Tr. J. of Botany 23 (1999) 97–112 © TÜBİTAK Research Article
Lichen Flora of Trabzon*
Kenan YAZICI Karadeniz Technical University, Fatih Education Faculty, Biology Dept., 61335 Akçaabat, Trabzon-TURKEY
Received: 09.10.1997 Accepted: 19.11.1998
Abstract: This study was performed between May 1993 and November 1995 in Trabzon. A total of 230 species and 1 variety, which belongs to the 35 families in Ascomycotina, were defined. A total of 39 species were understood to be new records for Turkey.
Key Words: Lichen, Flora, Trabzon, Turkey
Trabzon İli Likenleri Özet: Bu çalışma Mayıs 1993–Kasım 1995 tarihleri arasında Trabzon il sınırları içerisinde gerçekleştirilmiştir. Ascomycotina sınıfı içerisinde 35 familyaya ait 230 tür ve 1 varyete tesbit edilmiştir. Bu türlerin 39’u Türkiye için yeni kayıttır.
Anahtar Sözcükler: Liken, Flora, Trabzon, Türkiye
Introduction Pseudomacchie and pure or mixed stands of Carpinus Although lichens are often treated as a separate plant L., Castanea Miller, Quercus L., Fagus L. and Picea Dietr. division most lichenologists now consider them part of were observed to be the dominant vegetation in the the fungi (1). Lichens have been used in monitoring district (11). surveys for radioactive pollution (2–4). After giving a collection number to each one of the The studies on lichens in Turkey are not as extensive samples collected, they were stored by putting them into as in other countries. So the lichen flora of Turkey is still paper purses and envelopes. largely unknown. In Trabzon, the first sudies on lichens A stereo microscope, light microscope and a color were performed by Steiner, Szatala and Anşin (5–7). The identification method using various reactives and flora studies on lichens in Trabzon are by Yazıcı and Gönülol books were used in the identification of the samples (8–10). This paper supplements these studies. (12–14). The lichen species defined as herbarium specimens Materials and Methods were placed in the herbarium of the Biology Department, Fatih Education Faculty, Karadeniz Technical University. The lichen samples were collected from 166 different stations in 1993–1995 in Trabzon. The study area is situated between lat 40°47’–41°07’ N and long Results 39°08’–40°28’ E (Figure 1). A total of 230 species and 1 variety belonging to the In the study area, there are many high plateaus and 35 families in Ascomycotina were defined. An asterisks streams flowing to the Black Sea. A lot of the lower fields (*) indicates a new record for Trabzon Province, while near the sea are agricultural areas. Especially, Maçka has double asterisks (**) indicates a nev record for Turkey. a richer plant cover than other districts.
* This study is a summary of my PhD thesis.
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41°07’ Çarşıbaşı BLACK SEA Trabzon Vakfıkebir Arsin Araklı Of Beşikdüzü Akçaabat Yomra GİRESUN Sürmene Şalpazarı Hayrat Düzköy Köprübaşı Tonya Maçka RİZE Çaykara 40°30’ 39°08’
N 0510 GÜMÜŞHANE km. 40°47’
TURKEY
Figure 1. Study area.
List of Taxa On calcareous rock. 1990 m.; Köprübaşı: Yangın yaylası, 23.09.1993, 40°48’17’’ N 48°08’00’’ E, * Acarospora fuscata (Nyl.) Arnold ** Alectoria sarmentosa (Ach.) Ach. 1650 m.; Sürmene: Lişit yaylası, Çaykara: Sultanmurat yaylası, Maçka: Şolma yaylası, 24.08.1994, 26.06.1995, 40°46’30’’ N 40°04’33’’ E, 18.08.1995, 40°45’00’’ N 40° 11’40’’ E, 40°52’45’’ N 39°33’00’’ E, 1665m.; 1725 m.; Hayrat: Sarıçiçek yaylası, 1935 m.; Şalpazarı: Sisdağı yaylası, Akçaabat: Kayabaşı yaylası, 24.08.1995, 04.09.1993, 40°51’20’’ N 40°28’30’’ E, 28.07.1995, 40°50’14’’ N 39°10’00’’E, 40°51’10’’ N 39°29’46’’ E, 1850 m. Yazıcı 1705 m., Büyükharman yaylası, 29.07.1995, 1850 m. Yazıcı 0334. 0330. 40°55’00’’ N 40°24’45’’ E, 1645 m., On calcareous rock. On Picea orientalis. Anabarda yaylası, 08.09.1993, 40°52’33’’ N 40°24’30’’ E, 1720 m. Yazıcı 0351. * Acarospora impressula Th. Fr. Anaptychia ciliaris (L.) Körb. ex A. Maçka: Kadırga yaylası, 21.07.1994, Massal. On Picea orientalis, Fagus orientalis and 40°43’10’’ N 39°23’00’’ E, 1950 m. Yazıcı Carpinus sp. Maçka: Hamsiköyü yaylası, 10.06.1993, 0341. 40°40’43’’ N 39°28’40’’ E, 1610 m. Zal * Anaptychia setifera Mereschk. ex Räs. On calcareous rock. yaylası, 03.08.1994, 40°41’’25’’ N Maçka: Hamsiköyü yaylası, 25.06.1993, ** Acarospora sinopica (Wahlenb.) 39°25’53’’ E, 1750 m., Kiraz yaylası, 40°40’43’’ N 39°28’40’’ E, 1610 m., Hanzar Körber 12.07.1995, 40°40’40’’ N 39°32’45’’ E, yaylası, 23.07.1995, 40°50’00’’ N Maçka: Çakırgöl dağı yaylası, 1695 m.; Akçaabat: Büyükoba yaylası, 39°30’02’’ E, 1760 m.; Tonya: Şarlı yaylası, 11.07.1995, 40°40’33’’ N 39°41’50’’ E, 11.08.1993, 40°53’25’’ N 39°31’46’’ E, 17.07.1993, 40°43’47’’ N 39°19’30’’ E, 2500 m. Yazıcı 0342. 1795 m.; Tonya: Karaptal yaylası, 1813 m.; Köprübaşı: İsmailağa yaylası, 14.08.1995, 40°52’15’’ N 39°16’38’’ E, 15.07.1993, 40°41’53’’ N 40°08’33’’ E, On siliceous rock. 1780 m., Akkise yaylası, 18.08.1993, 1790 m Yazıcı 0504. * Acarospora umbilicata Bagl. 40°44’39’’ N 39°20’20’’ E, 1830 m.; Araklı: On Picea orientalis and Fagus orientalis. Trabzon: Beşirli–Toklu köyü border, Erikli yaylası, 13.08.1993, 40°38’30’’ N 10.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 40°02’17’’ E, 2000 m., Üstinki yaylası, Aspicilia caesiocinerea (Nyl. ex Malbr.) m. Yazıcı 0320. 29.07.1994, 40°37’58’’ N 40°00’25’’ E, Arnold
98 K. YAZICI
Maçka: Altındere Valley Zalişki Yaylası, On siliceous rock. On Robinia pseudoacacia, Populus sp., 15.05.1995, 40°47’30’’ N 39°39’35’’ E, * Bryoria bicolor (Ehrh.) Brodo & D. Salix sp., Acer sp. and siliceous rock. 1500 m., Kozağaç köyü, 12.07.1995, Hawksw. Caloplaca decipiens (Arnold) Blomb. & 40°50’23’’ N 39°37’00’’ E, 850 m.; Fross Akçaabat: Helvacı köyü, 04.07.1994, Akçaabat: Hıdırnebi yaylası, 08.08.1993, 40°01’18’’ N 39°31’23’’ E, 230 m.; Trabzon: 40°58’55’ N 39°24’40’’ E, 1540 m., Trabzon: Akyazı köyü, 04.09.1995, Akyazı köyü, 04.09.1995, 41°01’30’’ N Kayabaşı yaylası, 24.08.1995, 40°51’10’’ N 41°01’30’’ N 39°36’04’’ E, 300 m.; 39°36’04’’ E, 300 m.; Aralıklı: Pervane köyü, 39°29’46’’ E, 1850 m.; Çaykara: Sıcakoba Şalpazarı: İzmiks yaylası, 09.07.1995, 16.08.1994, 40°55’02’’ N 40°00’10’’ E, yaylası, 19.08.1995, 40°46’57’’ N 40°56’30’’ N 39°12’00’’ E, 1400 m.; 1300 m., Hasköy, 14.10.1995, 40°57’40’’ N 40°10’50’’ E, 1930 m., Uzungöl yaylası, Beşikdüzü: Bozlu köyü, 19.09.1995, 40°00’27’’ E, 490 m.; Of: Kıyıboyu köyü, 28.08.1993, 40°44’37’’ N 40°19’30’’ E, 40°59’16’’ N 39°12’24’’ E, 410 m.; Araklı: 12.04.1994, 41°04’28’’ N 40°15’33’’ E, 250 1690 m.; Hayrat: Tuh yaylası, 27.07.1993, Dağbaşı Köyü, 05.10.1995, 40°48’06’’ N m.; Arsin: Çatak, 02.06.1994, 40°53’49’’ N 40°47’05’’ N 40°26’30’’ E, 1750 m. Düzköy: 39°55’38’’ E, 1550 m.; Çarşıbaşı: Kale köyü, 39°55’01’’ E, 700 m., Üçpınar köyü, Gürgendağ yaylası, 04.08.1993, 40°51’13’’ 05.07.1995, 41°04’28’’ N 39°20’30’’ E, 250 08.06.1995, 40°50’18’’ N 39°53’58’’ E, 870 N 39°24’45’’ E, 1700 m. Yazıcı 0400. m.; Arsin: Harmanlı köyü, 28.07.1993, m. Yazıcı 0323. On Picea orientalis. 41°00’04’’ N 39°54’05’’ E, 350 m., Çiçekli köyü, 07.08.1995, 41°OO’02’’ N 39°57’30’’ On siliceous and calcareous rock. * Bryoria fuscescens (Gyel.) Brodo & D. E, 500 m. Yazıcı 0411. Aspicilia calcarea (L.) Mudd Hawksw. On calcareous rock. Maçka: Kozağaç köyü, 17.07.1995, Maçka: Kulindağı yaylası, 15.07.1995, Caloplaca ferruginea (Hudson) Th. Fr. 40°50’23’’ N 39°37’00’’ E, 850 m.; Trabzon: 40°52’26’’ N 39°33’40’’ E, 1700 m.; Tonya: Beşirli–Toklu köyü border, 26.11.1995, Erikbeli yaylası, 28.08.1995, 40°43’45’’ N Trabzon: Dolaylı köyü, 11.09.1993, 41°01’37’’ N 39°38’08’’ E, 130 m., 39°21’03’’ E, 1850 m.; Şalpazarı: Geyikli 40°59’25’’ N 39°44’05’’ E, 350 m.; Karadeniz Technical University Campus, köyü, 01.08.1995, 40°52’47’’ N 39°12’30’’ Beşikdüzü: Bozlu köyü, 19.09.1995, 19.05.1993, 41°03’20’’ N 39°43’35’’ E, 130 E, 1310 m.; Akçaabat: Büyükoba yaylası, 40°59’16’’ N 39°12’24’’ E, 410 m.; Araklı: m.; Vakfıkebir: Güney köyü, 23.04.1995, 11.08.1993, 40°53’25’’ N 39°31’46’’ E, Dağbaşı köyü, 05.10.1995, 40°48’06’’ N 41°01’00’’ N 39°16’20’’ E, 400 m. Yazıcı 1795 m. Yazıcı 0324. 39°55’38’’ E, 1550 m.; Of: Koruluk köyü, 0326. On Picea orientalis. 04.09.1994, 41°01’27’’ N 40°17’52’’ E, 500 m. Yazıcı 0409. On calcareous rock. * Calicium viride Pers. * Caloplaca flavescens (Huds.) J.R. Aspicilia cinerea (L.) Körb. Maçka: Güzel yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ E, 1945 m. Yazıcı Laundon Trabzon: Beşirli–Toklu köyü border, 0420. Trabzon: center, 27.03.1995, 41°03’13’’ 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 N 39°40’50’’ E, 50 m., Ayasofya Church m., Karadeniz Technical University Campus, On Picea orientalis. Garden, 09.05.1993, 41°03’17’’ N 19.05.1993, 41°03’20’’ N 39°43’35’’ E, 130 * Caloplaca arenaria (Pers.) Müll. Arg. 39°39’50’’ E, 90 m.; Of: Uğurlu köyü, m.; Of: Gürpınar köyü, 21.09.1993, Akçaabat: Bozdoğan köyü, 10.05.1994, 19.09.1993, 40°58’36’’ N 40°18’00’’ E, 650 40°59’29’’ N 40°17’47’’ E, 700 m. Yazıcı 40°59’55’’ N 39°28’45’’ E, 350 m. Yazıcı m. Yazıcı 0357. 0328. 0481. On calcareous rock, dyke wall and brick. On siliceous and calcareous rock. On siliceous rock. * Caloplaca flavorubescens (Huds.) J.R. Aspicilia contorta (Hoffm.) Krempelh. * Caloplaca cerina (Ehrh. ex Hedwig) Laundon Trabzon: Beşirli–Toklu köyü border, Th. Fr. 26.11.1995, 41°01’47’’ N 39°38’08’’ E, 130 Şalpazarı: Sisdağı yaylası, 28.07.1995, Trabzon: Gürbulak köyü, 12.10.1995, m. Yazıcı 0501. 40°50’14’’ N 39°10’00’’ E, 1850 m. Yazıcı 41°00’13’’ N 39°36’09’’ E, 400 m., 0361. On calcareous rock. Karadeniz Technical University Campus, On Fagus orientalis. * Aspicilia laevata (Ach.) Arnold 13.03.1995, 41°03’20’’ N 39°43’35’’ E, 130 Trabzon: Beşirli–Toklu köyü border, m.; Çarşıbaşı: Kovanlı yaylası, 28.06.1995, Caloplaca flavovirescens (Wulf.) Dalla 26.11.1995, 41°01’47’’ N 39°38’08’’ E, 130 41°04’00’’ N 39°22’32’’ E, 850 m. Yazıcı Torre & Sarn. m. Yazıcı 0338. 0388. Maçka: Altındere Valley Altındere District, On calcareous rock. On Acer sp., siliceous and calcareous 13.08.1995, 40°46’51’’ N 39°40’23’’ E, rock. 1200 m.; Trabzon: Karadeniz Technical * Baeomyces rufus (Hudson) Rebent. Caloplaca citrina (Hoffm.) Th. Fr. University Campus, 21.10.1993, 41°03’20’’ Maçka: Altındere Valley Sumela N 39°43’35’’ E, 130 m. Yazıcı 0362. Monastery and surroundings, 02.05.1993 Maçka: Altındere Valley Sumela 40°46’25’’ N 39°39’15’’ E, 1200 m., Monastery and surroundings, 24.05.1993, On siliceous and calcareous rock. Hamsiköyü yaylası, 27.06.1993, 40°40’43’’ 40°46’25’’ N 39°39’15’’ E, 1200 m.; * Caloplaca holocarpa (Hoffm.) Wade N 39°28’40’’ E, 1610 m. Yazıcı 331. Beşikdüzü: Ağaçlı köyü, 01.09.1995, Akçaabat: Helvacı köyü, 04.07.1994, 41°00’04’’ N 39°11’57’’ E, 390 m.; On siliceous rock, stone and soil. 40°01’18’’ N 39°31’23’’ E, 230 m.; Trabzon: Vakfıkebir: Bozonü yaylası, 10.07.1993, Ayasofya Church Garden, 09.05.1993, * Brodoa intestiniformis (Vill.) Goward 40°56’45’’ N 39°21’04’’ E, 1100 m., 41°03’17’’ N 39°39’50’’ E, 90 m.; Hayrat: Maçka: Çakırgöl dağı yaylası, Esentepe, 21.10.1995, 41°00’56’’ N Ovacık köyü, 11.04.1995, 41°01’17’’ N 09.08.1995, 40°40’33’’ N 39°41’50’’ E, 39°14’28’’ E, 400 m. Yazıcı 0429. 2500 m. Yazıcı 0585. 40°23’45’’ E, 650 m. Yazıcı 0325.
99 Lichen Flora of Trabzon
On calcareous rock. Şalpazarı: Alaca yaylası, 17.08.1993, Şalpazarı, Tonya, Köprübaşı, Hayrat, Çaykara, 40°49’40’’ N 39°11’57’’ E, 1800 m. Yazıcı Akçaabat and Düzköy. Yazıcı 0602. * Caloplaca irrubescens (Arnold) Zahlbr. 0484. On mosses and soil. Maçka: Kozağaç köyü, 17.07.1995, On soil. 40°50’23’’ N 39°37’00’’ E, 850 m. Yazıcı Cetraria nivalis (L.) Ach. 0491. * Catapyrenium squamulosum (Ach.) Maçka: Altındere Valley Aksu Yaylası, Breuss On siliceous and calcareous rock. 20.06.1994, 40°45’58’’ N 39°39’50’’ E, Çaykara: Sultanmurat yaylası, 1650 m., Ardıçlı yaylası, 22.08.1995, * Caloplaca teicholyta (Ach.) J. Steiner 18.08.1995, 40°45’00’’ N 40°11’40’’ E, 40°47’32’’ N 39°36’40’’ E, 1620 m., Trabzon: Karadeniz Technical University 1935 m.; Şalpazarı: Kızılkaya yaylası, Lapazan yaylası, 16.06.1995, 40°41’35’’ N Campus, 13.08.1994, 41°03’20’’ N 14.06.1995, 40°46’15’’ N 39°12’00’’ E, 39°35’43’’ E, 1710 m., Zarışa yaylası, 39°43’35’’ E, 130 m., Center, 18.03.1995, 1650 m. Yazıcı 0479. 07.06.1994, 40°47’37’’ N 39°32’50’’ E, 41°03’13’’ N 39°40’50’’ E, 70 m. Yazıcı 1695 m.; Tonya: Sazalan yaylası, On soil. 0492. 09.07.1993, 40°43’30’’ N 39°16’40’’ E, * Cetraria aculeata (Schreber) Fr. On wall, dyke wall, limestone and brick. 1870 m.; Çaykara: Çayıroba yaylası, It was found on plateaus higher than 21.06.1993, 40°41’45’’ N 40°20’35’’ E, ** Caloplaca thallincola (Wedd.) Du 1680 m. in Maçka, Akçabat, Sürmene, 1840 m., Büyük yayla, 27.06.1994, Rietz Şalpazarı, Tonya, Köprübaşı, Hayrat and 40°40’45’’ N 40°27’15’’ E, 1900 m.; Araklı: Çaykara: Sultanmurat yaylası, Düzköy. Sometimes it was seen together with Kaleron yaylası, 17.07.1994, 40°40’00’ N 18.08.1995, 40°45’00’’ N 40°11’40’’ E, Cetraria muricata. Yazıcı 0601. 39°59’57’’ E, 1795 m., Üstinki yaylası, 1935 m., Vartan yaylası, 15.08.1995, 29.07.1994, 40°37’58’’ N 40°00’25’’ E, * Cetraria chlorophylla (Willd.) Vain. 40°46’00’’ N 40°10’05’’ E, 1945 m. Yazıcı 1990 m.; Köprübaşı: Vizera yaylası, 0540. Maçka: Şolma yaylası, 11.08.1994, 06.07.1993, 40°46’55’’ N 40°07’28’’ E, 40°52’45’’ N 39°33’00’’ E, 1665 m; Düzköy: 1700 m. Yazıcı 0515. On siliceous rock. Doğankaya yaylası, 11.09.1995, 40°51’57’’ On soil. * Candelaria concolor (Dicks.) B. Steiner N 39°22’29’’ E, 1690 m.; Çaykara: Karastel * Cetraria pinastri (Scop.) Gray Beşikdüzü: Bozlu köyü, 19.09.1995, yaylası, 25.06.1994, 40°43’42’’ N 40°59’16’’ N 39°12’24’’ E, 410 m.; Araklı: 40°19’00’’ E, 1712 m.; Köprübaşı: Vizera Maçka: Akboran yaylası, 16.06.1993, Dağbaşı köyü, 05.10.1995, 40°48’06’’ N yaylası, 06.07.1993, 40°46’55’’ N 40°41’25’’ N 39°44’55’’ E, 1830 m., Şolma 39°55’38’’ E, 1550 m; Sürmene: Çamburnu, 40°07’28’’ E, 1700 m., Taşlı yaylası, yaylası, 11.08.1994, 40°52’45’’ N 30.08.1994, 41°01’38’’ N 40°10’30’’ E, 100 06.09.1993, 40°44’20’’ N 40°06’00’’ E, 39°33’00’’ E, 1665 m.; Tonya: Taksorobası m., Muratlı köyü, 18.03.1994, 40°59’42’’ N 1775 m., Yangın yaylası, 23.09.1993, yaylası, 12.07.1993, 40°50’12’’ N 40°05’15’’ E, 250 m.; Vakfıkebir: Bahadırlı, 40°48’17’’ N 40°08’00’’ E, 1650 m. Yazıcı 39°16’55’’ E, 1600 m.; Çaykara: Sıcakoba 25.10.1995, 41°01’32’’ N 39°15’30’’ E, 370 0433. yaylası, 19.08.1995, 40°46’57’’ N m., Fol stream, Vakfıkebir–Tonya road, On Picea orientalis and Fagus orientalis. 40°10’50’’ E, 1930 m.; Köprübaşı: Cerah 23.09.1995, 41°00’13’’ N 39°16’00’’ E, 700 yaylası, 29.08.1993, 40°44’28’’ N ** Cetraria delisei (Bory ex Schaer.) m.; Of: Uğurlu köyü, 19.09.1993, 40°09’45’’ E, 1750 m. Yazıcı 0519. 40°58’36’’ N 40°18’00’’ E, 650 m. Yazıcı It was defined to grow on soil, soily rocks On Picea orientalis. 0340. and mosses on plateaus higher than 1700 m. in Maçka, Akçaabat, Köprübaşı, Hayrat, ** Chaenotheca brunneola (Ach.) Müll. On Picea orientalis, Fagus orientalis, Tonya, Şalpazarı, Çaykara, Düzköy, Sürmene Arg. Carpinus sp., Populus sp. and Acer sp. and Araklı. Mostly it was seen together with Maçka: Hamsiköyü yaylası, 22.08.1993, Candelariella aurella (Hoffm.) Zahlbr. Cetraraia ericetorum. Yazıcı 0503. 40°40’43’’ N 39°28’40’’ E, 1610 m.; It was defined on siliceous and calcareous ** Cetraria ericetorum Opiz Akçaabat: Hıdırnebi yaylası, 08.08.1993, rocks between 10–2000 meters at all 40°58’55’’ N 39°24’40’’ E, 1540 m. Yazıcı It was defined on soil, soily rocks and stations. Sometimes it was seen together with 0589. mosses in fields higher than 1700 m. in Candelariella vitellina growing on siliceous and Maçka, Akçaabat, Köprübaşı, Hayrat, Tonya, On Picea orientalis. calcareous rocks. Yazıcı 507. Şalpazarı, Çaykara, Araklı, Şalpazarı and Chaenotheca chrysocephala (Turn. ex * Candelariella medians (Nyl.) A.L. Düzköy. Mostly it was seen together with Ach.) Th. Fr. Cetraria delisei. Yazızı 0449. Trabzon: Center, 08.03.1995, This species was found on the bark of 41°03’13’’ N 39°40’50’’ E, 70 m. Yazıcı Cetraria islandica (L.) Ach. Picea orientalis in fields higher than 1500 m. 0600. This species was commonly defined on in Maçka, Düzköy, Akçaabat, Tonya, Çaykara, On limestone. mosses and soil in regions higher than 1400 Şalpazarı, Aarklı, Köprübaşı, Sürmene and Hayrat. Yazıcı 0456. Candelariella vitellina (Hoffm.) Müll. m. in Maçka, Akçaabat, Sürmene, Araklı, Arg. Düzköy, Şalpazarı, Tonya, Şalpazarı and ** Chaenotheca furfuracea (L.) Tibell Köprübaşı. Yazıcı 0511. It was found on siliceous and calcareous Maçka: Hamsiköyü yaylası, 22.08.1993, rocks between 10–2000 meters at all ** Cetraria muricata (Ach.) Eckfeldt 40°40’43’’ N 39°28’40’’ E, 1610 m.; stations. Yazıcı 0607. Sometimes it was found together with Çaykara: Malbet yaylası, 30.06.1994, 40°46’03’’ N 40°20’10’’ E, 1695 m.; * Catapyrenium rufescens (Ach.) Cetraria aculeata on soily rocks on plateaus higher than 1700 m. in Maçka, Sürmene, Şalpazarı: Karaçam yaylası, 06.06.1995, Breuss 40°47’31’’ N 39°14’20’’ E, 1687 m.;
100 K. YAZICI
Sürmene: Çamburnu, 30.08.1994, On soil and mosses. On mosses and soil. 41°01’38’’ N 40°10’30’’ E, 100 m. Yazıcı Cladonia furcata (Huds.) Schrad. * Cladonia subcervicornis (Vain.) 0453. Kernst. Maçka: Altındere Valley Taşköprü yaylası, On living root of Picea orientalis, Acer 02.03.1995, 40°45’00’’ N 39°40’14’’ E, Trabzon: Beşirli–Toklu köyü border, sp., soil, calcareous rock. 1250 m.; Akçaabat: Kuruçam yaylası, 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 ** Cladonia botrytes (Hagen) Willd. 06.08.1993, 40°58’17’’ N 39°26’45’’ E, 800 m. Yazıcı 0513. m.; Çaykara: Göl yaylası, 19.07.1993, Maçka: Hamsiköyü yaylası, 22.08.1993, On soil. 40°42’00’’ N 40°11’40’’ E, 1805 m. Yazıcı 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı * Cladonia uncialis (L.) Weber ex Wigg. 0545. 0610. Maçka: Kozağaç köyü, 12.07.1995, On mosses. On soil. 40°50’23’’ N 39°37’00’’ E, 850 m. Yazıcı * Cladonia gracilis (L.) Willd. 0409. ** Cladonia cenotea (Ach.) Schaer. Trabzon: center, 25.03.1995, 41°03’13’’ On mosses. Akçaabat: Kirazlık yaylası, 24.08.1993, N 39°40’50’’ E, 135 m.; Çaykara: Sıcakoba 40°54’00’’ N 39°31’15’’ E, 1785 m.; Araklı: * Collema auriforme (With.) Coppins & yaylası, 19.08.1995, 40°46’57’’ N Hendek yaylası, 03.08.1993, 40°43’47’’ N J.R. Laundon 40°10’50’’ E, 1930 m. Yazıcı 0439. 39°53’28’’ E, 1850 m.; Sürmene: Harman It was commonly defined on mosses and yaylası, 19.06.1995, 40°49’38’’ N On mosses and soil. soily rocks in lower shaded fields near the sea 40°05’55’’ E, 1680 m. Yazıcı 0603. in Akçaabat, Tonya, Trabzon (center), * Cladonia macilenta Hoffm. Beşikdüzü, Araklı, Sürmene, Yomra, It was defined on a total of 15 plateaus Akçaabat: Enginoba yaylası, 26.08.1993, Vakfıkebir, Of and Arsin. From time to time it above 1600 m. in Maçka. 40°52’34’’ N 39°29’00’’ E, 1690 m., Kalecik was seen with Collema crispum. Yazıcı 0595. On decayed bark of Picea orientalis, yaylası, 22.07.1994, 40°53’55’’ N * Collema crispum (Huds.) Weber ex mosses and soil. 39°29’02’’ E, 1770 m. Yazıcı 0441. Wigg. Cladonia coccifera (L.) Willd. On decayed and dried bark of Picea Mostly defined on dyke wall, mosses, soil orientalis. Maçka: Şolma yaylası, 11.08.1994, and calcareous rock in lower fields near the 40°52’45’’ N 39°33’00’’ E, 1665 m. Yazıcı Cladonia pyxidata (L.) Hoffm. sea in Trabzon (center), Sürmene, Vakfıkebir, 0335. Beşikdüzü and Akçaabat. Yazıcı 0551. Maçka: Altındere Valley Sumela On decayed bark of Picea orientalis, Monastery and surroundings, 24.05.1993, * Collema cristatum (L.) Weber ex mosses and soil. 40°46’25’’ N 39°39’15’’ E, 1200 m., Mile Wigg. yaylası, 27.07.1994, 40°43’30’’ N Maçka: Kozağaç köyü, 17.07.1995, Cladonia coniocraea (Flk.) Spreng. 39°25’30’’ E, 1790 m., Lisevur yaylası, 40°50’23’’ N 39°37’00’’ E, 850 m.; This species was abundant on decayed 19.06.1993, 40°43’00’’ N 39°28’40’’ E, Akçaabat: Helvacı köyü, 04.07.1994, barks of Picea orientalis and mosses on 1700 m.; Düzköy: Honofter yaylası, 40°01’18’’ N 39°31’23’’ E, 230 m. Yazıcı plateaus above 1500 m. in Maçka, Çaykara, 21.08.1995, 40°48’35’’ N 39°23’15’’ E, 0509. Sürmene, Akçaabat and Düzköy. Yazıcı 0359. 1925 m., Derinoba yaylası, 05.08.1995, On calcareous rock. Cladonia digitata (L.) Hoffm. 40°46’14’’ N 39°22’08’’ E., 1920 m.; Tonya: Sivritepe yaylası, 04.09.1995, 40°53’41’’ N Collema flaccidum (Ach.) Ach. Akçaabat: Hıdırnebi yaylası, 08.08.1993, 39°16’10’’ E, 1100 m., Sazalan yaylası, Maçka: Hamsiköyü yaylası, 22.08.1993, 40°58’55’’ N 39°24’40’’ E, 1540 m.; 09.07.1993, 40°43’30’’ N 39°16’40’’ E, 40°40’43’’ N 39°28’40’’ E, 1610 m., Seloba Akçaabat: Ormana yaylası, 10.07.194, 1870 m.; Akçaabat: Kalecik yaylası, yaylası, 13.06.1994, 40°43’00’’ N 40°54’40’’ N 39°35’19’’ E, 1702 m.; 22.07.1994, 40°53’55’’ N 39°29’02’’ E, 39°39’25’’ E, 1793 m.; Çaykara: Çahmut Çaykara: Eğridere yaylası, 23.06.1994, 1770 m. Yazıcı 0343. yaylası, 21.07.1993, 1745 m., 40°48’40’’ N 40°48’57’’ N 40°20’30’’ E, 1640 m. Yazıcı 40°19’00’’ E, 1745 m.; Trabzon: Gölçayır On mosses and soil. 0370. köyü, 11.06.1995, 41°00’30’’ N 39°40’55’’ On decayed bark of Picea orientalis, Cladonia rangiferina (L.) Weber ex E, 500 m. Yazıcı 0500. Wigg. mosses and soil. On Carpinus sp., Ulmus sp., mosses and ** Cladonia floerkeana (Fr.) Flk. Maçka: Kozağaç köyü, 17.07.1995, siliceous rock. 40°50’23’’ N 39°37’00’’ E, 850 m., Yazıcı * Collema furfuraceum (Arnold) Du Maçka: Hıdırnebi yaylası, 08.08.1993, 0430. Rietz 40°58’55’’ N 39°24’40’’ E, 1540 m. Yazıcı 0483. On mosses and soil. Trabzon: Karlık köyü, 22.10.1994, 40°59’30’’ N 39°39’40’’ E, 580 m.; On deyaced bark of Picea orientalis and Cladonia rangiformis Hoffm. Vakfıkebir: Yaylacık köyü, 06.08.1995, soil. This is a very common species. It was 41°59’53’’ N 39°14’34’’ E, 390 m., Yazıcı Cladonia foliacea (Huds.) Willd. abundant on mosses and soil at altitudes of 0650. 100 to 2500 m at all stations. Yazıcı 0431. Akçaabat: Arpacılı köyü, 07.05.1993, On Carpinus sp. and siliceous rock. * Cladonia squamosa (Scop.) Hoffm. 40°58’30’’ N 39°28’09’’ E, 300 m.; Trabzon: * Collema limosum (Ach.) Ach. Beşirli–Toklu köyü border, 26.11.1995, Akçaabat: Hıdırnebi yaylası, 08.08.1993, Trabzon: Beşirli–Toklu köyü border, 41°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı 40°58’55’’ N 39°24’40’’ E, 1540 m., 26.11.1995, 41°01’37’’ N 39°38’08’’ E 130 0604. Kayabaşı yaylası, 24.08.1995, 40°51’10’’ N m. Yazıcı 0505. 39°29’46’’ E, 1850 m. Yazıcı 0401.
101 Lichen Flora of Trabzon
On mosses and calcareous rock. 08.06.1995, 40°50’18’’ N 39°53’58’’ E, 870 40°50’30’’ N 40°15’58’’ E., 700 m.; m. Yazıcı 0443. Trabzon: Karadeniz Technical University * Collema nigrescens (Huds.) DC. On mosses and soil. Campus, 21.10.1993, 41°03’20’’ N Trabzon: Beşirli–Toklu köyü border, 39°43’35’’ E, 130 m.; Akçaabat: Hıdırnebi 26.11.1995, N 41°01’37’’ N 39°38’08’’ E Diploschistes scruposus (Schreb.) yaylası, 08.08.1993, 40°58’55’’ N 130 m. Yazıcı 0360. Norm. 39°24’40’’ E, 1540 m.; Sürmene: Kal yaylası, On mosses and calcareous rock. Maçka: Altındere Valley Taşköprü yaylası, 26.08.1995, 40°47’26’’ N 40°05’27’’ E, 02.03.1995, 40°45’00’’ N 39°40’14’’ E, 1650 m.; Yomra: Lapazalık yaylası, Collema polycarpon Hoffm. 1250 m., Kozağaç köyü, 17.07.1995, 14.06.1993, 40°49’30’’ N 39°48’25’’ E, Çaykara: Koldere köyü, 17.08.1995, 40°50’23’’ N 39°37’00’’ E, 850 m.; 1530 m. Yazıcı 0565. 40°50’30’’ N 40°15’58’’ E, 790 m. Yazıcı Vakfıkebir: Fevziye köyü, 03.11.1995, On Fagus orientalis, Carpinus sp., 0363. 40°58’37’’ N 39°15’52’’ E, 350 m.; Arsin: Populus sp. and Juglans sp. Üçpınar köyü, 08.06.1995, 40°50’18’’ N On siliceous rock. ** Gyalecta jenesis (Batsch) Zahlbr. 39°53’58’’ E, 870 m. Yazıcı 0408. * Collema subnigrescens Degel. Maçka: Hamsiköyü yaylası, 22.08.1993, On silice rock. 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı Trabzon: Beşirli–Toklu köyü border, * Ephebe lanata (L.) Vain. 0567 m. 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı 0364. Çaykara: Sıcakoba yaylası, 19.08.1995, On calcareous rock. 40°46’57’’ N 40°10’50’’ E, 1930 m, Vartan Heterodermia speciosa (Wulfen) On calcareous rock. yaylası, 15.08.1995, 40°46’00’’ N Trevisan * Cornicularia normoerica (Gunn.) Du 40°10’05’’ E, 1945 m. Yazıcı 0460 m. Maçka: Şolma yaylası, 11.08.1994, Rietz On siliceous rock. 40°52’45’’ N 39°33’00’’ E, 1665 m.; Maçka: Kiraz yaylası, 12.07.1995, * Evernia divaricata (L.) Ach. Çaykara: Uzungöl yaylası, 19.07.1995, 40°40’40’’ N 39°32’45’’ E, 1695 m., Omela Maçka: Hamsiköyü yaylası, 22.08.1993, 40°44’37’’ N 40°19’30’’ E, 1690 m., Kuti yaylası, 28.08.1994, 40°43’45’’ N yaylası, 29.06.1994, 40°41’48’’ N 39°45’00’’ E, 1770 m.; Hayrat: Gama 40°40’43’’ N 39°28’40’’ E, 1610 m., Güzel yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ 40°22’40’’ E, 1750 m., Büyük yayla, yaylası, 27.07.1995, 40°49’42’’ N 27.06.1994, 40°40’45’’ N 40°27’15’’ E, 40°23’00’’ E, 1680 m. Yazıcı 0392. E, 1945 m.; Çaykara: Uzungöl yaylası, 19.07.1995, 40°44’37’’ N 40°19’30’’ E, 1900 m.; Şalpazarı: Çağmanlı yaylası, On siliceous rock. 1690 m. Yazıcı 0471. 04.06.1995, 40°49’19’’ N 39°15’00’’ E, 1630 m.; Düzköy: Gürgendağ yaylası, Dermatacarpon miniatum (L.) Mann. On Picea orientalis. 04.08.1993, 40°51’13’’ N 39°24’45’’ E, Maçka: Hamsiköyü yaylası, 22.08.1993, ** Evernia mesomorpha Nyl. 1700 m., Karadağ yaylası, 15.08.1993, 40°51’10’’ N 39°23’52’’ E, 1700 m. Yazıcı 40°40’43’’ N 39°28’40’’ E, 1610 m.; Maçka: Şolma yaylası, 11.08.1994, 0641. Akçaabat: Kemaliye köyü, 04.06.1994, 40°52’45’’ N 39°33’00’’ E, 1665 m., Güzel 40°58’53’’ N 39°26’49’’ E, 890 m.; Trabzon: yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ On Picea orientalis and mosses. Beşirli–Toklu köyü border, 19.11.1995, E, 1945 m. Yazıcı 0465. 41°01’37’’ N 39°38’08’’ E, 130 m.; Hayrat: ** Hymenelia lacustris (With) M. Choisy Gama yaylası, 27.07.1995, 40°49’42’’ N On Picea orientalis. Maçka: Hamsiköyü yaylası, 22.08.1993, 40°23’00’’ E, 1680 m.; Şalpazarı: Dorukkiriş Evernia prunastri (L.) Ach. 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı köyü, 28.06.1993, 40°52’45’’ N 39°12’20’’ 0579. It was commonly defined on Picea E, 1210 m.; Arsin: Çatak, 02.06.1994, orientalis and Quercus sp. in fields above 700 On siliceous rock. 40°53’49’’ N 39°55’01’’ E, 700 m. Yazıcı m. in Maçka, Akçaabat, Çaykara, Düzköy, 0399. ** Hypogymnia austerodes (Nyl.) Räs. Tonya, Şalpazarı, Araklı, Köprübaşı, Sürmene Maçka: Çakırgöl dağı yaylası, On siliceous and calcareous rock. and Hayrat. Yazıcı 0550. 09.08.1995, 40°40’33’’ N 39°41’50’’ E, * Diploschistes caesioplumbeus (Nyl.) Sometimes it was seen together with 2500 m. Yazıcı 0572. Vain. Ramalina farinacea especially in Maçka, On mosses and siliceous rock. Trabzon: Beşirli–Toklu köyü border, Düzköy, Şalpazarı and Araklı. * Hypogymnia farinacea Zopf. 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 * Fuscopannaria leucophea (Vahl.) P.M. m.; Vakfıkebir: Fevziye köyü, 03.11.1995, Jørg. Maçka: Çakırgöl dağı yaylası, 40°58’37’’ N 39°15’52’’ E, 350 m., Bahadırlı, 09.08.1995, 40°40’33’’ N 39°41’50’’ E, Maçka: Hamsiköyü yaylası, 22.08.1993, 2500 m.; Tonya: Karaptal yaylası, 25.10.1995, 40°01’32’’ N 39°15’30’’ E, 370 40°40’43’’ N 39°28’40’’ E, 1610 m., İpsila m.; Of: Gökçeoba köyü, 01.09.1994, 14.08.1995, 40°52’15’’ N 39°16’38’’ E, yaylası, 17.08.1994, 40°43’15’’ N 1780 m. Yazıcı 0580. 41°04’40’’ N 40°19’30’’ E, 400 m. Yazıcı 39°36’45’’ E, 1690 m.; Çaykara: Uzungöl 0496. yaylası, 19.07.1995, 40°44’37’’ N On siliceous and mosses. On calcareous rock. 40°19’30’’ E, 1690 m. Yazıcı 0560. Hypogymnia physodes (L.) Nyl. * Diploschistes muscorum (Scop.) R. On calcareous rock. It was found on the branches of Picea Sant. Graphis scripta (L.) Ach. orientalis in regions above 1500 m. in Maçka, Akçaabat, Sürmene, Hayrat, Şalpazarı, Maçka: Kaskar yaylası, 02.09.1993, Maçka: Güzel yayla, 20.08.1994, Çaykara, Akçaabat, Düzköy, Tonya, Araklı and 40°39’40’’ N 39°34’31’’ E, 1790 m., 40°52’35’’ N 39°34’55’’ E, 1945 m.; Köprübaşı. It is especially common in Maçka Kamadağ yaylası, 02.09.1994, 40°40’12’’ N Çaykara: Koldere köyü, 17.08.1995, and Akçaabat. Yazıcı 0445. 39°29’45’’ E, 1750 m.; Arsin: Üçpınar köyü,
102 K. YAZICI
Hypogymnia tubulosa (Schaer.) Hav. Lecanora muralis (Schreb.) Rabenh. 39°59’42’’ E, 380 m., Hasköy, 14.10.1995, 40°57’40’’ N 40°00’27’’ E, 490 m. Yazıcı Maçka: Hamsiköyü yaylası, 22.08.1993, It is a very common species and was 0609. 40°40’43’’ N 39°28’40’’ E, 1610 m., Güzel found on calcareous rocks in the fields at yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ altitudes of 100 to 1800 m. at all stations. Leptogium gelatinosum (With.) J.R. E, 1945 m., Fırınoba yaylası, 06.07.1995, Yazıcı 0437. Laundon 40°40’30’’ N 39°35’20’’ E, 1780 m.; Lecanora polytropa (Hoffm.) Rabenh. Maçka: Hamsiköyü yaylası, 27.06.1993, Akçaabat: Acısu yaylası, 28.05.1993, 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı 40°57’55’’ N 39°27’00’’ E, 750 m.; Araklı: Akçaabat: Kayabaşı yaylası, 24.08.1995, 0630. Pinalar yaylası, 20.08.1993, 40°42’56’’ N 40°51’10’’ N 39°29’46’’ E. 1850 m.; 39°54’28’’ E, 1825 m., Demirtaş yaylası, Şalpazarı: Sisdağı yaylası, 28.07.1995, On mosses. 40°50’14’’ N 39°10’00’’ E, 1850 m.; 22.06.1994, 40°42’05’’ 40°01’07’’ E, 1815 ** Leptogium hibernicum Mitch. ex P. Çaykara: Sultanmurat yaylası, 18.08.1995, m. Yazıcı 0355. Jørg 40°45’00’’ N 40°11’40’’ E, 1935 m; It is very common in fields above 1500 Çarşıbaşı: Kovanlı yaylası, 28.06.1995, Maçka: Kadırga yaylası, 21.07.1995, m. in Maçka and Akçaabat. 41°04’00’’ N 39°22’32’’ E, 850 m. Yazıcı 40°43’10’’ N 39°23’00’’ E, 1950 m.; On Picea orientalis. 0414. Akçaabat: Kayabaşı yaylası, 24.08.1995, 40°51’10’’ N 39°29’46’’ E, 1850 m. Yazıcı Hypogymnia vittata (Ach.) Gas. On siliceous rock. 0619. Maçka: Altındere Valley Sumela * Lecanora pulicaris (Pers.) Ach. Monastery and surroundings, 24.05.1993, On siliceous rock and mosses. Maçka: Altındere Valley Zalişki yaylası, 40°46’25’’ N 39°39’15’’ E, 1200 m.; Araklı: Leptogium lichenoides (L.) Zahlbr. Acısu yaylası, 01.08.1993, 40°46’42’’ N 07.05.1995, 40°47’30’’ N 39°39’35’’ E, 40°00’25’’ E, 1810 m.; Köprübaşı: Kangel 1500 m.; Düzköy: Derinoba yaylası, Maçka: Altındere Valley Altındere District, yaylası, 05.08.1994, 40°49’40’’ N 05.08.1995, 40°46’14’’ N 39°22’08’’ E, 13.08.1995, 40°46’51’’ N 39°40’23’’ E, 40°08’15’’ E, 1670 m.; Sürmene: Büyük 1920 m; Akçaabat: Hıdırnebi yaylası, 1200 m.; Düzköy: Beypınar yaylası, yayla, 23.06.1995, 40°47’33’’ N 40°03’45’’ 08.08.1993, 40°58’55’’ N 39°24’40’’ E, 25.08.1995, 40°47’30’’ N 39°23’10’’ E, E, 1700 m. Yazıcı 0391. 1540 m. Yazıcı 0520. 1700 m., Doğankaya yaylası, 11.09.1995, 40°51’57’’ N 39°22’29’’ E, 1600 m.; On Carpinus sp. On Picea orientalis. Akçaabat: Hıdırnebi yaylası, 08.08.1993, ** Icmadophila ericetorum (L.) Zahlbr. Lecidella elaeochroma (Ach.) M. Choisy 40°58’55’’ N 39°24’40’’ E, 1540 m.; Beşikdüzü: Yeşilköy, 11.09.1994, 40°59’32’’ Maçka: Güzel yayla, 20.08.1994, Akçaabat: Hıdırnebi yaylası, 08.08.1993, N 39°11’27’’ E, 490 m.; Araklı: Kirazlı 40°52’35’’ N 39°34’55’’ E, 1945 m. Yazıcı 40°58’55’’ N 39°24’40’’ E, 1540 m., yaylası, 17.07.1994, 40°41’00’’ N 0371. Beşikdüzü: Beşikdüzü–Şalpazarı road, 39°59’20’’ E, 1780 m.; Sürmene: Ağaçbaşı 25.07.1995, 41°01’00’’ N 39°10’15’’ E, 190 On decayed bark of Picea orientalis. yaylası, 18.06.1995, 40°48’40’’ N m., Araklı: Araklı–Gümüşhane road, 40°06’17’’ E, 1695 m.; Köprübaşı: İsmailağa * Lasallia pustulata (L.) Mérat 30.07.1994, 40°56’45’’ N 40°00’45’’ E, 80 yaylası, 15.07.1993, 40°41’53’’ N m., Çukurçayır köyü, 22.08.1994, Maçka: Çakırgöl dağı yaylası, 40°08’33’’ E, 1700 m.; Hayrat: Aksu yaylası, 40°47’58’’ N 39°58’13’’ E, 1200 m.; 09.08.1995, 40°40’33’’ N 39°41’50’’ E, 23.07.1993, 40°48’57’’ N 40°27’10’’ E, Köprübaşı: Dağardı köyü, 06.09.1993, 2500 m. Yazıcı 0380. 1710 m., Sarıçiçek yaylası, 04.09.1993, 40°52’32’’ N 40°09’22’’ E, 1120 m.; 40°51’20’’ N 40°28’30’’ E, 1700 m. Yazıcı On siliceous rock. Çarşıbaşı: Şahinli yaylası, 03.07.1995, 0557. * Lecanora albella (Pers.) Ach. 41°03’00’’ N 39°23’30’’ E, 980 m. Yazıcı Akçaabat: Hıdırnebi yaylası, 08.08.1993, 0605. On mosses. 40°58’55’’ N 39°24’40’’ E, 1540 m.; On Fagus orientalis and Carpinus sp. * Leptogium saturninum (Dicks.) nyl. Sürmene: Kal yaylası, 26.08.1995, * Lepraria incana (L.) Ach. Maçka: Hanzar yaylası, 23.07.1995, 40°47’26’’ N 40°05’27’’ E, 1690 m. Yazıcı 40°50’00’’ N 39°30’02’’ E, 1700 m; Düzköy: 0412. This species was abundant on soily Honofter yaylası, 21.08.1995, 40°48’35’’ N calcareous and siliceous rocks and bark of On Fagus orientalis and Carpinus sp. 39°23’15’’ E, 1780 m.; Çaykara: Malbet Ulmus sp. in coastal areas of Trabzon yaylası, 30.06.1994, 40°46’03’’ N Lecanora campestris (Schaer.) Hue (center), Of, Sürmene, Yomra, Arsin, Araklı, 40°20’10’’ E, 1695 m.; Araklı: Hendek Akçaabat, Çarşıbaşı, Vakfıbekir, Beşikdüzü Tonya: Şarlı yaylası, 17.07.1993, yaylası, 03.08.1993, 40°43’47’’ N and in lower fields of Tonya, Hayrat, Maçka, 40°43’47’’ N 39°19’30’’ E, 1813 m.; 39°53’28’’ E, 1800 m.; Hayrat: Anabarda Çaykara and Köprübaşı. Yazıcı 0611. Trabzon: Beşirli–Toklu Köyü border, yaylası, 08.09.1993, 40°52’33’’ N 19.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 Leptogium cyanescens (Rabenh.) Körb. 40°24’30’’ E, 1700 m.; Sürmene: Hocasuyu m.; Yomra: Lapazalık yaylası, 14.06.1993, Maçka: Altındere Valley Sumela yaylası, 21.06.1995, 40°48’58’’ N 40°49’30’’ N 39°48’25’’ E, 1250 m., Tokat 40°04’30’’ E, 1650 m. Yazıcı 0353. yaylası, 09.09.1993, 40°49’24’’ N Monastery and surroundings, 24.05.1993, 39°50’04’’ E, 1200 m., Çınarlı köyü, 40°46’25’’ N 39°39’15’’ E, 1200 m., On siliceous rock and mosses. Altındere Valley Zalişki yaylası, 07.05.1995, 07.04.1994, 41°00’27’’ N 39°48’00’’ E, 230 * Lobaria amplissima (Scop.) Fross. m. Yazıcı 0397. 40°47’30’’ N 39°39’35’’ E, 1500 m.; Tonya, Sivritepe yaylası, 04.09.1995, 40°53’41’’ N Maçka: Şolma yaylası, 11.08.1994, On Fagus orientalis, Carpinus sp., 36°16’10’’ E, 1100 m.; Araklı: Pervane köyü, 40°52’45’’ N 39°33’00’’ E, 1665 m., Populus sp., dyke wall, stone, siliceous and 16.08.1994, 40°55’02’’ N 40°00’10’’ E, Fırınoba yaylası, 06.07.1995, 40°40’30’’ N calcareous rock. Taşönü köyü, 10.10.1995, 41°01’12’’ N 39°35’20’’ E, 1700 m, Zal yaylası,
103 Lichen Flora of Trabzon
03.08.1994, 40°41’25’’ N 39°25’53’’ E, Maçka, Şalpazarı, Tonya, Araklı, Düzköy and On Fagus orientalis, Carpinus sp., Acer 1750 m.; Akçaabat: Ormana yaylası, Köprübaşı. Mostly it was defined together sp. and Juglans sp. 10.07.1994, 40°54’40’’ N 39°35’19’’ E, with Nephroma parile and rarely with Ophioparma ventosa (L.) Norm. 1700 m. Yazıcı 0383. Nephroma resupinatum. Yazıcı 0493. Maçka: Çakırgöl dağı yaylası, On Picea orientalis. It was commonly defined especially in 09.08.1995, 40°40’43’’ N 39°41’50’’ E, Lobaria pulmonaria (L.) Hoffm. Maçka. 2500 m.; Çaykara: Vartan yaylası, 15.08.1995, 40°46’00’’ N 40°10’05’’ E, Maçka: Altındere Valley Altındere District, * Nephroma parile (Ach.) Ach. 1945 m., Göl yaylası, 19.07.1993, 13.08.1995, 40°46’51’’ N 39°40’23’’ E, It is a very common species like 40°42’00’’ N 40°11’40’’ E, 1805 m. Yazıcı 1200 m., Büyükoba yaylası, 08.08.1995, Nephroma laevigatum. It was commonly 0612. 40°51’40’’ N 39°33’15’’ E, 1750 m.; defined on living and decayed bark and dried Düzköy: Doğankaya yaylası, 11.09.1995, branches of Picea orientalis and on siliceous On siliceous rock. 40°51’57’’ N 39°22’29’’ E, 1690 m.; Tonya: rocks and mosses in forests at altitudes of ** Pannaria conoplea (Ach.) Bory Karaptal yaylası, 14.08.1995, 40°52’15’’ N 1600 to 1800 m. in Maçka, Şalpazarı, Tonya, Çaykara: Karastel yaylası, 25.06.1994, 39°16’38’’ E, 1780 m.; Çaykara: Eğridere Araklı, Çaykara, Köprübaşı, Düzköy and 40°43’42’’ N 40°19’00’’ E, 1712 m.; Maçka: yaylası, 23.06.1994, 40°48’57’’ N Akçaabat. Yazıcı 0451. Lisevur yaylası, 19.06.1993, 40°43’00’’ N 40°20’30’’ E, 1640 m.; Şalpazarı: Karaçam It was abundant especially in Maçka. 39°28’40’’ E, 1700 m.; Tonya: Şarlı yaylası, yaylası, 06.06.1995, 40°47’31’’ N * Nephroma resupinatum (L.) Ach. 17.07.1993, 40°43’47’’ N 39°19’30’’ E, 39°14’20’’ E, 1687 m.; Araklı: Acısu yaylası, 1813 m. Yazıcı 0530. 01.08.1993, 40°46’42’’ N 40°00’25’’ E, Maçka: Altındere Valley Sumela 1800 m.; Sürmene: Büyük yayla, Monastery and surroundings 02.05.1993, On Picea orientalis and mosses. 23.06.1995, 40°47’33’’ N 40°03’45’’ E, 40°46’25’’ 39°39’15’’ E, 1200 m., Ardıçlı ** Pannaria pezizoides (Web.) Trevis. 1700 m.; Hayrat: Suluhan yaylası, yaylası, 22.08.1995, 40°47’32’’ N Şalpazarı: İzmiks yaylası, 09.07.1995, 24.07.1993, 40°53’40’’ N 40°24’40’’ E, 39°36’40’’ E, 1620 m.; Akçaabat: Kirazlık 40°56’30’’ N 39°12’00’’ E, 1400 m.; Maçka: 1700 m. Yazıcı 0327. yaylası, 24.08.1993, 40°54’00’’ N Hamsiköy yaylası, 27.06.1993, 40°40’43’’ N On Picea orientalis and Quercus sp. 39°31’15’’ E, 1785 m.; Tonya: Taksorobası 39°28’40’’ E, 1610 m. Yazıcı 0591. yaylası, 12.07.1993, 40°50’12’’ N On decayed bark of Picea orientalis, * Lobaria scrobiculata (Scop.) DC. 39°16’55’’ E, 1600 m.; Şalpazarı: Kızılkaya mosses and soil. Maçka: Akboran yaylası, 16.06.1993, yaylası, 14.06.1995, 40°46’15’’ N 40°41’25’’ N 39°44’55’’ E, 1790 m., Güzel 39°12’00’’ E, 1650 m.; Köprübaşı: Vizera Parmelia caperata (L.) Ach. yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ yaylası, 06.07.1993, 40°46’55’’ N As regards to choosing substrata it was E, 1945 m. Yazıcı 0329. 40°07’28’’ E, 1700 m.; Sürmene: Aydınpınar understood to be one of the least sensitive On Picea orientalis and mosses. yaylası, 10.08.1995, 40°45’41’’ N species. It was defined on Cistus creticus, 40°03’10’’ E, 1760 m. Yazıcı 0622. Fagus orientalis, Carpinus sp., Quercus sp., Lobothallia radiosa (Hoffm.) Hafellner * Ochrolechia parella (L.) A. Massal. Pinus sp. mosses and siliceous rocks at Trabzon: Beşirli–Toklu Köyü border, altitudes of 100 to 1600 m. at all stations. 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 Trabzon: Beşirli–Toklu köyü border, Mostly it was found together with Parmelia m.; Beşikdüzü: Beşikdüzü–Şalpazarı road, 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 conspersa. Yazıcı 0581. m. Yazıcı 0627. 25.07.1995, 41°01’00’’ N 39°10’15’’ E, 80 Parmelia conspersa (Ach.) Ach. m. Yazıcı 0613. On siliceous rock. It is a very common species like Parmelia On calcareous rock. * Opegrapha lithyrga Ach. caperata. It was defined to grow on siliceous * Megaspora verrucosa (Ach.) Hafellner Şalpazarı: Sisdağı yaylası, 28.07.1995, rocks at altitudes of 100 to 1600 m. at all & V. Wirth. 40°50’14’’ N 39°10’00’’ E, 1850 m. Yazıcı stations. Yazıcı 0552. Maçka: İpsila yaylası, 17.08.1994, 0615. * Parmelia elegantula (Zahlbr.) Szat. 40°43’15’’ N 39°36’45’’ E, 1690 m. Yazıcı On siliceous rock. Maçka: Kozağaç köyü, 17.07.1995, 0541. * Opegrapha varia Pers. 40°50’23’’ N 39°37’00’’ E, 850 m.; Araklı: Ekin yaylası, 18.06.1994, 40°41’58’’ N On soil. Akçaabat: Hıdırnebi yaylası, 08.08.1993, 40°02’17’’ E, 1790 m. Yazıcı 0442. Menegazzia terebrata (Hoffm.) A. 40°58’55’’ N 39°24’40’’ E, 1540 m.; Massal. Çaykara: Koldere köyü, 17.08.1995, On Pinus sp. Maçka: Altındere Valley Sumela 40°50’30’’ N 40°15’58’’ E, 790 m. Yazıcı * Parmelia exasperata de Not Monastery and surroundings, 24.05.1993, 0393. Maçka: Hamsiköyü yaylası, 27.06.1993, 40°46’25’’ N 39°39’15’’ E, 1200 m., Şolma On Carpinus sp. and Juglans sp. 40°40’43’’ N 39°28’40’’ E, 1610 m.; yaylası, 11.08.1994, 40°52’45’’ N * Opegrapha vulgata (Ach.) Ach. Akçaabat: Hıdırnebi yaylası, 08.08.1993, 39°33’00’’ E, 1665 m.; Araklı: Acısuyu 40°58’55’’ N 39°24’40’’ E, 1540 m.; yaylası, 01.08. 1993, 40°46’42’’ N Akçaabat: Enginoba yaylası, 26.08.1993, Akçaabat; Kemaliye köyü, 04.06.1994, 40°00’25’’ E, 1800 m. Yazıcı 0548. 40°52’34’’ N 39°29’00’’ E, 1690 m.; Tonya: 40°58’53’’ N 39°26’49’’ E, 890 m.; Tonya: Erikbeli Yaylası, 28.08.1995, 40°43’45’’ N On Picea orientalis and mosses. Sivritepe yaylası, 04.09.1995, 40°53’41’’ N 39°21’03’’ E, 1850 m.; Çaykara: Çahmut 39°16’10’’ E, 1100 m.; Çaykara: Çayıroba * Nephroma laevigatum Ach. yaylası, 21.07.1993, 40°48’40’’ N yaylası, 21.06.1993, 40°41’45’’ N It was defined on living and decayed bark 40°19’00’’ E, 1745 m.; Hayrat: Gama 40°20’35’’ E, 1840 m. Yazıcı 0394. and dried branches of Picea orientalis and yaylası, 27.07.1995, 40°49’42’’ N 40°23’00’’ E, 1680 m. Yazıcı 0365. On Picea orientalis, Carpinus sp., Acer sp. mosses at altitudes of 1600 to 1800 m. in and Cistus creticus.
104 K. YAZICI
Parmelia glabratula (L.) Nyl. 1650 m., Hamsiköyü yaylası, 22.08.1993, 41°01’37’’ N 39°38’08’’ E, 130 m.; Akçaabat: Kemaliye köyü, 04.06.1994, 40°40’’43’’ N 39°28’40’’ E, 1610 m.; Araklı: Beşikdüzü: Ağaçlı köyü, 01.09.1995, N 40°58’53’’ N 39°26’49’’ E, 890 m. Yazıcı Taşönlü köyü, 10.10.1995, 41°01’12’’ N 41°00’04’’ N 39°11’57’’ E, 390 m.; 0617. 39°59’42’’ E, 380 m., Yılantaş yaylası, Çarşıbaşı: Şahinli yaylası, 03.07.1995, 01.07.1995, 40°47’37’’ N 40°02’00’’ E, 41°03’00’’ N 39°23’30’’ E, 980 m. Yazıcı On Juniperus oxycedrus. 1650 m.; Sürmene: Aydınpınar yaylası, 0532. * Parmelia loxodes Nyl. 10.08.1995, 40°45’41’’ N 40°03’10’’ E, On siliceous and mosses. 1760 m., Harman yaylası, 19.06.1995, Maçka: Kozağaç köyü, 12.07.1995, 40°49’38’’ N 40°05’55’’ E, 1680 m.; * Parmeliopsis ambigua (Wulf.) Nyl. 40°50’23’’ N 39°37’00’’ E, 850 m. Yazıcı Köprübaşı: Yangın yaylası, 40°48’17’’ N Maçka: Güzel yayla, 20.08.1994, 0636. 40°08’00’’ E, 1650 m.; Yorma: Üçpınar 40°52’35’’ N 39°34’55’ E, 1945 m.; On siliceous rock. yaylası, 04.06.1993, 40°51’10’’ N Akçaabat: Kayabaşı yaylası, 24.08.1995, * Parmelia pastillifera (Harm.) Hale 39°47’30’’ E, 1510 m. Yazıcı 0599. 40°51’10’’ N 39°29’46’’ E, 1850 m. Yazıcı 0563. Akçaabat: Kemaliye köyü, 04.06.1994, On Picea orientalis, siliceous rock and 40°58’53’’ N 39°26’49’’ E, 790 m.; Çaykara: mosses. On Picea orientalis. Koldere köyü, 17.08.1995, 40°50’30’’ N Parmelia stygia (L.) Ach. * Peltigera aphthosa (L.) Willd. 40°15’58’’ E, 790 m. Yazıcı 0415. Maçka: Çakırgöl dağı yaylası, Maçka: Hamsiköyü yaylası, 25.06.1993, On Carpinus sp. and Acer sp. 09.08.1995, 40°40’33’’ N 39°41’50’’ E, 40°40’43’’ N 39°28’40’’ E 1610 m., Yazıcı Parmelia perlata (Huds.) Ach. 2500 m.; Düzköy: Haçka yaylası, 0623. 26.08.1994, 40°50’00’’ N 39°26’10’’ E, 190 Maçka: Lapazan yaylası, 16.06.1995, On mosses. m.; Çaykara: Kuti yaylası, 29.06.1994, 40°41’35’’ N 39°35’43’’ E, 1210 m.; Peltigera canina (L.) Willd. 40°41’48’’ N 40°22’40’’ E, 1750 m.; Akçaabat: Kuruçam yaylası, 06.08.1993, Köprübaşı: Cerah yaylası, 29.08.1993, Tonya: Taksorobası yaylası, 12.07.1993, 40°58’17’’ N 39°26’45’’ E, 800 m., Arpacılı 40°44’28’’ N 40°09’45’’ E, 1750 m. Yazıcı 40°50’12’’ N 39°16’55’’ E, 1600 m., köyü, 07.05.1993, 40°58’30’’ N 39°28’09’’ 0475. Bodamişkiye yaylası, 26.07.1993, 40°43’47’’ E, 300 m.; Trabzon: Akyazı köyü, N 39°17’57’’ E, 1818 m.; Çaykara: Karastel On siliceous rock. 04.09.1995, 41°01’30’’ N 39°36’04’’ E, 300 yaylası, 25.06.1994, 40°43’42’’ N m.; Şalpazarı: Geyikli köyü, 01.08.1995, Parmelia subrudecta Nyl. 40°19’00’’ E, 1712 m.; Araklı: Hendek 40°52’47’’ N 39°12’30’’ E, 1010 m.; yaylası, 03.08.1993, 40°43’47’’ N Akçaabat: Bozdoğan köyü, 10.05.1994, Beşikdüzü: Yeşilköy, 11.09.1994, 40°59’32’’ 39°53’28’’ E, 1850 m.; Trabzon: 40°59’55’’ N 39°28’45’’ E, 350 m.; Trabzon: N 39°11’27’’ E, 490 m., Beşikdağı, Beşirli–Toklu köyü border, 26.11.1995, Beşirli–Toklu Köyü border, 26.11.1995, 22.09.1994, 41°01’05’’ N 39°12’00’’ E, 510 41°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı 41°01’37’’ N 39°38’08’’ E, 130 m.; m.; Hayrat: Ovacık köyü, 11.04.1995, 0583. 41°01’17’’ N 40°23’45’’ E, 650 m.; Sürmene: Yokuşbaşı köyü, 22.04.1994, Vakfıkebir: Göller yaylası, 23.08.1993, 40°58’30’’ N 40°05’45’’ E, 320 m.; Hayrat: It was defined on mosses at all stations in 40°58’30’’ N 39°21’34’’ E, 845 m.; Yomra: Ovacık köyü, 11.04.1995, 41°01’17’’ N Maçka. Yokuşlu köyü, 06.03.1994, 41°00’08’’ N 40°23’45’’ E, 650 m. Yazıcı 0617. * Peltigera degenii Gyelnik 39°52’07’’ E, 260 m., Seslikaya yaylası, On Cistus creticus, siliceous rock and Maçka: Şolma yaylası, 11.08.1994, 10.09.1993, 40°48’58’’ N 39°48’42’’ E, mosses. 40°52’45’’ N 39°33’00’’ E, 1665 m. Yazıcı 1150 m. Yazıcı 0423. Parmelia sulcata Taylor 0462. On Carpinus sp. and Quercus sp. Maçka: Altındere Valley Sumela On mosses. * Parmelia pokornyi (Körb.) Szat. Monastery and surroundings, 24.05.1993, Peltigera elisabethae Gyelnik Maçka: Kadırga yaylası, 21.07.1994, 40°46’25’’ N 39°39’15’’ E, 1200 m., Maçka: Şolma yaylası, 11.08.1994, 40°43’10’’ N 39°23’00’’ E, 1950 m. Yazıcı Altındere Valley Zalişki yaylası, 07.05.1995, 40°52’45’’ N 39°33’00’’ E, 1665 m.; 0489. 40°47’30’’ N 39°39’35’’ E, 1500 m.; Köprübaşı: Barma yaylası, 05.09.1994, On siliceous rock and stone. Akçaabat: Kuruçam yaylası, 06.08.1993, 40°49’20’’ N 40°10’30’’ E, 1875 m. Yazıcı 40°58’17’’ N 39°26’45’’ E, 800 m., Arpacılı 0554. * Parmelia pulla Ach. köyü, 07.05.1993, 40°58’30’’ N 39°28’09’’ On mosses. Maçka: Şolma yaylası, 11.08.1994, E, 300 m., Acısu yaylası, 28.05.1993, 40°52’45’’ N 39°33’00’’ E, 1665 m.; 40°57’55’’ N 39°27’00’’ E, 750 m.; Araklı: * Peltigera horizontalis (Huds.) Baumg. Düzköy: Haçka yaylası, 26.08.1994, Çukurçayır köyü, 22.08.1994, 40°47’58’’ N Maçka: Altındere Valley Sumela 40°50’00’’ N 39°26’10’’ E, 1900 m., 39°58’13’’ E, 1200 m.; Köprübaşı: Dağardı Monastery and surroundings, 02.05.1993, Doğankaya yaylası, 11.09.1995, 40°51’57’’ köyü, 06.09.1993, 40°52’32’’ N 40°09’22’’ 40°46’25’’ N 39°39’15’’ E, 1200 m., N 39°22’29’’ E, 1690 m.; Akçaabat: Kemaliye E, 1120 m.; Vakfıkebir: Bozönü yaylası, Altındere Valley Taşköprü yaylası, köyü, 04.06.1994, 40°58’53’’ N 39°26’49’’ 10.07.1993, 40°56’45’’ N 39°21’04’’ E, 02.03.1995, 40°45’00’’ N 39°40’14’’ E, E, 890 m. Yazıcı 0543. 1100 m. Yazıcı 0385. 1250 m., İpsila yaylası, 17.08.1994, On Picea orientalis, Fagus orientalis and On Carpinus sp. and Quercus sp. 40°43’15’’ N 39°36’45’’ E, 1690 m., Hanzar yaylası, 23.07.1995, 40°50’00’’ N Acer sp. Parmelia tiliacea (Hoffm.) Ach. 39°30’02’’ E, 1760 m., Kaskar yaylası. * Parmelia saxatilis (L.) Ach. Akçaabat: Helvacı köyü, 04.07.1994, 02.09.1993, 40°39’40’’ N 39°34’31’’ E, Maçka: Altındere Valley Aksu yaylası, 40°01’18’’ N 39°31’23’’ E, 230 m.; Trabzon: 1790 m.; Akçaabat: Hıdırnebi yaylası, 20.06.1994, 40°45’58’’ N 39°39’50’’ E, Beşirli–Toklu Köyü border, 26.11.1995, 08.08.1993, 40°58’55’’ N 39°24’40’’ E,
105 Lichen Flora of Trabzon
1540 m.; Tonya: Erikbeli yaylası, On siliceous and calcareous rocks. Akçaabat: Hıdırnebi yaylası, 08.08.1993, 28.08.1995, 40°43’45’’ N 39°21’03’’ E, Pesturasia amara (Ach.) Nyl. 40°58’55’’ N 39°24’40’’ E, 1540 m. Yazıcı 1850 m.; Yomra: Tokat yaylası, 09.09.1993, 0654. Düzköy: Haçka yaylası, 26.08.1994, 40°49’24’’ N 39°50’04’’ E, 1510 m. Yazıcı On Carpinus sp. 0526. 40°50’00’’ N 39°26’10’’ E, 1700 m.; Tonya: Sazalan yaylası, 09.07.1993, 40°43’30’’ N Phaeophyscia orbicularis (Neck.) On mosses. 39°16’40’’ E, 1770 m., Şarlı yaylası, Moberg * Peltigera malacea (Ach.) Funck 17.07.1993, 40°43’47’’ N 39°19’30’’ E, Maçka: Altındere Valley Sumela 1800 m.; Araklı: Acısu yaylası, 01.08.1993, Akçaabat: Bozdoğan köyü, 10.05.1994, Monastery and surroundings, 24.05.1993, 40°46’42’’ N 40°00’25’’ E, 1790 m. Yazıcı 40°59’55’’ N 39°28’45’’ E, 350 m.; Trabzon: 40°46’25’’ N 39°39’15’’ E, 1200 m.; 0535. Beşirli–Toklu Köyü border, 26.11.1995, Trabzon: Karadeniz Technical University 41°01’37’’ N 39°38’08’’ E, 130 m.; Çaykara: On Fagus orientalis and Carpinus sp. Campus, 19.05.1993, 41°03’20’’ N 39°43’35’’ E, 130 m.; Köprübaşı: Köşk Vartan yaylası, 15.08.1995, 40°46’00’’ N Pertusaria aspergilla (Ach.) J.R. yaylası, 03.07.1993, 40°45’30’’ N 40°10’05’’ E, 1945 m. Yazıcı 0614. Laundon 40°07’30’’ E, 1500 m.; Sürmene: Muratlı On mosses. Maçka: Altındere Valley Sumela köyü, 18.03.1994, 40°59’42’’ N 40°05’15’’ Peltigera polydactylon (Neck.) Hoffm. Monastery and surroundings 02.05.1993, E, 250 m.; Arsin: Harmanlı köyü, 40°46’25’’ N 39°39’15’’ E, 1200 m.; 28.07.1993, 41°00’04’’ N 39°54’05’’ E, 350 Akçaabat: Kayabaşı yaylası, 24.08.1995, Trabzon: Gürbulak köyü, 12.10.1995, m. Yazıcı 0452. 45°51’10’’ N 39°29’46’’ E., 1850 m.; Maçka: 41°00’13’’ N 39°36’09’’ E, 500 m., Physcia adscendens (Fr.) H. Oliv. Kamadağ yaylası, 02.09.1994, 40°40’12’’ N Beşirli–Toklu köyü border, 26.11.1995, 39°29’45’’ E, 1750 m.; Araklı: Pinalar 41°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı This is a very common species. yaylası, 20.08.1993, 40°42’56’’ N 0625. Sometimes it was determined together with 39°54’28’’ E, 1825 m.; Düzköy: Gürgendağ Xanthoria parietina on Populus sp., Robinia yaylası, 04.08.1993, 40°51’13’’ N On siliceous rock. pseudoacacia, Salix sp., Acer sp. and siliceous 39°24’45’’ E, 1700 m.; Tonya: Kalınçam, ** Pertusaria corallina (L.) Arnold rock in the fields at altitudes of 10 to 900 m. 25.09.1995, 40°46’15’’ N 39°17’40’’ E, in Çarşıbaşı, Vakfıkebir, Beşikdüzü, Akçaabat, Maçka: Altındere Valley Sumela 1570 m. Yazıcı 0444. Trabzon (center), Sürmene, Of, Arsin, Araklı, Monastery and surroundings, 24.05.1993, Yomra and was defined on siliceous rocks on On mosses. 40°46’25’’ N 39°39’15’’ E, 1200 m.; plateaus above 900 m. in Şalpazarı, Maçka, Peltigera praetextata (Flörke ex Şalpazarı: Sisdağı yaylası, 28.07.1995, Köprübaşı, Hayrat, Düzköy, Akçaabat, Tonya, Sommerf.) Zopf. 40°50’14’’ N 39°10’00’’ E, 1850 m. Yazıcı Çaykara and Araklı. Yazıcı 0618. 0574. Maçka: Altındere Valley Altındere District, Physcia aipolia (Ehrh. ex Humb.) Hampe 13.08.1995, 40°46’51’’ N 39°40’23’’ E, On siliceous rock. 1200 m., Lapazan yaylası, 16.06.1995, Maçka: Altındere Valley Sumela Pertusaria lactea (L.) Arnold 40°41’35’’ N 39°35’43’’ E, 1710 m., Seloba Monastery and surroundings, 24.05.1993, yaylası, 13.06.1994, 40°43’00’’ N Trabzon: Beşirli–Toklu köyü border, 40°46’25’’. N 39°39’15’’ E, 1200 m., 39°39’25’’ E, 1793 m.; Düzköy: Karadağ 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 Hamsiköyü yaylası, 22.08.1993, 40°40’43’’ yaylası, 15.08.1993, 40°51’10’’ N m., Yomra: Üçpınar yaylası, 04.06.1993, N 39°28’40’’ E, 1610 m.; Akçaabat: Kuruçam 39°23’52’’ E, 1700 m.; Akçaabat: Kirazlık 40°51’10’’ N 39°47’30’’ E, 1510 m. Yazıcı yaylası, 06.08.1993, 40°58’17’’ N yaylası, 24.08.1993, 40°54’00’’ N 0406. 39°26’45’’ E, 800 m.; Tonya: Sivritepe yaylası, 04.09.1995, 40°53’41’’ N 39°31’15’’ E, 1785 m.; Şalpazarı: Güdün ** Pertusaria leioplaca DC. yaylası, 03.08.1995, 40°44’43’’ N 39°16’10’’ E, 1100 m., Kalınçam, 39°12’20’’ E, 1900 m.; Trabzon: Akçaabat: Hıdırnebi yaylası, 08.08.1993, 25.09.1995, 40°46’15’’ N 39°17’40’’ E, Beşirli–Toklu köyü border, 26.11.1995, 40°58’55’’ N 39°24’40’’ E, 540 m.; Maçka: 1400 m. Yazıcı 0454. 41°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı Altındere Valley Aksu yaylası, 20.06.1994, On Prunus sp., Quercus sp and Tilia sp. 0502. 40°45’58’’ N 39°39’50’’ E, 1650 m. Yazıcı 0349. Physcia caesia (Hoffm.) Fürnr. On mosses. On Fagus orientalis and Carpinus sp. Maçka: Kozağaç köyü, 12.07.1995, * Peltigera rufescens (Weiss) Humb. 40°50’23’’ N 39°37’00’’ E, 850 m., Mile * Pertusaria pertusa (Weig.) Tuck. Maçka: Hamsiköyü yaylası, 22.08.1993, yaylası, 27.07.1994, 40°43’30’’ N 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı Maçka: Hamsiköyü yaylası, 22.08.1993, 39°25’30’’ E, 1600 m.; Düzköy: Beypınar 0369. 40°40’43’’ N 39°28’40’’ E, 1610 m., yaylası, 25.08.1995, 40°47’30’’ N Fırınoba yaylası, 06.07.1995, 40°40’30’’ N 39°23’10’’ E, 1935 m.; Tonya: Akkise yaylası, On mosses. 39°35’20’’ E, 1780 m. Yazıcı 0345. 18.08.1993, 40°44’39’’ N 39°20’20’’ E, * Pertusaria albescens (Huds.) M. 1650 m.; Trabzon: Dolaylı köyü, Choisy & Werner ** Phaeographis smithii (Leight.) B. de. 11.09.1993, 40°59’25’’ N 39°44’05’’ E, 350 Lesd. m.; Şalpazarı: Geyikli köyü, 01.08.1995, Trabzon: Beşirli–Toklu Köyü border, 40°52’47’’ N 39°12’30’’ E, 1310 m.; 26.11.1995, 41°01’37’’ N 39°38’08’’ E, Çaykara: Koldere köyü, 17.08.1995, Beşikdüzü: Yeşilköy, 11.09.1994, 40°59’32’’ 130m.; Beşikdüzü: Beşikdüzü–Şalpazarı road, 40°50’30’’ N 40°15’58’’ E, 790 m. Yazıcı N 39°11’27’’ E, 490 m., Beşikdağı, 25.07.1995, 41°01’00’’ N 39°10’15’’ E, 90 0319. 22.09.1994, 41°01’05’’ N 39°12’00’’ E, 510 m.; Araklı: Dağbaşı köyü, 05.10.1995, On Juglans sp. m.; Vakfıkebir: Güney köyü, 23.04.1995, 40°48’06’’ N 39°55’38’’ E, 1550 m. Yazıcı * Phaeophyscia ciliata (Hoffm.) Moberg 41°01’00’’ N 39°16’20’’ E, 400 m. Yazıcı 0402. 0486.
106 K. YAZICI
On siliceous and calcareous rocks. Trabzon: Beşirli–Toklu köyü border, On siliceous rock. 19.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 * Physcia dimidiata (Arnold) Nyl. * Porpidia crustulata (Ach.) Hertel & m. Yazıcı 0373. Knoph Trabzon: Beşirli–Toklu köyü border, On calcareous rock. 19.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 Şalpazarı: Kızılkaya yaylası, 14.06.1995, m. Yazıcı 0494. Physconia distorta (With.) Laundon 40°46’15’’ N 39°12’00’’ E, 1650 m. Yazıcı 0616. On siliceous rock. Maçka: Hamsiköyü yaylası, 22.08.1993, 40°40’43’’ N 39°28’40’’ E, 1610 m.; On siliceous rock. Physcia semipinnata (J.F. Gmel.) Akçaabat: Kuruçam yaylası, 06.08.1993, Moberg Porpidia macrocarpa (DC.) Hertel & 40°58’17’’ N 39°26’45’’ E, 800 m.; Kemaliye Schwab Maçka: Hamsiköyü yaylası, 10.06.1993, köyü, 04.06.1994, 40°58’53’’ N 39°26’49’’ 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı E, 890 m., Acısu yaylası, 28.05.1993, Köprübaşı: Barma yaylası, 05.09.1994, 0461. 40°57’55’’ N 39°27’00’’ E, 750 m.; Hayrat: 40°49’20’’ N 40°10’30’’ E, 1875 m. Yazıcı Ovacık köyü, 11.04.1995, 41°01’17’’ N 0582. On calcareous rock. 40°23’45’’ E, 650 m.; Vakfıkebir: Göller On siliceous rock. * Physcia stellaris (L.) Nyl. yaylası, 23.08.1993, 40°50’30’’ N Porpidia speirea (Ach.) Krempelh. Akçaabat: Kemeliye köyü, 04.06.1994, 39°21’34’’ E, 845 m.; Of: Gökçeoba köyü, 40°58’53’’ N 39°26’49’’ E, 890 m., Arpacılı 01.09.1994, 41°04’40’’ N 40°19’30’’ E, 400 Maçka: Çakırgöl dağı yaylası, köyü, 07.05.1993, 40°58’30’’ N 39°28’09’’ m. Yazıcı 0395. 09.08.1995, 40°40’33’’ N 39°41’50’’ E, E, 300 m.; Trabzon: Gürbulak köyü, On Acer sp., Quercus sp. and Prunus sp. 2500 m.; Düzköy: Haçka yaylası, 12.10.1995, 41°00’13’’ N 39°36’09’’ E, 400 26.08.1994, 40°50’00’’ N 39°26’10’’ E, * Physconia muscigena (Ach.) Poelt m., Karlık köyü, 22.10.1994, 40°59’30’’ N 1900 m.; Şalpazarı: Sisdağı yaylası, 39°39’40’’ E, 580 m., Pınaraltı köyü, Maçka: Kadırga yaylası, 21.07.1994, 28.07.1995, 40°50’14’’ N 39°10’00’’ E, 02.06.1995, 40°56’15’’ N 39°46’17’’ E, 890 40°43’10’’ N 39°23’00’’ E, 1950 m.; Tonya: 1850 m. Yazıcı 0434. m.; Araklı: Araklı–Gümüşhane road, Erikbeli yaylası, 28.08.1995, 40°43’45’’ N On calcareous rock. 30.07.1994, 40°56’45’’ N 40°00’45’’ E, 120 39°21’03’’ E, 1850 m. Yazıcı 0593. * Protoparmelia badia (Hoffm.) m.; Sürmene: Muratlı köyü, 18.03.1994, On mosses. 40°59’42’’ N 40°05’15’’ E, 250 m.; Yomra: Hafellner Yokuşlu köyü, 06.03.1994, 41°00’08’’ N Placynthium nigrum (Huds.) Gray Maçka: Çakırgöl dağı yaylası, 39°52’07’’ E, 260 m., Çınarlı köyü, Trabzon: Center, Boztepe, 25.03.1995, 09.08.1995, 40°40’33’’ N 39°41’50’’ E, 07.04.1994, 41°00’27’’ N 39°48’00’’ E, 230 41°03’13’’ N 39°40’50’’ E, 90 m. Yazıcı 2500 m.; Şalpazarı: Sisdağı yaylası, m.; Vakfıkebir: Fol deresi, Vakfıkebir–Tonya 0621. 28.07.1995, 40°50’14’’ N 39°10’00’’ E, road, 23.09.1995, 41°00’13’’ N 39°16’00’’ 1850m., Alaca yaylası, 17.08.1993, E, 700 m.; Of: Tekoba Köyü, 15.03.1994, On calcareous rock. 40°49’40’’ N 39°11’57’’ E, 1800 m. Yazıcı 40°59’26’’ N 40°17’45’’ E, 790 m., Kıyıboyu ** Plastismatia norvegica (Lynge) W. 0332. köyü, 12.04.1994, 41°04’28’’ N 40°15’33’’ Culb. & C. Culb. On siliceous rock. E, 250 m., Koruluk köyü, 04.09.1994, Maçka: Güzel yayla, 20.08.1994, 41°01’27’’ N 40°17’52’’ E, 500 m. Yazıcı Pseudephebe minuscula (Nyl. ex 40°52’35’’ N 39°34’55’’ E, 1945 m. Yazıcı 0403. Arnold) Brodo & D. Hawksw. 0555. Çaykara: Hanırmağı yaylası, 16.08.1995, On Robinia pseudoacacia, Populus sp., On decayed bark of Picea orientalis. Acer sp., Quercus sp. and Salix sp. 40°47’26’’ N 40°13’00’’ E, 1950 m.; * Polychidium muscicola (Swartz) Gray Akçaabat, Hıdırnebi yaylası, 08.08.1993, * Physcia tenella (Scop.) DC. 40°58’55’’ N 39°24’40’’ E, 1540 m. Yazıcı Çaykara: Sultanmurat yaylası, 0367. Trabzon: Beşirli–Toklu köyü border, 18.08.1995, 40°45’00’’ N 40°11’40’’ E, 10.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 1935 m. Yazıcı 0512. On siliceous rock. m., Center, Boztepe, 19.05.1995, 41°03’13’’ ** Pseudephebe pubescens (L.) M. N 39°40’50’’ E, 120 m., Karadeniz Technical On soil. Choisy University Campus, 13.08.1994, 41°03’20’’ Porpidia cinereoatra (Ach.) Hertel & N 39°43’35’’ E, 130 m., Gölçayır köyü, Khoph Çaykara: Vartan yaylası, 15.08.1995, 11.06.1995, 41°00’30’’ N 39°40’55’’ E, 500 40°46’00’’ N 40°10’05’’ E, 1945 m.; Maçka: Altındere Valley Sumela m., Dolaylı köyü, 11.09.1993, 40°59’25’’ N Köprübaşı: Kangel yaylası, 05.08.1994, Monastery and surroundings, 24.05.1993, 39°44’05’’ E, 350 m.; Şalpazarı: Dorukkiriş 40°49’40’’ N 40°08’15’’ E, 1670 m. Yazıcı 40°46’25’’ N 39°39’15’’ E, 1200 m., İpsila köyü, 28.06.1993, 40°52’45’’ N 39°12’20’’ 0458. yaylası, 17.08.1994, 40°43’15’’ N E, 950 m.; Beşikdüzü: Yeşilköy, 11.09.1994, On siliceous rock. 39°36’45’’ E, 1690 m., Kulindağı yaylası, 40°59’32’’ N 39°11’27’’ E, 490 m., Bozlu 15.07.1995, 40°52’26’’ N 39°33’40’’ E, Psedevernia furfuracea (L.) Zopf. var. köyü, 19.09.1995, 40°59’16’’ N 39°12’24’’ 1700 m.; Düzköy: Doğankaya yaylası, furfuracea E, 410 m.; Araklı: Taşönü köyü, 10.10.1995, 11.09.1995, 40°51’57’’ N 39°22’29’’ E, 41°01’12’’ N 39°59’42’’ E, 380 m.; Arsin: Maçka: Altındere Valley Sumela 1690 m., Haçka yaylası, 26.08.1994, Harmanlı köyü, 28.07.1993, 41°00’04’’ N Monastery and surroundings, 02.05.1993, 40°50’00’’ N 39°26’10’’ E, 1900 m.; Tonya: 39°54’05’’ E, 350 m. Yazıcı 0346. 40°46’25’’ N 39°39’15’’ E, 1200 m., Taksorobası yaylası, 12.07.1993, 40°50’12’’ Altındere Valley Zalişki yaylası, 15.05.1995, On Robinia pseudoacacia, Populus sp., N 39°16’55’’ E, 1600 m.; Şalpazarı: Karaçam 40°47’30’’ N 39°39’35’’ E, 1500 m., Salix sp., Acer sp. and Quercus sp. yaylası, 06.06.1995, 40°47’31’ N 39°14’20’’ Hamsiköyü yaylası, 25.06.1993, 40°40’43’’ * Physcia tribacia (Ach.) Nyl. E, 1680 m. Yazıcı 0506. N 39°28’40’’ E, 1610 m., Güzel yayla,
107 Lichen Flora of Trabzon
20.28.1994, 40°52’35’’ N 39°34’55’’ E, Sürmene, Çaykara, Köprübaşı, Şalpazarı: Düzköy: Haçka yaylası, 26.08.1994, 1945 m., Fırınoba yaylası, 06.07.1995, Yazıcı 0464. 40°50’00’’ N 39°26’10’’ E, 1900 m. Yazıcı 40°40’30’’ N 39°35’20’’ E, 1780 m., Kaskar Ramalina fastigiata (Pers.) Ach. 0635. yaylası, 02.09.1993, 40°39’40’’ N On siliceous rock. 39°34’31’’ E, 1790 m.; Düzköy: Haçka Maçka: Güzel yayla, 20.08.1994, yaylası, 26.08.1994, 40°50’00’’ N 40°52’35’’ N 39°34’55’’ E, 1945 m., * Rhizocarpon obscuratum (Ach.) A. 39°26’10’’ E, 1900 m.; Akçaabat: Kemaliye Kulindağı yaylası, 15.07.1995, 40°52’26’’ N Massal. 39°33’40’’ E, 1700 m.; Akçaabat: Hıdırnebi köyü, 04.06.1994, 40°58’53’’ N 39°26’49’’ Köprübaşı: Barma yaylası, 05.09.1994, yaylası, 08.08.1993, 40°58’55’’ N E, 890 m.; Tonya: Taksorobası yaylası, 40°49’20’’ N 40°10’30’’ E, 1875 m. Yazıcı 39°24’40’’ E, 1540 m. Yazıcı 0490. 12.07.1993, 40°50’12’’ N 39°16’55’’ E, 0633. 1600 m.; Çaykara: Malbet yaylası, On Picea orientalis and Carpinus sp. On siliceous rock. 30.06.1994, 40°46’03’’ N 40°20’10’’ E, Ramalina pollinaria (Westr.) Ach. 1695 m.; Şalpazarı: Kızılkaya yaylası, ** Rhizocarpon oederi (Web.) Körb. Maçka: Kozağaç köyü, 12.07.1995, 14.06.1995, 40°46’15’’ N 39°12’00’’ E, Çaykara: Hanırmağı yaylası, 16.08.1995, 1650 m.; Araklı: Kaleron yaylası, 40°50’23’’ N 39°37’00’’ E, 850 m.; Trabzon: Beşirli–Toklu köyü border, 10.11.1995, 40°47’26’’ N 40°13’00’’ E, 1950 m. Yazıcı 17.07.1994, 40°40’00’’ N 39°59’57’’ E, 0514. 1790 m.; Köprübaşı: Taşlı yaylası, 40°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı 06.09.1993, 40°44’20’ N 40°06’00’’ E, 0510. On siliceous rock. 1775 m. Sürmene: Hocasuyu yaylası, On siliceous rock. * Rhizocarpon richardii (Nyl.) Zahlbr. 21.06.1995, 40°48’58’’ N 40°04’30’’ E, * Ramalina siliquosa (Huds.) A.L.Sm. 1650 m., Aydınpınar yaylası, 10.06.1995, Şalpazarı: Alaca yaylası, 17.08.1993, 40°45’41’’ N 40°03’10’’ E, 1760 m.; Hayrat: Maçka: Hamsiköyü yaylası, 27.06.1993, 40°49’40’’ N 39°11’57’’ E, 1800 m., Sisdağı Suluhan yaylası, 24.07.1993, 40°53’40’’ N 40°40’43’’ N 39°28’40’’ E, 1610 m., yaylası, 28.07.1995, 40°50’14’’ N 40°24’40’’ E, 1705 m. Yazıcı 0561. Kozağaç köyü, 12.07.1995, 40°50’23’’ N 39°10’00’’ E, 1850 m. Yazıcı 0447. 39°37’00’’ E, 850 m. Yazıcı 0564. On Picea orientalis, Fagus orientalis, On siliceous rock. On siliceous rock. Carpinus sp. and Juglans sp. * Rhizocarpon viridiatrum (Wulf.) ** Ramalina subamplicata (Nyl.) Flk. * Ramalina calicaris (L.) Fr. Körb. Maçka: Hamsiköyü yaylası, 22.08.1993, Maçka: Altındere Valley Aksu yaylası, Akçaabat: Acısu yaylası, 28.05.1993, 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı 40°57’55’’ N 39°27’00’’ E, 750 m. Yazıcı 27.09.1994, 40°45’58’’ N 39°39’50’’ E, 0366. 1650 m., Şolma yaylası, 24.08.1994, 0495. On Prunus sp. 40°52’45’’ N 39°33’00’’ E, 1665 m., Güzel On siliceous rock. yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ * Ramalina thrausta (Ach.) Nyl. Rhizoplaca chrysoleuca (Smith.) Zopf E, 1945 m., Lapazan yaylası, 16.06.1995, Maçka: Büyükoba yaylası, 08.08.1995, 40°41’35’’ N 39°35’43’’ E, 1710 m., Lisevur 40°51’40’’ N 39°33’15’’ E, 1800 m.; Maçka: Çakırgöl dağı yaylası, yaylası, 19.06.1993, 40°43’00’’ N Akçaabat: Kayabaşı yaylası, 24.08.1995, 09.08.1995, 40°40’33’’ N 39°41’50’’ E, 39°28’40’’ E, 1700 m.; Tonya: Sazalan 40°51’10’’ N 39°29’46’’ E, 1850 m. Yazıcı 2500 m., Zal yaylası, 03.08.1994, yaylası, 09.07.1993, 40°43’30’’ N 0546. 40°41’25’’ N 39°25’53’’ E, 1750 m.; Araklı: 39°16’40’’ E, 1870 m.; Çaykara: Uzungöl Erikli yaylası, 13.08.1993, 40°38’30’’ N On Picea orientalis. yaylası, 28.08.1993, 40°44’37’’ N 40°02’17’’ E, 1950 m.; Hayrat: Aksu yaylası, 40°19’30’’ E, 1690 m., Malbet yaylası, Rhizocarpon geminatum Körber 23.07.1993, 40°48’57’’ N 40°27’10’’ E, 30.06.1994, 40°46’03’ N 40°20’10’’ E, Maçka: Kadırga yaylası, 21.07.1994, 1710 m. Yazıcı 0542. 1695 m.; Hayrat: Gama yaylası, 27.07.1995, 40°43’10’’ N 39°23’00’’ E, 1950 m.; On siliceous rock. 40°49’42’’ N 40°23’00’’ E, 1680 m.; Çaykara: Uzungöl yaylası, 25.07.1994, Sürmene: Ağaçbaşı yaylası, 18.06.1995, 40°44’37’’ N 40°19’30’’ E, 1690 m.; Rhizoplaca melanophthalma (Ram.) 40°48’40’’ N 40°06’17’’ E, 1695 m. Yazıcı Şalpazarı: Sisdağı yaylası, 28.07.1995, Leux 0405. 40°50’14’’ N 39°10’00’’ E, 1850 m.; Araklı: Maçka: Çakırgöl dağı yaylası, On Picea orientalis. Demirtaş yaylası 22.06.1994, 40°42’05’’ N 09.08.1995, 40°40’33’’ N 39°41’50’’ E, 40°01’07’’ E, 1815 m. Yazıcı 0590. 2500 m., Ardıçlı yaylası, 22.08.1995, Ramalina capitata (Ach.) Nyl. On siliceous rock. 40°47’32’’ N 39°36’40’’ E, 1800 m.; Maçka: Hamsiköyü yaylası, 10.06.1993, Çaykara: Sultanmurat yaylası, 18.08.1995, Rhizocarpon geographicum (L.) DC. 40°40’43’’ N 39°28’40’’ E, 1610 m., Yazıcı 40°45’00’’ N 40°11’40’’ E, 1935 m.; 0629. It was commonly defined on siliceous Şalpazarı: Güdün yaylası, 03.08.1995, On siliceous rock. rocks in fields above 1400 m. in Maçka, 40°44’43’’ N 39°12’20’’ E, 1950 m. Yazıcı Şalpazarı, Tonya, Düzköy, Sürmene, 0573. Ramalina farinacea (L.) Ach. Köprübaşı, Hayrat, Çaykara and Akçaabat. On siliceous rock. It was commonly defined on Carpinus sp. From time to time it was seen together with and Quercus sp. at altitudes of 800 to 1100 Umbilicaria cylindrica and rarely Umbilicaria * Rimularia insularis (Nyl.) Rambold & and on Picea orientalis at locations above deusta in fields above 1700 m. at the same Hertel stations. Yazıcı 0620. 1400 m. in Maçka, Akçaabat, Şalpazarı, Maçka: Çakırgöl dağı yaylası, Çaykara, Düzköy, Tonya, Hayrat, Köprübaşı, * Rhizocarpon lecanorinum Anders 11.07.1995, 40°40’33’’ N 39°41’50’’ E, Sürmene, Trabzon (Center). Sometimes it Maçka: Zal yaylası, 03.08.1994, 2500 m. Yazıcı 0524. was seen together with Evernia prunastri and 40°41’25’’ N 39°25’53’’ E, 1750 m.; Usnea spp. in Maçka, Akçaabat, Düzköy, On siliceous and calcareous rock.
108 K. YAZICI
* Rinodina teichophila (Nyl.) Arnold E, 890 m.; Araklı: Acısu yaylası, 01.08.1993, 26.11.1995, 41°01’37’ N 39°38’08’’ E, 130 40°46’42’’ N 40°00’25’’ E, 1810 m.; Araklı: m. Yazıcı 0390. Akçaabat: Helvacı köyü 04.07.1994, Pervane köyü, 16.08.1994, 40°55’02’’ N 40°01’18’’ N 39°31’23’’ E, 230 m.; Trabzon: On calcareous rock. 40°00’10’’ E, 1300 m.; Vakfıkebir: Fevziye Beşirli–Toklu köyü border, 26.11.1995, köyü, 03.11.1995, 40°58’37’’ N 39°15’52’’ ** Umbilicaria aprina Nyl. 41°01’37’’ N 39°38’08’’ E, 130 m. Yazıcı E, 350 m., Yaylacık köyü, 06.08.1995, 0584. Çaykara: Sultanmurat yaylası, 41°59’53’’ N 39°14’34’’ E, 390 m.; Of: 18.08.1995, 40°45’00’’ N 40°11’40’’ E, On calcareous rock. Tekoba köyü, 15.03.1994, 40°59’26’’ N 1935 m. Yazıcı 0480. * Sporastatia testudinea (Ach.) A. 40°17’45’’ E, 790 m.; Arsin: Çiçekli köyü, On siliceous rock. Massal. 07.08.1995, 41°00’02’’ N 39°57’30’’ E, 500 m. Yazıcı 0566. ** Umbilicaria arctica (Ach.) Nyl. Maçka: Çakırgöl dağı yaylası, 09.08.1995, 40°40’33’’ N 39°41’50’’ E, On siliceous and calcareous rock. Çaykara: Sultanmurat yaylası, 2500 m.; Çaykara: Sultanmurat yaylası, ** Thamnolia vermicularis (Sw.) Schaer. 18.08.1995, 40°45’00’’ N 40°11’40’’ E, 1935 m. Yazıcı 0470. 18.08.1995, 40°45’00’’ N 40°11’40’’ E, Maçka: Çakırgöl dağı yaylası, 1935 m. Yazıcı 0344. 09.08.1995, 40°40’33’’ N 39°41’50’’ E, Umbilicaria cylindrica (L.) Del. ex Duby On siliceous rock. 2500 m.; Çaykara: Sultanmurat yaylası, It was abundant on siliceous rocks at 18.08.1995, 40°45’00’’ N 40°11’40’’ E, * Squamarina cartilaginea (With) P. locations above 1600 m. in Maçka, Çaykara, 1935 m., Sıcakoba yaylası, 19.08.1995, James Tonya, Düzköy, Akçabat, Şalpazarı, Araklı, 40°46’57’’ N 40°10’50’’ E, 1930 m. Yazıcı Sürmene and Hayrat. Yazıcı 0473. Maçka: Kozağaç köyü, 17.07.1995, 0569. 40°50’23’’ N 39°37’00’’ E, 850 m. Yazıcı * Umbilicaria decussata (Vill.) Frey On soil. 0634. Maçka: Hamsiköyü yaylası, 22.08.1993, On soil and calcareous rock. Toninia sedifolia (Scop.) Timdal 40°40’43’’ N 39°28’40’’ E, 1610 m. Yazıcı ** Stereocaulon nanodes Tuck Maçka: Altındere Valley Sumela 0477. Monastery and surroundings, 24.05.1993, On siliceous rock. Çaykara: Sıcakoba yaylası, 19.08.1995, 40°46’25’’ N 39°39’15’’ E, 1200 m., 40°46’57’’ N 40°10’50’’ E, 1930 m. Yazıcı Kozağaç köyü, 17.07.1995, 40°50’23’’ N ** Umbilicaria deusta (L.) Baumg. 0508. 39°37’00’’ E, 850 m., Zarışa yaylası, It was defined on siliceous rocks at On calcareous rock. 07.06.1994, 40°47’37’’ N 39°32’50’’ E, locations above 1600 m. in Maçka, Çaykara, 1695 m.; Düzköy: Doğankaya yaylası ** Stereocaulon pileatum Ach. Tonya, Şalpazarı, Araklı, Sürmene, Düzköy, 11.09.1995, 40°51’57’’ N 39°22’29’’ E, Akçaabat and Hayrat. Yazıcı 0597. Akçaabat: Hıdırnebi yaylası, 08.08.1993, 1690 m.; Akçaabat: Helvacı köyü, 40°58’55’’ N 39°24’40’’ E 1540 m., Yazıcı 04.07.1994, 40°01’18’’ N 39°31’23’’ E, 230 On siliceous rock. 0553. m., Acısu yaylası, 28.05.1993, 40°57’55’’ N * Umbilicaria hirsuta (Swar. ex Westr.) On siliceous rock. 39°27’00’’ E, 750 m.; Trabzon: Akyazı köyü, Hoffm. 04.09.1995, 41°01’30’’ N 39°36’04’’ E, 300 * Sticta fuliginosa (Hoffm.) Ach. m., Beşirli–Toklu köyü border, 19.11.1995, Maçka: Kadırga yaylası, 21.07.1995, Maçka: Altındere Valley Sumela 41°01’37’’ N 39°38’08’’ E, 130 m., Dolaylı 40°43’10’’ N 39°23’00’’ E, 1950 m., İpsila Monastery and surroundings 02.05.1993, köyü, 11.09.1993, 40°59’25’’ N 39°44’05’’ yaylası, 17.08.1994, 40°43’15’’ N 40°46’25’’ N 39°39’15’’ E, 1200 m., Şolma E, 350 m.; Çaykara: Vartan yaylası, 39°36’45’’ E, 1690 m.; Çaykara: Sıcakoba yaylası, 11.08.1994, 40°52’45’’ N, 15.08.1995, 40°46’00’’ N 40°10’05’’ E, yaylası, 19.08.1995, 40°46’57 N 40°10’50’. 39°33’00’’ E, 1665 m.; Çaykara: Sıcakoba 1945 m., Kuti yaylası, 29.06.1994, E, 1930 m. Yazıcı 0377. yaylası, 19.08.1995, 40°46’57’’ N 40°41’48’’ N 40°22’40’’ E, 1750 m.; On siliceous rock. 40°10’50’’ E 1930 m. Yazıcı 0398. Şalpazarı: Dorukkiriş köyü, 28.06.1993, ** Umbilicaria nylanderiana (Zahlbr.) 40°52’45’’ N 39°12’20’’ E, 1210 m.; Araklı: On Picea orientalis and mosses. Magn. Demirtaş yaylası, 22.06.1994, 40°42’05’’ N * Sticta sylvatica (Huds.) Ach. 40°01’07’’ E, 1815 m.; Sürmene: Lişit Çaykara: Sıcakoba yaylası, 19.08.1995, Çaykara: Sıcakoba yaylası, 19.08.1995, yaylası, 26.06.1995, 40°46’30’’ N 40°46’57’’ N 40°10’50’’ E, 1930 m., Yazıcı 40°46’57’’ N 40°10’50’’ E, 1930 m. Yazıcı 40°04’33’’ E, 1725 m. Yazıcı 0517. 0529. 0608. On soil. On siliceous rock. On mosses. * Trapelia coarctata (Sm.) M. Choisy Umbilicaria subglabra (Nyl.) Harm. * Tephromela atra (Huds.) Hafellner ex Trabzon: Beşirli–Toklu köyü border, Çaykara: Sıcakoba yaylası, 19.08.1995, Kalb 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 40°46’57’’ N 40°10’50’’ E, 1930 m. Yazıcı Düzköy: Haçka yaylası, 26.08.1994, m., Ayasofya Church Garden, 09.05.1993, 0425. 40°50’00’’ N 39°26’10’’ E, 1900 m.; Tonya: 41°03’17’’ N 39°39’50’’ E, 90 m. Yazıcı On siliceous rock. Sivritepe yaylası, 04.09.1995, 40°53’41’’ N 0396. 39°16’10’’ E, 1100 m.; Trabzon: * Umbilicaria vellea (L.) Ach. em. Frey On calcareous rock and limestone. Beşirli–Toklu köyü border, 19.11.1995, Maçka: Kozağaç köyü, 17.07.1995, 41°01’37’’ N 39°38’08’’ E, 130 m., Pınaraltı ** Trapelia involuta (Taylor) Hertel 40°50’23’’ N 39°37’00’’ E, 850 m.; Tonya: köyü, 02.06.1995, 40°56’15’’ N 39°46’17’’ Trabzon: Beşirli–Toklu köyü border, Sazalan yaylası, 09.07.1993, 40°43’30’’ N
109 Lichen Flora of Trabzon
39°16’40’’ E, 1870 m.; Çaykara: 10.07.1994, 40°54’40’’ N 39°35’19’’ E, * Xanthoria candelaria (L.) Th. Fr. Sultanmurat yaylası, 18.08.1995, 40°45’00’’ 1700 m. Yazıcı 0587. Çaykara: Sıcakoba yaylası, 19.08.1995, N 40°11’40’ E, 1935 m.; Şalpazarı: Sisdağı In addition it was defined at a total of 20 40°46’57’’ N 40°10’50’’ E, 1930 m., Vartan yaylası, 28.07.1995, 40°50’14’’ N stations in Maçka. yaylası, 15.08.1995, 40°46’00’’ N 39°10’00’’ E, 1850 m., Karaçam yaylası, 40°10’05’’ E, 1945 m., Malbet yaylası, 06.06.1995, 40°47’31’’ N 39°14’20’’ E, On Picea orientalis. 30.06.1994, 40°46’03’’ N 40°20’10’’ E, 1687 m.; Köprübaşı: İsmailağa yaylası, * Usnea hirta (L.) Wigg. 1695 m.; Araklı: Yılantaş yaylası, 15.07.1993, 40°41’53’’ N 40°08’33’’ E, 01.07.1995, 40°47’37’’ N 40°02’00’’ E, 1790 m., Kangel yaylası, 05.08.1994, Maçka: Büyükoba yaylası, 08.08.1995, 1650 m.; Maçka: Omela yaylası, 28.08.1994, 40°49’40’’ N 40°08’15’’ E, 1670 m., Köşk 40°51’40’’ N 39°33’15’’ E, 1800 m., Kaskar 40°43’45’’ N 39°45’00’’ E, 1770 m. Yazıcı yaylası, 03.07.1993, 40°45’30’’ N yaylası, 02.09.1993, 40°39’40’’ N 0606. 40°07’30’’ E, 1720 m. Yazıcı 0427. 39°34’31’’ E, 1790 m.; Düzköy: Haçka yaylası, 26.08.1994, 40°50’00’’ N On decayed wood and calcareous rock. On siliceous rock. 39°26’10’’ E, 1900 m., Honofter yaylası, Xanthoria elegans (L.) Th. Fr. * Usnea articulata (L.) Hoffm. 21.08.1995, 40°48’35’’. N 39°23’15’’. E, 1925 m.; Akçaabat: Kayabaşı yaylası, Maçka: Çakırgöl dağı yaylası, Maçka: Hamsiköyü yaylası, 25.06.1993, 24.08.1995, 40°51’10’’ N 39°29’46’’ E, 11.07.1995, 40°40’33’’ N 39°41’50’’ E, 40°40’43’’ N 39°28’40’’ E, 1610 m., Güzel 1850 m. Yazıcı 0498. 2500 m., Altındere Valley Sumela Monastery yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ and surroundings, 02.05.1993, 40°46’25’’ N E, 1945 m.; Çaykara: Uzungöl yaylası, On Picea orientalis. 39°39’15’’ E, 1200 m., Hamsiköyü yaylası, 19.07.1995, 40°44’37’’ N 40°19’30’’ E, * Usnea lapponica Vain. 27.06.1993, 40°40’43’’ N 39°28’40’’ E, 1690 m. Yazıcı 0382. Akçaabat: Kayabaşı yaylası, 24.08.1995, 1610 m., Kozağaç köyü, 12.07.1995, On Picea orientalis. 40°51’10’’ N 39°29’46’’ E, 1850 m. Yazıcı 40°50’23’’ N 39°37’00’’ E, 850 m.; Tonya: Sazalan yaylası, 09.07.1993, 40°43’30’’ N ** Usnea diplotypus Vain. 0626. 39°16’40’’ E, 1600 m.; Şalpazarı: Güdün Maçka: Altındere Valley Aksu yaylası, On Picea orientalis. yaylası, 03.08.1995, 40°44’43’’ N 20.06.1994, 40°45’58’’ N 39°39’50’’ E, Usnea longissima Ach. 39°12’20’’ E, 1950 m.; Sürmene: Büyük 1650 m.; Akçaabat: Kayabaşı yaylası, yayla, 23.06.1995, 40°47’33’’ N 40°03’45’’ 24.08.1995, 40°51’10’’ N 39°29’46’’ E, Maçka; Şolma yaylası, 11.08.1994, E, 1700 m.; Vakfıkebir: Esentepe, 1850 m. Yazıcı 0624. 40°52’45’’ N 39°33’00’’ E, 1665 m., Güzel 21.10.1995, 41°00’56’’ N 39°14’28’’ E, 400 yayla, 20.08.1994, 40°52’35’’ N 39°34’55’’ On Picea orientalis. m.; Hayrat: Büyükharman yaylası, E, 1945 m. 1665 m.; Akçaabat: Hıdırnebi 29.07.1995, 40°55’00’’ N 40°24’45’’ E, ** Usnea esperantiana Clerc yaylası, 08.08.1993, 40°58’55’’ N 1645 m. Yazıcı 0362. 39°24’40’’ E, 1540 m.; Çaykara: Uzungöl Maçka: Hanzar Yaylası, 23.07.1995, yaylası, 19.07.1995, 40°44’37’’ N On calcareous rock. 40°50’00’’ N 39°30’02’’ E, 1760 m.; 40°19’30’’ E, 1690 m. Yazıcı 0350. Akçaabat: Hıdırnebi yaylası, 08.08.1993, Xanthoria parietina (L.) Th. Fr. 40°58’55’’ N 39°24’40’’ E, 1540 m. Yazıcı On Picea orientalis. This is the most common species between 0413. Usnea subfloridana Stirton 10 and 1800 meters. It was defined on On Picea orientalis. Robinia pseudoacacia, Corylus avellana, Maçka: Güzel yayla, 20.08.1994, Populus sp., Salix sp., Acer sp., Prunus sp., Usnea filipendula Stirton 40°52’35’’ N 39°34’55’’ E, 1945 m. Yazıcı Ficus sp., siliceous and calcareous rocks at all 0577. Çaykara: Uzungöl yaylası, 25.07.1994, stations. In addition as regards to choosing 40°44’37’’ N 40°19’30’’ E, 1690 m.; On Picea orientalis substrata, this species was understood to be one of the least sensitive. Sometimes it was Şalpazarı: Çağmanlı yaylası, 04.06.1995, Xanthoria calcicola Oxner 40°49’19’’ N 39°15’00’’ E, 1600 m.; Hayrat: seen together with Physcia adscendens and Semerdağ yaylası, 30.07.1993, 40°49’22’’ N Trabzon: Gürbulak köyü, 12.10.1995, rarely Physcia tenella. Yazıcı 0469. 41°00’13’’ N 39°36’09’’ E, 400 m., 40°28’30’’ E, 1755 m. Yazıcı 0389. ** Xanthoria polycarpa (Hoffm.) Th. Fr. Beşirli–Toklu köyü border, 26.11.1995, ex Rieb. In addition it was defined at a total of 15 41°01’37’’ N 39°38’08’’ E, 130 m., Center, stations in Maçka. Boztepe, 27.03.1995, 41°03’13’’ N Akçaabat. Hıdırnebi yaylası, 08.08.1993, On Picea orientalis. 39°40’50’’ E, 120 m., Karadeniz Technical 40°58’55’’ N 39°24’40’’ E, 1540 m.; University Campus, 13.03.1995, 41°03’20’’ Trabzon: Karadeniz Technical University Usnea florida (L.) Wigg. N 39°43’35’’ E, 130 m.; Araklı: Taşönü köyü, Campus, 21.10.1993, 41°03’20’’ N Hayrat: Gama yaylası, 27.07.1995, 10.10.1995, 40°01’12’’ N 39°59’42’’ E, 380 39°43’35’’ E, 130 m. Yazıcı 0531. 40°49’42’’ N 40°23’00’’ E, 1680 m., m.; Sürmene: Muratlı köyü, 18.03.1994, On Populus sp. and Acer sp. Semerdağ yaylası, 30.07.1993, 40°49’22’’ N 40°59’42’’ N 40°05’15’’ E, 250 m.; Hayrat: 40°28’30’’ E, 1755 m.; Araklı: Kirazlı yaylası, Ovacık köyü, 11.04.1995, 41°01’17’’ N ** Verrucaria glaucina auct. brit., non 17.07.1994, 40°41’00’’ N 39°59’20’’ E, 40°23’45’’ E, 650 m.; Yomra: Yokuşlu köyü, Ach. 1835 m.; Şalpazarı: Kızılkaya yaylası, 06.03.1994, 41°00’08’’ N 39°52’07’’ E, 260 Trabzon: Beşirli–Toklu köyü border, 14.06.1995, 40°46’15’’ N 39°12’00’’ E., m.; Çarşıbaşı: Kale köyü, 05.07.1995, 26.11.1995, 41°01’37’’ N 39°38’08’’ E, 130 1650 m.; Tonya: Bodamişkiye yaylası, 41°04’28’’ E 39°20’30’’ E, 250 m. Yazıcı m. Yazıcı 0333. 26.07.1993, 40°43’47’’ N 39°17’57’’ E, 0337. 1800 m.; Akçaabat: Ormana yaylası, On calcareous rock.
110 K. YAZICI
Findings in this region. Especially, Evernia prunastri, Ramalina In this study, a total of new 146 lichen species was spp., Usnea spp., Pseudevernia furfuracea var. furfuracea determined for Trabzon. 39 of which were new records and Hypogymnia spp. were observed to grow abundantly for Turkey. A total of 165 species belong to Lecanorales, on bark of Picea orientalis. In addition in the forests of 48 of which belong to the family Parmeliaceae. Maçka Cetraria islandica on mosses, Cladonia cenotea, Cladonia coniocraea on decayed bark of Picea orientalis Of the species, 91 are foliose, 73 are crustose, 51 are and Nephroma parile on mosses and dried branches of fruticose, 14 are squamulose, 1 is flamentous, 1 is Picea orientalis were abundant as well. placodioid and 1 is leprose. The substrata features of the investigated species are A total of 79 lichen species were observed to be consistent with the findings in the literature (12–14). epiphytic only, 91 as saxicolous, 14 as only terricolous, 19 as epiphytic and saxicolous, 9 as epiphytic and terricolous, 2 as terricolous and saxicolous. In addition, 4 Discussion of the species were found on decayed bark of Picea A total of 172 different lichen species have been orientalis (L.) Link., mosses and soil. One was determined defined in Trabzon so far. Steiner, in 1909, defined 51 to grow on decayed bark of Picea orientalis and dried lichen species with substrata. Szatala, in 1960, reported branches of Picea orientalis. Two species grew on decayed 39 lichen speices. Anşin, in 1979, defined 10 lichen and dried barks of Picea orientalis and mosses. One species in Maçka district. Now, the names of most of species was found to grow on Picea orientalis only. The these species are synonyms. Although Szatala gave other one was found on decayed and dried bark of Picea Trabzon as a station name unfortunately he has not orientalis, siliceous rock and mosses. presented any exact locations. Moreover, the station As regards to choosing substrata in the study area, names used in the studies by Steiner have been changed Xanthoria parietina, Lecanora campestris, Parmelia and they are impossible to pinpoint. In addition, some of caperata, Physcia tenella and Physica adscendens were the stations in these studies have been urbanised or understood to be the least sensitive. Xanthoria parietina, settled. Therefore some lichen species, which cannot Parmelia caperata and Lecanora campestris grew on 7 endure air pollution, could not be defined. These are different substrata. On the other hand Physcia Catillaria lenticularis, Pertusaria amara, Pertusaria lactea, adscendens and Physca tenella grew on 5 different Lecanora campestris, Lecanora gangaleoides, Cladonia substrata. In coastal areas commonly found species were parasitica, Collema flaccidum, Leptogium gelatinosum, Colema auriforme, Xanthoria parietina, Parmelia Lecanora allophana and Ramalina farinacea. However, at caperata, Physcia adscendens, Lepraria incana, Cladonia these stations, air pollution resistant species such as rangiformis and Lecanora muralis. Caloplaca decipiens, Caloplaca flavescens, Caloplaca The most common species was Rhizocarpon holocarpa, Caloplaca irrubescens, Xanthoria aureola, geographicum growing on siliceous rocks on plateaus Xanthoria parietina and Physcia adscendens were above 1500 m. in Maçka, Çaykara, Düzköy, Akçabat, determined. Sürmene, Tonya, Köprübaşı, Şalpazarı, Araklı and Hayrat. Anşin, in 1979, defined 10 lichen species in Altındere There were a few species as common as Rhizocarpon Valley Sumela Monastery and surroundings in Maçka: geographicum at 800–1200 m. in the same regions. Cladonia pyxidata, Cladonia rangiformis, Cladonia There were mainly Cetraria delisei, Cetraria ericetorum, rangiferina, Cladonia arbuscula, Lobaria pulmonaria, Cetraria aculeata, Cetraria muricata, Umbilicaria Peltigera polydactylon, Pseudevernia furfuracea var. cylindrica, Nephroma parile, Pseudevernia furfuracea var. furfuracea, Rhizocarpon geographicum, Usnea florida and furfuracea, Ramalina farinacea and Hypogymnia Xanthoria parietina. They have been defined in Maçka by physodes. Yazıcı and Cevahir except Cladonia arbuscula and Cladonia Peltigera horizontalis, Peltigera praetextata and rangiferina (16, 17). But Cladonia rangiferina was found Peltigera canina were fairly common on mosses at in Maçka in this study. altitudes of 130 to 1750 m. and are the most common Usnea subglabra and Usnea cylindrica, which were species in Maçka, Tonya, Çaykara, Akçaabat, Sürmene, defined by John in Maçka: Sumela and Çaykara: Uzungöl Hayrat, Şalpazarı and Araklı (15). in 1990 (18, 19), were defined in Maçka in this study. The fact that Maçka county has a richer plant cover In addition, Lobaria pulmonaria, Menegazzia than other districts enables many lichen species to grow terebrata, Heterodermia speciosa, Parmelia caperata and
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Anaptychia ciliaris, which were defined in Maçka: Sumela Acknowledgement and in Çaykara: Uzungöl in 1990 (20), were found in I would like to thank Assistant Professor Dr. Ayşen Maçka and Çaykara: Uzungöl in this study. TÜRK, Dr. Volker JOHN, Dr. Philippe CLERK and Dr. But Lasallia rossica Dombrovskaja defined by John in Andre APTROOT for their useful discussion concerning Maçka: Sumela in 1990 (18) was not found in this study. the identification of some lichens. I also thank Forest Engineer Yusuf ŞAHİN.
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