A Systematic Revision of Neotropical Lizards in the Clade Hoplocercinae (Squamata: Iguania)

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A Systematic Revision of Neotropical Lizards in the Clade Hoplocercinae (Squamata: Iguania)

Zootaxa 2752: 1–44 (2011) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2011 · Magnolia Press ISSN 1175-5334 (online edition) A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania) OMAR TORRES-CARVAJAL1,3, RICHARD ETHERIDGE2 & KEVIN DE QUEIROZ3 1Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre y Roca, Apartado 17-01-2184, Quito, Ecuador. E-mail: [email protected] 2Department of Biology, San Diego State University, San Diego, California 92182, USA. E-mail: [email protected] 3Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, MRC 162, Washington, DC 20560, USA. E-mail: [email protected] Table of contents Abstract . 1 Resumen . 2 Introduction . 2 Material and methods . 6 Results . 7 Clade and species accounts . 10 Hoplocercinae Alifanov 1996 . 10 Enyalioides Boulenger 1885 . 11 Enyalioides cofanorum Duellman 1973 . 12 Enyalioides heterolepis (Bocourt 1874) . 13 Enyalioides laticeps (Guichenot 1855) . 19 Enyalioides microlepis (O'Shaughnessy 1881) . 22 Enyalioides oshaughnessyi (Boulenger 1881) . 23 Enyalioides palpebralis (Boulenger 1883) . 25 Enyalioides praestabilis (O'Shaughnessy 1881) . 27 Enyalioides rubrigularis Torres-Carvajal et al. 2009 . 30 Enyalioides touzeti Torres-Carvajal et al. 2008 . 31 Hoplocercus Fitzinger 1843 . 32 Hoplocercus spinosus Fitzinger 1843 . 32 Morunasaurus Dunn 1933 . 33 Morunasaurus annularis (O'Shaughnessy 1881) . 33 Morunasaurus groi Dunn 1933 . 35 Morunasaurus peruvianus Köhler 2003 . 36 Key to the species of Hoplocercinae . 37 Acknowledgments . 38 References . 38 APPENDIX I. Specimens examined. 42 Abstract The clade Hoplocercinae includes 13 species assigned to three taxa traditionally ranked as genera—Enyalioides, Hoplo- cercus, and Morunasaurus—, and distributed on both sides of the Andes from Panama to Brazil. We provide a revised taxonomy of Hoplocercinae including synonymies, diagnoses, standardized descriptions, distributions, notes on color and natural history, and a key for species identification. We also propose vernacular names in English and Spanish for all species. Key words: Enyalioides, Hoplocercus, Iguanidae, lizards, Morunasaurus, Neotropics, taxonomy Accepted by S. Carranza: 23 Nov. 2010; published: 31 Jan. 2011 1.

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Checklist of Helminths from Lizards and Amphisbaenians (Reptilia, Squamata) of South America Ticle R A
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