Verlag Dr. Friedrich Pfeil ISSN 0936-9902 Excerpt from Ichthyological Exploration of Freshwaters An international journal for field-orientated ichthyology Volume 28 Number 2 This article may be used for research, teaching and private purposes. Exchange with other researchers is allowed on request only. Any substantial or systematic reproduction, re-distribution, re-selling in any form to anyone, in particular deposition in a library, institutional or private website, or ftp-site for public access, is expressly forbidden. Ichthyological Exploration of Freshwaters An international journal for field-orientated ichthyology Volume 28 • Number 2 • January 2018 pages 97-192, 40 figs., 14 tabs. Managing Editor Paulo H. F. Lucinda, Laboratório de Ictiologia Sistemática, Universidade Federal do Tocantins, P.O. Box 136, 77500-000 Porto Nacional, TO, Brazil E-mail
[email protected] Honorary Editor Maurice Kottelat, Rue des Rauraques 6, CH-2800 Delémont, Switzerland Associate Editors Ralf Britz, Department of Zoology, The Natural History Museum, London, United Kingdom Kevin W. Conway, Department of Wildlife and Fisheries Sciences, Texas A&M University, College Station, USA Jörg Freyhof, Leibniz-Institute of Freshwater Ecology and Inland Fisheries, Berlin, Germany Sven O. Kullander, Naturhistoriska Riksmuseet, Stockholm, Sweden Heok Hee Ng, Lee Kong Chian Natural History Museum, National University of Singapore, Singapore Editorial Advisory Board Helen K. Larson, Museum and Art Gallery of the Northern Territory, Darwin, Australia Lukas Rüber, Naturhistorisches Museum, Bern, Switzerland Ivan Sazima, Museu de Zoologia, Unicamp, Campinas, Brazil Paul H. Skelton, South African Institute for Aquatic Biodiversity, Grahamstown, South Africa Tan Heok Hui, Lee Kong Chian Natural History Museum, National University of Singapore, Singapore Ichthyological Exploration of Freshwaters is published quarterly Subscriptions should be addressed to the Publisher: Verlag Dr.