Gray-Bitter-cress. Rare. Botany 102 students 280 (DAV, NCC). Mixed evergreen forest. Bowcutt I 757. "'SPBRGULA ARVBNS IS l. ssp. ARVENSIS-Stickwon, starwort. Recently GUILLENIA l.ASIOPIIYU.A (Hook. & Am.) E. Greene-California mus­ burned coast live oak woodland. Native to Europe. Bowcutt /860. tard. Disturbed places. Reported by DiTomaso (1996), Stocking *SPBRGUI..ARIA RU9RA (L.) J. S. Prest & C. Presi-Sand-spurrey. (s.d.), and Wright (1975). Collected nearby in burned chaparral Common. Growing in middle of trail in annual grassland. Native on Mt. Hood. Baker I 1697 (NCC). to Europe. Bowculf & Mawdl·ley 2076. •HrRscHI-eu>tA JNCANA (L.) Lagr.-Fossat- Hedge mustard. Occasion­ •STEU.ARIA MEDIA ( L .) Vi liars-Common chickweed. Common. Oak al. Recently burned coast live oak woodland and grassland. Na­ woodland and disturbed places. Native to southwestern Europe. tive to Mediterranean. &wcutt 1857, Bullock & Beever 206. Botany 102 students 82, &wcrm & Mawdsley 2066b. LEPIDrUM STRICTVM (S. Watson) Rattan-shaq>-podded pepper-grass. STEI..t.ARtA -m-ENS Nutt.-Shining chickweed. Rare and inconspicu­ Locally abundant. Along trails in annual grasslands and roadsides. ous. Disturbed places. DiTomaso & Kaiser s.n. (private collec­ BowcuJt 184Ju. Bo'K.'CUII 2074, Lewis & Mutch 12. Adams et al. rion). Also reponed by Stocking (s.d.). 250 (DAV, NCC). •ltJNAJUA AI\NUA L.-Money plant. moonworL Rare. Mixed ever­ CISTACEA£ green forest along Adobe Canyon Road. Native to Europe. Bow­ HELIANTHEMUM SCOPAR ICM Nutt.-Pealc rush-rose. Occasional. cull & Grun 1797 (DAV, NCC). Charruse chaparral along Gray Pine trail. Bowcutt 1917. ROIUPI>A NAsnJRtUM·AOUATICUM (L.) Hayek-Water cress. Rare. Growing in seepy areas. Bowcutt, Kennedy, & Henricks 1786. CONVOLVULACEAE STR.EPTANTHUS GLANOUI..OSUS Hook. ssp. GLANDUU>SUs-Jewelflower. Rare. Serpentine grassland. Bury & Herzog 289. CALYS1COIA COLLtNA (E. Greene) Brurnmitt-Moming-glory. Rare. THYSANOCARPUS CURVIPES Hook.-Lacepod. fringepod. Occasional. Serpentine chaparral. Beever 213. Bury&: Herzog 275. Grassland. Observed by the author. Reported by DiTomaso CALYS1'FOJA occtDENTAUS (A. Gray) Brummin ssp. OCCIDENTAL.Js­ (1996), Stocking (s.d.), and Wright (1975). Collected "probably Morning-glory. Occasional. Annual grassland. Bowcutt 2163 in the vicinity of MI. Hood'' (Best et al. 1996). (DAV, NCC). CALYS1COIA PURPURATA (E. Greene) Brummitt ssp. PURPURATA­ Moming-glory. Occasional. Serpentine chaparral and white alder CALYCANTHACBAE riparian woodland. Bowcutt /749, Bowcutt /932. CAL. YCANTIIUS OCCIDF.NTAUS Hook. & Am.-Spicebush, sweet shrub. *CONVOLVULUS ARVf.NSIS L.-Bindweed. Occasional. Disturbed Oocasional. Coast redwood forest and mixed evergreen forest at places, e.g.. in campground and around the horse corrals. Native confluence of Sonoma and Bear Creeks. Bowcun 1764 (DAV, to Europe. Bowcutt, Mawdsley. & Knoll 2198. NCC). CRASSULACEA£ CAMPANUL.ACEA£ DUDLEYA CYMOSA (Lemaire) Britton & Rose-Live-forever. Occa­ sional. Rock outcrop in mixed evergreen forest. Bowcutt /877. liETEROCOOON RARJFLORUM Nun.-Heterocodon. Rare. Vernally wet places. DrTomoso & Kauer s.n (private coflecrion). CUCURBIT ACEA£ TluOOANIS BJFU)RA (Ruiz lopez & Pavon) E. Green~Venus look­ ing-glass. Rare Dasturbcd sites. DrTomoso & Kaiser s.n. (private MARAH FABACEUS (Naudtn) E. Greene va:r. AGRES11S (E. Greene) K. collection) M . Stockang-Callfomaa man-root. OccasionaL Opening in maxed evergreen forest. Bowcutt 1775. CAPRIFOUACEAE CUSClTI'ACEAE lONtCERA IIISPIDUL.A Douglas var. VACIU.ANS A. Gray-Honeysuckle. Common. Mixed evergreen forest. seep in oak woodland. and CUSClTI'A CALJfORNICA Hook. & Am. var. CAUFORi'HCA-Dodder. serpentine chaparral. Bowcutt. Henriclcs, & Kennedy 1748. Rare. In grasslands. para~itic on Hemizonia congesta. Bowcutt &: SAMBUCUS MF.XICANA C. Presi- Biue elderberry. Occasional. Annual Mawdsley 2193. grassland. Bowcutt 1822. ERICACEA!l SYMPHORICARPUS ALBUS (L.) S. F. Blake V3I. LAEVIGATUS (Fern.) S. F. Blake- Snowberry. Occasional. Mixed evergreen forest. Bow­ ARBUTUS MENt.tllSII Pursh- Pacific madrone. Common. Mixed ev­ cull, Kennedy, & Henricks 1763. ergreen loresr. Bury et a/. 266 ( DA V, NCC). 28 Bowcutt ALISO

ARCfOSTAPHYLOS CANESCENS Eastw. SSp. CANESCENS-Hoary man­ RUPERTIA PHYSODES (Hook.) Grimes--California-tea, Rupert's scurf­ zanita. Occasional. Chamise chaparral. Bowcutt 2135. pea. Rare. Along a small tributary to Sonoma Creek near group ARCTOSTAPHYtOS MANZANITA C. Parry ssp. MANZANITA- Manzanita. camp in riparian vegetation on edge of mixed evergreen forest. Common. Charnise chaparral, serpentine chaparral, and oak Bowcutt 1903. woodland. Bowcutt 2065 (DAV & NCC), Lee 23, Botany 102 THERMOPSIS MACROPHYLLA Hook. & Am. var. MACROPHYLLA- False­ students 97. lupine. Occasional. Along seasonal creek on edge of grassland. ARCfOSTAPHYLOS VISCIDA Parry ssp. PULCHELLA (J. Howell) Wells­ Bowcutt 1821, Botany 102 smdents 102, Bury & Herzog 273. White-leaved manzanita. Serpentine chaparral. Collected in Son­ TRIFOLIUM ALBOPURPUREUM Torrey & A. Gray var. DICHOTOMUM oma Creek Canyon near Mt. Hood (Best et al. I 996). Also re­ (Hook. & Am.) lsely--Common Indian clover. Occasional. ported by Stocking (s.d.) and Wright ( 1975). Grassland, rocky slopes, and serpentine. Botany 102 sllldents 242a. EUPHORBICEAE TRIFOLIUM BIFIDUM A. Gray var. DEC!PLENS E. Greene-Notch-leaved clover. Occasional. Grassy opening in oak woodland. Lee 91. EREMOCARPUS SETIGER (Hook.) Benth.-Doveweed, turkey mullein. *TRIFOLIUM CAMPESTRE Schreber- Hop clover. Occasional. Ruderal, Occasional. Annual grassland. Bowcutt 1886. along Meadow Trail. Native to Europe. Mawdsley 45. TR!FOLfUM CfLIOLATUM Benth.-Tree clover. Occasional. Opening in FABACEAE mixed evergreen forest. Loredo-Prendeville & Canon 65, Beever ASTRAGALUS GAMBELIA.NUS E. Sheldon-Dwarf locoweed. Occasion­ 230, Bullock & Togioka 242b (DAV, NCC). al. Annual grassland. Collected in Adobe Canyon (Best et aL TRIFOLIUM DEPAUPERATUM Desv. var. DEPAUPERATUM-Dwarf sack 1996). Also reported by DiTomaso (1996) and Wright (1975). clover. Occasional. Annual grasslands, grassy openings in oak *Cvnsus SCOPARIUS (L.) Link-Scotch broom. Rare. Mixed ever­ woodland and mixed evergreen forest. Bowcutt 2075, Lewis & green forest. Native to southern Europe and northern Africa. Lo­ Mutch 4. redo-Prendeville & Canon 62. *TRIFOLOJM DUBIUM Sibth.-Shamrock. Common. Annual grassland. *GE.NISTA MONSPESSULANA (L.) L. Johnson-French broom. Rare. Native to Europe. Lewis & Mutch 8. Six foot tall, single plant found along Adobe Canyon Road just *TRIFOLIUM FRAGIFERUM L.-Strawberry clover. Roadside. Native to east of Goodspeed Trail head. Native to Mediterranean. Bowcutt Europe. Bowcutt & Knoll 2186. 2093 (DAV, NCC). "TRIFOLIUM HtRTUM AIL-Rose clover. Occasional. Grasslands and "LATHYRUS CICERA L.-Red pea. Common. Annual grassland, open roadcuts in serpentine. Native to Eurasia. Bowcutt 1854, Bowcutt oak woodland, opening in mixed evergreen forest, and disturbed 2/22, Bury & Herzog 287, Bowcutt & Kennedy 1823. places. Native to Europe. Bowcutt 1843, Botany 102 students 42, TR!FOUUM MICROCEPHALUM Pursh-Small-headed clover, maiden Botany 102 students 89, Daehler & Lu 252 (DAV, NCC). clover. Occasional. Opening in mixed evergreen forest. Loredo­ *LATHYRUS HIRSUTUS L.--Caley pea. Rare. Annual grassland. Native Prendeville & Canon 63. to Europe. Bowcutt 2164. TRIFOLOJM MICRODON Hook. & Am.-Valparaiso clover, square-head LATHYRUS VESTITUS Nutt. var. OCHROPETALUS (Piper) lsely--Com­ clover. Common. Opening in oak woodland. Lee 90. mon Pacific pea. Common. Oak woodland and white alder ripar­ TRIFOLIUM OLIGANTHUM Steudei-Few-ftowered clover. Common. ian woodland. Lee 85, Beever et at. 233 (DAV, NCC). Annual grassland. Botany 102 students 98. LoTUS HUMISTRATUS E. Greene-Short-podded lotus. Common. An­ *TRIFOLIUM STRIATUM L.--Ciover. Rare. Near horse corral in dis­ nual grassland and serpentine chaparral. Bowcutt 1913, Bowcutt turbed area. Native to Europe. This recently introduced taxon is 2084, Bullock et al. 207. not reported by Isley (1993), however, Best et al. (1996) include LOTUS MICRANTHUS Benth.-Small-flowered lotus. Occasional. it in their Sonoma County flora without a voucher. Bowcuu 2141. Grassy opening in oak woodland. Botany 102 students 93, Bow­ *TRIFOLIUM SUBTERRANEUM L.-Subterranean clover. Occasional. cult 1861. Disturbed places. Native to southern Europe and northern Africa. LoTUS PURSHIANUS (Benth.) Clements & E. G. Clements var. PUR­ Botany 102 students 271. SHIA.NUS-Spanish clover. Common. Annual and perennial grass­ TRIFOLOJM VARIEGATUM Nutt.-White-tipped clover. Occasional. lands. Bowcutt & Green 1801. LoTUs SCOPARrus (Nutt.) Ottley-Deerweed. Occasional. Grassy Seasonal drainage. Bury eta/. 277 (DAV, NCC). opening in mixed evergreen forest, sun flecks in oak woodland, TRIFOLIUM WILLDENOV!l Sprengel-Tomcat clover. Occasional. roadcuts. Bowcutt 1871. Grassland. Adamr et al. 272. LoTUs WRANGELIANUS Fischer & C. Meyer-California lotus. Oc­ *VICIA LUTEA L.- Yellow vetch. Rare. Annual grassland. Native to casionaL Open oak woodland and disturbed places. Botany 102 Europe. Bowcutt 1849, Bowcutt 2152. students 31. *VICIA SATIVA L. ssp. SATIVA-Narrow-leaved vetch. Common. LUPINUS ALBlFRONS Benth. var. ALBIFRONs-Silver lupine. Common. Opening in oak woodland. Native to Europe. Botany 102 students Grassy opening in mixed evergreen forest and oak woodland. 79 (DAV, NCC). Botany 102 student.~ 104, Bowcutt 2158 (DA V, NCC). *VICIA VILLOSA Roth ssp. VARIA (Host) Corbiere-Vetch. Common. LUPINUS BICOLOR Lindley-Lindley's annual lupine, miniature lu­ Disturbed places. Native to Europe. Bullock & Beever 221 (DAV, pine. Common. Open oak woodland. Botany 102 students 37. NCC). Lewis & Mutch 15. LUPINUS LATIFOL!US J. Agardh var. LATIFOLIUS-Lupine. Oak wood­ land. Botany 102 students 95 (DAV, NCC). FAGACEAE LUPINUS NANUS Benth.-Douglas' annual lupine. Common. Annual grassland. Bowcutt 2083, Lewis & Mutch 2, Botany 102 students LITHOCARPUS DENSIFLORUS (Hook. & Am.) Rehder- Tan oak. Com­ 232. mon. Coast redwood forest and mixed evergreen forest. Bowcutt, *MEDICAGO POL YMORPHA L.-Burclover. Abundant. Ruderal areas Henricks, & Kennedy 1765. and white alder riparian woodland. Native to Mediterranean. Lew­ QUERCUS AGRIFOLIA Nee var. AGRIFOL!A--Coast live oak. Abundant. is & Mutch 24, Bury & Herzog 294 (DA V, NCC), Botany 102 Coast live oak woodland, chamise chaparral, and mixed ever­ students 209. Botany 102 students 210. green forest. Bowcutt 1909b, Lee 87, Adams, Weick, Daehler & PlCKERINGIA MONTAl'

woodland. Observed along . Bald Mountain Trail. Also in oak Occasional. Opening in oak woodland. Loredo-Prendeville & scrub along Brushy Peak Trail. Bowcutt 1920. Canon 70. QUERCUS CHRYSOLEPIS Liebm.--Canyon live oak. Common. Mixed PHACELIA CAUFORNICA Cham.-California phacelia. Occasional. evergreen forest. Bowcutl 1872, Bowcutt 1909a, Bowcutt, Hen­ Road cut in oak woodland. Bowcutt 1808. ricks, & Kennedy 1774. PHACEUA IMBRICATA .E. Greene ssp. IMBRICATA. Occasional. Grassy QUERCUS DURATA Jepson var. DURATA-Leather Oak. Abundant. Ser­ opening in oak woodland. Bowcuu 2157. pentine chaparral. Bowcutt 1752b ( DA V, NCC), Bury & Her<.og PH.ACEtiA OJSTANS Benth.-Wild-heliotrope. Rare. Collected in Son­ 256. oma Creek Canyon by Howell in 1935 (Best et al. 1996). Also QUERCUS GARRYANA Hook. var. GARRY ANA--Oregon oak. Abundant. reported in park by Stocking (s.d.) and Wright (1975). Mixed evergreen forest and oak woodland. Bowcutt 2138 (DA V, NCC), Bowcutt & Knoll 2201. HYPERICACEAE QUERCUS KELLOGGD Newb.-Black oak. Common. Oak woodland. Canon & Loredo-Prendeville 73 (DAV, NCC). HYPERICUM CONCINNUM Benth.-Gold-wire. Occasional. Serpentine QUERCUS WJSLIZENI A. DC. var. FRUTESCENS Engelm.-lnterior Jive chaparral. Bowcuu 1884. oak. Rare. Oak scrub. Bowcutt 1921. JUGLANDACEAE GENTIANACEAE CENTAURWM TRICHANTHUM (Griseb.) Robinson--Centaury. Locally + JUGLANS CALrFORNICA S. Watson var. HINDSII Jepson-Northern California black walnut. Rare. Oak woodland at old homesites. common. Mesic meadow called Columbine Mcadow. Bowcull, Native to north-central California, frequently used as root stock Mawdsley, & Knoll 2195. for Juglans regia. Botany 102 students 81, Bowcuu & Knoll2202. GERANIACEAE LAMIACEAE *ERODIUM BOTRYS (Cav.) Bertol.-Filaree. Occasional. Annual grassland. Native to southern Europe. Bowcutt 1796a, Bowcuu *LAMIUM PURPUREUM L.- Red henbit. Rare. Annual grassland and 2071. white alder riparian woodland. Native to Europe. Botany 102 stu­ *ERODHJM cj BRACHYCARPUM (Godron) Thell.-Filaree. Occasional. dents 23, Bmany 102 students 61. Annual grassland along Lower Bald Mountain trail. Native to LEPECHINIA CALYCfNA (Benth.) Epling-Pitcher sage. Rare. Oak southern Europe. Bowcuu 1887, Bowcutt 2123. scrub along Brushy Peak Trail. Bowcutt 1924. *ERODJUM CICUTARIUM (L.) L'Her.- Red-Stemmed filaree. Common. *MARRUBIUM VULGARE L.- Horehound. Rare. Disturbed places. Na­ Annual grassland. Native to Eurasia. Bowcutt 1796, Bowcutt tive to Europe. Bowcutt 2063. 1824. *MELISSA OFFICINALJS L.-Bee balm, lemon balm. Rare. Trailhead *ERODlUM MOSCHATUM (L.) L'Her.-8torksbill. Common. Ruderal for Goodspeed trail on edge of dirt parking area, coast redwood situations. Native to Europe. Bowcuu 2072. forest. Native to southern Europe. Bowcurt 1943. *GERANWM DISSECTUM L.--Cranesbill. Common. Annual grassland. *MENTHA PULEGIUM L.-Pennyroyal. Rare. Outflow area from an old Native to Europe. Botany 102 students 22, Bowcutt 1784, Bowcutt stock pond in a grassy opening in oak woodland. Along Vista 1825b. trail, near junction with Bald Mountain trail. Native to Europe. *GERANIUM MOLLE L.--Cranesbill. Occasional. Annual grassland. Bowcutt 2059. Native to Europe. Bowcutt 1825. MONAROELLA VIRIDIS Jepson-Coyote mint. Occasional. Serpentine chaparral. Bowcuu 1879. GROSSULARIACEAE SALVIA COLUMBARIAE Benth.--Chia. Rare. Roadcut along Adobe R1BES CNERME Rybd. var. INERME-White-stemmed gooseberry. Oc­ Canyon Road. Bowcutt 2167. casional. Mixed evergreen forest. Bowcurt 1780. SCUTELLARIA CALIFORNJCA A. Gray- California skullcap. Rare. RIBES ROEZLII Regel var. CRUENTUM (E. Greene) Rehder-8ierra Grassland along Mcadow Trail. Bowcutt 1904. gooseberry. Rare. Oak woodland. Reported by Wright (1975). STACHYS AJUGOIDES Benth. var. RIGJDA Jepson & Hoover-Hedge­ Collected on Mt. Hood and on east slope of Mt. Hood (Best et nettle. Common. Mixed evergreen forest, oak woodland and al. 1996). white alder riparian woodland. Bowcutt 1807, Adams & Weick 283, Waraich & Togioka 234. Hll'POCASTENACEAE STACHYS ALBENS A. Gray-White hedge-nettle. Rare. Spring in oak woodland along Vista Trail, seep at Columbine Meadow, and AESCULUS CALIFORNICA (Spach) Nutt.--California buckeye. Com­ along pond on old Harr Ranch. Bowcuu 1893, Bowcull, Maw­ mon. Coast Jive oak woodland. Bowcull 1865. dsley, & Knoll 2172.

HYDRQPHYLLACEAE LAURACEAE EMMENANTHE PENDULIFLORA Benth.-Whispering bells. Rare. Re­ cently burned or otherwise disturbed chamise chaparral. Bowcutt UMBELLULARlA CALIFORNICA (Hook. & Am.) Nutt.--California bay. 2125. Common. Coast redwood forest, mixed evergreen forest, and ERIODICTYON CAUFORNICUM (Hook. & Am.) Torrey-Yerba santa. white alder riparian woodland. Botany 102 students 18, Botany Occasional. Chamise chaparral. Bowcutt 1918. 102 studems 259 (DAV, NCC). NEMOPHILA HETEROPHYLLA Fischer & C. Meyer-Variable-leaved ne­ mophila. Common. Mixed evergreen forest. Bowcuu 1753, Bow­ LINACEA£ cull & Mawdsley 2087. NEMOPHILA MENZlESII Hook. & Am. var. ATOMARIA (Fischer & C. HESPEROLINON MICRANTHUM (A. Gray) Small-Small-flowered dwarf Meyer) Chandler-Baby white-eyes. Occasional. Mixed ever­ flax. Occasional. Serpentine chaparral. Bowcull & Kennedy 1817. green forest and grassland. Adam & Weick 254, Lewis & Mutch HESPEROLINON SPERGUU:NlfM (A. Gray) Small-Dwarf flax. Rare. 3, Bowcutt & Green 1794. Chamise chaparral and serpentine chaparral. Bowcutt & Kennedy NEMOPHILA MENZfESU Hook. & Am. var. MENZIESU-Baby blue-eyes. 1882, Bowwtt & Kennedy 1910. 30 Bowcutt ALISO

LYfHRACEAE POLEMONIACBAE

*LYfHRVM HYSSOPIFOUUM L.-Hyssop loosestrife. Rare. Disturbed GILIA CAPITATA Sims. SSp. CAPITATA-Blue field gilia. Rare. Recently places. Native to Europe. Bowcutt 1839. burned coast live oak woodland. Bowcutt 1864. GII.IA TRICOJ.OR Benth. ssp. TRICOtOR-Tricolor gilia. Common. Pe­ MALVACEAE rennial grassland. serpentine chaparral. Bowcuu 1799. Lee 106, Botany 102 students 215. SJDALCEA DIPWSCYPHA (Torrey & A. Gray) Benth.-Fringed sidal­ LJNANTHUS ANDROSACEUS (Benth.) E. Greene-Mustang-clover. cea. Occasional. Perennial grassland. Bowcutt 1800, Botany 102 Common. Grassland. Beever 214. studellls 239. LINANTHUS BICOLOR (Nutt.) E. Greene--Bicolored linanthus. Uncom­ mon. Grassland. DiTomasso s.n. MORACEAE LINANTHUS PARVIfLORUS (Benth.) E. Greene-Linanthus. Occasional. *Ficus CARICA L.-Fig. Rare. Along seasonal creek in mixed ever­ Annual grassland and perennial grassland. Bowcu/1, Keeney, & green forest within Pony Gate Gulch, appears to be an adventive. Green 1802, Bowcutt & Green 1788, Botany 102 students 100, Native to Mediterranean. Bowcutt /867. Beever 214, Adams et al. 253 (DA V, NCC). NAVARRETIA LEUCOCEPHALA Benth. ssp. LEUCOCEPHAl.A-Wbite-fiow­ MYRTACEAE ered navarretia. Rare. Collected in Adobe Canyon between Mt. Hood and Sugarloaf Ridge (Best et al. 1996). +EUCALYY!1JS GLOBULUS Labill.- Biue gum. Rare. Roadside near NAVARRETIA SQUARROSA (Esch.) Hook. & Am. Skunkweed. Occa­ park employee home in northwestern portion of park. Bowcutt & sional. Dirt roads. Bowcutt 1915. Knoll 2185. PHLox GRACILIS E. Greene--Slender phlox. Rare. Grassy opening in oak woodland. Botany 102 studellls 92. Bowcutt & Kennedy OLEACEAE 1833. FRAXINUS LATIFOLIA Benth.-Oregon ash. Rare. Toe of Sugarloaf Ridge in mixed evergreen forest within campground. Bowcutt POLYGALACEAE 1777, Bowcutt & Knoll 2178. POLYGALA CALlFORNICA Nutt.-California m.ilkwort. Occasional. Mixed evergreen forest and serpentine chaparral. Togioka & War­ ONAGRACEAE aich 246, Botany 102 students 265. CLARKIA AMOENA (Lebm.) Nelson & ] . F. Macbr. ssp. HUNTIANA (Jep­ son) Harlan Lewis & M. Lewis-Farewell-to-spring. Common. POL YGONACEA£ Annual grassland and serpentine chaparral. Bowcutt 1810b. CLARKIA GRACILIS (Piper) Nelson & J. F. Macbr. ssp. GRACILis­ ERIOGONUM LUTEOI,UM E. Greene-Buckwheat. Locally common. Clarkia. Occasional. Serpentine chaparral. Bowcutt 1803, Bowcutt Serpentine chaparral. Bowcu/1 1912. Also collected on trail to Mt. 1810a, Botany 102 studems 274. Hood by Ross in 1984 (Best et al. 1996). CLARKIA PURPUREA (Curtis) Nelson & J. F. Macbr. ssp. QUADRIVUL· ERIOGONUM NUDUM Benth. var. NUDUM-Naked buckwheat. Rare. NERA (Douglas) Harlan Lewis & M. Lewis-Purple clarkia. Oc­ Coast live oak woodland, opening in mixed evergreen forest, and casional. Coast live oak woodland. Bowcu/1 & Green 1813, Bow­ serpentine chaparral. Bowcutt 1898, Bowcutt 1907. cute & Green 1834, Bowcutt 2159. *POLYGONUM ARENASTRUM Boreau.-Common knotweed, doorwecd. CLARKIA UNGUICULATA Lindley-Red ribbons. Rare. Grassy opening Occasional. Disturbed places. Native to Eurasia. Bowcutt 2120 in mixed evergreen forest. Bowcutt 1868. (DAV, NCC). EPILOBIUM CANUM (E. Greene) Raven ssp. l,ATIFOL£UM (Hook.) Ra­ *POL YGONUM HYDROPlPER L.-Smartwecd. LocaJly abundant in pond ven-Hummingbird's trumpet, California fuchsia. Rare. Large in northwestern portion of park. Native to Europe. Bowcutt, Maw­ rock outcrop along Vista Trail. Bowcutt 206/a. dsley, & Knoll 2174. EPILOBHIM MINUTUM Lehm.-Minute willow-herb. Occasional. Ser­ *RUMEX ACETOSELLA L.-Shecp sorrel. Common. Annual grassland. pentine chaparral. Bowcun & Green 1818. Native to Europe. Bowcu/1 1827, Bowcutt & Green 1787, Botany 102 students 10, Botany 102 students 62, Loredo-Prendeville & OROBANCHACEAE Canon 72 (DAV, NCC). *RUMEX CRISPUS L.-Curly dock. Rare. Annual grassland and ru­ 0ROBANCHE BULBOSA G. Beck-Broom-rape. Chamise chaparral. deral. Native to Eurasia. Bowcutt 1880, Bowcutt 2121. Reported in the park on the south slope of Sugarloaf Peak. Refer *RUMEX PULCHER L.-Fiddle dock. Occasional. Annual grassland. to Best et al. 1996, pp. 180 and 208 under Ceanothus sonomensis. Native to Mediterranean. Bowcu/1 1845, Bowcutt /88/. RUMEX SALICIFOLIUS J. A. Weinm. cf var. SALIClFOLIUS-Willow PAPAVERACEAE dock. Common. Spring in oak woodland along Vista Trail. Bow­ DENDROMECON RIGIDA Benth.-Bush poppy. Chaparral. Reported by cull /896. Stocking (s.d.) and Wright (1975). Collected on Mt. Hood (Best et al. 1996). PORTUL.ACACEAE EsCHSCHOLZIA CALlFORNICA Cham.-California poppy. Common. Grasslands. Bullock 245. CALANDRlNI.A CILIATA (Ruiz Lopez & Pavon) DC.-Red maids. OC­ casional. Annual grassland. Bowcutt 2080. CLAYTONI.A EXIGUA Torrey & A. Gray ssp. EXIGUA. Occasional. Ser­ PLANTAGINACEAE pentine chaparral. Adams et a/. 286. PLANTAGO ERECTA E. Morris-California plantain. Occasional. An­ CLAYTONtA PARVlFLORA Hook. ssp. PARVll-l..ORA-Small-leaved mon­ nual grassland and serpentine chaparral. Beever et al. 219 (DAV, tia. Common. Mixed evergreen forest and whlte alder riparian NCC), Adams et al. 263, Bowcu11 & Green 1805, Bowcutt & woodland. Bury et al. 291 (DAV, NCC). Mawdsley 2086. CLAYTONI.A PERFOLI.ATA Willd.-Miner's lettuce. Common. White al­ *PLANTAGO LANCEOLATA L.-English plantain. Occasional. Annual der riparian woodland. Lewis & Mutch 19, Waraich. Togioka, & grassland. Native to Europe. Lewis & Mutch 5. Beever 228. VOLUME 18, NUMBER I Sugarloaf Ridge State Park Flora 31

MoNTIA FONTANA L.-Water-chickweed. Occasional. Annual grass­ HETEROMELES ARBUTIFOLIA (Lindley) Roemer-Toyon. Common. land in campground. Bowcutt 1792, Bowcuu & Mawdsley 2067. Mixed evergreen forest. Bury et al. 257 (DAV, NCC). MONTIA PARVIFOLIA (DC.) E. Greene-Montia. Rare. Growing on HOLODISCUS DISCOLOR (Pursh) Maxim.-Oceanspray. Rare. Mixed mossy bank along Sonoma Creek in mixed evergreen forest. Bow­ evergreen forest. Bowcutt 2148. cult 1874. POTENTR.LA GLANDULOSA Lindley ssp. GLANDULOSA-Sticky cinque­ foil. Rare. White alder riparian woodland. Bowcutt 1842. PRlMULACEAE *PRuNus CERASIFERA Ehrh.-Cherry plum. Occasional. Near camp­ ground. Native to southeastern Europe. Bowcutt 1929, Loredo­ *ANAGALLIS ARVENSIS L.-Scarlet pimpernel. Common. Ruderal sit­ Prendeville & Canon 75, Bowcutt & Mawdsley 2066a (DAV, uations. Native to Europe. Loredo-Preruieville & Canon 69 (DAV. NCC). NCC), Beever 218, Botany /02 students 297 (DAV, NCC). PRUNUS VTRGlNIANA L. var. DEMISSA (Nutt.) Torrey- Western choke DoOECATHEON HENDERSOND A. Gray-Shooting star. Occasional. cherry. Rare. White alder riparian woodland. BowcuN 2161 (DAV, Oak woodland. Bowcuu 209/, Lee 96. NCC). TRiENTALIS LATIFOLIA Hook.-Star flower. Common. Coast redwood ROSA GYMNOCARPA Nutt. in Torrey & A. Gray-Wood rose. Com­ forest and mixed evergreen forest. Bowcuu 1756. mon. Coast redwood forest and mixed evergreen forest. Bowcutt 1766b (DA V. NCC). RANUNCULACEAE *Ruaus DISCOLOR Weihe & Nees-Himalaya-berry. Occasional. AQUll.EGIA FORMOSA Fischer-Columbine. Locally common. Mesic Seep in opening within oak woodland along Hillside Trail. Native meadow called Columbine Meadow. Bowcutt, Mawdsley, & Knoll to Europe. Bowcutt 1836. 2194. RUBUS PARVIFLORUS Nutt.-Thimbleberry. Rare. Coast redwood for­ DELPHINIUM NUDICAULE Torrey & A. Gray- Scarlet larkspur. Occa­ est and mixed evergreen forest. Bowcutt /766a. sional. Oak woodland. Botany 102 students 94, Beever, Togioka, RUBUS URSINUS Cham. & Schldi.-Califomia blackberry. Occasion­ & Bullock 235. al. Opening and seep in oak woodland. Loredo-Preruieville & RANUNCULUS CALIFORNICUS Benth.-California buttercup. Common. Canon 77. Grasslands and oak woodland. Lewis & Mutch 6, Waraich, To­ gioka, & Bullock 223. RUBIACEAE *RANUNCULUS MURICATUS L.-Prick!e-fruited buttercup. Common. GALIUM APARINE L.-Bedstraw, goosegrass. Common. Draw within Annual grassland in campground, mixed evergreen forest, and grassland at edge of coast live oak woodland. Although listed as ruderal habitats. Native to Europe. Lewis & Mutch 7, Lewis & native here by Dempster (1993), she suggests that it may be na­ Mutch I/, Loredo-Prendeville & Canon 60, Botany 102 students tive to Europe. Botany 102 students 20, Botony 102 students 222, 300. Botany 102 students 267 (DAV. NCC), Bowcutt 185/b. *GALIUM DIVARICATUM Lam.-Lamarck's bedstraw. Occasional. An­ RHAMNACEAE nual grassland. Native to the Mediterranean. Bowcutt 1812. *GALIUM MURALE (L.} AlL-Tiny bedstraw. Occasional. Meadow in CEANOTHUS ClJNEATUS (Hook) Nutt. var. CUNEATUS-Buckbrush. Oc­ campground. Native to Europe. Bowcutt & Mawdsley 2069. ca<;ional. Chamise chaparral. Bowcutt 191/, Bowcutt 2079, Bow­ *GALIUM PARISIENSE L.- Wall bedstraw. Occasional. Annual grass­ cutt 2134. land. Native to Mediterranean. Bowcutt 1888, Bowcutt 2124, CEANOTHUS FOLIOSUS C. Parry var. rULIOSUS-Wavyleaf ceanothus. Bowcutt 2181. Rare. Chaparral. Reported by Stocking (s.d.) and Wright (1975). GALIUM PORRIGENS Dempster var. TENUE (Dempster) Dempster­ Collected on Mt. Hood in 1893 (Best et al. 1996). Ciimbing bedstraw. Common. Serpentine chaparral, chamise CEANOTHUS JEPSONIT E. Greene var. JEPSONII-Muskbrush. Common. chaparral. and oak woodland. Bowcutt 1752d, BowcuN 1831, Bot­ Serpentine chaparral. Adams & Weick 269, Bowcutt & Green any /02 students 49, Botany 102 students 268 (DAV, NCC). 1752c. CEANOTHUS OUGANTHUS Nutt. var. SOREDIATUS (Hook. & Arn) Hoo­ SALICACEAE ver- Jim brush. Occasional. Chamise chaparral. Bowcutt 2133 SALIX LASIOLEPIS Benth.-Willow. Occasional. White alder riparian (DAV, NCC). woodland. Bowcutt 2073, Bowcutt 2137b. CEANOTHUS PARRY! Trei.-Parry's ceanothus, lady bush. Occasional. Mixed evergreen forest. Bowcutt & Green 1798. SAX!I'RAGACEAE CEANOTHUS SONOMENSIS J. Howell-Sonoma ceanothus. Locally common. Serpentine chaparral. CNPS List LB. Bowcuu & Covert BOYKINIA OCCIDENTALIS Torrey & A. Gray-Coast boykinia. Rare. 2139, Bury & Herzog 264. Coast redwood forest along Sonoma Creek. Bowcutt 1940. RHAMNus CA.LlFORNlCA E.~chsch.-California coffeeberry. Occasion­ HEUCHERA MICRANTIIA Lindley-Aiumroot. Rare. Mixed evergreen al. Mixed evergreen forest. Bowcul/ 1899. forest. Bowcutt 2147. RHAMNUS CROCEA Nutt.-Spiny redberry. Rare. Reported by Wright LITHOPHRAGMA AFFINE A. Gray-Woodland star. Occasional. Oak ( 1975). Collected near Sonoma Creek at foot of Mt. Hood in 1902 woodland. Botany 102 students 17, Botany 102 students 71 (DA V. (Best et al. 1996). NCC). LITHOPHRAGMA HETEROPHYLLUM (Hook. & Am.) Torrey & A. Gray­ ROSACEAE Woodland star. Occasional. Mixed evergreen forest. Bowcutt & Keeney 1759. ADENOSTOMA FASClCULATUM Hook. & Arn.-Chamise. Abundant. SAXIFRAGA CALIFORNICA E. Greene-California saxifrage. Rare. Oak Chamise chaparral. Daehler, Lu, Bury, & Herzog 270 (DA V. woodland. Bowcutt & Mawdsley 2092. NCC). APHANES OCCIDENTALIS (Nutt.) Rydb.-Western lady's mantle, dew SCROPHULARIACEAE cup. Rare. DiTomo.so & Kaiser s.n. (private collection). CERCOCARPUS BETUL01DES Torrey & A. Gray var. BETULOIDES­ ANTlRRHINUM VIRGA A. Gray- Tall snapdragon. Rare. Collected in Mountain-mahogany, birch-leaf mountain-mahogany. Occasional. Adobe Canyon and at the foot of Mt. Hood (Best et al. 1996). Chamise chaparral. BowcuN 1908. CASTILLEJA ATTENUATA (A. Gray) Chuang & Heckard-VaHey tas- 32 Bowcutt ALISO

sels. Locally abundant in May 1998. Nonnative grassland. Maw­ VISCACEAE dsley 30. PHORADENDRON VILLOSUM (Nutt.) Nutt.-Oak mistletoe. Occasional. CASTILLEJA FOLIOLOSA Hook. & Arn.-Wooly Indian paintbrush, Growing on a coast live oak tree within a grassland. Bowcutt, chaparral paintbrush. Occa~ional. Oak scrub along Brushy Peak Mawdsley, & Knoll 2191, Bowcutt & Knoll 2203. Trail and serpentine outcrop in mixed evergreen forest. Bowcutt 1926. Bowcutt 2168. Loredo-Prendeville & Canon 64. VITACEAE COLLINSIA HETEROPHYLLA Buist-Chincse houses. Locally common. Oak woodland. Bowcutl 1830. Vms CAL!FORNICA Benth.-Califomia wild grape. Occasional. Ob­ COLLtNSIA SPARSifLORA Fischer & C. Meyer var. ARVENSIS (E. served by the author within riparian zone along Sonoma Creek Greene) Jepson-Few-flowered blue-eyed Mary. Locally com­ near Goodspeed Trail head. Also collected in "Adobe Canyon mon. Annual grassland. Botany 102 students 99. adjacent to Mt. Hood" (Best et al. 1996). MIMULUS AURANTIACUS Curtis-Sticky monkey flower. Common. Serpentine chaparral and openings in mixed evergreen forest. Lo­ redo-Prendeville & Canon 66, Bullock 216, Bury & Herzog 281. MONOCOTYLEDONEAE MIMULUS CARDINALIS Benth.-Scarlet monkey flower. Rare. White CYPERACEAE alder riparian woodland. Bowcult 1935. MIMULUS CONGDONII Robinson-Congdon's monkey flower. Rare. CAREX AMPL!FOLIA Boon-Sedge. Occasional. Mixed evergreen for­ Probably in burned chaparral area. Collected on "southeast slope est and white alder riparian woodland. Bowcutt, Kennedy, & Hen­ of Mt. Hood" (Best et al. 1996). ricks 1776, Bowcutt 1934 (DA V, NCC). MIMULUS DOUGLASII (Benth.) A. Gray-Purple mouse ears. Rare. CAREX BUXBAUMil WahL-Sedge. Occasional. Small draw in Elymus Probable on serpentine, collected on "southeast slope of Mt. glaucus dominated grassland. Bowcutt 1851a. Hood" (Best et al. 1996). CAREX DENSA L. Bailey-Dense sedge. Locally common. Mixed ev­ MIMULUS GUTTATUS DC.-Common monkey flower. Rare. Spring in ergreen forest. Bowcutt, Kennedy, & Henricks 1785. oak woodland. Botany 102 students 276. CAREX GLOBOSA Boott- Sedge. Occasional. Mixed evergreen forest. MtMULUS KELLOGG!I (E. Greene) A. Gray-Kellogg's monkey flower. Bowcutt 1760 (DA V, NCC). Rare. Collected on "southeast slope of Mt. Hood" (Best et al. CAR.EX OVALIS Good-Sedge. Occasional. Mixed evergreen forest. 1996). Bowcutt 1782a. MIMULUS MOSCHATUS Lindley-Musk monkeyflower. Rare. Along CAREX TUMULICOLA Mkze.-Foothill sedge. Occasional. Meadow in pond. Bowcutl, Mawdsley, & Knoll 2173. campground. North 12, Botany 102 students 292. *PARENWCELLIA VISCOSA (L.) Caruel-Yellow parentucellia. Ruder­ CYPERUS ERAGROSTIS Larn.-Sedge. Common. Moist disturbed plac­ a!, between horse stables and the day use parking lot. Native to es and white alder riparian woodland. Bowcutt 1837. Europe. Mawdsley s.n. (private collection). PENSTEMON HETEROPHYLLUS Lindley var. HI:."TEROPHYLLUS-Foothill UUDACEAE penstemon. Occasional. Serpentine chaparr-dl. Bowcutt, Keeney, llus FERNALDII R. Foster-Iris. Occasional. Mixed evergreen forest. & Green 1809 (DAV, NCC). Bowcutt 1761 (DA V, NCC), Beever 227. SCROPH'ULARIA CALIFORNICA Cham. & Schldl. ssp. CAL!FORNICA-Cal­ SISYRtNCHRJM BELLUM S. Watson-Blue-eyed grass. Occasional. An­ ifornia figwort, bee plant. Occasional. Seep in opening in oak nual grassland and open oak woodland. Botany 102 students 39, woodland. Bowcutt 1835. Bullock, Togioka, & Waraich 244 (DAV, NCC). TRIPHYSARIA ERIANTHA (Benth.) Chuang & Heckard ssp. ERJANTHA­ Butter-and-eggs, Johnny-tuck. Occasional. Grassland. Reported JUNCACEAE by DiTomaso (1996), Stocking (s.d.), and Wright (1975). Col­ lected along Adobe Canyon Road near park (Best et al. 1996). JUNcus BALTICUS Willd.-Wire rush. Occasional. Moist open places. Bowcutt 2182b, Botany 102 students 251. SOLANACEAE JUNCUS BUFONRJS L. var. BUFONIUs-Toad-fiax. Common. Disturbed places. Bowcutt 1840, North 14, Bury & Herzog 251b, Botany SOLANUM XANTI A. Gray-Purple nightshade. Rare. Opening in oak 102 students 281. woodland. Loredb-Prendeville & Canon 76. ]UNCUS EFFUSUS L. var. PACIFICUS Fern. & Wieg.-Rush. Common. Seasonally moist swales. Bowcutt 1781. URTICACEAE JUNCUS NEVADENSIS S. Watson-Rush. Locally common. Freshwater marsh at stock pond. Bowcutt & Knoll 2182a. URTICA OJOICA L. S~"j). HOLOSERJCEA (Nutt.) Thorne-Hoary nettle, JuNCUS PATENS E. Meyer-Rush. Occasional. Annual grassland. stinging nettle. Occasional. White alder riparian woodland. Bow­ Bowcutt 1848, Bowcutt 2153a. curt 1901. JUNcus XIPHIOrDES E. Meyer-Rush. Occasional. Spring in oak woodland along Vista Trail and white alder riparian woodland. VALERIANACEAE Bowcutt 1895, Bowcutt 1936, Bowcutt 2179. LuzuLA COMOSA E. Meyer-Wood rush. Occasional. Meadow in Pl..ECTRmS BRACHYSTEMON Fischer & C. Meyer-Picctritis. Occa­ campground and openings in oak woodland. North 11. sional. Oak woodland. Bowcutt 1829. LEMNACEAE VERBENACEAE LEMNA MlNUSCULA Herter-Duckweed. Locally abundant in pond in VERBENA LASIOSTACHYS Link-Western verbena. Occasional. White northwestern portion of park. Bowcutt, Mawds/ey, & Knoll 2175. alder riparian woodland. Bowcutt 1933. LILIACEAE VIOLACEAE BRODIAEA ELEGANS Hoover ssp. ELEGANs-Harvest brodiaca. Rare. VIOLA OCELLATA Torrey & A. Gray-Western heart's ease. Occa­ Annual grassland. Bowcutt 1847. DiTomaso & Kaiser s.n. (pri­ sional. Coast redwood forest. Bowcutt 2143, Baker 955 (CAS). vate collection). VOLUME 18, NUMBER I Sugarloaf Ridge State Park Flora 33

CALOCHORTIJS AMASILIS Purdy-Yellow globe lily, Diogenes' lan­ BROMUS CARINATUS Hook. & Am. var. CARINATUS-California tern. Common. Serpentine chaparral. Botany 102 students 217. brome. Common. Mixed evergreen forest. North 3. North 8. CALOCHORTUS LUlCUS Lindley-Yellow mariposa lily. Occasional. *BROMUS DIANDRUS Roth-Ripgut grass. Abundant. Disturbed places Annual grassland. Bowcutl I 850. and rocky openings in serpentine chaparral. Native to Europe. CHLOROGALUM POMERIDIANUM (DC.) Kunth.-Soap plant. Common. North 1, Bowculf & Kennedy I815. Annual grassland and openings in mixed evergreen forest. Ob­ *BROMUS HORDE.ACEus L.-Soft chess. Common. Disturbed places served by the author and reported by DiTomaso (1996), Stocking and serpentine chaparral. Native to Eurasia. North 24, Bowculf, (s.d.), and Wright (1975). Kennedy, and Henricks 1751. DICHELOSTEMMA CAPITATUM Alph.-Wood ssp. CAPITATUM-Biue BROMUS LAEVIPES Shear-Brome. Common. Coast live oak wood­ dicks. Common. Grasslands and openings in mixed evergreen for­ land and serpentine chaparral. Bowcutt 1914, Bowcutt 2150, Bow­ est. Loredo-Prendeville & Canon 68, Bullock, Togioka, & War­ cult, Kennedy, & Henricks 1752, North 20. aich 237. *BROMUS MADRITENSIS L. SSp. RUBENS (L.) Husnot-Foxtail chess, DICHELOSTEMMA CONGESTUM (Sm.) Kunth.-Ookow. Occasional. Spanish brome. Occasional. Disturbed places. Native to Europe. Grassland. Bowcutt 2I49, Bowcutt 2170. North 18. DISPORUM HOOKER! (Torrey) Nicholson-Fairy bells. Occa~ional. *CYNOOON DAC.TYLON (L.) Pers.-Bennuda grass. Occasional. Road­ Coast redwood forest and mixed evergreen forest. Bowcutt, Ken­ side. Native to Africa. Bowculf 2165. nedy. & Henricks 1769. *CYNOSURUS ECHlNATUS L.-Hedgehog dogtaiJ. Abundant. Grass­ FRITILLARIA AFFINIS (Schultes) Sealy var. AFFINls-Checker lily. lands, openings in oak woodland, and in disturbed places. Native Common. Serpentine chaparral. Botany 102 students 48. to Europe. North 21, North 47. SMtLAClNA STELLATA (L.) Desf.-Slim Solomon's seal. Common. DANTHOMA CALIFORNICA Bolander var. CALIFQRNICA-Califomia oat­ Coast redwood forest and mixed evergreen forest. Bowcutt 1779, grass. Rare. Meadow in center of campground. North 37, Bow­ Bowcutt, Kennedy, & Henricks .1768. curt, Kennedy, and Henricks 1793. TRU.LIUM AL!IIDUM Freeman-Wake robin. Rare. Oak woodland. ELYMUS ELYMOIDES (Raf.) Swezey ssp. CAUFORNICUS (J. G. Smith) Bowcutt & Mawdsley 2088. Barkworth-Squirreltail. Occasional. Serpentine grassland along TRU.LIUM OVATUM Pursh-Western trillium. Rare. Vegetative plant Stern Trail. Bowcull /878. observed by author in coa~t redwood forest. Reported by Stocking ELYMUS GLAUCUS Buckley ssp. GL.AUCUS-Biue wild rye. Common (s.d.) and Wright (1975). to locally abundant. Annual grassland on edge of coast Live oak TRrraEIA LAXA Benth.-lthuriel's spear. Common. Recently burned woodland. Bowcutt .1846, Bowcu/1 2156. coast live oak woodland. Bowcutt 1863. ELYMUS TRACHYCAULUS (Link) Sinn. var. SUBSECUNOUS (Link) Gould ZIGADENUS FREMONTIJ (Torrey) S. Watson- Fremont's death camas. vel aff.-Slender wheatgrass. Occasional. Annual grassland Occasional. Chamise chaparral/coast live oak woodland ecotone. burned in July 1995. Bowculf 206/b. Bowcutt & Mawdsley 2078. FESTUCA CALIFORNICA Vasey-California fescue. Locally abundant. ZIGADENUS MICRANTHUS Eastw. var. MICRANTHUs-Small-flowered Oak woodland. Bowcutt 2131 (DAV, NCC}, North 29. death camas. Reported by DiTomaso (1996), Stocking (s.d.), and Wright (1975). Collected on Mt. Hood in 1902 (Best et al. 1996). FESTUCA ELMER! Scribner & Merr.-Elmer's fescue. Occasional. Var­ ious habitats. North 9. FESTUCA IDAHOENSIS Elmer-Idaho fescue, blue bunchgrass. Locally ORCHIDACEAE common. Campground meadow. North 35. PIPER!A ELEGANS (Lindley) Rydb.-Elegant rein orchid. Rare. Two *GASTRIDIUM VENTRICOSUM (Gouan) Schinz & The! I.-Nit grass. Oc­ plants observed by the author along Brushy Peaks Trail and sea­ casional. Disturbed sites and chamise chaparral. Native to Europe. sonal creek which feeds into Maim Fork of Sonoma Creek. No Bowcutt 2I27a, DiTomaso & Kaiser s.n. (private collection). specimens gathered due to small population size. Also reported *HOLCUS LANATUS L.-Common velvet grass. Locally common. An­ from the park by R. Coleman (Best et aL 1996). nual grassland. Native to Europe. Bowcutt I852. PIPERIA ELONGATA Rydb.-Piperia. Collected in the park by R. Mor­ HORDEUM BRACHYANTHERUM Nevskj SSp. BRACHYANTHERUM-Mead­ gan (Best et al. 1996). ow barley. Occasional. Annual grassland. Bowcutt 1853, Bowcutt PIPERIA TRANSVERSA Rydb.-Piperia. Collected near Adobe Canyon 2142a (NCC), North 38. by Baker in 1928 and observed more recently in the park by R. *HORDEUM MARJNUM Hudson ssp. GUSSO!'IEANUM (Pari.) Thell.­ Morgan (Best et al. 1996). Mediterranean barley. Common. Annual grassland. Native to Eu­ rope. Bowcu/1, Kennedy, and Henricks I791. *HORDEUM MURINUM Hudson ssp. LEPORINUM (Link) Arcang.-Bar­ POACEAE ley. Common. Disturbed places. Native to Europe. North 2. AGROSTIS PALLENS Trin.-Thin grass. Occasional. Recently burned *LAMARCIA AUREA (L.) Moench-Goldentop. Rare. Ruderal situa­ coast live oak woodland. Bowcutt I 856. tions on Bald Mountain. Native to Mediterranean. Mawdsley s.n. *AIRA CARYOPHYLLA L.-Silver European hairgrass. Common. An­ (private collection). nual grassland. Native to Europe. North 31. *LoLIUM MULTIFLORUM Lam.-ltalian ryegrass. Abundant. Ruderal *AVENA BARBATA Link-Slender wild oats. Abundant. Annual grass­ situations. Native to Europe. North 17. land. Native to southern Europe. Bowcutt, Henricks, & Kennedy *LoLIUM PERENNE L.-Percnnial ryegrass. Common. Annual grass­ I789. land. Native to Europe. Bowcutt I84I, Bowcutt I844, Bowcult *AVENA FATUA L.-Wild oat. Locally abundant. Annual grasslands. 2162. Native to Europe. North 50, Bowcutl 1790. MELICA CALIFORNICA Scribner-California mclic. Occasional. An­ *JlRACHYPODfiJM DISTACHYON (L.) Beauv. Rare. Coast live oak wood­ nual grassland and grassy opening in oak woodland. Bowcutt land. Native to southern Europe. Bowcutt 215I. 2155. *BRIZA MAXIMA L.- Rattlesnake gra~s. Common. Roadsides and MELICA SUBULATA (Gris) Scribner-Alaska onion-grass. Common. other disturbed places. Native to southern Europe. North I 5, Mixed evergreen forest. Bowcutt 1782b (DAV, NCC). North 44. MELICA TORREYANA Scribner-1brrey's melic grass. Common. Ser­ *BRIZA MINOR L.-Little quaking grass. Common. Annual grassland. pentine chaparral and mixed evergreen forest. Bowcutt 1883, Native to southern and western Europe. North 42. North 5, North 25 (NCC). 34 Bowcutt ALISO

NASSELLA CERNUA (Stebb. & Love) Barkworth-Nodding needle­ A Hora of Sonoma County: manual of the flowering plants and grass. Common. Chamise chaparral. North 51. fe.ms of Sonoma County, Califo.mia. First Edition. California Na­ NASSEU.A LEP(DA (A. Hltchc.) Barkworth-Foolhill needlegrass. tive Plant Society. 347 p. Common. Chamise and serpentine chaparral. Bowcun 2127b, CALIFORNIA DEPARTMet-IT OF PARKS AND REcREATION. 1992. Sugar­ Bowcun 2140 (NCC), North 27. loaf Ridge State Park. Map, with natural and cultural history, pub­ NASSEU.A PULCHRA (A. Hltchc.) Barkworth-Purple needlegrass. lished by the Califo.mia Department of Parks and Recreation, Sac­ Common. Annual and perennial gra.'ISiands. North 32. ramento. California. *PASPALUM OlLATATUM Poiret- Dams grass. Occasional. Roadside CONSTANCE, L. 1993. Apiaceae, pp. 13~ 166 . In 1. C. Hiclcman (ed.), in campground. Native to South America. Bowcutt 2144. The Jepson manual: higher plants of California. Univ. of Calif. *PHAU.RIS AQUATlCA L.- Harding grass. Rare. Chamise chaparral Press, Berkeley, California. 1400 p. along Brushy Peak trail. Native to Meditemmean Europe. Bow­ Of:Ml'sTER, L. 1993. Rubiaceae, pp. 97~986 . In J. C. Hickman (ed .}, cun 1927. The Jepson manual: higher plants of California. Univ. of Calif. *PtPTATHERUM M!I..,IACEUM (L.) Cosson-Srnilo grass. Occasional. Press, Berkeley, California. 1400 p. White alder riparian woodland. Native to Eurasia. Bowcutt 1931. DITOMASO, J. 1996. Species list for Sugarloaf Ridge State Park *POA ANNUA L.-Annual bluegrass. Common. Disturbed places. Na­ grassland areas. Unpublished list obtained from the author. Uni­ tive to ~urope. North 4. versity of California, Davis, California. *PoA BULBOSA L.-Bulbous bluegrass. Occasional. Meadow in cen­ &.FoRo, C. R. 1964. Climate of Sonoma County. U.S. Department ter of campground. Native to Europe. Nonh 40. of Commerce, Weather Bureau. 37 p. POA SECUNDA J. S. Pres! ssp. SECUNDA-One-sided bluegrass. Oc­ HASTINGS, M. S. AND J. M. DITOMASO. 1996. Fire controls yellow casional. North 34, Nonh 46. star thistle in California grasslands. Restoration & Managemenr PoA TRIVIAUS L.- Trivial poa. Occasional. Coast redwood forest. Notes 14(2): 124- 128. Bowcutt /942. HICKMAN. J. C. (ed.) 1993. The Jepson manual: higher plants of *POLYPOGON MONSPELIENSIS (L.) Desf.-Rabbitfoot grass. Common. California. Univ. of Calif. Press, Berkeley, Califo.mia. 1400 p. Disturbed places. Native to Europe. Bowcutt, Keeney, & Green HOLMGREN, P. K .. N. H. HOt.MGREN, and L. C. BARNETT (eds.). 1990. 1838. Index herbariorum Part I: The Herbaria of the World, 8th edition. *TAENIATHERUM CAPUT-MEDUSA£ (L.) Nevski- Medusa head. Local­ Regnum Veg .• Vol 120. ly abundant. Annual grassland. Native to Europe. Bowcutt 1843b, JsLEv, D. 1993. Trifoliwn. pp. 646-654./n J. C. Hlckman (ed.) 1993. North 56 (NCC). The Jepson manual: higher plants of California. Univ. of Calif. TRISET\JM CANESCENS Buckley-Tall trisetum. Occasional in moist Press, Berkeley, California. 1400 p. places. Botany I 02 students 9. MASON. H. L. 1957. A Hom of the marshes of California. Univ. of *TRrnCUM AESTIVUM L.-Wheat. Rare. Roadside between horse cor­ Calif. Press, Berkeley. California. 878 p. ral and group campground. Native of E.urope. Bowcutt, Ma w­ MUNZ, P. A., and D. KF.CK. 1973. A California Hom with supplement. dsley, & Knoll 2188. Univ. of Calif. Press. Berkeley, California. 1905 p. *VULPIA BROMOIDES (L) S. F. Gray-Brome vulpia, six-week fescue. NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION (NOAA). Common. Ruderal situations. Native to Europe. North /6, North 1993. C limatological data. annual summary California. VoL 97, 39 (NCC). No.3. *VULPIA MYUROS (L) Gmelin var. HIRSUTA (Hackel) Asch. & Graeb­ SHELTON, M . L. 1978. Calibrations for computing Thornthwaite's ner- Rattail fescue. Common. Ruderal situations. Native to Eu­ potential evapotranspiration in California. The Professional Ge­ rope. North /3, North 19 (NCC). ographer 30(4): 389- 396. SKJNNER. M . W. , and B. M . P AVLIK. 1994. Inventory of r.ue and TYPHACEA£ endangered vascular plants of California. California Native Plant Society Special Publication No. I. (5th Edition) 338 p. TYPHA cf. DOM INGENSIS Pers.--CaUail. Locally abundant. Along edge STEBBINS, G. L 1998. San Fr.rncisco bay region excursions. Fre­ of pond in northweste.m portion of park. Bowcutt & Knoll 2183. montia 26(2): 9-17. STOCKING, K. s.d. Some of lhe more conspicuous vascular plants of LITERAlURE CITED Sugarloaf Ridge S tate Park. Unpublished li$t in Department of Parks and Recreation files. Sonoma, California. ABRAMS, L. 1940. Lllusttated flora of the Pacific States. Vol. I. Stan­ U.S. DEPA.RTMEI'fT OF AGRICULTURE. 1972. Soil survey: Sonoma ford Univ. Press. Stanford, Cali fo.mia. 538 p. County, California. Produced by the U.S. Forest Service and Soil ---. 1944. Illustrated flora of the Pacific States. VoJ. 2. Stanford Conservation Service in cooperation with the Univ. of Calif. Ag­ Univ. Press, Stanford, Califo.mia. 635 p. ricultural Experiment Station. U. S. Gov. Print. Off., Washington, ---. 1951. Ulustrated tlora of the Pacific States. VoL 3. Stanford D.C. 182. 120 p. Univ. Press. Stanford, California. 866 p. WAGNER, D. L., and E. J. BORTUGNO. 1982. Geologic map of the ---AND R. S. FERRIS. 1960. Illusttated flora of lhe Pacific States. Santa Rosa Quadrangle, California, I :250,000. Regional geologic Vol. 4. Stanford Univ. Press, Stanford. California. 732 p. map series. Map No. 2A. Sheet I of 5. California Division of BARBOUR, M. G., J. H. BURK, AND W. D. PITTs. 1987. TeiTCStrial Mines and Geology. plant ecology. Second Edition. The Benjamin/Cummings Publish­ WHATFORD, J. C. 1994. Vegetation typing/mapping of Sugarloaf ing Company, Inc. Menlo Park, California. Ridge State Park. Unpublished letter repon for the California De­ BARNHART, S. 1. , 1. R. McBRIDE, and P. WARNER. 1996. Invasion of partment of Parks and Recreation. Sonoma, California. northern oak woodlands by Pseudotsuga mentie~·ii (Mirb.) Franco WRIGHT, K. 1975. Sugarloaf Ridge State Park resource inventory. in the Sonoma Mountains of Califo.mia. Madroiio 43(1): 28-45. Unpublisb ed list in the California Department of Parks and Rec­ BEST. C., J. T. HOWELL, W. I