DOCSLIB.ORG

Safety of Bacopa Monnieri (Brahmi) for Cognitive and Brain Enhancement

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Safety of Bacopa Monnieri (Brahmi) for Cognitive and Brain Enhancement Evidence-Based Complementary and Alternative Medicine Mechanisms, Efficacy, and Safety of Bacopa monnieri (Brahmi) for Cognitive and Brain Enhancement Guest Editors: Con Stough, Hemant Singh, and Andrea Zangara Mechanisms, Efficacy, and Safety of Bacopa monnieri (Brahmi) for Cognitive and Brain Enhancement Evidence-Based Complementary and Alternative Medicine Mechanisms, Efficacy, and Safety of Bacopa monnieri (Brahmi) for Cognitive and Brain Enhancement Guest Editors: Con Stough, Hemant Singh, and Andrea Zangara Copyright © 2015 Hindawi Publishing Corporation. All rights reserved. This is a special issue published in “Evidence-Based Complementary and Alternative Medicine.” All articles are open access articles distributed under the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. Editorial Board Mona Abdel-Tawab, Germany Il-Moo Chang, Republic of Korea Brett Froeliger, USA Jon Adams, Australia Chun-Tao Che, USA Maria pia Fuggetta, Italy GabrielA.Agbor,Cameroon Kevin Chen, USA Joel J. Gagnier, Canada Ulysses P. Albuquerque, Brazil Evan P. Cherniack, USA Siew Hua Gan, Malaysia Samir Lutf Aleryani, USA Salvatore Chirumbolo, Italy Jian-Li Gao, China Ather Ali, USA Jae Youl Cho, Korea Mary K. Garcia, USA A.-Shtayeh, Palestinian Authority K. Christensen, Denmark S. Garcia de Arriba, Germany Gianni Allais, Italy Shuang-En Chuang, Taiwan D. G. Gimenez,´ Spain Terje Alraek, Norway Y. Clement, Trinidad And Tobago Gabino Garrido, Chile Shrikant Anant, USA Paolo Coghi, Italy Ipek Goktepe, Qatar Isabel Andujar,´ Spain Marisa Colone, Italy Michael Goldstein, USA Letizia Angiolella, Italy Lisa A. Conboy, USA Yuewen Gong, Canada Virginia A. Aparicio, Spain Kieran Cooley, Canada Settimio Grimaldi, Italy Makoto Arai, Japan Edwin L. Cooper, USA Gloria Gronowicz, USA Hyunsu Bae, Republic of Korea Olivia Corcoran, UK Maruti Ram Gudavalli, USA Onesmo B. Balemba, USA Muriel Cuendet, Switzerland Alessandra Guerrini, Italy Winfried Banzer, Germany RobertoK.N.Cuman,Brazil Narcis Gusi, Spain Panos Barlas, UK Vincenzo De Feo, Italy Svein Haavik, Norway Vernon A. Barnes, USA Roco De la Puerta, Spain Solomon Habtemariam, UK Samra Bashir, Pakistan Laura De Martino, Italy Abid Hamid, India Purusotam Basnet, Norway Nunziatina De Tommasi, Italy Michael G. Hammes, Germany Jairo Kennup Bastos, Brazil Alexandra Deters, Germany K. B. Harikumar, India Arpita Basu, USA Farzad Deyhim, USA Cory S. Harris, Canada Sujit Basu, USA Manuela Di Franco, Italy Jan Hartvigsen, Denmark George David Baxter, New Zealand Claudia Di Giacomo, Italy Thierry Hennebelle, France Andre-Michael´ Beer, Germany Antonella Di Sotto, Italy Lise Hestbaek, Denmark Alvin J. Beitz, USA M. Dijoux-Franca, France Eleanor Holroyd, Australia Louise Bennett, Australia Luciana Dini, Italy Markus Horneber, Germany Maria Camilla Bergonzi, Italy Tieraona L. Dog, USA Ching-Liang Hsieh, Taiwan Anna R. Bilia, Italy Caigan Du, Canada BennyT.K.Huat,Singapore Yong C. Boo, Republic of Korea Jeng-Ren Duann, USA Roman Huber, Germany Monica Borgatti, Italy Nativ Dudai, Israel Helmut Hugel, Australia Francesca Borrelli, Italy Thomas Efferth, Germany Ciara Hughes, UK Gloria Brusotti, Italy Abir El-Alfy, USA Attila Hunyadi, Hungary Arndt Bussing,¨ Germany Tobias Esch, USA Sumiko Hyuga, Japan Rainer W. Bussmann, USA Giuseppe Esposito, Italy H. Stephen Injeyan, Canada Andrew J. Butler, USA Keturah R. Faurot, USA Chie Ishikawa, Japan Gioacchino Calapai, Italy Yibin Feng, Hong Kong Angelo A. Izzo, Italy Giuseppe Caminiti, Italy Nianping Feng, China Chris J. Branford-White, UK Raffaele Capasso, Italy Patricia D. Fernandes, Brazil Suresh Jadhav, India Francesco Cardini, Italy Josue Fernandez-Carnero, Spain G. K. Jayaprakasha, USA Opher Caspi, Israel Antonella Fioravanti, Italy Stefanie Joos, Germany Subrata Chakrabarti, Canada Fabio Firenzuoli, Italy ZeevLKain,USA Pierre Champy, France Peter Fisher, UK Osamu Kanauchi, Japan Shun-Wan Chan, Hong Kong Filippo Fratini, Italy Wenyi Kang, China Shao-Hsuan Kao, Taiwan Albert S Mellick, Australia Elia Ranzato, Italy Juntra Karbwang, Japan A. G. Mensah-Nyagan, France Ke Ren, USA Kenji Kawakita, Japan Andreas Michalsen, Germany Man-Hee Rhee, Republic of Korea D. A. Kennedy, Canada Oliver Micke, Germany Luigi Ricciardiello, Italy Youn C. Kim, Republic of Korea Roberto Miniero, Italy Daniela Rigano, Italy C. Kim, Republic of Korea Giovanni Mirabella, Italy JoseL.R´ ´ıos, Spain Yoshiyuki Kimura, Japan David Mischoulon, USA Paolo Roberti di Sarsina, Italy Toshiaki Kogure, Japan Francesca Mondello, Italy Mariangela Rondanelli, Italy Jian Kong, USA Albert Moraska, USA Omar Said, Israel Tetsuya Konishi, Japan Giuseppe Morgia, Italy Avni Sali, Australia Karin Kraft, Germany Mark Moss, UK Mohd Z. Salleh, Malaysia Omer Kucuk, USA Yoshiharu Motoo, Japan A. Sandner-Kiesling, Austria Victor Kuete, Cameroon KamalD.Moudgil,USA Manel Santafe, Spain Yiu W. Kwan, Hong Kong Yoshiki Mukudai, Japan Tadaaki Satou, Japan Kuang C. Lai, Taiwan Frauke Musial, Germany Michael A. Savka, USA Ilaria Lampronti, Italy MinKyun Na, Republic of Korea Claudia Scherr, Switzerland Lixing Lao, Hong Kong Hajime Nakae, Japan G. Schmeda-Hirschmann, Chile Christian Lehmann, Canada Srinivas Nammi, Australia Andrew Scholey, Australia Marco Leonti, Italy Krishnadas Nandakumar, India Roland Schoop, Switzerland Lawrence Leung, Canada Vitaly Napadow, USA Sven Schroder,¨ Germany Shahar Lev-ari, Israel Michele Navarra, Italy Herbert Schwabl, Switzerland Chun G. Li, Australia Isabella Neri, Italy Veronique Seidel, UK Xiu-Min Li, USA Pratibha V. Nerurkar, USA Senthamil Selvan, USA Min Li, China Karen Nieber, Germany Felice Senatore, Italy Ho Lin, Taiwan Menachem Oberbaum, Israel Hongcai Shang, China Bi-Fong Lin, Taiwan Martin Offenbaecher, Germany Ronald Sherman, USA C. G. Lis, USA Junetsu Ogasawara, Japan Karen J. Sherman, USA G. Litscher, Austria Ki-Wan Oh, Republic of Korea Kuniyoshi Shimizu, Japan I-Min Liu, Taiwan Yoshiji Ohta, Japan Kan Shimpo, Japan Yijun Liu, USA Olumayokun A. Olajide, UK Yukihiro Shoyama, Japan V. L opez,´ Spain Thomas Ostermann, Germany Morry Silberstein, Australia T. Lundeberg, Sweden Siyaram Pandey, Canada K. S. Sirajudeen, Malaysia Filippo Maggi, Italy Bhushan Patwardhan, India Graeme Smith, UK V. Ma g g i n i , It a l y BeritS.Paulsen,Norway Chang-Gue Son, Korea Gail B. Mahady, USA Philip Peplow, New Zealand Rachid Soulimani, France Jamal Mahajna, Israel Florian Pfab, Germany Didier Stien, France Juraj Majtan, Slovakia Sonia Piacente, Italy Con Stough, Australia Francesca Mancianti, Italy Andrea Pieroni, Italy Annarita Stringaro, Italy Carmen Mannucci, Italy Richard Pietras, USA Shan-Yu Su, Taiwan Arroyo-Morales Manuel, Spain Andrew Pipingas, Australia Barbara Swanson, USA Fulvio Marzatico, Italy Jose M. Prieto, UK Giuseppe Tagarelli, Italy Marta Marzotto, Italy Haifa Qiao, USA O. Taglialatela-Scafati, Italy James H. McAuley, Australia Waris Qidwai, Pakistan Takashi Takeda, Japan Kristine McGrath, Australia Xianqin Qu, Australia Ghee T. Tan, USA James S. McLay, UK E. Ferreira Queiroz, Switzerland Hirofumi Tanaka, USA Lewis Mehl-Madrona, USA Roja Rahimi, Iran Lay Kek Teh, Malaysia Peter Meiser, Germany Khalid Rahman, UK Norman Temple, Canada Karin Meissner, Germany Cheppail Ramachandran, USA Mayank Thakur, Germany Menaka C. Thounaojam, USA Giuseppe Venturella, Italy Jenny M. Wilkinson, Australia Evelin Tiralongo, Australia Pradeep Visen, Canada D. R. Williams, Republic of Korea Stephanie Tjen-A-Looi, USA Aristo Vojdani, USA C. Worsnop, Australia MichaThlfj Tomczyk, Poland Dawn Wallerstedt, USA Haruki Yamada, Japan Loren Toussaint, USA Shu-Ming Wang, USA Nobuo Yamaguchi, Japan Yew-Min Tzeng, Taiwan Chong-Zhi Wang, USA Ling Yang, China Dawn M. Upchurch, USA Yong Wang, USA Junqing Yang, China Konrad Urech, Switzerland Jonathan L. Wardle, Australia Eun J. Yang, Republic of Korea Takuhiro Uto, Japan Kenji Watanabe, Japan Ken Yasukawa, Japan Sandy van Vuuren, South Africa J. Wattanathorn, Thailand Albert S. Yeung, USA Alfredo Vannacci, Italy Michael Weber, Germany Armando Zarrelli, Italy Subramanyam Vemulpad, Australia Silvia Wein, Germany C. Zaslawski, Australia Carlo Ventura, Italy Janelle Wheat, Australia Ruixin Zhang, USA Contents Mechanisms, Efficacy, and Safety of Bacopa monnieri (Brahmi) for Cognitive and Brain Enhancement, Con Stough, Hemant Singh, and Andrea Zangara Volume 2015, Article ID 717605, 2 pages Bacopa monniera (CDRI-08) Upregulates the Expression of Neuronal and Glial Plasticity Markers in the Brain of Scopolamine Induced Amnesic Mice, Arpita Konar, Akash Gautam, and M. K. Thakur Volume 2015, Article ID 837012, 9 pages Molecular and Functional Characterization of Bacopa monniera:ARetrospectiveReview, Koilmani Emmanuvel Rajan, Jayakumar Preethi, and Hemant K. Singh Volume 2015, Article ID 945217, 12 pages ASpecialExtractofBacopa monnieri (CDRI-08) Restores Learning and Memory by Upregulating Expression of the NMDA Receptor Subunit GluN2B in the Brain of Scopolamine-Induced Amnesic Mice,RakeshRai,HemantK.Singh,andS.Prasad Volume 2015, Article ID 254303, 13 pages CDRI-08 Attenuates REST/NRSF-Mediated Expression of NMDAR1 Gene in PBDE-209-Exposed Mice Brain,PriyaVerma,RajaneeshK.Gupta,BehroseS.Gandhi,andPoonamSingh Volume 2015, Article ID 403840, 10 pages Bacopa monnieri Extract (CDRI-08) Modulates the NMDA Receptor Subunits and nNOS-Apoptosis Axis in Cerebellum of Hepatic Encephalopathy Rats, Papia Mondal and Surendra Kumar Trigun Volume 2015, Article ID 535013, 8 pages Bacopa
Load more

Recommended publications
  • Sutera Cordata C.E.O. Kuntze Bacopa (Chaenostoma Cordatum, Bacopa Cordata, Sutera Diffusa)
    Sutera cordata C.E.O. Kuntze Bacopa (Chaenostoma cordatum, Bacopa cordata, Sutera diffusa) Other Common Names: Sutera. Family: Scrophulariaceae. Cold Hardiness: Cold tolerant and used as a winter annual or perennial in USDA zones 9(8b) through 11; often used as a transition or summer annual in cooler climates. Foliage: Evergreen; opposite; simple; rhombic-ovate to nearly cordate; small, dO to eO(¾O) long, and thickish; tips acute; margins distally dentate-serrate; palmately veined, faintly impressed above and lightly raised beneath; base cuneate, rounded, or nearly cordate; the blade is medium to dark green; petioles are ¼O to ½Olong, green, and sometimes winged with tiny glandular pubescence. Flower: Perfect; terminal; five-petals fused at the base into a narrow throat and recurved distally to form a flattened small five-lobed white flower; the corolla is surrounded by a five-lobed green calyx; additional colors in the blue, pink, and lavender range are being developed via hybridization with other species. Fruit: Tiny capsules; not ornamental; deadheading is not required. 1 Stem / Bark: Stems — stiff; mostly prostrate; green with tiny glandular pubescence; Buds — tiny, /32O or less in length; foliose; green; Bark — not applicable in our region. Habit: Bacopa forms a flat spreading semi-evergreen rounded herbaceous mat of trailing stems and under cultivation in our region plants are only 2O to 4O(8O) tall with a 2N to 4N spread; in its native land it may develop into a woody subshrub mounding to 24O tall with an indefinite spread; the overall texture is fine. Cultural Requirements: Morning sun and afternoon shade is best, but plants can tolerate full sun to partial shade if a steady moisture supply is available; plants frequently tend to slump or succumb to the heat of our summers, but if they survive, they may return to flower in autumn; plants require a moist well drained fertile soil and are not particularly drought tolerant; recovery from severe drought stress is poor.
    [Show full text]
  • Ethnomedicinal Profile of Flora of District Sialkot, Punjab, Pakistan
    ISSN: 2717-8161 RESEARCH ARTICLE New Trend Med Sci 2020; 1(2): 65-83. https://dergipark.org.tr/tr/pub/ntms Ethnomedicinal Profile of Flora of District Sialkot, Punjab, Pakistan Fozia Noreen1*, Mishal Choudri2, Shazia Noureen3, Muhammad Adil4, Madeeha Yaqoob4, Asma Kiran4, Fizza Cheema4, Faiza Sajjad4, Usman Muhaq4 1Department of Chemistry, Faculty of Natural Sciences, University of Sialkot, Punjab, Pakistan 2Department of Statistics, Faculty of Natural Sciences, University of Sialkot, Punjab, Pakistan 3Governament Degree College for Women, Malakwal, District Mandi Bahauddin, Punjab, Pakistan 4Department of Chemistry, Faculty of Natural Sciences, University of Gujrat Sialkot Subcampus, Punjab, Pakistan Article History Abstract: An ethnomedicinal profile of 112 species of remedial Received 30 May 2020 herbs, shrubs, and trees of 61 families with significant Accepted 01 June 2020 Published Online 30 Sep 2020 gastrointestinal, antimicrobial, cardiovascular, herpetological, renal, dermatological, hormonal, analgesic and antipyretic applications *Corresponding Author have been explored systematically by circulating semi-structured Fozia Noreen and unstructured questionnaires and open ended interviews from 40- Department of Chemistry, Faculty of Natural Sciences, 74 years old mature local medicine men having considerable University of Sialkot, professional experience of 10-50 years in all the four geographically Punjab, Pakistan diversified subdivisions i.e. Sialkot, Daska, Sambrial and Pasrur of E-mail: [email protected] district Sialkot with a total area of 3106 square kilometres with ORCID:http://orcid.org/0000-0001-6096-2568 population density of 1259/km2, in order to unveil botanical flora for world. Family Fabaceae is found to be the most frequent and dominant family of the region. © 2020 NTMS.
    [Show full text]
  • Medicinal Practices of Sacred Natural Sites: a Socio-Religious Approach for Successful Implementation of Primary
    Medicinal practices of sacred natural sites: a socio-religious approach for successful implementation of primary healthcare services Rajasri Ray and Avik Ray Review Correspondence Abstract Rajasri Ray*, Avik Ray Centre for studies in Ethnobiology, Biodiversity and Background: Sacred groves are model systems that Sustainability (CEiBa), Malda - 732103, West have the potential to contribute to rural healthcare Bengal, India owing to their medicinal floral diversity and strong social acceptance. *Corresponding Author: Rajasri Ray; [email protected] Methods: We examined this idea employing ethnomedicinal plants and their application Ethnobotany Research & Applications documented from sacred groves across India. A total 20:34 (2020) of 65 published documents were shortlisted for the Key words: AYUSH; Ethnomedicine; Medicinal plant; preparation of database and statistical analysis. Sacred grove; Spatial fidelity; Tropical diseases Standard ethnobotanical indices and mapping were used to capture the current trend. Background Results: A total of 1247 species from 152 families Human-nature interaction has been long entwined in has been documented for use against eighteen the history of humanity. Apart from deriving natural categories of diseases common in tropical and sub- resources, humans have a deep rooted tradition of tropical landscapes. Though the reported species venerating nature which is extensively observed are clustered around a few widely distributed across continents (Verschuuren 2010). The tradition families, 71% of them are uniquely represented from has attracted attention of researchers and policy- any single biogeographic region. The use of multiple makers for its impact on local ecological and socio- species in treating an ailment, high use value of the economic dynamics. Ethnomedicine that emanated popular plants, and cross-community similarity in from this tradition, deals health issues with nature- disease treatment reflects rich community wisdom to derived resources.
    [Show full text]
  • Pharmacognostic and Pharmacological Aspect of Bacopa Monnieri: a Review
    Vol 4, Issue 3, 2016 ISSN- 2321-6824 Review Article PHARMACOGNOSTIC AND PHARMACOLOGICAL ASPECT OF BACOPA MONNIERI: A REVIEW PUSHPENDRA KUMAR JAIN1, DEBAJYOTI DAS2, PUNEET JAIN3*, PRACHI JAIN4 1Department of Pharmacy, Naraina Vidyapeeth Group of Institutions, Panki-Kanpur, Uttar Pradesh, India. 2Department of Pharmacy, School of Pharmaceutical Sciences, Siksha ‘O’ Anusandhan University, Bhubaneswar, Odisha, India. 3Maharana Pratap Education Center, Kalyanpur, Kanpur, Uttar Pradesh, India. 4Dr. Virendra Swarup Education Centre, Panki, Kanpur, Uttar Pradesh, India. Email: [email protected] Received: 25 April 2016, Revised and Accepted: 29 April 2016 ABSTRACT It is said that the use of Bacopa monnieri (BM) for memory enhancement goes back 3000 years or more in India, when it was cited for its medicinal properties, especially the memory enhancing capacity, in the vedic texts “Athar-Ved Samhila” (3:1) of 800 BC and in Ayurveda. In the folklore of Indian medicine, several herbs have been used traditionally as brain or nerve tonics. One of the most popular of these neurotonics is BM, a well-known memory booster. Brahmi has been administered at religious institutions to help students to enhance their memory for learning ancient, religious hymns. It is also used as cardio-tonic, tranquilizer and sedative, improves the process of learning, restores memory, and enhances power of speech and imagination, diuretic and nervine tonic, antistress, for nervous and mental strain, use in insanity, epilepsy, hysteria, esthenia, nervous breakdown. It is a small, creeping succulent herb. The leaf and flower bearing stems are 10-30 cm long and arise from creeping stems that form roots at the nodes with pale blue or pinkish white flowers belonging to family Scrophulariaceae grown nearly banks of freshwater streams and ponds, paddy fields, and other damp places.
    [Show full text]
  • Towards Resolving Lamiales Relationships
    Schäferhoff et al. BMC Evolutionary Biology 2010, 10:352 http://www.biomedcentral.com/1471-2148/10/352 RESEARCH ARTICLE Open Access Towards resolving Lamiales relationships: insights from rapidly evolving chloroplast sequences Bastian Schäferhoff1*, Andreas Fleischmann2, Eberhard Fischer3, Dirk C Albach4, Thomas Borsch5, Günther Heubl2, Kai F Müller1 Abstract Background: In the large angiosperm order Lamiales, a diverse array of highly specialized life strategies such as carnivory, parasitism, epiphytism, and desiccation tolerance occur, and some lineages possess drastically accelerated DNA substitutional rates or miniaturized genomes. However, understanding the evolution of these phenomena in the order, and clarifying borders of and relationships among lamialean families, has been hindered by largely unresolved trees in the past. Results: Our analysis of the rapidly evolving trnK/matK, trnL-F and rps16 chloroplast regions enabled us to infer more precise phylogenetic hypotheses for the Lamiales. Relationships among the nine first-branching families in the Lamiales tree are now resolved with very strong support. Subsequent to Plocospermataceae, a clade consisting of Carlemanniaceae plus Oleaceae branches, followed by Tetrachondraceae and a newly inferred clade composed of Gesneriaceae plus Calceolariaceae, which is also supported by morphological characters. Plantaginaceae (incl. Gratioleae) and Scrophulariaceae are well separated in the backbone grade; Lamiaceae and Verbenaceae appear in distant clades, while the recently described Linderniaceae are confirmed to be monophyletic and in an isolated position. Conclusions: Confidence about deep nodes of the Lamiales tree is an important step towards understanding the evolutionary diversification of a major clade of flowering plants. The degree of resolution obtained here now provides a first opportunity to discuss the evolution of morphological and biochemical traits in Lamiales.
    [Show full text]
  • Potential Herb–Drug Interactions in the Management of Age-Related Cognitive Dysfunction
    pharmaceutics Review Potential Herb–Drug Interactions in the Management of Age-Related Cognitive Dysfunction Maria D. Auxtero 1, Susana Chalante 1,Mário R. Abade 1 , Rui Jorge 1,2,3 and Ana I. Fernandes 1,* 1 CiiEM, Interdisciplinary Research Centre Egas Moniz, Instituto Universitário Egas Moniz, Quinta da Granja, Monte de Caparica, 2829-511 Caparica, Portugal; [email protected] (M.D.A.); [email protected] (S.C.); [email protected] (M.R.A.); [email protected] (R.J.) 2 Polytechnic Institute of Santarém, School of Agriculture, Quinta do Galinheiro, 2001-904 Santarém, Portugal 3 CIEQV, Life Quality Research Centre, IPSantarém/IPLeiria, Avenida Dr. Mário Soares, 110, 2040-413 Rio Maior, Portugal * Correspondence: [email protected]; Tel.: +35-12-1294-6823 Abstract: Late-life mild cognitive impairment and dementia represent a significant burden on health- care systems and a unique challenge to medicine due to the currently limited treatment options. Plant phytochemicals have been considered in alternative, or complementary, prevention and treat- ment strategies. Herbals are consumed as such, or as food supplements, whose consumption has recently increased. However, these products are not exempt from adverse effects and pharmaco- logical interactions, presenting a special risk in aged, polymedicated individuals. Understanding pharmacokinetic and pharmacodynamic interactions is warranted to avoid undesirable adverse drug reactions, which may result in unwanted side-effects or therapeutic failure. The present study reviews the potential interactions between selected bioactive compounds (170) used by seniors for cognitive enhancement and representative drugs of 10 pharmacotherapeutic classes commonly prescribed to the middle-aged adults, often multimorbid and polymedicated, to anticipate and prevent risks arising from their co-administration.
    [Show full text]
  • Lemon Bacopa (Bacopa Caroliniana)
    Lemon bacopa (Bacopa caroliniana) For definitions of botanical terms, visit en.wikipedia.org/wiki/Glossary_of_botanical_terms. Lemon bacopa is a low-growing, herbaceous wildflower that occurs naturally in very moist to aquatic habitats such as along pond and stream margins, and in swamps, marshes and shallow ditches. It typically blooms late spring through fall, but can bloom year- round. Its nectar attracts a variety of small pollinators. Lemon bacopa’s small but showy, purplish-blue flowers are 5-lobed, tubular and copious. Seeds are borne in inconspicuous capsules. Stems are succulent and hairy. Leaves are succulent, clasping and oppositely arranged. They emit a lemony scent when bruised or crushed, giving the plant its common name. They can also be steeped in water to make a flavorful, lemony tea. A close relative of lemon bacopa is the more commonly Photo by Eleanor Dietrich occurring herb-of-grace (aka waterhyssop) (Bacopa monnieri). Although they look similar and are found in similar habitats, herb-of-grace does not emit a lemon scent. As well, its flowers are pale lavender to white and its leaves are not clasping. Neither Bacopa species are related to hyssop (Hyssopus sp.), which is in the mint (Lamiaceae) family. Family: Plantaginaceae (Plantain family) Native range: Nearly throughout Florida To see where natural populations of Lemon bacopa have been vouchered, visit www.florida.plantatlas.usf.edu. Hardiness: Zones 8–11 Lifespan: Perennial Soil: Moist to wet (saturated or inundated), slightly acidic soils Exposure: Full sun to minimal shade Growth habit: up to 6” tall but widespreading Propagation: Cuttings, division Garden tips: Lemon bacopa makes an excellent groundcover in wet or saturated landscapes.
    [Show full text]
  • Hairy Water Hyssop (Bacopa Lanigera)
    JULY 2012 TM YOUR ALERT TO NEW AND EMERGING THREATS. 1. 2. 3. 4. 1. Habit with numerous spreading stems. 2. Close-up of paired leaves and stem with dense spreading hairs. 3. Piece of stem rooting at the joints. 4. Small bluish-purplish flowers. Hairy water hyssop (Bacopa lanigera) Introduced Not Declared AQUATIC Quick Facts Hairy water hyssop is a small aquatic plant native to South America (i.e. Brazil and Paraguay) that is also known as hairy bacopa. It is > A creeping aquatic plant that forms very dense mats in mud or shallow sometimes grown as an aquarium plant and is becoming established water in wetter sites in the coastal parts of eastern Australia. > Stems produce roots where they come into contact with soil Distribution > Stems densely covered in spreading Hairy water hyssop has been in Australia for some time, but was mistaken for the similar Water hairs hyssop (Bacopa caroliniana) until 2009. The first known occurrence of it becoming established > Small leaves borne in pairs and outside cultivation in Australia was in 1999, when it was recorded growing all over a small almost rounded in shape farm dam in the Beaudesert area. It has since been recorded at a couple of locations in Logan City, in a waterway in Moreton Bay Regional Council, and along a creek near Port Douglas in > Small bluish-purple flowers borne northern Queensland. More recently, infestations have been noted growing around a farm dam singly on slender stalks in Redland City and in a small wetland at the Gold Coast. Habitat Description This species is mostly commonly found growing in This plant is capable of growing underwater, on the water surface, or on land.
    [Show full text]
  • Lamiales – Synoptical Classification Vers
    Lamiales – Synoptical classification vers. 2.6.2 (in prog.) Updated: 12 April, 2016 A Synoptical Classification of the Lamiales Version 2.6.2 (This is a working document) Compiled by Richard Olmstead With the help of: D. Albach, P. Beardsley, D. Bedigian, B. Bremer, P. Cantino, J. Chau, J. L. Clark, B. Drew, P. Garnock- Jones, S. Grose (Heydler), R. Harley, H.-D. Ihlenfeldt, B. Li, L. Lohmann, S. Mathews, L. McDade, K. Müller, E. Norman, N. O’Leary, B. Oxelman, J. Reveal, R. Scotland, J. Smith, D. Tank, E. Tripp, S. Wagstaff, E. Wallander, A. Weber, A. Wolfe, A. Wortley, N. Young, M. Zjhra, and many others [estimated 25 families, 1041 genera, and ca. 21,878 species in Lamiales] The goal of this project is to produce a working infraordinal classification of the Lamiales to genus with information on distribution and species richness. All recognized taxa will be clades; adherence to Linnaean ranks is optional. Synonymy is very incomplete (comprehensive synonymy is not a goal of the project, but could be incorporated). Although I anticipate producing a publishable version of this classification at a future date, my near- term goal is to produce a web-accessible version, which will be available to the public and which will be updated regularly through input from systematists familiar with taxa within the Lamiales. For further information on the project and to provide information for future versions, please contact R. Olmstead via email at [email protected], or by regular mail at: Department of Biology, Box 355325, University of Washington, Seattle WA 98195, USA.
    [Show full text]
  • The Linderniaceae and Gratiolaceae Are Further Lineages Distinct from the Scrophulariaceae (Lamiales)
    Research Paper 1 The Linderniaceae and Gratiolaceae are further Lineages Distinct from the Scrophulariaceae (Lamiales) R. Rahmanzadeh1, K. Müller2, E. Fischer3, D. Bartels1, and T. Borsch2 1 Institut für Molekulare Physiologie und Biotechnologie der Pflanzen, Universität Bonn, Kirschallee 1, 53115 Bonn, Germany 2 Nees-Institut für Biodiversität der Pflanzen, Universität Bonn, Meckenheimer Allee 170, 53115 Bonn, Germany 3 Institut für Integrierte Naturwissenschaften ± Biologie, Universität Koblenz-Landau, Universitätsstraûe 1, 56070 Koblenz, Germany Received: July 14, 2004; Accepted: September 22, 2004 Abstract: The Lamiales are one of the largest orders of angio- Traditionally, Craterostigma, Lindernia and their relatives have sperms, with about 22000 species. The Scrophulariaceae, as been treated as members of the family Scrophulariaceae in the one of their most important families, has recently been shown order Lamiales (e.g., Takhtajan,1997). Although it is well estab- to be polyphyletic. As a consequence, this family was re-classi- lished that the Plocospermataceae and Oleaceae are their first fied and several groups of former scrophulariaceous genera branching families (Bremer et al., 2002; Hilu et al., 2003; Soltis now belong to different families, such as the Calceolariaceae, et al., 2000), little is known about the evolutionary diversifica- Plantaginaceae, or Phrymaceae. In the present study, relation- tion of most of the orders diversity. The Lamiales branching ships of the genera Craterostigma, Lindernia and its allies, hith- above the Plocospermataceae and Oleaceae are called ªcore erto classified within the Scrophulariaceae, were analyzed. Se- Lamialesº in the following text. The most recent classification quences of the chloroplast trnK intron and the matK gene by the Angiosperm Phylogeny Group (APG2, 2003) recognizes (~ 2.5 kb) were generated for representatives of all major line- 20 families.
    [Show full text]
  • Lemon Bacopa: Bacopa Caroliniana1 Lyn Gettys and Carl J
    SS-AGR-388 Lemon Bacopa: Bacopa caroliniana1 Lyn Gettys and Carl J. Della Torre III2 Introduction Lemon bacopa is an herbaceous aquatic perennial native plant that commonly grows along shorelines, in “wet feet” areas, and in water that is less than 3' deep. The leaves of lemon bacopa are small (less than 3/4" in length), nearly round, thick, fleshy, and succulent. When the species grows as an emergent plant (with most or all of the plant out of the water), the stems are often covered with fine hairs and may appear velvety. In contrast, the stems of submersed plants (with most or all of the plant under water) are hairless or nearly hairless. Crush the leaves of this delightful plant to release a fragrance reminiscent of lemon, lime, anise, or licorice. The species produces lovely, bright-blue flowers, which make it an attractive native addition to water gardens, aquascapes, and restoration and mitigation Figure 1. Lemon bacopa flower. projects. Lemon bacopa is a host for larvae of the white Credits: Lyn Gettys, UF/IFAS peacock butterfly (Anartia jatrophae) (Florida Native Plant Society 2015). Lemon bacopa is widely available from a Description and Habitat variety of sources, including stores that carry aquarium Lemon bacopa is very common in Florida and occurs in plants, water garden supply shops, and nurseries that sell 56 of 67 counties, but the species is also widely distributed aquatic or native plants. throughout the southeastern United States. Populations of lemon bacopa have been reported in eastern Texas, South Classification Carolina, southern Louisiana, Mississippi, Alabama, and Common Name: Lemon bacopa, blue waterhyssop in a handful of counties in Georgia and North Carolina (USDA NRCS 2015).
    [Show full text]
  • Westminsterresearch the Botany and Macroscopy of Chinese Materia Medica: Sources, Substitutes and Sustainability Leon, C
    WestminsterResearch http://www.westminster.ac.uk/westminsterresearch The botany and macroscopy of chinese materia medica: sources, substitutes and sustainability Leon, C. This is an electronic version of a PhD thesis awarded by the University of Westminster. © Ms Christine Leon, 2017. The WestminsterResearch online digital archive at the University of Westminster aims to make the research output of the University available to a wider audience. Copyright and Moral Rights remain with the authors and/or copyright owners. Whilst further distribution of specific materials from within this archive is forbidden, you may freely distribute the URL of WestminsterResearch: ((http://westminsterresearch.wmin.ac.uk/). In case of abuse or copyright appearing without permission e-mail [email protected] THE BOTANY AND MACROSCOPY OF CHINESE MATERIA MEDICA: SOURCES, SUBSTITUTES AND SUSTAINABILITY CHRISTINE JUNE LEON A thesis submitted in partial fulfilment of the requirements of the University of Westminster for the degree of Doctor of Philosophy by Published Work April 2017 ABSTRACT Interest in Traditional Chinese Medicine (TCM) is global. The burgeoning international trade in its crude and processed plant ingredients (Chinese materia medica - CMM) reflects demand across all sectors of healthcare, yet the identification of source plants and CMM has been overlooked for many years leading to problems in safety, quality, efficacy and sustainable sourcing. The Guide (Chinese medicinal plants, herbal drugs and substitutes: an identification guide, Leon & Lin, Kew Publishing, 2017), which forms the core of this dissertation by publication, presents a fresh approach to the identification of 226 internationally traded CMM (officially recognised in the Chinese Pharmacopoeia, CP2015) along with their 302 official source plants.
    [Show full text]