Philippine Journal of Science 142: 5-49, Special Issue ISSN 0031 - 7683 Date Received: 2 May 2013

A Verification of Reports of Marine Algal from the

Put O. Ang, Jr., Sin Man Leung, and Mei Mei Choi

Marine Science Laboratory School of Life Sciences, Chinese University of Hong Kong Shatin, N.T., Hong Kong SAR, CHINA

Records of marine macroalgae reported from the Philippines were checked against AlgaeBase, the international database for algal nomenclatures, and Index Nominum Algarum (INA) Bibliographia Phycologica Universalis of the University of California at Berkeley Silva Center for Phycological Documentation to verify their present nomenclature, status of and bibliographic reference. To date, 306 names of taxa (including species, varieties and forms) of greens (), 234 names of taxa of browns (, Phaeophyceae) and 751 names of taxa of reds (Rhodophyta), or a total of 1291 published names of taxa have been reported from the Philippines. Of these, 231 taxa representing 197 species in 20 families for green algae, 171 taxa representing 153 species in 10 families for , and 564 taxa representing 543 species in 52 families for are considered valid records listed with their currently accepted names. All in all, 966 currently accepted taxa, representing 893 species in 82 families of marine macroalgae have been reported from the Philippines. Among the greens, 15 taxa have their type localities in the Philippines. This number is 40 for the browns and 33 for the reds. Proportionally, this is 6.5% of the total for the greens, 23.4% for the browns and 5.9% for the reds.

Key Words: Philippine marine macroalgae, algal taxonomic status, algal type species from the Philippines, AlgaeBase, Index Nominum Algarum (INA)

Introduction et al. (2005) on Padina, Belleza and Liao (2007) on Caulerpa, Geraldino et al. (2010) on Hypnea and Payo Philippines is known for its rich marine flora. Over the et al. (2013) on . years, several efforts have been attempted to compile and annotate the records of marine algae collected from the With the extensive application of molecular techniques Philippines, including those by Silva et al. (1987), the two in studies on algal systematics in recent years, major volume works by Trono (1997, 2004), and more recently, revisions have been made in many groups of tropical the checklist with a cut off date at 1999 by Ganzon-Fortes marine algae. There is therefore a need to update the (2012). The last mentioned literature compiled a list nomenclatures of Philippine marine algal records. This of 949 algal species, including 61 Cyanobacteria, 212 nomenclature validation is important to algal biodiversity, Chlorophytes, 154 Ochrophytes (Phaeophyceaens) and conservation and biogeographical studies in order to 522 Rhodophytes. Several recent works since 1999 have provide a more accurate picture of floristic connection contributed to the long list of Philippine marine algal and the relatedness of Philippine flora with those of records. Notable of these included the works of Geraldino other neighbouring regions. The main objective of this current work is therefore to compile all known records Corresponding Author: [email protected] of Philippine macroalgae from the literature till April of

5 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

2013 and check them against AlgaeBase (Guiry and Guiry 2), and 564 taxa representing 543 species in 52 fami- 2013), the international database of algal nomenclatures, lies for red algae (Table 3). Among the reds, four taxa and Index Nominum Algarum (INA) Bibliographia are generic records only (Euptilota sp., Callithamnion Phycologica Universalis of the University of California sp., Dudresnaya sp. and Rhabdonia sp.). These genera at Berkeley Silva Center for Phycological Documentation are unique records, although with no specific names re- (Silva 1996-to date) to come up with the most updated ported for the Philippine specimens, and are therefore Philippine records. Furthermore, species with their type counted as part of the 543 species recorded. Family locality within the territorial waters of the Philippines are Rhabdoniaceae, where Rhabdonia sp. belongs, also rep- also checked and listed. resents a unique family with only one known species re- corded from the Philippines. The family of Callophycus serratus cannot be ascertained, hence, is classified un- der “Rhodophyta incertae sedis” by AlgaeBase. All in Materials and Methods all, 966 currently accepted valid taxa, representing 893 species in 82 families of marine macroalgae have been This compilation work relied initially on the main cata- reported from the Philippines. logue of marine algae reported from the Philippines by Silva et al. (1987). Further updates were based on works Synonyms or invalid names recorded for different spe- by Trono (1997, 2004) and Ganzon-Fortes (2012). In ad- cies under the three major groups of algae are also listed dition, all relevant, more recent (post 1999) literatures respectively in Tables 1 to 3, across their currently ac- were consulted. In case of doubts, original literatures cepted names. Some invalid names are clearly due to were also consulted for confirmation of the records. spelling error (e.g. siliquosum vs. S. siloquo- These literatures are all included in the Reference list. sum), some are simply due to incorrect gender agreement with the generic name (e.g. Kappaphycus striatus vs. The compiled list of algae was then double checked for K. striatum). These variations of nomenclature in print duplication. The revised list was further checked for are listed here for record purposes. Among the greens, synonymy and invalidity of taxa based on AlgaeBase Enteromorpha aragoensis Bliding is considered illegiti- (Guiry and Guiry 2013) and INA Database (Silva 1996- mate due to lack of a type citation (Table 1). Under each to date). The most recent information on family clas- phylum, there are published records of taxa which have sification, name of authorities, and type localities were no matching records in AlgaeBase. These are separately also checked and accepted to come up with the final list. listed. These taxa have also been checked against INA Where type localities of species are reported to be in the Database. Those which are not also found in INA Da- Philippines, the protologues were consulted, or reference tabase probably represent invalid names. Furthermore, was made to Silva (1996-to date), for verification. some taxa are classified under “Preliminary Entry” in Al- Results and Discussion gaeBase. These are taxa where detailed studies on their taxonomic status are not available. These are all listed This compilation includes only taxa of marine macroal- in the respective Tables for record purposes and are not gae belonging to the three major phyla, the Chlorophyta included in the total count of species. (Green), Ochrophyta (Pheophyceae, Brown) and Rho- dophyta (Red) listed in Tables 1 to 3 respectively. For Among the greens, 15 taxa have their type localities in ease of reference, the scientific names of listed taxa are the Philippines. This number is 40 for the browns and 33 arranged alphabetically first by the family classification, for the reds. Proportionally, this is 6.5% of the total for followed by the generic and species epithets. The abbre- the greens, 23.4% for the browns and 5.9% for the reds. viation of the authority names of all binomials follows The high proportion of type species (taxa) in browns is that used in AlgaeBase. For consideration of nomencla- mainly due to the relatively high number of Sargassum tural priority, the original year of publication was added taxa reported as being new to science by Grunow (1915, at the end of each currently accepted taxon listed. 1916) (16 taxa) and Trono (1994) (12 species). Two taxa (Sargassum cinctum var. jagorii Grunow and S. crassi- In total, 306 published names of taxa (including species, folium var. subduplicatum (‘subduplicata’) Grunow) de- varieties and forms) of greens, 234 names of browns and scribed by Grunow (1915) based on materials from the 751 names of reds, or a total of 1291 names of algal taxa Philippines are not recorded in AlgaeBase (see Table 2). have been reported from the Philippines. Of these, there To include these would increase the proportion of brown are 231 validly published taxa representing 197 species algae with type or syntype locality in the Philippines to in 20 families for green algae (Table 1), 171 taxa repre- 24.3%. On the other hand, the green alga Cladophora senting 153 species in 10 families for brown algae (Table

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Table 1. Listing of green (Chlorophyta) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed. List Family Currently Accepted Name Synonym / Invalid Name Type Locality No. Dalupiri Island, Babuyan Islands, 1 Anadyomenaceae Anadyomene eseptata Gilbert 1961 Cagayan, Luzon, Philippines (Gilbert 1961: 425). Syntype localities: , Min- Microdictyon clathratum G. 2 Anadyomene leclancheri Decaisne 1844 danao, Philippines; Seram and Flores, Martens Indonesia (Martens 1866: 66-67). 3 Anadyomene plicata C. Agardh 1823 Anadyomene stellata (Wulfen) Anadyomene brownii 4 C. Agardh 1823 Schroder Anadyomene wrightii Harvey ex J. E. Gray 5 1866 6 Microdictyon boergesenii Setchell 1925

7 Microdictyon japonicum Setchell 1925

8 Microdictyon okamurae Setchell 1925 Microdictyon agardhianum 9 Microdictyon tenuius J. E. Gray 1866 Decaisne coacta (Dickie) 10 G. Murray & De Toni 1889 11 Boodlea composita (Harvey) F. Brand 1904 Boodlea montagnei (Harvey ex J. E. Gray) 12 Egerod 1952 13 Boodlea struveoide M. A. Howe 1918 Cladophora herpestica (Montagne) zollingeri 14 Kützing 1849 (Kützing) Reinbold Cladophoropsis fasciculata (Kjellman) Cladophoropsis sundanensis 15 Wille 1910 Reinbold Cladophoropsis gracillima 16 E. Y. Dawson 1950 Cladophoropsis membranacea (Hofman 17 Bang ex C. Agardh) Børgesen 1905 Little Santa Cruz Island, opposite Cladophoropsis philippinensis 18 Zamboanga, The Philippines (Taylor W. R. Taylor 1961 1961: 58). Cladophoropsis dichotoma (Zanardini) Papenfuss; Cladophoropsis vaucheriiformis 19 Cladophoropsis neocaledon- (Areschoug) Papenfuss 1958 ica (Grunow ex G. Murray & Boodle) Papenfuss anastomosans anastomosans (Harvey) 20 (Harvey) Piccone & Grunow Kraft & M. J. Wynne 1996 ex Piccone Microdictyon curtissiae W. Phyllodictyon pulcherrimum 21 R. Taylor; J. E. Gray 1866 Struvea ramosa Dickie Chamaedoris orientalis 22 Struvea okamurae Leliaert 2007 Okamura & Higashi 23 Bryopsidaceae Bryopsis corticulans Setchell 1899 Bryopsis indica 24 A. Gepp & E. S. Gepp 1908

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25 Bryopsis pennata J. V. Lamouroux 1809 Bryopsis pennata var. secunda (Harvey) 26 Collins & Hervey 1917 Bryopsis plumosa (Hudson) 27 C. Agardh 1823 Caulerpa arenicola W. R. 28 Caulerpaceae Caulerpa antoensis Yamada 1940 Taylor 29 Caulerpa brachypus Harvey 1860 Caulerpa brachypus f. parvifolia (Harvey) 30 Caulerpa parvifolia Harvey A. B. Cribb 1958 Caulerpa cupressoides (M. Vahl) C. 31 Agardh 1817 Caulerpa cupressoides f. amicorum 32 (Harvey) Weber-van Bosse 1898 Caulerpa cupressoides f. disticha Weber- 33 van Bosse 1898 Caulerpa cupressoides var. Caulerpa cupressoides var. [lycopodium] lycopodium f. elegans (P. L. 34 f. elegans (P. L. Crouan & H. M. Crouan) & H. M. Crouan) Weber-van Weber-van Bosse 1898 Bosse Caulerpa cupressoides var. ericifolia 35 (Turner) Weber-van Bosse 1898 Caulerpa cupressoides var. lycopodium 36 Weber-van Bosse 1898 Caulerpa cupressoides var. urvilleana Caulerpa urvilleana Mon- 37 (Montagne) L. J. Hodgson, Pham Huu Tri, tagne Lewmanomont & McDermid 2004 38 Caulerpa elongata Weber-van Bosse 1898

39 Caulerpa fastigiata Montagne 1837

40 Caulerpa fergusonii G. Murray 1891

41 Caulerpa filicoides Yamada 1936

42 Caulerpa lentillifera J. Agardh 1837 Caulerpa lentillifera var. compacta G. C. Bugsuk, Palawan, Philippines (Trono 43 Trono & Ang 1982 and Ang 1982: 7) Caulerpa lessonii 44 Bory de Saint-Vincent 1828 Caulerpa mexicana Sonder ex Kützing 45 1849 Caulerpa mexicana var. pluriseriata 46 W. R. Taylor 1975 Caulerpa microphysa (Weber-van Bosse) 47 Feldmann 1955 Caulerpa racemosa var. laetevirens (Montagne) Weber-van Bosse; 48 Caulerpa peltata J. V. Lamouroux 1809 Caulerpa racemosa var. peltata (J. V. Lamouroux) Eubank Caulerpa peltata var. macrodisca Caulerpa macrodisca 49 (Decaisne) Weber-van Bosse 1898 Decaisne Caulerpa prolifera (Forsskål) 50 J. V. Lamouroux 1809 Caulerpa racemosa (Forsskål) 51 J. Agardh 1873

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Caulerpa racemosa var. lamourouxii 52 (Turner) Weber-van Bosse 1898 Caulerpa racemosa var. macra Weber-van 53 Bosse 1913 Caulerpa racemosa var. macrophysa 54 (Sonder ex Kützing) W. R. Taylor 1928 Caulerpa racemosa var. occidentalis 55 (J. Agardh) Børgesen 1907 Caulerpa racemosa var. turbinata 56 (J. Agardh) Eubank 1946 Caulerpa reyesii Menez & H. P. 57 Calumpong 1982 58 Caulerpa selago (Turner) C. Agardh 1817 Caulerpa serrulata (Forsskål) 59 J. Agardh 1837 Caulerpa serrulata f. lata 60 (Weber-van Bosse) C. K. Tseng 1936 Caulerpa serrulata f. spiralis 61 (Weber-van Bosse) Gilbert 1942 Caulerpa serrulata var. boryana (J. 62 Agardh) Gilbert 1942 Caulerpa serrulata var. boryana f. longifolia 63 Gilbert 1942 Caulerpa serrulata var. boryana f. oc- 64 cidentalis (Weber-van Bosse) Yamada & Tanaka 1938 Caulerpa serrulata var. pectinata 65 (Weber-van Bosse) W. R. Taylor 1960 Caulerpa sertularioides (S. G. Gmelin) 66 M. A. Howe 1905 Caulerpa sertularioides f. brevipes 67 (J. Agradh) Svedelius 1906 Caulerpa sertularioides f. farlowii 68 (Weber-van Bosse) Børgesen 1907 Caulerpa sertularioides f. longipes 69 (J. Agardh) Collins 1909 Caulerpa sertularioides f. longiseta 70 (Bory de Saint-Vincent) Svedelius 1906 71 Caulerpa subserrata Okamura 1897 Caulerpa taxifolia (M. Vahl) 72 C. Agardh 1817 Caulerpa urvilleana var. vitiensis 73 Weber-van Bosse 1898 74 Caulerpa verticillata J. Agardh 1847 Caulerpa vesiculifera (Harvey) 75 Harvey 1863 76 Caulerpa webbiana Montagne 1837 Caulerpa webbiana var. pickeringii (Har- 77 vey & Bailey) Eubank 1946 Caulerpella ambigua (Okamura) 78 Caulerpa ambigua Okamura Prud'homme van Reine & Lokhorst 1992 Chaetomorpha aerea 79 Cladophoraceae (Dillwyn) Kützing 1849

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Chaetomorpha antennina 80 (Bory de Saint-Vincent) Kützing 1847 81 Chaetomorpha brachygona Harvey 1858 Chaetomorpha crassa 82 (C. Agardh) Kützing 1845 83 Chaetomorpha gracilis Kützing 1845

84 Chaetomorpha inflata Kützing 1849 Panay Island, Philippines (Howe 85 Chaetomorpha kellersii M. A. Howe 1932 1932: 169) Chaetomorpha ligustica (Kützing) 86 Kützing 1849 Chaetomorpha linum (O. F. Müller) 87 Kützing 1845 Dapdap, Siquijor, Chaetomorpha philippinensis 88 Philippines (Leliaert et Leliaert 2011 al. 2011: 385) 89 Chaetomorpha spiralis Okamura 1903 Chaetomorpha vieillardii 90 (Kützing) M. J. Wynne 2011 91 Cladophora albida (Nees) Kützing 1843

92 Cladophora aokii Yamada 1925 Cladophora catenata (Linnaeus) 93 Kützing 1843 Cladophora conferta P. L. Crouan & H. M. 94 Crouan 1865 95 Cladophora crispula Vickers 1905

96 Cladophora crucigera Grunow 1867

97 Cladophora cymopoliae Børgesen 1925

98 Cladophora dalmatica Kützing 1843

99 Cladophora japonica Yamada 1931 Cladophora laetevirens (Dillwyn) Kützing 100 1843 101 Cladophora liebetruthii Grunow 1884 Cladophora pellucida (Hudson) Kützing 102 1843 103 Cladophora prolifera (Roth) Kützing 1843 Batan Island, Batanes, Luzon, Philip- 104 Cladophora quisumbingii Manza 1939 pines (Manza 1939: 109) Cladophora rupestris (Linnaeus) Kützing 105 1843 106 Cladophora sakaii I. A. Abbott 1972 Cladophora sericea (Hudson) Kützing Cladophora crystallina 107 1843 (Roth) Kützing 108 Cladophora sibogae Reinbold 1905 Cladophora vagabunda (Linnaeus) 109 Cladophora inserta Dickie Hoek 1963 110 Cladophora wrightiana Harvey 1860

111 Rhizoclonium africanum Kützing 1853

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Rhizoclonium crassipellitum var. robustum 112 G. S. West 1904 113 Rhizoclonium grande Børgesen 1935 Rhizoclonium kerneri Stock- 114 Rhizoclonium riparium (Roth) Harvey 1849 mayer Rhizoclonium riparium var. implexum Rhizoclonium kochianum 115 (Dillwyn) Rosenvinge 1893 Kützing 116 Rhizoclonium setaceum Kützing 1847

117 arabicum Kützing 1856 Puerto Galera Bay, Mindoro Oriental, Codium bartlettii C. K. Tseng & W. J. 118 Philippines (Tseng and Gilbert 1942: Gilbert 1942 291-293). 119 Codium contractum Kjellman 1857

120 Codium cylindricum Holmes 1896 Codium decorticatum (Woodward) M. A. Codium elongatum Aucto- 121 Howe 1911 rum 122 Codium edule P. C. Silva 1952

123 Codium fragile (Suringar) Hariot 1889 Codium geppii O. C. 124 Codium geppiorum O. C. Schmidt 1923 Schmidt 125 Codium intricatum Okamura 1913

126 Codium muelleri Kützing 1856

127 Codium ovale Zanardini 1878 Puerto Galera, Mindoro Oriental, Codium papillatum C. K. Tseng & W. J. 128 Philippines (Tseng and Gilbert 1942: Gilbert 1942 293-295). 129 Codium platyclados R. Jones & Kraft 1984

130 Codium pungniforme Okamura 1915

131 Codium tenue (Kützing) Kützing 1856

132 Codium tomentosum Stackhouse 1797 Bornetella nitida Munier-Chalmas ex 133 Dasycladaceae Sonder 1880 134 Bornetella oligospora Solms-Laubach 1892 Bornetella sphaerica (Zanardini) 135 Bornetella ovalis Yamada Solms-Laubach 1892 Chloroclados philippinensis Gilbert; Chlorocladus australasicus 136 Chloroclados australasicus Sonder 1871 Sonder; Chlorocladus philippinensis Gilbert Cymopolia van-bosseae Solms-Laubach Cymopolia vanbosseae 137 1892 Solms-Laubach Dasycladus vermicularis (Scopoli) Krasser 138 1898 139 Neomeris annulata Dickie 1874

140 Neomeris bilimbata Koster 1937

141 Neomeris vanbosseae M. A. Howe 1909

142 Derbesiaceae Derbesia attenuata Funk 1955

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143 Derbesia marina (Lyngbye) Solier 1846 Derbesia tenuissima (Moris & De Notaris) 144 P. L. Crouan & H. M. Crouan 1867 Pedobesia ryukyuensis (Yamada & T. 145 Tanaka) Kobara & Chihara 1984 Dichotomosiphona- Avrainvillea erecta (Berkeley) Philippine Islands (Berkeley 1842: 146 ceae A. Gepp & E. S. Gepp 1911 157). Avrainvillea lacerata Harvey ex 147 J. Agardh 1887 Avrainvillea longicaulis (Kützing) 148 G. Murray & Boodle 1889 149 Avrainvillea nigricans Decaisne 1842 Avrainvillea obscura (C. Agardh) Avrainvillea capituliformis 150 J. Agardh 1887 Tanaka Gayralia oxysperma (Kützing) Monostroma oxyspermum 151 Gayraliaceae K. L. Vinogradova ex Scagel 1989 (Kützing) Doty 152 Gomontiaceae Monostroma latissimum Wittrock 1866

153 Monostroma nitidum Wittrock 1866 Porphyra crispata Kjellman Halimeda copiosa 154 Halimedaceae Goreau & E. A. Graham 1967 155 Halimeda cuneata Hering 1846

156 Halimeda cylindracea Decaisne 1842

157 Halimeda discoidea Decaisne 1842

158 Halimeda fragilis W. R. Taylor 1950

159 Halimeda gigas W. R. Taylor 1950

160 Halimeda goreauii W. R. Taylor 1962 Halimeda gracilis 161 Harvey ex J. Agardh 1887 Halimeda incrassata (J. Ellis) J. V. 162 Lamouroux 1816 163 Halimeda lacunalis W. R. Taylor 1950

164 Halimeda macroloba Decaisne 1841

165 Halimeda macrophysa Askenasy 1888

166 Halimeda micronesica Yamada 1941 Halimeda orientalis Gilbert Halimeda opuntia (Linnaeus) 167 J. V. Lamouroux 1816 Halimeda batanensis W. R. 168 Halimeda renschii Hauck 1886 Taylor 169 Halimeda simulans M. A. Howe 1907

170 Halimeda stuposa W. R. Taylor 1950

171 Halimeda taenicola W. R. Taylor 1950 Halimeda tuna (J. Ellis & Solander) 172 J. V. Lamouroux 1816 Sta. Ana, Cagayan, Luzon Island, 173 Halimeda velasquezii W. R. Taylor 1962 Philippines (Taylor 1962: 177). Cadadalman, Camiguin Island, Halimeda velasquezii var. robusta P. A. 174 Cagayan, Philippines (Cordero 1974: Cordero 1974 115).

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Ostreobium quekettii Bornet & Flahault 175 Ostreobiaceae 1889 Phaeophila dendroides (P. L. Crouan & H. 176 Phaeophilaceae M. Crouan) Batters 1902 caliculus J. V. Lamouroux 177 1824 Acetabularia crenulata J. V. Lamouroux 178 1816 179 Acetabularia dentata Solms-Laubach 1895

180 Acetabularia major G. Martens 1866 Acetabularia philippinensis Gilbert Acetabularia roxasii G. C. Acetabularia ryukyuensis Okamura & Trono, Santiago & Ganzon- 181 Yamada 1932 Fortes Acetabularia ryukyuensis var. philippinensis (Gilbert) Valet 182 Halicoryne wrightii Harvey 1860 Parvocaulis clavatus (Yamada) S. Berger, U. Fettweiss, S. Acetabularia clavata 183 Gleissberg, L. B. Liddle, U. Yamada Richter, H. Sawitzky & G. C. Zuccarello 2003 Acetabularia minutissima Okamura; Acetabularia moebii Solms- Parvocaulis parvulus (Solms-Laubach) S. Laubach; Berger, U. Fettweiss, S. Gleissberg, L. B. 184 Acetabularia parvula Solms- Liddle, U. Richter, H. Sawitzky & G. C. Laubach; Zuccarello 2003 Acetabularia velasquezii G. C. Trono, Santiago & Ganzon-Fortes Parvocaulis polyphysoides (P. L. Crouan & H. M. Crouan) S. Berger, U. Fettweiss, Acetabularia exigua Solms- 185 S. Gleissberg, L. B. Liddle, U. Richter, H. Laubach Sawitzky & G.C. Zuccarello 2003 186 Rhipiliaceae Rhipilia crassa Millar & Kraft 2001 Rhipilia orientalis A. Gepp & E. S. Gepp 187 1911 188 Rhipilia tomentosa Kützing 1858 Bulusan, Sorsogon, Philippines (Kraft 189 Rhipiliopsis carolyniae Kraft 1986 1986: 51) Boergesenia forbesii (Harvey) Feldmann 190 1938 Dictyosphaeria cavernosa (Forsskål) 191 Børgesen 1932 Dictyosphaeria cavernosa var. bullata 192 Børgesen 1952 Dictyosphaeria intermedia Weber-van 193 Bosse 1905 Dictyosphaeria ocellata (M. A. Howe) 194 Olsen-Stojkovich 1985 Dictyosphaeria versluysii Weber-van Bosse 195 1905 Ernodesmis verticillata (Kützing) Børgesen 196 1912

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Boodleopsis pusilla (F. S. Collins) W. R. 197 Udoteaceae Taylor, A. B. Joly & Bernatowicz 1953 198 Boodleopsis verticillata E. Y. Dawson 1960

199 Chlorodesmis caespitosa J. Agardh 1887 Chlorodesmis fastigiata (C. Agardh) S. C. Chlorodesmis comosa Har- 200 Ducker 1969 vey & Bailey Chlorodesmis hildebrandtii A. Gepp & E. 201 S. Gepp 1911 202 Chlorodesmis major Zanardini 1874 Udotea javensis (Montagne) 203 Rhipidosiphon javensis Montagne 1842 A. Gepp & E. S. Gepp 204 Tydemania expeditionis Weber-van Bosse 1901

205 Udotea argentea Zanardini 1858 Syntype localities: Tanah Djampeah Udotea argentea var. spumosa A. Gepp & and Salayer Islands, Indonesia; Pearl 206 E. S. Gepp 1911 Bank, Tawitawi, Sulu Archipelago, Philippines (Coppejans et al. 2001: 429) Udotea flabellum (J. Ellis & Solander) M. 207 A. Howe 1904 208 Udotea geppiorum Yamada 1930 Udotea geppii Yamada

209 Udotea glaucescens Harvey ex J. Agardh 1887

210 Udotea indica A. Gepp & E. S. Gepp 1911 Udotea occidentalis A. Gepp & E. S. Gepp 211 1911 Syntype localities: various in Indian Udotea orientalis A. Gepp & E. S. Gepp and Pacific Oceans; Indonesia; Philip- 212 1911 pine Islands (Gepp and Gepp 1911: 119-120). 213 Ulvaceae Ulva australis Areschoug 1854 Ulva pertusa Kjellman Ulva chaetomorphoides (Børgesen) Enteromorpha chaetomor- 214 Hayden, Blomster, Maggs, P. C. Silva, M. phoides Børgesen J. Stanhope & J. R. Waaland 2003 Enteromorpha clathrata (Roth) Greville; 215 Ulva clathrata (Roth) C. Agardh 1811 Enteromorpha ramulosa (Smith) Carmichael Enteromorpha compressa 216 Ulva compressa Linnaeus 1753 (Linnaeus) Nees Enteromorpha flexuosa (Wulfen) J. Agardh; Enteromorpha lingulata J. 217 Ulva flexuosa Wulfen 1803 Agardh; Enteromorpha tubulosa (Kützing) Kützing Ulva flexuosa f. submarina (F. S. Collins & Enteromorpha flexuosa f. 218 Hervey) M. J. Wynne 2005 submarina Collins & Hervey Enteromorpha flexuosa Ulva flexuosa subsp. paradoxa (C.Agardh) 219 subsp. paradoxa (C. Agardh) M. J. Wynne 2005 Bliding Enteromorpha intestinalis 220 Ulva intestinalis Linnaeus 1753 (Linnaeus) Nees Ulva kylinii (Bliding) Hayden, Blomster, 221 Maggs, P. C. Silva, M. J. Stanhope & J. Enteromorpha kylinii Bliding R.Waaland 2003

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222 Ulva lactuca Linnaeus 1753 Ulva fasciata Delile Enteromorpha prolifera (O. 223 Ulva prolifera O. F. Müller 1778 F. Müller) J. Agardh 224 Ulva reticulata Forsskål 1775

225 Ulva rigida C. Agardh 1823

226 Valoniaceae aegagropila C. Agardh 1823

227 Valonia fastigiata Harvey ex J. Agardh 1887

228 Valonia macrophysa Kützing 1843

229 Valonia utricularis (Roth) C. Agardh 1823 Ventricaria ventricosa (J. 230 J. Agardh 1887 Agardh) Olsen & J. A. West pachynema (G. Martens) Valonia confervoides Za- 231 Børgesen 1934 nardini Invalid Species Name

Family Taxon Recorded Remarks Reference *Enteromopha has now been included under Ulva. This Enteromorpha aragoensis Bliding 1960, Silva et al. 1987; Ulvaceae species is currently consid- nom. illeg. Silva 1996-to date ered illegitimate because no type material was designated. Species with No Matching Record in AlgaeBase Family Taxon Recorded Remarks Reference Caulerpaceae Caulerpa cupressoides var. cupressoides Not listed also in INA Belleza and Liao 2007 Database Same as Caulerpa cupres- Caulerpa cupressoides var. lycopodium f. soides f. disticha Weber-van Belleza and Liao 2007 disticha Weber-van Bosse Bosse 1898 Not listed also in INA Caulerpa mexicana var. mexicana Belleza and Liao 2007 Database Listed in INA Database as Caulerpa racemosa var. racemosa Caulerpa racemosa var. (Forsskål) J. Agardh racemosa J. Agardh Caulerpa racemosa var. serrulata (Fors- Not listed also in INA skål) J. Agardh Database Same as Caulerpa serrulata Caulerpa serrulata var. serrulata f. lata f. lata (Weber-van Bosse) C. Belleza and Liao 2007 (Weber-van Bosse) C. K. Tseng K. Tseng Same as Caulerpa sertulari- Caulerpa sertularioides var. sertularioides oides f. brevipes (J. Agardh) Belleza and Liao 2007 f. brevipes (J. Agardh) Svedelius Svedelius Cladophoraceae Cladophora filipendula No authority of taxon listed; Ganzon-Fortes 2012 Not listed also in INA Database Species Listed as Preliminary Entry in AlgaeBase

Family Taxon Recorded Remarks Reference Caulerpaceae Caulerpa sertularioides f. flagellata Listed in INA Database Silva et al. 1987 (Weber-van Bosse) Weber-van Bosse Cladophoraceae Cladophora luzoniensis V. Martens Listed in INA Database; Guiry and Guiry 2013; Type locality: San Mateo Silva 1996-to date River, Luzon, Philippines (Martens 1866: 20) Halimedaceae Halimeda bikinensis W. R. Taylor Listed in INA Database Ganzon-Fortes 2012

15 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

Table 2. Listing of brown (Ochrophyta, Phaeophyceae) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed. List Family Currently Accepted Name Synonym / Invalid Name Type Locality No. Asteronema breviarticulatum (J. Hincksia breviarticulata (J. Agardh) 1 Acinetosporaceae Agardh) Ouriques & Bouzon 2000 P. C. Silva 2 Ectocarpus rallsiae Vickers 1905 Hincksia rallsiae (Vickers) P. C. Silva Feldmannia columellaris (Børgesen) 3 Islam 1976 Feldmannia formosana (Yamada) 4 Itono 1973 Feldmannia indica (Sonder) Womers- 5 ley & A. Bailey 1970 Feldmannia irregularis (Kützing) G. 6 Hamel 1939 Feldmannia mitchelliae (Harvey) H. Hincksia mitchelliae (Harvey) P. C. 7 S. Kim 2010 Silva Leathesia marina (Lyngbye) Decaisne 8 Chordariaceae Leathesia difformis Areschoug 1842 Mutimo cylindricus (Okamura) H. 9 Cutleriaceae Cutleria cylindrica Okamura Kawai & T. Kitayama 2012 Canistrocarpus cervicornis (Kützing) cervicornis Kützing; 10 De Paula & De Clerck 2006 Dictyota indica Sonder ex Kützing San Pio Quinto, Camiguin Island, Ba- Dictyopteris camiguinensis Tanaka 11 buyan Islands, Cagayan, Luzon, The 1967 Philippines (Tanaka 1967: 16) Dictyopteris delicatula J. V. Lamour- 12 oux 1809 Dictyopteris divaricata (Okamura) 13 Okamura 1932 Dictyopteris jamaicensis W. R. Taylor 14 1960 Dictyopteris pacifica (Yendo) I. K. 15 Spatoglossum pacificum Yendo Hwang, H. S. Kim & W. J. Lee 2004 Dictyopteris polypodioides (A. P. De 16 Candolle) J. V. Lamouroux 1809 Dictyopteris repens (Okamura) Børge- 17 sen 1924 18 Dictyopteris undulata Holmes 1896 Dictyota bartayresiana J. V. Lamour- 19 Dictyota patens J. Agardh oux 1809 20 Dictyota ceylanica Kützing 1859 Dictyota ciliolata Sonder ex Kützing 21 1859 Dictyota bartayresiana var. denticulata 22 Dictyota crenulata J. Agardh 1847 Kützing Dictyota crispata J. V. Lamouroux 23 Dictyota bidentata Harvey & Bailey 1809 Dictyota dichotoma (Hudson) J. V. 24 Lamouroux 1809 25 Dictyota friabilis Setchell 1926

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Dictyota implexa (Desfontaines) J. V. Dictyota divaricata J. V. Lamouroux 26 Lamouroux 1809 Dictyota linearis (C. Agardh) Greville 27 Dictyota lata J. V. Lamouroux 1809

28 Dictyota major W. R. Taylor 1945 Dictyota mertensii (Martius) Kützing 29 Dictyota dentata J. V. Lamouroux 1859 Lobophora variegata (J. V. Lamou- Pocockiella variegata (J. V. Lamou- 30 roux) Womersley ex E. C. Oliveira roux) Papenfuss 1977 Padina antillarum (Kützing) Piccone 31 1886 32 Padina arborescens Holmes 1896

33 Padina australis Hauck 1887 Padina boryana Thivy ex W. R. Taylor 34 1966 35 Padina distromatica Hauck 1887 Padina fernandeziana Skottsberg & 36 Levring 1941 Padina fraseri (Greville) Greville 37 1830 Padina gymnospora (Kützing) Sonder 38 Padina crassa Yamada 1871 39 Padina japonica Yamada 1931

40 Padina jonesii Tsuda 1972

41 Padina minor Yamada 1925 Padina moffittiana I. A. Abbott & 42 Huisman 2003 Padina pavonica (Linnaeus) Thivy 43 1960 44 Padina sanctae-crucis Børgesen 1914

45 Padina tetrastromatica Hauck 1887 Rugulopteryx okamurae (E. Y. Daw- 46 son) I. K. Hwang, W. J. Lee & H. S. Dilophus okamurae E. Y. Dawson Kim 2009 47 Spatoglossum asperum J. Agardh 1894 Spatoglossum variabile Figari & De 48 Notaris 1853 flabelliforme Weber-van 49 Bosse 1913 Stypopodium zonale (J. V. Lamouroux) Stypopodium lobatum (C. Agardh) 50 Papenfuss 1940 Kützing 51 Zonaria diesingiana J. Agardh 1841 Zonaria flabellata (Okamura) Papen- 52 fuss 1944 Saccharina latissima (Linnaeus) C. 53 Laminariaceae E. Lane, C. Mayes, Druehl & G. W. Ulva latissima Linnaeus Saunders 2006 Mesospora negrosensis J. A. West & Lalaan, San Jose, Negros Oriental, Phil- 54 Mesosporaceae H. P. Calumpong 1996 ippines (West and Calumpong 1996: 5).

17 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

55 Ralfsiaceae Ralfsia fungiformis (Gunnerus) Setch- ell & N. L. Gardner 1924 Ralfsia verrucosa (Areschoug) Are- 56 schoug 1845 Hormophysa cuneiformis (J. F. Gme- Hormophysa triquetra (C.Agardh) 57 lin) P. C. Silva 1987 Kützing Myagropsis myagroides (Mertens ex 58 Turner) Fensholt 1955 Pulong Bakaw, Calatagan, Batangas, 59 Sargassum abbottiae G. C. Trono 1994 Philippines (Trono 1994: 5). 60 Sargassum agardhianum Farlow 1889 Barrio Pangil, Currimao, Ilocos Norte, 61 Sargassum angii L. M. Liao 1998 Sargassum yamadae G. C. Trono Philippines (Trono 1994: 4). Sargassum binderi Sonder ex J. Agardh; Sargassum biserrula J.Agardh; Sargassum aquifolium (Turner) C. 62 Sargassum crassifolium J. Agardh; Agardh 1820 Sargassum heterocystum (Kützing) Montagne; Sargassum oocyste J. Agardh Sargassum baccularia (Mertens) C. 63 Agardh 1824 Sargassum balingasayense G. C. Barrio Balingasay, Bolinao, Pangas- 64 Trono 1994 inan, Philippines (Trono 1994: 5). Sargassum bataanense G. C. Trono Pulong Bato, Orion, Bataan, Philippines 65 1994 (Trono 1994: 6). Syntype localities: Sunda Strait and Sargassum belangeri Bory de Saint- 66 Java, Indonesia; Philippine Islands Vincent 1834 (Silva et al. 1996: 659). Sargassum binderi var. angustifolium 67 Sargassum cervicorne Greville 1849 Sonder 68 Sargassum cinctum J. Agardh 1848 Syntype localities: Coast of Australia: Sargassum cinctum var. gracilentum Sargassum cinctum var. gracilenta Goodie Isl., Balina, Rockinham; near 69 Grunow 1915 Grunow Manila, Philippines (Grunow 1915: 419). Sargassum cinctum var. mixtum Near Manila, Luzon, Philippines (Grun- 70 Sargassum cinctum var. mixta Grunow Grunow 1915 ow 1915: 419). 71 Sargassum confusum C. Agardh 1824 Sargassum corderoi R. B. Modelo Jr., Polo, New Washington, Aklan, Panay 72 I. Umezaki & L. M. Liao 1998 Is., Philippines (Modelo et al. 1998: 79). 73 Sargassum crispifolium Yamada 1931 Sargassum currimaoense G. C. Trono Pangil, Currimao, Ilocos Norte, Philip- 74 1994 pines (Trono 1994: 6). Sargassum cystocarpum C. Agardh 75 1820 Sargassum cystophyllum Montagne Manila, Luzon, Philippines (Montagne 76 1842 1842: 249). 77 Sargassum distichum Sonder 1845 Sargassum aemulum Sonder Honduras, Puerto Galera, Mindoro 78 Sargassum dotyi G. C. Trono 1994 Oriental, Philippines (Trono 1994: 3). Sargassum droserifolium var. spathu- Sargassum droserifolium var. spathu- Strait Bernardino, Philippines (Grunow 79 latum Grunow 1915 lata Grunow 1915: 411).

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Syntype localities: Philippine Islands; Sargassum duplicatum var. rotunda- Sargassum duplicatum var. rotundata 80 Java, Indonesia; Seychelles (Grunow tum Grunow 1915 Grunow 1915: 392-393). 81 Sargassum esperi C. Agardh 1820 Sargassum feldmannii Pham-Hoang 82 Ho 1967 Sargassum filifolium (C. Agardh) C. 83 Agardh 1824 "in insulis Philippinensibus” (Montagne 84 Sargassum filiforme Montagne 1844 1844: 660) (Silva et al. 1996: 672). Sargassum fluitans (Børgesen) Børge- 85 sen 1914 Sargassum fulvellum (Turner) C. 86 Agardh 1820 87 Sargassum furcatum Kützing 1843 Syntype localities: Mauritius; Philip- Sargassum gaudichaudii Montagne 88 pine Islands (Montagne 1842: 249) 1842 (Silva et al. 1996: 673). Sargassum giganteifolium Yamada 89 1925 90 Sargassum gracile J. Agardh 1848

91 Sargassum gracillimum Reinbold 1913 Sargassum granuliferum C. Agardh 92 1820 Sargassum hemiphyllum (Turner) C. 93 Agardh 1820 Sargassum hombronianum var. ma- Sargassum polycystum var. manilense Cavite, Luzon, Philippines (Grunow 94 nilense (Grunow) Grunow 1916 Grunow 1916: 8) Sargassum horneri (Turner) C. Agardh 95 Sargassum filicinum Harvey 1820 Sargassum berberifolium J. Agardh; Sargassum cristaefolium C. Agardh; Sargassum droserifolium Bory de Saint-Vincent; Sargassum duplicatum Bory de Saint- Sargassum ilicifolium (Turner) C. 96 Vincent; Agardh 1820 Sargassum duplicatum (J. Agardh) J. Agardh; Sargassum sandei Reinbold; Sargassum turbinatifolium C.K.Tseng & Lu Syntype localities: Pulau Tikus, near Sargassum ilicifolium f. benkulense Sargassum ilicifolium f. benkulensis 97 Bengkulu, Sumatra, Indonesia; Philip- Grunow 1915 Grunow pine Islands (Grunow 1915: 407). Sargassum ilicifolium var. compactum 98 (Bory de Saint-Vincent) Grunow 1915 Sargassum ilicifolium var. pseudospi- 99 nulosum Grunow 1915 100 Sargassum kushimotense Yendo 1907 Sargassum latifolium (Turner) C. 101 Agardh 1820 Syntype localities: "in marfi Indico"; 102 Sargassum microcystum J. Agardh 1848 Moluccas, Indonesia; Manila, Philip- pines; (Silva et al. 1996: 685).

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Sargassum microcystum f. luzonense Sargassum microcystum f. luzonensis Near Manila, Luzon, Philippines (Grun- 103 Grunow 1915 Grunow ow 1915: 413) Sargassum myriocystum var. eury- phylla (Grunow) Grunow; Sargassum microcystum var. euryphyl- Sargassum myriocystum var. euryphyl- Ticao Island, Masbate, Philippines 104 lum (Grunow) Grunow 1915 lum (Grunow) Grunow; (Grunow 1915: 440) Sargassum polycystum var. euryphyl- lum Grunow 105 Sargassum microcystum var. microtis Syntype localities: Java, Indonesia; Grunow 1915 , Philippines (Grunow 1915: 414). Sargassum microphyllum C. Agardh 106 1820 107 Sargassum miyabei Yendo 1907 Sargassum natans (Linnaeus) Gaillon 108 1828 109 Sargassum nigrifolium Yendo 1907

110 Sargassum notarisii Zanardini 1858 Big Balatero, Puerto Galera, Mindoro 111 Sargassum ohnoi G. C. Trono 1994 Oriental, Philippines (Trono 1994: 7). Sargassum oligocystum Montagne 112 1845 "in freto Bernardino" (San Bernardino Sargassum oligocystum var. bernar- Strait between Sorsogon, Luzon, and 113 dium Grunow 1915 Northern Samar, Philippines) (Grunow 1915: 386) “Fretum Bernardium, Macassar” (San Sargassum oligocystum var. subflexuo- Sargassum oligocystum var. subflex- 114 Bernardino Strait, Philippines; Makas- sum Grunow 1915 uosa Grunow sar, Indonesia) (Grunow 1915: 385) Sargassum oocyste var. bernardinum Sargassum oocyste var. bernardina "in freto Bernardino" (San Bernardino 115 Grunow 1915 Grunow 1915 Strait, Philippines)(Grunow 1915: 437). Sargassum paniculatum J. Agardh 116 1848 Sargassum parvifolium (Turner) C. 117 Agardh 1820 118 Sargassum patens C. Agardh 1820 Sargassum patens var. schizophyllum 119 (Kützing) Yendo 1905 Sargassum philippinense Grunow San Bernardino Strait, Philippines 120 1916 (Grunow 1916: 166) Sargassum piluliferum (Turner) C. 121 Agardh 1820 Sargassum polyceratium Montagne 122 1837 Sargassum polycystum C. Agardh 123 Sargassum myriocystum J. Agardh 1824 124 Sargassum pteropleuron Grunow 1868 Sargassum pteropleron Grunow

125 Sargassum pulchellum Grunow 1888

126 Sargassum sagamianum Yendo 1907 Sargassum samarense G. C. Trono Borangan, Eastern Samar, Philippines 127 1994 (Trono 1994: 4) Sargassum serratifolium (C. Agardh) 128 C. Agardh 1820

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129 Sargassum siliquosum J. Agardh 1848 Sargassum siloquosum J. Agardh Sargassum siliquosum var. basilani- Sargassum siliquosum var. basilanica Basilan Strait (between and 130 cum Grunow 1916 Grunow Basilan Island) (Grunow 1916: 175) Sargassum siliquosum var. bicornutum Sargassum siliquosum var. bicornuta China Sea; Ticao Island, Masbate, Phil- 131 Grunow 1916 Grunow ippines (Grunow 1916: 175) Sargassum siliquosum var. mani- Sargassum siliquosum var. manipaense paensis Grunow; "ad insulas Philippinas, Manipa" (Grun- 132 Grunow 1916 Spelled in AlgaeBase as Sargassum ow 1916: 175) siliquosum var. manipense Grunow 133 Sargassum spinifex C. Agardh 1820 Sargassum subspathulatum (Grunow) 134 Grunow 1916 Palawig, Zambales, Philippines (Trono 135 Sargassum sullivanii G. C. Trono 1994 1994: 6) 136 Sargassum tenerrimum J. Agardh 1848

137 Sargassum tristichum Sonder 1845 Sargassum biforme Sonder Sargassum turbinarioides Grunow 138 1915 Medio Island, Puerto Galera, Mindoro 139 Sargassum umezakii G. C. Trono 1994 Oriental, Philippines (Trono 1994: 7) Sargassum velasquezii G. C. Trono San Isidro, Puerto Galera, Mindoro 140 1994 Oriental, Philippines (Trono 1994: 4) 141 Sargassum vulgare C. Agardh 1820 Sargassum vulgare var. indicum C. 142 Agardh 1820 Sargassum yendoi Okamura & Ya- 143 mada 1938 Cabugao Bay, Virac, Catanduanes, 144 Sargassum yoshidae G. C. Trono 1994 Philippines (Trono 1994: 5) Sirophysalis trinodis (Forsskål) Kütz- Cystoseira trinodis (Forsskål) 145 ing 1849 C.Agardh Stephanocystis hakodatensis (Yendo) Cystoseira hakodatensis (Yendo) 146 Draisma, Ballesteros, F. Rousseau & Fensholt T. Thibaut 2010 147 Turbinaria condensata Sonder 1860 Turbinaria conoides (J. Agardh) Kütz- 148 ing 1860 Turbinaria conoides f. laticuspidata 149 Cebu, Philippines (Taylor 1964: 481) W. R. Taylor 1964 Turbinaria conoides f. retroflexa W. R. Tandayong Island, Luzon, Philippines 150 Taylor 1964 (Taylor 1964: 481-482) Turbinaria decurrens Bory de Saint- 151 Vincent 1828 Turbinaria denudata Bory de Saint- 152 Vincent 1828 153 Turbinaria filamentosa Yamada 1925 Turbinaria luzonensis W. R. Taylor Mulanay, Luzon, Philippines (Taylor 154 1964 1964: 482). Turbinaria murrayana E. S. Barton 155 1891 Turbinaria ornata (Turner) J. Agardh 156 1848

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Turbinaria ornata f. evesiculosa (E. S. 157 Barton) W. R. Taylor 1964 Turbinaria trialata var. capensis Kütz- 158 ing 1860 Turbinaria turbinata (Linnaeus) 159 Turbinaria trialata (J. Agardh) Kützing Kuntze 1898 Turbinaria vulgaris J. Agardh 1848 160 nom. illeg. 161 Scytosiphonaceae Chnoospora implexa J. Agardh 1848 Chnoospora minima (Hering) Papen- 162 fuss 1956 163 Chnoospora pannosa J. Agardh 1848 Colpomenia sinuosa (Mertens ex 164 Roth) Derbes & Solier 1851 Hydroclathrus clathratus (C. Agardh) 165 M. A. Howe 1920 Hydroclathrus tenuis C. K. Tseng & 166 Lu 1983 Rosenvingea intricata (J. Agardh) 167 Børgesen 1914 Hab. in mari Indico ad Manillam (Hb. Rosenvingea orientalis (J. Agardh) 168 Binder!) [Manila, the Philippines] Børgesen 1914 (Agardh 1848: 78). Sphacelaria novae-hollandiae G. 169 Sphacelariaceae Sonder 1845 170 Sphacelaria rigidula Kützing 1843 Sphacelaria tribuloides Meneghini 171 1840 Species with No Matching Record in AlgaeBase

Family Taxon Recorded Remarks Reference No authority of taxon listed; Sphacelariaceae Sphacelaria mucifera Ganzon-Fortes 2012 Also not listed in INA Database Sargassaceae Sargassum cinctum var. jagorii Type locality: Luzon, Philippines Grunow 1915; Silva 1996-to date Grunow (Grunow 1915: 421) Syntype localities: Makassar, Indo- Sargassum crassifolium var. subdupli- nesia; Strait Bernardino, Philippines Grunow 1915; Silva 1996-to date catum ('subduplicata') Grunow (Grunow 1915: 421) Species Listed as Preliminary Entry in AlgaeBase Family Taxon Recorded Remarks Reference Sargassaceae Sargassum elongatum No authority of taxon listed; Phillips 1995; Ganzon-Fortes 2012 Not listed in INA Database

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Table 3. Listing of red (Rhodophyta) algae reported from the Philippines, with their valid names, authorities and previous record reported under their synonym(s) or invalid name(s) (if available). Type localities of the species, if found in the Philippines, are also listed. List No. Family Currently Accepted Name Synonym / Invalid Name Type Locality Acrochaetium hancockii (E. Y. Daw- 1 Acrochaetiaceae son) Papenfuss 1845 Acrochaetium liagorae Børgesen 2 1915 Balabac Island, Gnat Reef, Pala- Acrochaetium nitidulum I. A. Abbott 3 wan, Philippines (Abbott 1962: 1962 95) Acrochaetium papenfussii I. A. Abbott Cagayan Sulu Island, Palawan, 4 1962 Philippines (Abbott 1962: 97) Acrochaetium robustum Børgesen 5 1915 Acrochaetium sinicola (E. Y. Dawson) 6 Papenfuss 1945 Acrochaetium trichogloeae Børgesen 7 1952 Acrochaetium tuticorinense Børgesen 8 1937 Colaconema gracile (Børgesen) 9 Ateweberhan & Prud'homme van Acrochaetium gracile Børgesen Reine 2005 Colaconema hallandicum (Kylin) Acrochaetium hallandicum (Kylin) 10 Afonso-Carillo, Sanson, Sangil & Hamel Diaz-Villa 2007 Colaconema hypneae (Børgesen) A. 11 Acrochaetium seriatum Børgesen A. Santos & C. W. N. Moura 2010 Ahnfeltiopsis concinna (J.Agardh) P. 12 Ahnfeltiaceae Ahnfeltia concinna J. Agardh C. Silva & DeCew 1992 Bangia atropurpurea (Mertens ex 13 Bangiaceae Roth) C. Agardh 1824 14 Bangia yamadae Tanaka 1944 Porphyra atropurpurea (Olivi) De 15 Toni 1897 Dirique Bay, Burgos, Ilocos Porphyra marcosii P. A. Cordero 16 Norte, Philippines (Cordero 1976 1976: 15) 17 Porphyra umbilicalis Kützing 1843 Ulva umbilicalis Linneaus Pyropia denticulata (Levring) J. A. Porphyra denticulata Levring 18 Philips & J. E. Sutherland 2011 Pyropia suborbiculata (Kjellman) 19 J.E.Sutherland, H. G. Choi, M. S. Porphyra suborbiculata Kjellman Hwang & W. A. Nelson 2011 Pyropia vietnamensis (Tak. Tanaka & Porphyra vietnamensis T. Tanaka & 20 P.H. Ho) J. E. Sutherland & Monotilla P. H. Ho 2011 Wildemania variegata (Kjellman) De Porphyra variegata (Kjellman) 21 Toni 1890 Kjellman Asparagopsis taxiformis (Delile) Falkenbergia hillebrandii (Bornet) 22 Bonnemaisoniaceae Trevisan de Saint-Leon 1845 Falkenberg

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Crouania attenuata (C. Agardh) J. 23 Callithamniaceae Crouania minutissima Yamada Agardh 1842 Catenella caespitosa (Withering) L. 24 Caulacanthaceae M. Irvine 1976 Catenella impudica (Montagne) J. 25 Agardh 1852 26 Catenella nipae Zanardini 1872 Caulacanthus ustulatus (Mertens ex 27 Turner) Kützing 1843 Acrothamnion preissii (Sonder) E. M. 28 Ceramiaceae Wollaston 1968 Antithamnion lherminieri (P. L. Antithamnion antillanum Børgesen 29 Crouan & H. M. Crouan) Bornet 1917 ex Nasr Balliella subcorticata (Itono) Itono & 30 T. Tanaka 1973 Centroceras clavulatum (C. Agardh) 31 Montagne 1846 32 Centroceras minutum Yamada 1944 Ceramium affine Setchell & N. L. 33 Gardner 1930 Ceramium cimbricum H. E. Petersen 34 Ceramium fastigiatum Harvey 1924 Ceramium cruciatum F. S. Collins & 35 Hervey 1917 Ceramium gracillimum (Kützing) Ceramium diaphanum (Lightfoot) Griffiths & Harvey; 36 Roth 1806 Ceramium tenuissimum (Roth) Areschoug Ceramium equisetoides E. Y. Dawson 37 1944 38 Ceramium loureiri C. Agardh 1824 Ceramium luetzelburgii O. C. 39 Schmidt 1924 Ceramium marshallense E. Y. Daw- 40 son 1957 Ceramium maryae Weber-van Bosse 41 1923 Ceramium mazatlanense E. Y. Daw- 42 son 1950 43 Ceramium multijugum Jaasund 1970 Ceramium nitens C. Agardh (J. 44 Agardh) 1851 Ceramium personatum Setchell & N. 45 L. Gardner 1930 Ceramium procumbens Setchell & N. Ceramiella procumbens (Setchell & 46 L. Gardner 1924 N. L. Gardner) Diaz-Piferrer Ceramium serpens Setchell & N. L. 47 Gardner 1924 Ceramium sinicola Setchell & N. L. 48 Gardner 1924 Ceramium tenerrimum (G. Martens) 49 Okamura 1921

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50 Ceramium vagans P. C. Silva 1987 Ceramium zacae Setchell & N. L. 51 Gardner 1937 Corallophila kleiwegii Weber-van 52 Centroceras apiculatum Yamada Bosse 1923 Gayliella flaccida (Harvey ex Kütz- Ceramium flaccidum (Harvey ex 53 ing) T. O. Cho & L. J. McIvor 2008 Kützing) Ardissone Gayliella taylorii (E. Y. Dawson) T. 54 Ceramium taylorii E. Y. Dawson O. Cho & S. M. Boo 2008 Herpochondria elegans (Okamura) 55 Microcladia elegans Okamura Itono 1977 Microcladia glandulosa (Solander ex 56 Turner) Greville 1830 57 Champiaceae Champia bifida Okamura 1901 Champia caespitosa E. Y. Dawson 58 1944 59 Champia compressa Harvey 1838 60 Champia disticha E. Y. Dawson 1944 61 Champia japonica Okamura 1931 Champia parvula (C. Agardh) Harvey 62 1853 63 Champia salicornoides Harvey 1853 Champia spathulata Weber-van 64 Bosse 1928 65 Champia vieillardii Kützing 1866 Coelothrix irregularis (Harvey) 66 Børgesen 1920 Amphiroa anastomosans Weber-van 67 Corallinaceae Bosse 1904 Amphiroa anceps (Lamarck) Decaisne 68 1842 69 Amphiroa annulata M. Lemoine 1929 Amphiroa beauvoisii J. V. Lamouroux 70 Amphiroa zonata Yendo 1816 Amphiroa crassa J. V. Lamouroux 71 1824 72 Amphiroa cumingii Montagne 1844 Amphiroa dimorpha M. Lemoine 73 1929 Amphiroa ephedraea (Lamarck) 74 Decaisne 1842 Amphiroa foliacea J. V. Lamouroux 75 1824 Amphiroa foliacea f. erecta Weber- 76 van Bosse 1904 Amphiroa fragilissima (Linnaeus) J. 77 V. Lamouroux 1816 Amphiroa fragilissima f. cyathifera 78 (J. V. Lamouroux) Weber-van Bosse 1904 Amphiroa hancockii W. R. Taylor 79 1942

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80 Amphiroa pacifica Kützing 1858 Amphiroa rigida J. V. Lamouroux 81 1816 Amphiroa subcylindrica E. Y. Dawson 82 1953 Amphiroa tribulus (J. Ellis & Solan- 83 der) J. V. Lamouroux 1816 84 Amphiroa valonioides Yendo 1902 Arthrocardia palmata (J. Ellis & 85 Solander) Areschoug 1852 Bossiella frondescens (Postels & Corallina frondescens Postels & 86 Ruprecht) E. Y. Dawson 1964 Ruprecht Hydrolithon farinosum (J. V. Lamour- Fosliella farinosa (J. V. Lamouroux) 87 oux) D. Penrose & Y. M. Chamberlain M. A. Howe 1993 Hydrolithon reinboldii (Weber-van 88 Bosse & Foslie) Foslie 1909 Jania acutiloba (Decaisne) J. H. Kim, Cheilosporum jungermannioides 89 Guiry & H. G. Choi 2007 Ruprecht ex Areschoug Jania adhaerens J. V. Lamouroux 90 1816 91 Jania capillacea Harvey 1853 Jania cubensis Montagne ex Kützing Haliptilon cubense (Montagne ex 92 1849 Kützing) Garbary & H. W. Johansen Jania cultrata (Harvey) J. H. Kim, Cheilosporum cultratum (Harvey) 93 Guiry & H. G. Choi 2007 Areschoug Jania decussato-dichotoma (Yendo) 94 Yendo 1905 95 Jania longiarthra E. Y. Dawson 1953 Jania micrarthrodia J. V. Lamouroux 96 1816 97 Jania pacifica Areschoug 1852 98 Jania pumila J. V. Lamouroux 1816 Jania rubens (Linnaeus) J. V. 99 Lamouroux 1816 Jania sagittata (J. V. Lamouroux) Cheilosporum sagittatum (J. V. 100 Blainville 1834 Lamouroux) Areschoug Jania spectabilis (Harvey ex Grunow) Cheilosporum spectabile Harvey ex 101 J. H. Kim, Guiry & H. G. Choi 2007 Grunow 102 Jania tenella (Kützing) Grunow 1874 Jania tenella var. zacae E. Y. Dawson 103 1953 104 Jania ungulata (Yendo) Yendo 1905 Jania ungulata f. brevior (Yendo) 105 Yendo 1905 Lithophyllum byssoides (Lamarck) 106 Foslie 1900 Lithophyllum pallescens (Foslie) 107 Foslie 1900 Lithophyllum pygmaeum (Heydrich) Lithophyllum moluccense (Foslie) 108 Heydrich 1897 Foslie

26 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

Lithoporella indica (Foslie) W. H. 109 Adey 1970 Lithoporella melobesioides (Foslie) 110 Foslie 1909 Mastophora rosea (C. Agardh) 111 Setchell 1943 Mesophyllum simulans (Foslie) Me. 112 Lemoine 1928 Neogoniolithon brassica-florida (Har- Neogoniolithon frutescens (Foslie) 113 vey) Setchell & L. R. Mason 1943 Setchell & L. R. Mason Neogoniolithon megalocystum 114 (Weber-van Bosse & Foslie) Setchell & L. R. Mason 1943 Porolithon onkodes (Heydrich) Foslie 115 1909 Bulusan, Sorsogon, Philippines 116 Corynocystaceae Corynocystis prostrata Kraft 1999 (Kraft et al. 1999: 28). Cubiculosporum koronicarpis Kraft Bulusan, Sorsogon, Philippines. 117 Cubiculosporaceae 1973 (Kraft 1973: 881). 118 Cystocloniaceae Hypnea boergesenii T. Tanaka 1941 Intertidal zone off Panglao Is- Hypnea caespitosa Geraldino & S. M. 119 land, Bohol, Philippines (Ger- Boo 2010 aldino et al. 2010: 338) 120 Hypnea cenomyce J. Agardh 1851 Hypnea charoides J. V. Lamouroux 121 1813 Hypnea cornuta (Kützing) J. Agardh 122 1851 Hypnea divaricata (C. Agardh) 123 Greville 1830 Hypnea musciformis (Wulfen) J. V. 124 Lamouroux 1813 Hypnea musciformis var. esperi J. 125 Agardh 1851 Hypnea musciformis var. hippuroides 126 (Kützing) Weber-van Bosse 1928 127 Hypnea nidulans Setchell 1924 128 Hypnea pannosa J. Agardh 1847 129 Hypnea saidana Holmes 1896 Hypnea spinella (C. Agardh) Kützing 130 Hypnea cervicornis J. Agardh 1847 Lectotype locality: Manila, Phi- Hypnea stellulifera (J. Agardh) Yama- Hypnea cornuta var. stellulifera J. 131 lippines (Yamagishi et al. 2003: gishi & M. Masuda 2003 Agardh 535). Hypnea valentiae (Turner) Montagne 132 1841 Rhodophyllis peltata var. lacunosa 133 Grunow 1874 Dasya adhaerens Yamada; Dasya anastomosans (Weber-van 134 Dasyaceae Eupogodon pilosus (Weber-van Bosse) M. J. Wynne 2002 Bosse) P. C. Silva Dasya antillarum (M. A. Howe) Eupogodon antillarum (M. A. 135 A.J.K .Millar 1996 Howe) P. C. Silva

27 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

Dasya baillouviana (S. G. Gmelin) 136 Montagne 1841 137 Dasya caraibica Børgesen 1919 138 Dasya mollis Harvey 1853 Dasya ocellata (Grateloup) Harvey 139 1833 Dasya punicea (Zanardini) Meneghini 140 ex Zanardini 1841 141 Dasya sessilis Yamada 1928 142 Dasya villosa Harvey 1844 Eupogodon planus (C. Agardh) Kütz- 143 ing 1845 Heterosiphonia crispella (C. Agardh) Heterosiphonia crispella var. laxa 144 M. J. Wynne 1985 (Børgesen) M. J. Wynne Heterosiphonia muelleri (Sonder) De 145 Toni 1903 Tapeinodasya bornetii Weber-van 146 Bosse 1904 Apoglossum spathulatum (Sonder) Hypoglossum spathulatum (Sonder) 147 Womersley & Shepley 1982 Kützing 148 Botryocarpa prolifera Greville 1830 Caloglossa bengalensis (G. Martens) Caloglossa adnata (Zanardini) De 149 R. J. King & Puttock 1994 Toni Caloglossa leprieurii (Montagne) G. 150 Martens 1869 Caloglossa ogasawaraensis Okamura 151 1897 Caloglossa vieillardii (Kützing) Caloglossa leprieurii var. hookeri 152 Setchell 1924 Post Basco, Batan Island, Batanes, 153 Claudea batanensis Tanaka 1967 Luzon, Philippines (Tanaka 1967: 18). 154 Claudea multifida Harvey 1854 Hypoglossum attenuatum N. L. 155 Gardner 1927 Hypoglossum harveyanum (J. Agardh) 156 Hypoglossum serrulatum J. Agardh Womersley & Shepley 1982 Hypoglossum heterocystideum (J. 157 Agardh) J. Agardh 1898 158 australis Harvey 1855 Martensia speciosa Zanardini Martensia flabelliformis Harvey ex J. 159 Agardh 1863 160 Martensia fragilis Harvey 1854 an der küste der Philippinen’ Martensia martensii (F. Schmitz) S. 161 Opephyllum martensii F. Schmitz [Philippines] (Schmitz and M. Lin, Fredericq & L. M. Liao 2001 Hauptfleisch 1897: 410). Taenioma perpusillum (J. Agardh) J. 162 Agardh 1863 163 Vanvoorstia spectabilis Harvey 1854 164 Zellera tawallina G. Martens 1866 Dicranema revolutum (C. Agardh) J. 165 Dicranemataceae Agardh 1852

28 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

166 Dumontiaceae Gibsmithia hawaiiensis Doty 1963 167 Rhodopeltis borealis Yamada 1931 Stenopeltis gracilis (Yamada & Rhodopeltis gracilis Yamada & 168 Tanaka) Itono & Yoshizaki 1992 Tanaka Gloiopeltis complanata (Harvey) 169 Endocladiaceae Yamada 1932 Gloiopeltis tenax (Turner) Decaisne 170 1842 Erythrocladia irregularis Rosenvinge 171 Erythropeltidaceae 1909 Erythrocladia pinnata W. R. Taylor 172 1942 Erythrotrichia bangioides Levring 173 Erythrotrichiaceae 1955 174 Erythrotrichia biseriata Tanaka 1944 175 Erythrotrichia parietalis Tanaka 1952 176 Faucheaceae Gloiocladia ramellifera Hauck 1886 Leptofauchea leptophylla (Segawa) Mas. Suzuki, Nozaki, R. Terada, Kita- 177 Fauchea leptophylla Segawa yama, Tetsu. Hashimoto & Yoshizaki 2012 Actinotrichia fragilis (Forsskål) 178 Galaxauraceae Børgesen 1932 Dichotomaria apiculata (Kjellman) 179 Galaxaura apiculata Kjellman A. Kurihara & Masuda 2005 Dichotomaria falcata (Kjellman) A. 180 Galaxaura falcata Kjellman Kurihara & Masuda 2005 Dichotomaria marginata (J. Ellis & Galaxaura marginata (J. Ellis & 181 Solander) Lamarck 1816 Solander) J. V. Lamouroux Dichotomaria obtusata (J. Ellis & Galaxaura obtusata (J. Ellis & 182 Solander) Lamarck 1816 Solander) J. V. Lamouroux Dichotomaria spathulata (Kjellman) 183 Galaxaura arborea Kjellman A.Kurihara & Huisman 2006 184 Galaxaura contigua Kjellman 1900 Galaxaura divaricata (Linnaeus) 185 Galaxaura fasciculata Kjellman Huisman & R.A.Townsend 1993 Galaxaura filamentosa R. C. Y. Chou 186 1945 Galaxaura kjellmanii Weber-van 187 Bosse 1921 Galaxaura subfruticulosa R. C. Y. Galaxaura rugosa (J. Ellis & Solan- 188 Chou; der) J. V. Lamouroux 1816 Galaxaura subverticillata Kjellman 189 Galaxaura striata Kjellman 1900 Tricleocarpa cylindrica (J. Ellis & Lectotype (Galaxaura fastigata 190 Solander) Huisman & Borowitzka Galaxaura fastigiata Decaisne Decaisne): Manila, Luzon, Phil- 1990 ippines (Huisman 2006: 25). Tricleocarpa fragilis (Linnaeus) Huis- Galaxaura oblongata (J. Ellis & 191 man & R. A. Townsend 1993 Solander) J. V. Lamouroux amansii (J. V. Lamouroux) 192 J. V. Lamouroux 1813 Gelidium capense (S. G. Gmelin) P. 193 C. Silva 1987

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194 Gelidium coulteri Harvey 1853 Gelidium crinale (Hare ex Turner) 195 Gaillon 1828 Gelidium crinale var. perpusillum 196 Piccone & Grunow 1884 Gelidium divaricatum G. Martens 197 1866 198 Gelidium isabelae W. R. Taylor 1945 199 Gelidium kintaroi Yamada 1941 Gelidium pulchellum (Turner) Kütz- 200 ing 1868 Gelidium pusillum (Stackhouse) Le 201 Jolis 1863 Gelidium pusillum var. pacificumW. 202 R. Taylor 1945 Gelidium rigens (C. Agardh) Greville 203 ex Kützing 1849 Ptilophora scalaramosa (Kraft) R. E. Bulusan, Sorsogon, Luzon, Phil- 204 Beckerella scalaramosa Kraft Norris 1987 ippines (Kraft 1976: 85). Gelidiella acerosa (Forsskål) Feld- 205 Gelidiellaceae mann & G. Hamel 1934 Parviphycus adnatus (E. Y. Dawson) 206 Gelidiella adnata E. Y. Dawson B. Santelices 2004 Chondracanthus tenellus (Harvey) 207 Gigartinaceae Gigartina tenella Harvey Hommersand 1993 208 Gracilariaceae Gracilaria arcuata Zanardini 1858 Gracilaria arcuata var. snackeyi 209 Weber-van Bosse 1928 210 Gracilaria blodgettii Harvey 1853 Gracilaria bursa-pastoris (S. G. Gracilaria compressa (C.Agardh) 211 Gmelin) P. C. Silva 1952 Greville Gracilaria crassa Harvey ex 212 Gracilaria canaliculata Sonder 1871 J.Agardh Gracilaria changii (B. M. Xia & I. A. 213 Abbott) I. A. Abbott, J. Zhang & B. M. Xia 1991 Gracilaria coronopifolia J. Agardh 214 1852 215 Gracilaria cylindrica Børgesen 1920 Gracilaria damaecornis J. Agardh 216 1852 Gracilaria debilis (Forsskål) Børge- Polycavernosa debilis (Forsskål) 217 sen 1932 Fredericq & J. N. Norris Gracilaria disticha (J. Agardh) J. 218 Agardh 1852 Gracilaria firma C. F. Chang & B. M. 219 Xia 1976 Gracilaria foliifera (Forsskål) Børge- 220 sen 1932 Gracilaria foliifera f. aeruginosa 221 Børgesen 1938 222 Gracilaria gigas Harvey 1860

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223 Gracilaria incurvata Okamura 1931 Gracilaria isabellana Gurgel, Fred- Gracilaria lacinulata (M. Vahl) M. 224 ericq & J. N. Norris 2004 A. Howe Hali Beach, Parañaque, Manila Gracilaria manilaensis Yamamoto & 225 Bay, Philippines (Yamamoto and G. C. Trono 1994 Trono 1994: 95). Gracilaria minor (Sonder) Durairat- 226 nam 1961 Gracilaria papenfussii I. A. Abbott 227 1983 Gracilaria punctata (Okamura) 228 Yamada 1941 Manila, Luzon, Philippines Gracilaria salicornia (C. Agardh) E. (Dawson 1954: 4; Millar and 229 Y. Dawson 1954 Xia 1997; see also Ganzon-Fortes 2012). Gracilaria spinigera E. Y. Dawson 230 1949 Gracilaria spinulosa (Okamura) C. F. 231 Chang & B. M. Xia 1976 Gracilaria srilankia (C. F. Chang & 232 B. M. Xia) Withell, A. J. K. Millar & Kraft 1994 Matabungkay, Batangas, Luzon Gracilaria sullivanii Yamamoto & G. 233 Island, Philippines (Yamamoto C. Trono 1994 and Trono 1994: 99). Gracilaria tenuistipitata C. F. Chang 234 & B. M. Xia 1976 Gracilaria tenuistipitata var. liui J. 235 Zhang & B. M. Xia 1988 Gracilaria textorii (Suringar) De Toni 236 1895 237 Gracilaria turgida E. Y. Dawson 1949 Gracilaria vanbosseae (I. A. Abbott) Polycavernosa vanbosseae I. A. 238 I. A. Abbott 1991 Abbott Gracilaria venezuelensis W. R. Taylor 239 1942 240 Gracilaria vieillardii P. C. Silva 1987 Gracilariopsis bailiniae J. Zhang & Gracilaria heteroclada J. Zhang & 241 B.M.Xia 1991 B. M. Xia 1988 nom. Illeg. Gracilariopsis heteroclada J. Zhang 242 & B. M. Xia 1991 Gracilariopsis lemaneiformis (Bory Gracilaria lemaneiformis (Bory de 243 de Saint-Vincent) E. Y. Dawson, Saint-Vincent) Greville Acleto & Foldvik 1964 Gracilariopsis longissima (S. G. Gracilaria verrucosa (Hudson) 244 Gmelin) M. Steentoft, L. M. Irvine & Papenfuss W. F. Farnham 1995 Gracilaria edulis (S. G. Gmelin) P. C. Silva; Hydropuntia edulis (S. G. Gmelin) 245 Gracilaria lichenoides Greville; Gurgel & Fredericq 2004 Polycavernosa fastigiata C.F.Chang & B.M.Xia Hydropuntia eucheumatoides (Har- Gracilaria eucheumatoides Harvey; 246 vey) Gurgel & Fredericq 2004 Gracilaria eucheumoides Harvey

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247 Hydropuntia urvillei Montagne 1842 Carpopeltis capitellata (Sonder) F. 248 Halymeniaceae Schmitz 1897 Carpopeltis maillardii (Montagne & 249 Millardet) Chiang 1970 Cryptonemia crenulata (J. Agardh) J. 250 Agardh 1851 Cryptonemia denticulata J. Agardh 251 1851 Cryptonemia seminervis (C. Agardh) Cryptonemia luxurians (C. Agardh) 252 J. Agardh 1846 J. Agardh Halymenia acuminata (Holmes) 253 Grateloupia acuminata Holmes 1896 Okamura Carpopeltis angusta (Harvey) Grateloupia angusta (Okamura) S. Okamura; 254 Kawaguchi & H. W. Wang 2001 Prionitis angusta (Okamura) Okamura Grateloupia articulata (Okamura) S. Carpopeltis articulata (Okamura) 255 Kawaguchi & H. W. Wang 2001 Okamura Grateloupia chiangii S. Kawaguchi & 256 Carpopeltis divaricata Okamura H. W. Wang 2001 Grateloupia conferta (Kützing) Kütz- 257 ing 1867 Prionitis cornea (Okamura) E. Y. 258 Grateloupia cornea Okamura 1913 Dawson Grateloupia crispata (Okamura) Y. P. 259 Carpopeltis crispata Okamura Lee 2008 Grateloupia dichotoma J. Agardh 260 1842 Grateloupia divaricata Okamura 261 1895 Grateloupia doryphora (Montagne) 262 M. A. Howe 1914 Grateloupia filicina (J. V. Lamour- 263 oux) C. Agardh 1822 Grateloupia ramosissima Okamura 264 1913 Grateloupia schmitziana (Okamura) Cryptonemia schmitziana (Oka- 265 S. Kawaguchi & H. W. Wang 2001 mura) Okamura 266 Halymenia dilatata Zanardini 1851 Halymenia durvillei Bory de Saint- 267 Vincent 1828 268 Halymenia elongata C. Agardh 1822 Halymenia floresii (Clemente) C. 269 Agardh 1817 Halymenia formosa Harvey ex Kütz- 270 ing 1866 271 Halymenia harveyana J. Agardh 1892 272 Halymenia maculata J. Agardh 1885 Halymenia microcarpa (Montagne) P. 273 C. Silva 1987 274 Halymenia stipitata I. A. Abbott 1998

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Polyopes affinis (Harvey) S. Kawagu- Carpopeltis affinis (Harvey) Oka- 275 chi & H. W. Wang 2002 mura Polyopes prolifer (Hariot) S. Kawa- Carpopeltis prolifera (Hariot) S. 276 guchi & H. W. Wang 2002 Kawaguchi & Masuda Spongophloea procumbens (Weber- van Bosse) Huisman, De Clerck, Thamnoclonium procumbens Weber- 277 Prud'homme van Reine & Borowitzka van Bosse 2011 Spongophloea treubii (Weber- van Bosse) Huisman, De Clerck, Thamnoclonium treubii Weber-van 278 Prud'homme van Reine & Borowitzka Bosse 2011 Carpopeltis formosana Okamura; Yonagunia formosana (Okamura) S. 279 Prionitis formosana (Okamura) S. Kawaguchi & Masuda 2004 Kawaguchi & Nguyen Lithothamnion australe f. brachiatum 280 Hapalidiaceae Foslie 1904 Lithothamnion australe f. minutulum 281 Foslie 1904 Lithothamnion australe f. ubianum 282 Foslie 1904 Lithothamnion indicum f. subtile 283 Foslie 1907 Mesophyllum erubescens (Foslie) Me. 284 Lemoine 1928 Mesophyllum imbricatum (Dickie) W. 285 H. Adey 1970 Mesophyllum pulchrum (Weber-van 286 Bosse & Foslie) Lemoine 1928 Mesophyllum siamense (Foslie) W. H. 287 Adey 1970 Phymatolithon calcareum (Pallas) W. 288 H. Adey & D. L. McKibbin 1970 Phymatolithon purpureum (P. L. 289 Crouan & H. M. Crouan) Woelkerling & L. M. Irvine 1986 Asteromenia exanimans G. W. Saun- 290 Hymenocladiaceae ders, C. E. Lane, C. W. Schneider & Kraft 2006 Hymenocladia dactyloides (Sonder) J. 291 Gracilaria dactyloides Sonder 1845 Agardh 1870 292 Kallymeniaceae Callophyllis adhaerens Yamada 1932 293 Callophyllis adnata Okamura 1932 Callophyllis fastigiata (J. Agardh) J. 294 Gracilaria fastigiata J. Agardh Agardh 1876 Kallymenia callophylloides Okamura 295 & Segawa 1935 296 Kallymenia pacifica Kylin 1956 297 Kallymenia perforata J. Agardh 1872 Kallymenia rosea Womersley & R. E. 298 Norris 1971 Kallymenia rubra Womersley & R. E. 299 Norris 1971 300 Kallymenia sessilis Okamura 1934

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301 Kallymenia tasmanica Harvey 1860 302 Pugetia japonica Kylin 1941 Callophyllis okamurae P. C. Silva Dermonema virens (J. Agardh) Pedro- Dermonema frappieri (Montagne & 303 Liagoraceae che & Ávila Ortíz 1996 Millardet) Børgesen Ganonema farinosum (J. V. Lamour- 304 Liagora farinosa J. V. Lamouroux oux) K. C. Fan & Yung C. Wang 1974 Helminthocladia australis Harvey 305 1863 Izziella orientalis (J. Agardh) Huis- 306 Liagora orientalis J. Agardh man & Schils 2002 307 Liagora boergesenii Yamada 1938 308 Liagora canariensis Børgesen 1927 Liagora ceranoides J. V. Lamouroux Liagora ceranoides f. leprosa (J. 309 1816 Agardh) Yamada 310 Liagora fragilis Zanardini 1851 311 Liagora hawaiiana Butters 1911 312 Liagora japonica Yamada 1938 313 Liagora robusta Yamada 1938 314 Liagora segawae Yamada 1938 Liagoropsis schrammii (P. L. Crouan 315 & H. M. Crouan) Doty & I. A. Abbott 1964 Neoizziella divaricata (C. K. Tseng) 316 Liagora divaricata C. K. Tseng S. M. Lin, S. Y.Yang & Huisman 2011 Titanophycus setchellii (Yamada) S. 317 Liagora setchellii Yamada M. Lin, S. Y. Yang & Huisman 2011 Titanophycus validus (Harvey) Liagora tenuis J. Agardh; 318 Huisman, G. W. Saunders & A. R. Liagora valida Harvey Sherwood 2006 Trichogloea requienii (Montagne) 319 Kützing 1847 Yamadaella caenomyce (Decaisne) I. Manila, Luzon, Philippines (De- 320 Liagora caenomyce Decaisne A. Abbott 1970 caisne 1842: 119). Ceratodictyon intricatum (C. Agardh) Gelidiopsis intricata (C. Agardh) 321 Lomentariaceae R. E. Norris 1987 Vickers Ceratodictyon repens (Kützing) R. E. Gelidiopsis repens (Kützing) Weber- 322 Norris 1987 van Bosse Ceratodictyon spongiosum Zanardini 323 1878 Ceratodictyon variabile (J.Agardh) R. Gelidiopsis variabilis (Greville ex J. 324 E. Norris 1987 Agardh) F. Schmitz Lomentaria articulata (Hudson) 325 Lyngbye 1819 Lomentaria baileyana (Harvey) 326 Farlow 1876 327 Lomentaria hakodatensis Yendo 1920 328 Lomentaria pinnata Segawa 1938 Palmaria palmata (Linnaeus) Weber 329 Palmariaceae & Mohr 1805 Peyssonnelia conchicola Piccone & 330 Peyssonneliaceae Grunow 1884

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Peyssonnelia distenta (Harvey) 331 Yamada 1930 Pearl Bank, Tawitawi, Sulu Peyssonnelia evae Weber-van Bosse 332 Archipelago, Philippines (Weber- 1921 van Bosse 1921: 279) Peyssonnelia foveolata (Weber-van 333 Bosse) Denizot 1968 Syntype localities: Philippine Peyssonnelia indica (Weber-van Islands; various in Indonesia, 334 Bosse) Denizot 1968 including Semau and Solor (Silva et al. 1996: 212). Peyssonnelia involvens Zanardini 335 1858 Quezon Is., Hundred Is., Pan- Peyssonnelia luzonensis P. A. Cordero 336 gasinan, Philippines (Cordero 1977 1977: 80). Peyssonnelia mariti (Weber-van 337 Bosse) Denizot 1968 Syntype localities: Philippine Peyssonnelia obscura Weber-van Islands; various in Indonesia, 338 Bosse 1921 including Semau and Solor (We- ber-van Bosse 1921: 274-275). Syntype localities: Various lo- Peyssonnelia orientalis (Weber-van Peyssonnelia rubra f. orientalis calities in Indonesia and Philip- 339 Bosse) Cormaci & G. Furnari 1987 Weber-van Bosse pine Islands (Weber-van Bosse 1921: 272). Peyssonnelia rubra (Greville) J. 340 Agardh 1851 Peyssonnelia squamaria (S. G. Gme- 341 lin) Decaisne 1842 342 Polystrata dura Heydrich 1905 Ramicrusta calcea (Heydrich) K. R. 343 Peyssonnelia calcea Heydrich Dixon 2013 Ahnfeltiopsis divaricata (Holmes) 344 Phyllophoraceae Gymnogongrus divaricatus Holmes Masuda 1993 Ahnfeltiopsis flabelliformis (Harvey) Gymnogongrus flabelliformis 345 Masuda 1993 Harvey Ahnfeltiopsis okamurae P. C. Silva & 346 Ahnfeltia furcellata Okamura DeCew 1992 Ahnfeltiopsis pygmaea (J. Agardh) P. 347 Gymnogongrus pygmaeus J. Agardh C. Silva & DeCew 1992 Gymnogongrus dilatatus (Turner) J. 348 Agardh 1851 Phyllophora submaritima E. Y. 349 Dawson 1949 Plocamium cirrhosum (Turner) M. J. Plocamium costatum (C. Agardh) J. 350 Plocamiaceae Wynne 2002 D. Hooker & Harvey Syntype localities: Timor, Indo- nesia; Zamboanga, Mindanao, 351 Plocamium patens G. Martens 1868 Philippines (Silva et al. 1996: 343). Plocamium telfairiae (W. J. Hooker & 352 Harvey) Harvey ex Kützing 1849 Plocamium serrulatum var. pectina- Tajojora, Batan Is. Batanes, 353 tum P. A. Cordero 1977 Philippines (Cordero 1977: 140).

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Pterocladiella caloglossoides (M. A. Pterocladia caloglossoides (M. A. 354 Pterocladiaceae Howe) Santelices 1998 Howe) E. Y. Dawson Pterocladia capillacea (S. G. Gme- Pterocladiella capillacea (S. G. Gme- 355 lin) Bornet; lin) Santelices & Hommersand 1997 Pterocladia densa Okamura Pterocladiella nana (Okamura) Shi- 356 Pterocladia nana Okamura mada, Horiguchi & Masuda 2000 Pterocladiella taylorii (Joly) Santeli- 357 Gelidiella taylori Joly ces 2007 Chondrococcus hornemannii (Lyng- Portieria hornemannii (Lyngbye) P. 358 bye) F. Schmitz C. Silva 1987 Desmia hornemannii Lyngbye Portieria japonica (Harvey) P. C. 359 Silva 1987 Renouxia antillana Fredericq & J. N. 360 Rhodogorgonaceae Norris 1995 361 Rhodomelaceae Acanthophora aokii Okamura 1934 Acanthophora muscoides (Linnaeus) 362 Bory de Saint-Vincent 1828 Acanthophora orientalis J. Agardh; Acanthophora spicifera (M. Vahl) 363 Acanthophora thierryi J. V. Lamour- Børgesen 1910 oux 364 Acrocystis nana Zanardini 1872 365 Alsidium pusillum E. Y. Dawson 1963 Amansia pumila (Sonder) J.Agardh Melanamansia pumila (Sonder) R. 366 1892 Norris Aneurianna lorentzii (Weber-van Lenormandiopsis lorentzii (Weber- 367 Bosse) L. E. Phillips 2006 van Bosse) Papenfuss Bostrychia calliptera (Montagne) 368 Montagne 1842 Bostrychia intricata (Bory de Saint- 369 Vincent) Montagne 1852 370 Bostrychia kelanensis Grunow 1936 Bostrychia moritziana (Sonder ex 371 Kützing) J. Agardh 1863 Bostrychia radicans (Montagne) 372 Montagne 1842 Bostrychia simpliciuscula Harvey ex 373 J. Agardh 1863 Bostrychia tenella (J. V. Lamouroux) 374 Bostrychia binderi Harvey J. Agardh 1863 Chondria armata (Kützing) Okamura 375 1907 376 Chondria crassicaulis Harvey 1860 Chondria curvilineata F. S. Collins & 377 Hervey 1917 Chondria dasyphylla (Woodward) C. 378 Agardh 1817 Chondria polyrhiza F. S. Collins & 379 Hervey 1917 380 Chondria repens Børgesen 1924 381 Chondria sedifolia Harvey 1853

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Chondria seticulosa (Forsskål) C. 382 Agardh 1822 Syntype localities: Borneo Bank Chondria sibogae Weber-van Bosse and Lombok, Indonesia; Sulu 383 1923 Archipelago, Philippines (Silva et al. 1996: 485). Chondrophycus carolinensis (Y. 384 Laurencia carolinensis Y. Saito Saito) K. W. Nam 2007 Chondrophycus cartilagineus (Ya- 385 Laurencia cartilaginea Yamada mada) Garbary & J. T. Harper 1998 Chondrophycus ceylanicus (J. 386 Agardh) M. J. Wynne, Serio, Cormaci Laurencia ceylanica J. Agardh & G. Furnari 2005 Chondrophycus columellaris (Børge- 387 sen) E. Coppejans & A. J. K. Millar Laurencia columellaris Børgesen 2000 Chondrophycus glandulifer (Kützing) Laurencia glandulifera (Kützing) 388 Lipkin & P. C. Silva 2002 Kützing Chondrophycus intermedius (Ya- 389 Laurencia intermedia Yamada mada) Garbary & J. T. Harper 1994 Chondrophycus parvipapillatus 390 (C.K.Tseng) Garbary & J. T. Harper Laurencia parvipapillata C.K. Tseng 1994 North of Pulong Bakaw, Ca- Chondrophycus tronoi (Ganzon- 391 Laurencia tronoi Ganzon-Fortes latagan, Batangas, Philippines Fortes) K. W. Nam 1999 (Ganzon-Fortes 1982: 404). Chondrophycus undulatus (Yamada) 392 Laurencia undulata Yamada Garbary & J. T. Harper 1998 Digenea simplex (Wulfen) C. Agardh 393 1822 Enantiocladia okamurae Yamada 394 1930 Endosiphonia spinulosa (Harvey) 395 Endosiphonia spinuligera Zanardini Womersley & Parsons 2003 Erythrocolon podagricum J. Agardh 396 1874 Syntype localities: Stat. 58. Ile de Savu récif; Stat. 80. Banc de Bornéo 40-50 m; Stat. 99. Ile Ubian du nord, Archipel de Sulu Exophyllum wentii Weber-van Bosse 397 12-33 m [North Ubian Island, 1911 Sulu Archipelago, Philippine Islands; Indonesia: Borneo Bank and Sawu] (Weber-van Bosse 1911: 29). Herposiphonia delicatula Hollenberg 398 1968 Herposiphonia dendroidea Hollen- 399 berg 1968 Herposiphonia dendroidea var. minor 400 Hollenberg 1968 401 Herposiphonia nuda Hollenberg 1968 Herposiphonia obscura Hollenberg 402 1968 Herposiphonia pacifica Hollenberg 403 1968

37 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

404 Herposiphonia parca Setchell 1926 Herposiphonia plumula (J. Agardh) 405 Falkenberg 1901 Herposiphonia secunda (C. Agardh) 406 Ambronn 1880 Herposiphonia secunda f. tenella (C. 407 Agardh) M. J. Wynne 1985 Herposiphonia subdisticha Okamura 408 1899 Herposiphonia tenella (C. Agardh) 409 Ambronn 1880 Herposiphonia trichia Hollenberg 410 1968 Laurencia brongniartii J. Agardh 411 1841 412 Laurencia caraibica P. C. Silva 1972 Laurencia chondrioides Børgesen 413 1918 414 Laurencia clavata Sonder 1853 415 Laurencia composita Yamada 1931 416 Laurencia decumbens Kützing 1863 Laurencia obtusa var. dendroidea (J. 417 Laurencia dendroidea J. Agardh 1852 Agardh) Yamada Laurencia distichophylla J. Agardh 418 1852 419 Laurencia flexilis Setchell 1926 Laurencia forsteri (Mertens ex 420 Turner) Greville 1830 Laurencia intricata J. V. Lamouroux 421 Laurencia implicata J. Agardh 1813 Laurencia majuscula (Harvey) A. H. 422 S. Lucas 1935 Laurencia mariannensis Yamada 423 1931 424 Laurencia nidifica J. Agardh 1852 425 Laurencia nipponica Yamada 1931 Laurencia obtusa (Hudson) J. V. 426 Lamouroux 1813 Laurencia obtusa var. densa Yamada 427 1931 428 Laurencia okamurae Yamada 1931 Laurencia japonica Yamada 429 Laurencia pinnata Yamada 1931 Laurencia similis K. W. Nam & Saito 430 1991 Laurencia snackeyi (Weber-van Laurencia obtusa var. snackeyi 431 Bosse) M. Masuda 1997 (Weber-van Bosse) Yamada Laurencia subsimplex C. K. Tseng 432 1943 433 Laurencia tropica Yamada 1931 434 Laurencia venusta Yamada 1931

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Leveillea jungermannioides (Hering 435 & G. Martens) Harvey 1855 Lophocladia lallemandi (Montagne) 436 F. Schmitz 1893 Lophosiphonia cristata Falkenberg 437 1901 Melanamansia glomerata (C. Agardh) 438 Amansia glomerata C. Agardh R. E. Norris 1995 Murrayella periclados (C. Agardh) F. 439 Schmitz 1893 Neosiphonia apiculata (Hollenberg) 440 Polysiphonia apiculata Hollenberg Masuda & Kogame 2003 Neosiphonia beaudettei (Hollenberg) 441 Polysiphonia beaudettei Hollenberg M. S. Kim & I. A. Abbott 2006 Neosiphonia ferulacea (Suhr ex J. Polysiphonia ferulacea Suhr ex J. 442 Agardh) S. M. Guimarães & M. T. Agardh Fujii 2004 Neosiphonia gorgoniae (Harvey) S. 443 Polysiphonia gorgoniae Harvey M. Guimarães & M. T. Fujii 2004 Neosiphonia hawaiiensis (Hollen- Polysiphonia hawaiiensis Hol- 444 berg) M. S. Kim & I. A. Abbott 2006 lenberg Neosiphonia poko (Hollenberg) I. A. 445 Polysiphonia poko Hollenberg Abbott 2002 Neosiphonia sertularioides (Grate- Neosiphonia flaccidissima (Hollen- 446 loup) K. W. Nam & P. J. Kang 2012 berg) M. S. Kim & I. K. Lee Neosiphonia sphaerocarpa (Børge- Polysiphonia sphaerocarpa Børge- 447 sen) M. S. Kim & I. K. Lee 1999 sen Neosiphonia tepida (Hollenberg) S. 448 Polysiphonia tepida Hollenberg M. Guimarães & M. T. Fujii 2004 Neosiphonia tongatensis (Harvey ex Polysiphonia mollis var. tongatensis 449 Kützing) M. S. Kim & I. K. Lee 1999 (Harvey & Kützing) Hollenberg Neosiphonia upolensis (Grunow) M. 450 Polysiphonia upolensis Grunow S. Kim & Boo 2008 Neurymenia fraxinifolia (Mertens ex 451 Turner) J. Agardh 1863 Osmundaria obtusiloba (C. Agardh) Vidalia obtusiloba (Mertens ex C. 452 R. E. Norris 1991 Agardh) J. Agardh Osmundea pinnatifida (Hudson) Laurencia pinnatifida (Hudson) J. V. 453 Stackhouse 1809 Lamouroux Palisada capituliformis (Yamada) K. 454 Laurencia capituliformis Yamada W. Nam 2007 Palisada corallopsis (Montagne) Laurencia corallopsis (Montagne) 455 Sentíez, M. T. Fujii & Díaz 2008 M. A. Howe Palisada flagellifera (J. Agardh) K. 456 Laurencia flagellifera J. Agardh W. Nam 2007 Palisada parvipapillata (C. K. Tseng) Chondrophycus parvipapillatus (C. 457 K. W. Nam 2007 K. Tseng) Garbary & J. T. Harper Chondrophycus patentiramea (Mon- Palisada patentiramea (Montagne) tagne) Nam; 458 Cassano, Sentíes, Gil-Rodríguez & Laurencia patentiramea (Montagne) M. T. Fujii 2009 Kützing

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Chondrophycus papillosus (C. Agardh) D. J. Garbary & J. T. Palisada perforata (Bory de Saint- 459 Harper; Vincent) K. W. Nam 2007 Laurencia papillosa (C. Agardh) Greville, Setchell & N. L. Gardner 460 Palisada robusta K. W. Nam 2007 Laurencia palisada Yamada Palisada surculigera (C. K. Tseng) K. 461 Laurencia surculigera C. K. Tseng W. Nam 2007 Palisada thuyoides (Kützing) Cas- Laurencia paniculata (C. Agardh) 462 sano, Sentíes, Gil-Rodríguez & M. T. J. Agardh Fujii 2009 Palisada yamadana (M. A. Howe) K. 463 Laurencia yamadana M. A. Howe W. Nam 2007 464 Polysiphonia flabelluta Harvey 1860 465 Polysiphonia forfex Harvey 1859 466 Polysiphonia fragilis Suringar 1867 467 Polysiphonia howei Hollenberg 1945 Polysiphonia japonica var. savatieri 468 Polysiphonia savatieri Hariot (Hariot) Yoon 1986 Polysiphonia mollis J. D. Hooker & 469 Harvey 1847 Polysiphonia pacifica Hollenberg 470 1942 Polysiphonia pacifica var. delicatula 471 Hollenberg 1942 Polysiphonia scopulorum Harvey 472 1855 Polysiphonia scopulorum var. villum 473 (J. Agardh) Hollenberg 1968 Polysiphonia sparsa (Setchell) Hol- 474 lenberg 1968 Polysiphonia subtillisima Montagne 475 1840 476 Polysiphonia triton P. C. Silva 1987 Polysiphonia tsudana Hollenberg 477 1968 Tolypiocladia calodictyon (Harvey ex 478 Kützing) P. C. Silva 1952 Tolypiocladia condensata (Weber-van 479 Bosse) P. C. Silva 1952 Tolypiocladia glomerulata (C. 480 Agardh) F. Schmitz 1897 Womersleyella setacea (Hollenberg) 481 Polysiphonia setacea Hollenberg R. E. Norris 1992 Yuzurua poiteaui (J. V. Lamouroux) Laurencia poiteaui (J. V. Lamour- 482 Martin-Lescanne 2010 oux) M. A. Howe Botryocladia botryoides (Wulfen) 483 Rhodymeniaceae Feldmann 1941 Botryocladia leptopoda (J. Agardh) 484 Kylin 1931 Botryocladia pyriformis (Børgesen) 485 Kylin 1931

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Botryocladia skottsbergii (Børgesen) 486 Levring 1941 Botryocladia uvarioides E. Y. Daw- 487 son 1944 Chrysymenia kaernbachii Grunow 488 Chrysemenia kaernbachii Grunow 1898 489 Rhodymenia californica Kylin 1931 Rhodymenia coacta Okamura & 490 Segawa 1935 Rhodymenia decumbens W. R. Taylor 491 1945 Rhodymenia divaricata E. Y. Dawson 492 1941 Rhodymenia intricata (Okamura) 493 Okamura 1930 Sarcodia montagneana (J. D. Hooker 494 Sarcodiaceae & Harvey) J. Agardh 1852 Cottoniella filamentosa (M. A. Howe) 495 Sarcomeniaceae Børgesen 1920 Titanophora incrustans (J. Agardh) 496 Schizymeniaceae Børgesen 1949 Titanophora mauritiana Børgesen 497 1949 498 Titanophora weberae Børgesen 1943 499 Scinaiaceae Scinaia aborealis Huisman 1986 500 Scinaia hormoides Setchell 1914 501 Scinaia japonica Setchell 1914 502 Scinaia latifrons M. A. Howe 1911 503 Scinaia moniliformis J. Agardh 1885 Scinaia tsinglanensis C. K. Tseng 504 1941 Lesleigha yamadae (Okamura & Sebdenia yamadae Okamura & 505 Sebdeniaceae Segawa) G. W. Saunders & Kraft Segawa 2011 Sebdenia flabellata (J. Agardh) P. G. 506 Parkinson 1980 Sebdenia limensis (Kützing) M. A. 507 Howe 1914 Eucheuma gelatinae (Esper) J. Betaphycus gelatinus (Esper) Doty ex 508 Solieriaceae Agardh; P. C. Silva 1996 Eucheuma gelatinum (Esper) Doty Betaphycus philippinense Doty; Holotype: Eastern Sorsogon, 509 Betaphycus philippinensis Doty 1995 Betaphycus philippinensis var. Philippines (Doty 1995: 237). philippinensis Doty Eucheuma farm, about 1 km Eucheuma alvarezii var. ajakii-assii southwest of the southern tip of 510 Doty 1985 Tumindao Island, Sulu Archipel- ago, Philippines (Doty 1985: 42). Eucheuma amakusaense Okamura 511 1902 Eucheuma arnoldii Weber-van Bosse 512 1928

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Eucheuma arnoldii var. alcyonida Bulusan, Sorsogon, Philippines 513 Kraft 1972 (Kraft 1972: 323). 514 Eucheuma crassum Zanardini 1878 Eucheuma crustiforme Weber-van 515 Bosse 1928 Eucheuma denticulatum (N. L. Bur- Eucheuma muricatum (S.G.Gmelin) 516 man) F. S. Collins & Hervey 1917 Weber-van Bosse Eucheuma denticulatum var. endong Dancalan, Sorsogon, Philippines 517 Trono & Ganzon-Fortes 2012 (Ganzon-Fortes et al. 2012: 1110) Eucheuma edule (Kützing) Weber-van 518 Bosse 1926 519 Eucheuma horridum J. Agardh 1852 Eucheuma isiforme (C. Agardh) J. 520 Agardh 1847 Eucheuma leeuwenii Weber-van 521 Bosse 1928 522 Eucheuma perplexum Doty 1988 Eucheuma serra (J. Agardh) J. 523 Agardh 1847 Eucheuma alvarezii Doty; Kappaphycus alvarezii (Doty) Doty Eucheuma cottonii Weber-van 524 ex P. C. Silva 1996 Bosse; Eucheuma cottonii var. erecta ? farm at Calatagan, Kappaphycus alvarezii var. tamba- Eucheuma alvarezii var. tambalang 525 Batangas, Philippines (Doty lang (Doty) nom. inval. Doty 1985: 41). Kappaphycus cottonii (Weber-van 526 Eucheuma okamurae Yamada Bosse) Doty ex P. C. Silva 1996 Semirara Islands of the northern Kappaphycus procrusteanus (Kraft) Eucheuma procrusteanum Kraft; 527 , Philippines (Kraft Doty 1988 Kappaphycus procrusteanum Kraft 1970: 215). Kappaphycus striatus (F. Schmitz) Eucheuma striatum F. Schmitz; 528 Doty ex P. C. Silva 1996 Kappaphycus striatum F. Schmitz 529 Meristotheca coacta Okamura 1930 Meristotheca papulosa (Montagne) J. 530 Agardh 1872 Sarconema filiforme (Sonder) Kylin Dicranema furcellatum (Montagne) 531 1932 J. Agardh Solieria dura (Zanardini) F. Schmitz 532 1895 Solieria pacifica (Yamada) Yoshida 533 1989 Solieria robusta (Greville) Kylin 534 1932 Wurdemannia miniata (Sprengel) 535 Feldmann & G. Hamel 1934 Sporolithon erythraeum (Rothpletz) 536 Sporolithaceae Kylin 1956 Sporolithon schmidtii (Foslie) G. D. 537 Gordon, Masaki & Akioka 1976 Pearl Bank, Sulu Archipelago, Sporolithon sibogae (Weber-van Archaeolithothamnion sibogae 538 Philippines (Silva et al. 1996: Bosse & Foslie) P. C. Silva 1987 Weber-van Bosse & Foslie 277)

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Sporolithon timorense (Foslie) P. C. 539 Silva 1987 Spyridia filamentosa (Wulfen) Har- 540 Spyridiaceae vey 1833 541 Spyridia velasquezii G. C. Trono 1972 Chroodactylon ornatum (C. Agardh) 542 Stylonemataceae Basson 1979 543 Rhodosorus marinus Geitler 1930 Stylonema alsidii (Zanardini) K. M. Goniotrichum alsidii (Zanardini) M. 544 Drew 1956 A. Howe Anotrichium tenue (C. Agardh) Nageli 545 1862 546 Dasyphila plumarioides Yendo 1920 Gordoniella yonakuniensis (Yamada 547 & T. Tanaka) Itono 1977 548 Griffithsia ovalis Harvey 1855 Griffithsia rhizophora Grunow ex 549 Weber-van Bosse 1923 Griffithsia subcylindrica Okamura 550 1930 elegans (Schousboe 551 ex C. Agardh) J. Agardh 1892 552 Haloplegma duperreyi Montagne 1842 Pleonosporium globuliferum Levring 553 1941 Ptilothamnion cladophorae (Yamada 554 & T. Tanaka) G. Feldmann-Mazoyer 1941 Wrangelia argus (Montagne) Mon- 555 tagne 1856 556 Wrangelia bicuspidata Børgesen 1916 Wrangelia penicillata (C. Agardh) C. 557 Agardh 1828 558 Wrangelia plumosa Harvey 1844 Wrangelia velutina (Sonder) Harvey 559 1855 Rhodophyta incertae Callophycus serratus (Harvey ex 560 sedis Kützing) P. C. Silva 1957 Taxon with Generic Record Only List No. Family Taxon Recorded Remarks Reference 561 Callithamniaceae Callithamnion Lyngbye Kraft et al. 1999 562 Euptilota (Kützing) Kützing Kraft et al. 1999 Dudresnaya P. L. Crouan & H. M. 563 Dumontiaceae Kraft et al. 1999 Crouan 564 Rhabdoniaceae Rhabdonia J. D. Hooker & Harvey Kraft et al. 1999 Species with No Matching Record in AlgaeBase Family Taxon Recorded Remarks Reference Corallinaceae Amphiroa fragilissima f. fragilissima (Linaeus) (Lamoroux) Weber van Also not listed in INA Database Trono 2004 Bosse

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Jania tenella var. tenella E. Y. Also not listed in INA Database Trono 2004 Dawson Silva et al. 1987, Ganzon-Fortes Halymeniaceae Halymenia japonica? Domantay Also not listed in INA Database 2012 Kallymeniaceae Kallymenia denticulata No authority of taxon listed; Ganzon-Fortes 2012 Also not listed in INA Database Species Listed as Preliminary Entry in AlgaeBase Family Taxon Recorded Remarks Reference Corallinaceae Jania constricta Kützing, nom. illeg. Also not listed in INA Database Trono 2004 Peyssonnelia rubra var. orientalis As Peyssonnelia rubra f. orientalis Silva et al. 1987; Peyssonneliaceae Weber-van Bosse Weber-van Bosse in INA Database Silva 1996-to date

luzoniensis Martens was also described based on Phil- National University of Singapore for his comments and ippine materials (Table 1). But this taxon is listed as a editorial suggestions on the manuscript. Special thanks are preliminary entry in AlgaeBase so its current taxonomic also given to Dr. Lawrence Liao of Hiroshima University, status remains unchecked. Japan, for his very valuable comments, corrections and additions on our earlier list of algal records from the Although the number of species (893) recorded from the Philippines. This work benefited from funding from the Philippines compiled in the present work remains com- Research Grant Council of Hong Kong GRF Project No. parable with that compiled by Ganzon-Fortez (2012) for CUHK460012. records up to 1999 (888 species, excluding Cyanobacte- ria), the details of the species list are quite different. This is so mainly because of the large number of taxa that has been synonymized in recent years. On a purely nu- REFERENCES merical sense, there is a net increase of 21 species among the reds in the new list, compared to a drop of 1 and 15 ABBOTT IA. 1962. Some Liagora-inhabiting species species respectively among the browns and the greens. of Acrochaetium. Occasional Papers Bernice Pauahi Payo et al. (2013) recently suggested naming 21 new Bishop Museum 23: 77-120. cryptic species of the red alga Portieria from the Philip- ABBOTT IA. 1985. Gracilaria from the Philippines: List pines based on three molecular markers they used. These and distribution of the species. Taxonomy of Economic “new” species have not been considered in the present 1: 89-90. compilation. ABBOTT IA. 1988. Some species of Gracilaria and As in other attempts to compile a listing of algal records, Polycavernosa from the Siboga expedition. Taxonomy no actual examination of the algal materials was done. of Economic Seaweeds 2: 151-156. This listing should therefore be viewed simply as a listed record with the validity of the published species names ABBOTT IA. 1994. New records and a reassessment verified. It does not, however, verify the correct identifi- of Gracilaria (Rhodophyta) from the Philippines. cation of each original record. Readers which find mis- Taxonomy of Economic Seaweeds 4: 111-118. takes in the list are most welcomed to inform the authors ABBOTT IA. 1999. Notes on some species of Halymenia so an improved list can be made in the future. Further in the southwestern Pacific. Taxonomy of Economic studies involving actual examination of the original algal Seaweeds 7: 163-172. materials are expected to verify the species identity. AGARDH JG. 1848. Species genera et ordines algarum, seu descriptiones succinctae specierum, generum et ordinum, quibus algarum regnum constituitur. Volumen Acknowledgements Primum. Algas fucoideas complectens. Lundae [Lund]: C.W.K. Gleerup. pp. [i-vi], [i]-viii, [1]-363. This paper is dedicated to the memory of Leonardo L. Co, a foremost Philippine plant taxonomist, conservationist AJISAKA T, NORO T, YOSHIDA T. 1995. Zygocarpic and botanical activist whose fruitful life was cut short by Sargassum species (Subgenus Sargassum) from Japan. the wanton gun fire in the forest of Leyte, Philippines Taxonomy of Economic Seaweeds 5: 11-44. on November 15, 2010. We thank Dr. Benito Tan of the ANG PO, TRONO GC. 1987. The genus Sargassum

44 Philippine Journal of Science Ang et al.: A verification of Philippine marine macroalgae Vol. 142: Special Issue

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