Hymenoptera, Platygastridae)
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
The Parasitoid Complex Associated with the Invasive Swede Midge
The parasitoid complex associated with the invasive swede midge, Contarinia nasturtii Kieffer (Diptera: Cecidomyiidae) in Europe: prospects for classical biological control in North America. By Paul K. Abram A thesis submitted to the Faculty of Graduate Studies and Research in partial fiilfillment of the requirements for the degree of Master of Science in Biology Carleton University, Ottawa, Ontario, Canada ©2012, Paul K. Abram Library and Archives Bibliotheque et Canada Archives Canada Published Heritage Direction du Branch Patrimoine de I'edition 395 Wellington Street 395, rue Wellington Ottawa ON K1A0N4 Ottawa ON K1A 0N4 Canada Canada Your file Votre reference ISBN: 978-0-494-87830-9 Our file Notre reference ISBN: 978-0-494-87830-9 NOTICE: AVIS: The author has granted a non L'auteur a accorde une licence non exclusive exclusive license allowing Library and permettant a la Bibliotheque et Archives Archives Canada to reproduce, Canada de reproduire, publier, archiver, publish, archive, preserve, conserve, sauvegarder, conserver, transmettre au public communicate to the public by par telecommunication ou par I'lnternet, preter, telecommunication or on the Internet, distribuer et vendre des theses partout dans le loan, distrbute and sell theses monde, a des fins commerciales ou autres, sur worldwide, for commercial or non support microforme, papier, electronique et/ou commercial purposes, in microform, autres formats. paper, electronic and/or any other formats. The author retains copyright L'auteur conserve la propriete du droit d'auteur ownership and moral rights in this et des droits moraux qui protege cette these. Ni thesis. Neither the thesis nor la these ni des extraits substantiels de celle-ci substantial extracts from it may be ne doivent etre imprimes ou autrement printed or otherwise reproduced reproduits sans son autorisation. -
Hymenoptera: Platygastridae) Parasitizing Pauropsylla Cf
2018 ACTA ENTOMOLOGICA 58(1): 137–141 MUSEI NATIONALIS PRAGAE doi: 10.2478/aemnp-2018-0011 ISSN 1804-6487 (online) – 0374-1036 (print) www.aemnp.eu SHORT COMMUNICATION A new species of Synopeas (Hymenoptera: Platygastridae) parasitizing Pauropsylla cf. depressa (Psylloidea: Triozidae) in India Kamalanathan VEENAKUMARI1,*), Peter Neerup BUHL2) & Prashanth MOHANRAJ1) 1) National Bureau of Agricultural Insect Resources, P.B. No. 2491, Hebbal, 560024 Bangalore, India; e-mail: [email protected]; [email protected] 2) Troldhøjvej 3, DK-3310 Ølsted, Denmark; e-mail: [email protected] *) corresponding author Accepted: Abstract. Synopeas pauropsyllae Veenakumari & Buhl, sp. nov., a new species of Synopeas 23rd April 2018 Förster, 1856 (Hymenoptera: Platygastroidea: Platygastridae: Platygastrinae), is recorded from Published online: galls induced by Pauropsylla cf. depressa Crawford, 1912 (Hemiptera: Psylloidea: Triozidae) 29th May 2018 on Ficus benghalensis L. (Moraceae) in India. It is concluded that S. pauropsyllae is a pa- rasitoid of this psyllid species. This is the fi rst record of a platygastrid parasitizing this host. Key words. Hymenoptera, parasitoid wasp, Hemiptera, Sternorrhyncha, psyllid, taxonomy, gall, host plant, Ficus, India, Oriental Region Zoobank: http://zoobank.org/urn:lsid:zoobank.org:pub:5D64E6E7-2F4C-4B40-821F-CBF20E864D7D © 2018 The Authors. This work is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs 3.0 Licence. Introduction inducing plant galls are mostly scale insects, aphids and With more than 5700 species and 264 genera, Platy- psyllids. Among psyllids (Hemiptera: Sternorrhyncha: gastroidea is the third largest superfamily in the parasitic Psylloidea), several families are known to induce galls; Hymenoptera after Ichneumonoidea and Chalcidoidea gall-making species are particularly numerous in Triozidae, (AUSTIN et al. -
Iranian Aphelinidae (Hymenoptera: Chalcidoidea) © 2013 Akinik Publications Received: 28-06-2013 Shaaban Abd-Rabou*, Hassan Ghahari, Svetlana N
Journal of Entomology and Zoology Studies 2013;1 (4): 116-140 ISSN 2320-7078 Iranian Aphelinidae (Hymenoptera: Chalcidoidea) JEZS 2013;1 (4): 116-140 © 2013 AkiNik Publications Received: 28-06-2013 Shaaban Abd-Rabou*, Hassan Ghahari, Svetlana N. Myartseva & Enrique Ruíz- Cancino Accepted: 23-07-2013 ABSTRACT Aphelinidae is one of the most important families in biological control of insect pests at a worldwide level. The following catalogue of the Iranian fauna of Aphelinidae includes a list of all genera and species recorded for the country, their distribution in and outside Iran, and known hosts in Iran. In total 138 species from 11 genera (Ablerus, Aphelinus, Aphytis, Coccobius, Coccophagoides, Coccophagus, Encarsia, Eretmocerus, Marietta, Myiocnema, Pteroptrix) are listed as the fauna of Iran. Aphelinus semiflavus Howard, 1908 and Coccophagoides similis (Masi, 1908) are new records for Iran. Key words: Hymenoptera, Chalcidoidea, Aphelinidae, Catalogue. Shaaban Abd-Rabou Plant Protection Research 1. Introduction Institute, Agricultural Research Aphelinid wasps (Hymenoptera: Chalcidoidea: Aphelinidae) are important in nature, Center, Dokki-Giza, Egypt. especially in the population regulation of hemipterans on many different plants.These [E-mail: [email protected]] parasitoid wasps are also relevant in the biological control of whiteflies, soft scales and aphids [44] Hassan Ghahari . Studies on this family have been done mainly in relation with pests of fruit crops as citrus Department of Plant Protection, and others. John S. Noyes has published an Interactive On-line Catalogue [78] which includes Shahre Rey Branch, Islamic Azad up-to-date published information on the taxonomy, distribution and hosts records for the University, Tehran, Iran. Chalcidoidea known throughout the world, including more than 1300 described species in 34 [E-mail: [email protected]] genera at world level. -
Twenty Three Species of Platygastrinae (Hymenoptera: Platygastridae) New to the Fauna of Poland
Acta entomologica silesiana Vol. 26: (online 016): 1–7 ISSN 1230-7777, ISSN 2353-1703 (online) Bytom, April 4, 2018 Twenty three species of Platygastrinae (Hymenoptera: Platygastridae) new to the fauna of Poland http://doi.org/10.5281/zenodo.1212271 PETER NEERUP BUHL1, Paweł Jałoszyński2 1 Troldhøjvej 3, DK-3310 Ølsted, Denmark, e-mail: [email protected] 2 Muzeum Przyrodnicze Uniwersytetu Wrocławskiego, ul. Sienkiewicza 21, 50-335 Wrocław, e-mail: [email protected] ABSTRACT. Twenty three species of Platygastrinae (Hymenoptera: Platygastridae) new to the fauna of Poland. New distributional records of twenty three species of Platygastrinae (Hymenoptera: Platygastridae) are given, all reported for the first time from Poland: Gastrotrypes caudatus Brues, Leptacis coryphe BUHL, Platygaster betularia kieffer, P. damokles (BUHL), P. frater BUHL, P. germanica BUHL, P. gracilipes HUGGERT, P. microsculpturata BUHL, P. philinna walker, P. robiniae Buhl & Duso, P. signata (foerster), P. soederlundi BUHL, P. splendidula RUTHE, P. striatithorax BUHL, P. varicornis BUHL, Prosactogaster erdosi szelenyi, Synopeas convexum thomson, S. doczkali BUHL, S. fungorum BUHL, S. jasius (walker), S. noyesi BUHL, S. osaces (walker) and Trichacis pisis (walker). The new records increase the number of Platygastrinae known to occur in Poland to 124 species. KEY WORDS: Hymenoptera, Platygastroidea, Platygastridae, Platygastrinae; faunistics, new records, Poland. INTRODUCTION Since the synopsis of GarBarczyk (1997), who listed from Poland 56 species of Platygastrinae (i.e., Platygastridae excluding Scelionidae and Sceliotrachelidae, as accepted by most authors today), a substantial progress has been made in the faunistic study of this group of tiny parasitoid wasps. Two species, Synopeas bialowiezaensis BUHL, 2005 and Platygaster polonica Buhl & Jałoszyński, 2016a were described based on specimens known only from Poland, and Inostemma kaponeni BUHL, 2005 was described from Finland and Poland. -
Whirligig Mite Anystidae
The species collected in your Malaise trap are listed below. They are organized by group and are listed in the order of the 'Species Image Library'. ‘New’ refers to species that are brand new to our DNA barcode library. 'Rare' refers to species that were only collected in your trap out of all 64 that were deployed for the program. BIN Group (Scientific Name) Species Common Name Scientific Name New Rare BOLD:ABW5642 Mites (Arachnida) Whirligig mite Anystidae BOLD:AAD7604 Beetles (Coleoptera) Lady beetle Coleomegilla BOLD:AAB5640 Beetles (Coleoptera) Multicolored Asian lady beetle Harmonia axyridis BOLD:AAH0130 Beetles (Coleoptera) Clover weevil Sitona hispidulus BOLD:AAF3428 Beetles (Coleoptera) Click beetle Melanotus BOLD:ABA6320 Beetles (Coleoptera) Pleasing fungus beetle Erotylidae BOLD:AAH0256 Beetles (Coleoptera) Minute brown scavenger beetle Corticarina BOLD:AAG5778 Beetles (Coleoptera) Sap-feeding beetle Glischrochilus BOLD:ABA6362 Beetles (Coleoptera) Rove beetle Amischa decipiens BOLD:ABW2870 Beetles (Coleoptera) Rove beetle Meronera venustula BOLD:AAL5087 Beetles (Coleoptera) Large rove beetle Philonthus cognatus BOLD:ABX2484 Beetles (Coleoptera) Crab-like rove beetle Tachyporus atriceps BOLD:ACL5932 Beetles (Coleoptera) Ocellate rove beetle Omaliinae BOLD:ACK6363 Beetles (Coleoptera) Rove beetle Staphylinidae BOLD:AAN6148 Beetles (Coleoptera) False metallic wood-boring beetle Trixagus carinicollis BOLD:AAG4782 Flies (Diptera) Leaf miner fly Cerodontha longipennis BOLD:AAM6324 Flies (Diptera) Leaf miner fly Pseudonapomyza europaea BOLD:AAL7534 Flies (Diptera) Wood gnat Sylvicola BOLD:AAH3035 Flies (Diptera) Cluster fly Pollenia BOLD:AAP2825 Flies (Diptera) Cluster fly Pollenia BOLD:AAN5178 Flies (Diptera) Gall midge Cecidomyiidae BOLD:ACX8294 Flies (Diptera) Gall midge Cecidomyiidae BOLD:AAN5148 Flies (Diptera) Biting midge Forcipomyia BOLD:AAB7030 Flies (Diptera) Midge Chironomus BOLD:AAG0994 Flies (Diptera) Midge Corynoneura BOLD:AAI8449 Flies (Diptera) Midge Heterotrissocladius sp. -
ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000). -
A Short History Regarding the Taxonomy and Systematic Researches of Platygastroidea (Hymenoptera)
Memoirs of the Scientific Sections of the Romanian Academy Tome XXXIV, 2011 BIOLOGY A SHORT HISTORY REGARDING THE TAXONOMY AND SYSTEMATIC RESEARCHES OF PLATYGASTROIDEA (HYMENOPTERA) O.A. POPOVICI1 and P.N. BUHL2 1 “Al.I.Cuza” University, Faculty of Biology, Bd. Carol I, nr. 11, 700506, Iasi, Romania. 2 Troldhøjvej 3, DK-3310 Ølsted, Denmark, e-mail: [email protected],dk Corresponding author: [email protected] This paper presents an overview of the most important and best-known works that were the subject of taxonomy or systematics Platygastroidea superfamily. The paper is divided into three parts. In the first part of the research surprised the early period can be placed throughout the XIXth century between Latreille and Dalla Torre. Before this period, references about platygastrids and scelionids were made by Linnaeus and Schrank, they are the ones who described the first platygastrid and scelionid respectively. In this the first period work entomologists as: Haliday, Westwood, Walker, Forster, Ashmead, Thomson, Howard, etc., the result of their work being the description of 699 scelionids species which are found quoted in Dalla Torre's catalogue. The second part of the paper is devoted to early 20th century. This vibrant work is marked by the work of two great entomologists: Kieffer and Dodd. In this period one publish the first and only global monograph of platygastrids and scelionids until now. In this monograph are twice the number of species than in Dalla Torre's catalogue which shows the magnitude of the systematic research of those moments. The third part of the paper refers to the late 20th and early 21st century. -
Diptera, Drosophilidae) in an Atlantic Forest Fragment Near Sandbanks in the Santa Catarina Coast Bruna M
08 A SIMPÓSIO DE ECOLOGIA,GENÉTICA IX E EVOLUÇÃO DE DROSOPHILA 08 A 11 de novembro, Brasília – DF, Brasil Resumos Abstracts IX SEGED Coordenação: Vice-coordenação: Rosana Tidon (UnB) Nilda Maria Diniz (UnB) Comitê Científico: Comissão Organizadora e Antonio Bernardo de Carvalho Executora (UFRJ) Bruna Lisboa de Oliveira Blanche C. Bitner-Mathè (UFRJ) Bárbara F.D.Leão Claudia Rohde (UFPE) Dariane Isabel Schneider Juliana Cordeiro (UFPEL) Francisco Roque Lilian Madi-Ravazzi (UNESP) Henrique Valadão Marlúcia Bonifácio Martins (MPEG) Hilton de Jesus dos Santos Igor de Oliveira Santos Comitê de avaliação dos Jonathan Mendes de Almeida trabalhos: Leandro Carvalho Francisco Roque (IFB) Lucas Las-Casas Martin Alejandro Montes (UFRP) Natalia Barbi Chaves Victor Hugo Valiati (UNISINOS) Pedro Henrique S. F. Gomes Gustavo Campos da Silva Kuhn Pedro Henrique S. Lopes (UFMG) Pedro Paulo de Queirós Souza Rogério Pincela Mateus Renata Alves da Mata (UNICENTRO) Waira Saravia Machida Lizandra Jaqueline Robe (UFSM) Norma Machado da Silva (UFSC) Gabriel Wallau (FIOCRUZ) IX SEGED Introdução O Simpósio de Ecologia, Genética e Evolução de Drosophila (SEGED) é um evento bianual que reúne drosofilistas do Brasil e do exterior desde 1999, e conta sempre com uma grande participação de estudantes. Em decorrência do constante diálogo entre os diversos laboratórios, os encontros têm sido muito produtivos para a discussão de problemas e consolidação de colaborações. Tendo em vista que as moscas do gênero Drosophila são excelentes modelos para estudos em diversas áreas (provavelmente os organismos eucariotos mais investigados pela Ciência), essas parcerias podem contribuir também para o desenvolvimento de áreas aplicadas, como a Biologia da Conservação e o controle biológico da dengue. -
(Hymenoptera: Platygastridae). III
© Entomologica Fennica. 17 June 2009 New or little known Palaearctic species of Platygastrinae (Hymenoptera: Platygastridae). III. Peter Neerup Buhl Buhl, P. N. 2009: New or little known Palaearctic species of Platygastrinae (Hy- menoptera: Platygastridae). III. — Entomol. Fennica 20: 65–83. Fifteen new species of Platygastrinae are described and their affinities discussed, viz. Platygaster alpina sp.n.(Austria),P. bayankhongorensis sp. n. (Mongolia), P. breviscapa sp.n.(Croatia),P. kazakhstanica sp. n. (Kazakhstan), P. nove- marticulata sp. n. (Mongolia), P. platygaster sp.n.(Kyrgyzstan),P. schwarzi sp. n. (Austria), P. sylveni sp.n.(Sweden),P. viklundi sp.n.(Sweden),Pyrgaspis striativentris sp. n. (Mongolia), Synopeas latvianum sp.n.(Latvia),S. micro- sculpturatum sp. n. (Mongolia), S. pinnei sp. n. (Latvia), S. schwarzi sp. n. (Mon- golia), and S. wangsjoi sp. n. (Sweden). The hitherto unknown females of Platygaster aberrans Buhl, 1998 and Synopeas hansseni Buhl, 1998, and the hitherto unknown male of Synopeas acutispinus Buhl, 1998, are described. P. N. Buhl, Troldhøjvej 3, DK-3310 Ølsted, Denmark; E-mail: pnbuhl @hotmail.com Received 23 January 2007, accepted 13 December 2007 1. Introduction dard website about the group with an annotated bibliography etc. can be found at http://atbi. In the past 15 years I have described about 150 biosci.ohio-state.edu:210/hymenoptera/eol_sce- new Palaearctic species of Platygastridae, nearly lionidae.home, under the editorship of N. F. John- all of them in the subfamily Platygastrinae, in son. which all species are koinobiont egg-larval para- This paper is a continuation of my previous sitoids on Cecidomyiidae (Diptera). And again, papers (Buhl 2003, 2005). -
Assemblage of Hymenoptera Arriving at Logs Colonized by Ips Pini (Coleoptera: Curculionidae: Scolytinae) and Its Microbial Symbionts in Western Montana
University of Montana ScholarWorks at University of Montana Ecosystem and Conservation Sciences Faculty Publications Ecosystem and Conservation Sciences 2009 Assemblage of Hymenoptera Arriving at Logs Colonized by Ips pini (Coleoptera: Curculionidae: Scolytinae) and its Microbial Symbionts in Western Montana Celia K. Boone Diana Six University of Montana - Missoula, [email protected] Steven J. Krauth Kenneth F. Raffa Follow this and additional works at: https://scholarworks.umt.edu/decs_pubs Part of the Ecology and Evolutionary Biology Commons Let us know how access to this document benefits ou.y Recommended Citation Boone, Celia K.; Six, Diana; Krauth, Steven J.; and Raffa, Kenneth F., "Assemblage of Hymenoptera Arriving at Logs Colonized by Ips pini (Coleoptera: Curculionidae: Scolytinae) and its Microbial Symbionts in Western Montana" (2009). Ecosystem and Conservation Sciences Faculty Publications. 33. https://scholarworks.umt.edu/decs_pubs/33 This Article is brought to you for free and open access by the Ecosystem and Conservation Sciences at ScholarWorks at University of Montana. It has been accepted for inclusion in Ecosystem and Conservation Sciences Faculty Publications by an authorized administrator of ScholarWorks at University of Montana. For more information, please contact [email protected]. 172 Assemblage of Hymenoptera arriving at logs colonized by Ips pini (Coleoptera: Curculionidae: Scolytinae) and its microbial symbionts in western Montana Celia K. Boone Department of Entomology, University of Wisconsin, -
Biological-Control-Programmes-In
Biological Control Programmes in Canada 2001–2012 This page intentionally left blank Biological Control Programmes in Canada 2001–2012 Edited by P.G. Mason1 and D.R. Gillespie2 1Agriculture and Agri-Food Canada, Ottawa, Ontario, Canada; 2Agriculture and Agri-Food Canada, Agassiz, British Columbia, Canada iii CABI is a trading name of CAB International CABI Head Offi ce CABI Nosworthy Way 38 Chauncey Street Wallingford Suite 1002 Oxfordshire OX10 8DE Boston, MA 02111 UK USA Tel: +44 (0)1491 832111 T: +1 800 552 3083 (toll free) Fax: +44 (0)1491 833508 T: +1 (0)617 395 4051 E-mail: [email protected] E-mail: [email protected] Website: www.cabi.org Chapters 1–4, 6–11, 15–17, 19, 21, 23, 25–28, 30–32, 34–36, 39–42, 44, 46–48, 52–56, 60–61, 64–71 © Crown Copyright 2013. Reproduced with the permission of the Controller of Her Majesty’s Stationery. Remaining chapters © CAB International 2013. All rights reserved. No part of this publication may be reproduced in any form or by any means, electroni- cally, mechanically, by photocopying, recording or otherwise, without the prior permission of the copyright owners. A catalogue record for this book is available from the British Library, London, UK. Library of Congress Cataloging-in-Publication Data Biological control programmes in Canada, 2001-2012 / [edited by] P.G. Mason and D.R. Gillespie. p. cm. Includes bibliographical references and index. ISBN 978-1-78064-257-4 (alk. paper) 1. Insect pests--Biological control--Canada. 2. Weeds--Biological con- trol--Canada. 3. Phytopathogenic microorganisms--Biological control- -Canada. -
Hymenoptera: Braconidae: Microgastrinae) Comb
Revista Brasileira de Entomologia 63 (2019) 238–244 REVISTA BRASILEIRA DE Entomologia A Journal on Insect Diversity and Evolution www.rbentomologia.com Systematics, Morphology and Biogeography First record of Cotesia scotti (Valerio and Whitfield, 2009) (Hymenoptera: Braconidae: Microgastrinae) comb. nov. parasitising Spodoptera cosmioides (Walk, 1858) and Spodoptera eridania (Stoll, 1782) (Lepidoptera: Noctuidae) in Brazil a b a a Josiane Garcia de Freitas , Tamara Akemi Takahashi , Lara L. Figueiredo , Paulo M. Fernandes , c d e Luiza Figueiredo Camargo , Isabela Midori Watanabe , Luís Amilton Foerster , f g,∗ José Fernandez-Triana , Eduardo Mitio Shimbori a Universidade Federal de Goiás, Escola de Agronomia, Setor de Entomologia, Programa de Pós-Graduac¸ ão em Agronomia, Goiânia, GO, Brazil b Universidade Federal do Paraná, Setor de Ciências Agrárias, Programa de Pós-Graduac¸ ão em Agronomia – Produc¸ ão Vegetal, Curitiba, PR, Brazil c Universidade Federal de São Carlos, Programa de Pós-Graduac¸ ão em Ecologia e Recursos Naturais, São Carlos, SP, Brazil d Universidade Federal de São Carlos, Departamento de Ecologia e Biologia Evolutiva, São Carlos, SP, Brazil e Universidade Federal do Paraná, Departamento de Zoologia, Curitiba, PR, Brazil f Canadian National Collection of Insects, Ottawa, Canada g Universidade de São Paulo, Escola Superior de Agricultura “Luiz de Queiroz”, Departamento de Entomologia e Acarologia, Piracicaba, SP, Brazil a b s t r a c t a r t i c l e i n f o Article history: This is the first report of Cotesia scotti (Valerio and Whitfield) comb. nov. in Brazil, attacking larvae of the Received 3 December 2018 black armyworm, Spodoptera cosmioides, and the southern armyworm, S.