Zootaxa 2752: 1–44 (2011) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2011 · Magnolia Press ISSN 1175-5334 (online edition)
A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)
OMAR TORRES-CARVAJAL1,3, RICHARD ETHERIDGE2 & KEVIN DE QUEIROZ3 1Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre y Roca, Apartado 17-01-2184, Quito, Ecuador. E-mail: [email protected] 2Department of Biology, San Diego State University, San Diego, California 92182, USA. E-mail: [email protected] 3Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, MRC 162, Washington, DC 20560, USA. E-mail: [email protected]
Table of contents
Abstract ...... 1 Resumen ...... 2 Introduction ...... 2 Material and methods ...... 6 Results ...... 7 Clade and species accounts ...... 10 Hoplocercinae Alifanov 1996 ...... 10 Enyalioides Boulenger 1885 ...... 11 Enyalioides cofanorum Duellman 1973 ...... 12 Enyalioides heterolepis (Bocourt 1874) ...... 13 Enyalioides laticeps (Guichenot 1855) ...... 19 Enyalioides microlepis (O'Shaughnessy 1881) ...... 22 Enyalioides oshaughnessyi (Boulenger 1881) ...... 23 Enyalioides palpebralis (Boulenger 1883) ...... 25 Enyalioides praestabilis (O'Shaughnessy 1881) ...... 27 Enyalioides rubrigularis Torres-Carvajal et al. 2009 ...... 30 Enyalioides touzeti Torres-Carvajal et al. 2008 ...... 31 Hoplocercus Fitzinger 1843 ...... 32 Hoplocercus spinosus Fitzinger 1843 ...... 32 Morunasaurus Dunn 1933 ...... 33 Morunasaurus annularis (O'Shaughnessy 1881) ...... 33 Morunasaurus groi Dunn 1933 ...... 35 Morunasaurus peruvianus Köhler 2003 ...... 36 Key to the species of Hoplocercinae ...... 37 Acknowledgments ...... 38 References ...... 38 APPENDIX I. Specimens examined...... 42
Abstract
The clade Hoplocercinae includes 13 species assigned to three taxa traditionally ranked as genera—Enyalioides, Hoplo- cercus, and Morunasaurus—, and distributed on both sides of the Andes from Panama to Brazil. We provide a revised taxonomy of Hoplocercinae including synonymies, diagnoses, standardized descriptions, distributions, notes on color and natural history, and a key for species identification. We also propose vernacular names in English and Spanish for all spe- cies.
Key words: Enyalioides, Hoplocercus, Iguanidae, lizards, Morunasaurus, Neotropics, taxonomy
Accepted by S. Carranza: 23 Nov. 2010; published: 31 Jan. 2011 1