Taxonomy, Distribution and Ecology of Bolboschoenus in Europe

Total Page:16

File Type:pdf, Size:1020Kb

Taxonomy, Distribution and Ecology of Bolboschoenus in Europe Ann. Bot. Fennici 44: 81–102 ISSN 0003-3847 Helsinki 26 April 2007 © Finnish Zoological and Botanical Publishing Board 2007 Taxonomy, distribution and ecology of Bolboschoenus in Europe Zdenka Hroudová1, Petr Zákravský1, Michal Ducháček2 & Karol Marhold3,4 1) Institute of Botany, Academy of Sciences of the Czech Republic, CZ-252 43 Průhonice, Czech Republic (e-mail: [email protected], [email protected]) 2) National Museum, Department of Botany, CZ-252 43 Průhonice, Czech Republic (e-mail: [email protected]) 3) Institute of Botany, Slovak Academy of Sciences, Dúbravská cesta 14, SK-845 23 Bratislava, Slovak Republic (e-mail: [email protected]) 4) Department of Botany, Charles University, Benátská 2, CZ-128 01 Praha, Czech Republic Received 15 July 2005, revised version received 24 May 2006, accepted 29 Aug. 2006 Hroudová, Z., Zákravský, P., Ducháček, M. & Marhold, K. 2007: Taxonomy, distribution and ecol- ogy of Bolboschoenus in Europe. — Ann. Bot. Fennici 44: 81–102. The five species of Bolboschoenus — B. glaucus, B. laticarpus, B. maritimus, B. planiculmis and B. yagara — occurring in Europe were studied. A detailed taxonomic and nomenclatural account is provided for these taxa, together with an identification key, distribution data and maps based on a revision of herbarium specimens from 36 herbaria and on field observations. Because ecological differentiation influences the distribution of the taxa concerned, detailed information on their ecology is also pro- vided. Key words: Bolboschoenus, Cyperaceae, distribution, Europe, nomenclature, tax- onomy Introduction ican (Browning et al. 1995) and Australasian (Browning et al. 1997) taxa. They reported B. The taxonomy of Bolboschoeneus (Cyperaceae; yagara to be a new taxon for Europe, as well as formerly included in Scirpus) has advanced con- its putative hybrid with B. maritimus (Browning siderably over the last decades. DeFilipps (1980) et al. 1996). treated the taxa now placed in Bolboschoenus Browning and Gordon-Gray (2000) provided within the broadly conceived genus Scirpus. an overview of the Bolboschoenus fruit morphol- Since the treatment by DeFilipps (1980), who ogy. Hroudová and her co-authors dealt with the recognized in Europe only Scirpus maritimus. inflorescence structure (Hroudová et al. 1998a), with two subspecies (subsp. maritimus and fruit anatomy (Hroudová et al. 1997, 1998b), subsp. affinis), the taxonomy has been stud- growth response to different trophic conditions ied extensively in different parts of the world. (Zákravský & Hroudová 1996), chromosome Browning and her co-authors dealt in detail with numbers (Jarolímová & Hroudová 1998) and the South African (Browning & Gordon-Gray ecological differentiation related to plant com- 1993, 1999, Browning et al. 1998), North Amer- munities (Hroudová et al. 1999a) in the central 82 Hroudová et al. • ANN. BOT. FENNICI Vol. 44 European taxa. They reported B. glaucus as a B, BIL, BP, BRA, BRNM, BRNU, GJO, GLM, new species for the Czech Republic (Hroudová GZU, JE, KL, KRA, KRAM, LBL, LD, LE, LI, et al. 1999b) and described a new species, B. LISI, LOD, M, MSB, P, PR, PRC, SAV, SLO, laticarpus (Marhold et al. 2004), corresponding SO, SOM, SZCZ, TRN, UGDA, W, WA, WRSL, to the previously reported alleged hybrid of B. WU, and ZV. Specimens from the private col- yagara and B. maritimus. Hroudová et al. (2001) lections of K. F. Günther, H. Manitz, H. Melzer, and Hroudová (2002) reported for the first time H.-J. Zündorf, specimens collected by us in for central Europe B. koshewnikowii (currently the Czech Republic, Slovakia, Hungary, Austria, synonymous with B. planiculmis; see Egorova & Poland, Germany and Portugal, and specimens Tatanov 2003). collected by our colleagues, namely O. Clever- For the area of the former Soviet Union ing, E. Collias, R. Hrivnák, M. Janišová, V. Egorova and Tatanov (2003) provided detailed Jarolímová, Z. Kaplan, P. Koutecký, F. Krahulec, data on B. planiculmis, and Tatanov (2003a) on W. Lazowski, L. Moravcová, J. Sádlo and J. B. yagara. They also solved some nomenclatural Štěpánková were also studied. The localities problems concerning these two taxa. Tatanov from the Czech Republic included in the maps (2003b) mapped the distribution of B. glaucus were found during a detailed study of the dis- in parts of eastern Europe that belonged to tribution of Bolboschoenus species in the coun- the former Soviet Union. Additionally, Tatanov try (Ducháček 2002, Ducháček et al. 2006). (2003c) provided a short account of Bolbosch- Selected localities representing the distribution oenus in the former Soviet Union, evaluated the of Bolboschoenus species in European countries taxonomic importance of the carpological char- are given in the Appendix. Centres of distribu- acters (Tatanov 2004a), and proposed an infra- tion as well as individual isolated localities are generic systematic arrangement of the genus included. A full list of herbarium specimens (Tatanov 2004b). Nevertheless, a comprehensive studied is available on request from the first treatment of Bolboschoenus for Europe (or Eura- author. In all lists only the localities, from which sia) is still lacking and relevant data are scattered the herbarium specimens with fruiting plants throughout numerous separate papers. were available are included (determination of The present paper aims to provide a syn- specimens without fruits is not reliable). Addi- thesis of the taxonomy and nomenclature of tionally, more detailed surveys of the occurrence Bolboschoenus in Europe (excluding B. affinis of Bolboschoenus in some European countries (= B. popowii), occurring in Europe only mar- were published with full lists of known locali- ginally, in Russia; see Egorova 1976), together ties; such surveys were made for Austria (Hrou- et al. et al. with the distribution data based on revision of dová 2006) and Poland (Hroudová 2005), and are in preparation for Germany, Slo- herbarium specimens and studies of plants in the vakia and Hungary. field. Because ecological differentiation plays Morphological and anatomical characters of a major role in the expansion of plants in new the studied species, and their quantitative ranges, localities and in their disappearance from some are based on measurements of plants from natu- areas, detailed data on the ecology of the taxa are ral populations from the Czech Republic, Slo- also provided. Our study concentrates on central vakia and Hungary, and of cultivated plants Europe, thus the distribution data for the other (results published by Hroudová et al. 1997, parts of Europe are not presented in detail; nev- 1998a, 1998b, 1999a, 1999b) and also on the ertheless, even the scattered data on their occur- studied herbarium material. rence provide important information reflecting their general European distribution pattern. Identification key to European species of Bolboschoenus Material and methods 1. Inflorescence branched, consisting of a central group of The distribution maps are based on the exam- sessile spikelets and of (1–)2–7(–12) rays bearing single ined specimens kept in the following herbaria: spikelets or fascicles of spikelets; rays mostly more ANN. BOT. FENNICI Vol. 44 • Taxonomy, distribution and ecology of Bolboschoenus in Europe 83 than twice longer than sessile spikelets; total number of spikelets on rays higher than or same as number of sessile spikelets; perianth bristles mostly persistent on ripe fruits; fruits (achenes) trigonous or convex with slightly developed edge on abaxial side; trigonous in cross section (rarely nearly flattened or slightly convex to subtrigonous on abaxial side); outer layer of pericarp (exocarp) thinner than mesocarp, formed by isodiametric or slightly elongated cells not always air-filled; achene surface smooth or with fine network of cell outlines (at 20¥ magnification) ....................................................... 2 1. Inflorescence head-like or branched, consisting of a central group of sessile spikelets and of 1–2(–4) rays bearing single spikelets or fascicles of spikelets; rays usually less than twice as long as sessile spikelets; total number of spikelets on rays lower than number of sessile spikelets; perianth bristles caducous; fruits (achenes) not trigonous, but concave, flattened, convex to subtrigonous on abaxial side; biconcave, lenticular, plano-convex to subtrigonous in cross section; outer layer of pericarp (exocarp) thicker than or as thick as mesocarp, formed by elongated cylindrical cells always air-filled; achene surface with a highly visible polygonal network of cell outlines (at 20¥ magnification) .................................... 4 2. Achenes mostly with a clear edge on abaxial side, trigonous in cross section; exocarp layer thin but visibly developed; floral scales never brownish-red or deep pur- plish-red ....................................................................... 3 Fig. 1. Bolboschoenus glaucus (from Košíře, Prague, 2. Achenes with only slightly developed edge on abaxial Czech Republic, (collected in 1983 and cultivated in side, plano-convex to subtrigonous in cross section; Průhonice), 2003 Z. Hroudová, PRA). — a: Inflores- exocarp slightly visible; floral scales frequently brown- cence. — b: Achene (abaxial view). — c: Achene ish-red or deep purplish-red .......................... B. glaucus cross-section. — d: Pericarp layers (ex = exocarp, m = 3. Achenes narrow (1.6–1.8 mm wide), equilaterally trigo- mesocarp, en = endocarp). nous in cross-section; exocarp very thin, formed of more-or-less isodiametric cells .....................
Recommended publications
  • Bolboschoenus Glaucus (Lam.) S.G. Smith, a New Species in the Flora of the Ancient Near East
    Veget Hist Archaeobot DOI 10.1007/s00334-011-0305-3 ORIGINAL ARTICLE Bolboschoenus glaucus (Lam.) S.G. Smith, a new species in the flora of the ancient Near East Miche`le M. Wollstonecroft • Zdenka Hroudova´ • Gordon C. Hillman • Dorian Q. Fuller Received: 5 October 2010 / Accepted: 23 May 2011 Ó Springer-Verlag 2011 Abstract Taxonomic advancement in the genus Bolbo- Bolboschoenus in present-day Turkey, indicating that it has schoenus (Cyperaceae, formerly included in the genus a long history of occurrence in this region. The environ- Scirpus) have resulted in the re-classification of the plant mental, ecological and economic implications of this new previously known as Bolboschoenus maritimus (synonym information suggest that it is entirely feasible that this plant Scirpus maritimus) into several closely-related but distinct provided late Pleistocene and Holocene Near Eastern Bolboschoenus species This improved taxonomy is of people with a dependable and possibly a staple food source. importance for archaeobotanical investigations of ancient sites within the temperate zones, where this genus fre- Keywords Bolboschoenus glaucus Á Epipalaeolithic Á quently occurs, because it allows more precise definitions Near East Á Neolithic Á Taxonomy Á Nutlet characteristics of the ecological requirements and growing habits of each species. Moreover, it details the distinct morphological and anatomical characteristics of the fruit (nutlets) of each Introduction species. Using these new nutlet classification criteria, we re-examined charred archaeological specimens which had Bolboschoenus maritimus (sea club-rush) is a semi-aquatic previously been identified as B. maritimus (or S. mariti- species of the Cyperaceae that produces edible nutlets, mus), from five Near Eastern late Pleistocene and early tubers and shoots (Fig.
    [Show full text]
  • Phylogeny of Abildgaardieae (Cyperaceae) Inferred from ITS and Trnl–F Data Kioumars Ghamkhar University of New England, Armidale, New South Wales, Australia
    Aliso: A Journal of Systematic and Evolutionary Botany Volume 23 | Issue 1 Article 12 2007 Phylogeny of Abildgaardieae (Cyperaceae) Inferred from ITS and trnL–F Data Kioumars Ghamkhar University of New England, Armidale, New South Wales, Australia Adam D. Marchant Royal Botanic Gardens, Sydney, New South Wales, Australia Karen L. Wilson Royal Botanic Gardens, Sydney, New South Wales, Australia Jeremy J. Bruhl University of New England, Armidale, New South Wales, Australia Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation Ghamkhar, Kioumars; Marchant, Adam D.; Wilson, Karen L.; and Bruhl, Jeremy J. (2007) "Phylogeny of Abildgaardieae (Cyperaceae) Inferred from ITS and trnL–F Data," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 23: Iss. 1, Article 12. Available at: http://scholarship.claremont.edu/aliso/vol23/iss1/12 Aliso 23, pp. 149–164 ᭧ 2007, Rancho Santa Ana Botanic Garden PHYLOGENY OF ABILDGAARDIEAE (CYPERACEAE) INFERRED FROM ITS AND trnL–F DATA KIOUMARS GHAMKHAR,1,2,4 ADAM D. MARCHANT,2 KAREN L. WILSON,2 AND JEREMY J. BRUHL1,3 1Botany, Centre for Ecology, Evolution, and Systematics, University of New England, Armidale, New South Wales 2351, Australia; 2National Herbarium of New South Wales, Royal Botanic Gardens, Sydney, Mrs Macquaries Road, Sydney, New South Wales 2000, Australia 3Corresponding author ([email protected]) ABSTRACT Within the tribe Abildgaardieae, the relationships between Fimbristylis and its relatives have not been certain, and the limits of Fimbristylis have been unclear, with Bulbostylis and Abildgaardia variously combined with it and each other.
    [Show full text]
  • Outline of Angiosperm Phylogeny
    Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese
    [Show full text]
  • Ifps Business Congress (Ipc) / Page 2: Presentation of New Ifps C (Ipc)
    PPAALLYYNNOOSS ISSN 0256-1670 Volume 41 (1) – 2018 NEWSLETTER OF THE INTERNATIONAL FEDERATION OF PALYNOLOGICAL SOCIETIES 15TH INTERNATIONAL CONTENT OF PAL YNOS VOL. 40 (2) PALYNOLOGICAL P PAGE 1: IFPS BUSINESS CONGRESS (IPC) / PAGE 2: PRESENTATION OF NEW IFPS C (IPC) / COUNCILLORS XITH IOPC PAGE 3: IFPS SO CIETIES’ REPORTS PAGE 8: FUTURE MEETINGS JOINT MEETING, PRAGUE, PAGE 9: PALYNO LOGICAL HOMAGE PAGE 10: NEWS F ROM PALYNOLOGY CZECH REPUBLIC, RELATED FIELDS PAGE 19: CURRENT IFPS AFFILIATED SEPTEMBER 12–19, 2020 SOCIETIES AND COUNCILLORS PAGE 20: IMPRESSUM We would like to invite you to the XVth International Palynological Congress and XIth International Organization of Palaeobotany Conference from September 12th to 19th 2020 in Prague, Czech Republic. An international joint meeting of the whole community is dedicated to the 200th Anniversary of Modern Palaeobotany, as IIFFPPSS BBUSINESS 1820 was the starting point of palaeobotanical nomenclature infered from the the first issue of “Flora der Vorwelt” the principal work of the Czech “Father of Palaeobotany“ Kaspar Maria von Sternberg. Palynology and Palaeobotany have a long tradition in the Czech and Slovak Republics 1820 – 2020: 200 YEARS OF with several famous pioneers including 1820 – 2020: 200 Kaspar Maria von Sternberg, Karl and Otakar MODERN PALAEOBOTANY Feistmantels, August Joseph Corda, Dionýz Štúr, Karel Bořivoj Presl, as well as world wide known Czech palynologists and palaeobotanists of the modern era e.g. Blanka Pacltová, Milada Vavrdová, Zlatko Kvaček, František Němejc,
    [Show full text]
  • December 2012 Number 1
    Calochortiana December 2012 Number 1 December 2012 Number 1 CONTENTS Proceedings of the Fifth South- western Rare and Endangered Plant Conference Calochortiana, a new publication of the Utah Native Plant Society . 3 The Fifth Southwestern Rare and En- dangered Plant Conference, Salt Lake City, Utah, March 2009 . 3 Abstracts of presentations and posters not submitted for the proceedings . 4 Southwestern cienegas: Rare habitats for endangered wetland plants. Robert Sivinski . 17 A new look at ranking plant rarity for conservation purposes, with an em- phasis on the flora of the American Southwest. John R. Spence . 25 The contribution of Cedar Breaks Na- tional Monument to the conservation of vascular plant diversity in Utah. Walter Fertig and Douglas N. Rey- nolds . 35 Studying the seed bank dynamics of rare plants. Susan Meyer . 46 East meets west: Rare desert Alliums in Arizona. John L. Anderson . 56 Calochortus nuttallii (Sego lily), Spatial patterns of endemic plant spe- state flower of Utah. By Kaye cies of the Colorado Plateau. Crystal Thorne. Krause . 63 Continued on page 2 Copyright 2012 Utah Native Plant Society. All Rights Reserved. Utah Native Plant Society Utah Native Plant Society, PO Box 520041, Salt Lake Copyright 2012 Utah Native Plant Society. All Rights City, Utah, 84152-0041. www.unps.org Reserved. Calochortiana is a publication of the Utah Native Plant Society, a 501(c)(3) not-for-profit organi- Editor: Walter Fertig ([email protected]), zation dedicated to conserving and promoting steward- Editorial Committee: Walter Fertig, Mindy Wheeler, ship of our native plants. Leila Shultz, and Susan Meyer CONTENTS, continued Biogeography of rare plants of the Ash Meadows National Wildlife Refuge, Nevada.
    [Show full text]
  • C6 Noncarice Sedge
    CYPERACEAE etal Got Sedge? Part Two revised 24 May 2015. Draft from Designs On Nature; Up Your C 25 SEDGES, FOINS COUPANTS, LAÎCHES, ROUCHES, ROUCHETTES, & some mostly wet things in the sedge family. Because Bill Gates has been shown to eat footnotes (burp!, & enjoy it), footnotes are (italicized in the body of the text) for their protection. Someone who can spell caespitose only won way has know imagination. Much of the following is taken verbatim from other works, & often not credited. There is often not a way to paraphrase or rewrite habitat or descriptive information without changing the meaning. I am responsible for any mistakes in quoting or otherwise. This is a learning tool, & a continuation of an idea of my friend & former employer, Jock Ingels, LaFayette Home Nursery, who hoped to present more available information about a plant in one easily accessible place, instead of scattered though numerous sources. This is a work in perpetual progress, a personal learning tool, full uv misstakes, & written as a personal means instead of a public end. Redundant, repetitive, superfluous, & contradictory information is present. It is being consolidated. CYPERACEAE Sauergrasgewächse SEDGES, aka BIESIES, SEGGEN Formally described in 1789 by De Jussieu. The family name is derived from the genus name Cyperus, from the Greek kupeiros, meaning sedge. Many species are grass-like, being tufted, with long, thin, narrow leaves, jointed stems, & branched inflorescence of small flowers, & are horticulturally lumped with grasses as graminoids. Archer (2005) suggests the term graminoid be used for true grasses, & cyperoid be used for sedges. (If physical anthropologists have hominoids & hominids, why don’t we have graminoids & graminids?) There are approximately 104 genera, 4 subfamilies, 14 tribes, & about 5000 species worldwide, with 27 genera & 843 species in North America (Ball et al 2002).
    [Show full text]
  • Ficha Catalográfica Online
    UNIVERSIDADE ESTADUAL DE CAMPINAS INSTITUTO DE BIOLOGIA – IB SUZANA MARIA DOS SANTOS COSTA SYSTEMATIC STUDIES IN CRYPTANGIEAE (CYPERACEAE) ESTUDOS FILOGENÉTICOS E SISTEMÁTICOS EM CRYPTANGIEAE CAMPINAS, SÃO PAULO 2018 SUZANA MARIA DOS SANTOS COSTA SYSTEMATIC STUDIES IN CRYPTANGIEAE (CYPERACEAE) ESTUDOS FILOGENÉTICOS E SISTEMÁTICOS EM CRYPTANGIEAE Thesis presented to the Institute of Biology of the University of Campinas in partial fulfillment of the requirements for the degree of PhD in Plant Biology Tese apresentada ao Instituto de Biologia da Universidade Estadual de Campinas como parte dos requisitos exigidos para a obtenção do Título de Doutora em Biologia Vegetal ESTE ARQUIVO DIGITAL CORRESPONDE À VERSÃO FINAL DA TESE DEFENDIDA PELA ALUNA Suzana Maria dos Santos Costa E ORIENTADA PELA Profa. Maria do Carmo Estanislau do Amaral (UNICAMP) E CO- ORIENTADA pelo Prof. William Wayt Thomas (NYBG). Orientadora: Maria do Carmo Estanislau do Amaral Co-Orientador: William Wayt Thomas CAMPINAS, SÃO PAULO 2018 Agência(s) de fomento e nº(s) de processo(s): CNPq, 142322/2015-6; CAPES Ficha catalográfica Universidade Estadual de Campinas Biblioteca do Instituto de Biologia Mara Janaina de Oliveira - CRB 8/6972 Costa, Suzana Maria dos Santos, 1987- C823s CosSystematic studies in Cryptangieae (Cyperaceae) / Suzana Maria dos Santos Costa. – Campinas, SP : [s.n.], 2018. CosOrientador: Maria do Carmo Estanislau do Amaral. CosCoorientador: William Wayt Thomas. CosTese (doutorado) – Universidade Estadual de Campinas, Instituto de Biologia. Cos1. Savanas. 2. Campinarana. 3. Campos rupestres. 4. Filogenia - Aspectos moleculares. 5. Cyperaceae. I. Amaral, Maria do Carmo Estanislau do, 1958-. II. Thomas, William Wayt, 1951-. III. Universidade Estadual de Campinas. Instituto de Biologia. IV. Título.
    [Show full text]
  • Witaj Und 2Plus – Eine Herausforderung Für Die Zukunft Witaj a 2Plus – Wužadanje Za Pr ˇichod
    wužadanjewužadanje ˇichod zaza prprˇichodˇichod Serbske šulske eine Heraus- towarstwo z. t. Sorbischer forderung Schulverein e. V. Witaj a 2plus – wužadanje za pr für die Zukunft Witaj a 2plus Witaj a 2plus Witaj und 2plus und Witaj Witaj und 2plus – eine Herausforderung für die Zukunft 08-1582 Schulverein 17.03.2009 6:57 Uhr Seite 1 Maientanz in der sorbischen Kinder- tagesstätte «Dr. Jurij M¶ynk» Ralbitz 08-1582 Schulverein 20.03.2009 8:15 Uhr Seite 2 Serbske [ulske towarstwo z.t. Sorbischer Schulverein e.V. 08-1582 Schulverein 20.03.2009 8:16 Uhr Seite 3 wu{adanje za p≠ichod + eine Heraus- forderung für die Zukunft 08-1582 Schulverein 16.03.2009 15:40 Uhr Seite 4 «Witaj a 2plus – wu{adanje za p≠ichod» b| tema cy¶odnjow- skeje mjezynarodneje fachoweje konferency Serbskeho [ulskeho towarstwa z.t. a Sakskeho statneho ministerstwa za kultus, kotra{ wotm| so 12. apryla 2008 w Chrósåicach. Na njej wobd≤|lichu so n|hd≤e 270 hornjo- a delnjoserb- skich zajimcow, star[i, kub¶arki, wu∑erki a wu∑erjo, studen- åa, zastupjerjo serbskich institucijow a rozd≤|lnych politi- skich gremijow, mjez nimi zapós¶an∑a Zwjazkoweho sejma Marja Micha¶kowa (CDU), zapós¶ancy Sakskeho krajneho sejma Astrid Günther-Schmidt (Zwjazk 90/Zeleni), Marko ]iman (CDU), Hajko Kozel (L|wica), statny sekretar w Sakskim ministerstwje za kultus Hansjörg König, p≠edsyda Domowiny Jan Nuk a dal[i. Postrowne s¶owa p≠ednjesechu zapós¶anc Marko ]iman, statny sekretar Hansjörg König a p≠edsyda Jan Nuk. B| to mjeztym t≠eåa w|domostno-kub- ¶anska konferenca, kotru{ je Serbske [ulske towarstwo p≠e- wjed¶o.
    [Show full text]
  • Native Plant List, Pdf Format
    Appendix A: City of Bellingham Native Plant List December 2020 The City of Bellingham Native Plant List (Figure 1) includes plant species that are native to Bellingham watersheds (Figure 2). The native plant list applies to all habitat types, including riparian, upland, and wetland areas. The list was developed using specimen records from the Consortium of Pacific Northwest Herbaria and Bellingham plant checklists curated by Don Knoke, a volunteer at the University of Washington Herbarium. To improve plant establishment and protect the genetic resources of our local plant populations, the City recommends using native plants that were grown from seeds or cuttings collected from the Puget Trough Ecoregion (Figure 3). Obtaining native plants grown from material collected from the Puget Trough Ecoregion will help ensure the plants are adapted to the unique environmental conditions of Bellingham watersheds and are genetically similar to our local plant populations. A more thorough discussion of the rational and selection process is provided in the City of Bellingham Public Works Department Native Plant Materials Selection Guidelines, December 2020. Figure 1. City of Bellingham Native Plant List Ferns Common Name Scientific Name Family Bracken fern Pteridium aquilinum var. pubescens Dennstaedtiaceae Bristle-like quillwort Isoetes tenella Isoetaceae Common horsetail Equisetum arvense Equisetaceae Deer fern Struthiopteris spicant (Blechnum spicant) Blechnaceae Dream fern Aspidotis densa Pteridaceae Giant horsetail Equisetum telmateia ssp. braunii
    [Show full text]
  • District Heating in Lusatia
    Brandenburg University of Technology Cottbus-Senftenberg District Heating in Lusatia Status Quo and Prospects for Climate Neutrality Fernwärme in der Lausitz Status quo und Perspektiven für Klimaneutralität Thesis for the degree of Master of Science in Environmental and Resource Management submitted by Jan Christian Bahnsen 1st Examiner: Prof. Dr. Bernd Hirschl 2nd Examiner: Katharina Heinbach Submitted: 29th May 2020 Statement of Authentication I hereby declare that I am the sole author of this master thesis and that I have not used any other sources other than those listed in the bibliography and identified as references. I further declare that I have not submitted this thesis at any other institution in order to obtain a degree. The content, either in full or in part, has not been previously submitted for grading at this or any other academic institution. ________________________________ _____________________________________ (Place, Date) (Signature) Abstract The master thesis at hand examines the potential of district heating in Lusatia. The thesis follows the approach of first identifying technical and economic potentials in general and then transferring them to the study region. For the quantitative determination of district heating potential in Lusatia, the status quo is determined and a GIS-based analysis is carried out with regard to minimum heat demand densities. The extent to which district heating is suitable for climate-neutral heat supply will be investigated using the potential of renewable and waste heat energy sources. Furthermore, the regional economic effects of developing these potentials are examined. The results show that despite an overall decline in heat demand, there is potential to increase the relative share of district heating in Lusatia.
    [Show full text]
  • The Vascular Flora of Rarău Massif (Eastern Carpathians, Romania). Note Ii
    Memoirs of the Scientific Sections of the Romanian Academy Tome XXXVI, 2013 BIOLOGY THE VASCULAR FLORA OF RARĂU MASSIF (EASTERN CARPATHIANS, ROMANIA). NOTE II ADRIAN OPREA1 and CULIŢĂ SÎRBU2 1 “Anastasie Fătu” Botanical Garden, Str. Dumbrava Roşie, nr. 7-9, 700522–Iaşi, Romania 2 University of Agricultural Sciences and Veterinary Medicine Iaşi, Faculty of Agriculture, Str. Mihail Sadoveanu, nr. 3, 700490–Iaşi, Romania Corresponding author: [email protected] This second part of the paper about the vascular flora of Rarău Massif listed approximately half of the whole number of the species registered by the authors in their field trips or already included in literature on the same area. Other taxa have been added to the initial list of plants, so that, the total number of taxa registered by the authors in Rarău Massif amount to 1443 taxa (1133 species and 310 subspecies, varieties and forms). There was signaled out the alien taxa on the surveyed area (18 species) and those dubious presence of some taxa for the same area (17 species). Also, there were listed all the vascular plants, protected by various laws or regulations, both internal or international, existing in Rarău (i.e. 189 taxa). Finally, there has been assessed the degree of wild flora conservation, using several indicators introduced in literature by Nowak, as they are: conservation indicator (C), threat conservation indicator) (CK), sozophytisation indicator (W), and conservation effectiveness indicator (E). Key words: Vascular flora, Rarău Massif, Romania, conservation indicators. 1. INTRODUCTION A comprehensive analysis of Rarău flora, in terms of plant diversity, taxonomic structure, biological, ecological and phytogeographic characteristics, as well as in terms of the richness in endemics, relict or threatened plant species was published in our previous note (see Oprea & Sîrbu 2012).
    [Show full text]
  • Raport Częściowy - Etap I, II I III
    Zweckverband Verkehrsverbund Oberlausitz-Niederschlesien Opracowanie Koncepcji Publicznego Transportu Pasażerskiego Komunikacji Regionalnej w korytarzu Bautzen - Gorlitz - Lubań - Jelenia Góra - Kamienna Góra w celu rozwoju turystyki w Euroregionie Neisse - Nisa - Nysa Raport częściowy - etap I, II i III Wydanie | 13 września 2012 Niniejszy raport uwzględnia instrukcje i wskazówki naszego Klienta i w związku z tym nie jest on przeznaczony dla osób trzecich. Zrzekamy się odpowiedzialności z tytułu używania niniejszego raportu przez osoby trzecie. Nr projektu 223412 Ove Arup & Partners International Ltd Sp. z o. o. Oddział w Polsce ul. Królewska 16 00-103 Warszawa Poland www.arup.com IVAS - Ingenieurbüro für Verkehrsanlagen und -systeme Alaunstraße 9 01099 Dresden (Neustadt) Deutschland / Germany www.ivas-ingenieure.de Weryfikacja dokumentu Nazwa projektu Opracowanie Koncepcji Publicznego Transportu Nr projektu Pasażerskiego Komunikacji Regionalnej w 223412 korytarzu Bautzen - Görlitz - Lubań - Jelenia Góra - Kamienna Góra w celu rozwoju turystyki w Euroregionie Neisse - Nisa - Nysa Nazwa dokumentu Raport częściowy - etap I, II i III Numer pliku w katalogu Numer katalogu Weryfikacja Data Nazwa pliku Report 2012_06_25_JSa.docx Draft 1 25 cze Opis First draft 2012 Przygotowany przez Sprawdzony przez Zatwierdzony przez Joanna Sarbiewska, Nazwisko Dirk Ohm Krzysztof Pogłód Martin Schüffler Podpis Wydanie 31 lip Nazwa pliku Report 2012_07_31_PL_final.docx 2012 Opis Przygotowany przez Sprawdzony przez Zatwierdzony przez Joanna Sarbiewska, Nazwisko
    [Show full text]