The African copper connection Rienk de Jong & Karen van Dorp Apart from their colourful appearance, Copper butterflies are intriguing because of their highly National Museum of Natural History Naturalis disjunct distribution. Almost all species occur in PO Box 9517 the Holarctic region, but a few are found isolated 2300 RA Leiden in the Southern Hemisphere. To learn more about The Netherlands the interrelationship of the southern taxa and
[email protected] their distribution history, an analysis of the phy- logeny of the group was carried out. Although morphological characters are sufficient to diagno- se the species, they proved insufficient to build a phylogeny on. Theref o r e, we had recourse to mo- lecules, using two genes. Here we report on the represented in Africa south of the Sahara by L. phlaeas (s o u t h results with reg a r d to the African rep re s e n t a t i v e s . to Malawi) and two species in South Africa, L. orus Stoll, 1780, and L. clarki Dickson, 1971, and surprisingly in New Zealand Entomologische Berichten 66(4): 124-129 by four species. The other tribe of the subfamily, Heliophorini (Bozano & Weidenhoffer 2001), also has a remarkable distribu- tion, with one genus in the Oriental region, one in Papua New Key words: copper butterflies, Africa, DNA, phylogeny, dis- Guinea and one in Guatemala. To understand these distribu- tribution tions we need to know the phylogeny of the group. In this paper we focus on the question whether the South African taxa Search for the phylogeny of copper butterflies are closely related to the other taxon in Subsaharan Africa, L.