Middle Triassic Ammonoids from Silesia, Poland
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Middle Triassic ammonoids from Silesia, Poland ANDRZEJ KAIM and RoBERT NmoŹwBDZKI Kaim, A. & NiedŹwiedzki, R. 1999. Middle Tiiassic ammonoids from Silesia, Poland. - Acta Palaeontologica Polonica 44, I, 93-115. Adescription of new ammonoid specimens (Acrochordiceras aff.damesi,cf . Acrochordi- ceras sp. indet., DiscopĘchites cf. dux, ?Paraceratites sp., cf. Balatonites sp. indet., Beneckeia buchi) from the Lower Muschelkalk (Anisian, Middle Tńassic) of Silesia (southern Poland) is presented. The detailed stratigraphic position of the new finds is given. The description is supplemented with a list of all species of ammonoids found hitherto in Silesia, which was a southern part of the Germanic Basin during the Anisian. Beneckeia and Noetlingites were typical of epicontinental seas and usually appeared in the early part of fransgressions. Other ammonoids entered from the Tethys into the epicontinental sea of the Germanic Basin during the maxima of fransgressions. Some of them (balatonitids, paraceratitids, bulogitids, Discoptychites and probably Acrochord- iceras) were successful colonizers which established their own populations in the Ger- manic Basin and evolved towards morphologies typical of epicontinental seas. Other (Beyrichites sp., Paraceratites binodosus, and'Trachyceras' sp.) are regarded as un- successful immigrants or empĘ shells drifted post-mortem from the Tethys. Key words: Ammonoids, Triassic, Anisian, Muschelkalk, Silesia, Poland, Acro- chordiceras, DiscopĘchite s, Balatonite s, B enecl<eia,P arac eratites. Andrzej Kaim [kaim@nłarda.pan.pl], InsĘtut Paleobiologii PAN, ul. Twarda 5l/55, P L-00 -8 I 8 Wars zaw a, P oland. Robert Niedźwiedzki I rnied@ ing.uni.wroc.pl ], InsĘtut Nauk Geolo gicznych, Uniw er- sytet Wroctawski, ul. Cybulskiego j0, PL-50-205 Wroctaw, Poland. Introduction Ammonoids are extremely rare in the Polish Lower Muschelkalk (Lower Anisian, Middle Triassic). The Lower Muschelkalk in Poland crops out in Upper and Lower Silesia, as well as in the Holy Cross Mountains (Fig. 1). From the latter area only Beyrichites sp., Beneckeia buchi, and Beneckeia sp. are known (Trammer t97f; Senkowiczowa 1991). More species are known from Silesia, mainly owing to the pre-SecondWorldWarworks of Noetling (1880),HoldefleiB (1915), Rassmuss (1915) for Lower Silesia and of Assmann (1937)for Upper Silesia. More recentĘ a few new 94 Atnmonoids from Silesia; KAIM & MEDŹWIDZKI finds of ammonoidshave been published by Leśniak(1978a) from Lower Silesia and by Podstolski (1980) from Upper Silesia. Dzik (1990) described a new specimen of Acrochordiceras from Strzelce Opolskie in Upper Silesia. He also redescribed and refiguredsome of Noetling's (1880)and Rassmuss'(1915) specimens of ammonoids from Raciborowice (formerly Alt GroB-Hartmannsdorf) in Lower Silesia, which are housed at the Museum ftir Naturkunde of the Humboldt University in Berlin. During field studies on the Silesian Lower Muschelkalk by Kaim (1995, 1997), Chrząstek (|996) and Niedźwiedzkj (1998) additional specimens have been found, with well defined stratigraphicpositions. The purpose of the presentpaper is to describe these new specimens,to review the knowledgeof Lower Muschelkalk ammonoidsin Silesia, and to examine their relations to the ammonoidsof the Tethys. The collection is housed at the Instituteof Paleobiology of the Polish Academy of Sciences,Warsaw (abreviatedzPLL) and at the Geological Museum of the Instituteof Geological Sciences of WrocławUniversity (abreviatedMGUWr). Geological setting The ammonoids described here come from the Upper and Lower Silesia regions in southernPoland. The area of Upper Silesia (Fig. 1) was located during Anisian time directly at the mouth of the Silesian-Moravian Gate connecting the Germanic Basin with the Tethys.According to Urlichs & Mundlos (1985)the areaof Upper Silesia was a place where ammonoids enteredfrom the ocean into the Germanic Basin. Lower Silesia (Fig. 1) is located somewhatto the north of the Sudety Mountains, which were a northernpart of the Vindelician High duńng the Middle Triassic. Upper Silesia. - The Middle Triassic (Muschelkalk) deposits of Upper Silesia are generallyfully marine with the maximum transgressioncoinciding with the Terebratu- la Beds (Fig. 2; Szulc 1993). They are underlain by shallow-water dolomites and evaporitesof the Rót, and overlain by continentaldeposits of the Keuper (Szulc 1993). The lithostratigraphicscheme of the Lower Muschelkalk was establishedby Assmann (1944) and it remains valid to this day. He divided the Lower Muschelkalk into four hthological units underlain by the shallow-waterdolomites of the RÓt (Fig. 2). The fully marine sedimentsof the Gogolin Beds startwith organodetritallimestone composed of crinoid columnals and bivalve shells (so called 'layer with Dadocrinus and Pecterz').The rest of the Gogolin Beds is developed as micritic limestones and marls, commonly deformed in a wavy manner ('Wellenkak' sensu Assmann t944). There are two distinctive marker-bedsin this part of succession:a 'cavemous lime- stone' (see Bodzioch & Kwiatkowski 1992),and an intraformationalbreccia. Fossils are uncommon in the micritic limestones and marls, and are representedmainly by bivalves and trace fossils. In the upper part of the Gogolin Beds nautilids were found and in some places layers of gastropodcoquinas and crinoidal limestone occur. The Górazdze Beds ale composedof threeunits of oncolitic limestoneseparated by beds of micritic limestone.In the oncolitic limestonebivalves, gastropods,brachiopods and cńnoid elements are common. The highest faunal diversity of the whole Lower Muschelkalk is found in thesebeds (Niedźwiedz|<t1998). The micńtic limestonesare ACTA PALAEONTOLOGTCA POLOMCA (44) (1) 95 lffilt"rrigenous marineand continental facies ffi carbonatefacies -t - .r_ areasor non-deposiuon Ę!, -l Tethyan Realm I I o.-J"porit, ero<i,.d Muschelkalk outcrops ffi Upper Triassic ffi Muschelkalk l,owerTiiassic Palaeozoic fault tffil wn ,/ Fig. 1.A. Middle Triassic palaeogeographyof Poland.Modified afterDzik & Trammer(1980). B. Geologi- cal sketch map of the western part of Upper Silesia. Modified after Bodzioch (1997). 96 Ammonoidsfrom Silesra: KAIM & MEDZWIEDZKI Depth curve Strzelce OpolsOpolskie Góra ŚwiętejAnny Górażdże Upper Silesian (Szulc 1993) lithostratigraphy relative ](ARCHOWICE)WICEBEDS (Assmann1944) shallowing <- E'+ p deepening Keuper rr-\..........e\ 'oruszolltllc€ v o Seds v i'..'|'.....' E i, Wlkowice 6 Beds il o il c ..i.{.........' o o ą Tamowice o ą it o ffi f Beds :l ..i..ł.. .'''. .. E = !\ o (E ui< Diplopora = 6 i€ Beds p ż3 c ZPALAm.Xlll.2 o ..;.....\. o MGUWr.5283.s :l CI .E it Beds :l o :\ o :l U) "i""""\" o o Terebratula I.JJ il d) ! Beds 6 ..i......../... v E il I E GórażdŻe f o a Beds :l Ę = t o "t""1""' @ = it IrJ JJ it lr :t uJ F wr II iI EE ot MGUWr.5282.s Gogolin il iI IJJJ Beds it :iE T--r--r rl il = iI '----- / o ZPALAm.Xlll.l (S :-:- Rót I E MGUWr.5281.s (l'= _9 l'' GÓRMDZE BEDS E= marlsand mar|y|imestone tffi#El grainstone Ltl crinoida||imestone FT micńtic|imestone W coquinas and nodularlimestone amronoidfinds ffiwavy f?=- intraformationalbreccia @ Fig. 2. Lithostratigraphy and depth curve of the Upper Silesian Muschelkalk Basin and localisation of the ammonoids in the Terebratula Beds. For correlation with the TeĘan Realm see Fig. 4. poor in fossils and yield only badly preserved,uncommon trace fossils. Wave ripple- marks are commonly preservedand in the uppermostpart hummocky cross stratifica- tion is developed. The TerebratulaBeds (Fig. 2) are tripartite,with the lower part composedof poorly fossiliferous laminated marl or marly limestone.There is a crinoidal limestone in the middle part of the unit, and its upper part consists of nodular.andwavy limestoneand marls with intercalationsof brachiopod and bivalve coquinas (Fig. 2); the reader is referred to Kaim (1997) for a detailed description of the TerebratulaBeds and their benthic assemblages. ACTA PALAEONTOLOGTCA POLOMCA (44) (1) 97 The Karchowice Beds predominantlycomprise crinoidal limestones,coquinas and oncolitic limestones,with subordinatemicritic intercalations.This unit is famous for its sponge-coral-echinodermbioherms (Morycow a I974; B odzioch 1997). The Upper Silesian ammonoidsdescribed herein come from the upper part of the Gogolin Beds, the TerebratulaBeds and the Karchowice Beds of the Strzelce Opolskie Quarry; from the TerebrafulaBeds of the Góra Świętej Anny Quarry and the GóraŻdŻe Quany; and from the Lower Gogolin Beds of Gogolin (Figs 18,2). Lower Silesia. - The first descriptionof the Muschelkalk in Lower Silesia was made by Noetling (1880), who establishedits lithostratigraphicsubdivision and descńbed the palaeontologicalcontent of thę strata.Additionalpalaeontologicaldata and a more detailedlithostratigraphic subdivision were provided by HoldefleiB (1915).Additional data on lithology, sedimentology and palaeontologyfrom outcrops and/or borehole cores were provided by Leśniak(1978a, b), Szulc (1991) and Chrząstek& Niedź- wiedzki (1998).Chrząstek (1996)has provided a microfacies analysis as well as new data on the palaeontology and sedimentology of the Lower Silesian and Sudetian Muschelkalk. The best outcrop of the Lower Silesian Muschelkalk is that of Raciborowice (Fig. 1A). The lithological units in the Raciborowice quarry fit the schemeapplied to the Lower Muschelkalk in Utriper Silesia. Following the correlation of Chrząstek (1996:table 4) we also apply Assmann's (1944)scheme to Lower Silesia. The main differencein comparison with the Upper Silesian Muschelkalk